DOCSLIB.ORG

A Phylogeny and Classification of Turtles

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
A Phylogeny and Classification of Turtles A PHYLOGENY AND CLASSIFICATION OF THE HIGHER CATEGORIES OF TURTLES EUGENE S. GAFFNEY BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY VOLUME 155.: ARTICLE 5 NEW YORK : 1975 __ - --- -. -"I-.,'11, ., '_., I,.. II""",.-.,,,.__.;;_:.e.;i:,;, ",.i,,;,,._ I .r....I!. ," _A !I. "' v--..1! -I "I I!,.;* I,. .....-""," Ml;,_. ,4%'. -,",.-,.A-r--'...-;,',,_-, ",,ill;,',:,. .,,,,- I.,- ," .. ., , 1, "_,,1,.,..,.'-"" _.,,... ". .1._.; ,,",".B..`."17!1_1'1,.`,if',1- ..,,w ._ ....,.",--I 11 lel- 4, 1_1.'11, I,;., ._k_;v.--,-,,.SftI_-1. I.,.Z,.-w-% .141 ", "k: ,l., I,. ,-I.,Iiii.Mll..-,.V_-_II,..',,-1 ,"'.."4..",_-",,.,AIIiti., 47.,,.`_.'..-.-i.--.,,'..,,,.,."',-:.', ..,,,'.,",'..,",-.,:",-,-"--.,",,, -.;,,, '.'.!",I..-, - ..,,9I.-, Pr-,.`!,,.,,ll,...*;-,..,.`_._-.iI., i..,,., ..,,-- ,-;_"".'_iW.J'14'11; ..,..I_, ., .", " ", 1,iw.`-i.11., 0 ,,%._.-iftcx,,, ik,.kN 1,. ,_,_'.'.. .;,. .",.I- - _.,-.;`Ml -I'_'-,.-' 4 ,_.,--.-.., .,,,, --I,"!.;,..,',-,.A4- ._,Av- -,, ll. I..;.e,, ".1I"_i. -.1 ,.. ....._.-1.11 -,".,'.;.-4.4, -, --, ,''N,"II`"_I"".§11 ,.. ,RvF., ", .,,.i., ..,t ;I. ,17.- ,,,..',' .-f _,.., ._-,,-,...,"...,.'-.' ," '--..-_".-RK." ,..wtlil_. .I.'',-"I "'.'.1-1, ..,,.;.7,. ,"'.,, .-,-%".`,.,.Iklr..I,_."'i,11-.,.11....-I-."..:,.....,,,.,,,-_ ,.' ,i.,a7 1, I.,,.-.., -, -,..1,,. ,,:- " rIrrvk"'1.1.1...,, ,,. 4.4,.. jj!kv.,"I'l--1..._-,-.jt .,,',., 1,";.,,. ., 1"',- _-,"'n ,,, "'! ,..-.,.1.-.N.:"A ,,, "'.." ._.,".1. ,--. 'A"1 ''.1 -'. .' .,: '.I-,_..,,.1,I,.4, .;' <-. "-'r ,- -, !!, I_`;-,i-i.,-, , -1II..,.,I..,l.-,,1,.--..: .....". ,-,.f,-,.,`.'..,. 11,-1-.-,.r..,, i. i`_I_" .l -- .- ,.,..,;i-._1, .,.-'1111_1;. 4..,I...I- N,.',,^,4i,"lm'.1.11 .".. ,..`.''_ 1-1114_111..,I .",.,.- I-, -,1.-1.,-,.,.7,,.., .; A,,.,_I .-., ,.. 1! _111, I-,,'.. .. .;..?- ,,,..,,,...11.,I.;.l 1,;. ",,.. ._,.lz,. A!Pi ..; `FW11.., --I ` .',,7,.----,-.;Z,q4S,,..'i,-,71. .",' .."`.....'.l,..z ,-II1Ii,Z..rI._.-.i--.-"I'l,"j".",I,.li,.,.,,..l Ili 1; 1.1.11. -t-,_.11,' I .. 11W -rl-.lc_X..4),.0',,,,,`- 11 1.11.",..,I,.,-, ,,. ,.,.,. A. ..,'..l-,,.,,..i.l-"..:,:.ll..t-I I-- -_.--,-,. .,-.--- , -4_z.-,IV h. -:.lz .4&#-'kf "-,-, `.-,.,I.i',,,; b. .,,,-'-',. ",, .""'.1..O.-..,.-1,. i, -1 .,-- ., ;, ,,V-., ..,j',"' -.-.,,._6X..'l..,..:,, ,Jr -1-I,I"." .,.- -.., "' """-,3'...,,",,,""- N,.'f,.;---...;,",,-""%__.---1-,,,"'`.7,I",,,,,..l., .. .....-.- -f.".. -1 -1 `31 _7 ,:--.,,,.,;,I-l ","w,.j.,- ,t. I,-,Z- .N,"....,,, ',,.r,,.%,t.. fg. -',t .-,. J-P;i.., ."-4"i,I,-1-", ,"pZ:71 1;I.,li,I.I_,&`:, -..,I.l,,&_I_1.,.I!,., .,-1-,",-,.", - ., 1. ..",-,.. -.. ,.;I,,--.: ,,-,-, .l ., --,,,.%-,,.. ,.. .,Y?l.-;ll-'l-'ln,t,,",.6..t',4'r. ,;,., ,i.!." .4'. ..III..,_.- ,-,. ,.'..O'w ,"4,-A ,.,,,,,, _,P:.-. 4.1 ,.4.I-i-I,,,_4 ,.,,.,.,.,,-..--. .I ., "',1111 -.- _i. ., ,,',-...- ..! ..,.l-A .1,,',.., Z"', __k.ll. ',.,,. ..J.'i,_-.-,-.' i"I,", ,,!.I..'t..,..': 1:,,,",.,,.-2,-,A,,-,".I..-.1" ."..-I---,I-_.'4..-.",,"',,.,..,..I-1"4.,,.i1-`,-,..",..-.,",,, %, ,..T.1;, ., ..":,I.1;,i.--,,.I.i,..., ;,_k,. .,,.. .. ;I f, .o -",, "I..;. ... ,# r.... ",_.: -.- .f ._..'3i;!_--,,.VW*..- ;,".,*_ T`.1_1'11 ,4"-_-J. .,?.J...`.`-.-.`,,,,",,i- - ... .._,.,.-....I_.-:--:01" -.-.,":'.., illtl..111.'.`_..:.,._,;.,,4..,i,! lf, tit,n .F_,"..I,1:-.. -`j. :.-,i,--.,_-,-,-,."', 11.1-1,.1"7, "I .',,,,-.L-,,, .,cN-. v, .-4 "' ., '__, ,_v,.,,.. 4.11111' -, r",`-, .,.:-...,,-,",k.-;...1, "' - - " ., _ llf., .l,I,,..-4,.t,.,,..., -.j,. ,, .,.,.,-,,." .,,4, ,":. --.1,I'lliI.,..-, " .,_ i.. ,,,, '" ON. I.-li,,11 ,:. ''. 'y ., ,-F,,.-0%_.".I-I.",,,.."I-,-,-ILA,,,m,gN "I-,.l -..-".I.,R.,.t,.,;-.-,-:;. - ,_,,-,5"'.3," ,,-:, ",.,'..,..1114 1..-":C""' JPT..":.",."j,.k,-," _ .,,,,-,".,),-1, ,,.,..-..,._i.t .."1'.. -, .1. .-. ,.'t ";II;, .!. .,,.-" -- .,; -.. 1. -:.;!-,_."'I_-,-1, . ,,, ,%;..w.,4.-..,I....,I".,I.."."_4.I.4,. Ell_. ., :,7', ..-,:'' _-.. 4'_,t,.- -t- -- O-_14,'.-'.A., 1--.'' .4. .",14 A ._-.-..".1,,,..,.,-..",-..,.-'r,-,.,...,,..'-,-?.,,_ A, ,,,.A ,., i;'44A, .;!!.-,..,,-,,t.If;,,t.",.. _. ,,.""'.1".. V?4,`,.;-,t..11, --,,--,-.-.,."-I` -iI ,_,%A..,-.z,,.t_-.I"-..,.::,"..",".0 '.'..f.- ..-4'11..,.1.11,...'I'.,.'.."".,".-"'-I, I,-,-,-,.:w ,*A, ,iW..:.-` 11,,-_7i',-.i.-w-'-;,1)".. _-,.;;..,,, ;,- "6;,i" ,,!',.4, -1-it'., *,..11;'1% ,,". -, -kII"I. -,.,P,7'w'I.- _1,I.1I.o ..,;,,.,, J`-,i4 ]1-1,W.,.7,::,.-fk.S:..._ll ,, 1.,. .; ,A. ,4, .,.,. ,.. I_. .._ " ` -,-,'5 ,."'11,.-."'',-yl'.."I.%.,.-`- ., `pf'.',- ,;S"-..-:.,,;,,_.,_,-,., IIA .I."i.';9,.C"',* 1..0.4,,10- ,- .i ",-''*.io. I"II---.:. If %TA i, oe..A,w. 'Z.,, .1I,1.7;.....i,,,-'t0"... ,.lk -,,).,,,,-,,0l,.,,,._....'.1.7;4-"",71,. 'T.",It --,l,,l,-,l 1, "* ,11-1-11.11 R I...,;,j .,.,,.. k-, .'4-,,,,,--,'%,. 4c2l`.,xz",,-.,.11.,,.I,,1- 4, t-i -1171.,,-.1i..-.`.;,.A-llfl--,-..,,--,-1,1- ,.-, -., 1Y, li,_I1-.h,?..'.1.-_,; 11 U,,. .X-,.0, _x .- -1 4 i,:.'l"-,!'",..'..,I,..'r I1.-.ii.-- ,vik. '.I""60-.vI--.-..,",7 "' ,,,,,,,..,.,..".'.,.el.,...."E.., ,.,.,,-..-,.', :,,..'-l't. .,. 1, -.:...,. ,V,,,- ., ._11-.',I.",'..... .,,vl.M,.,,-.1,-., ",:' ,,,,' "" ". -.` I'.,'s,,,A, ,"x?_,"".--. .,; -,- .., _'.'O,_-,.. -,1._X, I,.j;.,...- 4- :_ "o. .. ':"-7. ," .4. ...-._.,I,i..,,.1--7-.,.--.,,.,W,,...,.,..'74",_ . ..: "' ,-'N-,:... -11,,1. .,'.',,;,.,..,.,7 .,: ,,:`..'--.r'-.N"":',-'-,.'' .i t, ..W";0i.I,,,, ,. 4.. _,. ,..','.. -'_ ....,.Zlit -,,, ,,_",-,", ,.`j,.C,;,,_I .; ,;,., _,4 ..t..,?.l"IN,,,.1,.,,.i'l._..t...,.-. " ,.";.fi.* 4..,,-,,.,, .,-,-.__ -.:.,:- _-.I,, -1--.I'll. '*'..., 4.,.,'A',,-,,.,.,:-f*-.-'-,'NI...,'.4,,-,I,i:zl.R 1_t .,,..!,,.,'. "'. 'l- F".,., -U;,. " ., "fI..__,,,:. .v"-1 W, .,4 ,,,,_," ,11 ",,.'.-,-A,4,,.*!!l.,':_.i'_ .-..11,_..,-IP1..,,,,, W.. .,,- ,, -i .. ..t7 Jw",I,-,..-F*.',,....,,:;p,.;.-, ,,,.' '.Y,'.".1 ..:.;, ,.__i:,,._'!, -. .", ,e -..,.,-_..,.,- -,,,I .-I.I.-N,,1114I.-I'iII'll``^.-., ` -,',Ckv-,A'......,..'4 ., i", 1....:,I;-,Ilg.,...,.r.I,..,.-j-...,I..4I*I-,.11i-.,.I.,:,.. ..",J.. -.' ".-",,, ., ,'.'ll .1' q-,"t.-_ ,r.%.Ii.'I, . 1.,,,V, -,.,.-,,, " C., ;Yl"_-Is -1,_,I""!iv, ,'.O,.,r."" ` _,,,.,_1:., I,1.It,...Wl,-- .- .11,.,. -1,.,'i-4s,.-,11, 1.1 -k.i".1 `.-%','A.;.-.' .'. _-- ..-...4.'_,,._-,N, -,--,".,.-!. -.-,..., ., -_,.1, -I -,f,:.,-_ ,,..I1. Z -2- ,I-,,I.q'. I.;,I_'T.. 1. .,-,,,,-;, -_.'':,".", .:,a I-,I. .N.--,.id-.2a.,-4.4.Aj"I"',"'.. 0.-*',A',,',..:.-,.N.,..,,W_,%'--',,:._';l.:,", ., ? "", ,.,. 10" A.;- .. I 7 R, X., .I,;,4,,,- .." ..:;,.,I""I'll,.I..,..liI.;:,-. A$1- ",4"t,', ., II -.;-..-.r--,,- -..,;., I"II...,1. "Z", -`,,4. -- ,-.. _-,., ,.`1Z-A1'i -_.-.,-1I---.-,!.-.,,., .,,`..-,. ,,_ ,",i,,.o.,.. -, Z,, .,'f..l""..','.,,."e!4i....,.:,I.'.,; 1( ._ .lI.i I., - -, .1-.:' -. ,I.. &,_,.v.I",I,".I1, .I,..," "',,-', ,11. .,'.,.. ,- I'll '. I-1 "_!.."';.,.,'*.,, ."'I., -.,. 1". ..,,II.11-li'l ,,.._,.-"Ii,.";.."4".".I..."v.-.".,-..,_,.'L". -".Cl..";,-,:, ."_-":,,.,. ,,,±, ., 1, ,. ,-.. ,4- *,I.I-.,..I.,.,,,-.,7. .', V , ,C,.., !"".. :',.' i.'q.-,.1-.."-, .. .'-,,;-.,k.,r,.. V.,--, 310- t,:Ak-.!. -- "":.,-.---..'_'- :4,,-I` -, ,,, - 1-11" 11 ..I ,,,, ,---. .,;,41.-..!, I.-, .!,"A.A.4 lF .-I11 M-,IV.t.., k-.1 ..,., ---,...,e. "..,*'-..`.'.-. ,-`.,` 4F..,I-..-,",, ol.II--., I, -,,,.1-..,,;*e. ,-..,. ..,,,._.t---li ..* .',.",P,,,". 4.,ft I,",.,,.1 I,-"'i-lal-I-1.-t.. wr" "I I.,-..-ot,., lvll I.: ,1: 'r, "-.'t .w'-, ,,,-F; .,.. ,.-_,.,.i,"-.".,.".,....-,: cA",..,--.,--;-'-;...'., ,, #...,"..1."..,- _.1 11*I..'kN.I."I,-,,I,,.i. ".11,Ii.U.,r,-,."771%kA.1-l"_N.j", _,.,.,.:_.',,";Q.".2,..-'' -A, ,._., .I1, I,..., .,J, .,_.;k2l _I.4 .. .,! ";,..__ .. ---.:., _14`_ _'_.._..OT,.-1111". V.,:.,, 1,41 -.1%, 7,-.I.., '6.".._ J--,,.,.,..1 4, .l-4.1 ..""., 1,I.n.,'i,vl.l, ,.,.l ._ -41 .-- I..,-`-`,_7""T,- ,.` .- X.F1*7,,lK.- -I;41IA.;, F, .-,.. Z"...`.,-,-.I. I_,",-, .,.-' v.-I:.--,I- ,.W,,,-_....,""Z..,-,.z...t,i....'..'.'.',..,,Pl.,.4',-'4.-.l,.i,..._lll ,7, x. I--,lllll..1-11.7111,.,-,I,'.l.". ,.tl "., .,. 'j, i.,i,.,..,," ., -, 1,-I,.,-;..IIk..,...-I.G ''-"'j.I.,.,'_..,-_71, .,.1-4"..%.-4--1'1. --Ikl- '!., ,.- .,_,',.7,,..., .: _,.-; -..- -1 1:..i., 31'r-,..--..-. II I- ,.4,..,.Zf, -AT."N' 1. 11 11 1,)0._I.;-*-_. li-1, ,.I .,,t,."", I, ..-7111-_.,.,.."Z' AA1,, '.VI,"4"'j, 4, .`;"!",g, -.,-,I,,,.1, ._` "t1 .1-:,,,,, ..'.I. .< LW,-.i0'lti-.. .;-_,.,-,_%...,..,",.:-,,'?$ .- 1.'.'.. "', T", .11. I;. -:Z.-*'.,Z, M--,.A...I.,,-,- -, - . 41_.l1,,iil t-I ...,t. -_ ,.- 1. l,I....-,., ,..-.,,--,,.,,,,,:.,I..I.I-4." -,,,-;- ,..4.',`.%'_ ". e',,.7-,..I":,,'',,,-'-.,.,I-I',---, '40-, -5" ,4.4 .r.,,,. ,,:; ,.N 7y. ".,.. -.,,'4,_`.*A,..:-..-I,t_.",a'..4 ",.,.,I:-i ._- 11 . ., .,.Ttz AM-..m P'll., '". ".;I.,I-I", ?W .."_,.:..-'. ". -`,'N", .", ,'.'!,., ,' ,-",.*- " -,.,.F,,..11- ..;,"""',- ".,.,., .-lj.. ,., ,,I,.-1 ;i%.,'k"._'.,4 " -,. ,,.-.F .* .,-_I._,.,.11--...,.."r --,'',-. ,-,,.,,I-4,7-,,-, n"""I,,. ,I. ,.7V, `Z'-.ll,.-'4,.,-,..,.,,.. e..II._,.....,".,.,,,.,-lt.:, _P'.,,,.'_,,-. _. _-.,,'7Zf "41 ,_',,'."!.:. f,*;'.f,.q ,II '- -,,,; 1.1WA-17-1. .1, ._lk.,..,.;11,,-,,-.x--- --,.._.4 _.. -, ,, I.-1.,",__..g.,II_,., -I'-, ,,'.;-.-'-',,i;k,:,0i. e.-,.--_!...., -, I""i..-..-.-,.,f,..,.;.,..,i:.,,..",..,.,..- .,f,,I%;-01,,"...,.4"'.-Ijp.',-.,.,---1_910, j.--,;----,.,-$04-."--ATZA,.. 1_"..tlil-11-0,...Z,-,-; .,.,t. ..,-,t_,.,,I...,14.."",. .41".. 5,.,-,,,.,,,"" A",-411`,.e","I., 1..x'',.I,,,..,.,,,,,r,.:..*,,,-,.1.A-`.:;.7'-,-'.--`K'-,V.-'-.:V",."--,__.-..R$-,,..,1,-,,,.',1,...,,.-4._,.,.,.,.-.,-1,"'.',-',1._,,.,.1-"__,til,14.L A PHYLOGENY AND CLASSIFICATION OF THE HIGHER CATEGORIES OF TURTLES EUGENE S. GAFFNEY Assistant Curator, Department of Vertebrate Paleontology The American Museum of Natural History Adjunct Professor, Department of Geological Sciences
Load more

Recommended publications
  • A New Xinjiangchelyid Turtle from the Middle Jurassic of Xinjiang, China and the Evolution of the Basipterygoid Process in Mesozoic Turtles Rabi Et Al
    A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Rabi et al. Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 RESEARCH ARTICLE Open Access A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Márton Rabi1,2*, Chang-Fu Zhou3, Oliver Wings4, Sun Ge3 and Walter G Joyce1,5 Abstract Background: Most turtles from the Middle and Late Jurassic of Asia are referred to the newly defined clade Xinjiangchelyidae, a group of mostly shell-based, generalized, small to mid-sized aquatic froms that are widely considered to represent the stem lineage of Cryptodira. Xinjiangchelyids provide us with great insights into the plesiomorphic anatomy of crown-cryptodires, the most diverse group of living turtles, and they are particularly relevant for understanding the origin and early divergence of the primary clades of extant turtles. Results: Exceptionally complete new xinjiangchelyid material from the ?Qigu Formation of the Turpan Basin (Xinjiang Autonomous Province, China) provides new insights into the anatomy of this group and is assigned to Xinjiangchelys wusu n. sp. A phylogenetic analysis places Xinjiangchelys wusu n. sp. in a monophyletic polytomy with other xinjiangchelyids, including Xinjiangchelys junggarensis, X. radiplicatoides, X. levensis and X. latiens. However, the analysis supports the unorthodox, though tentative placement of xinjiangchelyids and sinemydids outside of crown-group Testudines. A particularly interesting new observation is that the skull of this xinjiangchelyid retains such primitive features as a reduced interpterygoid vacuity and basipterygoid processes.
    [Show full text]
  • A Phylogenomic Analysis of Turtles ⇑ Nicholas G
    Molecular Phylogenetics and Evolution 83 (2015) 250–257 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev A phylogenomic analysis of turtles ⇑ Nicholas G. Crawford a,b,1, James F. Parham c, ,1, Anna B. Sellas a, Brant C. Faircloth d, Travis C. Glenn e, Theodore J. Papenfuss f, James B. Henderson a, Madison H. Hansen a,g, W. Brian Simison a a Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA b Department of Genetics, University of Pennsylvania, Philadelphia, PA 19104, USA c John D. Cooper Archaeological and Paleontological Center, Department of Geological Sciences, California State University, Fullerton, CA 92834, USA d Department of Biological Sciences, Louisiana State University, Baton Rouge, LA 70803, USA e Department of Environmental Health Science, University of Georgia, Athens, GA 30602, USA f Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720, USA g Mathematical and Computational Biology Department, Harvey Mudd College, 301 Platt Boulevard, Claremont, CA 9171, USA article info abstract Article history: Molecular analyses of turtle relationships have overturned prevailing morphological hypotheses and Received 11 July 2014 prompted the development of a new taxonomy. Here we provide the first genome-scale analysis of turtle Revised 16 October 2014 phylogeny. We sequenced 2381 ultraconserved element (UCE) loci representing a total of 1,718,154 bp of Accepted 28 October 2014 aligned sequence. Our sampling includes 32 turtle taxa representing all 14 recognized turtle families and Available online 4 November 2014 an additional six outgroups. Maximum likelihood, Bayesian, and species tree methods produce a single resolved phylogeny.
    [Show full text]
  • Sea Turtles in the East Pacific Ocean Region
    Sea Turtles in the East Pacific Ocean Region IUCN-SSC Marine Turtle Specialist Group Annual Regional Report 2019 Editors Juan M. Rguez-Baron, Shaleyla Kelez, Michael Liles, Alan Zavala-Norzagaray, Olga L. Torres-Suárez, Diego F. Amorocho, Alexander R. Gaos Photo: Olive ridley (RMU LO-EPO) at Ostional, Costa Rica Photo credit: Roderic Mast Recommended citation for this report: Rguez-Baron J.M., Kelez S., Lilies M., Zavala-Norzagaray A., Torres-Suárez O.L., Amorocho D., Gaos A. (Eds.) (2019). Sea Turtles in the East Pacific Region: MTSG Annual Regional Report 2019. Draft Report of the IUCN-SSC Marine Turtle Specialist Group, 2019. Recommended citation for a chapter of this report: AUTHORS (2019). CHAPTER-TITLE. In: Rguez-Baron J.M., Kelez S., Lilies M., Zavala- Norzagaray A., Torres-Suárez O.L., Amorocho D., Gaos A. R. (Eds.). Sea Turtles in the East Pacific Region: MTSG Annual Regional Report 2019. Draft Report of the IUCN- SSC Marine Turtle Specialist Group, 2019. 2 TABLE OF CONTENTS REGIONAL OVERVIEW ............................................................................................................ 7 1. RMU Dermochelys coriacea (DC-EPO) ....................................................................................... 8 1.1. Distribution, abundance, trends ........................................................................................................... 8 1.2. Other biological data ............................................................................................................................. 8 1.3.
    [Show full text]
  • A New European Albian Turtle That Extends the Known Stratigraphic Range of the Pleurosternidae (Paracryptodira)
    Cretaceous Research 55 (2015) 74e83 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes A new European Albian turtle that extends the known stratigraphic range of the Pleurosternidae (Paracryptodira) * A. Perez-García a, b, , E. Espílez c, L. Mampel c, L. Alcala c a Centro de Geologia, Faculdade de Ci^encias da Universidade de Lisboa (FCUL), Edificio C6, Campo Grande, 1749-016 Lisbon, Portugal b Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, Paseo Senda del Rey, 9, 28040 Madrid, Spain c Fundacion Conjunto Paleontologico de Teruel-Dinopolis (Museo Aragones de Paleontología), Avda. Sagunto s/n, E-44002 Teruel, Spain article info abstract Article history: Postcranial material corresponding to three specimens of freshwater turtles, from the lower Albian Received 23 December 2014 (upper Lower Cretaceous) of Arino~ (Teruel Province, Spain), is analysed in this paper. This study allows us Accepted in revised form 18 February 2015 to identify the presence of Pleurosternidae (Paracryptodira) outside its known stratigraphic range, from Available online Kimmeridgian to Barremian, and extends its distribution to the Albian. The species from Arino~ represents a new taxon, Toremys cassiopeia gen. et sp. nov., which is the only pleurosternid described so far in post- Keywords: Berriasian levels. Toremys cassiopeia is closely related to other taxa from Europe, to which the Cretaceous Pleurosternidae pleurosternids are restricted. Knowledge about the European freshwater turtle faunas distributed be- Toremys cassiopeia, gen. et sp. nov. fi Lower Cretaceous tween the Barremian and the uppermost Cretaceous is very limited. The new nding provides relevant Albian data on these poorly understood faunas.
    [Show full text]
  • Testudines, Chelonioidea) from the Middle Turonian of Northwest Germany
    Netherlands Journal of Geosciences / Geologie en Mijnbouw 82 (2): 161-167 (2003) Turtle remains (Testudines, Chelonioidea) from the Middle Turonian of northwest Germany C. Diedrich1 & R. Hirayama2 1 Nansenstr. 8, D-33790 Halle/Westphalia, e-mail: [email protected] 2 Faculty of Information Teikyo Heisei University Uruido, 2289-23 Ichihara, Chiba 290-0193, Japan; e-mail: [email protected] Manuscript received: February 2001; accepted: January 2003 Abstract Turtle remains ascribed to the family Cheloniidae (Testudines, Cryptodira, Chelonioidea), collected from the lamarcki zone (Middle Turonian) at Wullen (NW Germany) are described. The material consists of a right humerus, a scapula, a complete costalia, and costalia fragments of a single individual with the humerus indicating a primitive cheloniid of the 'toxochelyid grade'. The present material, as well as previously recorded chelonioid humeri from the Cenomanian and Turonian of Ger­ many illustrate a progressive diversification of chelonioids during the early Late Cretaceous. Keywords: Humeri, Turtles, Cheloniidae, Upper Cretaceous, Europe Introduction ogisch-Palaontologische Institut und Museum der Westfalischen Wilhelms-Universitat Miinster. The turtle remains described in the present paper were collected in the 1970s by Mr. A. Duckstein (Miinster), from the Middle Turonian as formerly ex­ posed at the Hollekamp quarry (topographical map, North Sea i'' 1 : 25 000 sheet 3907 Ottenstein, co-ordinates R 256755 and H 577120, Fig. 1), which is now flooded and actually a freshwater lake. This important fossil KT Hamburg site is very well known in North Germany especially for its rich echinid faunas (Ernst et al. 1998). WULLEN sokm Turtle remains are quite rare in the Cenomanian/ Turonian of Europe and of special interest.
    [Show full text]
  • Membros Da Comissão Julgadora Da Dissertação
    UNIVERSIDADE DE SÃO PAULO FACULDADE DE FILOSOFIA, CIÊNCIAS E LETRAS DE RIBEIRÃO PRETO PROGRAMA DE PÓS-GRADUAÇÃO EM BIOLOGIA COMPARADA Evolution of the skull shape in extinct and extant turtles Evolução da forma do crânio em tartarugas extintas e viventes Guilherme Hermanson Souza Dissertação apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto da Universidade de São Paulo, como parte das exigências para obtenção do título de Mestre em Ciências, obtido no Programa de Pós- Graduação em Biologia Comparada Ribeirão Preto - SP 2021 UNIVERSIDADE DE SÃO PAULO FACULDADE DE FILOSOFIA, CIÊNCIAS E LETRAS DE RIBEIRÃO PRETO PROGRAMA DE PÓS-GRADUAÇÃO EM BIOLOGIA COMPARADA Evolution of the skull shape in extinct and extant turtles Evolução da forma do crânio em tartarugas extintas e viventes Guilherme Hermanson Souza Dissertação apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto da Universidade de São Paulo, como parte das exigências para obtenção do título de Mestre em Ciências, obtido no Programa de Pós- Graduação em Biologia Comparada. Orientador: Prof. Dr. Max Cardoso Langer Ribeirão Preto - SP 2021 Autorizo a reprodução e divulgação total ou parcial deste trabalho, por qualquer meio convencional ou eletrônico, para fins de estudo e pesquisa, desde que citada a fonte. I authorise the reproduction and total or partial disclosure of this work, via any conventional or electronic medium, for aims of study and research, with the condition that the source is cited. FICHA CATALOGRÁFICA Hermanson, Guilherme Evolution of the skull shape in extinct and extant turtles, 2021. 132 páginas. Dissertação de Mestrado, apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto/USP – Área de concentração: Biologia Comparada.
    [Show full text]
  • Universidad Nacional Del Comahue Centro Regional Universitario Bariloche
    Universidad Nacional del Comahue Centro Regional Universitario Bariloche Título de la Tesis Microanatomía y osteohistología del caparazón de los Testudinata del Mesozoico y Cenozoico de Argentina: Aspectos sistemáticos y paleoecológicos implicados Trabajo de Tesis para optar al Título de Doctor en Biología Tesista: Lic. en Ciencias Biológicas Juan Marcos Jannello Director: Dr. Ignacio A. Cerda Co-director: Dr. Marcelo S. de la Fuente 2018 Tesis Doctoral UNCo J. Marcos Jannello 2018 Resumen Las inusuales estructuras óseas observadas entre los vertebrados, como el cuello largo de la jirafa o el cráneo en forma de T del tiburón martillo, han interesado a los científicos desde hace mucho tiempo. Uno de estos casos es el clado Testudinata el cual representa uno de los grupos más fascinantes y enigmáticos conocidos entre de los amniotas. Su inconfundible plan corporal, que ha persistido desde el Triásico tardío hasta la actualidad, se caracteriza por la presencia del caparazón, el cual encierra a las cinturas, tanto pectoral como pélvica, dentro de la caja torácica desarrollada. Esta estructura les ha permitido a las tortugas adaptarse con éxito a diversos ambientes (por ejemplo, terrestres, acuáticos continentales, marinos costeros e incluso marinos pelágicos). Su capacidad para habitar diferentes nichos ecológicos, su importante diversidad taxonómica y su plan corporal particular hacen de los Testudinata un modelo de estudio muy atrayente dentro de los vertebrados. Una disciplina que ha demostrado ser una herramienta muy importante para abordar varios temas relacionados al caparazón de las tortugas, es la paleohistología. Esta disciplina se ha involucrado en temas diversos tales como el origen del caparazón, el origen del desarrollo y mantenimiento de la ornamentación, la paleoecología y la sistemática.
    [Show full text]
  • M a R Y L a N D
    ' o J x l C a JJ¿ ' u¿»... /io hC i M A R T ü R J LES Y L w : A • i v & v ' :À N sr\lài«Q3/ D FRANK J. SCHWARTZ *IES 23102 Vlaams Instituut voor dl Zu Planden Marina Instituts MARYLAND TURTLE FRANK J. SCHWARTZ, Curator C h e s a p e a k e B i o l o g i c a l L a b o r a t o r y S o l o m o n s , M a r y l a n d University of Maryland Natural Resources Institute Educational Series No. 79 J u n e 1967 FOREWORD The 1961 publication of MARYLAND TURTLES resulted in an increased awareness of these interesting members of Maryland’s vertebrate fauna. New in­ formation stemming from this effort has been incorporated into this revision. The researches of Dr. J. Crenshaw, Jr. on the genus Pseudemys, especially Pseudemys floridana, Florida Cooter, has resolved much of the confusion regarding this species’ true distribution and systematica in the state. Its occurrence must now be relegated to an "introduced” or "escape” category. Additional information is also on hand to confirm the Bog Turtle’s tenacious survival in swampy-bog habitats adjacent to the Susquehanna River. Recent information has helped delineate the occurrence of the Atlantic Ridley turtle in the upper Chesapeake Bay. A new section has been added which discusses fossil turtles. It is hoped this edition will maintain interest in and further expand our knowledge of the turtles of the area.
    [Show full text]
  • An Early Bothremydid from the Arlington Archosaur Site of Texas Brent Adrian1*, Heather F
    www.nature.com/scientificreports OPEN An early bothremydid from the Arlington Archosaur Site of Texas Brent Adrian1*, Heather F. Smith1, Christopher R. Noto2 & Aryeh Grossman1 Four turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of the Lewisville Formation (Woodbine Group) in Texas. Herein, we describe a new side-necked turtle (Pleurodira), Pleurochayah appalachius gen. et sp. nov., which is a basal member of the Bothremydidae. Pleurochayah appalachius gen. et sp. nov. shares synapomorphic characters with other bothremydids, including shared traits with Kurmademydini and Cearachelyini, but has a unique combination of skull and shell traits. The new taxon is signifcant because it is the oldest crown pleurodiran turtle from North America and Laurasia, predating bothremynines Algorachelus peregrinus and Paiutemys tibert from Europe and North America respectively. This discovery also documents the oldest evidence of dispersal of crown Pleurodira from Gondwana to Laurasia. Pleurochayah appalachius gen. et sp. nov. is compared to previously described fossil pleurodires, placed in a modifed phylogenetic analysis of pelomedusoid turtles, and discussed in the context of pleurodiran distribution in the mid-Cretaceous. Its unique combination of characters demonstrates marine adaptation and dispersal capability among basal bothremydids. Pleurodira, colloquially known as “side-necked” turtles, form one of two major clades of turtles known from the Early Cretaceous to present 1,2. Pleurodires are Gondwanan in origin, with the oldest unambiguous crown pleurodire dated to the Barremian in the Early Cretaceous2. Pleurodiran fossils typically come from relatively warm regions, and have a more limited distribution than Cryptodira (hidden-neck turtles)3–6. Living pleurodires are restricted to tropical regions once belonging to Gondwana 7,8.
    [Show full text]
  • Primeres Jornades D’Arqueologia I Paleontologia Del Pirineu I Aran Coll De Nargó I La Seu D’Urgell, 29 I 30 De Novembre De 2013
    PRIMERES JORNADES D’ARQUEOLOGIA I PALEONTOLOGIA DEL PIRINEU I ARAN COLL DE NARGÓ I LA SEU D’URGELL, 29 I 30 DE NOVEMBRE DE 2013 DEL PIRINEU I ARAN PRIMERES JORNADES D’ARQUEOLOGIA I PALEONTOLOGIA DEL PIRINEU I ARAN COLL DE NARGÓ I LA SEU D’URGELL, 29 I 30 DE NOVEMBRE DE 2013 D’ARQUEOLOGIA I PALEONTOLOGIA I PALEONTOLOGIA D’ARQUEOLOGIA PRIMERES JORNADES ORGANITZACIÓ PRIMERES JORNADES D’ARQUEOLOGIA I PALEONTOLOGIA DEL PIRINEU I ARAN COLL DE NARGÓ I LA SEU D’URGELL, 29 I 30 DE NOVEMBRE DE 2013 COMISSIÓ ORGANITZADORA Josep Gallart Fernàndez Robert Querol Butia Arqueòleg Territorial Auxiliar administratiu Serveis Territorials de Cultura a Lleida Serveis Territorials de Cultura a Lleida Àngel Galobart Lorente Xavier Reñé Escué Cap del Grup de Recerca del Mesozoic Administratiu Institut Català de Paleontologia Serveis Territorials de Cultura a Lleida Marta Monjo Gallego Montserrat Sorolla Raedo Arqueòloga Territorial Administrativa Serveis Territorials de Cultura a Lleida Serveis Territorials de Cultura a Lleida Marina Aresté Dolcet Júlio Quílez Mata Col·laboradora Director de l’Arxiu Comarcal de l’Alt Urgell Serveis Territorials de Cultura a Lleida Mònica Ferré Moreno Marc Coca Moreno Monitora Col·laborador Dinosfera Serveis Territorials de Cultura a Lleida Carles Gascon Chopo Joaquim Minguell Cardeñes Tècnic de patrimoni Tècnic Consell Comarcal de l’Alt Urgell Serveis Territorials de Cultura a Lleida Lluís Obiols Perearnau Institut d’Estudis Comarcals de l’Alt Urgell DIPÒSIT LEGAL B 9692-2015 DISSENY I MAQUETACIÓ: cr3ativa.com IMPRESSIÓ: Arts
    [Show full text]
  • Mesozoic Marine Reptile Palaeobiogeography in Response to Drifting Plates
    ÔØ ÅÒÙ×Ö ÔØ Mesozoic marine reptile palaeobiogeography in response to drifting plates N. Bardet, J. Falconnet, V. Fischer, A. Houssaye, S. Jouve, X. Pereda Suberbiola, A. P´erez-Garc´ıa, J.-C. Rage, P. Vincent PII: S1342-937X(14)00183-X DOI: doi: 10.1016/j.gr.2014.05.005 Reference: GR 1267 To appear in: Gondwana Research Received date: 19 November 2013 Revised date: 6 May 2014 Accepted date: 14 May 2014 Please cite this article as: Bardet, N., Falconnet, J., Fischer, V., Houssaye, A., Jouve, S., Pereda Suberbiola, X., P´erez-Garc´ıa, A., Rage, J.-C., Vincent, P., Mesozoic marine reptile palaeobiogeography in response to drifting plates, Gondwana Research (2014), doi: 10.1016/j.gr.2014.05.005 This is a PDF file of an unedited manuscript that has been accepted for publication. As a service to our customers we are providing this early version of the manuscript. The manuscript will undergo copyediting, typesetting, and review of the resulting proof before it is published in its final form. Please note that during the production process errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain. ACCEPTED MANUSCRIPT Mesozoic marine reptile palaeobiogeography in response to drifting plates To Alfred Wegener (1880-1930) Bardet N.a*, Falconnet J. a, Fischer V.b, Houssaye A.c, Jouve S.d, Pereda Suberbiola X.e, Pérez-García A.f, Rage J.-C.a and Vincent P.a,g a Sorbonne Universités CR2P, CNRS-MNHN-UPMC, Département Histoire de la Terre, Muséum National d’Histoire Naturelle, CP 38, 57 rue Cuvier,
    [Show full text]
  • As an Indeterminate Plesiochelyid Turtle
    Reinterpretation of the Spanish Late Jurassic “Hispaniachelys prebetica” as an indeterminate plesiochelyid turtle Adán Pérez-García Acta Palaeontologica Polonica 59 (4), 2014: 879-885 doi: http://dx.doi.org/10.4202/app.2012.0115 A partial postcranial skeleton (carapace, plastron, and other poorly preserved elements) of a turtle, from the late Oxfordian of the Betic Range of Spain, has recently been assigned to a new taxon, Hispaniachelys prebetica. This is one of the few European turtle taxa reported from pre-Kimmeridgian levels, and the oldest turtle so far known from southern Europe. The character combination identified in that taxon (including the presence of cleithra, and single cervical scale) did not allow its assignment to Plesiochelyidae, a group of turtles very abundant and diverse in the Late Jurassic of Europe. The revision of the single specimen assigned to this taxon led to the reinterpretation of some of its elements, being reassigned to Plesiochelyidae. This study confirms the presence of Plesiochelyidae in the Oxfordian. However, because the Spanish taxon does not present a unique combination of characters, it is proposed as a nomen dubium. Key words: Testudines, Plesiochelyidae, “Hispaniachelys prebetica”, Oxfordian, Jurassic, Spain. Adán Pérez-García [[email protected]], Centro de Geologia, Faculdade de Ciências da Universidade de Lisboa (FCUL), Edificio C6, Campo Grande, 1749-016 Lisbon, Portugal; Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, C/ Senda del Rey, 9, 28040 Madrid, Spain. This is an open-access article distributed under the terms of the Creative Commons Attribution License (for details please see creativecommons.org), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
    [Show full text]