A, Supplement to the American Land and Lresh-Water Isopod Crustacea

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A, Supplement to the American Land and lresh-Water Isopod Crustacea

BY WILLARD G. VAN NAME

BULLETIN

THEIAMERICAN MIUSEUM OF 4NATURAL HISTORY Vo LXX II. ART. ii, pp. 19-142 New York Issued June 20, 1940

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Y7 -Vu1I4 Article II. A SUPPLEMENT TO THE AMERICAN LAND AND FRESH- WATER ISOPOD CRUSTACEA BY WILLARD G. VAN NAME Figures 1 to 32 During the period that has elapsed since water forms. There are a number of addi- the publication of "The American Land and tions in the family Asellidae, which has Fresh-Water Isopod Crustacea" in May, lately been the subject of an increased 1936,1 a number of additional species have amount of study. been added to the American list and new Trichoniscus (Androniscus) dentiger Verhoeff, information published regarding many 1908 Philoscia bonariensis Giambiagi de Calabrese, others. It therefore seemed desirable to 1935 prepare a supplement to that work to cover Neotroponiscus carolii Arcangeli, 1936 the additional species, correct errors and Platyarthrus hoffmannseggii Brandt, 1833 omissions and record changes in nomen- Porcellio spinicornis occidentalis Miller, 1936 clature and notes and references of im- Porcellio dilatatus Brandt, 1833 ?Porcellio ragusae Dollfus, 1896 portance on other American forms. Porcellio littorina Miller, 1936 This article is to be regarded as purely Circoniscus p1alidus Arcangeli, 1936 supplementary to my work of 1936 named Nerocila armata Dana, 1853 above, and is designed for use only in Asotana splendida (Leigh-Sharpe), 1937 Exosphaeroma platense Giambiagi, 1922 connection with it. No attempt is made to Asellus montanus Mackin and Hubricht, 1938 repeat matter that was given in that work. Asellus dentadactylus Mackin and Hubricht: The present article consists of the fol- 1938 lowing three parts: Caecidotea macropopoda Chase and Blair, 1937 Caecidotea ozarkana Chase and Blair, 1937 I.-Descriptions of new species and other Mancasellus ouachitaensis Mackin and Hub- species added to the American list. richt, 1938 II.-Additional notes and references, Mancasellus louisianae Mackin and Hubricht, changes of names, corrections, etc., apply- 1938 ing to species dealt with in my 1936 work. Through the kindness of Dr. E. C. III.-Additions to the bibliography (in- Williams, Jr., of Northwestern University, cluding works omitted in 1936). I have had the opportunity of examining The following four species are described a large collection of isopods recently made as new to science: at Barro Colorado Island, Panama Canal Trichoniscus (Miktoniscus) medcofi, new species Zone, but failed to find any undescribed Philoscia floridana, new species species. I am also under special obliga- Philoscia avrilensis, new species tions to Dr. J. C. Medcof, of the Canadian Exosphaeroma insulare, new species Fisheries Research Board, Mr. Stanley The following species and subspecies Mulaik of Edinburg, Texas, and Prof. described by other writers are added to the T. D. A. Cockerell of Boulder, Colorado, list of known American land and fresh- for interesting material sent me. PART I.-DESCRIPTIONS OF NEW SPECIES AND OTHER SPECIES ADDED TO THE AMERICAN LIST SUBORDER ONISCOIDEA, LAND ISOPODS Trichoni GENUS OR SUBGENUS ANDRONISCUS European species and races, the females of VERHOEFF, 1908 which are practically alike, though the A division of Trichoniscus in the com- males are distinguishable by characters of prehensive sense composed of a number of the pleopoda and legs. They are remark- 1 Bull. American Museum of Natural History, able for their coloration, usually a striking LXXI, pp. i-vii, 1-535, Figs. 1-323, by Willard G. pink or orange-pink, in life. (See Ver- Van Namne. hoeff, 1908c.) 109 110 Bulletin American MusBeum of Natural History [Vol. LXXVII Most of them have been included under numerous densely crowded tubercles ar- the name Trichoniscus roseus (Koch). The in true ranged transversal rows. Cephalon roseus was described by Koch (as Itea transversely oval, with the lateral lobes rosea) from Bavaria. front Among the most distinctive well-marked, denticulate, nearly character- straight. Lateral parts of mesosome more istics of the males of this group are the expanded than in the other peculiar modification of the merus of the rather species, 1st seventh pair broad and partly flanking the legs and the spoonlike termination cephalon, the 3 posterior recurved of endopodite of the first pleopoda, both pairs and acuminate. Metasome (in male) ex- of which are well exemplified in the illus- ceeding half the length of the trations of the species that follows. and mesosome, but little more than half as broad; terminal expansion of last segment trans- Trichoniscus (Androniscus) dentiger versely truncate, with 4 small apical spi- Verhoeff, 1908 nules. Antennulae with the last joint much Figure 1 longer than the 2nd, and carrying 6 sen- Androniscus dentiger VERHOEFF, 1908C, P. sory filaments, 5 apical and 1 lateral. 139, Fig. I0.-VANDEL, 1933, p. 131.-WAECHT- Antennae comparatively slender, attaining

Fig. 1. Trichoniscus (Androniscus) dentiger %Verhoeff, 1908. After Sars, 1899 (T. "roseus"). LER, 1937, p. 258, Fig. 50.-MEDCOF, 1939, P. 115. 1/3 of the length of the body, flagellum Philougria rosea KINAHAN, 1857, Nat. Hist. composed of 4 articulations. Last pair of Rev., V, p. 197, P1. XXIII, fig. 3.-BATE AND legs in male peculiarly modified, having WESTWOOD, 1868, Hist. Brit. sessile-eyed Crust., the meral II, p. 460, Fig. joint strongly dilated, and produced inside a Trichoniscus (Androniscus) dentiger ARCAN- to large conical prominence, GELI, 1931a, p. 12. in against which the succeeding joint Trichoniscus roseus BUDDE LUND (in part admits of being bent. Inner ramus of 1st only), 1879, p. 9; 1885, p. 247.-SARS (in part pair of pleopoda in male, with the terminal only), 1899, p. 163, P1. LXXIII, fig. 1. dilated at the "Body oblong oval, joint end in the form of a greatest width al- little bowl of a spoon; that of 2nd pair most attaining half the length, dorsal face rather but strong and distinctly 3-articulate, slightly convex, and rough, owing to terminal joint knife-shaped and denticu- 1940] Van Name, Supplement to American Isopod Crustacea ill late inside. Uropoda with the outer ramus in the belief that it would be found in about twice the length of the basal part, Norway also. inner ramus somewhat narrower, but The first American record of this species scarcely shorter. Color of dorsal face in is due to Dr. J. C. Medcof. He reports it life of a clear minium-rose. Length of (Medcof, 1939) as occurring in many adult male 5 mm. greenhouses in the region of London, "This species, first described by Koch, Ontario, and Toronto.

Fig. 2. Trichoniscus (Miktoniscus) medcofi, new species. is easily distinguished by its comparatively Trichoniscus (Miktoniscus) medcofi, broad and flattened body, the dorsal face new species of which is densely granular, and by its Figure 2 beautiful colour, which, however, is only Body proportionately narrower than usual in seen in fresh specimens." (Sars, 1899, pp. the Trichoniscidae (ratio at least 4 to 1 if the 163-164.) intersegmental muscles are relaxed), the back This is the common, if not the only, considerably arched transversely, its surface with scattered short hairs and many small scabrous representative of the roseus group in tubercles which form at least four more or less northwestern Europe (British Islands, Bel- definite transverse rows on most of the thoracic gium, Luxemburg and parts of France), segments. The tubercles are very conspicuous including, at least in greenhouses, localities on the head and anterior segments becoming less and prominent toward the rear but do not disappear in northern Germany Denmark. Sars, entirely even on the abdomen. The frontal 1899, included and illustrated it (under the outline seen from above is convex, with fairly name roseus) in his Crustacea of Norway large lateral lobes extending obliquely down and 112 Bulletin American Museum of Natural History [VOl. LXXVII forward and having a rounded anterior margin. basal joints of the uropoda are short and have Eyes composed of a single large ocellus sur- the insertion of the inner branch removed by a rounded by a somewhat larger area of black small lateral interval from, and somewhat far- pigment. Antennae with a flagellum of four ther forward than, that of the outer branch, distinct articles and a terminal group of bristles. which is considerably the longer and stouter. Thoracic segments I to IV inclusive, with the The largest example is a male measuring rear corners rounded (IV only slightly), V to somewhat over 5 mm. long when the inter- VII have the rear corners quite acute and in- segmental muscles are well relaxed. creasingly extended backward. The legs are all The specimens, except very young ones, are quite long with well-developed spines. Leg VII strongly pigmented, having the upper surface of the male has the merus widened posteriorly handsomely variegated with brown or purplish- and compressed from side to side. brown pigment on a light background which is The abdomen is abruptly somewhat narrower whitish or yellowish in the preserved material. than segment VII of the thorax, and tapers There is usually a narrow dark streak across the considerably. The rear corners of segments 3 forehead extending through and behind the eye to 5 are extended back into sharp points rather on each side. closely appressed to the following segment. The pleopoda of the first two pairs of the male LOCALITIES.-Native habitat unknown. conform quite closely to Kesselyak's (1930) de- Dr. J. C. Medcof, for whom the species is scription of the type species Miktoniscus linearis. named, kindly sent me about 40 specimens,

Fig. 3. Philoscia bonariensis Giambiagi de Calabrese, 1935. After that author.

The outline of the median sexual appendage a of from may be described as Kesselyak does, by calling including large proportion males, it cigar-shaped. It has a slightly acuminate tip Urbana, Illinois, collected October 27, 1936, on which a few fine cross striations are visible on on the moist floor of the Botanical Green- high magnification. This appendage 'is quite house of the University of Illinois where it long and is not normally carried between, but is said to be abundant and a few additional extending ventral to, and independent of the first pair of pleopoda. ' That pair has the exopo- ones from another greenhouse in that city, dites plate-like, narrowed in the di,tal half also a single specimen from the Garfield and rounded at the tip; the endopo1ites are Park Conservatory, Chicago, Illinois, col- styliform and composed of two narrow elongate lected February 9, 1939. The is segments. The distal segment may be described type in as a very narrow, slightly twisted tapering The American Museum of Natural History lamella stiffened by an axial midrib, which is (Cat. No. A.M.N.H. 9069). itself slightly curved dorsally and 'nedially. The extreme, slightly curved tip, when highly magnified, is seen to bear a small file-like area Oniscidae of sharply defined furrows on its convex aspect. In the second pleopoda the exopodite is plate- Philoscia bonariensis Giambiagi de like, short and wide, with a narrow posterior Calabrese, 1935 extension; the endopodite has a short basal Figure 3 and a segment long styliform distal 'segment Philoscia bonariensis GIAMBIAGI DE CALA- which tapers into a very slender seta that reaches BRESE, 1935, pp. 495, 496, 1 Fig. (misspelled far back toward the end of the abdomen. Phyloscia in legend of The telson has the median extension rather figure). narrow. The rear end is without spines and is Only briefly described, and the figure truncated in a distinctly convex curve. The here reproduced given in the above paper. 19401 Van Name, Supplement to American Isopod Crustacea 113 A more detailed description in a future parts of the outline of the head. Eyes unusually large, almond-shaped, wide in front and very number of Anal. Mus. Argentino Cien. Nat. bulging but composed of rather few large con- is promised. spicuously developed ocelli, 13 in the larger The species is said to resemble P. longi- specimen, 11 in the smaller, in addition to one or cornis Budde-Lund of Mediterranean coun- more poorly developed ones in each case. No tubercle between the first antennae. tries but to have longer and differently Second antennae moderately long, quite slen- proportioned antennae. The integument der and very slightly pubescent. In the male is of soft consistency, minutely punctate specimen their length if straightened out would and covered with a short, scattered pubes- exceed half the body length. The first article of the flagellum, which exceeds the fifth joint cence. The uropoda are as long as the of the peduncle, is slightly the longest; the last abdomen. Color dusky ("pardusco"). one, exclusive of the terminal bristle, is almost Length 8.5 mm., width 3.5 mm. as long as the first.

Fig. 4. Philoosca floridana, new species. LOCALITY.-Quequ6n, Province ofBuenos Thoracic segments I to III have the rear Aires, Argentina. Types in Mus. Argen- corners rounded, in IV they form about a right angle; the last three segments are increasingly tino, Cien. Nat., No. 13051. extended back with the rear corners angular (differing conspicuously from the rounded cor- Philoscia floridana, new species ners in P. inquilina). All the thoracic segments have the lateral ends cut off in a convex curve, Figure 4 especially the more anterior ones. Legs moder- Closely allied to Philo8cia inquilina from ately long, rather weakly spinous. Sexual British Guiana and P. richmondi from Puerto differences in them were not observed. Rico, especially to the former. The abdomen is fairly long and not greatly Body seen from above elliptical, the surface tapered. Segments 3 to 5 have the extended unusually even and glossy, the head abruptly rear corners small, appressed and acute. Telson narrower below the eyes, rather large, with the with the sides slightly converging in the anterior front border convex (slightly sinuous when the part; the main part is widely triangular with a head is tilted up); with scarcely a vestige of slight suggestion of sinuosity in the sides; lateral lobes, the eyes forming the antero-lateral the apex slightly prominent but not acute. 114 Bulletin American Museum of Natural History [Vol. LXXVII

Uropoda mutilated or missing in the available larger eyes, the acute instead of rounded specimens. The extended endopodites of the first pleop- rear corners of segments V, VI and VII, oda of the male are stout and noticeably bent differently shaped pleopoda in the male as outward at the abruptly narrowed tips. Their well as by the coloration. There appear to exopodites have a rather long backward exten- be other minor differences in the present sion rounded at the tip. In the two specimens at hand the coloration of the upper parts is quite species, such as the wider head, less definite striking and probably somewhat characteristic. frontal line, the tip of the telson less acute, On the main part (tergum) of each thoracic etc., but if so it is difficult to be sure with- segment the anterior part is light colored, the out more material. pigment being confined to the posterior part, Two male and while on each of the epimera there are two specimens, female, both rather narrow dark antero-posterior stripes on a more or less mutilated and incomplete, light (unpigmented) background. This pro- were obtained by Dr. F. E. Lutz at Winter

Fig. 5. Philoscia avrilersis, new species. duces the effect of seven light and seven wide Park, Florida, in February, 1939. The dark stripes across the back between the epimera, with a longitudinally striped border on male specimen is the type (Cat. No. each side comprising three light and two dark A.M.N.H. 9023). stripes on each epimeron. The coloration of the head and abdomen presents nothing striking Philoscia avrilensis, new species except the presence of a conspicuous dark band between the eyes. There is some dark pigment Figures 5, 6 on the legs and lower parts. The body, seen from above, is rather broadly The species appears to be a small one. The oblong, with lateral ends of the first thoracic male, though fully adult and with firm integu- segment and to a decreasing extent also of the ment is only 5 mm. long and not quite 2 mm. second and third, turned horizontally outward wide, the female is immature and smaller and so as to form a projecting border. proportionately narrower-bodied. The head is narrow and deeply set back in the thorax. The lateral lobes are unusually This species is distinguished from P. prominent for a member of the Philoscia group. inquilina, apparently its closest ally, by They are of rounded outline, but are turned 194011Van Name, Supplemen-t to American Isopod Crustacea 115 into an obliquely vertical position so that as seen turned directly backward. The endopodite of from above they appear quite narrow. The pleopod 1 of the male has three teeth at its tip. upper border of the epistome is quite closely The telson is wide and has only a very short appressed to the front of the head and dips backward extension, which is triangular with downward in a deep V at the median line as concave sides qnd an acute apex which does not seen from in front. Above this border the reach as far back as the tips of the fifth epimera. median part of the forehead is slightly concave The telson has a shallow median depression or or impressed. cavity on the surface. Both uropoda are missing The eyes are large, with at least 22 well- in the only specimen. The color is the usual developed ocelli. The anteininae are loing anid brown with light (unpigmented) markings char- slender, reaching alonig the fourth thoracic seg- acteristic of the Philoscia group. ment when drawni back. Their flagellum is Length of the type and only specimen (a male): likewise long and slender; its proximal article is 11.2 mm. It is Catalogue No. 8101 in the the longest and the terminal one a little the A.M.N.H. collection. shortest, if its short terminal spine is not in- LOCALITY.-Bois d'Avril, Haiti, altitude cluded. 6000 feet, collected by Bird and Winkhans, March 7, 1935. This is quite different from any other American species of Philoscia that I have seen. The broad body with small tubercles on the surface, the head deeply set back in the thorax, the long posteriorly directed abdominal epimera and the prominent lateral lobes of the front of the head give it some superficial likeness to an Oniscus. There are, however, no traces of the fluting of the external plates of the first and second plopoda nor any tracheae in them that I could discover after removal of the appendages and clearing in glycerine. It agrees quite closely in many characters with the southern European subgenus Tiroloscia Verhoeff, 1926 (Mitt. Bulg. Ent. and Zool. Fig. 6. Philoscia avrilensis, new species, details. Gesell., III, p. 155, 1929, Jahrb. Syst., LVI, p. 136, Figs. 30, 39).

The entire dorsal surface as well as that of the antennae is noticeably rough and scabrous as NEOTROPONISCUS ARCANGELI, 1936 seen with a hand lens; in addition there are, on This genus was established for the fol- the head and thoracic segments, especially the forward ones, scattered small tubercles which for lowing peculiar species: the most part are not arranged with any regut- larity. On the thoracic segments there is a very Neotroponiscus carolii Arcanigeli, 1 936 slightly impressed transverse furrlow a little way in front of the rear margin. Figure 7 Even the first thor acic segmenit shows, by the Neotroponiscus caroltii ARCANGELI, 1936, p. sinuous curve of its rear margin, a slight tend- 201, PI. Iv, figs. 1-4. ency to a backwar d extension of the l ear General form of the body that of a lateral corners. Theii extension increases progressively and becomes very great in the three Porcellio; the relatively small convexity posterior segments. In these last three seg- of the dorsal surface is limited to the ments the reai corner is acute; in segments I, tergites; the epimera extending out hori- II and III it is r-ounded; in segment IV the zontally, especially those of the first seg- apex of the angle is slightly rounded off. The legs are rather weak and slender though fairly ment. long. There does not appear to be very much Under high magnification integument of sexual differentiationi in the legs, though the the dorsal surface appears finely scaly, but more numnerous spinies pr esenit oni the merutis and(1 the chief characteristic is the presence of carpus of the three anfterior pairs of legs of the male may be a sexual character. high conical tubercles r-egularly arranged The abdominal epimera are long, narrow and in transverse rows on the head and body 116 Bulletin American Museum of Natural History [Vol. LXXVII

segments. They do not extend on to the LoCALITY.-Two female specimens were epimera. There are also small scattered obtained at Pirajil, State of Sao Paulo, scale-like setae, some of which form a Brazil by Prof. E. Caroli. series along the rear margin of the seg- Arcangeli considers this genus related to ments. Porcellio although no tracheae are present The head is completely immersed in the in the external plates of any of the ple- concavity of the front thoracic segment; it is opoda. remarkable for the very long lateral and median lobes as may be seen from the PLATYARTHRUS BRANDT, 1833 illustration. The eyes have 17 to 18 The broad, depressed body of oval out- ocelli, the antennae are stout and reach line with rather large expanded epimera, beyond the rear margin of the second the short but wide flattened antennae segment of the thorax when drawn back. which present the unusual feature of having

Fig. 7. Neotroponiscus carolii Arcangeli, 1936. After Arcangeli. Lower right-hand figure shows the posterior margin of segment I greatly enlarged.

The flagellum has the proximal article the flagellum reduced to a single article; almost half the length of the second. the short thick legs, absence of eyes and The epimera of segments 3 to 5 are long the white or whitish coloration due to and curved backward, the telson has a absence of pigment, distinguish this genus, long, narrow, oblong median extension which is of subterranean habits, living espe- rounded at the end and provided with a cially in ants' nests. Of the few species prominent median keel for two-thirds of its constituting the genus, only the following length. The uropoda are very small but is widely distributed. of the form usual in Porcellio and its allies. COLORATION.-Variegated with brownish Platyarthrus hoffmannseggii Brandt, and pallid areas, becoming whitish on the 1833 epimera, uropoda and antennae. Length: 4.4 mm.; width (at the 4th thoracic seg- Figure 8 Platyarthrus hoffmannseggii BRANDT, 1833, ment): 2.11 mm. p. 12, P1. iv, fig. 10.-BUDDE-LUND, 1879, p. 2; 1940] Van Name, Supplement to American Isopod Crustacea 117

1885, p. 199.-SARS, 1899, p. 175 (-seggi), P1. part rather large, oblong, outer ramus Lxxvi, fig. 2.-ARCANGELI, 1921, p. 190 (-geggi), about same length and lanceolate in form, P1. VII, fig. 1.-WAECHTLER, 1937, p. 274, Fig. 75. inner ramus much narrower, and scarcely Sars, 1899, p. 175, gives the following to the middle of the outer. description: extending "Body broadly oval, and much depressed, Colour pure white. Length of adult with the dorsal face smooth, though slightly female 3 mm." granulose all over. Cephalon partly flanked A species widely distributed in southern by the side-plates of 1st segment of meso- and central Europe. It was not included some, frontal edge arcuate and minutely in my work of 1936 as I could then find crenulated, lateral lobes narrowly rounded no record of its occurrence in America. at the tip, and obliquely produced in front, Since that time Dr. G. E. Hutchinson ob- edges crenulated. Side-plates of meso- tained it near Lindsley Pond, North Bran- some rather large, lamellar, subcontiguous, ford, Connecticut, in the nest of an ant 1st pair the largest, edges finely denticulate. "apparently Lasius flavus nearcticus

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Fig. 8. Platyarthrus hoffmannseggii Brandt, 1833. After Sars, 1899. Metasome scarcely exceeding 1/3 of the Wheeler," and also found one in a nest of length of the mesosome, the 2 anterior seg- Formica fusca var. subseracea Say. He ments very small, epimeral plates of the 3 sent me two fine specimens fromui the North succeeding segments narrow, recurved; Branford locality, and says it is apparently last segment comparatively short, tri- well established in the vicinity of New angular, resembling that in Philoscia Haven, Connecticut. muscorum. Antennulae with -the basal joint fully as long as the other 2 combined. Porcellio spinicornis occidentalis Antennae scarcely exceeding 1/3 of the Miller, 1936 length of the body, and hirsute all over Figures 9A, B with short hairs; last peduncular joint Porcellio spinicornis occidentalis MILLER, very large and dilated, flagellum shorter 1936, p. 170, Figs. 8, 12, 20, 25, 26. than that joint, and lanceolate in form. Differs from the typical spinicornis Legs densely spinous inside, propodal joint according to Miller in the following conical in form. Uropoda with the basal respects: 118 Bulletin American Museunm of Natural History [Vol. LXXVIf

1.-Articles of flagellum subequal; the Porcellio dilatatus first not longer than the second. 2.-Me- Brandt and Ratzeburg, 1833 dian lobe of the head not so expanded. Figure 10 3.-Ischiopodite of seventh legs of male Porcellio dilatatus BRANDT AND RATZEBUJRG, more bowed. 4.-Telson spatulate rather 1829-1833, II, p. 78, P1. xii, fig. 6.-BRANDT, than triangular. 1833, p. 14, P1. iv, figs. 7, 13.-BUDDE-LUND, LoCALITIES.-Common in the San Fran- 1879, p. 2; 1885, p. 106.-METNERTZ, 1934, p. 237, Figs. 11, 12, etC.-WAECHTLER, 1937, cisco Bay region and along the coast of p. 283, Fig. 88. California, also found at Davis near Allied to P. spinicornis an(l similar to it Sacramento (Miller). "P. spinicornis" has in most respects, but conspicuously been reported as occurring in California wider- "with bodied, and lhaving the extendled part of the P. laevis" by Essig, 1926, p. 3, but telson rounded at the end insteadl of to what extent this record may apply to pointed. Head and anterior part of the the present subspecies I do not know. body tuberculated, becoming smoother toward the posterior end. Color of upper parts more or less marbled or variegated, but the middle and often the lateral lobes of the head dark colored. Median lobe of head somewhat triangular and rather prominent when seen from above. DISTRIBUTION.-A native of the westernii Mediterranean region, now widely dis- persed about human habitations in various parts of the world. As noted under P. s. occidentalis, it may occur in California, but the only American specimen to come into my hands is one from Indian Gardens, Grand Canyon, Arizona, collected bv Dr. P. C. Geiser, June 23, 1938. ?Porcellio ragusae Dollfus, 1896 c Figures 11, 12 B Porcellio ragusae DOLLFUS, 1896f, p. 2, Fig. 2.-VERHOEFF, 1933a, pp. 3, 19, 25, 28; 1938, Fig. 9. A, B. Porcellio spinicornis occi- pp. 102, 119. dentalis Miller, 1936. After Miller. C. Por- cellio spinicornis, typical form, outline of telson. I assign to this species provisionally, After Sars, 1899. and with some hesitation because of certain points of uncertainty mentioned below, Miller's figures indicate that the telson four specimens from Texas, that apparently and the frontal outline of his new sub- represent an Old World form that has been species are very different from those of the accidentally introduced. No Europeani typical spinicornis as illustrated by Sars specimens have been available for com- an(l Waechtler, and as I find them in parison. eastern American examples. The pleopoda In the Texas specimens the body, seen are similar to those of the typical form of from above, is oblong, rather parallel spinicornis according to Miller. Were it sided, with the head deeply set back into not that the closely related species P. dila- the thorax, and the abdomnen short though tatus Brandt, 1933 (see below), is consider- not abruptly narrower than the last seg- ably wider-bodied than spinicornis and ment of the thorax. It is considerably therefore not likely to be confused with it, arched transversely. I would be inclined to suspect that P. s. The dorsal surface of the head and occidentalis might be that species, which is thorax are quite thickly covered with small now known to occur in America. rounded tubercles which are most numer- 1940]1Van Name, Supplement to American Isopod Crustacea 119 ous on the head and anterior thoracic segment V when well drawn back. The segments, becoming fewer and less con- females have the antennae considerably spicuous posteriorly, and disappearing on shorter and weaker. The flagellum of the the abdomen. They show a tendency to antennae is long and has the basal article arrangement in transverse rows, about five much exceeding (but not twice as long as) rows on the first thoracic segment decreas- the terminal one. The antennae have the ing to three and finally to two on the segments more or less grooved or longi- posterior ones. Besides these there is a tudinally carinated, but with poorly de- row of very much smaller tubercles along veloped dentiform processes. or close to the rear border of the segments. Only the last three thoracic segments The general surface of the back between (nos. V, VI and VII) have the rear angles the tubercles bears scattered, very minute actually sharp. Segment IV has only the scabrous setae or stiff and very short hairs, extreme apex of the angle rounded a little; visible only with some magnification. those in front have it more so. Even the The head is rather narrow and well set first thoracic segment shows a slight sinu- back in the thorax. The eyes are large ous curvature of the rear margin of the with many well-developed ocelli not all of segment. This curvature becomes pro- which are well pigmented. Below them gressively greater towar(l the rear; in

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Fig. 10. Porcellio dilatatus Brandt, 1933. After Waechtler, 1937, and Meiinertz, 1934. the frontal margin forms two large lateral segments III and IV it has already become lobes extending obliquely forward; they well marked. are of somewhat square outline with the The legs of segments I-III, inclusive, corners much rounded. Between them and VII show sexual modifications in the the frontal margin is continuous but up- male, those of 1, II and to a less extent III, turned and much less prominent, except having brush-like areas of short spines as that in the middle it is extended forward usual in the genus. The legs of segment into a small lobe (widely triangular with VII have the basipodite considerably in- straight sides as seen from above) whose flated or swollen, and the ischium of some- slightly upturned surface is somewhat what concave outline below and with a concave. This median lobe extends even large rather shallow depression on the farther forward than the lateral ones when external aspect of the distal half of that the head is held level. Above its median segment, as if it had been pinched or projecting point the front outline of the pressed in. forehead as seen fronm above is slightly The outlines of the male pleopoda ap- re-entrant. proximate to those of Porcellio spinicornis. The antennae are long and fairly stout in The telson has its posterior extension un- the male, reaching the anterior part of usually large and wide, triangular and 120 Bulletin American Museum of Natural History [Vol. LXXVII acute at the tip with rather straight sides. verging toward the telson, which is pig- It has an oval but only very slightly con- mented, as are also parts of the large basal cave median depression on the upper segments of the uropoda, the antennae and surface. The uropoda have flattened ta- the upper surface of the lateral and median pering external branches which are quite lobes of the head. short in the female as shown in the figure, Body length of the largest specimen, but which are at least one and one-half which is a female, about 11 mm. One of times as long in the only one of the males the male specimens is not very much in which they are not missing. smaller. The coloration of the upper surface is LoCALITIEs.-Two females and a male

Fig. 11. Specimen supposed to be Porcellio ragusae Dollfus, 1896. rather pale, and consists of a light marbling were received from Mr. Stanley Mulaik of gray or blackish on the light yellow who collected them at Corpus Christi, background formed by the unpigmented Texas, March 21, 1936, with specimens of parts. The pigmented areas are so placed Porcellionides mulaiki, Porcellionides prui- as to give the back a somewhat indistinctly nosus andArmadillidium vulgare. A smaller yet noticeably longitudinally striped effect male was received from L. I. Davis from as shown in the figures, leaving a rather La Joya, Texas. The type locality of P. wide lateral border unpigmented. On the ragusae Dollfus is Ficuzza, near Palermo, abdomen the stripes are continued, con- Sicily. 19401 Van Name, Supplement to American Isopod Crustacea 121 P. ragusae Doilfus is intermediate be- nent, almost vertical; median lobe large tween the typical subgenus of Porcellio and rounded. Eyes small, composed of (represented by P. scaber) and the sub- about six to eight ocelli, situated at base of genus Mesoporcellio Verhoeff (represented anterolateral lobes. First pair of anten- by P. laevis) exhibiting an unusual combina- nae rudimentary. Second pair of antennae tion of characters of both, such as the extend to posterior margin of second thoracic segment. First article of flagellum short, one-third the length of second. "Segments of thorax subequal. First segment with anterolateral angles only pl slightly produced around head. Posterior I margins of first three thoracic segments 'N straight, rounded at lateral angles. Pos- terolateral angles of remaining thoracic b segments become progressively more and VIIi more produced posteriorly, but not ex- so. Fig. 12. Specimen supposed to be Porcellio tremely Legs slightly spiny. Ischiopo- ragusae Dollfus, 1896. Details of male. dite of seventh walking legs in male with a pronounced humplike process. granulated body surface and triangular "Abdomen as usual in the genus, with median lobe of the head resembling that of lateral parts of first and second segments P. scaber, with the only slightly sinuous covered by last thoracic segment. Pos- rear border of the first body segment char- terolateral parts of third, fourth and fifth acteristic of Mesoporcellio. segments acuminately produced almost at Dollfus' description is brief and his fig- right angles to transverse axis. Telson ures show few details. As compared with triangular, only slightly produced. Exopo- the Texas specimens the European form is dite of first pleopods in male bears a dis- much larger, females reaching 20 mm. in length, with a wider body, and plain coloration (gray with three light spots at the base of the telson), and if the figures are accurate the external angles of the lateral frontal apNexp lobes appear to be less rounded. It is, however, quite possible that the Texas specimens available are not full grown, which may account for some discrepancies, )~~~~~~~~(% and with the present material I do not feel justified in describing the Texas form as new.

Porcellio littorina Miller, 1936 Fig. 13. Porcellio littorina Miller, 1936. Figure 13 Adapted from Miller, 1936.

Poreellio littorina MILLER, 1936, p. 168, Figs. 3, 7, 14, 15, 19.-HATCH, 1939, p. 257. tinctive knob-like process on its outer distal "Body smooth, nearly two and a half margin. times as long as wide, 6.0 mm.: 2.5 mm. "LOCALITIEs.-Twelve specimens, in- Color brown, with a band of irregular light cluding one female with embryos in marsu- spots on either side of median dorsal line. pium, collected under rocks above high Lateral margins of thorax somewhat tide at Bay Farm Island, Alameda, Cali- lighter. fornia (type locality), along with two "Head but slightly immersed in first species of Actoniscus." (Miller, 1936, p. thoracic segment. Front produced in 168.) Actoniscus is a syn. of Armadil- three lobes, anterolateral lobes not promi- loniscus. One specimen recorded by Hatch, 122 Bulletin American Museum of Natural History [Vol. LjXXVII

1939, from Bay Center, Washington, just The head is rather wide; its anterior above high tidle line. border forms an even curve as seen from above except beneath each eye, where it is Porcellionides saussurei (Dollfus), produced into small rounded lobes extend- 1896 ing obliquely downward, so that they show Figure 14 but little in a dorsal view of the head un- Metoponorthats saussurei DOLLFUS 1896d, p. 48, less the latter is tilted up. The lobes are Fig. 2.-RICHARDSON, 1905, p. 626, Fig. 673 concave on their outer (after Dollfus). (antero-lateral) surface. Porcellionides saaissurei VAN NAME, 1936, p. 245, Fig. 137 (after Dollfus). The upper margin of the epistome forms The American Museum of Natural His- a narrow upturned border which extends tory has two small specimens which seem across the face between the lateral lobes very probably referable to this species de- at about the level of the lower third of the scribed in 1896 by I)ollfus from an incom- eyes, dipping down in the middle a little plete (headless) specimen, and which does in a gradual sinuous curve. The eyes are not appear to have been found since then. rather long oval, large and prominent, well Dollfus' description and figures are, how- pigmented and have about 17 well-developed ocelli. The antennae are moderately long with a comparatively stout flagellum of two articles, the terminal one very slightly longer than the other. The flagellum fully if not more than equals the last segment of the peduncle in one speci- 9 men, but not in the other, owing to the greater length of that segment. The terminal spine is short. The body surface on magnification, especially when dry, is seen to be covered with a rather coarse scabrous granulation. There is a slightly impressed transverse furrow on the thoracic segments, near the rear margin in their median part, but curving more forward on the lateral parts of the segments. The thoracic segments have the rear lat- Fig. 14. Specimen believed to be Porcellio- eral angles rounded off in the first and sec- nides saussurei (Dollfus), 1896. ond but little in the third, and successively more angular in succeeding ones, though ever, too inadequate to thus identify only those of VI and VII can be called them with a great degree of assurance, al- actually acute. Beginning very slightly though their locality and the correspond- with the fourth, the rear angles are suc- ence with his description and figures seem cessively more produced backward. too close to justify treating these speci- The thoracic epimera are directed down- mens as new. I (lo not understand what a ward in the anterior part of the body. "posterior dlepression" on each abdominal Toward the rear, the epimera including segment mentioned by Dollfus means. I those of the abdominal segments 3, 4 and 5, find a depression (noticeable but very shal- which are long, acute and curved backward, low) only on the telson. The telson is spread out more in a lateral or horizontal somewhat more acute and it, as well as the direction. uropoda, is not proportionately as long as The telson is almost triangular and is in Dollfus' figure, possibly because of a sex large and long, reaching beyond the ends of difference, as both the present specimens the epimera of segment 5, the uropoda and are females. their branches are also rather elongated. 19401 Van Name, Supplement to American Isopod Crustacea 123 The coloration is quite conspicuous, the to C. bezzii and C. gaigei and has a smooth upper parts being brown variegated with body surface and very short but stout an- light (unpigmented) spots so distributed as tennae. Among the specific characters to leave four quite noticeably darker given by Arcangeli are the outline of the stripes, two of these located each side the epimera of the first thoracic segment and median line, separated by a confluent row the frontal line; as seen from above the of light spots, the other two are on each latter forms a slightly re-entrant angle at side along the basal part of the epimera. its median point; the epistome below it The upper part of the epistome and the projects forward slightly as a very obtuse median part of the forehead just above it triangle. are conspicuously dark. The antennae are The best distinguishing characters of the dark, ringed with whitish at the joints. species are, however, the telson which, The "three little light spots at the base of the terminal segment" (telson), mentioned by Dollfus, are quite noticeable in both of the specimens. The smaller of these specimens is 6 mm. long, the other if straightened out might measure nearly 8 mm. LOCALITY.-The locality of the specimens (Cat. No. A.M.N.H. 8104) here described is Vera Cruz, Mexico, which is on the coast. The type locality of Dollfus' species is at Cordova, not far distant from Vera Cruz, but at an altitude of over 2800 feet. Fig. 15. Circoniscus pallidus Arcangeli, 1936. Lower right hand figure, lateral view of Cubaridae segment I (lateral portion). After Arcangeli. Circoniscus pallidus Arcangeli, 1936 Figure 15 though rounded at the tip as in the other Circoniscus pallidus ARCANGELI, 1936, p. 204, species named above, is triangular with P1. IV, figs. 5-9. practically straight converging sides, and This appears to have been described by the uropoda, the large basal joint of which Arcangeli from a single small specimen has the external border much more curved which, from its color, entirely a somewhat than in the other species and both endo- translucent white, against which the black podite and exopodite extending directly color of the eyes and the brown of the teeth backward and tipped with a group of setae. of the mandible stand out, may perhaps SIZE.-2.84 mm. long and 1.3 mm. wide. have been quite immature. LOCALITY.-Pirajll, State of Sao Paulo, C. pallidus is a typical Circoniscus allied Brazil. Collected by Prof. E. Caroli. SUBORDER FLABELLIFERA Cymothoidae Monod's article (1931a) in which he treats N. fluviatilis as synonymous with Nerocila armata Dana, 1853 Dana's armata was unfortunately over- Nerocila fluviatilis SCHIOEDTE AND MEINERT, 1881, p. 66, P1. v, figs. 6-9; 1884, p. 414.- looked in my work of 1936. Dana's speci- VAN NAME, 1936, p. 431, Fig. 267.-CORDERO, men was from Rio Janeiro, and was evi- 1937, p. 4, Figs. 1-8. dently marine. Monod states, however, Nerocila armata DANA, 1853, p. 761, P1. L, that he had examined a number of speci- figs. lOa-lOd.-MONOD, 1931a, p. 6, Figs. 2-4; 1937, p. 465. mens from Guiana in the British Museum (For other synonyms and discussion see from the fishes Leporinus fasciatus, Creni- Monod, 1931a.) cichla saxatilis, Chichla ocellaris and Pseud- 124 Bulletin American Museum of Natural History [Vol.[ LXXVII

Monod, 1937, points out that Leigh- Sharpe's new genus is a synonym of Asotana and that his species splendida, described from fresh water (Rio Napo) in Equador is very possibly identical with A. formosa Sch. and M., 1881, a conclusion that can hardly be questioned as far as the genus is concerned, and which I am much A-1 inclined to accept for the species also. PgA22 But Leigh-Sharpe's figures show that his specimen is much smoother above than the highly ornamented one depicted (possibly rather too fantastically) in Schioedte and Meinert's work, and as the localities are -widely separated, splendida from a Fig. 16. Asotana splendida (Leigh-Sharpe), tributary of the Amazon in northern Equa- 1937. After Leigh-Sharpe. dor, and formosa from a Pacific slope river of southern Peru, a decision on that ques- auchenipterus nodosus and that it occurs tion may be postponed. both in salt and fresh water. He has given much attention to the study of this genus Two of Leigh-Sharpe's figures are repro- of isopods, and his view should carry much duced here. His type and only specimen is weight. a female 21 mm. long and is preserved in "N. fluviatilis" was also redescribed and the British Museum. The first antennae figured by Cordero, 1937, from salt water in have but four articles while formosa has the vicinity of Montevideo, where it is a eight, but such a character would have to common parasite of various fishes. be demonstrated in several cases at least in order to have much weight. Asotana splendida (Leigh-Sharpe), 1937 "Colour (in alcohol); a pale straw tint: Figure 16 on the first and second free thoracic seg- ?Asotana formosa SCHIOEDTE AND MEINERT, ments dorsally there are roughened patches 1881, p. 155, P1. x, figs. 10-12.-MONOD, 1937, of dark brown denticles, and on the second p. 465. Badroulboudour splendida LEIGH - SHARPE, segment two larger areas of black den- 1937, Parasitology, XXIX, p. 391, Figs. 1-4. ticles." (Leigh-Sharpe, loc. cit.)

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Fig. 17. A, Exosphaeroma insulare, new species, dorsal view (length of, specimen 8 mm.). B, Same, frontal margin, bases of first antennae (a') and frontal lamina (flm). C, Exosphaeroma oregonense (Dana), 1853, same details as in B. 19401 Van Name, Supplement to American Isopod Crustacea 125 Sphaeromidae most easily observed character distinguishing it from E. oregonen8e. The slightly prominent Exosphaeroma insulare, new species frontal line, which forms the margin, dips down in oregonense to form three very obtuse yet Figures 17A, B distinct V-shaped angles, one median just above Very closely related to the well-known marine the minute oval downwardly projecting oval species E. oregorense (Fig. 17C) of the Pacific rostrum, and one on each side above the distal coast, and resembling it in most of its characters end of the wide flattened basal segments of the though it may not reach as large a size as that first antennae, and the edge is raised into a dis- species. tinct tubercle on the outer or lateral side of the The largest of the specimens available for lateral V. Beyond (lateral to) this tubercle is description, is almost 8 mm. in body length a small notch. when well straightened out, and a little over In the present species the angles are less well 4 mm. wide. The other specimens are consider- defined and the tubercle is hardly noticeable, ably smaller and more or less closely rolled up. so that the margin approaches more nearly to

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Fig. 18. Exosphaeroma platense Giambiagi, 1922. Adapted from Giambiagi.

The general form of the head, body, thoracic forming a gently curved sinuous line. The and abdominal segments including the broad rostrum, which does not project forward, is of and considerably arched telson, is as in E. oblong outline rather than oval. There appears oregonen8e, though the lateral ends of the also to be a slight difference in the uropoda of thoracic segments are more obtuse in their the two species. In both they are of rather termination. Seen from above the body is wide blade-like flattened form, with the exterior oblong, nearly straight sided, and broadly margin considerably curved and the end rounded rounded before and behind; the upper surface truncate and not provided with hairs. The is only slightly rough and granular (less so than outer branches are the longest and largest, in oregonense). The head is less than half the reaching when extended directly back at least width of the first thoracic segment. The eyes as far as the broadly rounded end of the telson. are large, well developed and very convex, with In the present species both branches appear numerous ogelli; the flagella of the antennae to be proportionately a little narrower than in have in the type specimen 8 articles in the first oregonen8e; the inner branch extends about pair and 11 in the second pair. four-fifths of the length of the outer branch. The anterinor margi of the head furnishes the The color of the specimens in alcohol is 126 Bulletin American Museum of Natural History [VOl. LXXVII blackish gray above with small oblong yellowish Exosphaeroma platense Giambiagi, markings on the dorso-lateral regions of the back corresponding to the muscular insertions. 1922 LoCALITY.-Eleven specimens were re- Figure 18 ceived from Prof. T. D. A. Cockerell, who Exosphaeroma platense GIAMBIAGI, 1922, p. collected them in fresh water in company 234, P1. ii. with a mollusk of the genus Physa (iden- This is a species much resembling E. tified as P. virgata Gould by Dr. W. J. thermophilum Richardson, 1905, in, form Clench) on San Nicolas Island, off the but at once distinguishable by the very coast of southern California. Type: Cat. short ovate external branches of the urop- No. A.M.N.H. 8092. oda, which are much less than half the E. thermophilum (Richardson), 1897 from, length of the internal branches. The first a warm spring near Socorro, New Mexico, antennae have 6 to 8 articles in the flagel- appears to be a near ally of this species, lum, the second antennae 7 to 10 articles. but not identical with it. It has the telson The species attains a body length of about narrower behind and the outer branch of 4.5 mm. the uropoda is half as long and half as It was omitted from my 1936 work in the broad as the inner one, so there is evidently belief that it was found only in salt or a considerable difference in that respect be- strongly brackish water, but a letter from tween the two species. In E. thermophilum Dr. Giambiagi de Calabrese of December the inner branch is much wider and the ex- 17, 1936, states that it occurs also in fresh ternal one proportionately much smaller water in the Rio Santiago, Province of and narrower than in the present species. Buenos Aires, Argentina. SUBORDER ASELLOTA Asellidae gnathopoda. The most important charac- Asellus montanus ter of the present species is, however, to be Mackin and Hubricht, 1938 found in the tip of the endopodite of the Figure 19 second pleopoda of the male. The tip is Asellus montanus MACKIN AND HUBRICHT, curved inward (toward the median line) 1938, p. 629, Figs. 1, 2, 7, 9, 10. and has the appearance of being twisted Very closely resembling A. intermedius, into a small spiral. The exopodite has but differing in that the latter has only one long setae (up to 10 in number) and many distal process on the palmar margin of the small ones along the inner margin. There

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Fig. 19. Asellus montanus Mackin and Hubr:icht, 1938. After Mackin and Hubricht. 1940] Van Name, Supplement to American Isopod Crustacea 127 is much variation in the proportionate Arkansas and Leflore and Latimer Coun- length of the telson and the uropo(la, and their branches. ties, Oklahoma. "This species attains about 12 mm. as Asellus dentadactylus maximum body length. The color pattern Mackin and is a mottled, rich Hubricht, 1938 reddish-brown with Figure 20 lighter yellowish spots on the head and thoracic segments, and a Asellus dentadactylus MACKIN AND HUBRICHT, uniform brown on 1938, p. 629, Figs. 3-6, 8. the pleotelson. Sometimes with a triangu- "A small mottled species, lar dark area on the head, but rather vari- in greyish-brown able in detail. color, not exceeding 10 or 12 mm. in length. The head is without lateral "The first antennae vary in flagellar seg- of the ments up to about 12. flanges; flagellum antennules with The second an- about a maximum of 13 segments. An- tennae are about as long as the body, but tennae not may exceed that quite as long as the body, with length. The largest about 70 segments in the of the specimen (a male) had 97 segments in the flagellum flagellum." largest specimen. "The gnathopoda are very much like

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Fig. 20. Asellus dentadactylus Mackin and Hubricht, 1938. After Mackin and Hubricht. The figure shows the armature of the those of Asellus communis male gnathopod (left inside view). (see Racovitza, 1919). The dactyl has a large triangular "Note the thickened basal portion of the near the dactyl, two distal process base, which is the most processes of the palmar distinctive characteristic. Note in the edge of the propodus, one blunt and the armature of the other acute and triangular. propodus the wide thin Usually 2 flange or cutting of the mar- heavy spines at the base of the palm, edge palmar but gin; the larger of the two palmar processes somewhat variable in this respect. Other is a continuation of this characters of the gnathopod flange. Also the are not sig- diminishing series of heavy spines at the nificant. The dactyl of the last peraeo- heel of the poda with palm." (Mackin and about 2 short ventral spines 1938, p. 629.) Hubricht, proximal to the claw, and usually 2 setae The details figured by the describers are on the dorsal side." (Mackin and Hub- here richt, 1938, p. 630.) reproduced. They state that the margins of all the segments are LoCALITIES.-Type locality: creek in and that thickly spined, Scott County, in some specimens the uropoda Arkansas, 4 miles south of are proportionately longer than in the Boles. Also localities in Lawrence County, figure. 128 Bulletin American Museum of Natural History [Vol. LXXVII

"This species is much like Asellus inter- LoCALITIEs.-Type locality: small creek, medius and probably will be confused with 5 miles south of Locust Cottage, Jefferson it. The male pleopoda furnish the best County, Arkansas (Cotype Cat. No. characters for separation. In the first 74841, U. S. National Museum). This pleopoda, the very long setae of the tip, species was also obtained in a small creek 2 so conspicuous in A. dentadactylus, are ab- miles south of Saline, Natchitoches Parish, sent in A. intermedius. The tip of the Louisiana. endopodite of the second pleopoda in A. intermedius is equipped with 2 processes, Caecidotea macropropoda absent in A. dentadactylus. A. montanus of Chase and Blair, 1937 this paper is also closely related, but that Figure 21 species is distinguished from any other by Caecidotea macropropoda CHASE AND BLAIR the spiral-like tip of the copulatory organ." 1937, p. 221, P1. I. (Mackin and Hubricht, 1938, p. 630.) The description and figures given by

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Fig. 21. Caecidotea macropropoda Chase and Blair, 1937. All figures based on male specimens. After Chase and Blair. 19401 Van Name, Supplement to American Isopod Crustacea 129

Chase and Blair are from male individuals. No figures or descriptions of the male Many of the characters given as specific pleopoda are given. are based on the relative lengths of the seg- LOCALITY.-A small stream in a cave and ments of the body, antennae and uropoda; at the mouth of it, five miles south of Kan- characters very variable within a species sas, Oklahoma. in this family. They are sufficiently shown in the figures here reproduced in Caecidotea ozarkana outline, so that it is not necessary to quote Chase and Blair, 1937 those parts of the description here. Body 13-17 mm. long; 3 mm. wide. Figure 22 Head 2 mm. wide and about 1 mm. long. Caecidotea ozarkana CHASE AND BLAIR, 1937, Flagellum of first antenna with 13-18 p. 221, P1. ii. articles; second antenna, 91-123 articles. According to the description and figures The propodus of the first pair of legs this is a smaller, slenderer form than C. (gnathopod) is very large and has in its in- macropropoda. Besides such differences as ferior margin two long triangular processes the figures, whose outlines are reproduced and two short ones. here, clearly show, the following distinc-

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Fig. 22. Caecidotea ozarkana Chase and Blair, 1937. All figures based on male specimens. After Chase and Blair. 130 Bulletin American Museum of Natural History [Vol. LXXVII tions between the two forms are stated in There are 2 or 3 propodal processes, the the description. one of the medial region being always well Body 6-8 mm. long, 1-3 mm. wide. developed and the distal one small.... The Head as wide as the body and about as basal portion of the dactyl is thickened long as wide. sometimes to form a process and may be Flagellum of first antennae of 11 articles; much more definite than shown in the of the second antennae 60-70. figure....In M. macrourus the basal or The terminal abdominal segment is proximal processes, while in M. ouachi- "three halves as long as wide" instead of taensis this process is absent or at most "four thirds" as in C. macropropoda. represented by only a low spine-tipped The uropoda are "a little longer than" elevation." (Mackin and Hubricht, 1938, the terminal abdominal segment instead p. 632.) of twice as long as in C. macropropoda. "The figure shows the form of the first "The triangular processes of the propo- male pleopoda. Note that the basal seg- dus on the first pair of legs, vary from one ment is nearly as long and wide as the dis- large one on the distal portion of the in- tal one. There are 1 or 2 coupling hooks. ferior margin with spines along the edge, The setae of the distal segment are small to one large triangular process located dis- terminally and increase in size onto the tally and two located proximally with spine sides. There is a more or less abrupt tran- along the margin." sition between the short and long setae.

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LoCALITY.-Spring five miles south of The maximum number of the long setae Kansas, Oklahoma, about one-half mile is about 15, and 18 for the short ones, but from the cave where C. macropropoda was the total may be much less." (Mackin found. and Hubricht, 1938, p. 633.) LoCALITIES.-Streams in Leflore, Lati- Mancasellus ouachitaensis mer and Pushmataha Counties, Oklahoma. Mackin and Hubricht, 1938 Type locality: tributary of Kiamichi Figure 23 River near Big Cedar, Leflore County. Mancawellu8 ouachitaensis MACKIN AND HUB- Cotype, No. 74844, U. S. National Mu- RICHT, 1938, p. 632, Figs. 13, 15, 17, 18, 20. seum. In general appearance similar to M. macrourus except that the uropoda are Mancasellus louisianae somewhat longer in proportion. The head Mackin and Hubricht, 1938 may or may not be laterally incised. Figure 24 Length up to 11 mm. The antennae may Mancasellus louisianae MACKI5 AND HUB- attain a length equal to the body and have RICHT, 1938, p. 634, Figs. 11, 12, 14, 16, 19. as many as 90 articles in the flagellum. Closely similar to Asellus hoppinae but "The male gnathopoda are distinctive. differing in lacking the mandibular palp, 1940] Van Name, Supplement to American Isopod Crustacea 131 in having relatively shorter uropods, and "In the first pleopoda of the male, the in the details of the armature of the gnatho- basal segment is much narrower than the pods. distal one, and there are 2 pairs of coupling "The body length in this species does not hooks. The setae of the distal segment exceed 7.5 mm. in any of our specimens. number up to 30. The second pleopoda of The color is brown with yellow mottlings. the male have the following characteristics: The head is deeply incised and the incision (1) the internal distal angle of the basal may be narrow or very broad. The an- segment not at all cut away, so that this tennules have up to 7 flagellar segments, segment is practically square, (2) the exo- and the antennae 56 in the largest speci- podite is straight on the outer margin, men. The antennae are not as long as the hardly curving to the tip at all, (3) the body by a considerable margin. setae number up to about 14, (4) the endo- "The dactyl of the male gnathopoda is podite is without even a suggestion of slightly thickened at the base. The pro- apophysis at the base, (5) the copulatory

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zi gn(i 71)

Fig. 24. Mancasellus louisianaeMackin and Hubricht, 1938. After Mackin and Hubricht. podus has a large central process on the or bulbar portion of the exopodite is palmar margin, a small distal one and a nearly spherical in outline. Note also spine at the base, but no process. Other the form of the tip of this appendage." details of the armature are shown in the (Mackin and Hubricht, 1938, p. 634.) figure. The dactyl of the last peraeopoda LoCALITY.-Reported only from among possesses up to 5 spines below, the last dead leaves in a small creek 2 miles south (accessory claw) large and curved. There of Saline, Natchitoches Parish, Louisiana. is a row of 4 or 5 setae above. Cotype, No. 74843, U. S. Nat. Museum.

PART II.-ADDITIONAL NOTES AND REFERENCES, CHANGES OF NAMES, CORRECTIONS, ETC. (Arranged alphabetically) Actoniscus, genus. See Armadilloniscus. ADDITIONAL LoCALITY.-San Nicolas Is- AUoniscus cornutus Budde-Lund, 1885 land, off southern California, obtained by ADDITIONAL LoCALITY.-San Nicolas Is- Prof. T. D. A. Cockerell. Essig, 1926, land off southern California, obtained by gives it as common on ocean beaches in Prof. T. D. A. Cockerell. California, feeding on kelp and other sea weeds. Miller, 1938, p. 117, gives notes on Alloniscus perconvexus Dana, 1856 habits. 132 Bulletin American Museum of Natural History [Vol. LXXVII Armadillidium nasatum Budde-Lund, This species is recorded (as Actoniscus 1885 tuberculatus) by Miller, 1938, pp. 114, 117, Figure 25 under rocks on beaches at Alameda in the Two specimens were collected by J. K. San Francisco Bay region. Karlovic under pieces of wood at Glenview, Illinois, northwest of Chicago, Oct. 3, 1936. Artystone trysibia Schioedte, 1866 In America this species is usually confined Mentioned by Leigh-Sharpe, 1937, with to hothouses and warmed buildings. the statements that it was recorded "from It was described by Budde-Lund in 1885,- the body cavity" (!) of 2 fish, Geophagus having been previously listed without de- brasiliensis from the Humboldt River, scription in his "Prospectus" published in Brazil, and that it was "the only hitherto 1879. In both cases the name is nasatum, recorded parasitic isopod from fresh wa- not nasutum, a frequent error. It may be ter."(!) He says that "there are six excel- worth while to state here that Budde- lent specimens in the British Museum." Lund's Prospectus, which is a rare work, is merely a list of species, and contains no ASELLOTA, SUBORDER CORRECTION.-In Van Name, 1936, p. 453, in the diagnosis of this group, the I words "not cheliform" should be replaced by "sub-cheliform." Asellus brevicauda Forbes, 1876 Asellus brevicaudus MACEIN AND HTi- BRICHT, 1938, p. 631. Many localities in Illinois, Missouri and Arkansas reported. Specimens from the caves in St. Clair and Monroe Counties, Illinois, are reported as broader in proportion to their length than in the typical form, and having much reduced pigmentation. VIE The American Museum of Natural History has examples from Wet Cave near Anna, southern Illinois, collected by J. H. Fig. 25. Armadillidium nasatum Budde- Karlovic, May 3, 1936, and many speci- Lund, 1885. After Waechtler, 1937, and Mein- mens from Missouri localities received from ertz, 1934. Dr. Hubricht. descriptions or figures whatever. The new names in it were nomina nuda, and had no Asellus communis Say, 1818 standing in nomenclature until republished Mackin and Hubricht, 1938, p. 629, re- with descriptions in 1885. port this from various localities in the Middle West from Illinois to Oklahoma, Armadillidium vulgare (Latreille), 1804 inclusive. CORRECTION.-The name Latreille in the CORRECTION.-In caption of Fig. 1 in captions of Figs. 157-159 in Van Name, Van Name, 1936, p. 7, representing Asellus 1936, should be in parentheses. communis, read "appendages" instead of "segments." Armadilloniscus holmesi Arcangeli, 1933 New name for A. tuberculatus (Holmes Asellus hoppinae Faxon, 1888 and Gay), 1909, proposed by Arcangeli, Reported by Mackin and Hubricht, 1938, 1933, p. 59, on account of preoccupation by p. 632, from localities in Arkansas. The A. tuberculatus Dollfus from the Azores. American Museum of Natural History has 1940] Van Name, Supplement to American Isopod Crustacea 133

a number of examples kindly donated by 3 figures), describes and illustrates the Dr. Hubricht, from springs and creeks female and young male of this species from (not subterranean waters) in Johnson specimens in the National Museum of County, Arkansas, collected in the spring Buenos Aires obtained on fishes in the Rio of 1936. Some of these specimens are of San Santigo at La Plata, Argentina, doubt- large size: males up to 19 mm., females to less from quite salt water. Cordero, 1937 16 mm. in length. Most of the specimens (p. 8, Figs. 9, 10), figures and records it from have the uropoda, both base and branches, Montevideo and from a pond near the Rio quite long. Some of them have about 100 San Francisco, Jatobt, State of Pernamii- articles in the flagellum of the second buco, Brazil, from fishes of the family antennae. Loricariidae. The females are of broad rounded outline as in Braga cichlae Schi- Asellus incisus Van Name, 1936 oedte and Meinertz, 1883, and Cordero ex- CORRECTION.-The description, Van presses the opinion that B. fluviatilis may Name, 1936, page 496, lines four and five be the same as that species. This seems of last paragraph, should state that the to me quite probable. internal, not the external branch of the uropoda is the longer. Caecidotea alabamensis Stafford, 1911 Additional localities reported by Mackin A large male (16 mm. long) from Monte and Hubricht, 1938, p. 630: Stone County, Sano, Madison County, Alabama, agrees in Missouri, spring 300 yards from entrance most respects with the description and to Marvel Cave; Barry County, Missouri, but the outer Roaring River State Park. figures, tooth on the pinching edge of the chela has a small acute process not described or figured by Stafford. This Asellus intermedius Forbes, 1876 process extends parallel to the dactylus Additional locality reported by Mackin which closes against (beside) it. The first and Hubricht, 1938, pp. 630, 631: Bayou antenna of this specimen has 12 articles in Boeuf, Lafourche Parish, Louisiana. the flagellum; the second antenna has the In view of the insufficient descriptions flagellum broken off, but with 86 articles and figures of this species that are avail- in the part remaining. able the statements of these authors regarding it that are made under A. denta- Caecidotea antricola 1931 dactylus and A. montanus in the above Creaser, article are of importance. ADDITIONAL LoCALITY.-Osage Hills Country Club, Kirkwood, St. Louis County, Asellus Missouri, in "outlet of drain near hole 17." tomalensis Harford, 1877 Three specimens, one about 20 mm. long, Reported by Carl, 1938, p. 451, as oc- received from Dr. Leslie Hubricht. curring in water of slight salinity at Van- couver, B. C. Caecidotea californica (Miller), 1933 Asellus MILLER AND Badroulboudour splendida Leigh-Sharpe, californicus Hoy, 1937 1939, p. 347, etc., Fig. 1. See under Asotana splendida, page 124 of this article. Calvigeroniscus Typographical error for Clavigeroniscus, Van Braga cichlae Schioedte and Meinertz, Name, 1936, p. 85. 1881 Circoniscus See remarks under Braga fluviatilis. gaigei Pearse, 1917 I refer here with a little hesitation a few Braga small specimens in The American Museum fluviatilis Richardson, 1911 of Natural History collection received from Giambiagi de Calabrese, 1933 (p. 512, Iquitos, Peru. 134 Bulletin American Museum of Natural History [Vol. LXXVII

Cirolana anops Creaser, 1936 Haplophthalmus danicus Budde-Lund, Figure 26 1879 These figures are reproduced from Found in Allen Gardens greenhouse at Creaser's 1936 article to supplement the Toronto, Ontario, by Dr. J. C. Medcof. brief notice of this species in Van Name, 1936, p. 523. Ligia baudiniana Milne Edwards, 1840 (syn. Ligyda b.) Cirolana cubensis Hay, 1903 A series of articles by T. C. Barnes, in Compared with the new species C. anops the Biological Bulletin for 1932 and later in Creaser, 1936, p. 119. years, deal with experimental work done at Bermuda upon this species. A figure in Cubarhina, subgenus. See under Tri- his article of 1935 (Biol. Bulletin, LXIX, chorhina. pp. 259-268, Fig. 1) showing a young indi- vidual just emerged from the marsupium Cubaris tuberosa (Budde-Lund), 1904 and not having the seventh pair of legs Verhoeff, 1937, pp. 410, 411, disagrees developed should, however, be mentioned. with Kesselyak (1930) who transferred this This species is recorded from Yucatan by species from Cubaris (syn. Armadillo) to Pearse, 1936, p. 22, and Creaser, 1936, p. Reductoniscus Kesselyak. 120.

a-t

Fig. 26. Cirolana anops Creaser, 1936. Adapted from Creaser. Cylisticus convexus (de Geer), 1778 Ligia exotica Roux, 1828 Reported from vicinity of Buenos Aires, Reported (as Ligyda e.) from Quequen, Argentina, by Giambiagi de Calabrese, Prov. of Buenos Aires, Argentina, by Giam- 1936a, p. 509. biagi de Calabrese, 1935a, p. 509. Exosphaeroma oregonense (Dana), 1853 See page 125, under E. insulare, new Ligia occidentalis Dana, 1853 species. Miller, 1938, p. 117, gives note on habits. Exosphaeroma papillae Bayliff, 1938 Ligia pallasii Brant, 1833 Exosphaeroma papillae BAYLIFF, 1938, Miller, 1938, pp. 117, 118 (habits, etc., p. 213, Figs. 1-11. and illustration, showing both sexes, the This should, I think, be excluded as wider-bodied). properly a marine species, though it some- males being much times ascends streams into water "practically fresh" according to Bayliff. Ligidium elrodii (Packard), 1873 LoCALITY.-Cold Spring Harbor, Long This is very probably not distinct from Island, New York. L. longicaudatum Stoller. If this could be 19401 Van Name, Supplement to American Isopod Crustacea 135

proved Packard's name would have prior- species, though with some doubt, as in ity. certain respects they appear intermediate in character between L. symmetrica and Ligidium gracile (Dana), 1856 L. guianensis Van Name, 1925. For this CORRECTION.-In caption of Fig. 22, reason he raises the question whether these Van Name, 1936, authority should be two species are not really the same, a (Dana), not (Stimpson). conjecture that may prove to be correct in Hatch, 1939, p. 256, reports this species view of the great variability of species of from moist situations in dense woods from this group with age, sex, etc., and con- sea level up to 4500 feet in Washington sidering the small amount of material State. Miller, 1938, p. 119, gives notes on (three specimens) on which the latter spe- habits and distribution. cies was based.

Ligidium latum Jackson, 1923 Mancasellus brachyurus, Harger, 1876 A number of specimens of this species were Reported by Mackin and Hubricht, received from Mr. R. M. Bond who 1938, p. 632, from a stream on Blanket collected them in a small perennial stream Mountain, Tennessee, in Smoky Mountain on Mission Peak (elev. 1750 feet), southeast National Park. of Mission San Jos6, Alameda County, California. The process on the tip of the styloid Mancasellus macrourus Garman, 1890 process of the second pleopod of the male Localities in Arkansas and Oklahoma is movably articulated. The small scales reported by Mackin and Hubricht, 1938, of the body surface are easily rubbed off; p. 632. According to Marcus, 1930, it is between them the surface is smooth and fairly abundant in the vicinity of Urbana, shining. Illinois.

Ligidium longicaudatum Stoller, 1902 Nerocila fluviatilis Schioedte and Mein- Specimens of this species were received ertz, 1881 from Dr. Leslie Hubricht of the Missouri See under N. armata Dana, page 123 of Botanical Garden, St. Louis, in the spring this article. of 1936, from localities in St. Louis County, Missouri, and Natchitoches Parish in northwestern Louisiana. Among the latter Phalloniscus anomalus (Dollfus), 1890 are examples (females) of 9 to 10 mm. This species is mentioned in the paper body length. The number of articles in by Bowley, 1935, p. 46, which otherwise the flagellum reaches 13 to 14. These deals exclusively with the New Zealand specimens have large light spots on the and Australian species of the genus, no basal part of the epimera, giving the effect specimens or new information about the of a light border to the thorax. Males South American form (anomalus) being from the Missouri localities have the available. Its position in this genus seems styloid process of pleopod 2 with only a open to question. slight undulation, hardly to be called a notch, near the end. Philoscia incerta Arcangeli, 1932 Livoneca guianensis Van Name. 1925 One specimen about 3 mm. long which See appeared to be this species, mounted entire remarks under Livoneca symmetrica. in Canada balsam on a microscope slide, was sent me for examination by Prof. Livoneca symmetrica Van Name, 1925 Monod of Paris. It was collected at Foret Cordero (1937, p. 9) refers two specimens de Fumee, Guadeloupe, West Indies. The believed to be from fresh water of the Rio species has thus far been known only from Tocatins, State of Pars, Brazil, to this the nearby island of Dominica. 136 Bulletin American Museum of Natural History [Vol. LXXVII

Philoscia (Ischioscia) mineri Van Name, based on those of Sars) I am supplementing 1936 them here with some others from the recent An immature specimen apparently of works of Waechtler, 1937, and Meinertz, this or a very closely allied species from 1934. Foret de Fum6e, Guadeloupe, West Indies, was sent me for examination by Prof. Porcellio formosus Stuxberg, 1875 Monod. The species has been recorded See under P. virgatus, page 138. only from Dominica Island. Porcellio laevis Latreille, 1804 Philosica muscorum (Scopoli), 1783 Figure 27 Dailey and Hatch, 1940, report taking Porcellio laevis MILLER, 1938, p. 171, specimens that appear to be this species at Figs. 4, 9, 16, 22, 27. two points in Seattle, Washington, several Reported from various places in Cali- miles from salt water. They say that the fornia from the San Francisco Bay region manner of its occurrence indicates that it southward. See also under Porcellionides may have been introduced in nursery stock. virgatus.

Fig. 27. Porcellio laevi8 Latreille, 1804. After Waechtler, 1937, and Meinertz, 1934.

Philoscia richardsonae Holmes and Gay, Porcellio scaber Latreille, 1804 1909 Figure 28 ADDITIONAL LoCALITIEs.-Obtained by Porcellio scaber, MILLER, 1936, p. 166, Miller (1938, p. 117) in grassy situations on Figs. 2, 6, 10, 17, 24; GIAMBIAGI DE CALA- the shores of San Francisco Bay, at Ala- BRESE, 1936a, p. 509. meda, California, and by Prof. T. D. A. Reported from Moss Beach, San Mateo Cockerell on San Nicolas Island, off south- County, California, by Miler and from ern California. Quequ6n, Prov. of Buenos Aires, Argentina, by Giambiagi de Calabrese. Porcellio, genus The intention in my work of 1936 was Porcellio scaber var. americanus Arcan- to provide means of distinguishing the few geli, 1932 American representatives of this genus Porcellio scaber americanus MILLER, from each other, not necessarily from the 1932, p. 166, Figs. 1, 5, 11, 18. many allied Old World forms not known to Reported from around San Francisco occur in America. But as Verhoeff, 1938, Bay (commonly), and Carmel, California. p. 121, has commented unfavorably on the See Van Name, 1936, p. 229. illustrations I gave of this group (mostly Miller states the distinguishing charac- 19401 Van Name, Supplement to American Isopod Crustacea 137

ters of this subspecies as follows: "1. Sur- mens (both females) from Texas and face granulations more pronounced. 2. Florida, respectively. The less prominent Pigmentation greater, especially in males, lateral lobes and more even curvature of and extends to ventral side of body, not the front of the head, also the slenderer even excluding the pleopods in females. antennae of P. mulaiki are noticeable. 3. Frontal lobe of head more acute. 4. That species also has the first article of the Lower margin of ischipodite of seventh leg flagellum the longest; in virgatus the re- in male more distinctly bowed. 5. Ex- verse is the case. Moreover in mulaiki opodites of first two pairs of pleopods in the abdomen is proportionately smaller both male and female exhibit slight and shorter than in virgatus. differences." A number of additional specimens of P.

a000 13 00

I pl1i

Fig. 28. Porcellio scaber Latreille, 1804. IAfter Waechtler, 1937, and Meinertz, 1934.

Porceilio spinicornis Say, 1818 mulaiki have been received from Mr. Figures 9C, 29 Mulaik, collected at Corpus Christi, Texas. Porcellio spinicornis MILLER, 1938, p. 170, F.ig. 21. Porcellionides saussurei Dollfus, 1896 See remarks under P. s. occidentalis on See page 122 of this article. page 117 of this article. Porcellionides virgatus (Budde-Lund), Porcellionides mulaiki Van Name, 1936 1885 Figure 30B Figure 30A In order to show the differences sepa- The figures and descriptive notes (origi- rating this species from P. virgatus (Budde- nating from Arcangeli, 1932) given by Lund), the outline of the head seen from Miller, 1936, p. 170, Fig. 21, apparently above and the second antennae of both represent P. virgatus (Budde-Lund) which species are figured here drawn from speci- probably does not occur in California at 138 Bulletin American Museum of Natural History [Vol. LXXVII all, not P. formosus Stuxberg, which is mento de Treinta y Tres, Uruguay, is re- probably a synonym of P. laevis. See Van corded by Cordero, 1937, p. 9 (spelling Name, 1936, p. 244, and remarks and henseli). figures in this paper under P. mulaiki. Tracheoniscus rathkei (Brandt), 1833 Probopyrus bithynis Richardson, 1904 Figure 31 Reported by Mackin and Hubricht, Reported (as Porcellio r.) Fig. 27, from 1938, p. 634, from the Mississippi River, Quequen, Prov. of Buenos Aires, Argen-

I It carp. VII

Fig. 29. Porcellio spinicornis Say, 1818 (syn. P. pictu8). Details after Waechtler, 1937, and Verhoeff. Hillcrest, Missouri, parasitic on Macro- tina, by Giambiagi de Calabrese, 1935a, brachium ohionis (Smith). p. 509. Cordero, 1937, p. 10, records specimens from the branchial cavities of shrimps of the genus Macrobrachium bought in the Trichoniscus demivirgo Blake, 1931 market at Belem, State of Para, Brazil, and Dailey and Hatch, 1940, assign tenta- also (footnote on p. 11) others possibly of tively to this species some specimens (fe- this species from fresh water near Buenos males only) taken with examples of Ligi- Aires. dium gracile, a native species, in damp localities in Seattle, Washington. Probopyrus creaseri Pearse, 1936 Some small female specimens from the Closely related to P. floridanus Richard- Public Garden greenhouse, Halifax, Nova son, 1904. It is parasitic on Palaemonetes Scotia, received from Dr. J. C. Medcof, carolinus Stimpson, which occurs in more may also belong here. or less brackish water, but cannot be con- sidered as properly a fresh-water species. See Pearse, 1936a, p. 51, Figs. 16-21. Rhyscotus texensis (Richardson), 1905 A mutilated and incomplete specimen, A B apparently of this species, was found in a lot of isopods collected by Mr. Stanley Mulaik at Corpus Christi, Texas, March 21, 1936, and is in The American Museum of Natural History collection. A< Telotha henselii (von Martens), 1889 A specimen in the National Museum at Fig. 30. Dorsal view of head and second antenna. A, Porcellionides virgatus (Budde- Montevideo from the gills of the fish Lund), 1885. B, Porcellionides mulaiki Van Hoplias malabaricus (Bloch) from Departa- Name, 1936. 19401 Van Name, Supplement to American Isopod Crustacea 139 Trichoniscus (Clavigeroniscus) riquieri Trichorhina pittieri (Pearse), 1921 Arcangeli, 1930 The type of this species is in the Univer- NEW LoCAILITY.-Barro Colorado Is- sity of Michigan Museum, Catalogue No. land, Canal Zone, Dr. C. E. Williams, Jr., 52002. A comparison of the British many specimens collected. The body Guiana specimens identified with T. pittieri surface in this species is, in spite of the Pearse by Van Name, 1925,1926 and 1936, tubercles, quite glossy. The minute crenu- with this type would be desirable, in view lation of the anterior margin of the frontal of the brevity of Pearse's description. lobes mentioned by Arcangeli is conspictu- ous on magnification. Trichorhina barbouri (Van Name), 1926 CORRECTION.-Parentheses inadvert- Trichoniscus pygmaeus Sars, 1899 ently omitted in Van Name, 1936, p. 190, I have received a number of specimens line 29. (including males) from Dr. J. C. Medcof, collected in the Garfield Park Conserva- Trichorhinella, subgenus. tory, Chicago, Ill. The large lateral lobes See under Trichorhina. of the head, somewhat square with the Tylos latreillii Audouin and Savigny 1826 CORRECTIONS.-The original spelling is latreiliji and the authority Audouin, not Adouin as given in Van Name, 1936, pp. 409-416. Arcangeli, 1938, has attempted to deal with the perplexing forms of the genus Tylos by including most of them in latreillii, but dividing it into a number of subspecies. As far as the American forms are concerned the value of his proposed revision is greatly diminished by being based only on the literature available which is very inade- )pleM8 VIE quate, not on new American material, which will be absolutely needed before the Fig. 31. Tracheoniscu8 rathkei Brandt, 1833. After Meinertz, 1934, and Waechtler, 1937. subject can be satisfactorily dealt with. In the species latreillii Arcangeli includes not only the European and North African anterior border slightly concave, are an aid in this species. The tubercles forms, but also those of eastern America, recognizing of West Indies and Bermuda. of the dorsal surface of the body are rather inclusive the inconspicuous unless the specimen is dry. He reduces Tylos niveus Budde-Lund to a subspecies of latreillii, having (perhaps cor- no in the constancy and Trichorhina, genus rectly) confidence importance of the differences reputed to Verhoeff, 1937, pp. 422-428, gives a key exist between the two. He does not make to the known species, adding one new one it entirely clear whether or not he would (T. vannamei) of unknown origin, found in include all the eastern American records of the building of the Berlin (Germany) Tylos under the subspecies niveus. Aquarium (misspelling T. barbourii for Whether the Pacific species punctatus barbouri, on p. 425). He divides Trichor- Holmes and Gay and insularis Van Name hina into the following subgenera: should also be reduced to subspecies Ar- 1. Trichorhinella (T. simoni only) cangeli is doubtful, apparently on geo- 2. Cubarhina (T. bequaerti only) 3. Trichorhina, sens. str. (the remaining spe- graphical grounds rather than on any cies) known differences in their characters. T. 140 Bulletin American Museum of Natural History [Vol. LXXVII spinulosus Dana (see below) he regards as Tylos punctatus Holmes and a distinct species. Gay, 1909 NEW LocALITY.-Guaymas, Mexico, on the Gulf of California. Specimens in The American Museum of Natural History. ?Tylos spinulosus Dana, 1853 Figure 32 If the very large specimens from Chile mentioned in Van Name, 1936, p. 416, are correctly referred to Dana's species, there is no question that Arcangeli is correct in considering it distinct. The figures here given, drawn from these specimens, in addition to the statements in my work referred to, will make this clear. It should be added to the statements made regarding them that the lateral ends of abdominal segments 4 and 5 (especially 5), also the end of the telson, are somewhat bent out- Fig. 32. Tylos sp. from Chile (perhaps Tylos ward, but this is scarcely at all the case in spinulosus Dana, 1853). segment 3.

PART III.-ADDITIONS TO THE BIBLIOGRAPHY (Experimental and physiological articles usually not included) ARCANGELI, A. CARL, G. C. 1933. Per una migliore conoscenza di alcune 1937. Flora and fauna of brackish water. specie . . di isopodi terrestri . Boll. Ecology, XVIII, pp. 447-453, Figs. Mus. Zool. Anat. Comp. Torino, 1-3. XLIII, pp. 47-62. CARL, J. 1936. Un genere e due specie nuove di isopodi terrestri del Brasile. Arch. 1908. Monographie der schweizerischen Iso- Zool. Ital., XXIII, pp. 201-208, P1. poden. Denk. Schweiz. Nat. Gesell. IV. Ziurich, XLII, pp. 113-242, Pls. i-vi. 1938. Tylo8 latreillii Aud. e Sav. Suoi biotipi, (Important monograph.) sua area di diffusione. Boll. Mus. CHASE, H. D., AND BLAIR, A. P. Zool. Anat. Comp. Torino, XLVI, 1937. Two new blind isopods from north- pp. 139-151, Pls. I-vI. eastern Oklahoma. Amer. Midland BARNES, T. C. Nat., XVIII, pp. 220-224, Pls. I, ii. 1935. Salt requirements and orientation of CORDERO, E. H. Ligia in Bermuda. III. Biol. Bull., 1937. Nerocila fluviatilis y otros LXIX, pp. 259-268, Fig. 1. de las familias parAsitos (See remarks under Ligia baudiniana, Cymothoidae y Bopyri- above.) dae del Uruguay y del Brazil. Anal. BAYLIFF, W. H. Mus. Hist. Nat. Montevideo, (2) IV, No. 12, pp. 1-11, 1 1938. A new isopod crustaceanl (Sphaero- Figs. 1-12, PI. midae) from Cold Spring Harbor, Long CREASER, E. P. Island. Trans. Amer. Micr. 1936. Crustaceans from Yucatan. Cenotes Soc., of LVII, pp. 213-217, Figs. 1-11. Yucatan, Carnegie Inst. of Wash- BLAIR, A. P. See Chase and Blair. ington, Pub. No. 457, Part XI, pp. BOWLEY, E. A. 117-132, Figs. 1-43. (Included in 1935. A survey of the oniscoid genus Phal- Supp. to Bibliography, Van Name, loni8cus Budde-Lund, with a descrip- 1936, p. 508.) tion of a new species. Journ. Roy. DAILEY, E. F., AND HATCH, M. H. Soc. West. Austral., XXI, pp. 45-73, 1940. Records of two species of terrestrial 4 Pls. Isopoda from Washington. Amer. BRANDT, J. F., AND RATZEBURG, J. T. C. Midland Nat., XXIII, p. 252. 1829-1833. Medicinische Zoologie, pp. i-iv, DR CALABRESE, D. See Giambiagi de Cala- 1-361, Pls. I-XXXVI. brese. 19401 Van Name, Supplement to American Isopod Crustacea 141

DOLLFUs, A. MEDCOF, J. C. 1896f. Crustaces Isopodes de la Sicile, pp. 1939. On the occurrence of the terrestrial 1-6, Figs. 1-3. Paris, 1896. isopod Androniscus dentiger in Canada. EssiG, E. O. Canad. Field-Nat., LIII, p. 115. 1926. Insects of western North America, MEINERTZ, I. pp. vii-xi, 1-1035, Figs. 1-766. 1932. Die Landisopoden Danemarks. Part GEISER, S. W. I: Ligiiden u. Trichonisciden. Zool. 1934. Further observations on the sex-ratios Jahrb. Syst., LXIII, pp. 352-406, of terrestrial isopods. Field and Lab., Figs. 1-18. III, No. 1, pp. 5-10. 1934. Die Landisopoden Danemarks. Part GIAMBIAGI DE CALABRESE, D. II: Die Onisciden. Op. cit., LXV, 1933. Description compleinentaria de un pp. 211-284, Figs. 1-31. (Many il- isopodo de agua dulce, Braga fluviatilis lustrations of details of species com- Richardson. Anal. Mus. Nae. Hist. mon to Europe and America.) Nat. Buenos Aires, XXXVII, pp. 1938. Isopoda Terrestria. In: Zoology of 511-516, 3 Figs. Iceland, III, No. 28, pp. 1-11. 1935. Un is6podo nuevo "Philoscia bonari- MILLER, M. A. ensis." Physis, XI, pp. 495-496, 1936. California isopods of the genus Por- 1 Fig. cellio with descriptions of a new 1935a. Is6podos nuevos para la fauna argen- species and a new subspecies. Univ. tina. Physis, XI, p. 509. of California Pub. Zool., XLI, pp. GIBBES, R. W. Figs. 1-29. fauna of South 165-172, 1848. Catalogue of the 1938. Comparative ecological studies on the Carolina. In Report on the geology terrestrial isopod Crustacea of the San of South Carolina by M. Toumey, pp. Francisco Bay region. Univ. of Cali- i-xxiv. (Crustacea on p. xvii.) fornia Pub. Zool., XLIII, pp. 113-142, HATCH, M. H. 1-3. (Includes check list of 1939. Records of terrestrial Isopoda or sow Figs. bugs from North America. Amer. California species.) 256-258. MILLER, M. A., AND Hoy, E. A. Midland Nat., XXI, pp. in a 1940. See also Dailey and Hatch. 1939. Differential growth and evolution HOFFMAN, C. H. subterranean isopod. Amer. Natural- 1933. The biology of Caecidotea tridentata ist, LXXIII, pp. 347-364, Figs. 1-4. Hungerford (Isopoda - Crustacea). MONOD, T. Journ. Kansas Ent. Soc., VI, pp. 26- 1931a. Sur quelque Crustacs aquatiques 33, 1 Fig. d'Afrique (Cameroun et Congo). Rev. HOWARD, L. 0. Zool. Bot. Afric. XXI, pp. 1-36, Figs. 1883. A list of the invertebrate fauna of 1-24. South Carolina. In: Resources of 1937. Sur une isopode parasite du genre South Carolina, pp. 265-359. Charles- Asotana Sch. et M., 1881 (=Badroul- ton. (Isopoda on p. 295.) boudour W. H. Leigh-Sharpe, 1937). HoY, E. A. See Miller and Hoy. Ann. Parasitol., XV, pp. 465, 466. HUBRICHT, L., AND MACKIN, J. G. See Mackin NIERSTRASZ, H. F. and Hubricht. 1931. Die Isopoden der Siboga-Expedition. KATO, K. III, Part 2 (Flabellifera). Siboga- 1936. Secondary sexual character in some Exp., XXXIIc, pp. 1-111p, Figs. 1-129, legs of the male Ligia exotica. Zool. Pls. X, XI. Mag. Tokyo, XLVIII, pp. 333, 334, NIERSTRASZ, H. F., AND MAREES VAN SWIN- 3 Figs. DEREN, J. W. LEIGH-SHARPE, W. H. 1931. Sftsswasser-Isopoden der deutschen 1937. Badroulboudour splendida n. g. et sp., Limnologischen Sunda-Expedition. a new parasitic isopod from Equador. Suppl. Bd. 9, Trop. Binnengewasser, Parasitology, XXIX, pp. 391-394, II, pp. 394-402, Figs. 1-5. Figs. 1-4. PEARSE, A. S. MACKIN, J. G., AND HUBRICHT, L. 1936. Results of survey of the cenotes in 1938. Records of distribution of species of Yucatan. Cenotes of Yucatan, Car- isopods in Central and Southern negie Inst. of Washington, Pub. No. United States, with descriptions of 457, Part II, pp. 17-28, Pls. I, ii. four new species of Mancaseltus and 1936a. Parasites from Yucatan. Cenotes of Asellus (Asellota, Asellidae).' Amer. Yucatan, Carnegie Inst. of Washing- Midland Nat. XIX, pp. 628-637, ton, Pub. No. 457, Part VII, pp. 45- Figs. 1-20. (The names of the au- 59, Figs. 1-21. thors appear as above at the begin- 1938. Fauna of the caves of Yucatan. Car- ning of the article. On the covers and negie Inst. of Washington, Pub. No. in the table of contents they appear as 491, pp. 1-17, Figs. 1-8. Hubricht and Mackin.) RATZEBURG, J. T. C. See Brant and Ratzeburg. 142 Bulletin American Mutseiuum of Natural History [Vol. LXXVII

STEPHENSEN, K. 1937. Ueber einige neue unid bekaunlte Iso- 1913. (This should be the date of the Con- poda terrestria (61. Isopoden-Aufsatz.). spectus Crustaceorum et Pyenogon- Sitzungsb. Naturf. Freurnde Beirlin, idorum Groenlandiae, niot 1917 as ann., 1936, pp. 411-430, Figs. 1-22. given in Van Naine, 1936, p. 506.) 1938. Zur Kenritniss der Gattuing Porcellio 1936. A tanaid (Tanais stanfordi Richard- und ueber Isopoda-Oniscoidea der son) found in fresh water in the Insel Cherso. Arch. f. Naturg., (n.f.) Kurile Islands, with taxonomic re- VII, pp. 97-136, Figs. 1-21. maiks oni the genius Tanais. Annot. 1938a. Weltstellung der Isopoda terrestria, Zool. Japon., XV, pp. 361-373, Figs. neue Familiein derselbein und neues 1-5. System. Zool. Jahrb. Syst., LXXI, TOUMEY, M. See under Gibbes, R. W. pp. 253-264. (New classification pro- VAN NAME, W. G. posed.) 1936. The Amnerican land and fresh-water WAECHTLER, W. isopod Crustacea. Bull. American 1937. Isopoda (Asselio) in: Die Tier welt Mus. Nat. Hist., LXXI, pp. i-vii, Mitteleur opas. II, part 2, pp. 227-- 1-535, Figs. 1-323. 317, Figs. 1-121, 1 P1. (This im- VERHOEFF, K. W. portant German work deals with anid 1908c. Androniscus, n. gen. (13. Isopoden- illustrates details of a number of Aufsatz.). Zool. Aniz. XXXIII, pp. species common to Europe and 129-148, Figs. 1-12. America, and has an extensive bibliog- 1933b. Zur Systematik, Geographie u. Oeko- raphy.) logie der Isopoda terrestria Italiens u. WEBB, W. M., AND SILLEM, C. ueber einige Balkan-Isopoden. (49. 1906. The British woodlice. Essex Natural- Isopoden-Aufsatz.). Zool. Jahrb., ist (Stratford, England), XIV, pp. Syst., LXV, pp. 1-64, Pls. i, ii. 38-56, 81-111, Pls. I-xxv, Figs. 1-59.

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