A, Supplement to the American Land and Lresh-Water Isopod Crustacea
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.,I .4 m A, Supplement to the American Land and lresh-Water Isopod Crustacea BY WILLARD G. VAN NAME BULLETIN THEIAMERICAN MIUSEUM OF 4NATURAL HISTORY Vo LXX II. ART. ii, pp. 19-142 New York Issued June 20, 1940 m ImmIIIIIIIIIJ. II II 1. r.m 11 A1~~~~~~~~~~~~~A ~~~~~41~~ ~ ~~~~~~u -K 1,4k 1 1 X 1 'Y4 AIJl~4 14 Y7 -Vu1I4 Article II. A SUPPLEMENT TO THE AMERICAN LAND AND FRESH- WATER ISOPOD CRUSTACEA BY WILLARD G. VAN NAME Figures 1 to 32 During the period that has elapsed since water forms. There are a number of addi- the publication of "The American Land and tions in the family Asellidae, which has Fresh-Water Isopod Crustacea" in May, lately been the subject of an increased 1936,1 a number of additional species have amount of study. been added to the American list and new Trichoniscus (Androniscus) dentiger Verhoeff, information published regarding many 1908 Philoscia bonariensis Giambiagi de Calabrese, others. It therefore seemed desirable to 1935 prepare a supplement to that work to cover Neotroponiscus carolii Arcangeli, 1936 the additional species, correct errors and Platyarthrus hoffmannseggii Brandt, 1833 omissions and record changes in nomen- Porcellio spinicornis occidentalis Miller, 1936 clature and notes and references of im- Porcellio dilatatus Brandt, 1833 ?Porcellio ragusae Dollfus, 1896 portance on other American forms. Porcellio littorina Miller, 1936 This article is to be regarded as purely Circoniscus p1alidus Arcangeli, 1936 supplementary to my work of 1936 named Nerocila armata Dana, 1853 above, and is designed for use only in Asotana splendida (Leigh-Sharpe), 1937 Exosphaeroma platense Giambiagi, 1922 connection with it. No attempt is made to Asellus montanus Mackin and Hubricht, 1938 repeat matter that was given in that work. Asellus dentadactylus Mackin and Hubricht: The present article consists of the fol- 1938 lowing three parts: Caecidotea macropopoda Chase and Blair, 1937 Caecidotea ozarkana Chase and Blair, 1937 I.-Descriptions of new species and other Mancasellus ouachitaensis Mackin and Hub- species added to the American list. richt, 1938 II.-Additional notes and references, Mancasellus louisianae Mackin and Hubricht, changes of names, corrections, etc., apply- 1938 ing to species dealt with in my 1936 work. Through the kindness of Dr. E. C. III.-Additions to the bibliography (in- Williams, Jr., of Northwestern University, cluding works omitted in 1936). I have had the opportunity of examining The following four species are described a large collection of isopods recently made as new to science: at Barro Colorado Island, Panama Canal Trichoniscus (Miktoniscus) medcofi, new species Zone, but failed to find any undescribed Philoscia floridana, new species species. I am also under special obliga- Philoscia avrilensis, new species tions to Dr. J. C. Medcof, of the Canadian Exosphaeroma insulare, new species Fisheries Research Board, Mr. Stanley The following species and subspecies Mulaik of Edinburg, Texas, and Prof. described by other writers are added to the T. D. A. Cockerell of Boulder, Colorado, list of known American land and fresh- for interesting material sent me. PART I.-DESCRIPTIONS OF NEW SPECIES AND OTHER SPECIES ADDED TO THE AMERICAN LIST SUBORDER ONISCOIDEA, LAND ISOPODS Trichoni GENUS OR SUBGENUS ANDRONISCUS European species and races, the females of VERHOEFF, 1908 which are practically alike, though the A division of Trichoniscus in the com- males are distinguishable by characters of prehensive sense composed of a number of the pleopoda and legs. They are remark- 1 Bull. American Museum of Natural History, able for their coloration, usually a striking LXXI, pp. i-vii, 1-535, Figs. 1-323, by Willard G. pink or orange-pink, in life. (See Ver- Van Namne. hoeff, 1908c.) 109 110 Bulletin American MusBeum of Natural History [Vol. LXXVII Most of them have been included under numerous densely crowded tubercles ar- the name Trichoniscus roseus (Koch). The in true ranged transversal rows. Cephalon roseus was described by Koch (as Itea transversely oval, with the lateral lobes rosea) from Bavaria. front Among the most distinctive well-marked, denticulate, nearly character- straight. Lateral parts of mesosome more istics of the males of this group are the expanded than in the other peculiar modification of the merus of the rather species, 1st seventh pair broad and partly flanking the legs and the spoonlike termination cephalon, the 3 posterior recurved of endopodite of the first pleopoda, both pairs and acuminate. Metasome (in male) ex- of which are well exemplified in the illus- ceeding half the length of the trations of the species that follows. and mesosome, but little more than half as broad; terminal expansion of last segment trans- Trichoniscus (Androniscus) dentiger versely truncate, with 4 small apical spi- Verhoeff, 1908 nules. Antennulae with the last joint much Figure 1 longer than the 2nd, and carrying 6 sen- Androniscus dentiger VERHOEFF, 1908C, P. sory filaments, 5 apical and 1 lateral. 139, Fig. I0.-VANDEL, 1933, p. 131.-WAECHT- Antennae comparatively slender, attaining mx Fig. 1. Trichoniscus (Androniscus) dentiger %Verhoeff, 1908. After Sars, 1899 (T. "roseus"). LER, 1937, p. 258, Fig. 50.-MEDCOF, 1939, P. 115. 1/3 of the length of the body, flagellum Philougria rosea KINAHAN, 1857, Nat. Hist. composed of 4 articulations. Last pair of Rev., V, p. 197, P1. XXIII, fig. 3.-BATE AND legs in male peculiarly modified, having WESTWOOD, 1868, Hist. Brit. sessile-eyed Crust., the meral II, p. 460, Fig. joint strongly dilated, and pro- duced inside a Trichoniscus (Androniscus) dentiger ARCAN- to large conical prominence, GELI, 1931a, p. 12. in against which the succeeding joint Trichoniscus roseus BUDDE LUND (in part admits of being bent. Inner ramus of 1st only), 1879, p. 9; 1885, p. 247.-SARS (in part pair of pleopoda in male, with the terminal only), 1899, p. 163, P1. LXXIII, fig. 1. dilated at the "Body oblong oval, joint end in the form of a greatest width al- little bowl of a spoon; that of 2nd pair most attaining half the length, dorsal face rather but strong and distinctly 3-articulate, slightly convex, and rough, owing to terminal joint knife-shaped and denticu- 1940] Van Name, Supplement to American Isopod Crustacea ill late inside. Uropoda with the outer ramus in the belief that it would be found in about twice the length of the basal part, Norway also. inner ramus somewhat narrower, but The first American record of this species scarcely shorter. Color of dorsal face in is due to Dr. J. C. Medcof. He reports it life of a clear minium-rose. Length of (Medcof, 1939) as occurring in many adult male 5 mm. greenhouses in the region of London, "This species, first described by Koch, Ontario, and Toronto. Fig. 2. Trichoniscus (Miktoniscus) medcofi, new species. is easily distinguished by its comparatively Trichoniscus (Miktoniscus) medcofi, broad and flattened body, the dorsal face new species of which is densely granular, and by its Figure 2 beautiful colour, which, however, is only Body proportionately narrower than usual in seen in fresh specimens." (Sars, 1899, pp. the Trichoniscidae (ratio at least 4 to 1 if the 163-164.) intersegmental muscles are relaxed), the back This is the common, if not the only, considerably arched transversely, its surface with scattered short hairs and many small scabrous representative of the roseus group in tubercles which form at least four more or less northwestern Europe (British Islands, Bel- definite transverse rows on most of the thoracic gium, Luxemburg and parts of France), segments. The tubercles are very conspicuous including, at least in greenhouses, localities on the head and anterior segments becoming less and prominent toward the rear but do not disappear in northern Germany Denmark. Sars, entirely even on the abdomen. The frontal 1899, included and illustrated it (under the outline seen from above is convex, with fairly name roseus) in his Crustacea of Norway large lateral lobes extending obliquely down and 112 Bulletin American Museum of Natural History [VOl. LXXVII forward and having a rounded anterior margin. basal joints of the uropoda are short and have Eyes composed of a single large ocellus sur- the insertion of the inner branch removed by a rounded by a somewhat larger area of black small lateral interval from, and somewhat far- pigment. Antennae with a flagellum of four ther forward than, that of the outer branch, distinct articles and a terminal group of bristles. which is considerably the longer and stouter. Thoracic segments I to IV inclusive, with the The largest example is a male measuring rear corners rounded (IV only slightly), V to somewhat over 5 mm. long when the inter- VII have the rear corners quite acute and in- segmental muscles are well relaxed. creasingly extended backward. The legs are all The specimens, except very young ones, are quite long with well-developed spines. Leg VII strongly pigmented, having the upper surface of the male has the merus widened posteriorly handsomely variegated with brown or purplish- and compressed from side to side. brown pigment on a light background which is The abdomen is abruptly somewhat narrower whitish or yellowish in the preserved material. than segment VII of the thorax, and tapers There is usually a narrow dark streak across the considerably. The rear corners of segments 3 forehead extending through and behind the eye to 5 are extended back into sharp points rather on each side. closely appressed to the following segment. The pleopoda of the first two pairs of the male LOCALITIES.-Native habitat unknown. conform quite closely to Kesselyak's (1930) de- Dr. J. C. Medcof, for whom the species is scription of the type species Miktoniscus linearis. named, kindly sent me about 40 specimens, Fig. 3. Philoscia bonariensis Giambiagi de Calabrese, 1935. After that author.