'8 891 D(rlrolsotlJ!4 d '1305 L9I a?qpdg t s9l Dua$auot d L 09r DraS!ru ds ,9 d 'I L9l qDqdaJonw d 'n 991 DID^OI)d n9l Dqlldopq ',J 't z9l nptrlttdas 'd ,z 6rl DalaulJ '1J0S z 'I 8tl oumBIS d 8il soraqluoaaH 'I 'lJOS ZNI Dalawq I WI luarxlBeJJctl€tuelsfs WI ,(qdaSoeSotg 8€l uIelr(s SurpeoJg 8€l lgqunN auosouolqJ 8€I lrnJd Ltl t3,r\olg 9EI slocrpedpu? ecuorsotouul 'slxals 9El slc?Jg[?r3nlo^q pue se^eo"I ,€l unluolunpul pu? llqeH ,EI ?J3U0Ceql JOssrlsrJolr?JeqJ uO soloN E'I uotlesrJrss?lJluosald eqJ ztI spoqleN pu? sprJel?ntr I'I saqluoraqJ pue oapuq Jo ,{rolsrH IfI Ielou0c III uorlcnpoJlul 6ZI 1J?JlsqY aAsd slualuoc 'saqeu€,{ o,1AlpuE sortedsqns oelql loJ epeu o& suortEurqutoaA\ou pu? .neJI pequ.sopoje sorcodsqns^\ou a^lC ep€u ole en4lorafu\qns dsqnsq.2 o1|{qdopo uo pxeq t,lU (neftajq! 'z.tnlqpuowuup 'ata 4 oralul^.qns d pue o.Eatsold,qrJ uo prseq qittnlj ipuowutrup A suoqeurquo, Mou aql o,{U qrssas pue .eil .d ,ei\ .d 'etd .d 'eid ,el]{ A a^d suapuad d olltldotD| Dtocutu,B DeaD suarDD d sauauoto d, :peqwry ele serads l$u 8u^\olloJ eql pue ep€rusr a,{U Balsorcol .t}r-i lolleurqluoc ^\eu oql 'peqFa6"p fzclnJ)-tras.Dapw[ { dl,t\eo aJEoIU sfr/rrtsodlrs' pes pue ail soraqluoolaq tJr Wnwter ore o.reqpxru8ocaJ s cedsu€q?Ilsnv ute$aA{ ftlul?ruel oql pue snuo8e sEpalE}sulor 'saqlunaqJ St sr'sdcedi u€llE4snvulets€ld oelql qtl{ paluosordsDalar,/Hf muo8 aql-roj uoeEcrJrss.elrJutua8ejlul ^\ru p puEposr^al el! eseceeelou,{qf -Ig,e,{u u?rle4snvurolsold (gg6ll gLz.6zl:\zlgetst{nN JsrcEaplrLuiqluprlplrsnv uratsJ/A Jo uorsr^oJV

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ISI9'e{€Isnv ul?lsolA'qlJod qlnos,loarts oBJooC,ornuBqloH ue{ellsnv LualsolA orutg aBar?aslauql usll€rlsnvurolsal[ Jo uols!ar v

6al (886t)8/U.6Zi 1Z)9 ersr{nN 130 NuytsiaVol. 6. No. 2 {1988)

9. P. argentea 169 10.P. micrantha 173 sect.4. Calyptrostegin 175 ll. P. ammocharis 175 12.P. grcniticola 177 13.P. imbricata 178 14.P. subvillifera 185 15.P. villifera 187 16.P. erecta 189 17.P. sylvestris 190 18.P. calcicob 193 19.P. longifora 194 20. P. preissii 197 2l. P. angustifolin 199 22. P. floribunda 201 23.P. sulphurea 203 24. P. pendens 205 25.P. aeruginosa 208 26. P. cracens 209 27. P. tinctoria 212 28. P. suaveolens 214 29.P. drummondii 219 secl.5. Macrostegia 222 30. P. physodes 222 sect.6. Stipostachys 224 31. P. hohoydii 226 sect.7 . Heterolaena 227 32.P. lehmanniana 228 33. P sessi/rs 34.P. rara 234 35. P. spectabilis 236 36.P. Iercantha 238 3'1. P. avonensis 239 38. P. brevistyla 240 39.P. ciliata 244 40. P. rosea 248 41.P. ferruginea 251 42.P. hispida z) 1. 43. P. Ianata 253 44.P. brevifolia 255 45.P. brachyphylla 259 2. Thecanthes 262 1. T. conueta zoz 2. T. punicea 264 3. T. sanguinea 267 Discussion 269 Generaand Sections 269 Phylogeny 270 FutureStudies 270 NominaNuda probablyapplied to WesternAustralian Taxa 271 NominaDubia 271 'sttlzoiwoqJ puz 'stqrolsttow 'aSollDdg'ouaqo ,{qd 'ouaqoJalaH porueuEuroq sdno$ Suureuloraql puesaqiuoJaqJ'oat olurt\ds se,r\uorlc'Js tsJ$ s,u,norgleql ldecxa(0181) u^\org,{q ua^6 suorpesaql qll,r papuodsolocsdnoJE sseqJ IUBJ perycadsun sdnoJSJueua8E4ur xrs posruEocalpu" oapruH qlJ1(saqluo1aqJ pe:.JlqwoJeJ ;o 'snuet (rt8l) reqcqpug ,rou eq1ol ralaa..rdlopun poqursopuooq ^lsnor^erd p?q l"ql salcods o,{rlperaJsu€rl pue saqiuoJaqJsnua8 eql poquosep(818 1) ruorts{r^\ Jal?'I uortJosqc?o 'EX?tJo uorlducsop € 3ur^r8pu? suorlrospeu?uun lnq psrequnu oArJur uaql SurcEId Jauq'sonods tt posruSoJeleH uellellsnv Jo tuorulBol lBquBtsqnslsrrJ oqt paqsrtqnd(0l gl) uaojg 'olotlsotd '88/l g selc?d:spuBI?aZ ,rr{aN aql uo pesBq ut porx€usp^^ rJUtroBC xo IoS? s{uBg Dapwld Duuassodsnuo8 ol?u[u?]s-lq3re ,{ ?ur8uoaql urqll,r Erl"rlsnv pue pueleoz,rsN uro4 o?ocBeBlouqJ al?Uuxsls'o.rleqlJo uorsnlcuroql poldeJcEsJoqlnE,ral fJeA'ro^e,roH '(66 :0t61) en3?rds^q pe^rosuorfll8urJoJ sE,aA tl pu? DreluDge:uJeu,rau eqt palderoB ''I J.I sJoqlneluenbesqns tsol l DuuassDJsnue8 peqsrTq"lsa ,{peorl€ oql ot Dleluog se u.rou{ ^lmohord mue8 oql polroJoJpuE esrcealord aqt u mueE€ $J DtsluDgeweu eql pesnel(28fl) 'J sna"uurl lnq retsroC C ry relslod ot$luog se S//t ur poqucsepse,r snueEoqJ o8e.{o^ puocoss,{ooJ uo JotsJoCC f pue Iq I ur palrallor serJJdspuplBoz ,{\aN eolql U f Z/l 'saquoJallJ Jo ora,r adorna ur o^rrJe ol Dapaw lo suourcedslsJrJ oqJ puo Dapurd lo fuolsH 'pesruSoJol 'a?OJeo?leu.4qJ are sarytuDJaqJJo seorJs oaJr+pueDapluld Jo sanodsSt qcrq,r u u?rlalsnv urotsolA uorsr^eJiruouox?l B sluesoldlod?d srqJ ecutssoroeds u?llellsnv Jo '(t861SI6l uralseA\cruepue JoJ paqsrtqnd uoeq o^€q seueu a\ouou efd) uorSouquad aqt Jo €Joldaqt JoJuoneJedord ur poqsqqndseuEu sorJods naou JnoJ urog UedV (€/gl) uBqluog Jo l"ql sr erlBJlsnvuJolsed\ ur eem?a?laur^r{Jeql Jo elBp ol lueuJlearlelrsueqadruoc lsouIoql tnq slleJlsnv uretseiAolul pualxel?qt $rcads,roJ e sapnlourtuoruteert srqJ (Eg6l poqsllqnduaaq fllumar sBq?uotcr^ pu? solel[ qtnos ,reN 'pu€lsuaenb'ert€rlsnv IlqlerqJ 'lJes rllnos rJJo+\saqtuocaqJ Dalauld se)saq iunaqJ puBDapluld lo saraadsJo uorsr^elV '-l ',(lrrueJ Dtptut ^q [Vo ilqeqo:draqrunu serceds u pepa"Jxo eql ul ?louo8asJo^rp lsoul'tse8J?l eql auo Dapwv (l arn8rC)seulddrJrqd oql ol qllou Supuetxopu? Brl?llsnv ulaqllou Jo 'pu€lsJ u Suurnooosabeds aArJ q!^\ saqiunaqJ tu?qleqJ ol lseoSurpuelxe puE ?rl?llsnv lnoqEnoJqllsoule Suuncco sercods001 ro^o qlr,r oapwld'e\er$ny q pat?rluocuoJ er? eJeueaqlog'oBspruC oqrJturqth\ e?urelaurdoqrllqns sjequetu,{luo aql elB'(tt6l) 'pu" Jo 'suaru?ls e{uoc JouonecrJrss€lc eql uI o?eprooBlou,{qI^Iulqqns ol Suolaq,{oqJ lnoJ 'guo ls?ell? 8ur^eqErauaE roqlo [? ,{lorer,{re^ (Dapuw ur) Jo o,r1Jo laqunu uauets lroql ^q ,{lrur?Jeql Jo sJequeu leqto uro4 poqsrnEurlslp,{llppol el? saqiuoraqJpue oapwld

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IfI eEoceoelour^r{IusrlEllsnv uJelsry'{'a,{d -J.g 132 NuyrsiaVoi. 6. No. 2 (1988) Three new genera,Gymnococca Fischer & C. Meyer, HeterolaenaFischer & C. Meyer and Cal)ptrostegiaC. Meyer.which togetheraccounted for a largemajority of the species previouslyp\aced, in ,were named in two publicationsof 1845.These and a few later publications resulted in numerous recombinations,particularly in the large Calyptrostegia.Infrageneric groups, still ol unspecifiedrank, were recognised by Meyet (1845) under the generaGymnococca and Calyptrostegia.In the latter genushe includedtwo of Endlicher'sgrotps, Malistachys and Epallage. Presumably the new genusllererolaena was alsomeant to conespondwith Endlicher'sinfrageneric group Heterolaena. However, Fischer andMeyer (1845) did not acknowledgeEndlicher and the only specieslisted, -&znelea specmbilis lasHeterolaena spectarllls), had not beennamed at the time Endlicherlisted his speciesunder the inftagenericgrotp Heterolaena.Hence the genusand inlragenericgroup of the same nameare regardedas having different types.

Another new genus,Macrostegia, was describedby Turczaninow(1852). It consistedof the singlespecies M. erubescens,which had shortly beforebeen named Hook. Although P physodesis a very distinctivespecies, Macrosteglo was neither accepted as a separategenus nor usedas an infragenericcategory by any later authors.

A revisedclassification by Endlicher(1848) retained Glmnococca and Calyptrostegiaas separategenera. However, Heterolaeno Endl. and Phyllolaenawere treatedas groupsol unspecifiedrank underhmelea whtle Thecanthesand Choristachyswere treatedas groups uji'derCalyptrostegio bLrl with Choristachy,sat a higher level than Thecanthes.Meissner (1857) recognisedonly.Prmelea al lhe genericleveland listedthree sections, Thecanthes, Eupimelea andGymnococca. Under sect.Eupimeled he listedeight groups of unspecifiedrank, consisting of the lastfive of Endlicher'soriginal groups and threegroups of his own.

Subsequentauthors have followed Meissner in recognisingPlnelea as the only genusin thiscomplex. However, Kuntze (1891), who consideredthat theolder name Ba.'lksra should be usedrather than Pimelea,published recombinations for mostof the then known species of Pimeleafollowed by a new classificationof infragenericgroups under (Kuntze 1903).This clasificationwas essentially the sameas that givenby Bentham(1873). Bentham's classi{icationis comparedin Table 1 with the more recentclassifications provided by Gilg (l 894)and Threlfall (1983).Gilgk classificationdiffered from Bentham'sin that Thecanthes wasrecognised as a subgenusrather than a section.The treatrnentol Thecanthesas a subgenus wasfollowed by Domke (1934)in his monographof the Thymelaeaceaeand by Ding Hou (l 960)in Flora Malesianabut not by Threlfall(1983) in her revisionof the speciesoccurring in four Australianstates. Threlfall followed Bentham's classification except that sheregarded oneol Bentham'ssubsections under Calyptrostegia as a separatesection while not recognising the other two subsections.

Nomenclatureat the specificand varietal\evels rn Pimeleahas beencomplicated by the publicationof many new namesbased on cultivatedmaterial. Cultivation ol Pimeleaspectes in Europeappean to havecommenced in 1793with the easternAustralian species Pimelea lini,folia.By 1860,about 20 specieshad beencultivated, including the following Western Australianspecies: P rosea,P. ferruginea, P. clavata,P. sylvestris,P. hispida,P. IongiJlora, P. lanata,P. imbricata,P. spectabilisand probablyP. floribunda. In most casesdried specimens werenot referredto when new nameswere published for cultivatedplants and often there wasalso no illustrationavailable to nominateas type.Most of thesenames are regarded here as nominadubia. Materials and Methods The presentclassification of WesternAustralian into generaand sections wasderived by examiningspecimens of all Australianspecies and by makingan assessment 'o8alsottoTy aJEstq)usodtis lces pue sooqiultaqH ;nes DualpnjaH pue sfq)D:tsodtts 'mEa$oadtpJ 'a8olodg 'oa1aq4 'sofiqjuonjaH :posruSooorsuorpas ue^es SulaolloJ oql qtlr 'pept^oJd DalawrdIo uortBrrJrss?i3pasl^er e pue ejsq pelEtsurarsr saqiuDJarlJ snuJB eqf uop?JulssslJluasard aqJ 'oseeler ^ uellooJo 0i81seql ol asolsu3q,r urldsJp ,{llsnsnere,r suouetslenpl^rpur puu sra,roJJalEu alrq,r pauesxe,{llnJse,r ol,{lsoqt uar{,r a8€tsJqt re u,\\?rporo^\ sle,^Aou al€rual pu? lenxasrgpa8r€luo ara,r sl.redlerolJ lo sro^\ou Fnpr^rpuJJZrs oJrl le u,r?lpse,r qcuerq SunrnrJ ro Suua,roL]E'petErtsnll uoxet qo?e roC 'peprocer uoaq s€q uoxet aql qcrq,r ur eoJBoorSop % ,{q oer8ep% qceeSurlEorpur 'paJederdsE,r d€ru uorlnqutsrp pumas € '?rTerlsnvuletselA aprstno Surpuolxo uoxel qJm roj 'patrellor uooqpeq uoxel oql qJrq,r uorJ EeJ?apntfuol oefep % fq epnlu?laal8op % qcse luasarderot Joqu,{se Sursnpalold s?,r BrlprlsnvulelsalA ur uox?tqJEe Jo uortnqutsp aqf 'alu ^q uaosqdeJSotoqd aqt dluo pue pa,rorroq tou s?,r uaurrcods0d,41 eql ssolunluouleorl Jrl€luels,{saqt ur €xet eleudoJdd€aq} Jspun polectpur10u sr HIUEd ur sqdP$oloqdJo ecuosold oql Hlugd ur peSpolueeq a^sq ?u?qroq sEesle^oro atEtsrelurrxo4 pe,roJloqsad,{t llE Jo sqdBrSolor{ded,{loloq e aq ol peunserd sz,r ueurrceds]eq1 Surtr-rrApu€r{ s,roussrol,.\{ oloq uatul3adsauo,{luo areql& rurq,tqpolelouue 'suourceds uoeqpEq ueurJods euo uEr{tororu Jr uasoqrse,r od,{lolcole ssrorduroJJ 198l ol roud Jeussrat\i,{qpequjsep pxet loj ed,{totJels s€uasoq3 s?,r uaur3eds el?ual oql'saroeds snorooorpe Jo uorlJollo,ad.{l aqt ur sueulcodsalEual pue elpu qloq ers,ri\ersql areqd\ 'srato?l?qc luelsuo3eseql teoder lou op dno$ qcm uqtl ^ sercedseq1;o suoudrJ3sep 'dnoJB lBnpnpul oql sratc?reqJlu?tsuo. eql sa^l3,{JUsrur.rd uorlJes ro snua8qJBa uoudrJcsapeql 'uonnodolJo Jo Jsrruruu oJ sleqeluou.rrcods oql uo uo^r8uolleuroJul ruolJfured pup lellel€u peup,{lluecer ,fue^ ro qse{ uorJ,{llftd pourujJotopolo,^^ rnoloc ro^\o[ pue JnolocDe]q tnq luuoteu par.rprxo4 ,(luopolou are.r Jnolm JEel pu€ JnoloJuals FUalBu ^JpuoJJ uelEt aro,r sluetxenssorxllv elerlsnv uratselAur uoxn aqtJo e8u?l I€crqdeJSoo8 pu€ IE rSoloqdroueql luesordarol pepr^ordsr uortJeios? asr,rJoqlo palrcoJB Ile'uoxel JelnarupdE roJ pauruexo s?,,t\suauiDeds Jo rsqurnulleus e ^luo oJor{l(\erl?rlsnv uJatsalAur petcallocsuourrJods uo ,{larrtuepes?q aJe seuo8et?c ar}roeds?rJur puE s3rcods suortduJsoc y,t\n pu? 'JN ',\\SN'tgl 'yNO'!IDl'DgJ'SNVJ'Idfl '4S'/d11V'11vJo AN I'Ct') :?u€qreq8ur,,$o1oJ aqr uorJ pe,roroq aJe,r{epar?eBleu ql Josar3ods u€rl€rlsnv urelso^\ Jo suaurJadsunu€qraH 'peuJJuoJ sJrJJdso,rt Jql JJpunpJssncsrp erE IUEJ I?lauE,\ Jo ecroq3eql roJ suoseargqJ pesruSooeroJ? serlarJe^ qalq,r ur saroedso,rl ar? eraql lnq seliodsqnsse peleorl eJe ?xEl JrJrcodse{ur soseJ lsorx uJ la^olsortods oq} spJ€.ldolecuo8Jo^rp ,{rBuortnlo^aJo aar8apJorlSlq B tuasordaJ,{oqlt?qt ut ,{ldurrssortou€^ ruorJ SuuaJJlp sP paprEBeraJp sar3adsqns 'l€uelelu qsa{ uoueuruexopatrurt dq paluoruolddnssueruroeds Jo '.lo^ol unueqreq Jo ,{Soloqdrourssor8 oqt lo uortEurruExoaqt uo peseqera,r cuoue8oql ol le^ell€teu?^ eqt uo{'uox€l qcealoJ {ueJ Jrrllouoxq etpudordd€tsou aql uo suorsrJe( '^llBcrt€ulats.,tspatsrl osl? aJe serlauB^pu€ sarcodsqnslueulEorl 3rlB[uols,4seql ur poletlpuror? uoxel qceo serlrurJJe 'se Jo alq€qordaqt l€lar lsesoloflr eq ot pareprsuocexEl aql eprsaquorloos ro sorJeds,{lo^e ac€idot e3uenbasr?eurl p ur oJqrssodlou srtr teqt uolqoJdaql eruorJa^o,{l ed oJ eruenbos e^rsueqerduoceqt ur uorlrsodreql ol EurploDpparaqunu el€ ledEdsrqt ur poquasopsorJads pu? suorlceseqJ dn u,r?Jps?,r 'sa^rlelortsesolc p3urnserd rraqt ot Jlqrssodsp,{losol, se poreldere,r sercodsqlq,r ur 'sol3odsu€Jlellsny aql loJ oJuenbescuEuetsfs rBaurl 'elnlElalll Jo ll? V alqBlr€,{perfl uJoJ!oapwV lo sarcodsuerl?JlsnV-eIXo Jo sorlsuolr?leqo rol?ru Jql Jo

tt I J?o:rea€lalu,\qf ugrlBrlsnv uretsJi( ald l g 134 NuytsiaVol.6, No.2 (19881 new whtlesecl. Macroslegra is a new combination.Table I outlinesthe relationshipsbetween thesesections and the groupsused in severalearlier classifications, further detailsbeing provided under the descriptionsof eachsection in the systematictreatment. Table 2 givesthe main characteristicsof thesections as recognised here. All of the sectionsare reprcsented in Western Australiaand all WestemAustralian species can be readilykeyed to their resp€ctivesections.

Table l: Comparisonof the more recentclassifrcations.

Bentham1873 Gilg 1894 Threllall1983 Rye

sect. s\Jbgen.Thecanthes sect,Thecanthes 2. Thecanthes sobgen.Eupimelea L Pimelea seu. 1. Hetemnthercs sect. Autopimelea sect. Pimelea secl. 2. Pimelea secl. Dithalamia sect. Dithalamia sect. Dithalamia sect. Heterolaena secl. Heterolaena sect. 7. Heterclaena sect.5. Macrostegia sect. CalyDtrostesia secl.Calyptrosrcgia secL Calyptrostegio seCI.1. Calypuostegio srbse€t.C a I y p'tr idi un |. Col)iridiun s]]bsect.Phyllolaena 2. Phyllolaena srbsect. Chotistachys 3. Choristachys sect. Chotistechys sect. 6. Stipostachys sect-Malistachys sect. Malistachys sect.not menloneo sect. Epallage sect. Epallage sect. 3. Epallage

Notes on Characteristicsof the Genera

Habit and Indumentum.Thecanthes is comprisedof annualherbs up to 0.8 metrehigh. AII speciesof Thecanthesare completelyglabrous except for onefrom the Northern Territory, whichhas simple hairs on the basalportion of the floral tube.As far as is known,all species arestdctly annual.One floweringspectmen of T. puniceawas observed to havethe remains of old flower heads,but thesewere probablyproduced earlier in the sameyear rather than in the previousyear's flowering season. Certainly this sp€cimenwas much smallerthan many otherspecimens which showedno signof previousflowering. However, T. puniceahasbeen collectedin flower in all monthsof the yearand may be capableof survivingfor morethan one year when conditionsare exceptionallyfavourable.

Pimeleais a much more specioseand more variablegenus than Thecanthes.It contains plantsvarying in habit from annual herbsand prostrateor dwarf undershrubsto very tall shrubsor eventrees. Some specimens o'l P. lanatamay be largeenough to be regardedas trees,the greatestrecorded height being about 4 metres.Pimelea trichostachya, which is one of the very few annual herbs in the genus!occun predominantlyin arid or semi-arid environments.Most specieshave axillary tufts of hairs,at leastin the axilsof immatureleaves. Leavesare sometimeshairy and stemsmore commonlyso. In most taxa the flowen have at leasta iew hain. Although the hairs arealways simple, they are very usefulfor distinguishing the taxa becausethey vary greatly in distribution,number, orientation,size and other characten.The variation in the indumentumfrom the baseof the floral tube to the apex of the sepalsis particularlyimportant for identification.

While mostof the shrubspecies appear to be readilykilled by fire, a few speciesare able to regeneratefrom swollenunderground parls. is a prime exampleof a fire-tolerantspecies. Its undergroundstock is elongate,extending well belowthe surfaceof the ground,and producesmultiple stemsafter fires. A few speciesol Pimeleaare able to reproducevegetatively by adventitiousroots from the nodesof prostratestems but noneof the WesternAustralian species appears to have this capacity. : t k R a !)- a 3 SiEl $a r"8 q-*e + $-i.5-;-A !" .d Eei-i. =a- =E- qsdz= eqd F f F g g Fg i6=F+5 * r* * * a*'g* <2 d F ;-8 E: EEi : E; 'a{E -x A i=::)zv 6 466"-;

'o ==7e gr *E ;t "=E *€ cg=' g+i;g BE e s E a;x=eF a 'eE - I a [ 8= ;*:q a -< € E3FE3g n >. i e€€u*. *E a o:e

6n €4 = €o

e E+ =? ? t2 r eiB f{3 e <-.a& :5FP P eE E r8i :=: !1 Z- z1 zZ V; =?"-.; 2i 7+;*E!. ;; a 48 "s?=t; J. ni =_'*-E p= + 32. + ?+ '*i 'rCt s.d. ;= -8e =9 z=.t 5"L *= e i5 =;:r 1:- R = *o3_ =k Tl :l e is F_q6*5E 3€

E7 E= i e signE_= + s E iE:; ; ; i i'gtt iigr"iiel[i; gf aEF fli?B aiF3iAFs giicFai n'i t3? !€Eg 3F :ies:i;e Ei ;;1;'iiiti 7= 6p.: Ftf' d igg " ilr :1Qt6 r" -?1 H! a' I s€l eeaceoqeu,{qluBrlp4snv uralso/A 'a,{U 'I g i36 NuytsiaVol. 6. No. 2 {1988) Stems,Leaves aruI Involucral .In commonwith the family as a whole,pimelea and Thecantheshave simple entire leaves and the shrubspecies tend 1o have very stringy bark. Nodesare prominent in mostgroups. Leaves are oppositeand decussatein Thecaniiesand most speciesof P[rt elea,the remarningspecies ol pimelea having altemate leaves.The is very short and mmmonly inconspicuousor rarely absent. pimeka sessilisis notablein Lavins uniformlysessile, somewhat stem-clasping leaves (Figure 53A,B). The leafmidrib is promineni on the abaxial.sudaceand usuallydistinct adaxially. Although the intra-marginaiveins are sometimessimilarly developed, only a few species,in particularthe eastemAustralian species Pimelealigustrina, have distinctly reticulate-veinedleaves.

Many Pimeleaspecies have adaxiallyconcave leaves, which sometimesappear to be mmpletely flat after they havebeen dried and pressed.some specieshave recurvedleaf margins and drying sometimescauses the marginsto becomemore recurvedor revolute.In oiher speciesof Pimeleaand. in Thecanthesthe leavesare flat.

All speciesof rhecantheshave four involucralbracts while in pimeleathe bractsrange from one to numerousor are absent.Involucral bractsin both generaare usuallybroadir than the leavesand frequentlymore colourful but in somepimelea spectesthey are very similarto, or intergradewith, the leaves.Throughout this treatment,thi numberi givenfoi involucralbracts refer only to the sessile(rarely subsessile) bracts irnmediately surr-ounding the llowers and excludeany rather similar but shortly petiolatestructures that mav occui shortlybelow. The latter arereferred to as'bractlike leaves'.characteristics of the involucral bracts,including their number, colour, size and hairiness, are valuable for distinguishingspecies and sometimessections.

InJlorescenceand Pedicels.rn Thecanthesthe inflorescenceis terminal,erect and head-like, with a concaverec€ptacle and numerous pedicellate flowers. The pedicers,illustmted in Figu; aredoniventmllv compressed ]5C-E. with an apicalallicurarion point. rney tenato ue toi!"i tnan tn ftmereoand are glabrous.As illustratedin Figure 75B, the four involucralbraits appearto form a continuationof the receptacle.Bentham (1g73) and other early authori regardedthe structurebelow the four lobesas the connatelower portion of the bractsbut Threlfall (1983)pointed out that it bearspedicels rhroughout its lengthand should therefore be regardedas a concavereceptacle. The inflorescenceof pimeleais basically racemosebut is occasionallyreduced to one or two flowers,as in the eastern pi^"t"i Austrarianspecies pys;ii'iid p. biJlora.In its commonform, the raceme mosr is many flowered,terminal, erbit to penjulous,very condensed andresembles a head,the axis reducedto aconvexor nearlyflat ieceptacle.ln somespecies, the axiselongates as flowering proceeds andthe inflorescenie;eco-;"esspite_like. Sometimes the flowen andfruits are densilvnackeo and .ne in{roriicencer. a.i.riu.o i,.r. u, .ontinuous, as rn P. holroydii (Figure50A). in ottru casesiireorcifi;;;;"; i."., Decomeseparated, (Figlret2C), and ttre innorescence aes".iiJJi, in,"rrrpted. varrarron_,?."1i^!.^{*:::!rf"occuning p tn another oniy in gilgiana,the inflorescenceis superficially,ir"il"r-i;-ih"t Thecanrhesin that ;i the recepracFii distin"trv"oniuu. uuid,ri#]" ,rrr, tn. or lessbasal. bractsare more In Pimeleathe pedicels are tereteor.nearly so, articulate at the apex,usually very shoft and usuallydensely hairy, a characteristicexample illustrated in nigrre rzg io, u iemui. flower of P argentea. Male flowersof the ,1mq ip.;i, ;, iilr.t.ul.i in eierr. rio, i;";. an exceptionallylong pedicel. The pedicelsof all speclespersisi toriome tlme a1lerthe frui6 and involucralbracts have beenshed but are usub y no'tu, p"tiirt"nt as the axis.The axis doesnot grow out from the old ;instead any.new branch{es) arise f; ;; ;pp;; node of the stembelow. The p€rsistentremains of the old innoriicencJano its originaabr;;; are useful for identifying Pimerm from otherwestem Austraiiun pruni g.n.ru ,n the vegerative state. -----

'8uol 'spuEIS 'aldlu€xo rutu S Z ol dn iuloJrJulq8re lnoqe seLlsunon d roC,{Je^oaqlJoos?qroodrlseqlSurpunoxns8uuEulspuelS,{JelJeupedolo^opllo,ra^€q 'ro^a,,doH sonadsaruos ueas,4[s?e tou sr spuelS 8uu ro csrp,{r€lrsu eqt selcodslsou ul ',{re^o Jo oql Surpunornsaqnl lEroUaql Jo uoluod pruxord aql ulqtr,r selulnulnm€r€tJoN '\JeqluE Jo sed{l uoulLuoJajouJ oqt ur lEJetel-[uosJo l?rolel pu? Jeq]u?Jo od,4tJollel oql ur lElx?p?s? paqursepsr pos?elaJ uasqseq ue1lod aqt qcrq,,[q8norq] strTs aqt Jo uoursodoqt e:aque,rr8 suoqdr:csep sar3ads aql ul esrorlurfllruts SureqoJu$slqop oqt',{ll?Jotel qlec oqt spoocxe,{llq8rls lr t?ql p?ojq os e^rtoeuuoJe qlr^\ sreqtu€(luou?lrJ Uoqs ^Ja^ e qtrM e t efssasqnsa^€q teqt'(C'aSZ em8rC) 'sorceds nto11[i3uo1Surpnlrur uellBJlsnyujelsei[ eft araqt'auortxe rsqlo aqt ly 'suaubodsd Iere^as peupur osSuueedde ls?al tB Jo osJoJl?l fllnJ 01l?q,r auos reqtue oql puee^rl3euuoJ ,rorr"u B 'luern?lrJ e qlr,r suorx?lse^?qsaqluoJaqJ se]'.,eds pueoapwld ',(seqolFdolo^op IIo,r Jo IIE Jo ser3eds.(uel,\I oerJro) sFdas,s? ot parreJarel? saqolaqt tnq oqnt lelou aqt seol polJaJarIIts s renaouoqt ed Jplnqnloql qfuol eJrluooql sorcodsesaqt suorldu3soprrJ 'eqnl Jo Jo 'DK1.p'isolpltiJo uoqs e ur eseqaqt tE el€uuocaq ol ftedd? qcrq,r qedaspare e^Eq d Butptliu'aBD DdE l3osur sorJodseuos srsoqtuBle 8up?ardsflaplr.{ll€nsn oJesl?das oqJ ',roleq 'slBdos ^ltq8rlsJo al€rEdas sl?das eql e:eq,r1e,raq aqt lz ,tll?nsn Jolnoo,,hl eql alrsoddo po esuroJe suouels eqJ pnq u red eleu:et1eeq1 sdel:a,to qedes Jo Jrpd euo ?rouo8 qloq uI

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'JJqlueeql oJuoJsrqappue rdBqs eL{l pup sledJs 'eqnl Jo 'rnoloJ oql ot o^rl?lersuau?ls eql Jo rllSuelaql l?rou eql ]o odeqsoql pu€ ezlsJo,rou er? anlB^JusouS?rp lset?eJ8 Jo sralceJEqcIuJolJ oqt'JorTrBe palou srep€reqJunluounpur aql uo4 trpdv SuluodolelJe u?ql pnq ul orou.rr?odde uolJo sro,rou ogJ '^Aollo,{ 'etrq,r,{lJBnsn lnynoloJ'saqu2JaqJ ro {urd oJB^aql,a/aww ur alrlt t.llpeJdaepfua^ot ollq,r ruoJJe8ueJ feqJ Jnolocpue ornlxat ur J?lrurs,{[ensnor? sledos pue eqnl lerol] aqJ'l€dot Ietyepeaqt alrsoddoualuEts auo fluo seqt1crym'sttuto!1tt A sercodsuElu?rus?J eql uI ldoJxe 'suau?]so,rl pu?seqol €lloJoc ou 'sledesJnoJ o^eq sle,roIJ eLIt saqJunaqJ puB Dapwtd uI

'uorle8rlsa^ur 're.ta,ro11 raqunJ speeuo?ocpaelorufqJ ur sojnpnjls l€JoUJo ur8uo oql seqolellojoc s? saqolreuu oql puEq€des s? ereq01 parreJar orB soqol umu aql'le8ruung Sur,rolloCseqol I?rolJur?ru aql peuroJpua onssrt lErxs Jq] Jo lruuns aql ts Jso.lPx{lec eqt t?ql paJaprsuoJ oH ?llorm eqt Jo u€d eo4 eqt Suleqssqol Jouur oqt'unrcoorpue puu ellorooaql qloq Jo pelsrsuooanssll reuu eql 1€r.llpu€ aulJ^lecueql lJqlel sE^\Jqnl aql anssrlrolno 'eroue8lErxe Jo aqt t?qt pepnlcuocJe?tuurLfl DapwU Surpnlourur?8e lera^esJo ,{uolBuplo,rou oql ,{pn1scrlauaSotuo $q ur (Z/61)ro8luung ^q pour?tqose,{A ,{lluleJqns aqt ur ra,rolJeqt ;o 'xfl€, Jo arnlcn:1saq1 o1 Surteler ecuapha Jaqunj solecsprolelod se .rallq eql 01SuuFJer oql Jo suorleuese soqol Jouur aqt pu€saqol x^F3 s€saqol ulelu eql papr€Seloqs urnrcooJpuBaqt Io suotuodeleups 3q1 qtlr raqloSolx^lec Jo,rol oql SuqsrsuoJ'ur8uo ur l?lnolpuoddp,{lelrlua Jo'Dapau s?,\\aqnl l?rou aql 1eqlpepnlcuoc (l 961)8luroH SutpnlJ[r'?reuo8Jo lequnu ? Jo sre,^AolJeqt uI saJ?rlr€lncs?^ oqt uortBuruexolaq uorl 'solBJsro se8?puedde'soqol elloloJ 'slBlod Jo se (luesaJduaq,r) aprsu oql uo saqolI€uortrppB oql ot pu€ qlueuodlo x,{lm oqt sE soqolur?rx str pu? oqnl sql 01 poJloJere^uq sroqln?lsohtr eqnl pJdol3^Jp I?JoU 'eeeprooBlou,{qJ lErou llJ,r e e^ErlsaqiuDJaqJ pue oapwld ueJqnsJo sJaquaurraqlo ur sV JaMolJ .I.A lfl a?a.?3ei.ui,{qluerlerlsnV uratse/A'aid 138 NuytsiaVol. 6, No. 2 (1988) Both generahave a two'caryellategynoecium but the is functionallyoneloculate at matuity becauseone carpelaborts at a very earlystage. The stylearises laterally shortly belowthe apexof the ovary, is straightexcept near the insertionpoint and alwaysglabrous. It is filiform and prominentlyexserted from the throat of the floral tube in Thecanthesand oltenin Pimelea.However, it is sometimesvery shortin the lattergenus, particularly in some dioeciousspecies such as P. serpyllifolia.The stigmais terminal,small and with shortpapillae or, in somePimelea specres, prominently enlarged and with long papillae.

Fruit. Thecanthesand Pimeleahave an indehiscentone-seeded . The floral tube may be persistentor partly to almost completelydeciduous. The sepalsremain attached to the floral tubeor very rarely,for examplein someplants of P forrestiana,are shed. A regularly circumscissilefloral tube characterisesThecanthes and alsooccun in most membersof sections Calyptrostegia,Epalkge and Stipostachys.Circumscission almost always occurs above the summitof the fruit and the fruit remainsenclosed in the persistentbase of the floral tube, as in P. suaveolenssubsp./ava (Figure 39F). In one or two species,such as P holroydii, the floral tube becomesbrittle and may be easily,but iregularly, brokenabove the fruit in someflowers. Suchspecies are not regardedhere as having circumscissileflowen. A peffistent floral tube occursin most speciesof sect.Heterolaena and is also fairly commonin sect. Pimelea.Inthis casethe fruit becomesdetached from the planttogether with the entirefloral tube and usuallythe .A deciduousfloral tube is commonin sect.Plmelea and also occursin sect.Heterantheros. In this casethe floral tubesplits iregularly in the swollenbasal portion as the fruit enlargesand is shed(apafi from an irregularbasal ring) beforethe fruit reachesmatudty, the maturefruit beinggreatly enlarged and ofted succulent.In the other casesdescribed above, the fruit is dry and not so greatlyenlarged. In sect.Heterolaena, which has a persistentfloral tube,the tubeis alwayshairy and usually hasa bandof longhairs above the ovary-portion.[n contrast,most species with a circumscissile floral tubehave the longesthairs below the circumscissionpoint, that is mainlyon the ovary- portion.Others are either glabrous or havedeciduous hairs. If the function of the long hairs were purely for the protectionof the seedthey might be expectedto be bestdeveloped on the ovary-portionin all species.It appearsthat they assistin the dispersalof the , their higher positionin fruits with a penistentfloral tube being relatedto this function.

The seedin dry-fruited specieshas a black seedcoat with a pattem of pits, shallow protrusionsor iregular features.Threlfan (1983) has described and provided good illustrations of the variouspatterns on the seeds.

ChromosomeNumber. The basechromosome number for the Thymelaeaceaeis x : 9 (Federov1974). In Pimeleathe lowestand mostcommonly recorded number is the tetraploid n : 18.Tetraploidy has been recorded, almost invariably from somaticcells, from tbreespecies in New Zealand(Elaise 1959, Rattenbury 1957)and 11 speciesin (Cruickshank 1953).The other numbersrecorded, each from one speciesin Tasmania(Cruickshank 1953) aren : 36, 45 and 54, thesebeing octoploid, decaploid and dodecaploidrespectively. There do not appearto have beenany countspublished lor Pimeleaspecies endemic to mainland nor for any speciesof Thecanthes.

BreedingSystem. All speciesof Thecanlhesare hermaphrodite.In Pimelea,hermaphroditism. dioecyand gynodioecyare all common.There are minor deviationsfrom thesethree main statesin somespecies. For examplea basicallygynodioecious species may haveoccasional plantsproducing both femaleand bisexualflowers.

Dioeciousspecies appear to have a 1:l ratio of male to femaleplanrs, judging irom the approximatelyequal numbers of maleand female herbarium specimens of eachtaxon, from the publishedfigures for two dioeciousspecies from Tasmania(Cruickshank 1953) and irom ----

,{cuonbe4orrlpue so,eds uerr?rrsnv uopalcnpuoc uaeq r"r ,^*["rH.til,H,rX?il',j,flii[tl# 'uorsserSo4ursB pepleSo: oqol sosmaruos ur elrsuetxo^lluercrgni se,,lt uorleirpuqxq gb 5b:so! ' aq1 sorcedspu€l?oz ,{roN rnoJ eql uoe,rlequo4esrpr:q,{r1 fiu'e:osopr,r pag6itei s,uo::ng ,slue8? , lqS4fep Surmp lsBolle Surleuqlodureu oql oJo,rsebeds ,{lJ pue seloedseaq papnlruuJ IeJe^as e l?ql eH.oalawtdJo setcodspuepaz ,roN moJJo sjo^\olJ JI?useql Sunrsr^s8nq pue se[ooq,sq]orupu€ saruralinq ,s6rglseetipep:biar 'sloostn leru! 1g96fl s,rorJng Jo ,{leue^ ? fq popu{lod ,{lqeqoJdeJ? ,,{le,r.o eq1 a,ioqe papualxe ,tpoqi.{lu6 ,sleaolJ aqnt l?rolJsqt qll,{_esoql,{[?I'edso IIeuIsqtl{\ sonodsoapu.tld.an1uot s,prq e,tq pat€rtouadoq-ot qfuol slr lnoq8nolqt p"orq ,{lluerorns oqnt eq1lo uoa:od,e1,nisaqj .saposttld1e:og e^eq ot sJeoddeteql sarceds,{Juo oql srsapostqd oaplug .4 jo'snn61y eqt buursri sorcadsprrq I peprocoros[? oH Brlertrsnyu:a1se16 ur aaprz6'1osirce,ti pe:e,uofi_oaiel eql jo ouru lnoqe sJo,rouSurlrsr^ setuJollnq seroads 'rer\oU Jo Jo tq8lo ts?el1z pep:oce:19111; ,mq8re; eql Jo os?qeqt tB J?pauoqt qreol 01oqnt eql uor od_el,{tsrupuq,{c ,{l,rolrBu I?lo[ .sro,rogJo oqt aterloued01 olqe ere secsoqordJepuols auol reqJ lopue1i aa:ei i41rina1aut14 go sor.edsasoqt pue saqunaqJ Jo sroteurlod pdrcuud eql ,{lqeqo:Oie sqtou pu? sorgteltng

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bl aEoc1?.elsu.lfqluerlprlsnv ulets3iA a,{d 'I g 142 NuytsiaVol. 6, No. 2 (1988) L PIMELEA Banks& Sol. ex Gaerrner

PimeleaBanks &Sol. ex Gaertner,Fruc! S9m.Pl l:186(1788),nom.cons. Type: p. laevigata Gaertner(: P. prostrata (Forster& G. Forster)Willd.). BanksiaForster & G. Fomter,Char. Gen. Pl.'7. t. 4 (1775),nom. rej., non Linn. f. (17g2), - p. nom. cons. CookiaJ. Gmelin,Syst, Nat.2: 24 (ljgl), nom.rllegit. Type: \Forster & G. Forster)WiIld. (asBanksia gnldia Forster& G. Forsterin former genusand as Cookz gnidia (Forcter& G. Fomter)J. Gmelin in Iarter genus)(lecto: fide S. Threlfall. Brunonia 5: ll8 rlq8lrr. GymnococcaFischer & C. Meyer,Index Sem. Hort. Petrop.l0: 46 (1845).Type; p. tlrupaceo Labill. (as G. drupacea(tabill.) Fischer& C. Meyer). Heterolaenalischer & C. Meyer,Index Sem. Hort. Petrop.l0: 47 (1845\. Type.p. spectabilis Lindley (asH. spectabilis(Lindley) Fischer & C. Meyer). C:lyptrostegiaC. Meyer,Bull. Cl. Phys.Math. Acad.Imp. Sci.Saint.petersbourg 4: 12 (lg4l). Type: C. hypericina(A. Cunn. ex Hook.) C. Meyer (: pimelealisustrina Labill. subso. hypericina(A. Cunn. ex. Hook.) Threlfall).

MacrostegiaTvcz., Bull. Soc.Imp. NaturalistesMoscou 252: 177 (1852). Type: M. erubescens Turcz. (: P. physodesHook.l.

Prostrate lo tall or rarely herbs with a woody base, usually erect, usually hermaphrodite,gynodioecious or dioecious;hairs simple, usually colourless and silky. Brancftes with a stringybark; upperstems usually with ratherinconspicuous tufts of hairs in the leaf axils.fuayes usually opposite and decussate. shortly petiolare or rarelysessile; midrib prominent on abaxialsurface and often yellowish.Inflorescence ustally a simpleterminal raceme, rarely of almostaxillary racemes each terminating in a veryshort axillary branchlet; raceme(s) usually very condensed,often head{ike,rarely reducedto 1 or 2 flowers,often with an involucre of bracls.Pedicels terete or nearlyso, usually very shortand hairy,articulate al apex.Flowers white to red or yellow;bisexual flowers protandrous or appeartngso. Floraltube often hairy outside,glabrous or sometimeshairy in the distal part inside,of 2 usuallydistinct portions surroundingthe ovary and style respectively;ovary-portion usually fusiform; style-portion usuallyeither narrowly cylindricand expandingslightly to the summitor very short.Sepals 4; outer pair ove appinginner pair in bud, often more hairy than inner pair. Corollalobes absent. 2 (in W.A.), insertedat the summit of the tube oppositethe outer sepals; anther usuallyyellow or orangeat maturity, then becomingbrown. Ovary 2-carpellateat first, effectivelyI'loculate at maturity, usuallygreen; 1. ,SO//esubterminal. filiformt stigmausually globular or scarcelyenlarged, small and papillose or largerand brushlike. Frul indehiscent,dry or sometimessucculent, usually remaining enclosed in the baseof the floral tube. Recordedchromosome numbers.. n : 18, 36, 45 and 54.

A genusol c. 107 species,I speciesendemic to Lord Howe Island, 17 speciesendemic to New Zealandand ChathamIsland, and the remainderendemic in Australia.45 soecies occurringin WestemAustralia. Pimelea species extend throughout Australia except foisome areasin the extremenofth, occurringin a wide rangeof habitats,from alpineto desert.In WesternAustralia, Pimelea is absentfrom the Northern BotanicalProvince except for p ammocharis,which occursonly in the southemparts of the province.

Notes.The namePimelea is derivedfrom the Greek wordpimele goft fat), in referencero the oily seedsor to the fleshycotyledons. Chromosome numbers have not beencounted from WesternAustralian material of the genusbut 2n : 36 hasbeen recorded from a Tasmanian populationof P. serpylliJolia,which also occursin WesternAustraua. '8uo[ 'g ruur 9 oruosqtua s^e,rp'8uol urur ,{llsoruso^ee'I " " "eplsul 'aprslno 0Z-01 D qdolDrlil snorq€I8 ^rr?q sla,rold snorqelSsurels g t " .' " 'aprstno DllotrytidnsA Z" eqnt lerou eprsurtureq ,{re^ 01 tEq,rorxos os ^lrBeu Jo snorqeF sra,&old 'JueJsoJolJUrqJee rBeu ,fureqsuals g 'Euol ruur (9)'SI so^so'I '8uol S ruru I-2 0 slocped oIEII lalqcu"rq ,{J?llrxeolrurJep lnq oqs " uo ro leuruJol Jolsnlcro,rolC t "" " " " " .' oa1ua8n 4 6" Euol urru 9,9'l slecpodaleli{ ^l?llrxe ,{lrurs suupadd?'letqou?rq tuBIIrx" lroqs ^ro^ e uo "trffirj3t:lg^j

ueql re oqs uoruod'olfts :3uolrntu (g-)Sg-9 sro,rol] ol?rrroJ oqnl " " " "" 1 Jo I?JolJ Z oua.SyB flunleur 1? po{eu lrnJC snoJq€18so^?ol pu? suals 4 I .' .'' " € slrDq)ouaod ll eqnl I?rol+Io eseqlualsrsrod oqt ur pasolouoInr.I ,{rr?q**".rlJ,lrjffl:J

ueql roauolqcnu uoqrodel^ls :Euoluu -g sJe.roueFueJ oqnt I I 'snorcoopJo l?rold Z slu?ld I salradsol fa)I

...... uoruod aEalsoIdKlDJt 1c0s ol,{lsueql:alror{s eqnl lejol] Jo uonlod,,{Je^o lueseld slJslq slrssos.g ^ " .' " ...... uouJodol,{ls lernlo^ur adDodg t lJos ueqlJo8uol eqnt uoruodfue^O tuosqesto€lq olrsses.9 I€JoU'e^oq?Jo lejcnlo^ur sBlou (luasalduoq,r) aqnt lelou Jo slr€r{:lrn{ ur ,fuE^oo^oqp olrssr3srunclrc aqnl uaq] ,lou ,lo ^Jr?rl{JB^O .S ".. IeJo[ Jr ouaDlolal)Hl, lsos ,...... e^oo? sJreqluoled Jo8uolqcnu e pue uoq:od,,(re,rouo s.Ireq 'os Joouoz t.roqs qlr,r uoqt Jrlo olrsslosunrlrclou oqnl lBlolj snolq€I8,UEAO 'iuolS u]tx 0l suau?ls crldrllooi alB^o.4l,roJlEUsledas.lurod uorssrssuncJD ,os ,lo o^oqEeptsur ,{Jr?q ueql pel3ulsuoc^lueuuroJd sqnl ., - 'lc€s8uol Jl lou lelolj aBalsotcny4lg ruruZI sueu?ls :eF8u?u] ,{l,roJleu,{la^ slpOegaprsiii ., snolq?IS'luloduorssrcsurncjra w polculsuoc,{lluoultuojd eqnl lEJolC uoruod.,tuB^ouor{1,ruolsrsrod,,,.,jll"lili rHii:13'{;:i:,T1,fiff,T: eqnt l?lolC snorcoorp,{lol€J Jo snotceorpouf8lo olpolqd?uleq .t " slu?Id oalaald lJes .' ' ...... tup 'uorZ lo luelnccnslmlC od-olflsu€rll ls8uol uor od ,{rc^o:olrssrJsuncrr3 fleler ,1ua1srs:sdro lrn{ ur asBqlp flleJn8eJr AurlllldsfJl€nsn aqn} .t 'epl$no lelolC snorcootpstu?ld fJreq pu" t lo tuosq?slcelq leJonlo^urueq] ale8uolo lczduroi J ,qenuue ,{[ensn oruaoselouq soloqodse l3rql se aor,ratlnoqB ,{1ensn tdaoxotueulruojd fll?rx?qB sopou uets :slEnuue^lotBl lo sleluueJed.Z _ _ul ,/-t - strqJusodljs 9 lcos aprstnosnolq?F sloeJqlelJnlo^ur :,{tunteu 1a ale8uo1i sJuecselouqselorlad puofoq,{lFrx?qe BurpnJlold sopou .Z 'aJuorsalouur tou ue s:spruuoled lenxosrqJo eleu uBqt tourBq,{lqeecqou lou (luesaldJr) eruacsolouulepual el€3uolosrxE eql lo xo^UorJo ssolJo aJoluel3eldeJad .I .' leu sotaqlu alaH J 1c0s .... otuocselorrur aleu u?ql leurzq qcnu erumsalollural€uraj a^muoc.{llueuturord alrz1o:iX .1

suorpas ol ,{aI

a?at?aElau^qIuprtBllsnv ulalsolA .,{d -.I g NuytsiaVol. 6, No. 2 (19881

7. Flowersdensely hairy outside.Fruit succulent. 8. Stemshairy at first. Fruit enclosedin persistentbase of floral tube ...... 4.p. clavata -- &_Stemsglabrous. Fruit nakedat matudty...... 5.p. microcephala 7. Flowersglabrous. Fruit dry. 9. Pedicelsdensely hairy. Inflorescencecompact or sometimeselonsate at matudty, rarely appearingbranched, subtended bv 2 or 4 seJsile leaflike bracrs...... e.Pedicels glabrou, "r;;;;d;;;;. i;f1;;il;;i;#i; ,branch' i| ;:r":i;:""r*' commonly appearingbranched, each subtendedbv a deciduous brownbract and petiolate1eaves...... 7.i'. forrestiana l. Planrshermaphrodire or gynodioecious. 10. Stemsvery pale yellow-brown in distal100-200 mm. Inflorescenceelongate at maturity; .^ involucralbracts 4t, glabrousourside...... 1f.p. hotroyiii 10. Stemseither becoming brown or grey to blackshortly below apexor (in p trichostac,hya)becoming reddish. Inflorescence compact or, if elongate,then bractseither 4 and hairy outsideor absent. I L Ovary glabrous.Floral tube not circumscissileor, if so, then with short _Irays qn ovary- portion and a zone of much longerpatent hairs above. 12. Leaf marginsflat or incurved. 13.Stamens shorter than sepals;filament 0.3-1.5 mm 1ong...... 44.p. brevifolia 13. Stamensequalling to geatly exceeding(very rarely shortei than) sepals; lllament 2-7 mm long. 14. Sepalshairy inside.Ovary-portion of floral tubewith antrorsehairs ...35. p. spectabilis 14. Sepalsglabrous inside. Ovary-portion of floral tubeglabrous or with reflexedto retrorse(rarely patent) hairs. 15. Floral tubeglabrous inside, the long hairson outsidemixed with short hairs.Sepals moderately densely hairy, with both long and short hairs. 16. Outerinvolucral bracts with distinctyellow margins 0.5 1.5mm wide.Flowers circumscissile...... 43.p. tanata 16. Outerinvolucral bracts not asabove. Flowers not circumscisile n"oa' 15.F6;;i';;b;;iil;i;h ii;i;;iffi; ;; th;;;i;ih; i;; ii;i;;12;{ outsideusually not mixedwith shorthairs. Sepals sparsely hairy, the hairs all of a similar size. I 7. Ovary-portionof floraltube glabrous or with patenrro anrorse hairs. Leavesdiscolorous, with a sheenon adaxial surface __ :...... 32.p. lehmanniana t/. Ovaryportion of floral tube with reflexed hairs. Leaves concolorous,duII...... _ ...... 34.p. rara 12. Leaf marginsrecurved to revolutelaterally or recurvedar apcx only. 18. Anthers subsessile,with strictly adaxialslits; cellsslightly ixceeded laterally by the connective. not or scarcelvexserted ...... :...... 3g.P' brevistyta 18. Anthers with a distinctfilament, the slits lateralor semi,lateralafter dehiscence;cells laterally exceedingthe connective.Stigma prominently exsefted. fllolqDz '81 "" "" "" " """" "JIeq UIpepuedxe d 'lurd l€tsrp fllq8ls ,tluo eqnt IBrolC]erx?p€ ^lputs sus roqtuv.ZE slrsa^fs " " " " "" ptslp ur popu?dxo,{ltcuqslp d fl"'{urd 'ro JIeq 'lero}el-ruas ueql J otrqr{oqnl IBrolC stls loq]s:}'eelg aqt uro4 lcurtsp,{Ipnsn'(8)9 stJ?Jg 'eprstno snorqep s:a,uo1j 'Fale1E pBeqro^\old 'eqnl plolJ eql ur aprsu flteq ste^{old '8uol Jo JI?tilelslp 0t uru Z-90 raqlu? :?uol turu , ot dn luoul?lrJ 'crd '8utp?atds .62 :suour?tsSuol rutu 9-SI a o1ol€ao ^l,l\oreu slBdos " " "" " " " sary(qd d 0€" 3uo1Iutu €-ZreqtuB:Euol utu gl-g.Jl luauEIU:suaursts Suol turu 'Jeln8ueulflnd.oll?u fle^'icete slBdos.62 6-S '1U0S0J(1 S13?JqlerCnlo^ur elsses're^\ou u uonlod-,lB^o ueq] te8uoleqnl ploll uorlJod-ot,tts " """' .' " " " Jo LZ otl1uotzlwJ01 3uo1tuur g'0 01 dn sr?q qtl,r'Euol urar eqnl p?dtrloc "JurJsoJoUuI.gZ " "" " " "" S'Z-Z" " " l€rolC'Euol D[tpDlsoq)lr1d'8 uur sreq lsaSrol '8uol f-gz .gZ aql ruur t-E oqnl l?Jold ^lrmleur 1" atz8uolo"JusJsaJoUuI .1u3sq"spsJq .17 lelcnlo^u alsses uorUod,,{JB^oueql lauoqs eqnl uoqJod-s1,{19 'o^oq? I?toU Jo sp lou (luasoldueq,r) eqnl plou Jo srl?q :1rn4ur ^l€^o e^oq?olrssrosuncrc eqnl ueql 'lou 'lo ,fur?q,{J?AO .l 'd ' """""""""" " " "" "" lelolJ Jr I slsuauo^D 18 3uo1{xur (€-)Z-9.I sr€q tsatuol aql'8uol turu ZI-g eqnl :eltq,{\sra.^aolC gZ otlluoJnal " .' "" " " " .' 'Auolturx gt-€ slt?q A'9€'3uol tso8uoloql ruul 8l-Z I oqnt :urEeJcJo ,rollo,(opd sre,lrold 9Z 'perJp uaq^\ olnlo^oJ nururoJaq .4psousa^pa-I ,{lunl?ur l€,{o[e,t oJEdoi altq,r sle^\olC'tZ " oau8rutal'{'lt " "" "" " "8uo1uru ot dn olcunped 'JJlls € roqi?rse^Erl isru t?,trr?qlBq,{oruos ueuo sulots " ' SZ Dasord 0r" 'moIq€I8 .gZ Suol turu S€-i elcunped alqrxolJ$^ee-l sueis 'atrnlo^al '{urd Surujoceqtou so^?e-I daep01 opd sJaaoH '?Z 'esnlqo '8uol so^?o[ueq] Suol urut €-g'Z IUru Jr .tZ (g-)SZ-r'0 tueuelJ :qBdesEurp$Jxe ol uBqlrouoqs suauqs '6€ " " " " " " " "" 'olnc? DiD!l!2d so^?sl'8uol ruu 9 S-92'gluou?lrJ:slEdas Eulpoacxe ,{llEe-6 ol ,{nq8ls suotuels tZ 'e4ue3rc osBqol pourJuocsJr?q qlr^\ lo aprsu snolqeF spBlq leuq 'd '€t 'ur8J?ur l?Jrnlo^ur ZZ DIDaDI uo ldecxeoprsur ,trEq sp?lq plcnlo^u ]€uul -ZZ 'Iutd sle^{ogueql ,?uol ruur g-l uq oqnl Suol ruru sraqluv .lZ " d"' !-d l?rolC S I-9 0 Dll{tldtqtDrqi1St"" etlq.r'8uol rutll g-t oqnl p:oyg.3uo1ufiu sreqluv.lZ 'eqnl S.0,t.0 'd Jo sr?q tsouoqs usql reEuol qonru tou (DtttqdttlJorq ur .61 sourrlo{uos)ro srrBq oqs qll,r! poxlu oqnl lptou srEq tso8uo.I " " " "" "" .' "oprswJo stissrsa't["" sno.rq"J8 ,e[sses .02 0qn1 lprold ewq lE SurdsBJc-uolsl?q^\ouos sa^ml 'd '2t"" " " " " " "" "" 'l?OJql DuotuuDulqal le eplsu srBq '4€lortod Jo 8uu e qI^{ aqnt SurdsBlc-uolslou ^Juoqssa^€e-J IBJold 'aqnl 0Z Jo srrerllse oqs u?ql la8uol seurl .61 t 1se0l1?'srreq uoqs qlp\ poxru tou oqnl l€rol} Jo s.rrBqlso8uo.I

'e{U ''J S'I apacsaslaur^qJuprl?llsnv u.ratsa^\ g )46 NuylsiaVol.6, No. 2 (1988)

31. Flowershairy outside.Bracts (6-)8-numerous. usuallv similar to and oftengrading inro the ieaves. 33. Leavesalternate, often crowded,glabrous. Involucral bracm c. 40...... 12.p. groniticola 33. Leavesoppositeor, if alternate,then usuallyhairy. Involucral bracts8-20. 34. Ovary,portion of floral tube denselyhairy throughour. Leavesusually alternate. 35. Hairsof floral tubenot longeron ovary-portionor slightly longerand gradinginto somewhatshorter hairs on distal part of style-portion.kaves usuallymedium green to dark bluishgreen or deepgreen...... l3 . p. imbricata 35. Hairs of floral tube long on ovary-portion,suddenly becoming distinctly shorter on style-portion.Leaves - . _.usuallypale to mediumgrey-green...... 14. p. subviltifera 34. Floral tube glabrousat extreme base of ovary-portion, denselyhairl above.Leaves opposire. 36. Young stemshairy. Bracts ieflexed in fruir...... 15.p. villifera 36. Stemsglabrous. Bracts not reflexedin fruit...... 16.p. eiecta 30. Ilowers glabrousinside. Flower headpendulous or erect. 37. Young stemshairy. Leavesalternati or opposite,hairy when young. 38. Longesthairs of floraltube 7-9mm long.Involucral bracts 6-12 ...... I 1.P. ammocharis 38. Hairs of floral tube up to 2 mm long. Involucral bracts4-6 " re P lonsiflora 37.st;;;r;b;;;;. L";;;;;ii;; ;il,"*. 39. Antherssubsessile at throatof floral tube,strictly introrse;celh laterally exceededby connective.Stigma not or scarcelv exsenedfrom throa1...... 20.p. preissii 39. Anthersdefinitely exserted, the slitsusually semi lateral after dehiscence;cells laterally exceedingthe connective.Stigma distinctlyexserted. 40. Inner involucralbracts densely ciliate around aDex. 41. longestcilia of bracts0.5-l mm long.Longeit hairs of the floral tubeconcentrated towards the base.Ovary with a terminal tuft of hairs...... 23. p. sulphurea 41. Longestcilia of bracts(l)2-3.5 mm long.Longest hairs of floral tube occurringtboughout style-portion.Ovary hairy bur hairsnot in a lerminalrufr 42. Ovary,portionof floral tube with both long and shon hairs throughout.Inner involucralbracts glabrous or sometimesslightly hairy inside...... 1...... 2j. p. nnctoria 42. Ovary,portionof floral tube with short hairs,the long hairs (whenpresent) confined to distal part. Inner bracti usuallyhairy inside...... '...... 29.p. suayeolens 40. Involucral bractsnot or scarcelyciliate. 43. Ovary-portionof floral tube with densedeciduous hairs 3-4mm 1ong...... 29.p. drummondii 43. Ovary,portionof floral tube glabrousor with sparseto densepersjstenr hairs I 3.5 mm lons. " """' sudJDtJd 92 ,{{o ef apd ol uaoJ8^uBoJ3 sla,[olg 'o]n* sa BeT - qr+ru ot pourJuoclou srr?q:eprstno^r?q q?dos /t DsoudnJao'd 'SZ''"'-"""""" " Arolle^ sJa.rolC'omlqo ,{[,rto.u8uJo osnlqoso^eo.l 'ep$no qupnl Euole sJl?q ,{aJ ? qlr^\ to snoJq?lEspdss /t 'lnoqSnoJqt.{JrBq,{J?^O,r,o1o,{ uoql snorqep J'spoos uo sJI?q,reJ? ls?all" qlh{ ueuo sJotrolc gt suapuad'J 'n2" " ' "" " 01 poulJuoJ 'uoo"6JI?q I€s?q srEqro snoJq?JE,fi?^O al€d'snoJqEIEsJo,lrold 9t 'molnpueop?aq 'sreq :elrrolg Suprards Jo urruueulnpu e$?dsl{loA4BIoJ E q1r.[ ro snorq?[3ot snoJq?lEqnstaqlra ste^\old'tt " ""esro4uB npunqtogf'g ZZ" ol tuelBdaaoq? sr?q eqt lulod uorssrcsunorrcoql e^oq€ rrltu ?-Z JoJSuuJnmo sreq esro4er etnuur qlr,{\ oqnl plolC SuddBI 'eprslno -ro^o ,{[1BoJE ossq oql lE qsrppeJsp?Jg'gt a1olusn?uo'd '12"""""-"""" esro4ue ol 'Eurddello^o tuolBd aqnt srr?H ^llq8rJs 'eseq II? l€rol} Jo ol pourJuoclou rnoloJ eqt Jo qsrypoJtou slc?Jg gt 'snoFpuedlo pelo peeqla^{olC e^oqe sr€q 'oqni Jeuoqs Jo Suua^oc esuap esolc ? qlr^{ I€loU Jo uoB.rod-,{l?^ooql uo sJreq8uo[ qll^\ lar{ou Jo eplslno tt

aeecPoeloufqluertPllsnv ur.lsolA 'a^u .-I g 148 NuytsiaVol. 6, No. 2 (1988)

Sect.1. Het€rantheros

Pimeleasect. Heterantheros Rye, sect.nov.

Fruticesdioici. caules glabri; nodi prominentes.Involucri bracteae4 vel 6. Inflorescentia mascula compacta;receptaculum concavum, supra insertionembractearum extensum. Inflorescentiaefemineae a masculisindumento in pedicelliset floribus abortivis vel in receptaculodensiore et longiore.Tubus floralis proportionestylari quam ovariali longiore, basi irregulariterfissus sub fructu exutus.Ovarium glabrum.Fructus siccus. Typus: Pimelea gilgiana.

. SlmD; dioecious.Srems glabrous except for inconspicuoushairs in upperleaf axils; nodes abaxiallyprominent, up to twiceas thick asWtiole. Leaves opposite, shortiy petiolate, glibrous. Involucral bracts4 or sometimes6, well differentiatedfrom leaves,eieit, glabrois.Male inflorercencehead-like, terminal, many-flowered; receptacle concave, extended as a rather thin annular structureabove insertion point of involucralbracts, the basalpedicels arising more or lesslevel with the baseof eachinner .Female inflorescence as describedfor maleinflorescence, differing from maleinflorescence in having additionaland longerhairs, the longerhairs occu.ing on pedicelsand abortedflowers or on receptacle.pediiels haiy or glabrous.F/owers medium-sized to large,hairy outside,glabrous iniide. Ftorat tube with styleportion much longer than ovary-portion, penistent in immaturefruit, splittingirregularly in ovary-portionas fruit expandsand tard y shed.sepcls spreading. stamens 2;-coniective narrowerthan anther;slits lateralafter dehiscence.ovary glabrous.sry/e filiform; stigma large,brushJike. Fruit dry, nakedat matudty

A sectionof I species,occurring in south,westernAustralia.

Nores.The, single species of this sectionwas not describeduntil after Bentham's(1g73) treatment.It is uniquein having a stronglyconcave receptacle extended above the baseoi the involucralbracts. A secondunique feature is the presencein the femaleinflorescence of either sterileflowers or bractlike structureswith h;ils longerthan thoseon the female flowers.In other respectsthis sectionappears similar to sect. pimelea exceptthat the involucral bractsare betterdifferentiated and the flowers largerthan is typical ior sect.pimelea. l. Pimeleagilgiana E. Pritzelin Diels& E. p tzel,Bot. Jahrb. Syst.35: 396.39j,t.46 (1904). lyoe: Geraldton,July 1901,E hitzel 429(lecto here designated: PERTH); S of Geraldton, 27 June 1901,L. Diels 319'7(isosyn: PERTH).

Shrub,erecI,0.35-7.2 m high,single-stemmed at ground level, many branched above. S/ems palegreen near eachinflorescence, usually becoming medium brown to blackfurther from alT'x,_eventually becoming grey. Leaves usually patent or widely antrorse;petiole usually 0.3-1.5mm long; lamina concolorous,medium gieen, usually nirrowly ovaie to narrowly obovate,sometimes more or lesselliptic, 3-23 x 1.5-6(-7)mm, flat or adaiially concave,more or lessa..tg Peduncleup to 1.5mm long. Male inJlorescencainvolucral bracisgreen, broadly ovateto almostcircular and 8.5-10.5 x 7-10.5mm or the outermostbracts (when 6 oreseni) narrower;receptacle extended c. 1 mm aboveinsertion point of bracts,glabrous, pedicels , .(LiFJ) .lr8ellrulou oapuldng uaaqnsDalaud L6t: rpord..JC ur ,roussratr{Dapwdng 'ttas .... .utou,.lpu3 oalaatd (gtgl) 19 :t lddns.ld uog,.ll8ellr oalatudng'.ena1awfi .tsss Balarrld z

, - po[ollsep s?,r unlJeqlsq leq] Jo ]lnq eql ueq,r lsol s€,r slalc ,{q potcalloc .grc ' ed,{lu,{sg aql .{lq?runserd ouot7lt7oapwtl JoizrietdruaO,tl ou,trou s, e:of1

.srolnp"ro requnJJopa3u ur sr ocuorsarouur aqrJo empruls aqJ lnou"lootLiil::ril:lJiri,lH: ^\aJeql lo uollJolordeuos opr^orda3uecsaro1lur oleuoJ eqt ur srmqer{l sdzq:edlj-rntcnrts aIII-lJelq alou B ot sle,rog aluolsoqt ur uorlJnpalloqUnJ ? slueseJdoJ^lq€urnsaid uorlenlrs JellelaqJ truuns oqtt?,{Jr€q pu? elplecEl euooeq ol slmdde,{loJousaullJr,uos lnq }tuuns eqt sJe,ro[Jeluols :sa7o1,,. 1? snor^qosleoq sourlauros saouocsaJoUur oprxoJ Jo aloetdoca:aq1 T so^rlElarosolo ou qlr,r solJadstcuttsrp ^Jo^ y saglutl/y : 'laquotdas .po ielN ad SuuaMoH J ( slarrodpues u,,auolseurr s",od",;;L?",iT,.1;jl:lj:#j,3x';l:T'*T; $fi.,H':#ff# 'tseoc ,(fl ,S .r) eql Jo ur{ 0,€urqlr,r s,{?,,hp ,gsl)flI .1506Z u?ruoo.I reeuol qtnos(A 'S,S0092) ,0[og I I pu€lslBolt?H IrC ruolJspuotxA 1.g antu j1:uoanqulstg 'h! '19.61 .r:(HJUEd $es'er?BuocJoS trut 9ll rru0t t 6ZLguostr,44.'5.5",,1rt")qffi4 urr./:(HJugd) gLL ptatualso 'uolpterec:(Hiaai.liwl ,qrrc .t Hl ,pu€tsl 7gggh arlqaw _y-qT.qsloorroH :(HJugd.).Estr qtoauuax J) aouelllrlq:ddxalt qbs uAJug V UDIg :HJXgd) plnag E'uawealJoS 'uonBls LI69 ST,q8nouaorgpue eura:oeg uai,,r.r1iq :\HIdEd) 0789poag ST etEtueJ:((V) tZSt Kqqsy 6r peoruo 7r11.V,k:oaa:D.pautanxa}:od uolu,{-IJo S :(uaes0t ro^oruo4 pelrelos) vl.IvuJsnv Nuaisa,t\ suiunndg ,urur s,rorleurpnrrguor r'r^ z x t .cpaas.Buor u.uE "rrqlrz.aTlr'iil ,;lljrj;tJ :'r"ffij3 srpq:eqnt s ,iiuo1 uo lnoq8norqttureq ,{lesuop,Buol rutx [.0 ot dn sledesaql unu 'apollttsldpue t S.0,ollq?rt sapounuetstnoqlr,r lnq syedasueuo pueeqnl Je:ogpesse:drubc p""-npei " ,j1i^ arueJsero$uraleuo; stauopf .c Jo aa4toqV.urur 9 Z I ,{q pqJasxaal^rs :iuol riru f ,t:i,;,ro :urx x [ 0 Z0 c Jeqluepue Suol ulur S.0 ot dn lueu?lu u qll,r sepou[x?tsieqnl fred uo osoql ,8uol .J .ele^o Jo I€lslp ol ]ellulls sJieqqll.^ ruru S.l 01 o1e^ofl,,\{oreu sledes:Buoy {xru -o sllEq ,uotuod ,Buol .c I tsoSuoloql al,{lsuo sJrEqasJeo3 q1t,r uru t pue JUpurl,{J uoruod 'snoJqplS .lluuns .uep 31,(1saql pueuur g.0 J x-z-s l uotuod-,{l€^oeql lE txu .3 '3uol trur g :auol .J 8.0 S c,aqnllerou :etrq,rs.ir,14orr lotu aT uru t.0 {.iunua,rryoqe ue qlr,\\lo luesqeepollrlsrd:ruur 9 0,t 0 x 9 0,t.0 lol.llu?pue 3uo1 urru g.3-;.1 1ueule1.rj eqj,sledas ueql reauolro rouoqs sueurels:aqnt uo esoqfot :elrurs sireq qirnr;aio1 wut ,e,roqe i.g 7,c!lor1a ss61 ro e:ou qedes:8uol ulru I-9.0 sJr?qtsa8uol eql s:req peFuet seu-ruarubs;tulizO:o eslojtue'eslEmloq]€l qtr,^\,ruur ur sno:qep,l[uluns .c,Auol 'JrJpurT.,{J Z-l leseq lp.rx?rprulx I luru /_9 ,{l,,ho!euaqnl le:og iqsrlurd:o at\l^ ua^oLf ururg of dn srreqq'1r.,r,r ,aiq 4ow.Buol,trruuns O sloclped.:3uolruru S ol dn pxe asrortuBsrr?q aqt ]e pue aptsur,tureq Alasu0pTY ! l.S'luruns ,trn{ ,re,,r.rog t€ 1d$xo oprstnosnolqel8 ur Buoluut / ol dn ur slce:q;o luiod uorlrosure^oqe uur g Z I snlnuu?uq1, oJr]u€ uB sp popuelxoalcetdooer :uru E1_1x gj_g ot rtE^o 'r!n4 ur :ldrnd per ,ro,rou :i:Li^]l:'.)JuaJsatouul 1,".q: doop ur uea:Eqsppe: src?rqlBrrnlo^ur alqaa! Suol LUu.l ,auo1 .i €-t 0 slr€q qlr,r saurleuos lo snolqEl8 r;ur f.g .e,{d .l 6il aeaceaelalu,{qluPrl?osny urotst/\\ B 150 NuytsiaVol. 6, No. 2 (1988i

l:F!!: 2^.Piryele-a gilgand. A- flowermg-flower stem of male planr; B-flowering stem of female planl C- male flower (x t0);D femareflotei {x I2):E- srerle fromferiale ireaa ix iztltiiawnirom ni. crinfaa and JS. Beatd 6911 lB- D. E\. sooi tl, cl

Gilg, nom. illegit. sect.Autopimelea Gilg, nom. illegit. in A. Engler & K. Prantl, Nat. Pflanzenfam.lII,6a:243 (1894).- Banksia sect.Pimelea lBanks & Sol. ex Gaertner) Kuntze in T. Post & Kuntze, Lex. Gen. Phan. 59 (1903). r-1s889 Joqloul u,roulun tnq a3o11odgpas u uolxurocsr lrruJ ur peldruJoturoruoraq pu? ot?8uolo 01 ocuocsorouuleql .{q u,,t{oqs,4cuopueloqJ'uoltcos letlel eql ur peolsurpor?ld aq plnm pue aSollodg lnes o1 Dalawtd tcas lurl ot nadde ounpsauol d pue ota8lnalds napw4 'Jauueu srql ul Surlllds seqnl qtr^\ sorcadssepnlrur teqt 'sotaqJuDfiJaH 'loas 'dno6,{luolerolJ Jrd,{louou aql uorJ U€dB oql pup lualnccnssorurlauos sr ltn4 eql qrrq,r ur dnor8,{luo oqt sr oapatd ]:,es os?qeqt lp lllds ,4lJeln8arltaqnt oql ql!.r 'sle,rolJ 's^\oqs lero[ lln{ luelns.ns? oql ol uonrppeu (t9 :g16l) ul^ul pup oroolilrtrul ojDttsold 'olot]sotd 'pueIEeZ d Jo uortuJtsnllruV saroodsu?rl€llsnv ol .l?l[xrs fJe^ s oqauw ^\aN urorJserJads od,{l oqJ sorcodsu?rT?rlsnv ul upql ete8uole^luouruoc erou oJesJe,rolJ oql pu? snorcoopou,{3.{llueuruoperd ol€,{sqt lpql ldecxaserJeds uBIl?JtsnV ol IEIItursaq ol Jeedde,{oqJ uorl3assrql ot Suoleqsnuo8 oqt Jo srequrouruerT?Jlsnv ?rlxa eql IIV satoN 'lrosopol eurdp ruolJ SurBueJ 'pBardsopr,r slElrqBqur pu? solelsII? ur Suumcco er?sorcods uellellsnv aqJ puslsl ur?qteq] pue pueleaz,reN ot 3ruepuasorcods '(€/6t) sru€pv pue ot SurprocaE'pue pu€lsl 'erlellsnv/l Il€IAtr'sobads a,lAoHprol ot Jrrxapuaseroods [ o] cruapuaseroods gl 9[ Jo uortcosv ',{rp lo lualnccnslrruj|ecumsrqop reup l?rx?p?ssol lo aJou (?rT?ltsnvulatsel[ uI lou) ,{leJeJ tuo^ ro IeJelelol l?.Iotel-uuosstrls :reqlup uBql ro,{orleu e^lpauuoo:[ .{laJelfua^ (BrT?ltsnV urstseiAur tou) to suawDls Surpeojdsslrdas ,fue^oe^oqE ollssrJsunclro ,{lel?l 'luetsrsJod 'peqs Z sourlatuos ,{lrprel pu€ spu€dxelrruJ ss esBqlB flJelnSolJlSurltllds ,{ll?nsn 'uoqJod -el^tsu?ql JoSuoluoruod-fle^o eqr'll8rus aqnl IBJoU:(?rTeJlsnyuJa$e1t{.ur) staMoL[ pnxastq n alDuad snoJq?I3ro ArrBrlsp)ryad pe8jeluoJo lopualssqceJ :pslo,rol],{u?ru- 'tcojo ',{lr:n1eru l 1eele8uolo-poldnfiolur ,{leffr Jopedtum aruarsarcU[ule\rlpel ro olnuru 'luoseJd soulleuos spotq elsses olrssosqnsro olelorlod[1lsot4s sarcaT olortedsB 'luourluoJdluznplu! {Jrql sBool^\1 c sepou:^r?q Jo snorqElS.sualssnorJeolpou,{8 {BrT?llsnv ul tou) lo allpojr.ldeuraq(?rlBrtsnv uletsolA ur lou) .{lelej to snorJoorp'sqnqs 01ot sqruqsapun '((€861) 'llBJIarqJ 99l :s sruounrg eprJ'o13el)(raussral^tr DuDlptoMaq :) 'Ilanl^tr s 'oaun) d C DquDqrop 4 :adQ 1E9611 u?qd ueC xa-I lsod ur eztun) 'r3es 'CL}l09 T J l'\llu?g)DlwqDqtro DBluDg- 9Z:9 lerlmv Il' q$egotwDpqtlo ]Jesoalawld '(oroqpatuu8rsep otceD lg DttoJiUtdns 'd '0ggl) 'JC d :adiJ 019 :fl rpord ur leussretrAlaDqiuolrlw 9 oapudng \Jes o4arutd '(ro^el,^l JT roqcsrC(llrq?1 DnDdrup g se) llrqe'I oarDdrup (l,Sgl) rpord'JC ur raussret{(ro,{etr{ d:adtJ 'S: 'to,{otr^I.J J ? loqcsrd)DnoJouuitD txesoalaard - 1lc col lrSolrutou DJoJouopW 'l9r8ll 'le,{ol,{ l n)o)ouuttfD - 9r :0t dollad UoH ues xapul J T raLlxstiono)ouuitg 'rg 'd Dimds :adtq '\t061) u?qd uoC xo.I '0z1un) d 09 tsod ul04un) (pug) s,tqrwsuoqJr Dts\uoqodtJtlesnslu'g - 'IerrsnvT J G.L}I) IZ:9 Il'qtuog \ lpug.)stqro$uoqJ tcasqnsDrBaJso.ttdtl') lxesDapwV - ltggl6V:n 'rn8erC"lfonl{ ' C (Jpuo s[q)us:uoq) r3esDapuld - (gtgl) 19 :t lddns Id ua0 lpuE \ lpug,)s{qrolsuoqJ go8atsotldllo3 - (tt8f) It€ Id uo0'lpug s|tpolstroqJ'p oapwd '((€g6t)gII :S pruourug'lle]torqJ .S eprJ'otcal)(snuoE roltsl oql ur urlorxD t (rotsroCC T ralsroC)aptuB m:yoo)s? pup snua8 roruroJaql rn relsroC C T relsrod apuE DteluDgsE) p (ratsrog g p nlssogl orptuB 'trSolll.ruou,^l\ 'enun[ 4 :adKJ G06I) 69 ueqd uaD xeT'oztun) lsod ur D!$luoqodtJ 'l,es 'tl8elll.tuouT J 'urloruC oNIuDg- (16/l) :Z teN }s.{S DplooJ- suoc urou '(Z8 ' tZ '(9LLI) 'L 'relsroCf ) J uul'I uou fs|u]ou n t Id uoC rEqJ g T letsloC,isryar8r

I9I rsa9parlourlqJuprlprtsnv uralsalA o{U I B 152 NuyrsiaVol. 6. No. 2 u988) membersof secI.Pimelea. However, the dioecy,glabrous flowers and penistentfloral tube of the two speciesare more typical of sect.Pimelea. Furthermore, in their vegetativecharacters, P. forrestiana and P. spiculigera are difficult to distinguish from p. microcephala,a more tlpical memberof sect.Pimelea.

In easternAustralia, Pimeleapetrophila and Pimeleal/aua are somewhatintermediate betweensect. Pimelea and sect.Calyprrostegia bur appeaito be bestplaced in sect.pimelea becausethet bractsare shonly petiolate.

2 Pimeleaserpyllifolia R. Br., Prodr. 360 (1810).- Banksiaserpyltifotia lR. Br.) Kuntze, Re]is.Gen. PL 2: 583(1891). Ttpe:Bay,1 [Perrel Bay, Isle of St Francis],, 3 Feb. 1802,R. Brown (BM\.

_Description as for Pimeleaserpyllifolia subsp. occidentalis below but stemssometimes gla0rous.

Distribution.(Figures 5, 7.)In WesternAustralia extends along the southcoast from Israelite Bay to the SouthAustralian border. Also occursin SouthAustralia, , Tasmania and New SouthWales.

lJflities. Closestlo Pimeleahalophila, alsoshowing affinities to the easternAustralian species P. hewardiana.

Notes.Two subspeciesare recognised.

Key to Subspecies

l. Stemsglabrous. Flowers usually glabrous inside...... 2a. subsp. serpyltifolia l. Stemshairy for a few internodesbelow each inflorescence.Flowers at leastspanely hairy inside...... 2b.subsp. occidentalis 2a. subsp.serpyllifolia

Pimeleacluytioides Meissner in Lehm.,Pl. Preiss.2: 27| \1848). Calyptrostegiacluytioides (MeissnerJWalp., Annales Botanices Systematicae 3: 324{1852). Z}pe.-i,ort phllip, Vicroria, 1813,C.J. Latrobe (holo vel iso:NEU, n.v.,fide S. Threlfall,Brunonia 5: 163-41993).

In WesternAustralia this subspeciesis known only from Euclaand is asdescribed for the other subspeciesexcept that the stemsare glabrous,For a descriptionof this subspeciesin easternAustralia see Threlfall (1983:163).

Specimensexamined. WESTERN AUSTRALIA {selectedfrom 6 seen):Eucla.I Seot.1963. ME Pftil/ips{NSWI. SOUTH AUSTRALIA (selectedfrom over280 seen): c. 3.5 km S of Kingscote,Kangaroo 11.Eichbr (AD, 'S1,zon.Island, ll?52 BRI, CANB); lower Coorong,20 km S of Salt Creek,-DE 10416(ADW). ItqY lgyTH WALES:23 mi [37km] W of Euston,t8 Feb.1955, T. & J. t /harle(NSw). VICTORIA (selectedfrom c. 50 seen):Bellarine Peninsula, Barwon area,2 Jan. l9g3,J.C. Krssane(PERTH): Rye, lIl. Ross 2528 (AD. MEL): TASMANIA (selectedfrom c. 30 seen):King Island.26 Aug. 1951,E Srnrri (HO,pERTFI): EastSister Island, lS Whinray52,392 (AD, HO). '{E 'S 3^oJ rq8rrr,rfo} 'S,rt0gt) ,t009ZI .6l,,Zt)'E E,(,g"tZI .{Egatrloplsl ruo4 tspoJ eql Suolespuorx3 ('/ setntug)uoqnq lstg 'HJUAd) .re^oc lL6t la\uul ?'pun8r€J Jo s u)t l.t :(HJuEd) 9691uost!/Il9A rul lurodJo N 0l :(HJuEd) g96Suostlll 9 dr'punSrEJJo ,'\\srul 0E c :(HJdEd) €l6a uos tulodr?eu:(CV) suosrod ull .r:(H-tUEd)grgl 14 9 d'ra1'oQ'Ipprqol{3o:) 16l V,BunBreJJoS l.Z I atuoag 5'v JoS ut SI :(HJUEdyyggg a|toag y,o,to3 rqirfii:iiiif,i '9161 5 .paulwDxa s lootq d t rdas'.{Egatrteelsl :VIIVUJS1V NdgJS:l,11 suawfiaag ',{lp ',{tuntEur ,{lqBunsord tp uaestou 1r"{J sro,roUel?u ur Suol urru g 0 > Jo NasqEapollUsH olrl-qsnlq€uBus :uJurg l-l ,{q ',l^oleq .c palresxa,/(/S snorqElS'8uoliuu 8 0-i 0 srEqJo Unt l?aldeuz qlr,tr'3uo1ruru 1 hDt1,Ouw x loqtue :8uolurut tueuplrJ:sJe,^\ou alEuaJ sapoutwojs 'acuecsrqapZ 0-1 0 i 0-SI 0 E'0-900 Jo JaUelpJotpl-ruas $lls :u[u x Jeqtup:8uol uru g tuaruElrJ t 0,SZ0 9 0-t 0 '8uol 's]e^o0 S0 :sl?dosuBrlt rolJor{s sr,/arrr]s snorqelS'xad€1e pa^lnJur utul t I g 0 Jo al?^o ,{lp?orqslrdas Suoluu S0-2 0'peFuet pu? papncteq^\ouros'luetBd lo osloltu€slreq:aprsur 'truruns lnoq8nolqt,tureq'eprstno snoJq?13'as€q l€ Sul]]rTds ,{lrplet lo ltnlJur lualsrsJod 'ruBrp tE u.lrxZ I-8 0'uru g l I x S o^oq€ponurluo3 ^laJJers s/auogfa1nua{lo aqnl 'pel8u?lZ Z',{]e^o llual€d,{ltsoul l'Jol! EuoIuru t 0 I 0 puepoFnJ l?q,reuos slleq:u{.ul-9 0lt?lsrpur snorqelSpue e^oqsfureq ,{lesreds oJoru'esEq l€,{lrzq ,{lasuep,tua^uouo'os ^lasJeds ^luo 'sseq soiurlouros1nq eprsur ,tur€rl J?ouSuol ulul I slr?qluated ,reJ e qllr ,{lal?llo eplslno 'lruuns 'oseq t 0 0 '8uol snorq€18 le uprp ruru te lopuels uru uauog[ alowp aqnl ',uolle^ 8 0 S 0 9 7 7 lotol! ujzeJalllznsn qsruealSlo ,r\olla,{ol atrft yaMoll Suoluur I,9 slr€r.llse8uol 'tceduoJ0 'lalqcu€lq ,{rEq,{lesuop'3uol ruu ,0 t 0 ,{llensnsp)rpad pere^\otJZI t ,{lpnsn Iplateluoqs,{ral Euo uaq,{^J€llxe Surjeadd?saurlouos tnq arraJsalo/r4 sno:qe18 'urur 'moloc leutrl-ljol'V 9 t-Z Z x 9 9 Z'et€^oqo,{lpeorqot e}E^oqo ut elrf}eel roe stJDlq '8uol 'xodB lDDryo^ul ruur I ot dn apunpad asntqool etnJE t€ olcnu uu qll{{ pu? po^lncol rmpJrns lerx?qB'sor]Bl lerxppeuo e^ecuoJ,{ltq8rls ro leg'u.rur S Z-l 0 x 9 [ qtpr,r,rqtSuelerues aql urqtr,relppru aql o^oqE,{llqArTStsopeojq lo JUdrIa,{lppoJqot Jrtdllle,{l^\oreu 'uee:8,Ier3:o uoaJ8unrpour 01 opd 'snololoouooeurlu€l :8uol uru ot dn eloued:snoJqelS 'asJoJlup 'pJp,rola'alrsoddo t 0 Jo luated saxfiaTSnoI {lltu t 0-1.0sJreq:yode uo{.req]JnJ snoJq?I8 SuIuocequoos 'a3uacsaJoUur qceo rBou ,fureq ,{lal€lepou ol IIgsl€ds'xode uoJJ laqlJnl ,{el3 'lc?lq aledot unrpou Suruoceq ol ,(aJB usqt pup sJuorsajolJurr{ceo l?ou u,r olq {rBp 'lo^el lJ€p .,tll?nsnslua]s' e^oq€,{qsnq puB peqJuerq,{u€ur punot8 t? polxuats,ol8urs,peolq ur 'q8rq 'atel]sord 'qnJqs Z ol dn u I'1 0 tsourJe(suolt€nlrs posodxa uri saurtouosJo ]JeJe 'aqnt lerou arll Jprsur,{rprtSurJq ur pue JJuaoseJouurqf,?o i(oloq p s:pour?lur,\\al uo ,{rrpqSu]€,q ur otlollltd.tas dsqnsJB d.Duohlldias dr uo{ sroJJrC '{l:lJ,{ ') 'SNVJ :osr:HJUgd '9 '996i 'la^o(I :oloq)1691. uostlA d tdosi'elleilsnV ulotsriA lurodJo S rul 0l .sr7d,{J 'olnlo^a srlEJoulqnl slor.loturarJq ut 13ue[uocsaJouur ErJursrllnuuou srrpouJolur ur otuounplcloryoli\tdas dsqnsrg doxotlllt{dt s d e lJelJr(l '^ou dsqns'e,{dsleluoprJJo dsqns qZ 'soroeds ,rou e eq ol st?add€lueue^ srqJ €uolJl^ ur surEJd Jltlesequo ]upuE^tuaJseurds patunts ? Joecuerrncco oqt polou(t9l :fg6l) II?JIOIIJsrloN 's3l?l& qtnos ,reN tse,r-qtnos '(Bru?rus€J) Jo euarlxoeqt uo4 proroj euoqllr trEJtSswg eqt spuelsruo puB?uot'l^ tsotu 'erterlsnv '(E,tgogzI Jo Jo qlnosul31s3a qtnos u! s.€toJt)Blrna l? smcao1/ ernSu) uounqltlslQ

€El seaap:plal!{qj ueqerlsnvuretsar\A iIU l g 154 NuytsiaVol. 6, No.2 (1988) Habitat. Occursin limestoneareas close to the coast.

Floweringperiod. Jull November.

Deriyationof name Occidentalis\L.) - western,referring to its occurrencein the western part of the speciesrange.

3. Pimeleahalophila Rye, sp. nov. (Figure3.)

Affinis P serpyllifoliaeR. Br. sedfoliis alternisminoribus, floribus extustantum pilosis et ovario in floribus femineisglabro diversa.

Zypzs:take King saltlake, Western Australia, 4 Oct. 1982,B.L. Rye 82035A(holo: PERTH, iso:CANB, K, MEL).

A

E C

B

Fig\tre3 . Pineleahalop&i1d. A smallflowering female piant; B. femaleinflorescence and bracts (x 5): C maleflolver (x7.5);Dfemaleflower{xl2);Estamen(x10);F'ovary(x12).Drawnfromfreshmarerialcollecredatlypelocality.

Relatedto P serpyllifoliaR. Br. but differs in the alremateand smallerleaves. in beins denselyhairy on the outsideof the flowers and in the glabrousovary of femaleflowersl --1

'09 1uone4snlll .adq ,29-19 .lpuBlrog.cos .le^ .lsurj ..qpur.I (/9g l) 09 1 :6 ,1/?p1rlt DapwH ' ^u\c:'.trl araryto!qD7 ap .H..tr.e6Ll.coc.lprl?rrsnv urru.^,nn[T*u"Jr.*,H;i?ili .t.I6gl) ,{role^rosqol? polcolloc,{lq?qordl .. ueu,rl(-ue1 rtldpJ,, .ad gg9:7 .14.uag:sr,rdX 'ezlun) _ .ld .lloH .^oN ..llrqe.I ([lrqe]) oto^ol) Dts4uDg (S0gl) I I :l ulu,rz1tealaur; .t

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Srens greento greenishbrown andhairy neareach inflorescence, becoming medium to dark brown and glabrousfurther from apex;hairs appressedto patent,either all similar or somelong and othersmuch shorter,fine, the longesthairs 0.6 L5 mm long. leaves opposite,patent or antrorse;petiole 0.4 1.4mm long,hairy; lamina discolorous, pale to mediumgreen on abaxial surface,medium to dark greenon adaxialsurface, narrowly elliptic or nearlyso to almost linear,{6 )l t-43 x (1.5-)2-9mm, flat or themargins recurved, more or lessacute, not mucronate but with an apicaltuft of hairsresembling a mucro and up to 0.5 mm long, moderatelyto glabrouson adaxialsurface; hairs appressedto antroffe, denselyhairy on abaxialsurface, ''(or very fine, the longestusually 0.8-1.2mm long. length of stem between inflorescenceand uppermostpersistent leaves) up to 25 mm long. Uppermostlmues subtending inflorescenceusually 2, deciduous,shortly petiolate,similar in colour, shapeand vestiture to leavesbelow, 2-5 x0.6 1.5mm. Inflorescence tetminalbut often appearingalmost axillary at fint, compact,many'flowered. Pedicels 0.3'l mm long,densely hairy;hairs 1-1.3mm long. F/owerswhiie or cream,rarely paleyellow, denselyhairy outside,glabrous inside. Maleflowers: tube2-3.5( 4) mm long,0.2'0.5mm diam.at middle,expanded to 0.5'0.6mm diam.at summitl sepalsovate-elliptic, 1.2-2 mm long;hairs (of tubeand sepals)antrorse, 0. l-0.6 mm long,the longestones, which are 0.3-0.6mm long,either confined to sepalsor on both tubeand sepals; stamensshorter than sepals,the filament0.4-0.7 mm long and anther0.4-0.8 x 0.3-0.5mm, the slitslateral after dehiscence;pistillode present, the abortedovary < l mm long.Female flowers:tube 1.5-2 x c. 0.7mm, continued 0.2'0.3 mm aboveovary-portion, 0.3-0.4 mm diam. at summit, circumscissileabove ovary in fruit, the portion below the circumscissionpoint persistent;sepals elliptic or nearlyso, 0.6-1.2 mm long;hairs (of tubeand sepals)most densely arrangedon sepals,appressed to antrorse,c. 0.3 mm long;staminodes < 0.5 mm long;ovary with an apicaltuft of hairsup to 0.4 mm long;style exserted by 0.4 0.6mm, the stigmabrush- like. Fr"ailsucculent, black whendried but probablydark purplewhen fresh,3 4 x c. 2 mm, with a few hain on outsideof p€rsistentbase of floral tube. (Figure4.)

t\ \ i\\' Stiin\ N\N(r\

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\ nnl t -l I e*o','vl ot e"]'.'""$ \ a tl-l .tf ,i'\ IT l Figure 5. Distdbution of Pimeleagilgiana L, P. halophita a, P. clnvata a, P. foftestiana a' P. spiculigeruvar. spiculigera v. P- spiculigera -rar. thesioidesL and P. serplllifolia *.

5a. subsp.microcephala

Pimeleamicrocephala var. elongata Meissner, Linnaea 26: 350(1854). Type: South Australia, 1854,F. Mueller(lecto, fide S.Threlfall, Brunonia 5: l6l (1983):G-DC, n.v., microfiche seen). lsuelpru paup urorJumelc {92 x) lo,roLlal?u.ral -c :(a x) strul pawu t qtr^\ ocuocsarolJuroleuroJ J :(02x) r.^\ou alpLLI-g :luEldel€L! Jo uralsSuua^\olj Y olDqdnoDu o4awu g aJa?]l

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Habitat.occurs in shrublandsor sometimesin woodlands, usually in sand,sometimes in rocky groundor on dunesor floodplains. Floweringperiod. Mainly May-October,also February-April.

Nol€s.The tyr materialof thissubspecies, bonowed from BM, wasvery poor, lacking flowers andfruits. The microfiche photographs of the lectotypesof the two vlrietieslisted above werealso poor but they appearto be of this subspeCies. 5b. subsp.glabra (F. Muell.& Tateex J. Black)Threlfall

Pimelea-microcephalavar.glabra F. Muell.& Tateex J. Black,Fl. S. Australia - rst edn3: 3.9709.26) Pimelea_glabralF. Muell. & Tateex J. Black)Ciroiin in Jessop,Ff. Cenirai A-ustralia222 (1981\. lype: Mt Illbillie,Everard Range, South Austratia, 5 June1g91, R. Helms\lecto, fide S. Threlfall,Brunonia 5: 162(19g3): AD, n.v.;isolecto: MEL).

_. Fgl l.dgTltltion-of thissubspecies, which does nor occul in WestemAustralia, refer ro Threlfall{1983:163).

Specimensexamined. SOUTH AUSTRALIA (selectedfrom 12 seen):Camp 4 [nearMt Illillinnal,9 June1891, R. Helms(AD, MEL, NSW).

Distribution.(Figure 8.) Knownonly from a smallarea in north-westemsouth Australia.

6-Pimefea spicufigeraf . Muell., F-ragm.ll:46-47 (r81gl. - Banksiaspiculigera (F. Mue|.) Kuntze,Revis. Gen. Pl. 2: 583(1891). Type: near Russell Range, Western"Ausiralia, daie unknown,A. Forrest(holo: MEL). a otqpq dsqnsqoqdao.olw d p]ug a opqdaJootw dsqnsolD4dauJlu oalauu lo uo[nqusrc 8 oJnBIj

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Figtrre9. WesreraAustralian distribution ol pimeleamicrocephala. --T

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l. Mature inflorescencecompact, not interrupted...... 6a \ar' thesioides 'tar' L Mutrta inflotescenceelongate, often interrupted...... 6b spiculigera

6a. var. thesioides(S. Moore) Rye, comb. nov. (Figure10.)

pimeleathesioidess. Moore, J. Linn. Soc.Bot.34 224 225(1899). Ilpe: betweencoolgardte unJ frf" Darlot, WesternAustralia, 1895,S.M. Moore (lectoherc designated,female: BMi isolecto:NY; syn,male: BM; isosYn:NY)

Inflorescence(or rarely inflorescenceunit) compact'not interrupted;rachis l-2 mm long

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'Joqurotdos-,{lnIpouad SuuaMoll ',{BIc,{pues ro puBS,,r\oll€qsm [llensn'sa8pu Jrluolel puu sdolJlno olrue$ uo smmo .?r1lgrH 'pe^lo^ur ^lau?^ eqt alns eq ot q8nouopo3ue^pB lou sr suorunodsesoql ur 'It\Z Jo Suue,rol]tnq lDgC siuawal)'y'W) peaaedthl pu? (HJUEd .6811ultdv .H.g.Jl 'S ,S aaueuros?rg ot spuatxaoq? .{lqrssod(A ,SZ0€ZI ,tt00t) aal$punJ pue(A 'S ,[ ;LT,I.uo!fiqlusle ,Lt,,Ztl Irplnn) ot rs?e-qtnos(A ,9€0SI I ,l togz)e.rolln] I reeuuorJ spuotxE( 9 orntrg) '(HJ{Ad) 'rellErqro ,ueuesroN gLIL uosM4 9 ;r reeu:(HJ{gd) I Llt toY O.y g] .o JoN lurl Iu.v S :{HJdAd) I uosutqou }.'arooIAJa{e.I :(Hldgd',-IaINl Kalsaax a 'arpre8loo:) 'W'.nuedttqqEu n s 10, !^pw JoS lmt tZ] ry I :(lAI,{) lpox ooroqre^\:(HlX:lJ 'Irelrtnx S 0gSI 'll t ttnqnH V 7 :(HJXAd) 9t9l uaa.tg I,EMaIInt{ lur)t rru :HIdAd) 'aruore)I Jo E 9.1.yJI I ,uolol8ursI 1169aErcat S'tr Jog [ru{ I S]Iu ZE:fiJUAd) 6ZI ZI hupto) '{.oU ,8€61.Bnv ,uorrelsU I I(JAJ) 916dsuJ Oy'tl uosraqdct{reou :(HIXEd,) agprqng J N uJouatO:(SNVJ)/09 tappotg A t'purC souf?dJo S lrxl gt] :(HJXAd) 'uoslJef [u 0E .paulaoxaSSLIpDag Sf iW :(uaas0[ ro^o ruo{ palcelas)VI-IYUJSnV NXEJS:I,U suauDads 'r,{U s9i a?acsa?lelu,{qIuDrlDl]snv urotse/A 'Ig 166 NuytsiaVol.6, No. 2 0988) the axils of the uppermost(leaf or) pair of leaves,which are petiolateand not bractlike. Pedunclestpto3 mm longin maleplants, up to 8 mm long rn fiuit. Invotucralbracts absent or I perspike, early-deciduous, ovate, commonly translucent yellow-brown with much darker veins, 1.5-2x 0.5-1mm, glabrous.llpr'kes' compact at firsi, soon becomingelongate and interrupted,many-flovered; rachis usually 5-25mm long at maturity, with up to-3 mm-between groups of flowersor fruits. Pedicels0.1-0.4 mm long, glabrous. Male fowers'yellow, completely glabrous;floral tubenarrowly cylindric to cylindric(except at summit),3-6.5 mm long,0.2-0.3 mm diam. near middle,0.7-1.2 mm diam.at summitl sepalsovate to broadlvovatJ. 1.2-1.8 mm long:stamens shorter rhan sepals. the filamenr 0.7-I mm longand anrhei 0.8-I x 0.3-0.5 mm, the slits semi'lateralafter dehiscence;pistillode absent. Female Jlowers not seenexcept in fruit, probablysimilar to thoseof -&rnelea spiculigera;floral tube persistent in fruit; sepals erectin fruit, c.0.6 mm long,tardily shedfrom floial tube. Fruit dry. Seedc. 5 x 1.5mm, with a prominentpattem of irregulartransverse ridges. (Figure I l.)

D

|igute.l|. PtmeleaIo eslfuda.A- flo\aering (tem of maleplanl; B.male llower (x l0)lC tematelhrrrina inflorescence: * D-enclosed fruir and sripe rx er.Drawn Trom u.r. tiigi, 2z1i,a. e, ,no V,i s|".;ai;'iti d]*''* t-Ie88S pBduroJ'lcolosralsnlo Jo p?eq :leJqou?rq ^Jqpx€ uoqs ,{re^ ? Surl€urulolresnlJ qc?e'sJelmlc .{relllx? Jo peoq e acuacsatofu1tuesqe sQoJq ptmlo{r, olssas snojq?I3(?rl€!snv Jo Ieuutnt 'snorolocuot urelsai&ur tou) .{larpJ,4JeA Jo'8unof ueq,r ts€elle',furBq,{Jo^ 01 }Eq,rouos 'sorceds ua]Josa^Da'I Eu[xBI Jo es?qJo elorladsE lcrqt s3 ecr,rl ,{ll?nsn snomeqrequr s^e,{lelou lnq solcodsqnrqs ur luouruold f erxeqesepou :snorq?IB {?IlEllsnv ualsa^\ ur tou) fleJBrro Suno^ueq,,\^ ls?ol lB,{r?r{ vua1s snonoouolu(Brl?Isnv uralseldur lou) ]o snorcoop ,tlerEr'snol3eorpouf B Jo alrporqduturarl ^llBnsn 'sqDq ^IaJvJ Jo sqruLlsaafil o\ apt$o.td

'(palBu8rsoporeq otco Da u 'd ''J( :]rqe]r :adq (L98ll 60S :il rpord ur reussroJ^laD tqdtsDe I oalaudng tres oapatd 'tg 'eztun) oaluaato zdtJ ueqd uaC xel lsod ur ezlun){ A 'l.esd C06I) 09 ' ry J I lpua) sKtlrDisryDw oteluDg- (tL8l)t:9 I€rtsnv lJ qtuefl \'lpull s[qro1sryow'Fes DapLuld- (St8l :t Srnoqsrated-tur?Srrs drq pprv qtptr s,{qd ) i/ 'ld ' tr IJ llnfl'ra.{st\I J I lpu]J stwolsltDw Il D8alsorld(lD) - (1,t8I ) l €f uef,) lpug sKLlrDtsttDWe oapuL! .((tg6l) :g Eruounrg 'll€JleqJ 0/l S eprJ'olxaIlrg dDrob?un|d 2dtJ (S061)09 u€qd ueC x?-I'azrun) T lsod I ur aztun) l IpuE)aBqlDdE ]:es Dr$luDg (Sr}il nL:t Srnoqsretad-lurEsrrs dul p?rv 'qrel^l-sdqd 'III l a8DttDdg olSalsotidttD)- (r€SI) I€t Id uo9 'puaa8qlodgIJ llng'ro.{ot{ J lputr) J oapwrd- (S/81)S :9 IBrlsnVIJ'qruofl (lpug) aSollodgl.,es oapwq

aSBIIBdgt rces 'qlq Jo o8usr or{l qlltA pesnJuo3uooq seq eueu s,uosredre E[ oq] sdeqled'(01.61 le uolooC)^olsleusH '6991 Ie uloclpt{ ^q perueduocau ur sorJodsoqt Jo aBUEJu,rou{ aql qSnojqt uorlrpodxo u? el?lopun prp aq qSnoqll?'e8ueu ,{elsjatu?Heql ur susurrcodspellos lou prp,(lq?qoJd tsorroCuqol ]3eroo aJBe^oqe potri lorcolloJpu?,{trTEOol od.{l or{l leql }qnoparuos sr oloqJ

'3rum ro sruocqo ,{l,rorJeuoq ot Surpuot'tlnrJur pe8lulueq3nu auooeqsloctped oqJ snue8oql lo sloqurour reqlo lle ,{lJeeuur sp p?eqro o{rds ,{J?trlose u?ql toqlel soltdspel?rcoss? Ilosolc aelql Jo slsrsuoJ,{ller6ue8tr esnmaq lsaJotul J€lncrued Jo srserceds srql Jo ocuocsoJol]utaqJ saJoN 'sobeds uqt lopun sotou eag nta8nntds Daputld ot 1saso13'sawu$V

'roq{.uoldesaunl pouad ?uuauo1tr 'soprqlrq ,{{oor pu? sdoJrlnoolru€I8 uo sJn)xo tulqoH '(A 'S ,lto8l I ,L0,EZ)elog sule8uvruo{ uortrallo.tuerar ? sr aloql pu? 'S 'S {g .0zollI ,EgolZ)uolr{d tt^{ urorJ^lpouoder sr uortrellocad^t eqJ (A ,ror^llrH 's,90082) ,Ztogl's,9s062I ,tS09Z) r€ouol ts€a(g.8€0s I I SuuellBJltr\tr ruo{ pus (a,000/II 3)slool{ o{E-I 'S .uollnql4sle reau ol qlnos (E ,fl01l I ,rZo9Z x) urnolltrAll€ou rxorJ spualxA( S eJnArJ '(HJUSd) Dt ltj g'urqnl(\ Jo N lurl 90t] Iru 99 :(HJUAd) 9777 ua8pru1 g'ht'\ead suuollel :(HJUAd 'JN 'SNVJ 'CV) 'H'N 'elnetrl{ ru :(HJNgd 'tlNYJ 'CV) 1.66ryads JoiA [urI 9 t ] 0 t ,LB )trads71N'psslseuoH ,^eI^tlIH Jo g [u)t 9 6] IlU 9 :(HJUgd) 'oo8le^ '.Iahl)It€€l raup.tDDVJ rEau:(HJd gd,)ISZZ nupi,D p/,'urtsny al?l :(HJUAd 9Z' qllotqlDD lW:(HJUAd) dolt1ne ounf'sela8uvlsel&:HJUqd) 1'SuuollpJ'anJ N t'tB6I 6/ llDlrotg E 11 pue teu8?tr{tunol^l uoo,{\teq:(Oy) I6ZE tqqsv W'pautuoxa V'oo8rnw'ro^rx proJpu€Sreeu :(ueos0t o ruo4 patrolas)YITV{JSnV NUIIJSA,{I suaaDads -I L9I aeecea?l0Lu,{qluDrlBrtsnv uralsrl\'o/iu g 168 Nuytsia VoL 6, No. 2 (1988) at first, sometimesbecoming elongate and then commonlydiscontinuous, 2 many,flowered; rachisusually not enlargedlaterally more than the peduncleor upperpart of stem.Pedicels hairy. Flowersalmost always hairy outside,glabrous inside or sometimes(not in Westem Australia)hairy insidethe distalpart of floral tube. Floral tube circumscissileabove ovary; ovary-portionlonger or (not in WestemAustralia) shorter than style,pofiion.^tepals sometimes erectand apparentlyconnate in a definitebasal tube (andthen the whole structurereferred to as the limb), sometimesspreading and apparentlyfree. Stamens 2, sometimessubsessile in throatof floral tube;connective often broadbut laterallyexceeded by the c€llsislits adaxial to semi-lateralor rarelylateral after dehiscence.Ouary hairy at apexor very rarelyglabrous. Fruit dty, enclosedin the p€rsistentenlarged base of the floral tube.

A sectionof 19 species,represented in all Australian states.

Notes. The three infragenericgroups Epa llage, Choristachysand, Malistachys wereall named in the same publication. Severalspecies were listed for Epallage blut only P. spicata and P. argentea were listed for Choristachys and Malistacftrs respectively.These two speciesare both readilydistinguished from other speciesof Pimeleabut are not consideredhere to be sufficientlydistinctive to warranttheir separationinto individualsections. The formeris now includedwithin sect.Pimelea and the latrerwithin secr.Epallase. Bentham (1873) added to Malistachysthe rather distantlyrelated species, P clavaia. wiich is hereincluded in sect. nmelea. Benthamand other authoN addeda numberof extra speciesto Epallage,most of which are includedhere. Bentham (1873) and Threlfall (1983)used anther type as the main criterionfor distinguishingsect.Epallage, attempting to includewithin the sectionallspecies with a broadconnective and adaxialslits and excludespecies with a narrow connectiveand lateralor semilateralslits. This resultedin a very artificial group includingspecies which, apart from their anther type, were typical of two other sections,Calyptrostegia and Heterolaena.The criterion was also unsatisfactorybecause quite a few taxa, such as P brevifolia,are variablein anther type, somespecimens having semi-lateralslits and others adaxialslits.

A uniquebut not universalcharacteristic in this sectionis the occurenceof erectseDals. which are distinctlyconnate at the base.There are neverany sessileinvolucral bracts-and the rachis is not distinctly enlargedin comparisonwith the peduncle.The latter two characteristicsare not found in any other sectionexcept very rarely in sect.Pimelea.

8. Pimeleatrichostachya Lindley in Mitchell,J. Exped.Trop. Austral.355 (1348).- Calypttostegiatrichostachya lLindley) W alp., Annales Botanices Systematicae 3: 325(1g52). - Banksiatrichostachya (Lindley) Kuntze, Revis.Gen. Pl. 583 (1891).Iype: ,,subtropical New Holland" [Northern Australia], l8 Oct. 1846.T.L. Mitchell {holo: CGE).

Annual semi-woodyherb, ercc]'.0.l5-0.5 m high,single-sremmed and deeppurplish brown at groundlevel, hermaphrodite. Young stems pale green to paleyellow brown, with scafiered hairs at first but no axillary hairs, soonbecoming glabrous; hairs l-1.5 mm long.Leaves alternateor rarely subopposite;petiole 0.3,1.2 mm long; lamina concolorous,pale green, narrowlyelliptic or linear,2.5-16 x 0.5,3.5mm, acuteor obtuse,with scatteredhairs ai first, soonbecoming glabrous; hairs often confined to margin,especially near apex, 0.5-1 mm long, Inflorescenceterminal, compact and closelysubtended by leavesat first, becomingelongaie and interruptedat matudty, up to 90 mm long;rachis more densely hairy than youngstuns, the hairssimilar to thoseon stems.Pedicels 0.5-l.2mmlong; hairs0.5,0.8 mmlsng. Ftowers usuallygreen to gre€nishyellow, rarely white to red;tube 3.5-5 mm long,circumscissile usually 0.5-0.75.mmabove ovary portion. Ovary-portionoffloral tube 2-3 x c. I mm, with a very densemixture of long and much shorterhairs; Iong hairs antrorse at first, becomingpateni, 2.5-3mm long; small hairs also becomingpatent, usually0.2-0.25 mm long.StyE itonion '(lan '6881 'ellertsnv '(dnuofo) :osr:Cl :oloqpaunserd) I sslrld| tdes urolsolA 'dnre8leg) t9Z 7 t JoS ..dnlefteg,,rpau pu€ u,rolfl tIN.ad,{713p311 I9:t lddns Id ue3'lpuE {reusslel{) ot|luot w mEaTsotldKlD- $fgl) 109:l ssrejd Id'uqel ur rausstelNollluoulw Dapwtl '(olEur'IAI8 't{g 'erlel]snv :u^s:apuel :pat?uErsoporel{ olcal) uMotg U'(081 uef SI urelso^A 'lpreFl olpprl^tr',{EgpuElsl esooD] ( [ag:adq (1681)E8S:Z ueC sI^aU'ezlun)(rg .rcs .drulId d) oalua7tnqs:1ung - (St8t) :t Slnoqslaladrur?S p?ry qteyl s.{q4 13 11ng 'ro,{aW'J tt (tg d} DaluaBtDa8alsottdtqo) (0181)Z9tlpold'tg U salue8rsEalauld 6

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'?uolcr^ pue sole/d qtnos ,{\eN 'puelsueano'erlcrtsnV qtnos ',{JolrJJoI uroqlloN ur srnoco oslv Japroquellprtsnv qlnos eql ot ssorcppu? (A 'S.l_S,l€o) aBu€UresB.U aql ot ts?a-qrnospuB'laproq ,{lotureJ .8S,,ZZI 'S ulaquoN aqt ol ssorrepu? (g 3) uoD?lss8uuds ,raN ot lsee-quou(g 'S ,00002I ,0S"92 't c ,S9ol.I I ,l €ole J)enJ reoouo4 spuotxaerlprtsny ulelse,4l u] { t I I setn?tl) uounquNe p 'l1A uosaD rec'ua^no:(ueas ra^ouo{ potoeles) ) lryl AN'9161 SI 'uolSururodVIUOJJIA llug) KDtutID)DW :{CgJ) 'ertoseu 6Z8Z T O S/9i Suruun2try g orntpNSuoqte^ :(uaes0S c urorJpelceles) Sglv,,\A '(,^[sN HJnOS'lus) ,\\AN ') 'pU 'gNyJ 'ldfl) z6€I l/osl1,.14 sul?rueloJuo ,{,rH oruool,{Jo E ru)t6Z o :(JN 60tI MDqsuataO 'lEe,rooueJ :(u3os re^o uro4 pe133l3s) d Jo 3N [uI'(HJUEd 69] lru ',4A0y) tt 09 CNY'ISNAEnO 6t0Z uoLu[Sg'e'uolxo'] Jo EN [u]I IZI iur :{gNvl) :(uaes o uro4 patootas) f I 908IIH 7.4'?8urt€r€t{ 0/ VItvdJSnV' HJnOS \JgJ) uopDt 'cr '?8to 'JN ',{\sN 'tght 'f 'Iuf 'ctY) L8Lt 'uoilels ]w(Htdgd NYJ LsLS sapuozo'llv daac :(uoes ra^ouo{ petrelas) llalA 'nl]'i{')Jo S [u]f f ll lu S 8 0/ 'e8ueX IUOJIdUgI NUAHIdON s!tt!,AH t't96t rde59 resErCJo a [uI fS] rru g7 c',{,ri1 e:,4g :(HJUEd)6ZtS a8pruJg W'erlrooBv)Jo g lu)t S0€Zlruttt'{eerJ uorued:LtCv'Oy) uotuts sellC 'SNyJ) LSIZ 'p?or? Cr'uon?ls lmrSoloroetehl JoS lrul SZIIIu 8/:(HIdEd 6Z0t )pads ouruueNuo erreqt?Iool^tr tzl t}tf ntoY 'al€A HN JoS [ut] IurSI:(HJU!ld) 'lAW 'qV) eV leuoJ :(HJUgd,) artoY y'uorl?ls sguuds.raN :LtSN sapH '1681 'a8uEN Z00Z o 'uorl?ls A 8nV I Z .^\ou€g:(HJUAd) 0779a8rcag S U raqt?ei[solrC Jo r\,\ [uI r :(HJUAd) a8ioaD U'uo a^el [rx)ttE] IZ :(HIUgd) t0zl lru0ZI 'uorssrJ t60g S Jo AN lul 'ouotspues868S aS.ioagg y { oeleapunJr?au :(HIdAd) huptog V J't961 IJO9Z Joa ur)t :(AASN)plota8ztrg ,{tnf 'rat,rEl :{HJdAd) ultdy ldas 'upuesroN0i A,,u'6681 'e H E J'296t'anl S Jo E [ru)t8l I] Iiu t, :(HldEd) 86yH uosutqrnH T u Flv y Jo S lrul S /l]ru I I :(ueas0t 3 uro{ patcalas)VI]VUJSnV NUEJS3,4[pautut'xa suanoads 'slrd 'txtx Jo s,roJ leurpntrSuolI{1r.4^ I o x S e-SZ sprrs pepnJcul ,/.&S Suol urru I'9 0 srreq1se3uo1 aqt'srr8q l?rrdEqlr,r'8uol urut Z-Z I ttDto aJuecsrqap rau?leralel-lues slrls:ruu y S0-'0 reqtuz:3uol uru t0Z0 tuauElu:papnlrur 'etE^of'0-20 '13ejo suaLuoisEuoI urru g 0'1 0 ^lpeorq (seqolao4 ro) s/rdas Buol urru S I- l sjreq tsa8uolarlt'srrEri passeJdde qlr,u'truruns te uerpurx t 0-t 0'3uoluru 8 0-90 uoruodlersrp :srr€qtJor.{s tnoqlr,^ tnq uoluod fJs^o Josrl?q Suol eql u€rlt rouoqsuolJo tnq ol relrurssrru q auortueqtr,r (suouelsJo la^eluorlJosur ,roleq) uourod lerxrxoJd :8uol rurx / l c aqntlntofto 69r aeatrecElslu,(qluerlelsnv uretsart aid l g 110 NuytsiaVol.6, No.2 (1988) Pimeleashuttleworthiana Meissner in DC., Prodr.14: 513 (1857).Type.. south-western Australia,date unknoum, J. Drummond731 0ecto here designated: NY; isolecto:MEL); south- westernAustralia, date unknown, J. Drummond730 (isosyn:MEL, NY). Pimeleavestita Meissner in DC.,Prodr. 14:513 (1857). Type: Mt Matilda,York, Western Australia,1l Sept.1839, L. heiss 1265(lecto here designated: LD; isolecto:MEL, NY). "LakeLeven" Pimeleaargentea var. racemosaF. Muell.,Fragm. 7:7 (1869\.Type: [ocality unknown],Western Australia, date unknown,G. Maxwell (holo:MEL).

D

4, Figute-r2Pimelea aryerrea. A- floweringsletu or mareplam: B.flowering srem prana(- mle flower.and-p€dicet of femare reafrx 5r:D- {x l0t; E-fematt floweJ. ar.rd pedicet rx l5r: F- oiry r* iSi. O.u*iiTiirn f,ir-fioiuii.iir represenredby B.L.Rye 83002 {A. C. Dr andK. NeiybeytB,'E',Tr' 'reque^oN'^ln[ ilwelN pouad SuuaMoH

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tT Figurc1-1. Disrribution ol Pimeleaaryentea L andWcslem Auslralian distributior of P.micrantha *. P. Lti(hastuchrLl r. rlDicalvariant ol P. amnachar\ :l and Mt Leakevarjant of P ammochas a. rh Al.linities.No closerelatives but greatestaffinity is with P. micrantha. e) sh Notes.A very variablespecies. Specimens lrom the morehumid areas, including the Darling to Range.are much more lush than specimens from the drierareas. Although female flowers of ol P. argenteausually have a well developedstyle'portion to the floral tube,some specimens s in the centraland easternpart oi the southcoast have the styleportion very reduced.

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Figure 14. Distribution ol Pimelea trichostachva.

Figure Is..Dsxributionol Pimeleamicrantha o, typical variantof P ammocharisI and Mt Leakevarianl of P. ammocha s o I

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'se^?el 'lrnlJ ur paxauolSurujmoq Jo vad? l? pa^mJel fFuorls Jor{ltoseulleuos ruoJJ 'snorounu-t(Z) polaluojoJJrpIJo,{{ ,tlpnm siJtuq ptJryo^ul elsses ,{J?q soulqeruos:o mo:qep 'oleuralle ,{leJEJJo sllsoddosr ra1 euru€l}o eseqJo olorlodsE lslql sem1,1 1 t'lueuluold .{llerx?q8sopou :sr?q pEoJdsopr,raJoru qll,{\ Jo sllx€raddn eqt ur sJIsI{snoncldsuocul qll,t\ suats snoboorp^larBr 'snorcaorpoufSlo ollpojqdeuleq ,{lFnsn 'sqruqstapunJo sqruqs '(powu8rsep aroq olcol) Ilanhtr d sltDqzoawD 'd 'JC :adtJ'(LS8]) 909 :tl rpord ut Jsus$strAlaDpsuqal E Dalaatdng Des DalaLuH '(pelEu8rsap a.raqolcol) rg X olDruqwt d :adtJ '(f061) 6S ueqd uo0'xa'I'ezlunx T lsod Iul (DuaDpolltlldsE) azlun) (pud DuaDplltLld 'q DleluoqodtJ ljes ol$lu'{l - (t/SI) 0Z:9'l?rtsnv lC't{luag (;pua) ouao1o11,tq4 'l.esqns ' 'r o8alsolJdtlDJ txesoaputd - (/€81) I S€ ld uaC lpuA Duaop illd Dapwld '(ll?JlerqJ ( looH xe uunJJ DutuadKl.! 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Involucralbracts 6-12 immediatelysurrounding inflorescence, intergradingbelow w ith theleaves, the same colour as leaves or morebrownish. 7-14 x I .5-,1 mm, moredensely hairy than leaves;hairs oriented as on leaves.l-2 mm lons in bisexual .2-3 mm longin maleplants and 3 4 mm longin femaleplants. Infloreseice terntnal. head-like,erect or pendulous;rachis compact or (in variantfrom Mt Leake)eioneate and up to 9 mm long.Pedicels up to 0.5mm long;hairs up to 3 mm long.Fiowers usuilly pale to deepyellow, rarely white or a combinationof whiteand yellow,densely hairy outside, the indumentuma mixtureof hairsof greatlyvarying length, glabrous insidi except for th; ovary._Floral tube of maleJlowen narrorviy cylindric, 5.5 I I i 0.6 I mm, almostuniformly hairythroughout or with longerhairs in proximalhalf; Iongest hairs of proximalhalf antrorsi or sometimespatent, 2-4{,6} mm long;longest hairs of distalportion usually patent, l-1.5 mm long. Floral tube of lemale or bisexualflo--ers narrowly cylindric with only a slight enlargementaround ovary,6-ll x c. I mm. not circumscissilebut sometimesbreaking irregularlyabove the maturefruit, with much longerhairs in proximalhalf; hairs antrorsi to patent.the longesthairs 7-9 mm longin proximalportion and I 3 mm longin distalportion. sepalselliptic or broadlyelliptrc. 1.5-4 mm long.tending to belarger in malethan in lemale flowers.densely hairy outside. Stamens: filament up to 0.2 mm long;anther 0.g,1 .4 x 0.,1,1 mm; connectivevery broadbut slightlynanower than anther: sliti adaxial.Staminodes of femaleflowers: anther 0.5'0.7 x 0.2-0.3mm. ouary with subterminaland terminalhairs 1.75-2.5mm long.Stigmo scarcely exserted or exsertedby up to 3 mm. pistillodeol male flowersincluded, with hairsl-2.5 mm longon abortiveovary. Seed c. 6 x 2 mm, with a prominentpattern ol irregujarbumps and hollows, which are more or lessalligned horizontallv into wavy ridgesand furrowsrespectively.

Specimensexamined. WESTERN AUSTRALL{ gelectedlrom over70 seen):Mt Bannerman. .LS Beartl5612 (PERTH); Millstream. M.t.H. Brooker2088 (AD, pERTH);Anna plains Station.N. { Burbifge 1398{PERTH); Woodstock Srarion, N. Z Burbidge 5975 (C,\NB); BlackstoneRange, R.C Carolin6037 (NSW); Willianbury Station, Kennedy Range, R.j. CranJieldi905 (PERTH);Vtaming Head, H. Demarz6110 {pERTH): Mt"Leatei ll4 L1 Fitzgerald1200 (NSW. PERTH)t Millstream. C,4. Gaulner6288 (pERTH): pass in Blackstone Range. .,1.S.George 5248 (PERTH); 22 mi [35 km] W of Warburton Mission._,1.S George. !]Zf nf, PERTH); Mclarty Hills, A.S. George14720 (pERTH)l Roebourne,"/."N. (PERTH): !_ylcliytsgn78 nearEdgar Range, K.F. Rennealty5561 (pERTH); 2t mi tj4 kml SW of LuluiguiStation. M. Lazarides6538 (BRI. CANB, MEL, NSW,NT, PERIH; Ant"teti Ridge,..1.SMitchell l12l (NT, PERTH);Onstow. S.p. ffeiffer 33 {PERTH);Abydos Station. F. Richardsonl3 (PERTH);Rudall River area,P.G. Wikon 10544lAD. NT. pERTHt. NORTHERN TERRITORY (selectedfrom over30 seen):Lander River. NM Henry 644 (BRI.CANB.MEL.NT. PERTH);I I mi [17.5km] Nof HetenSprings Srarion, R.A.'peny 1905(AD. BRt. CANB. NT, PERTH).

Dislribution.(Figures 13. 15.)ln WesternAustralia exrends from KennedvRanse (2401,+, S, I l5'15'E)east to theborder. reaching Mt Leake(17033'S. 126002' Ej i; rhe;orth and extendingsouth to nearthe junction of WesternAustralia, Northem Territory and South Australia(2605. l29oEl.

Habitat.Mostly recordedfrom redsandy soils on dunesor from rockyrises in aridand semiarid regions.

Floweringperiod. Mainly May October.

AJJinities.closest to Pimeleapenicillaris, which occursin SouthAustralia, . Queenslandand NorthernTerritory. Fi D

L t;f:' ""ff;Ft (€l x) Irp^o g :(tt x) uaurrrsC :(6x) r.^rou f :(9x)Jsal .g :u.rs ?uu.^,o rrr ;',18Y,, il tl'; t C B ffi =fl lf{{k wt qlffiNiluA

'slJ?rqprrnlo^ur snorounu ejou pue sa^€elra,^dolrBu aqt ur srsJJrplnq lg u DtoJuqwt d 01 poleleu '(1!\SN"Iar{ ') 'flNVJ :osr:HJUAd :oloq) 't861 'prterlsnv 'llrH IISVI uoJSalUS U ^oN S urolsaiA Suldoocl?pp[lJJo edols aS :sr?d,{J 'snquorsor0unu snqrl?.rcnlo^ursretoBlq te snquoltsn8upslrloJ trqJrp pJS lfl d aDlDJltqut d slulJJv

(91 orn3rC)^ou ds'a,{Uslo)lllusr8 Brlaurld ZI 'ale8uolo ourocoqot Supuataouasselouul oql ur pue stJ?lqlelJnlo^ur eql uo sjreqJelroqs 8ul^eq ur'snor3aorpuBql leqlel olrpolqd?ruleqSulaq ur,{lurBru sloJJlp pu? suoq)oauo 7 3olueue,l ed,{teql Jo e8ueleql Jo quou o{eel lIAtrJo lruruns oqt uorJ ,4ln[ur ro,rolJur polrelloos?,r lI sonods,,!{eue sB pat€oll oq 01pullsrp flluorol.JJnssr tr laqtaq,,hJo stlDqJotutaoil Josorcodsqns Bs€ pepl€8oJ lsoq sr 1r tor{loq,^{ aurulatap ot pepnls pu?palJollo3 raqunJaq ol spaou(002I plota8zlrgA ll uoroxel]f,xelSuls e uro.4,{luo u,1oul tueu?^ V ',{JotllleJ urequoN pue erlertsnv uJelsaiArltr,r roploq uerleltsny qtnos aql Jo uoucunf oql oi osolcpolJollm ueaqs?q 1r esnpceq 01€1s l€ql Jo lse,r-quou3rx3J1x0 3q1 ur Jnscoll3,r [Ew suoqJouaryoaprutd'E\eJtsny qtnos [roJJsuort3alJoc ou oje eJoqlq8noqtlv sa/oN

LLI aem?Jrlrru,{qIuprlErlsnV urats3rlA arid l'B i78 NuyrsjaVol. 6. No. 2 lt988t Shrub, erect.spreading. 0.2-l m high,single.stemmed at groundlevel. Srezs paleyellow- greenor red'brown neareach inflorescence, becoming medi-um grey to almostbiack iurther from glabrous. apex, Leavesarlernate. antrorse to patent,glabrous except when immediately belowan inflorescence;petiole up to 0.3 mm long; laminiconcolorous,pale green to bluish green,linear,,4 l7 x 0.5-r mm, flat or adaxialyioncave, acuteor obiuse.F"aun"ie rplo I mm long. Involucrol braclsnumerous, usually c_ 40, becomingr.ffexeO ,n iirli, ,iniilai in colour.to leaves,narrowly g triargular to linea;,6 x t 2 mrn,aaUrou, o, spunetyttaiiy ciliare;hairs appressed toantror.", d.5 r.S rnm tong, trr. iong"ii flIa:*,9.; I r.)1.1:]t mm tong. !:iryinside. pedicek InJtoresrcnceerect, compact. 0.2 0.5 mm long; longesthiirs c. I mm long. Flowersbisexuar, cream or whiti abovecircumscission point; ti'orati oe i i.i mm long, circumscissile at summjt of ovary-portion.Oyary-portion ol tuUe t.i Z.Sx c l mm,.very ltoiat denservhairv oursideand rrark-coloureduui itre cotoui hioo"n oy dri nuiri, glabrousinside: hairs.an(ror:e.rhe longe.l ones 2 5 mm |ong.longir rhan and les: fine rhan narrsaDove crrcum\crs\ronpotnt. Style-po ion o1lloral tubecllindric. J.5-5 mm long.c. I y"liir. wirh,iongantrorse to paGnrhairs and imalter.patent nuirs o,iisia., :T,!:Tj.rlolqe;t.ones I-19 I l.J mm tongand_shortest ones c. 0.2mm long,with parentto antrorse l.ongoccuJJrnC lijll yj 1.: Tm inside. Sepals eltip_tic or narrowlyelliptic, 2,5 mm tong, with narrson oursrdesrmrrar to thoseon distalpart of floraltube, the longest glabrous ones1,2 mri'long, inside.Stamens \onger than or sometimes*ore-t-ua, oi le* equallingsepals; filameii 2.5-3mm long; anther 0.6-0.8;0.3 0.+.rnrn; .onn..tiu. ,lgfrtiynur.ower than anther; slitsadaxial. ovarv wtthhairs in distalpari; longest trairs o.o-iilm'long sryl".".iit ['uv 3-5mm. Seed not seen.

Specimensexamined. WESTERN AUSTR^AL_IA Gibbs,lS Norseman'Mt Vr Beard3111 (PERTH); HollandRd. C.A. Gardner* w.i Bt""ntil)+iipEnrnir Nit'cibb;,^F: Mi cibbE K. Newbey ty!![tl:?r!(PERTH): lpERrH); ojzifpeniHf; stlnnetRock, K. Newbey 7677 9.3km NNW of RoesRocr, k .\kr'bry-ii,i:i nii 1929,H. Steedman(PERTH). fpinrsf; cio*. ri.i

Distribution.(Figure 18.)Extends from ChiddarcoopingHill (10054,S,llg04l,E) nearRoes (33,55'S, southto {ock I19020..E).andt o, neriFi-ng;rinii[. ::,,i2; S, riS;iS; E)';# roSrenner Rock rJ20J5 S, l2l,ll tt.

Habitat. Occ]utson granite outcrops, in soil pocketsor shallowsoil over granite sheets. Floweringperiod. SeptemberDecember. Conservation pimelea status. graniticolahas a scattereddistribution oneplant was observed archildarcooping Hirr{A.s. weston pers. "#L:iXl[:#::1"",il:l populationsare alsosmall (K. Ner The speciesmav be rare and needsto be survevedro assessits .""r.,"J[? Fi1 l;l:comm') bra cra.niti ) - - - ?:i:::t:?:-!!:'" tL stanite cola (L.) to inhabit, referring to the occurrence ot tne\pectes on graniteoutcroo:. to pima* (bL r,,Clgr:f i^ttriranbut perhapsmore readily p. {:,!1 l.,o conlusedwi th vi ifera, frc from which it differs in having more numerouslnvolucral bracts. shr sor 13.Pimelea imbricata R. Br..prodr.J6 I (1g10)._ pe1 Banksiaimbricata lR. Br.) Kuntze, Revis. t, ,583rt8et)rype.KingGeorgeSound,liesiil"'a"lii.ilo.". laor,R.'Brrr; to( 3i;, ob1 ,{lesJ?dssaullauos 'socqJnsqloq uo ^Jr?q,{losuopot ,{lot?repou,{ ensn'olnr? ro esnlqo 'o^ecuoJ 'ruru ^ prxEp?ro t€[ (9Jt,9 x 'os ,{lr?oulo ortdua,{I,&olrEu .{lFnsn 'cndrTlo ol 0 9l- 'snoJoJoouocI J?aulT'uaor8qsmlq doop ol uoerEuinrpau ?urueJ:Euoluru S 0 ol dn oloned :alonadolruuep ? qtr,r Jo olrssosssel ro orolll'luot8d ^loJ?Jlo osjolluB'ollsoddo sourleuos '8uol ro oteuratlBsaADaT euole srreq Suol Jo sourlorxos rutu g 0-l'0 slrBqluelpd Jauoqs pu? Suol uur Z-l srEq luet?d ernlxru e ,tllensn(luaseJd uoq,r) unluaunpul :xedsuo4 Jo '(,{JBq lequnJ snorqeF pu€ ,{er8runrpour Surumoq flosuopJr ,{Ja^Istuu€odd? seunleuoslnq) ocuecsarogurqoeo Jeou osrrJ urdorq',rolo,{ ,{laJsl) u,rolq-pol Jo pel deoppu€ ^ll€q flasuep01 snorqelSsrrats la^elpunorS l? poruurols-nlnuueuo'q8rq ru (1,)90,2 0(,10)'peJe'qnlqs

'6201 uaqo) N iqperueseJdorlprrateu qserj ujorJ UMBrC 1Z L \) Fre^o I :lal xJu.Luers C :(/ x).lo^\ou-J:(S x) rJerq 8 :urots3uu3,rou .v tofoa Je^Dptlqar d durqrEolddE]!EuE\ E DtaSltldter otocuqut oalawtj.[1 an j

: C a\ \N

'0,{d 6LI o€oteeelaLu,{qfuDrlBr}snv urolsoiA l g --

t80 NuyrsiaVol.6. No.2 (1988) hairyor glabrous,rarely prominently ciliate but with no or few hairson thesurfaces: hairs anfrorseto appressed,the longestusually l-3 mm long.peduncle tp to 6 mm \ong.Involucral bractsl0 20 oJ sometimesmore, becoming reflexed in fruit, all sessileor somevery shortly petiolate,similar to leavesin colour or sometimespartially (especiallynear apex)deep red to purple,linear to ovate,usually narrowly ovate, usually 612 x 1.5mm, usuallydensely hairy on both surfaces,sometimes sparsely hairy or glabrousourside, rarely glabrousexcepi nearmiddle or baseinside; hairs appressed to antrorse,spreading rather widely laterally from themargin, the longestones on outsideor margin2-2.5 mm long.InJlorescence ercct, compacl. Pedicels0.2-0.6 mm long;longest hairs 1.3(-5.5) mm long.Floweri bisexualor female,white to cream-orsometimes pale to deeppink, with hairsusually 0.2-0.4 mm long insidestyle- portionof floraltube. glabrous inside sepals; tube 3.5 l0 mm long,usually tardilylircumsciisile 0.5-1mm- above ovary-portion, sometimes apparently persistent. Biiexual jlowers: ovary portionof floral tubeusually 2 2.5 x c. 0.7 mm, denselyhairy; hairsof olary porrionantrorie or sometimespatenr, the hairsin distalpart similarin lengthor gradinginto thoseon style- portion, the hairs below usuallyshorter, the longesthairs 2 3(.4)mm iong, the short h;in commonlyc. 0.2mm long;style,portion of floraltube cylindric or narrowlycylindric, ,l 7.5 mm long,0.8 1.2(.1.5)mm diam.at summit,either densely covered outside by a mixtureol longpatent hairs up to 2.5mm longand shorter patent hairs 0.1-0.3 mm longor moderately denselyto sparselycovered by longhairs outside; sepals elliptic to ovate,2.5 3.5 mm long. with a rathersimilar indumentum to that of distalpart of floral tube;stamens longer or rarely shorterthan sepals,the filament1.2.3.5 mm long,the anrher0.5-1.2 x 0.2,0.4mm, the connectrvenarower than anther,the slitsmore or lesssemilateral after dehiscence;ovary hairyat summit,the longesthairs c. 0.3mm long;styleexserted by 2.5 5 mm.Femalefloweri: ovary-portionof floraltube 1.5-2 x c. 0.7mm, densely hairy; hairs ofovarl-portion antrorse or patent,the longesthairs 0.3 2.5 mm long;style portion of floraltube shortly cylindric or cylindric,3.5-6(-7) mm long,c. I mm diam.at summit,densely covered by a mixtureof antrone to patenthairs 1.5 2.5 mm longand shorter patent hairs 0.1.0.3 mm long;sepals elliptic to ovate,1.5'2.5 mm long,densely hairy, the indumentum similar to thaton disralpart oi floral tube;staminodes shorter than sepals, with a filament0.4 0.6 mm lons and anther0.2,0.5 x 0.J0.,1 mm;ovary similar to thatof bisexualflowers but tending to besmaller; style exserred by 3-5(-7)mm. the stigmausually brush like. Seed 2 3 x c. I mm,with longitudinalrows of minutepits. (Figure 17. )

Distribution.(Figures 18, 19, 20.) Widespread in sourh-westemAustralia from the Murchison River to eastol Esperanceand inland to Sandstone.Also occursin South Australia. Floweringperiod. August March, especiallySeptember-January. Aflinities. Closestto Pimeleasubvillifera.

Notes.A very variabletaxon in WesternAustralia, with threemain variants occupying distinct geographicalareas. The variantscould be recognisedeither as subspecies or varieties,perhaps with equaljustification. Varietal rank hasbeen used by all previousauthors and hasbeen retainedhere because there appears to beconsiderable intergradation between the widespread P. imbricatavar. piligera and each of the othertwo variantsin WesternAustralia and because differencesbetween var. piligera and the South Australianvariant are minimal.

More field work is neededto determinehow completean intergradationthere is between WesternAustralian variants. Limited examinationof rhe specieson the coastalplain near Perth suggeststhat westernpopulations are of typical P. imbricatavar. maJbrbut that the plantsbecome progressively hairier towards the DarlingScarp and becometypical of P imbricatavar. piligera belore the scarpis reached.Indeed in WesternAustralia as a whole, thereis a tendencyfor plantsto be relativelyglabrous near the coastand progressivelymore hairy the further inland they occur.

L otaSrytd'ntr'c1 "" " " .' ' ...... 8uo1 ,Euol urur g-9 aqnl l?rolJuoql Suol se eptM s? ^lnou Jr uru 1.9.g 0. sroqluy lruruns ru[x urqll,r ot lo lrlurunsBurqo?ol '8uo,lJo I II0,r oprsu slmq qlr,r ulu slo,rolJ eqnl .€. ( 'po8uq-{urd0l-9 lenxeslqJo lBlolC fleJer ^Je^ 'ulBe$ lo ollq^\ sJo,rolC.Z .' " ...... IUId otnqqlu! r?^ qtl doopol al?dsJo,rolC .Z ll 'ocuecseJouur r{J?ol?eu ^JrBqs,{?,{AIe ^ll?eu sutols.slrsq lJoqs u pue Suol Jo omtxrru E ,{q pelo^oc ^lasuapoqnl plolJ uorlJod-al,{ls.I - ...... snotqq3 Jo.8uol IO|OU'Je^EII sue]s lle slrpq '.{Jreq .I ar.lt flasuop ,{lolBlepout01 ,{lasl?dsaqnt l?lou Jo uotuod-el^ls p T

urolselA pue V ,./arll/d te^ oloruqtut Dalautd uoruqlttslo atn8ll lo 6 J I

pLte Dur.tqtut .d ju,rl f rN^DtnlrEt!! d L toro!-!!]!!l!!!l a Drou Daputdro ""n"q",.,.a ?'l{'.,{i'u,g

3?ea?oelou^qlusrlprtsnv urals!/A o,{U-.I g 182 NuytsiaVol.6, No. 2 (1988) 3. Floral tube of bisexualflowers 8 I I mm long, with hairs inside absentor nearly so from uppermost1 mm of tube. Anthers c. 0.5mm long,nearly as wide as long...... 13d.yar.petraea

l3a. var. major(Meissner) Rye, comb. nov.

Pimeleamicrocephala var. major Meissnerin Lehm., Pl. Preiss1: 606 11845).- Pimetea imbricatavar. glabrataMeissner in Lehm., Pl. Preiss.2: 270 (1848). pimeleaimbricata var. gracillimaMeissner in DC., Prodr. l4: 507 {1857).- Pimeleaimbficata l. eracillima (Meissner)Benth., Fl. Austral.6:2l (1873).Zype: south,western Ausrralia, date unknown J. Drummond 552 (iso:MEL).

S/emsglabrous, deep red-brown.Zeaves opposite or alternate (usuallysome of each arangementon eachplant), medium green. linear to narrowly oblongor narrowlyovate to narrowly obovate,up to l6 mm long, glabrous.Involucral bractsmedium green, glabrous outside(very rarely sparsely hairy), hairy insidebut the hairssometimes confined to the middle and base,sometimes ciliate, when ciliatethe marginusually toothed, each tooth terminating in a cilium. F/owerswhite. Floral tube: ovary-portiondensely hairy, the longesthairs 3 4 mm long; style'portionsparsely to moderatelydensely hairy, the hairs all long, the longest ones2 2.5 mm long.

Specimensexamined. WESTERN AUSTRAI-IA (selectedfrom over 35 seen):Serpenrine. T.E.H.Aplin 1230(PERTH); Ellen Brook Sancruary , N.T. Burbidge7916 (CANBT; Jan

Distribution.(Figure 18.) Extends from north of Gingin (31021'S, 115054'E) south to Serpentine(33022' S, I15058'E). l Habltat Restrictedto the coastalplain, occurringin sandover clay or sandyclay in low- lying areasassociated with seasonallywaterlogged depressions or watercourses. L I Flovieringperiod. Mainly OctoberJanuary. also recordedFebruary. I A AJJinities.Closest to Pimeleaimbricata 'tar. piligelr. Apart from the charactersgiven in the key, var.major differsfrom var.piligera in the longerhairs on the ovary.portionof the floral ttbe. Pimeleaimbricata var. major tendsro havenarrower leaves than all the other varieties. UI or Note.!.All of the specimensexamined were bisexual. Specimens collected from Mahogany rh Creek have stronglyciliate and distinctly denticulateinvolucral bracts whereasin other de sp€cimensthe involucralbracts are not ciliateor not prominentlyso andare entire. Another pc differenceis that the hairson the ovary-pofiionof MahoganyCreek specimens are very short stl comparedwith those of other specimens.The Mahogany Creek variant needsfurther us investigation. Sp Rc l3b. var. imbricata 65 2, yar. Pimeleaimbricata barteri Ctnn. ex Meissnerin DC., Prodr. l4: 507 (1857).- pimelea on imbricataf. baxlerl(Cunn. ex Meissner)Benth., FL Austral.6: 21 (1873).Iype: southwestern Ke Australia,date unknown, W. Baxter (lecto here designated: NYi isolecto:MEL); Mt Clarence. 12, WesternAustralia, 25 Nov. 1840,l. Prerss1273 (syn: LD, NY). J.r, '^oN '.lle,ra{Eg ,oleptsorroC llAI:(HJUAd) 91 aBnag S.V g uuDW tt^r*, r*r^ '6061 ,Cy) I ,oolorooll\ ,.IAr\l)ii} rco'repurd:iHJdAd'.IEI I ,9LI ttrox l,f :(HJUed grzI Ilrox W'Sut,i.oxtoJ:(HJU:ld) Dqulx iittiliy 'ootrel€l[ 'gNyJ ,(fy)L8t Jd,dn8urTle/,puC:(HldEd) Eggg 7 9 :(HJdEd 6gLZrauuo1 N.N oruBrodsgadJoqrsue,rex uo Surssorrra^ru Suno^ url r :(-IEW)uluotJ .W,9691 ,ur8e^[ ,Z 'J 'e3-ue5 JoIAN t e1e1:(HfUAa)l sutng y ,{qsa:o11:(SNVI) aBprqtng .u?r\\oJo{€.I :tHfUEJfOSSg ttlz 1.1t1.^l ptoag'97'mau?y Jo iA [ru{0l] rurg :(ey) Iqqsv y,{oorqu?rJJo:lu-Il .o,pU Lg}S .paulwoxa,l ra^ru tles:(usos0gz ra^ouorJ pepelos)vl-IvuJsnv Nu:ll,sg,t suaw2ads

sr?rnsa'uoraqr;sr?q rarrorrs qonur pu' Buol ar",r,.u'Jlf;:rti*.iti3l#fir5,,iiJ[il1l Jo ,Buol :sro^\ouFnxosrq ur u?qt elpuoJur rauoqsqcnu ,{llBnsn uru S Z ot dn $rEqqtr,,h uoulod -^ru^o:aqn (po8uu flalel) ueerc ot altq,r s"raMofur.tnoq8no-rqr 'olrtualqtol! 'oldJnd lutd iririq ,tloiuep ,{ll?nsn ot poJdeap ,{[e0Jed Jo so^?eluBr{t fje^lrs Jloru lo Jnoloceu€s aql ,Judrlle gJotq l.onnlotul Buolu]tu ot dn ,{l,r^ollEu,{llpnsn ,uaaJ8 unrpeu pup snoJqel8lo ^rreq,{lasreds^laler'uoe$ dooplo uoel8qsrnlq ot unpou pue,{:req 'aeuretlB lr?p klasuopfilensn sa^oa7snolqelS,{laJeJ fro^,fur?q ,{loteJoporu,{lerej ,,{Jrprl .,{lasuep ^llensn suars

'(\ :uis) ptaljtpto .!.y 'u,roulun ',{JoSorC e1?p'ellellsnvualsel[ llod :((.I:u,tsosr) ss/a/dr'7,6€gl.tco 'erlertsnv 'ltunouuaetC] L/ZI 9l uralso/A :((.I :u,(sos0gLZI sslaq ,6€g 'erlErtsnv 'quad'o8ueu ,,urelunouuaorD,, 7 l ^oN I urelsei[ SurUeC:{) :palEu8rsopeleq ol3o .u,l .II]Jpuoanrue .lBrsnv .lC ,qluegtgs I'ert€rlsnv ulalsa,rqtnos :adtJ ltLgl) IZ :9 ora8\rd.JDjnuqw! oalauld- (t061) .lo8 ,lazlrrd.g .rB^.r€ /6t:9€ ts,{s qrqet ry qslc (qtuog)u ra8.rlld l

ala,ifipeurlnexo ojo)uqa|Je^ suelllrcadseql 'otadltld Ienxoslq Jo Jo IIV JE^DjDJUqu! sl tl asn?coqo^oqe polro lou sl srqJ (IN) l/aql, '?rl€ltsnv d '(,{BfpooJ)Jo T,uMou{un el€p ujetsalA ,,defpoof,, se,\\pauluiexo ed,{tu,{s laqtlnj y sa,?oN 'nta311td rcr DiDltqlu! Dapwtd ot $esol) sauruli[V

'qcJehtrur papjocoJosle ',4renuEf-laqotr0pouad Suua*ol! 'puus suorssJrdeppeSSollatB,r ,{ll?uos€es qu,r lo JtruBlSqtLr potercossp ur slncco /rrigrH '(A 'S,0l,tt) 'S,Z0oSt) ,9l0S dnllrNtp oruorrnroopet€losr up qtl^\'(A 'S ,ts,fll'uounqlt$le ,{uBqlv ol tseo(E .S009I I ,9t0t€) dnl?pnqJ ll{ rxo4 spualxE( gl arn8rl) '(CflJ) 's}^nN Ia13yr11 y'9961 lro turod ol ppor'.(.Dg))MDqsuanllo tottoJ.I 'dnuerEno14 fl d n 00gl :(HJdg pltis reou:(HJU artoY .e ,dnfirN:(lal!) 'LL8I d) EI,IZ'uouu?qs:HJdSd) V'urIoquofl d) ll V la anw'f roc ZI LZ; a^o7 IJUEN:(HJUgd) '/4 'e{E-I ON'IEed .t .rg6t 8Z0l-rxo uaa.tg ['al3d9,{\ f33og rzeu'.(-IAINIqjtztqpD ',{uEqlv Jo ,,\[S[uI0t] Iu.r'dnlurfu?t{9 :(HJdgd) Z6LLztouae 7i ur) :HJdAdl u!tdv 'dnlapnql Jo S 99 'paulu\xatt. H.g.J U^[ :(uoasSg ro^o ruo{ polcolos)VI.]vUJSnv NUIIJSIIIA suala)ads '8uol uu Z-t l sJrerltsa8uol eqt'slt€q JolJoqsqonu pu€Suol Jo orntxrruB,{q peJo^oc^losuep uoulod ollts :uortJod-olflsJo osorllu€ql Je8uol,{lerl€cs lo ro8uoltou slrerleqt tnq uoruod-el,{lsueqt ,tureq ,{losuop eloul ,{pq8rJssaurtouos uoulod,,{Je^o 'oprslno ,orrtuo,aldrnd fr?q ,{lesuepaqnJ lotoll lutd deepc1, epd sraMoll tnoq8norqt,{Ieq ot por ,{llenJBdual.Jo slJorq luJnlotul ,fureq,{losuop ot fleslpdsIIe ,{laJ?lJo olerlrcsauo lsouraddnorlt Josnorq€I8'8uol uru g ol dn'cqdllla^l,roreu ,{[lansn'uool8 runlpou 'aleulalle ro atlsoddosadra1 aruecsolouurqJee lEau qsrppalpue ,{lreq^lasuap ,{letelopour s /rts'

t8r aParDiBlaur^qJuuls]lsnv urltsay'd o,{d l g 184 NuyrsiaVol. 6, No 2 {1988) 1871. F. MuetterIMEL): Mt Gibbs,K Newbey6526 IPERTH), c. 2l km NNW of coast at Stokeslnlet,,4.E Orchard1614(AD,CANB, PERTH); Murchison Rivergorge,2T Sept. 1962.M.E. Phitlips ICBG):Cockleshell Guily, 24 Sept. 1968,M.E. Phillips (CGB, MEL); Pinjarrah,Oct. 1872,lS Price(MEL); Watheroo National Park, R D Rovce9544 {PERTH); Mt Burdett,B.L. Rye 82020(PERTH); Mt Caroline, 1886,G A. Sewell(MEL); c. 35 km N of Merredinf).1 E. llhibley 4116(AD, PERTH); Wickepin-Harrismith, E. Wirtwer 881 (PERTH).

Distribution.(Figure 19.) Extends from the MurchisonRiver gorge(27034' S, I 14027'E) south to MargaretRiver (33057'S, 115004'E), inland to Wogarl(31054'S, 118031'E) and east lo nearMt Ney (33034'S.12228'E\.

Habitat.Occtrs usually in sandor sandyclay, sometimes in ciayor with gravel,often associated with graniteoutcrops or sheetsor with seasonallywaterlogged depressions, sometimes in sandplain.

Floweringperiorl. Mainly SeptemberDecember, also recordedin August and February.

Affnities. Closeto and tendingto intergradewith eachof the other varietiesin Western Australiaand possiblyeven more closelyrelated to the South Australianvariety

Notes.This is by far the most widespreadand variableof the varieries Many herbarium specimensare female. It appearsthat thereare approximately equal numbers of bisexualand femaleplants in the wheatbeltareas but probablymore bisexual than maleplants in the more humid parts of the range.

A further syntypeexamined was PhillipsRiver, date unknown, G. Maxwell (K). This is not cited abovebecause it appearsto be closerto P. inbricata \ar ' imbricata-

Oneol the namesgiven in the nomina dubra,Pimelea imbricaru var. irana{Graham) C H. Ostenfieldwas publishedprior to P imbricata var.piligera and shouldpossibly be applied to thisuxon. However,no typ€specimen was found ior the formerand the originaldescription of P. nanu Grahamwas not sufiicientlydetailed to allow a definiteidentification.

13d.var. petraea(Meissner) Rye, comb. et stat.nov.

Pimeleapetraea Meissner, Lrnnaea 26:347 (1854).- Banksiapetrcea (Meissner) Kuntze, Revis.Gen. PI. 583{18 91). Pimeleaoctophylla subsp. perraea (Meissnerl Threlfall' Brunonia 5: 178(1983). fl.,pe: near Cudnaka, South Ausralia, 1851.F. Mueller llecto,fide S.Threlfall, loc.cit.: MEL, n.v.).

A descriptionof thistaxon, which does not occurin WesternAustralia, is givenby Threlfall, loc.clt.

emrnined.SOUTH AUSIRALIA (selectedfrom over40 seen):Mt Yardea,Gawler Specimens tt Range,PG. Wilson 558 {AD, PERTH); Yorke Peninsula,1819, Tapper (MELI Pi 20.)Occuls in southernSouth Australia in the Eyre Peninsula,Flinders Distribution.{Figure of Yorke Peninsula. Ransesand S.

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186 NuyrsiaVol.6, No.2ll988) Sftrub,usually erect,0.2-0.7 m high,single or multi-stemmedat groundlevel. Srems yellow brownto red-brownand densely hairy near each inflorescence (but the coloursomewhat hidden by hairs),becomrng medium brown to almostblack and glabrousfurther from apex.Leaves alternate,sessile or with a petioleup to 0.3 mm long; lamina concolorous,usually pale to mediumgrey-green or yellowishgreen, elliptic or sometimesnarrowly elliptic, 2-1 x 1-2.5 mm,flat or adaxiallyconcave, obtuse, densely hairy; hairs on abaxialsurface longer and coarser than those on adaxial surface,the longestones l-2 mm long. Peduncle0.5 4 mm long. Involuual bracts8- l8, the samecolour as leaves, narowly ovateto almostovate, 4-9 x 1-2.5 mm, denselyhairy; hairs similarto thoseon leaves.Inflorescence ercct, compact. Pedicels up to 0.3 mm long;longest hairs 1-1.5mm long.F1o"e/s female or bisexual,white, densely hairy outside,with hairs0.l'0.3 mm long insidedistal part of floral tube;tube 5-8 mm long in bisexualflowers and 4-5 mm long in femaleflowers, probably not circumscissile.Oyary portion of florul tube 1.5-2x c. 0.5 mm; hairs antrorse,3-5.5 mm long in bisexualflowers and 2.5-4mm long in femaleflowers. Style-portion offloral tube 3-6 mm long in bisexual flowersand 2 3 mm Iong in femaleflowers, 0.5-0.7 mm diam. at summit, with a mixture of long and much shorterhairs, the longesthairs 1-2.5mm long.Sepals ovate, 1.72.5 mm long in bisexualflowem and 1.32 mm long in femaleflowers, glabrous inside; outside hairs similarto thoseon style-portionof tube.Stamens shorter than or rarely slightly exceeding sepals;filamentI 1.8 mm long;anther0.6-l x 0.3-0.5mm; connective slightly narower than antherjslits usually more or lessadaxtal. Staminodes: filament 0.2-0.6 mm long;anther0.2-0.4 x 0.I -0.2mm. Or.,aryhairy at summit;longest hairs 0.8- 1.2 mm long.Sty/e exserted by 0.5-2 mm in bisexualflowers and 2-4 mm in femaleflowers. Seecl c. 2.5 x I mm. with faint longitudinalmarkings.

Specimensexamined. WESTERN AUSTRALIA: N of Esperance,Oct. 1903,C. Andrews (PERTH); 6 mi [9.5 km] W of Agnew, J.S. Beard6580 (PERTH); Lake Cowan, NW of Norseman,N.T. Burbidge2733 (CANB); 10 mi [16 km] E ro SE of Sandstone,1960, Cote (PERTH);near Lake Lefroy, 1893,E. Cronin (MEL|, near SourhernCross, A. Faira 2424 (CANB);between Red KangarooHill and Yilgarn, Nov. 1891,R. Helms (MELI; nearLake Lefroy,-Nov. 1891,R Helms (AD); 9 km NE of Norseman,K. Newbey 7530 (pERTH); 52^mi[84 km] W of 9oolgardle,l7 Sept.1962, M.E. Phi ips (CBG);65 mi tt05 kml NW of Sandstonetowards Wiluna, R.D. Royce10931(PERTH); N of Sandstone,W of Old Grdgee Homestead,R.D Royce 10453(PERTH); Coolgardie, 1900, L.C. WebsterIBRI):27 mtl43 kml S of Coolgardieon road to Norseman,3 Oct. 1961,J.H. Wittis \MEL). I SOUTHAUSTRALiA (selectedfrom c. 10seen):c.48 km W of Kimba.R. Iil/640 (AD). ( (Figure I Distribution. 19,2l.) In WesternAustralia recorded from north of Sandstone(27,10' ( S, 119040'E) to nearNorseman (32012' S, 121046' E\. Alsooccurs in SourhAustralia. t l Habitat. Recordedin sandplainor on rocky hillsides.

Floweringperiod. September-November. 5 t (l A"lJinities.Closest to Pimeleaimbricata. Apart from the differencesindicated in the key, Pimeleasubvillifera tends to bemore densely hairy throughoutthan p imbficata.Itsinvolucril 0 bractsare alwaysgreen, never having the red-purpletints p. I often occurringin imbricata, (I and femaleplants are more commonin P. subvilldera.In WesternAustralia p subvillifera tendsto have shorterstamens and the anther slits more adaxialthan in p. imbricati but Si this differenceis not so evidentin South Australia. T, Noles. Femaleand bisexualplants appeared to be approximatelyequal in frequencyin Western ul Australiancollections whereas only one of the SouthAustralian specimens examined was w biserual.the orhers all beingfemale. .s, kn rrt.r.'8 3:(Hrx'dszLtz pltjs n,,,,,"',11t'ond'r5llHi,lr;t:,?]t:'r"tjitrfiJ'r',:J ur :{HJusd) 'joV liroiJo-fi ,pf .f, :?l:'l,:J-J: qN 8r 'iururrueJloolqnas r rrior r ;rq;;ra :lHIUgd) 6tt6 ar^o{ O y Jo :(V,41O)uoaTiyg' ,ur"ril,uyii 'eqlng S [u{,slllH ?Z]lu SI ,ro,roJ ,14:y 9lep :HJIA6) LgggtttDa.uuax 3.y us8uol[ ro]BlsuerJuorsr^oloJ :fiJUAd) g7ggt oauuaxjl) ,slllH uBBuoA\,ra^\oJ 'g :ojelsue:j uom,talaj,toiaqdeE :H+dad 6sI/ Qloauuay y's11t11ueauo6ie,ro1 :*or"1sue.tguotrtnrlel aq1 pud airrpr5 l uqdra_uaa,,,uoq:(H 's111g 'rfirg L rr?J,i'sltH tggs tlDaiaax g, y uesuo1,1 11e4s1uo1rtr :(HJuad) ttrz qtuaauaxC Z ueauo4.:(glgg!11gg, rntfiiix 7 5 ;;r;e;j j"E;",ftj :(CV)l.S I 8r4V/1^? 's1tH u"Euol[ :1H1ggjf1anp]r9 y3 g7[ ,au,rliS I ;E6ii:,ro51 1:,to5 rf:ndifi 1t:1:{J _r!_sl.-rjueu 'setue^ralsutue( :(HJuEa).ptDDBziU t 7 ri,pr"p".liHigd;i 6001rtrlr W JoaN rul st :(HJugd)g1gI ptoig ;eqriee;gy6 0{€'J rpou pu€lur 57 .paautoia A|'ils-Ji lseocurorJ [ul{ tj liu Z :vIlyUJSnV N6EJSC,4 ii)utcaa!

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Ftgutc )) PimeleoI illilera. a'ta art marchingthc typc spccimen.A- floweringstcm: B porlionof stem(x l); C- outcr\lrlace of brscl (r 5);D. flo\acr(x l0)i E sramen(x 12)iF ovarylx l2). Drawnlrcm C.A. GatdnerlMt Srirling.Nov. I928).

Distribution.(Figure 18.) Extends from Gutha (29000'S,115057'E), south-west to near Cervantes(c. 30031' S, t 15004'E) and south to TamminFlora Reserve (c. 31039' S, I I 7'30'E).

Habitat. Occursin varied habitats.Recorded in sand,sometimes on limestonepavements, nearthe coast.Most commonlycollected from lateriticsoils, often with outcroppinglate te. in the WonganHills, the DarlingRange and at York. Alsooccurs on graniteoutcrops or in sandin moreinland localities.

Flowering period. October-Apnl,especially November February.

Closestto Pimeleaerecld, the differencesbetween the two speciesbeing noted under Alfnities. FJ P. erecta. E

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'sacsJJnsqloq uo .Je^ ,{r?q ,{lesuoporE l8ql qedoso^pq }seoJ oql J?auluo4 ,./a!1qt 4 1o suaunteds.uaS!t!d DlDJuqrutDapww'qlr,r pesnJuoa0q,{Ern pus'olquosal ^llElcrJJodns,{sqJ SEJJE JJr{lo uroJJ suounedsul ueql sreq Suoleroul s?qeqnl or{l uonlod-al,{lsoql pue so^€alp?olq 'sro,rou 'sa^?el'sruels IsJoU Jo ^lo^rleloJ pue slcerq ,{lreq^lesuop fJo,{ o^?q aseqJ tloql€oq,{Aaql ur sdoJJlnoalruar8 ruo{ suoulroodsseqot?u uarurJods ed,{leqJ sanadselqeue^,41e^ V saloN 't,{U .i 68r opr.e.el:Lu{ql u?rlsrtsnvuratsa/A B 190 NuytsiaVol.6. No.2 tl988l Sftrab,erect. often spreading, 0.3 l(-2)m high,single stemmed at groundlevel.5iems pale brownand usually partly reddish near each inflorescence, becoming dark grey or darkgrey brownfurther from apex,glabrous except for axillaryhairs or withoutaxillary hairs. leaves opposite.antrorse to patent,glabrous; petiole 0.,1-1 mm long;lamina concolorous. medium green,usually elliptic to ovateor narrowlyso. rarely very narrowly elliptic, 5 20x 1.55 mm, flat or adaxiallyconcave, acute or narrowly obrnse.Peduncle l-8t l5) mm long.Involucrol braclsusually 8 or 10,not becomingreflexed. dark reddish at basebut otherwisesimilar in colourto leaves.ovate to elliptic,5- I I x 1.34 mm,glabrous outside, densely appressed-hairy inside,densely ciliatet hairs (including the cilia) up to 0.4mmlong. Inflorescence erect. mmpact. Pedicels0.10.2 mm long;longest hairs I 1.5mm long.F/owers bisexual. usually whlte or off white.sometimes pale pink; tube6-8 mm long,circumscissiie and ratherconstricted c. 0.5 mm aboveovary portion. Ovary'portionofJloral tube 2 3 x c. I mm, glabrousat base outsideand densely hairy above. glabrous inside; hairs appressed to antrorse.L5 2 mm long. Styleportion of floral tube3.5-5 mm long,1.2 2 mm diam.at summit,densely hairy outside. ratherdensely hairy inside; hairs on outsideantrorse. all ratherunilorm and short or some minuteand othersmuch longer,the shortestusually 0.1 0.2 mm long.the longestup to I mm long;hairs on insideoccurring throughout the expanded part of thestyle portion. patent. the longesthairs c. 0.4 mm long..tepals ovate or nearlyso. 2.5-3.5 mm long.densely hairy outside.the indumentumsimilar to that on distalpart of floral tubeor with a greaterproportion oi longhairs. densely hairy in basalthird inside,rather densely hairy (aiongmiddle) to glabrous above.Stamens longer than sepals: filament 3 5 mm long;anther 0.8-l x 0.2-0.4mm; connective ratherbroad bur narrower than anther, slits semi lareral aiter dehiscence. Oro,')'sparsely to ratherdensely hairy. with minutehairs 0.1-0.2 mm longthroughout and larger hairs 0.4 1 mm long at summit.St-v/e exserted by 3 4.5 mm. MatureJeed not seen.

Specimensexamined WESTERN AUSTRALIA (selectedfrom over 40 seen):near Salmon Gums, tr{E. Blackall 1004(PERTH); c. 3 mi [5 km] SE of Mt Ragged,A.S. George2125 (PERTH):l5 km SW of Newdegate,"/.M Kocft89 (PERTH);6 mi [10km] S of Fonestiana crossroads. F. LullJitz39Tl(PERTH): Frank Hann National Park.D Monk 186(PERTH): 2.5mi[4km] NWoiOngerup,K NeweyT0 (PERTH); I mill.5 kmlNof Hopetoun.ME Phillips623004 (PERTH); Perkins Rock. Fitzgerald River National Park.. R.D Royce9212 lPERTH);Israelite Bay, 27 Nov. 1950,J.H. ,yillis(MEL); Hopetoun. 21 Oct. 1968.J.W. Wrigley(BRI, CBG): Esperance,near Pink Lake, 30 Oct. 1968.J. Wrigley(CBG. PERTH).

Distributk)n.(Figure 25.) Extends from Ongerup(33056' S, I 18'28'E) eastto IsraeliteBay (33037'S. 123052'E). 1 ( Habitat. Occursin sandor clay, sometimesrecorded in open malleervoodland. t

Floweringperiod. July March. especiallyOctober January. I a Derivationof name.Erectus lL.) - upright.referring to the erectinflorescence and involucral S bracts.the latter. unlike the bracts of severalrelated species. not becomingreflexed in fruit. I Affnities. A muchless variable species than its closestrelative, P/melea villifera. The involucral bractsof P e/ectadiffer from thoseof P villilera in berngless hairy, havinga lessprominent midriband being reddish at the baseon theoutside. Other differences are given in the diagnosis.

I( (R. 17.Pimefea sylvestris R. Br.,Prodr. 361 (1 810). Calyptostegiasylvestrb Br.iC Meyer, bt Bull.Cl. Phys.Math. Acad.lmp. Sci.Saint Petersbourg 4: 74 (l8zl5i. Bonksiasj'/vestris t (R.Br.) Kuntze, Revis. Gen. Pl. 2: 583(1891). Ipe. KingGeorge Sound, Western Australia, Dec. 1801.R. Brown (BM. MEL) IeJupurl,{caJou seMoqnl IEIoUeqt t?ql s1$33nsuortEJtsnllr oqt lnq s?./rsalv{sA Jo pcrd,{1e:a,,'r sueu€lspue molm ra,rou sll punos e8loeC 8ur) le polroflocpoos rxo4 u^\or8 ,{lpeuodars?,r uoxel oqJ soloodsoql ^Jnuaprol uodn perlaraq tsnu uortertsnll pue uorlducsopod.{l oql 'uolllltzotS d roJpatecoluoeq seq uou.rnadsod.,tlou sV (S861etd,) olnDp) d peu€u ueoq ,rou seqqJrq,r'soneds pelplel e Suraqs?'{0161) preeg se qcns'sroqlne ouros.{q peprB8aJ sE,rlI s. lsad1,{sA qll,r JuroedsuocSureq s? ejeq popJ?8elsr nto[11tcot7Dapald sa]oN

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Figorc 24. Pimeleasylvesrlb. A floweringsrem wirh bracrsreflexed; B- flower (x 8)i C- stamenand openedfloral tube (x 91:D ovary (x 12).Drawn from lR."/.CrunJield 19'70.

than usaal Pimeleasylvestris typically has pure white flowers, although some specimens are palepink or pink'tinged,and its floraltube shows a moreobvious expansion at thesummit thanin P calcicola.In P. sylvestris,the involucral bracts become reflexed in fruit andare oftendark purple or blackon the abaxialsurface. The anthersare more elongate than in P. calcicola,with a narrowerconnective and the antherslits semi-lateral after dehiscence 'V sr.fsa,1/,(s"i1d puv L oDaraDapud Jo uotlnqulslc SZ,tn8lj I I # ", J, xY'trlv.i \f 4 A fui"il A f 1".,1 -r- -t V fii" J r--! I \\ r '8uol 'qlSuel 'oprslnosnorqeF'lruluns te'rx?rp rxrx ulru sll lnoqEnolqtfllq8lls ,{luo 'rupuq,tc 9 l-l ZI-6 'tutu eupuedxa tltrtoueu aqnl ptolluo uotltod-a1t$ snoJqBIS 9 l- I x t-t aqruptoL/ uoglod-K.to^Ouor od-^re^oo^oq? uur g ellssrcsunolto'8uol luru oqnl:JI?q lo l-9 0 'l€nxaslq9l-ZI u?ql poJnolo3,tldoap erou uauo oql '{uld doopol elBd s/aMofur Ielsrp Jpq l?uIrxoJd '8uol 'Auolruru sJr?q:Euol ului slaJryad]xzduror'txere aJuacsatoll[u1sno:q?[8 Z-SI l-S'0 'poxolJoj ulru x -6 'os ,{ljeouJo o1B^o 'oJnlxol pu? Jnolocur so^?ol01 lqluls Eulurooaq 'sJo,rou8't 6l '8 lou SurpunoJms,{lasolc sollneuos Jo 9 siJ'lq lDnryo rl Euoltulu (9d02'9(Z) 'asnlqo 'a^Bcuoc 'ujld] apunpad tsorulBol alnrB ,{llsurlsrp,{Jl?lyep? lo l€lJ L-t(l) x LZ El(r) 'crldrll3 'snoJolo3uo3 013udrle ,{I,{\o!eu'uo3rE untpour ot ol?d suttxq :3uolurul 9 l-1 0 sloqod 'luoled 'olrsoddo :snorqelS ol osrojlu? sa,lra1 sJrBqfl?llx? roJ ]dacxesnorqelS'xod€ uoJJ 'eouacserouul Jor]unJ^al3 uoql u,,i{oJq-pgJEutruocaq qtee l?ou FBUIl-{uld sournouospuE 'lo^al 'qEIq 'qruqs ueoJEolEd suats o^oqe^qsnq punoJ8w pouruels-olSurs ur I Z'0'].ere 'urqeT t(St8l) 909l ssrordId ul reusslet\l3o:IooH uou !laneonlltllJolS DaPuUl '(ldsN"Iat{ ') 'gNvJ 't86t '1rO 'E 'S '?IlBrlsnY :osr:HJUgd :oloq) L}|I uaqo) N 0l ,StoSII ,0S0lE 'quad 'oulre3'fBiA '3,{d urotsolA proJurorJ:adq $861) 9-V:g els nN BloJblsrEolaulld 8l

'lseocqlnos aqt rBeuuo{ peproaojuoeq dluo o^?q suotubadspaJa,,(olJ-Iurd sle,,l.{olJ 'UoI 'llo,releg pue so^?eloBJ?I ^Ja^ a^?q r?ou U{ uo[ sltlsa^lKsoapwld Josu3t"utJeds 'tnoq8noJql 'atu$slqap oldrnd01 lurd doepSuruoooq ueql 18eplsul ot{l uo o6ue:o^luo oru sJe\|uee\\ qI}JDIDJa ur seeloqrho8u?ro oJ€ sjoqlue aql s!4saqKs d uI lerxepe u€ql J0q13l urorseiA'3fU 'I €61 3lrcua?lau^qJ uerlerlsnv B l,94 NuyNiaVol. 6. No. 2 11988) hairy insidein the distalhalf. the circumscissionproinr often distinctively coloured: hairs mostly patent, oltencurved, 0.3-0.6( 0.8) mm long."tepals oyate, 2.4 mm Jong,g\abro:Ls. Stameis longeror rarely.shorterthan sepals;filament 2-3 mm long;anther O_-j.d.g x 0.3.0 5 mm, connective.verytrroad but slightlynarrower than anther;sliis stdctly adaxial.Ovary hatry at summir,hairs 0.5,0,7 mm-long.,t/y/e exserted by 4 7 mm.Seedc.4 x i mm.with l";;,tr;;;; lurrows.(Figure - Rye 1984:5.)

Specimenseramined. WESTERN AUSTRALIA (selectedfrom over 25 seen):Mandurah, , J Burbidge(CANB): r^9 6]ry f4f,-kmlS oi Fremanrle,3 Oct. tglg. E U. Cann;ii (CBG);\\c:!9!) Mandurah,C. F. DaviesI 38 (PERTH);burns BeachXa,, U. Oeman' iSS {pERfFii MadoraBeach setrlement. B. Haberley201 (pERTH); Nofih Fremantle.: Nov. f SSZ,i. Helms(PERTH| Yanchep National_park,AM. James49 (pERTH);24 mi peg{3S ili ;; Yanchep--road,F. LullJitz 3700IPERTH); The plains.near Mandurah. V. Mo'nn& A.S. George32 {PERTH); near Coogee, dareunknown, F. Mueller (MEL);'tgeZ, ialgorup National P:rk. part, S Paasr 134^l-(PERTHI;Yanchep Nationat 27 Oct. U.A. pniiip, tCeq; Carine.B.L. (PERTHI; Rye82014 Nof Wanneroo,F.G. Smith1584. lpERTHi Vicro;i; Park. date unknown. J.L. SteedmanINS\N ). (Figure yanchep Distribution. 26.)Extends from Nationalpark (31032'S.I 15040, E) south of YalgorupNational Park {32050'S. I 15,40'E).

Habitot. Occvs closeto the coastassociated with outcroppinglimestone. growing in sand. Floweringperiod. SeptemberNovember.

Alfnities. Closesrro .

Abtes.See note un d,er Pimelea slhtestrls. The sepalsare distinctly concave and do not spread as widelyas in otherWestern Australian species in this section.

19.Pimefea fongiflora R. Br.,Prodr. 361 [810). CalyptrostegialongiJlora lR. Br.)Endl., - Gen.Pl. Suppl.4:6l (1848). Banksialongiflora (R. Br.)Kunrze. nivis. Cen. pl.2: 5gj (1891).Z_lpe: King GeorgeSound, Western Australia, Dec. 1801,R. Brown (BM, MEL).

Shrub.erect,0.3 L3 m high,usually spindly, single stemmed at groundlevel. Srenzs redrlish brown and verydenseiy hairy neareach inflorescence, becoming medium brown to yerydark brown and glabrousfurther lrom apex;hajrs antrorse to patent.very fine, the longesiones -2 I mm long.sometimes very variable in sizeand includingminute hairs 0.1 0.3 rim long. Zeare.sopposite or alternate.sometimes crowded; p€tiole 0.2 1 mm long;laminaconcoloroui, medjumto darkgreen. linear to elliptic,usually narrowly elliptic,4.lg x l-3 mm, adaxially concave.acute to obtuse.sparsely to denselyhairy outside(but often appearingglabroui) whenyoung, glabrous or denselyhairy inside; hairs l-4 mm long.pecluiile 2.20 mm long. lnvolucralbr-acts 4-6. becoming reflexed in fruit, similarto leaveiin colour,narrowly ovaie or ovate.5 l2 x I .5'3mm. sparsely to denselyhairy on bothsurfaces or glabrousinside; hairs similar.to thoseon leaves.Inflorescence erect, usually compactbut sometimesbecoming cylindric.the rachisup to 10 mm 1ong.Pedicels usually 0.4.0.5 mm long;hairs up to 2.5 mm long.very line. F/or':ersbisexual or very rarelyfemale, brown below cir-umscission ooint. white or rarelycream above, glabrous inside floral tube and sepals:rube 7l2 mm long, circumscissile0.5-l mm aboveovary-portion. Oyary-portion offloral tube1.5-3 x c. I md; hairsappressed to antrorse,the longestones 1.1.5 mm long,slightly thicker than thoseon styleportion. Style-portionoffloral tube very narrowly cylindric.5.5-9 mm long. c. I mm diam.-atsummit, dense ly anduniformly hairy;hairs appressed to antrorse,up to 1mm long, {xr veryfine. sepals narrowly ovate to ovate,5-6 mm long,with similarhairs to-those on distii Ire u,orJu.r'rc.(zr x)^r?^o x)uau?rsro ^\.,^ ,,r,,,?jf;ji1f3y",fl,::JJi:"i?ii',j1'"?:H rJeIq .)1ey,D:(91 *^., : Y:'it lorcprrnsrauLl IealtorJeunsrJtno-g:ur:riBur.u,,aog y,aio aiqa.orol/t{t oioalawg 7"una 1

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(/Z olnErC) slrd ,roll€qs ,{ra^ s,.r\ol qtl,r 'ruut x c pars' .sjanaou 'sle,rolJJo leupntl8uol l. l e oleual ur ruru J,tq peuasxa ur pouosxelou aql BurqceoJ 9'8uo[ l lenxastq 1nq teolqt a1,q5louIJ,{Je^ uru g 0 ol dn sJrBq:^\oloqsnoJq?l8 lsoule lo snolqzlA,slrBqourJ ^Jo^ nl p0r0 uB qlu ,&rdo '3uol ulut .c JoqtuB:sJal'olJ ele:UJ;a| sapoutut1js Jo'IeIX€pB I?uruJet E 0 tro fllcuts sllls:,{ll?Jatel slac aqt Surp€Jxa,{ltqBIS 'pEoJq ,{le^ o^rloouum:uru x .oqnt 9.0-S0 9 l Z I :8uol turu SZ0 c tuetuElrJ:lo,rol] Jo teolr{t le olrssesqnssraurls leloll Jo u?d 961 aeacsaeletrl.{qfU?rlsllsnv urelse/A 'a,{U -.I g 196 NuytsiaVol. 6, No. 2 (1988) Distribution. {Figure 26.) Extendsfrom near Bunbury to Cape Riche, with a disjunct occurrenceabout 140km to the north-eastof the latterlocality in FitzgeraldRiver National Park.

F loweringPerlod AugustFebruary.

A-fJinities.Closest to Pimeleapreissiibut very distinctive.It is the only WestemAustralian speciesof secl.Calyptrostegia in which the inflorescencesometimes becomes elongate.

No/es. Two subspeciesare recognised.

Key to Subspecies l. Young leavesdensely hairy on both sudaces.Bracts densely hairy inside subspevrei r. V;;;;l;;;;, h"l.v". ,u"-t"ii"J";;, ;a;;rilcr"b";;;;;;;;,.rrrre' ) hairy.Bracts glabrous inside...... 19b.srtbsp. longiflora I

19a.subsp. eyrei (F. Muell.)Rye, comb. et stat.nov. (Figure27.) l c Pimeleaeyrei F. Muell.^Fragm.5: 109(1866). - Banksiaeyrei (F. Muell.)Kuntze, Revis. Gen. PL 2: 583 (1891).Type: Eyre Range,Phillip River and FitzgeraldRiver, Western I Australia,date unknown, G. Maxwell (holo: MEL).

Leaves:lamina narrowly elliptic or sometimeselliptic, 5-9 x 1.5-2,5mm, conspicuously n hairy and appearingsilvery when young;hairs denseon both sudacesof leaf. up to 4 mm a long.Involucral bracrsslightly flatter than the leavesbelow, with similar hairsto thoseon leaves.Flowers white or cream;hairs up to 2 mm long. 2 0 Specimensexamined. WESTERN AUSTRALIA: N of HamersleyRiver estuary, A.S. George (PERTH); (PERTH); c 7222 TkmWof EastMt Barren,K. Newbey9470 HamersleyDrive, I 3kmNof trackof HamersleyInlet, I Taylor1130 & P. OllerenshawICBG, MEL, PERTH). (Figure (c. Distriburion. 26.)Extends from the BremerRiver 34010'S, 119004' E) to Hamersley ni Inlet (33057'S, I 19'55'E), in the FitzgeraldRiver National Park. fc Habitat. Recordedin sand.often on quartziteridges, in low shrublands. m m Flowerin g periotl. A ugustNovember. n2 flt l9b. subsplongiflora gli aF NaturalistesMoscou 2512: 178 (1852) Pimgley Calyptrostegiavillosa Turcz., Bull. Soc. Imp. Pt (Turiz.) in DC.,Prodr. l4: 508(1857). fyper south'western Australia, 1848, vlllosa Meissner Iel (iso K, MEL, NY). J. Drummond colL 5, n. 428 vel syn: o! de hmelealongdlora var. latifolia Benth.,Fl. Austral. 6: 34 (1873).7ype: south westeln Australia' sht isca, l. nii^^ond coli.5, n. 428 (holo:K, iso vel svn: MEL, NY) 1-. Leaves:lamina linear to narrowly elliptic, 4-18x I 3 mm, glabrouson adaxialsurlace or cir ,olIraii*.r with a few hairs in leavesclose below an inflorescence,spanely to moderately po glabrous tra"v on aUu*lutturface when young but usuallyappearing glabrous, becoming with tul uni.'frui^ up to 2.5 mm long.,lty olicral bractstending to be shorterand broaderthan leaves, xI trliiu- tioo'.ritn., sparsely)-ourside, glabrous inside. F/owers usually white; hairs up to 1.5 Ot, mm long. lon

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Pt da Figurc 19. Di\rnbu on 01Pimelea prelssii. F.g:'e I0 Dirtriburior ol Pin""lpartlotirt,tdd Pi Specimensexamined. WESTERN AUSTRALIA (selecredfrom over 45 seen):Ruabon Ludlour,T.E H. Aplln 1182 (PERTHJ; EI nearNannup, .,1.M Ashby 3694 (A,D);N 6any Hwy, ph 59.2mi [9.4.5km] S of Penh.EM Canning3l13 (CSC); tS.+ mi [25km] fiom pemberton towards_Nannup,-EM Canning6518 (CBG); tskmWof Dunsborough,R.l. CmnpitiSZS (PERTH);c.23km NW of NorthBanister, NM Donnert4'7|{AD); Frankland Ri7er,C.j. tl-Ot] BowRiver, (NSW,pERTH); fal lgrQer. !P\RTH); Dec.1912, S.W. Jacksan Wooioloo, yel M. Koch 1'755 (MEL); Mt Saddleback,l3 Nov. t904,A. Monbon \pERTH);Needitup Hilt, bla $_A"y?ry 1339(PERTH); 16 km W of Kybuluppool, West River, K NewbeyI t302 phittips per (P^!!fH);Alban_y !I_wy,28 mi [45km] Sof Perrh,M.E. 1916(CBG);pinjaira, Oct, (2) t^\1.2:!! !!!ce \MEL);Ambersate, R.D. Royce 3952 (pERTA); Helena V utiey,i. s"ii-ii 394(PERTH); 7 mi kml E of Nannup,l2 Oct.1961, (MEL.pERTHi. lon u1.5 J.H. I4/ilis me 5883 L 9-t088S at?^o,{l,roJl?u flarBr'el?^o fJIBnsn'qsrppor ,{llnJ o} ,4l Ed ,{lorer^ra^ ro ueerSumlpotu 'poxouor Sururoceqlou'sJo,rolJ Surpunorns .{lasoJ3'9 ,{larzJ ^Je,' Jo spqlq auol 'o^eruoc I 'urtulDnryo^ul wuJ al)unpad esnlqo,{l,roueu Jo olnce .{llelx?pero t?U x 9( Z) 'rsourlZ€'Z 'uaer8 'snorolocuoc S-l 0€ ol3udrllo^],{\orrEu ,{llensn unrpou Bu[xEl:8uol u]ur l-2 eloued 'esrortu?'etrsoddo 'srrBq 0 :snorqB[8 sadraT ,{JBIIrx?JoJ tdmxe snorqeF'yedE uro.U loqunJ 'ecua3sorolJur lc?Jq tsou{eol u,rolq-,4oJ8urnrpeu ro,{ol3 unrparuSuruoceq qoEormu qsl,rollef ,{lleuredsaruuoruos ro u,^Aorq-peldaep ot unrperuswals lo^olpuno6 lB pouurots-rllnur.{larpr 'q8rq 'uodo 'tcan 'qruqg ^ra^ ro pernulatseJ8un ur (g l I flal?r ,llqnsod)I- 1 0 ro ,{lpurds '('IEt\tr :osr:HIUEd ur oloqd ' ') 'So0€ 'uortrpadxE ^ u :oloq)slulrH ldos urolsol[ J] raplE Lr'1681 0Z'BrT?rlsny 'tocA[Eo?ZI eql Jo /9 duel :adtJ 1916l)/ t I urroJul rsrt{ llng o$1s13uo1re^ smuatDapatd '() :oloq) ptDlAols 'u,rou)lunotep '€llejlsny 'u?uoo8u?N:ad(J tII 'no.ssmuaiU uralsaiA (SI6l)tIt-9tI tuloJultsrN IIng oapal.l '\LN) uoqButuunJ u?f 'elleltsnvuratsslN'punos o8rooC 8uD{ 'rnoqr8H L,'8t81 ZZ'J(I rats^O:ad{J (/SSI) 66n : rpord ur roussrat\suaJsa^pJ j€rronolusn8uo oa1aw16 '(IN :oloq porunserd) 'Zi8l 'erleltsnv 'ezlun) aqllg I' ulatse,r-qlnos:adtJ \1681) t8S :Z ld uag sr^ad DsouauotsluDg - :€ eerllpruals,{Ssmru?tog seFuuv ' dlEI\ \reusstelN) lzgIi n7,€ ' {reussloll) Dso^tauo8alsotJdtlrJ - (8t8 1)0LT-692 :Z ssrerdld uqaT ur reus$al^{osoMau Daprut I ('IEIAI:osr) ',u 't puoawruQ 'ti8l 'BrT?rlsnvurelso,r qlnos LtZ ' IIor I' ttSl :adnJ glS1692 :Z ssrold Id uqo'I ur reussrat{ upuoluurup j3tt ouollsn8uo oapwt4 '(IN "IEt\l'Cl :osr) puouurue alBp'?ll?rlsnyurolso,r LgZ'urq3.l I'u,r\oulun -qtnos:ad,{J (gtgl) 692 :Z ssrerd ld ul roussret{.rolrut :ert mlot4sn8uooapwr4 '(IN :oloq perunsold)lasr"/jir J ilq?wnserdl,JazDry,,'u,rou{un et€p'[?rl?rlsnv urotso^\ ur pe].elloc,4llenlcel qlnos 'uqe'I ,,sepl[ ,roN,, .rd{J (8t81) 692 :Z ssrojd Id ur raussrol,{.loi.rilu mr m1olrn8uo na1aw14 'iNgl janog ie(I rerl?rtsnv 'punos 'ld '34un)I7J'1081 'd,) urarsald aAroo{)Bur>I:adiJ(1681) €8I :Z uoD sr^ou ( rg olto./i$n8uo DleluDg- SrnoqsrolodJur?S'ros dluJ pecy qtel\tr-s^qd 'J(rg lsngl) nL:t lJ IlnB're,{ot{ U) otlotusnBuooBatsoud,ln3 (0181)09€ rpord'rg U Bllolrlsnauuualaurd IZ 's]o,rou puEsl3erq rall?rus a^?q ot puol osl€puE 63uBl seroeds oql Jo tled ulatse,/r\oqt uoJJ sueurtedsu€qt trq?qur lollerusole eoJ?lo^lU lseiA - dnlo8uo oql uo4 suerurcods.sa.loN 'olo./j$n8uo 'sawug$V d ol tuelxe rassol? 01 pue otollABuolDalawtd o! aso13 'requocoq-loqualdogpouad Satauolg 's$eJoJ 'sosjnooJelB,r snid(lnng ul ,{llensn lo suorsseldepto,r-Jotut,t{ qlr,r polelross€ sourtatxospu? JnuolEIqtr,,h seurneutos'puES ur smrco a8ued Surll€( oql qtnos 'spuglr|co^A IeAeJB Jo sadllnw ur s€eJBJrllJol€l ur ,{ ensn',{EIc u smmoa8ueu 8uIIl?( e\j q'ullqoH '\E 'S ,S ,LV06I\ ,r?0t€)ro^rx tsoli\ or lsBa(A 'S 'S .6Z0gtI .r90tt)drua8uo ulo4 pue(E ra^rd,r og pue(g ur^\nea.I 'S,gtol€),SS09l I ,9E0?t) ,g0l)9I I ,ZZoi€) adeJol qtnos(g,6l09l I ooloroo^\rrro4lseol l? spualxa(62 etnFll) uotnE.tlste

661 aea.easllurtrr{lusrlsrtsnv urats.y'A o,{U l.B NuytsiaVol. 6, No. 2 (1988) or broadlyovate, 4-15 x 2.5-8mm, glabrousoutside, hairy inside at leastnear centre and bas€or the outerbracts rarely glabrous inside, not ciliate;hairs appressed or rarelyantrorse. Inllore_scenceustally erector semi-pendulous,rarely pendulous,io mpact.pedicels usually 0.3-0.5mm long hairsusually c. 1.2mm long.F/owers bisexual or lemale,usually whiti or cream,sometimes pale pink or yellow,glabrous inside floral tubeand sepals; tube (4.5_)6_16 mm long, circumscissilec. I mm above ovary. hoximal portion of floral nbe (lnlow circumscissionpointl 2-4 x c. I mm,rather densely to verydensely hairy, dark.coloured but the colourhidden by hairs;hairs all antrorseor the distalones patent, the longesthairs 1,3.5 mrn long,not asfine asthose occurring above circumscission pont. Distatportion offloral ) tube (abovecircumscission point) very narowly cylindricto iylindric, (2.i-14-12mrir"long, ( c. I mln_diam.at summit;hairs all antrorseor a smallminority patent,very fine, the longeii 1 ones_0.5-lqg !o!C,the shortestones c. 0.1mm long.Sepais ettiptii to ;vate,'(1.5-)2-3-f4) ( mm long,with hairssimilar to thoseon distalpart of tuG or wiih a few hairi tonger,ihe I longeston€s qp to 2 mm long-Stamens shorter or longerthan sepals;filament 1-4mm long; I anther0.5-1.3 x 0.2-0.7mm; connective narrow to verybroad but ahvaysat leastslighfy l naffower.than anther;slits adaxialto alrnostlateral after dehiscence,uiualty semi-laierai, l rarelystrictly adaxial.Staminodes: frlament commonly 0.3-0.5 mm long;antlier commonly I 0.1-0.5x c. 0.1mm. Ovary gabrous or rarelywith apicalhairs up to 0.6mm. Sry/e exserted I by 1.5mm. .Seed nor seen. {Figure 31.) I i I c T

Figule31. Pi elea-angusrrbrb. A, flowering_stem;_B- inner sudace of bract (x 6); C. flower (x I l ); D, (x lO. Drawn from fresh material . rcpresentedby R.I. Crsnfretd 2g5g (N ii N.-'iine, 1011 tB, i. b). 'ellsoddo ,{ll?rruo^turod llB ot Supuat'tuoled ro esro.uuE satDa:[ sJrBr.tr,{.r?lltx? JoJ tdmxe snorqelS'xaderuorj requnJu,roJq ,{er8 runrpou Burumeq 'aJu$serollur qc?e Jmu u,rojq-pol 'q8rq 'uedo']Jate 'qruqs Jo qsr,rolla,{slurls lo^al punolSte peuuets alSurs uu l SZ0 '(IN "IEI^{ rcl 't :osr) f u llocpuouwrue I 'ttSl 'zrtBrlsnyuretso,r qtnos :ad{J (168I)t8S:Z uaC sr^od'aztun) tt8l '.JC Id {reussrol,\) opunquo6fots:1uog - (/S8l) S09: rpord ur raussretr[ Bpunqlrou Balaurd ZZ 'srleq 'oatnl.ldlns oql qSnorqtolqISI^ ^lJ?alo sI uollJod-,{.Ie^ootll pue d ur esoqlelrl aJou'srreqrosreds qcnu o^eqsuotuloeds otxos ,|{aI,{luorJ pernJsqo st uotuod 'npunquouf 'esuop ,fuE^oaqt tEql os 4 ur su ^]o^ er?lulod uolssl3sruntllcaqr Moloq sjlPq oql sueunedsouros uI o8u8l sercodsoqt tnoq8norqlporollecs orB'strls roqlu? s? qons 'sortsuatJerBqo l€IxBpe lensnun,{la^o€ler laqto eore,{ueqlv eqt ur qtnos eql ur patEl]uocuocers sueurf,odspera,{\oU'Iurd olrq,r 'uotpl?ro3 01 eqq?aug ruoJJ Ilureru SurJJnrco'o8uer sertads aql Jo uBd uJJquouaql ol polcutsarore slJelq xls qtl^AsuaulJeds €xpt Julaadse{urotul Jlqrsr^p,{lrpEar tou lnq e8u?lIpclt{d?l8oa8 s}r tnoq8no:qt satJods slqeue^ ,{louerxe uy sa]oN 'Dflq/jult d sarcadsuBrl€Isnv urals?eeqt o't pre npunquoT[Dapluld ol ]saso17sautug[[y ',{J€nue[-tsn6nV[IljlelN poltad Suuafolf 'saerl 'sqnrqs puElpoo,lasourlouros Jo soallpru Jo slsrsuoouorl?le8e,{ tu€uuuopaql selrspaSSoFet?,r ,{llpuoseos ur ,{llEnsnueql pu€ f€lc ,{puEsul sauuoluos 'slror 'pu€s Jrtrralelqll.r lrosro pues,{lle^pJ8 ur saurteuos ul ,{lureursJlxao'lo1lqDH 'elona le loproq[?rleltsnv qtnoseqt ol JsolJ^la^ pJploJJluaeq sBq lr esnmaqotsts ]Bql olul 'S.Zt"6Z) spuatxa,{lqrssod tnq EllBIsnVqtnos roJ poprocoJ toN (g.Z0,llZI sarzual^trot pu?lur 'atels pu?lsua erlt ul (!l 'S elcngo1 0q1 tsa,rqtnos uortxeeq1 01 qtrou 'S.tqolz).tg,gzI ,tt0lt) Jo rql ur (a ,9Z0ilI IrBd l€uou?Nureqlu) ulo4 spuolya(82 ornAlg)uounquiste 'l'IllN) '988 '.elcng DurnJ Lr'9 t '.(Jg)) ftDqsuaqo url ',{B^\qBrH.riu?qlv :(HJU:ld) 'uloqurogd T 7IIZnfoJ T'surertluAJo 'W'9'896tli\N 6 't)tet,^[ tIgIL plris U reeu :(HJUEd).tro1s 1cOt Jo ,{\N [ru] :(rldsN)irors etep'3uIUu?18):(HJUAd) a.{y S 6tI lur Zl 7'u,rou)un'uoor8roqrrH 0Z8 78r'lserroj'.p,rerol^l uetO:(HJUAd) t9L9 toY O U :(rgJl sdqtllfd S W'2961 rro Z tsn1d aI?'I S II:(HJUAd pal JoS[r'CV) t908] tu6t:(HJ.Ugd) 888 S'ocerD JoN llul /t] Iui!{rBd l.t€ I ploq O g y'ieg sueolroJo oInC :{HJUAd)II99 ttaq(raN) l?uoqBN ,(lu 'IUEJ .I€.I uu€Hlu€rC elrhl06 Jo l[S rul IZ:(HJUad) n?Z Iaq^^aN,',4e9 rauorflJolA lru{ (HJ.dAd) taqntaN :(HJd!ld) Kaq^aN ,{ltr 'dnre8uo9tl lrrr0t ZIZZ )'eunI Jo A [rur{t] lu Z Z9 'o^roseuX Jo ,f\N lrul S 9] Iu t :(HJUAd) |LIL auDqolrw N X auoUDpW Cf €reqr?,roH :(HJUAd) zitJitnT 'peeuqsnfl :(TS,\l) ttrox 'Ijlpe aw aql llrH 'urs?gLISI U 8682 'rrtJegW :(HJ.USd)BZ9 rlrox 1 erlo] :{HJUad) 8LV9qtoauuax U ) runol Jo (rl u)t :(HJUgd) tnq8py 'tg6t 8nv 'aJJIIcqoN tuf :(CV) S t f D /Z'1681 Jo'eITrooBIe) ,^d.N,\t I I [0t€ lse: uoq)nf g'uo]dru?H tn :((v) swpH ^oN ur1 , S N '(IV) A Jo S 001 Suue8uere^\r€ou :(HJUAd pg11Eang 7'B].Jlnm,roJfteu :(HJUgd) 828u{lt9 V g '€qqBeuA ',4[SN'lu8.) ',{,rH lotl ru)tg :(HJUAd sulitlg lse.^,\qoN eql uo rreqle) JoS 169I 'IcoU11, sg uorJ lrul I I :(HJUSd) gggga8toag S U euorcl^ uoono :(HJUgd) t86l lnc S '11qqo3rJ] Iru 'ssorJ 'gNyJ) aBtoagg y oN :(gNVl) o.tlol &r'tr uroqlnosreou :(HJXAd ,\\o 'po33€d SZVZ 'ey) ',{rcntso 60V02ratqrlg 11 lr{ JolooJ reeu :(HJUgd 8662Duuoe NN ro^ru 9I' plolJplorul 3 :(HJXAd) s1a?O etep'solzuet\tr:.1AIN) Z(, uoq8auun3 y etlr 'punos EI 'rnoqreH Z'u,roulun 'p?or eBJoeC3ur) r4s,{O:(g NyJl 66gLa8prqrng J N Suuepuel[f e8urd AIII 'es. (HIdAd) nrotg tulod I :(Cv) Kqqsy 'Eroot{ tBgE D W'h^Ia) Jo t[N [rDI 8] lu 98ZI ,I'y Jo /1[ :(uaes00t o ruorJpalrelas) VIIYUJSnV NUAJS:I,U pautuDxasuaatrads pul 'l 107 a?otee?leu,{qluerlerrsnv ur.tsilA ..,{U g r88 202 NuytsiaVol. 6. No. 2 (1988) whenon an obliqueto horizontalportion of stem,glabrous; petiole 0.5- 1.3 mm long;lamina almostconcolorous, medium green or bluish greenon adaxialsurface, similar but slightly more glaucouson abaxialsudace, ovate to elliptic or more often narrowly so, l7)12-40 x {2-)6-15mm, rather thick, adaxiallyconcave, acute or narrowly obt:ose.Peduncle 2-2'7 mm long.Involucral bracls 4 or veryrarely 6, sometimesclosely subtended by 2 bractlike leaves, laterallyoverlapping considerably and closelysurrounding basal l/3 l/2 of inflorescence,often stronglyrecurved in distal 1-8mm, not becomingreflexed, largely green but red or pink on base,ovate or broadlyovate, l0-28 x 8-20mm, glabrousoutside, appressed'hairy inside but not near margins,not ciliateor only the innermostpair (when6 bractspresent) with some cilia 0.5 1 mm long. Inflorescencependulous, compact. Flowers bisexual or rarely female, white or creamabove circumscission point, glabrousinside; tube l4-17 mm long in bisexual flowers,c. 1I mm longin femaleflowers, circumscissile 2 3 mm aboveovary. hoximal portion of Jloral tube (belowcircumscission point) 3'4 x c. 1 mm, denselyhairy, darkcoloured but the colourhidden by hairs;hairs mostly antrorse to patentand c. 2 mm longbut the uppermost hain shorterand retrorse,not asline as hairsabove circumscission point. Distalportion of floral tube (abovecircumscission point) 10-12mm long in bisexualflowers, c. 7 mm long in femaleflowers, densely covered by veryfine retrorsefairs 0.1-0.3mm longin the proximal (2-)34 mm and oftenalso with a few longerantrorse hain similarto the longerhairs occurring higheron tube,the distal6-9mm with short,usually patent, hairs c. 0.2 mm long and longer antrorsehairs, the longesthain c. I mm long.Sepals narrowly elliptic,3.5 5 mm long,with short hairs and a few longer hairs 1.5-2mm long. StamensIonger than (usuallygreatly exceeding)sepalsi filament 4'7 mm long;anther 1.2 1.7 x 0.30.4 mm; connectivenarrouer than anther: slits semi'lateralafter dehiscence.Staminodes: filament 1-3mm long; anther I c. I mm long. Orary glabrous.Sry/e exserted by 9-10 mm. Seednot seen.(Figure 32.) a 2 Ji st

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L suawois"eqnt pJoU ,Buol ,rndlTlo Jo Ued Ietsrpuo osoql ot relrurs sjreq qtr,{ ruru S,Z lo c4dl1a,tlr'ro:'teus/rdas pspuel leqm:Lrosuayo s.:req aqi,auoi u.rur g.g_1.6 ureqlriered:jpoqs pue Euoluru g l-9 0 sJleqluoled ^la.leJ :o :sro.r1ue1o :rnlxitu e rir6o rionsrcsrunrrrcenoq'" unluounpur :luroduotssrcsurnJJD ,rolaq 8uu:nccouoruod-^lB^o uo esoqlol lplruts slr?r{ '3uol ,cupurt^J qll^\'lltuuns lp. ur?rplxur t l-l uur tl-i ,{l,rorJer ,{ro^io t1n6t:iu aqni lorcA uouod.al,{75 1u1oduorssrJsrrrncJn a^oq? BuulnJco ,Buo1 'osrorlup/o esoqlse aurJse 1ou u-ltu Z-[ ro pessaldd?slreq:tuu r x aqrupnL[lo uoirod Ktia,6.uor1:od-,{.re,ro 'Euol i t-g l e^oqsuu t S 0 ellsslcsuncrc ulu oqnl:sl?dospu€ oqnl oprsul ',rollo^ 'eleuroJ l,l .iiol9.9 l?rou snolqEl8 ^lorel ro l?nxoslqsr auofu urur s:itq l8iol ruu splpad 'lredruoc 'snoppuod .Buoyruu 7-1 9.02.0 acuatsatogfu1 eqn bqi ,xeoripuno-ie 'eprsur 'xed? 1 9.g lsa8uol iri1i," ilaiu.lp ^rreq-pessoldd€ p:e,rol slleq aal e qtr,r Jo oprslnosnoJq?l8 slce:q io reuut 'aprsur ,aprslno tr T :01?llclou ^raq-pessoldd€,{lpnsn snolq?l8sD?lq :uru .qsri lsoulalno 0l_€x SI-, !?lnrro ol crldua ,tl^\olJ?u o[0,{ olou seullorxoslo sa^?olse lnoloc arrrBseq},poxeuet 'sro,lrog ,srred Suuro$q lou Surpmorns flosolo g-t ut spDtq .Auo1v:w ,1cunl"a 'osntqo 'alecuoc ,ulu lotciloaul 9i ro elnre ,{llorxep?ro leu x ,ie1nc-:rcoi cnOrtta.tl,,noj;e-u 'uoor8 ,snololoruoc 6-S.l 9l-7 qsmlqlo ueeJ8rnnrpour eurruq :Buolrurui.O cjlOn alo4ed ,apssesqns ,elrsoddo eq1 ol olrssos'luolpd a$oJIrB sr,1ra?.s:leq,{:e1[xe:o; snojqel8,xedeuro.r.1 {J tu,rolq,po: ldacxa ror{unJu,rorq-,{oJ8 lo ,{e$ unrpou BuluoJaq ueqt lslrJqst,roJlo,{ jo ecuecsa.rogui qcBaJEau u,rolq,poJ \JEp swag.{cols punoJ8topun,{poo,r ol?Auolo ue ruo:; gurle:aue8e: 'lo^ol ,qEtq ,uado ,pesa ,qntqg punor8 le pauuels-qFrx ueuo u 9.0 gl.0 lo ,{lpurds ,.uqo.] .U .DnE [(Stgl) 909 :l ssroldld ur leusslel^l:rg uou D^Di Dapadl puou)anlerr'u.rou{un er?p ,Erl'rlsnv urerse,^{ qlnos :(IN ..Iaw:orrri[3H-, l;r',"r1;g3t ,erT?rtsnv ,qtrod oraq opel qLZI sslaq 7'6€g1 ount g utetse/ ,[spu?l,4ewul] -ourd,, '1oatn!1ns ,,elddv reeu :adtJ lI6g tgg :Z ld urC sr^eU iel azlun; (:eussrallybinqdps .dle^A'{laustay,trt ,lYuW {ZS8l)SZf :[ rpJl)eur]s{SsrJuElog selpuuV oanitilns ,. - .1og ,leussrol{ Dldalsollo^lDJ (gtgl) 96€ :9 (ult.jeg)SuUroZ errnrldlns zalaurr4:g7

'tlpuowurup ,eur€u d Jopunll?lep alou ur potou sE s[l] uon?Jrldd? .la^o,roH Jo 130JJoaaql lnoqp lqnop oruossl olaql sorcadseurEs oql ot serlddB,{lq?qold 'opunqgto6f ,.4oo11o1ot1itacottou] g o:oJoqsJeo,{ e^rJ parlsrTqnd oapu4 erueuoqI .saloN 'orloltlsn8uo Dapatd ot :saso13 sa4tuglly 'laqolco ,{ln[ .pouadEntauolg tluar?ruo ro puBS doep u, pue rs'oc aqr Buol' ouorseurl ,.^3X",H:fli1H.'6'ffil3f orqrnos (E,r€0rrr 's.zz08z)uordu'quoN r'ouruo4 ro,g,itJ'lJJ i;5,.ojit)i,l,i#i# .o.v,uorpteroc - (HJ_UAd)888 ntoY Jos [ru{Is] rurzt :(DfJ)90rrs9 sdltrqd E'w'BreBuo( yo g trul 9.911!!9 9 :(ijJdad)gg kiqhp .y Viioraurzriuo'N Jo:l!t:!q.i:-4.l,.u1s11j,yllq+uqd"tarN)rqoiD,:rniLri'irt;Fw 1t :HJEAd) sauDI .W.V ,\red [u]I t tl lur 8,Ioar] B8Z 1euor1e51deqcuza :(fffUaai 698011lnUDuH oll"urooSreou :{HJ u{Il) i7g ,uoopul 1 U ,oorouuel& UAd)626 a1u13o 11 rsnt :{HJUAd)6Z9Z aS.toaD Z j(HfXlaj l, up.tD9 .re^r; :(SNVJ)gggz^rsptqtns 'uotpvlagS tSZZf V ) llrH 'W'V'.ErraaurapagJoJ'N goN lurftngfr,n oiitHriraJ,ai vea iqiii A :(Ov)7LSI Kqqsv 741 V,uo1p1eng1er;:GJUAJ,Of|i7L 'W'V'erooIN:(uoas .pauuola &ttrv S, ro^ouro.r; potcalos) VITVUJSnV f.fuAfSg'1i su)windg aeacPaelalu,{qlu?rlertsnv utots:/d .a,{u .l.g 204 NuytsiaVol 6. No 2 (1988) shorteror longerthan sepals;filament 1.5-4 mm long,anther 0 6 1.4x 0.2'0.4mm: connective ratherbroad but narrowerthan anther; slits semi lateral a{ter dehisc€nce Staminodes:hlament 0.2 L5 mm long;anther 0.2 0.5 x c. 0.1 mm. Ovarywith a terminaltuft of hairs;longest hairs0.4 0.8 mm long.S/)/e exserted by 2.59 mm. Seednot seen.(Figute 33 )

.*L-,/ t) rJ('lv =#Kn

)lV ./'l

t 2 S l t S n k c o, fi ei ni Fwure 33.Pimelea sulp,tulea. A flowenngstem of femaleplant: B bisexualflower (x 10);C opcneduppcr part hr (\ 8): D old stanren l0): E ovary (x 20).Drawn lrom freshmaterial represcnled by N. Cahen olTemaleflowcr t\ X 1005lA. ( . tr ! d \. Cahen1006 ,8. D.

l I 'sdunq pesrsrfnq8ns qlr^\ pauJo ed ,{lleurpnl6uol'rxru g x 'tueled I g 3 paas'uur E-€,{q pouesxaatt$'AuoI uItx g 0 01 dn srl?q :JI?qFs?q ur ^r?q ,{losj8dsseurleuos ro snorqeF,(/r O aruaasrqopleu? IeFlef[xas slls :roqlueuprll Jo,roll?u q.nu e^rtreuuoc:ruur g'0,S2 x g Jeqtu?:8uol rutu tuaulEllJ:sFdos Sulpoocxo 0 'ale^ol-SZ I t E .lruruns flTJolfssuawois'8uol uur g't-S 01 at?^o,41$o:reuqrdas le'uelp uru '8uol 'lluurns t g l-t ruru S 0l-9 o1,tll€npEr8 AurpuBdxe pu? cupull,{t [l1'lloJJeuaqnl ptog$o uotttod-a$is ruln I c x S tt aqru ptog[ to uolltod tt ,1O uorlrod^J?^o a^oq€turx I J olrsslcsuncJrc'Euol ulu -6 eqnl:snorqelS'ueo$ ol€d'eleueJ florEj Jo s/aMorur'8uol 'snolnpuod lenxosrq 'ruu rurx $r?q :8uol ruru ot dn sla1rpadF?duoc atuacsatoll[uJsnoJq?I8 €-Z 'eppru Z 'qsrpper 0Z-Zl x 9Z- .^Aolaq^lq8rTs lsep?orq ro 3rldlllo^peorq fllBr ?d sorunoruos pue uea$,rolle^ ro uear8unrpour ot oled'.{JI?Je1EIsrenAou oql Surprq'pexel+eJ Suruocaq 'sre^\ol] 'srred tou Surpunorns,{Josolc V-Z( ut spotq lonryo^uJ Suol ruur 9-I apunpad 'osnlqool alnc?'snoJqEIS 'uur x 'crldrTle,{liaorreu ol ole^o'ue3$ qsr,\\oflo^sou}euros 'snoroloJuoJ8-E'alsses 8l-9'asJorlu€ 'a Jo ueej8 unrporu 01 luelEd soddosa,lra1 sJeq ,{JEIIrxp 'ocuocsejouur JoJldeJxe sno:qep'xede uo{ JeqlrnJu,rorq-fer8 unrpeu Sulrumoq qc?o 'q8rq ',{pulds'tcolo rEouu,roJq'poJ sua1s le^elpuno$ 1?peluluols-eFurs u I-1 0 ?n.lryS '^ft^o,{ri€r{ssel pu? qedesJo8uol'sre,rolJ 'sa^ml uaar8el€d eqt Jo molm ueor8oqt ur sroJJrplnq fianI l dDsow8rua, dr ot polelau '0y\SN'TAI^I ') 'SNVJ :osriHJUAd :otoq) uaqq'N 't\6t BnV 'g,ttoZZI 'S 'eIT?JtsnV 'ecu?redsg 'pdV6t0t 'pU EI ,€€08€ urelsei& JoE lor.roH Jol&S url 9 t dnEurloof :sndfi 'osolrd snururou€Ao lo snquor8uol 'snqlput^ srtedos oprqedsnquog'snqpuh srrToJlraJJrp pos 11en14g aosouBn.a,A srur.JJv

ftt em8rC)^ou ds'o,{Usuapuad Balaurd tZ

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'pues eqr.roraqreeqs crrror?r Bq*.r ro lo^er' qrua r.rr"'0";:oJ,"l?li;u; fi"'fr?itry:## '(a,8z06ll'S,9totfc) ssorJuloqtnos r?ou 01 pu?lul puE 'S r) le^ru plpra8zlrj 's.Et"6z){g ,S[06]I ,90lrlt {led lEuorteN 01ls?e-qlnos (a , osll Bqq?eugreou uo4 spuelxll(gE etn?rd).uounEtJs.rQ (CV) 'd 'llorue0 'DgJ) 60Z€.t uosl!/4 Jo,4A uDl 08 c :(HIUAd uoelrDl A T dslt) O W.E6Btot(oJ 'qllusurBH g ur1 :0UB) piDMoiS'l'{co{ ernrg l(HIUad) n[ou .e.X 'Ired Jo S0 'ouuv6Zl 'lrerun)6Z16 leuorleNre^ru pleroEztrg turodJo N :(.Igl{) ZLf.t.\nW grJ JoIANIA ru{ ruH 'uot{oorg gN onpur1 :(HJUAd 'rslurd08:(HJUEd) 9Ll06L ln! T Ur 'BqqeougJo / t Z6p^oHt o{e-IJo gN :(HJ{Ad) W6 u{fut tr? JoS ru{ r:(SNVJ) 6ZVuoauo) 7'ired s8ur):(CgJ) 3u1uuo2 prE,rotul8UIO ruorJ 99S€89'Eun^olqun0 l,ttr'g'uooputg [ru{ ,] FU.9.,11S.t :(SNVJ)t09S aSzqln€r Z N puBur8e^\ uao,,rloq :(HJUAd) f6rE Zo\rDtg I 'utqnnA:Hldgd 'N\SNI plDag 'rnonso-I :(HJUAd,(]VJ Kqqsv.W.V 'onfloqur{ VZBL 51 }W CZLZ I Jmu:(uses 96 re o ruo4 poliatas)\IITruJSnv NdaJSa,4&pamama suawnads asacsa?leur^qJuerlerlsnv ur.tselA 'i{U 'l g NuytsiaVol. 6, No.2 (1988)

r' ,',?M \'"'t'V" / ,f!t/ (x (x D- ovary(x l4). Fil.]te 34. Pimeleapenders. A flowefingstem; B .llower l0);C stamen l4); Drawnfrom fre'shmaterial repreiented by lhe type collection.N. Co&e, 1019.

Specimensexamined. WESTERN AUSTRALIA: Howick Hill, 20 Sept l968,,DG Austin (iO); Lucky Bay,E.M. Bennett8g7B (PERTH); 0.5 km-E of-C-lvde Hil' MA lyrgryan 1.197'&S.McNie(PERTH);Mt. Ney,M A. Clements1790 (CANB); tvPg localirv, N Cohg! 10198(PERTH); 39.5 km NE of Gibson,N. Coh9rl1021, l02la (PERTH);c 3 km NE of HowlckHill, H. Eichter19866 (AD, PERTH); Mt Afiq .4.S.George l4]! (P-ERTH); c. 25mi [40kml E of Esperance,M.E. Phillips3197 (CANB); Howick Rd., SW of Mt Nev' 'osou8ntaD oapuld lo uorlnqulslc 9t alntrj :

I I I I

'a -S€ suapuadd pue'V odnqdlnsoalarijd Jo uotnqutslc alnBrl I I L] i I-\ Y --Lr Y \i,\\ /4 ) ",p.i^€i )aa Lr w 1""fl""1_ l_ f V j ]'r ! w ll'g

J Y I I I 7\1 'suapuad d ut snorqeF,dlelalduor sfE,rlE eJ?sl"des eql lnq sledosfurBq ,{llerued amr4 nsoaEruao } So sueulrcodseulos speesoql uo ulallpdeq1ur pue slce:qpue sulolsBuno,{ eq1lo:no1oc eq-1 ur 'srsou8?tp mouSruao 4 utotlsJellp suapuadd eq1u uo rAsacue.ra;rp eql uo:1 geOy .ue,lto1i aql Jo oprslnoeql uo slr?q alalduor s! ^q suaJon urolJpeqsln8utlsp ,{ltp?ol oq Jo 'd{cel d uux suapuadDapwlt suaJuz ol Jelurs osle pue ,so u8ruao oalawt4 o11saso13.siutug[{y 'smua3ssJolJur ,u,{Aop - -Il .awou snolnpuadaq1 o1 8ur::e;ol Bu6u?q ( suapuad lo uopotuag 'lsn8ny ,{etrAlpouad Suuauolg peorpor'elr uo,{ero ^puBS,{ora rou,rorq u,oryn,r.oo1rtlj."3ilffi?t;ft:lrB:lffirffii 'la,zrotzt's 's ,69ott)plrv rI trot tseo(E ,0t.rzzt ,8S0€t){?ed uerxqcuelCuo{ lsBoJoq1 Suol? pu€ lsBoc qtnos aqt ot ,S,ZZotE) 'S lE,690ZZI IIIH opftJ pue lg ,6Z"ZU .1.20€t),(aN ltr{ reeu uorJ spuarxAfS€ atnir4 ioinqliiitg '(HJ{gd) 'IlHrdrssrssr],{ 9//9 uotsall S'y :(IUg) uosuqofquDrs ,sllrHutooua rlA reeu:(HJXgd) "pd 699 Z It1Zg a[A TS {cr,rAoHJo,t[S ru)t 9.t..pX dnurlooJ:(HJXAd) ;ZOZS5iy-7:g o?ea?oslaur,{qluerlErtsnV u.ratsalA'rf U'I g a 208 NuyrsiaVol. 6. No. 2 (1988) 25. Pimeleaaeruginosa F. Muell.. Fragm. 7: 2 (1869).- Pimeleasylvestris var. aeruginosa {F.Muell.) Benth., Fl. Austral.6: 11-12(1873). rype: south-western Australia, date unknown, J. Drummond (holo:MEL). Shrub,erect, spindly, 0.2 L5 m high, singlestemmed at groundlevel. Srems yellow'green neareach inflorescence, becoming red-brown then greyfurther from apex,glabrous except for axillary hairs.Zeaues opposite, patent or antrorse,sessile (but narrowly taperedat base) or with a petioleup to I mm long, glabrous;lamina concolotous.bluish to greyishgreen or sometimesyellowish, narrowly ovate to narowly obovate,'l22 x 2.5'7.5 mm,rather thick, adaxiallyconcave, obtuse or nearly actie. Peduncle1.5 mm long.Involucral bracts rtstally in 3-6 pairs,closely surrounding flowers, not becomingreflexed, broadly elliptic to almost circular, 11-25 x 7 I 7 mm, glabrouson both surfaces;outer bractsnot ciliate;inner bracts yellow or yellowish,sometimes with a few fine cilia c. 1 m long aroundapex or lower on margins.Inflorescence pendulous, cofipact. Pedicels 0.5-1 .5 mm long; hairsup to 2 5 mm long.F/owers bisexual or rarelyfemale, dark-coloured below the distinctcircumscission point, yellowabove; tube I l'1 5 mm long,circumscissile 0.7'l.2 mm aboveovary'portion, glabrous. ONaryportion of|lorul tube 3-4x 1-1.5mm. Style'pottion of Jloral tube narrowly cylindric but enlarginggradually to summit,8 I I mm long, c. I mm diam.at summit.Sepals elliptic- ovate or narrowly so to elliptic, 3-4 mm long, glabrousoutside or with a few hairs along midrib,glabrous inside! haim up to 1.25mm long,very fine. Stamens shorter than to exceeding sepals;filament 1.5-4 mm long;anther l-1.5 x 0.3-0.6mm; connective much narrowet than anther;slits semi-lateralto lateralafter dehiscence.Ovary haity throughoutbut sometimes only very sparselyso: hairsappressed to antrone, 0.3'0.5mm long,very fine. ,S/y/eexserted by 4'7 mm. Seedc. 4.5 x 1.5mm, with longitudinalfurrows. (Figure 37.) Specimensexamined. WESTERN AUSTRALIA (selectedfrom over 80 seen):between Perenjoriand Carnamah. J.C. Anway 533(PERTH); 3 mi [5 km] W of Kulja, T.E H. Aplin 536 (PERTH);3 mi [5 km] S of Wubin, T.E.H. Aplin 558(PERTH); Tammin, A.M. Ashbv 912 (AD); Muntagin,E. ?l Bailey 639(PERTH); 1l mi u8 kml Eof Narembeen,J.S. Beard 5908(CANB); near Walgoorlan, W.E. Blackall 4002 (PERTH); l0 mi [16 km]N of Trayning, M.l.H. Brooker 1891 (PERTH); 13 km E of Koorda, M.D. Crisp 6539 (CBG); S Yanneymooning,P. de Rebeira171 (PERTH);Wilroy ,4.R. Fairall 1483(PERTH); 5 mi [8 km] E of Burracoppin,R. Filson 212 \MELI; Wyalkatchem,C.A. Gardner 179 (MEL, PERTH); 39 mi [63 km] E of Pingaring,A.S. George9340 (PERTH);Ballidtt, E H. Ising 93 (AD); Cowcowing,M. Koch 1009(MEL, PERTH);Merredin, M. Koch 2111 (MEL); 20 km W of Tandejin,R.H. Kuchel 2063(AD, CANB);39 mi [63 km] E of LakeKing on Norseman road. F. Lullftz 5027 (PERTH); Frank Hann National Park, D. Monk 398 (PERTH):near Gabbin. B. Rosier82 (PERTH);I I mi [18 km] N of ltbberding,R A Saffrev 861(PERTH)I l5kmNof Trayning,P.S. ValentineT44 {PERTH); Eof Caron,F.W.Went 98 (PERTH);I mi u.5 kml E of Wyalkatchem,E. Witt^,er1225 (PERTH)

Distribution.(Figure 36.) Extends from nearMullewa (28036'S, I 15024' E) south'eastto north of StokesInlet (c. 33"06'S, 121010' E).

Habitat. Overmost of its rangethe speciesoccuts in sandor sandyclay, often mixedwith gravelor overlyinglaterite, and the soil colour is commonlyyellow. In the south-east,the ioil is sometimespale brown and overlyingclay. The vegetationis often mallee'dominated.

Floweringperiod. Mainly JuneOctober.

AJJinities.Closest to Pimeleaperulens and P. cracensSee notes under thosespecies

Notes.The specimenswith hairson the outsideof the sepalsare all from the south-eastern part of the speciesrange. 'srsold srE?ruslledos le eJol^olqlpue^o euoluodoJd 'oso[d 'snn3? 'C onburuold IIsJolJ oqnl sIIloJ lJoJJp pas IonW aosouldruaDd slulJJV (g€ amEld)'^ou'ds'o^U suar8lJBalaluld gZ

'q 's 'v) pu'(a izql uallocN ^qpecesetdorprrorpur qsorJ .rorJ u.^\Ero (g xi "*9 s 1?r"l)z9j#rJ-'e't x) tel\ou elauoJ -J :(8 x) ro^\oU l€nxosrq I :lu9ld IsnxosrqJo urels 8uue,{ou ,V DsouBuEDoalaard Lt aJnSt,I

V B

'e,{X 'I 60(, aEaceoeleu^qluqleJlsnv uralso^\ g 1rr11r^O-9qor1.rnd,8.3 km NE oJ Norwood Rd,NE of E.*r""*, J.];;::,u,ril iffif S, 122005'E,13 Aug. 1983.N. Cohen1023 (holo: PERTH; iso: CANB. K. MEL. NSW).

Relatedto P aeruginosaF.Muell. but differsin the acuteleaves, the usuaryhairy floral tube with a shorterovary-portion, and the more hairy sepals.

^ Sfrr"rb,erect, spindly, 0.4- 1.5 m high,single. or sometimesmulti,sremmed at ground level. Srens yellow to palegreen near each inflorescence,becoming reddish thenrnldiu* grey brown further from.apex,glabrous except for axillary hairs.Zea,es opposite, usually pitent or antrorse, sometimessomewhat reflexed, glabrous; petiole 0.3-i.3 mm longi timina concolorous,yellowish or mediumgreen to deepbluish green,narrowly elliptic to ovateor nearly so, 6 22 x 2-6 mm or rarely lessin the smallestleaves of a branch,ihinner than in I aeryt&inosa,flat or adaxiallymncave, actre. Peduncle),-4 mmlong. Involu*al bracts rs,ally 6 or 8, rarely 10, paired,closely surrounding flowers, not becomingreflexed, yellowish oi pale green,sometimes with reddishportions or the outer bractssometimes fuily or partly thesame.colour as leaves, elliptic, l2 28 x 6 l4 mm, glabrousor sometimeswith a feri,appiessei hairs insideat baseor along middle.Inflorescence pendulous, compact. pedicels uiually c. I mm long_;hairs 1 2.5 mm long.-Flowers bisexual or rarely female,crcam or creamygreen to-paleyellow, glabrousinside floral tube and sepals;tube 7.5,12mm long, circumsc-issile 0-52 mm above ovary Wrlion. Ovaryportion oi Jloral tube 2.2.5 x c. 0.b'mm, hairy or glabrous;hairs antrone, 1-3mm long, very fine, the longestones exceeding those on siyle Wrtion. Style-portionofJloral tube very narrowly cylindric-,7 9.5 mm long in-bisexualflow6is, 5-6mm longin femaleflowers,0.7 1.2 mm diam.at summit,hairy or glabrous;indumentum (whenpresent).of very fine, widely antrorseto patenthain, commonlya mixture of long and_shonhairs in the range0.3'2.5 mm long,sometimes consisting of a few longhairs restricted to distal part or sometimesthe hairs widespreadbut all short.sepals elliptic or nearly so, 2.5-4.5mm lo:rg,hairy outside;hairs 0_ 3-2.5 mm long.Stamens slightly oi Oefinit"tytong.i than sepals;filameat 2-4 mm long;anther c. I x 0.3 mm; connectivenarrower than anthir; slitssemi-lateral after dehiscence.Staminodes: filament c. 0.2 mm long; antherc. 0.35 mm long. Ovary hairy throughout;hairs appressed,very iine, the longestones 0.3-l mm long. Sty/eexserted by 4,6 mm. Seedc. 4 x 1.7 mm, with faint longitudinalmarkings.

Disttibution. (Figure42.) Extendsfrom the Donnelly River eastto near IsraeliteBay and inland north to Kumarl.

Floweringperiod. July Novemberand rarely December.

Deriyation of name. Cracens(L.) - graceful,slender, referring to the habit.

Affinities. Closestto Pimelea aeruginosaoveralL ln vegetativemorphology more similar to P tinctoriaexcept in the lack of cilia on the involucralbracts. The leavesof both p. cracens and P. tinctoria are not as thick as those of P. aeruginosa.

Notes.The two subspeciesrecognised are allopat c. They appearto be distinguishedonly by the presence/absenceand type of indumentumon the floial tube. Howevei, they show no intergradationand are regardedhere as beingjust distinct enoughto be recognisedas subspeciesrather than as varieties Key to Subspecies

l. Floraltube completely glabrous or with a few hairs1,2 mm long nearsulnmrt...... 26a.stlbsO. Slabra 1. Floral tube hairy throughout,the shortesthairs of the style-portion0.3'0.5 mm long...... 26b.subspcracens 6e0z q aa w,l drr .er l?rsnv u,r,,.16, on.,,,t#fl11"tfi T,;i .g*H"Tl;T-+i"tt #,?? 'suqBJ8qns .d le,t s:qe13snqtrolJ atua'on dsqnse,{g au DD e ue!!(I

'^ou dsqns'e^U srq?13.dsqns .e97

N {qpru.s.rder re!*reur "r.r.,rno.3 !._.g!,,u1,or')uslo lrsalJgllr ylglc;itr .sir:jrirlasqns:l/1'.Bi,ii,ig,l'[1if",]",lffi !?"d1"1 JleurqJo ru.tsBuLrrl\olJ B :rueldlenxrsrq 1o u:eliBiirie,nog _y vaioLi oalawti.gi atiat\

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212 NuytsiaVol 6. No 2 (19881 Differs lrom P. cracensRye subsp.oacens tn the glabrousor almostglabrous flowers.

Floral tubec0mplelely glabrous or with a few hairsI 2 mm longin distalhalf of style-portion.

Specimensexamined. WESTERN AUSTRALIA: Donnelly River plains,W ol Manjimup, A.M. Ashby 504(PERTH); Muir Rd, E of LakeMuir, W.R.Barker 2347 (AD); Mersea Lake, Wilgarup,Nov. 1962,W.A. Loneragan (PERTH).

Distibution.(Figure 42) Extendsfrom the DonnellyRiver (c.34013'S, 115055'E) east to LakeMuir (34026'S. I 16025'E).

Habitat: Apparcntlyassociated with wetlandsbut no habitat detailsprovided on specimen sheets.

F lowering period. August-November.

Derivation of name. Glaber(L.) - hairless,referring to floral tube.

26b.subsp. cracens

Floral tube hairy throughout;ovary-portion with fine antrorsehairs l-3 mm iong; style portion wirh a mixture of short and long hairs or with only short hairs. the shortesthairs 0.3-0.5mm longand longesthain up to 2.5 mm long.(Figure 38.)

Specimensexamined. WESTERN AUSTRA.LIA (selected from over90 seen):near Hamersley River, Oct. 1903,C. Andrews (PERTH)ILake King, J.S. Beard2185 (PERTH);Pingrup, W.E. Blackall 3025 (PERTH); Circle Valley, A.J. Cough l0 (PERTH); Buyi Billanak Homestead,SE of CondingupPeak, NN Donner 2662 (AD, CANB, PERTH); c. 58 km Nof mouthofOldfield River,11. Eichler20379 (AD, CANB, PERTH);near Truslove, Cl. Gardner 14190(PERTH); c. 80 km ENE of Lake King towardsNomeman, L. Haegi 991 (AD, PERTH); Kumarl, l.l. Horbury 69 (PERTH); BremerBay area,A. Kessell996 (PERTH)tSalmon Gums, R.H. Kuchel 1731(AD); W endof LakeKing, F. Lulnh 5538 (PERTH)Ic. l2 km N of lsraeliteBay, EC Nelson(PERTH): Frank Hann National Park, K. Newbey5564 (PERTH); c.3kmNEof HowickHill, LE Orchardll3l 1AD,PERTH); 25 mi [40 km] W of Ravensthorpe,S Paasr7l I (PERTH);35 km E of Gnowangerup,B.L. Rye 82015(PERTH); 6.5 km W of LakeBiddy. R.l. Sa|frey593 (PERTH); Frankland River, Nov. 1932.H. SteedmanI.PERTH); 10.5 km NNW of Ongerup,M.D. Tindale3892 (CANB, PERTH); Mt Madden,P.G. Wilson6828 (PERTH)r6 km from Bordentowards Amelup, J.W. Wrigley 68485I (PERTH).

Distribution(Figure 42.) Extends from nearMount Barker(c. 34038' S. I170,10'E) eastto nearIsraelite Bay (c. 33037'S, 123052'E) and inland as far north as Kumarl (32'47'S, l2l'33'E).

Habitat. Ocatrs in sand-vclay or rarely sand,often in shrublandsdominaled by scattered mallees.

Flowertngperiod. July'Novemberand rarely December. t f 27.Pimelea tinctoria Meissner in Lehm.,PL Preiss.l:603{1845) Calr-ptostegiatincLo a b (Meissner)End 1., Gen. Pl. Suppl.4: 6l (1848).- Pimeleasuaveolens yar. tinctoria(Meissner) a Benth.,Fl. Austral.6: l4-15(1873). - Banksiatinctoria (Meissner) Kuntze, Revis. Gen. b "Mt Pl. 2: 583(1891). Iype: Wuljenup"[Willyung Hill], WesternAustralia, 20 Oct. 1840. e L. Preiss1280 (presumed holo: LD; iso: MEL). o

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214 NuytsiaVol 6,No 21l98E) mm long. Inflorescencependulous, compact. Pedicels 0.5'2 mm long; hairs I 2 mm long. F/owers bisexual,apparently dark red-brownbelow the circumscissionpojnt, yellow or yellowish green above, glabrousinside floral tube and sepals;tube 17.522 mm long, circumscissilel-2 mm aboveovary'portion. Ovary'portion of Jloral tube 2.5 3 x l-1 5 mm, coveredby long hairsand morenumerous small hairs; Iong hairs antrorse to patent,2-4 mm Iong,usually rather coarse; short hairs commonly 0.1'0.3 mm long.Style portion offloral /rb€ narrowlycylindric but expandinggradually to summit,14 19mm long,1- 1.5 mm diam. at summit,with a moderatelydense mixture of long and short hairs;Iong hairs patent, 3-5 mm long,finer thanthose on ovary portion;shofihairs patent, mostly 0.1-0.3 mm long.Sepais elliptic,3.5-5.5 mm long,sparsely hairy; indumenrum mainly of smallhairs. with a fewlong hairssimilar to thoseon the floral tube.S/am€ns shorter than sepals;filament 1.5-4mm long; anther l-1.5 x 0.5-l mm; connectivemuch narowet than anther;slits semi lateral after dehiscence.Orzry ratherdensely hairy; hairsappressed, 0.2-0.4 mm long,fine. Sryle exserred by 3-4mm. Matureseed not seen.{Figure 39.)

Syxcimensemmined. WESTERN AUSTRALIA (selectedfrom over40 seen):Mt Manypeaks, A.M. Ashby 1950{AD, PERTH);between Mount Barkerand Albany, 13 OcL 1968,E.M. Canning(CBG),near Denmark,,4.B Cashmore 2l (PERTH);6mi [9.5km] Nof CapeRiche, C.A. Gardner2Il9 (PERTH);King GeorgeSound, -8.I Goadby38 (PERT[{);South Stirlings, F. Lullftz 3458 (PERTH); 6 mi I9.5 kml S of Narrikup, R. Melville 4400& R.D. Rovce (MEL);Toolbrunup, Stirling Range, ZB Muir 3937(MEL); 65 mi [104km] from Albanv towardsJeramungtp. S.Paust 543 (PERTH)I Stewart Road, 18.9 km from VasseHwy, lR Wheeler2ll1 (PERTH),65 km NE of Albany towardsJerramungup, D.l E. Whibley5240 (AD);Chester Pass, Stirling Range, 13 Oct. 1961,J.H. Willis(MEL); Millbrook Rd, N of Albany, E Wittwer 211 (PERTH).

Distribution.(Figure 43.) Extends from near Denmark (34057'S, I 17021'E) east to nearCape Riche(34035' S, I18043'E) and inlandto the StirlingRange (c. 34050'S, I l8'28' E), with a disjunctoccurrence south west of Nannup(c. 34'12'S, I I 5036'E).

Habitat. Recordedin sand,sometimes with gravel,in low shrublandsor in clearings.

F loweringperiod. August-October.

A|rtnifies. Most similar to Pimeleasuaveolens in floral morphologyand to P. cracenstn vegetativemorphology.

28. Pirneleasuaveolens Meissner in Lehm., PI, Preiss.I : 603'604(1845). - Calyptostegia "Greenmountain" suaveolens{Meissner) Endl., Gen. Pl. Suppl. 4: 6l (1848).Iype: [Greenmount],Perth, Westem Australia, 13 Sept. I 839,Z. Pens 1268(lecto here designated: LD; isolectoMEL, NY); southwestem Australia, J. Drummond548 (isosyn: MEL).

Shrub,erect,0.25- 1.2 m high,often multi'stemmed at groundlevel, unbranched or with few branchesand open above, regenerating vegetatively ftom a fusiformunderground swelling. Sternsyellowish first then red-brownnear each inflorescence, becoming grey brown or pale greythen mediumto dark greyfurther from apex,glabrous except for axillary hairs.Leaves opposite,antrorse or sometimespatent, tending to all point verticallywhen on an oblique to horizontalportion of stem,glabrous; petiole up to 2.3mm long,lamina concolorous, usually eithermedium green or glaucous,narrowly ovateto narowly obovateor ovateto obovate, 5 34 x 1.5-6mm, thick, adaxiallyconcave, acute or obtuse.Peduncle l'4mmlong. Involuoal bractsrn 4-7pairs,sometimes becoming reflexed in fruit, moreyellowish or rarelymore reddish than leaves,often pale brown in fruit, narrowly ovateto broadlyelliptic, 6 28 x 3-14mm; outermost2 or 4 bractsglabrous outside, sometimes ciliate, usually at leastpartially hairy

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'a,{U 'I apareoeleuqf uurlsrtsnvurelsolA B 216 NuytsiaVo1. 6, No.2 (1988) occurringnear middleof eachmargin, (1-)2-5 mm long, very fine. InJlorescencependulous or very rarely erecl,compact. Pedicels usually c. I mm long; longesthairs 1.5-2.5mm long. F/owersbisexual or very rarely female,dark-coloured below circumscissionpoint, pale to deepyellow above;tube (8-)9-18mm long, circumscissile1.5-4 mm aboveovary-portion, glabrousinside. Ovary-portion ofJloral tube 1-5-3x c. I mm, with appressedto patenthain 0.1-0.6mm long,which areoften mixedin distalpart with long hairs usually2,3 mm long. Style-portionoffloral tube very narrowly cylindric,scarcely enlarged at summit,7-15 mm long,0.8- l.l mm diam.at summit,with a mixtue of hairsof variedlengths, some hain minute and somemedium'sized to long;minute haim patent,usually 0.1,0.3 mm long; longerhairs patentor widely antro6e, becomingprogressively shorter from baseupwards, those below circumscissionpoint 2-5 mm long and sometimesdistinctly coarser than thoseabove, those at middleof tube 1-3.5mm longand very fine, thoseat summit (whenpresent) l-2 mm long. Sepalsnarrowly ovate to ovate,(2.5-)3-6 mm long, the indumentumon outsidesimilar to that of distal part of floral tube, glabrousinside or with a few minute inconspicuoushairs below apex.Stamens shorter than or as long as (rarelyexceeding) sepals; filament 1.5,3.5 mm long; anther 0.7-1.6x 0.3-0.4mm; connectivenarrower than anther;slits semi-lateral to almostadaxial after dehiscence.Ovary moderatelydensely hairy; hairsappressed, very fine, the longestones 0.3-0.4 mm long.SOr/e exserted by 2,6.5mm. Seedc.6 x 2 mm, with longitudinalrows of minute pits. (Figures40, 41.)

Distibution. (Figure44.) Extends from nearJurien Bay sourh-eastto Albany and inlandto near Coolgardie.

Floweringperiod. June-October.

Affnities. Closestto Pimeleatinctoria in floral morphologybut with leavesmore similar to P. aeruginosa. Nores.Two subspeciesare recognised. They appearto be geographicallyallopatric from the availablecollections. Specimens occurring at the edgesof the apparentdisjunction tend to be somewhatintermediate in morphologybetween the two subspecies.

Key to Subspecies L Leavesnot glaucous.Inner involucralbracts yellowish green or green or sometimespale brown or reddish.Floral tube 12-18mm long ...... 2 8a. subsp.suaveolens l. Leavesglaucous. Inner involucralbracts yellow or rarely yellowish green.Floral tube8-14 mm 1ong...... 28b.subsp. flava

28a.subsp. suaveolens

Pimeleamenkeana Lehm. ex Meissnerin Lehm.,Pl. Preiss.1: 604 (1845 ). - Calyptrostegia menkeana(Lehm. ex Meissner)Endl., Gen. Pl. Suppl.4:61 (1848).- Pimeleaymyeolens var.menkana (Ix,hm.ex Meissner)Diels & E. Pritzel,Bot. Jahrb.Sysr. 35: 395 (.1904). Type: south-westernAustralia, Oct. 1840,L.heiss 1269(lecto here designated: LD; isolecto:MEL, N\4. S7 Leaves:lamina not glaucous,narowly ovate or narrowly elliptic to elliptic, (5-)934 x n( (1.5)2.5-6 mm, acuteor narrowlyobtuse, sometimes mucrontlale. Involucral bractsUsu,ally N yellowishgreen, sometimes green or rarely somewhatreddish, narrowly ovateto narowly ll ellipticor ovateto elliptic, l0-28 x 5,14mm; innermostbracts glabrous outside. Floral tube of 12-18mm long.Sepals 3.5-5.5 mm long.S/amens shorter than sepals; anther 1.2,1.6 mm long. 2t (Figure40.) 20 .t, ./t/ ,oolorooiA ,aNVl ,cv) ,tz6 uosu)ohl llorsrul3) :(Iug) 06fl rlro) :(HJuEd 'rnanso-I 902 [Dp IoH V J'ru?C {oorg onSol :(HIdAd) (,961u{1r9 tsE tl{ ru{ ,olepwljefJo l[N Z :(CSJ) tt0t 33ng y1J'dnuueBa8plCJo ,4[ rul 8 :(Cy) LgtI .tauuo(JNN Jo ur)t r :(HJUAd pp{u't) y ' pou :(DgJ) 8uluuD) .W.g ,896I .tro 'dnuueNAN S ZW I H 9I spre,roluotraquod uro4 :(gNVl) aSprqrn8r ,ur8olleN 're{rpg [rx)I I ] Iiu , 01 9 t €Z J N :lldq) Dlolg lunot :(HJUAd '(JyJ iqLfsv ,dn8unrr?ldreou 'e8uexL9B0Z J S { 9992 l,t/ t' SurTruS:(ures 09 [ lo^o tuo4 pepoles)VIIVUJSnV NUAJSAA pauwoxasuaa2ads

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LIZ aea.pa?laurlqluprtEltsnv ulatse/1 J{U l B 218 NuytsiaVol. 6, No. 2 (1988) (BRI); Gingin, A. Morrison 12173(PERTH); 6 km W of Lake King, K. Newbey 2703 (PERTH); 3 mi [5 km] from Tallanallatowards Ha ey, M.E. Phillips 4068 (CBG, MEL); Rosa Glen, Margaret River, R.D. Royce 2765 (PERTH);near Yarloop, F.W. Went 232 (PERTH).

Disfiibution,(Figure 44.) Extends from Mt Lesueur(30011'S, 115012'E) south to Cape Naturaliste(33032' S, 115000'E) and from thereto Albany (35002'S, I 17053'E) and eastof the Stirling Range(c. 34050'S, 118028'E).

Habitat. Usttally occurs in sandy clay, always recorded with gravel or laterite rocks or underlyinglaterite, in forestsor woodlandsor sometimesshrublands, often on hillsides.

Flowering per iod. June-October.

Noles. The hairs on the floral tube in this subspeciesare usually longer than thosein Pimelea suaveolenssubsp. flava but some specimensof P. suaveolenssubsp. szaueolensfrom the southernpart of the rangehave small hairs. The involucralbracts, especially the innermost ones,are usuallyless hairy than in P. suaveolensstbsp. flava. 28b. subsp.flava Rye, subsp.nov. (Figure 41.) Diffeft a P. suaveolenleMeissner subsp. suaveolente foliis glaucis,bracteis intensius flavis et plerumquefloribus minoribus.

Typus:9.6 km S of Bodallin on South BodallinRd, WesternAustralia, 27 Aug. 1983,N. Cohen 1027 (holo:PERTH; iso: CANB, K, MEL, NSW).

Differs from P. suaveolensMeissner subsp. suaveolens ln the glaucousleaves, more deeply yellow involucralbracts and usuallysmaller flowers.

Leaves:larrina somewhatto very glaucous,narrowly elliptic to elliptic or narrowly obovate to obovate,5"13 x 1.5-6mm, acute to obtuse,not mucronate.Involucral bractsyellow or rarely yellowishgreen or the outer onespartly green,elliptic or sometimesbroadly elliptic, 6-l6 x 3.11 mm; innermostbracts often hairy ou,tside.Floral tube (8')9-14mm long. Sepals (2.5-)3-4mm long.Stdmers shorter than to slightlyexceeding sepalst anther 0.7-1.4 mm long. I I Specimensexamined. WESTERN AUSTRALIA (selectedfrom over 55 seen):2 mi [3 km] Wof Gilgai Siding,?lEI{ Aplin 1972(BRI, PERTH); Koorarawalyee,,4.M. Ashby 284'l (AD, BRI, PERTH); Hyden, M. Barrow 44 (PERTH);between Buracoppin and Moorine Rock,lS Beard 6219 (PERTH); Frank Hann National Park, D. Butcher 308 (PERTH); c. 18.9km WSW of Coolgardie,R.l Chinnock3092 (AD); 8mi [3 km]N of Bodallin,CF. 1 Davies254 (PERTH);2 mi [3 km] SW of QueenVictoria Rock,R. Filson 8903(MEL); 0.5 ^ mi [0.8km] S of Lake King, E LullJitz 5530(PERTH); 80 km WNW of Kumatl, T.B.Muir 4374(MELI:22-23 mi[35'37km]Wof SouthernCross, 10 Sept. 1968,ME Phillips(CBG, PERTH); C amphorne,N of Narembeen,R .D. Royce 7849(PERTH); Mt Caroline,1892, ul M. Sewell(MEL); Mt Gibbs,Dec. 1929,H. Steedman(PERTH); Muntadsin, T.W. Stone th & E.T. Bailey 813 (PERTH);90 Mile Tank, c. 80 km W of Daniell,P. Wilson3206 (AD). to

Distribution (Figure44) Extendsfrom Mt Caroline(33048'S, 117038'E) in the west to near l-, Coolgardie(c. 3057' S, 12S58'E) in the north-eastand to nearPeak Charles (32046' S, 121017' gr E) in the south-east. gr SUl Habitdt. orcuts in sandor sandyclay, usuallywith gravelor lateriterocks or underlying probably mallee an laterite, in shrublandsor open woodlands. cili 'olBlllc'slc?lq eql l?Ipeu lo lsouJelno u"r{t Jol[?u]seq ot Bupual spelq $outouur :alerlr. 'ruur u po^Jnco].ruul ,oie^o,{lp?ojq lou ,-Z lelslp gl-1.x tz-l I ro e1e,roslce:q lerpour pue lsorurolno:oprslno pue oprsu snoJqel8.poxeger Aururocaq ,xade 'ecu"Jsorouur 1ou 1epa,r:nCi: ,ilueppns reqler ^[ensn Aursol3uo^llq8lt pu€,tlJelalelBurddelre,ro^,i11ea:B 'qsppo-J JoJIEq ,edulsl?seq ,{leJBJstc?lq Jeuu eq} I€4uoi ro as8qqsppal e qjr,tr slcerq :e1no aql ,ueeja -,no11e, ,perBd,etour o1ueerE apd ,{lpnsn sorurleuosJo g.9 slciq ptcryoaLzT.Buol-ruru y_ ,epprur 1 apunp_A etnm.qlensn leg,rxtu gl,S.i x gt-01 .roleq ,tlrqagsrsepeorq:o cnOig6 'ueara ,snoroloruoc ,opses .pexegel ol rrld o ,{l,roueu umpou ,tpldfis rlirar :6lueied 6l esJolu? 'elrsoddosa^oa7 'sJrcq ,xede ^Jellrxe JoJldacxo snolqel8 ruorg:oqili1 fe$ {JEp uaql u^{orq-por SurruoJoqpu? acurJserogw qc"a Jeeu uoeJ8oled ro u,norq_,ito1iefepd f'ypnsir suars punojS .q3q ,qih5 la^el l3 tusrpruur sz ol dn pu? poruuels-elturs u 2,r.6',neia :{,Tf#;d {3..,?,.,".rA):roq)sz,u

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,o,tu a?ajuonlaur,{qJusrlellsnv uletseid I g NuytsiaVol. 6, No.2 (1988) cifia2-2.5 mm long. Inflorescencependulous or sometimeserect, compact Pedicels)tpto 2 mm longi hairs iostiy 1'3 mm long. F/owersbisexual, white or cream,glabrous inside; tube l9-2imm long,without definiteovary. and style-portionsbut broaderbelow the slightly constrictedcircumscission point than above,the broaderproximal portion much longerthan 4 ovary.Proximal portion ;f Jlorat tube 8'10 x c. 1 mm' with deciduousantrorse hairs 3 *'o iong,glabrous in fruit. bistal portion offloral tube very narrowly cylindricexcept neal summit,-if-t Z mm long,c. 1.5mm diam.at summit,with patenlhairs: hain variablein length' the longesthairs 2-3.5 mm long,the smallesthairs usually c 0.5mm long Sepalsovate,4-5 mm lon-g,with hairssimilar to thoseon distalpart of floral lube.stamens slightly exceeding sepals;fi-lament 3-4.5 mm long;anther l 4-2x 0 5-0.6mm; connectivemuch narrowerthan anthe;; slits semi-lateralafter dehiscence.Oucry glabrous ,Sttleexserted by 3'5 mm Seed c.6.5 x 1.1mm, with longitudinalrows of smallbut well developedpits

Specimensexamined. WESTERN AUSTRALIA (selectedfrom over 40 seen):between Mt D B'aringand Mt Ragged,J.S. Beard 533'7(PERTH); second Legch W of Nanarup,1982, Daviison (PERTHJ:Duke of OrleansBay, tL Demarz6291 (PERTH); c. 8 km W of IsJaelite Bay,N.N. Donner 2827 (PERTH)Inear Cape l-e ply!3 ' p e. Qgrdner 14733{PERTH)i .. i0 k. w of Mt Ragged,R.H. Kuchet 1638(PERTH); 3 mi 15kml N of Esperance'K' (PERTH); Newbey2564 (PERTH); Dempster Head, Esperance, B. L, \ste 8Jp2l Esperance, B.L. TLrner 5523 (MEL); East Mt Barren,E. Wittwer 372 (PERTH); FrenchmansPeak, J.W. Wrigley 685400(CANB).

Distribution.(Figure 44.) Extends from Mt Gardner(35000' s, I l80l I' E) eastto neal Islaelite Bay (33037'S, 123049'E), alwayswithin 40 km of the coast.

Habitat. Mainly recordedon sanddunes or sandysoil overlyinggranite

Flowering period. June-August.

Affnities. No closerelatives but certainly has affinilies with.Pimeleosuaveolens and tts alhes, in6tuOingthe SouthAustralian species P macrostegia.Itsleaves are rather similarto those ol P. physodes.

Noles.The namePimelea macrocephala Hook.. which was published prtor to Calyptrostegia drummondiiTurcz.. has olten beenapplied to this taxon becauseboth descriptionscite the singlecollection ol J. Drummondcotl. 5, n.426.However, the description given by Hooker matchesthe accompanyingillustration ol cultivatedmaterial rather than the Drummond specimencited. No herbariumspecimen derived from the cultivatedmaterial has been located. sothe illustrationis takento be the type.Hooker's description and illustrationresemble P floribunda very well exceptperhaps in the descriptionof the flower colour as a very pale rose.Certainly the description and illustration do not matchP. drummondii.particularly in regardto the shapeand orientationof tbe leaves.Pimeleo macrocep,4ala is regardedhere as a nomendubium. u^\Ero(0rx) rr€^o s :(0r x) uouels.o :0x; ,"^ou.3,r""1,g€p.*J1!#,i?"{"llfiyg1j7;}ffi,.['l|}1,ffi]

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Sect.5. Macrost€gia

Pimeleasect. Macrostegia (Turcz.) Rye, comb. nov. - MacrostegiaTurcz., Bull. Soc.Imp. NaturalistesMoscou 2512: 1,11 (18521. Type: M. erubescensTurcz. (: P. physodesHook.)

S,&rubs,hermaphrodite or gynodioecious.S/ezr glabrousexcept for inconspicuoushairs in upperleaf axils;nodes abaxially prominent, c. twiceas thick asbase of leaf.Ieaves opposite, sessileor subsessile,glabrous. Involucral bracts highly differentiatedfrom leaves,closely surroundingflowers, not becomingreflexed, very large,colourful, hiding flowerslaterally, glabrous.Inflorescence head like, terminal,pendulous, compact, many flowered;receptacle well developed.Pedlcels with denseantrorse to appressedhairs. F/owels hairy outside,glabrous nside.Floral rubecircumscissile above ovary, prominently constricted at circumscissionpoint; proximal portion (belowcircumscission point) extendedwell aboveovary, with deciduous hairs;distal portion cylindric,longer or sometimesas long asportion belowconstriction and often broader,broad throughout rather than only towardssummit. Sepals erect, elongate. Stamens2,very Iong;connective narrower than anther;slits lateral after dehiscence.Ouary glabrous.Sry/e filiform, verylong; stigma small, papillose. Fnrlt dry, enclosedin the p€rsistent enlargedbase of the floral tube. A sectionof I species,occurring in south westemWestern Australia. Noles.This sectionis undoubtedlyclosest to sect.Calyptrostegiabvlhas a numberof unique charactem.It differs from all other sectionsprimarily in the shapeof the floral tube and sepals, in the sepalsbeing both erectand free and in the extremelylong stamensand style. The involucralbracis are the largestand most showy in the genus.The singlemember of sect. Macrostegia,P. physodes,is probablythe only bird'pollinatedspecies in the genusand this may accountfor its very distinctivefloral features. 30.Pimelea physodes Hook., Icon. Pl. 9: t. 865 (1851).- Banksiaphysodes (Hook )Kuntze, Revis.Gen.-Pi. 2: 583 (1891).fype: south'westernAustralia, 1848,J. Drummond cnll. 5, n. 424 (lectohere designated: K; isolecto:KW, n.v.);south westernAustralia, "/ Drummond suppl.n. 84 (syn:K; isosyn:MEL). Macrostesiaerubescens Turcz.. Bull. Soc.Imp. NaturalistesMoscou 2512: 171'\78 (18521. Type:soulh-western Australia, 1848,J. Drummond coll. 5, n 424 (holo:KW, n.v., photo in PERTHI iso: K). Shrub,neaiy alwayserect, spindly or open,0.2-1 mm high,single'stemmed at ground level. Stemsyellowish or deepred to purplenear each inflorescence, becoming medium grey-brown further from apex.Leaves opposite, antrorse, concolorous, medium green, ovate to elliptic or narrowly so, 18)12-32x (3-)5'11mm, flat or concaveon adaxialsurface, acute to obtuse. Peduncle3-14 mmlong. Involucral bracrs 6, 8 or rarely4, sometimesalmost completely pale greento cream,more commonly with reddishportions or completelyred to purple,the outer bractsusually with the mostred to purplecolouring, elliptic to broadlyelliptic,22'60 x 1l'45 mm, glabrots.Pedicels usually 0.7-l mm long; hairs l'2.5 mmlong. Flowers bisexualor rarely female,green or creamygreen with a reddishstyle; tube 6-9 mm long, circumscissileand stronglyconstricted c. 3 mm abovebase and c. 1 mm aboveovary. Poximal portion ofJloral t re (belowconstriction) usually 3-4 x c. 1.5mm, with deciduousantrorse hairs l 5-2.5mm F long,glabrous in fruit. Distalportion offloral tube (abovecnnstriction) cylindric, 3-5.5 x 1.5'2 6 mm, with very fine antrorseto patent hairs mostly 0.5-1'5mm long, the longesthairs up io 3 mm long. Sepalsvery narrowly triangular,5 9 mm long, with hairssimilar to thoseon distal portion of floral tube but sometimesless densely hairy. Stamensgteatly exceedingsepals; ol filament 11.5-16mm long;anther 2 3 x0.2'0.3 mm.Staminodes of femaleflowers: filament iF c. 8 mm long;anther c. I mm long.Style exserted by 18-23mm. Seed c.6 x 1.5mm, with (F longitudinalrows of shallowpits. (Figure46.) 6: 'uorEg l do e'S'&ror4tsuo^eIJo ldt\I]!{ ru\02'x i\-Igl/ot6Z ZooIJ UAIlsetd :(TAW) I99 a8toag'g'1 'uou?fltt{ lsBg:(HJUAd) aznag )podqumqJ :(HIUAd) Bll L S't,'eEu?X're^ru 9II9t )aupn) ',, lroqs lW :(HJUAd)LI6I rauptog'tz, sdn[qd :(HJdAd) aluD(L 'edroqlsuo^eu rul :((Jv) ulisnv 't'Q 'edroqlsue^?U tZ 't Z Jo 1Y\N 09 ,61 Jo lA rxl 8 :(uees9S ro^o uro4 palcelos) VI-IYUJSnV NUIIJSA/\(\paaluDxa suauoads

(v\ 'l'g) 'S . ntg6 raqkaN ) Iq peluasorderl?pat€(u qse{ pur (C €900t aa./oa9 y ruorJu^ApJC lZ I x) ^rp^o O :(g y) u.ulets -J :(g x) re^\olJ g :ruols8uua,\\olj "V sap.ntttl.lo4atuH gl eJntul :J3

'rtrd 'I tza eBecBaBlour,{qJuerlertsnv uralsaA B (1988) Z)1 Ntt))tia Val. 6, No. 2 887(PERTH)I Jerramungup, F. Lulljitz 35ll (PERTH);8 mi Il3 kml W ol PabelupLake' K. Newbet+S t (peRf H): No Tree Hill, I Nov l962' M.E Phillips(CBG); NW cornerof FitzgeraidRiver NationalPark, R.,4. Salfrey 1442 (PERTH); 39kmSof Ravensthorpealong roaJto HamersleyRiver estuary, I.R. Telford8621 (CBG): East Mt Barren'28 Oct l968' J.W. Wrigley (BRl. CBG).

Distibution.(Figure 47.) Extends from nearLake Pallarup (c. 33015'S, 119043' E) sout! t9 FitzgeraldRiverl'{ational Park i34004'S, I19026'E) and from Jerramungup(33056'S. I 18055' E) eastto Desmond(33'38'S. 120008'E).

Habitat. Occursin sand,often with gravelor rocks,on plainsor hillsides.

Flowering period. July'October.

AlJinities. Pimeleaphysodes is similar in vegetativecharacters to some specimensof P dilmmondii. Although verydistinctive in its floral morphology,P. physodesis clearlyclosest to P. drummondii.Like P. drummondil,it losesthe hairsof the proximalportion of the floral tubevery readilyand is glabrousoutside in lruit 'bells' No/es.The bell,likeinflorescen ce ol P. physodesbean a striking resemblanceto the of a numberol speciesoccurring mainly in the StirlingRange' particularly to D macrostegia(Turcz.) Benth. The parallelevolution in the two generaof pendulousflower headswiih largecolourlul bractshiding rhe elongateflowers from side view is apparently relatedto the alaptationof the speciesconcerned to bird pollination.Keighery (1975) recorded Phytidonyris."loropt. the Tawny-crownedHoneyeater, as a probablepollinator of P physodes.

Sect.6. Stipostachys

Stipostach),sRye, sect.nov.

caulespallidi, glabril nodi abaxialiternon ulira petiolumprotrudentes. Involucri bracteae 4-12,saepe deciduae. Inflorescentia maturitate elongata, contigua; pedicelli numerosi, in serietuspluribus longitudinalibus dispositi. Flores bisexuales. Tubus floralisextus tenuitel patenti-pilosus;portio stylarisquam ovarialis longior. Ovarium apice pilosum. Fructus siccus, basi tubi florali persistenteinclusum.

Typus;Pimelea haemostachya F. Muell.

Shrubsor partially herbaceousperenmals, hermaphrodite Stezs pale'coloured'glabrous; nodesnot protrudingabaxially beyond the petiole.leaues usually-opposite and decussate, shonlyFtiolate, gJabiousllamina concolorous. IDofu cral bracts4-12, often deciduous, sessile, paperyand pale-colouredat maturity, broaderthan leaves lnflo.rescenc€terminal, erect' compict at fint, elongateat maturity, continuous.Rachis and pedi.ce.ls with short hairsand long antrorseto pat.nt haitr, the pedicelsnumelous and in many longitudinalrows. Floral nb\ with fine paient hairsoutside and olten inside,cjrcumscissile or {in WesternAustralia) fully persistentbut sometimesiregularly tearingabove ovary;style portion much longerthan ovaiy portion.,97,o/.s spreading, hairy outside,glabrous inside. Stamens 2; connectivenarower tha|ranther;slits semi-lateral after dehiscence.Ovary hairy at apex.S/y/e exselted. filifolm; stigmasmall, papillose.Frulr dry. enclosedin penistentbase of floral tube'

A sectionof 3 species,oi rvhich I occursmainly in the Pilbara,western Australia and 2 occurin Queensland. 'acusJsologur ole8uoJasnonulluoc e s?qU sopoulueuuo:d-uou Sul,lequt lensnunsI qclqi 'uouoas lcurtsp ellnb B urroJsenods eserll'DnJap A qll.^AJoqlaSoJ seltods o,1Al eql ueo,\\loq drqsuorleloJelqrssod s palou soDodsad^l 3q1loJ ldoxxeaSo odg lJos tnq \stq)o$ttoq) Jo 'l.esqns Jopunpapnl3ur or3q oft sdnol8 qloq) fle^Ipodsel aSollDdg laes pug stllJols!rcqJ 'lxes 'd 'D4aruld o8a$oJidKlD) olur tlptotloq pre DtqJDtsowdrryd pelElEdesvJetllueg Jo suorlecrJrss?lccrJaueS€{ur 3uhr3 suouecllqnd lerlJeo eql ol .toudpoquJsop ueeq peq ouou lnq {€/81)tuerut?arl s,u?qlueg ot Joudpoqursop ote,,\{. s,{{rrlsodgs Itos Jo solcodso,,r\l saloN '| llptotlotl d pue a sapos[qdoalau4d Jo uor]nqulsrc li aJniIC - '1i+ Tf P* {s*rl : I I 1\ i-fil I 1 I lf I rfillf r+t1 l liiri lll;r

'+1tril-l t- vrvulsnvNUlsrM -' J l I1lI|rLli ctoc?oEleLu^qIupllErtsnV urotsa/t 'rtrd 'l g 226 NuytsiaVot. 6. No. 2 {1988) which can extendto over 200 mm long but remainsdense, the flowersin many longitudinal rows.The upperfloral tubeseems to be odd in that it tends,at leastwhen dried. to be square in cross-sectionbut this characterneeds to be examinedfurther. Its closestaffinities are with secl. Calyptrostegiawhile it also shows some similarity lo sect.Heterohena.

31. Pimeleaholroydii F. Muell.,Fragm. 6: 159-t60,t. 59 (1868).- Banksiaholroydii (F. Muell.)Kuntze, Revis.Gen. Pl. 2: 583 (\891). Type:Hamersley Range, Western Australia, date unknown, C. Harper 16 (holo: MEL).

Shrub,erect,0.3'l m high,single-stemmed at groundlevel. Srems very paleyellow-brown in the distal 100-300mm, becomingred-brown further from apex.Leaves opposite or subopposite;petiole 0.5-2 mm long;lamina more or lessmedium green or bluishgreen, often with blackpatches when dded, ovate or broadlyovate, 12-31 x 6-20mm, flat,obtuse; midvein and severallateral veins pale yellow-brownin the proximal half of the lamina,becoming indistinctabove, prominent on abaxialsudace. Peduncle :ust:,ally 5-30 mm long.Involucral bracts4-T,thesame colour as leaves at first but becomingpale yellow-brown in fruit, broadly ovateor rarelyovate, 9-20 x 7-16mm, glabrousoutside, usually ciliate especially around apex, hairy throughoutthe inner surfaceor with hairs concentratedin the centre-base,dry and chartaceousin fruit, persistentbut with portionssometimes breaking off; cilia I -4 mm long, very fine, siIky.Inflorescence initially compactand head-likebut elongatingas flowering proceeds;rachis narrowly conic and usually 13-30mm long at maturity. Pedicels0.5-l mm long,with mixedlong and shorthairs, the long hairs 1.5-3.5mm long,the shorthairs 0.2-0.5 mm long. Flowers white or sometimescream, not circumscissilebut sometimesbreaking irregularlyabove the ovary, glabrousinside; floral tube 10-12mm long. Ovary-portionof floral tube2-3 x 0.7 I mm, very denselyhairy; hairsantrorse, 0.5- 1.5 mmlong.Style-portion offloraltube 8 9 x c. 0.7 mm, with a mixtureof long hairs 2-2.5mm long and minute haim 0.20.3 mm longin proximalhalf, shortly hairy above.Sepals elliptic, 2.5-3 x l-1.7mtn. Stamens slightly shorterthan to slightly exceedingsepals; filament 2-2.5mm long; anther 1.2-1.5x 0.40.7 mm. Ovaryc.2 mm long;hairs l-1.3 mm long.,S/y/e exserted by 1.5-4mm. Seed 4-4.5x 1.5-1.8mm, with longitudinalrows of shallowpits. (Figure 48.)

Specimensexamined. WESTERN AUSTRALIA (selectedfrom over 15seen): Wanna Station. J.S.Beard 6084(NSW, PERTH); 2), mi[34 km] E of HamersleyStation, ]/. Demarz 2851 (PERTH); l0 km S of HamersleyHomestead, 1{ Demarz't 102(CANB, PERTH); between Port Hedlandand FortescueRiver, Oct. 1946,C.M. Donald (CANB, MEL); 2 km SE of Dingo Bore,Mt JamesStation, Zt. Sexer 378(ADW); 5 km E of Juna DownsHomestead. M.E. Trudgen 358 & G. Marton (PERTHI.

Disfiibution. (Figure47.) Extendsfrom HamersleyStation (c. 22015'S, I17050,E) eastto Yuna Downs(c. 22051'S, I18032'E) and south-westto WannaStation (23055' S. 116033' E) andMt JamesStation (24051'S, ll70l3'E).

Habitat. Recnrdedin red clayey soil.

Floweringperiod. RecordedJanuary-February and August,October.

Af|inities. A very distinct species,closest to the two other membersof sect.Stipostachrs, Pimelea haemostachyaand, P- decora, both of which occur in Queensland. '{r0^0IAl J T roqrsrC \ielpur-I) slttqopads ll s?) ^epur.I sllqopads .d :adq '\9n8il .UoH .rxos ,ro. Lr:01 dorted xapul aW J ? loqcsrCDuaDlolapH '(pel?u8lsep .d .\e061 .ua0 .x{ oroqolrol) rg U ,asor :adKJ .ezlun) .J ur .ttes69.u€qd d, }sod (DuaDpoDpHs€) aztun) ( Ipu1l DuaDpnjaq nsyuog _ ,(f .ld .iraC;.pud .ur8?rc,.Ionw .l SSt) tgS DuaqonlaH'qoaputu - (gggI)691 :g (IpugljDuaopralaH .ws Dapw] 'lres suaqoraleH /

'J 's 'vl . (J Jl IsSzztDua.l H pue{O Izgl pbag s l uro{u^{srcl (9 1 x)ocuecsorollur Suun{ -l :lat xl \rE^o .:l :(8 x) ururets C :(SI x) rJ^\olJ,f :1€x) Delq ,g :uloNSurle^\ou -y tlptotloq oapald .9, err.?tf

Laa leoceoeleul[ql uelJellsnvur0tsey' 'e,{d 'I g NuytsiaVol. 6. No. 2 {1988) Shrubsor undershrubs,hermaphrodite or gynodioecious,rarely someindividuals with bisexualand female flowers. Slezs glabrousexcept lor inconspicuoushairs in upperleaf axils or rarely(P. ferruginea) inconspicuously hairy when young; nodesabaxially prominent, c. twiceas thick aspetiole. Zeaves opposite, decussate, glabrousi petiole (when present) usually brownor brownish;Iamina often discolorots. Involucral bracts in l-3 (usually2) pairs,sessile, often with subsessilebract like leavesbelow, erect and closelysurrounding flowers, not becomingreflexed in fruit, broaderand usually more colourful than leaves,glabrous or subglabrousoutside, often ciliate laterally but not (exceptin P. brevistyla\around apex. Inflorescenceterminal, head-like, many'flowered; receptacle very compact,ovoid to more or lessflat. Pedicelswilh denseantrorse to appressedhairs. Flowershairy outside.F/oral tube wholly penistentin fruit or rarelycircumscissile above ovary, nearly always with 2 or moredistinctly different kinds of hairs,usually with a subbasalto medialband of longpatent hairs;ovary'portion usually prominently 8-dbbed; style-portion longer than ovary-portion, broadestat summit,the gradualexpansion occuning mainly in the distalhalf. Sepals spreading, p€rsistentor shedwith distalpart of floral t|be. Stamens2; cellsusually protruding laterally beyondconnective and with slitssemi-lateral to lateralafter dehiscence,Iarely \P. brevistrla) laterallyexceeded by connectiveand strictly introne. Ovary glabrous.Stigma small, papillose. Fruit dry, enclosedin the persistentenlarged base of floral tube, apparentlytending to be shedbefore seed matures.

A sectionof 14species, endemic in south-westemWestern Australia. It is closestto sect. Calyptrostegiabut also showssome similarities with sect.Pimelea and secl.Stipostachys.

No/es.The nameHeterolaena is derivedfrom the Greek wordshetero (different, unequal, variable)and chlaena (cloak), perhaps refening to the variablesize of the hairson the floral tube.When this sectionwas first named,a heterogenousassemblage of specieswas included, mostof which werelater referredto secl.Calyptrostegia. The lectotypechosen here for the groupis a speciesincluded by all subsequentauthors who havelisted species for this section. The genusHeterolaena was publishedafter the name Heterolaenahad been usedas an unspcifiedinfrageneric category but withoutany referenceto the previoususage of the name and without citing any of the speciesincluded in the earlierpublication.

As circumscribedhere, sect. Heterolaena is a relativelyuniform group.It is characterised by the compactflower headswith well definedinvolucral bracts, by the elongatefloral tube, usuallywith longand shorthairs, and by the usualabsence of circumscission.Its closestties are with sect.Calyptrostegia.

32. Pimelealehmanniana Meissner in Lehm.,Pl. Preiss.1: 603 (1845).- Calyptrostegio lehmanniana(Meissner) Endl., Gen. Pl. Suppl.4: 61 11848).- Banksialehmanniana (Meissner) "Mt Kuntze,Revis. Gen. Pl. 2: 583 (1891).Type: Wuljenup"[Willyung Hill], Western Australia,l4 Oct. 1840,L. heiss l27l {lectohere designated: LD; isolecto:MEL, NY).

Shrub,erect".0.3-1.2 m high, singlestemmed at ground level.Srens usuallydark reddish brownnear each inflorescence, dark grey-brownfurther from apx. laaves anlrorseor patent; petioleusualy 0.5-1.3 mm long; lamina discolorous,dull yellowishgreen to grey-greenon abaxialsurface, darker bluishgreen with a sheenon adaxialsurface, narrowly obovateto narrowlyovate in mostleaves but ovatein a few leavesimmediately below the inflorescence, l4)ll 26(34\ x l-7 mm, up to 8 mm widein uppermostovate leaves, flat or slightlyrecurved at apex,usually acute and/or mucronulate. Peduncle 2-17 mm long. Involucral bracts4 or 6, commonlypale yellow-green, sometimes reddish, rarely similarin colourto leaves,ovate or broadlyovate, 13-21x'7 -14mm; outer bractsoften partially hairy insidebut with fewer haimthan on innerbncts, not ciliateor rarelywith very few cilia;inner bractspartially hairy inside,sometimes ciliate, the longestcilia 1.33 mm long.Inflorescence erect or pendulous. '91g6 taq^aaN) ,{q poluesotdallpullsu qs).rJLuorJ u. erc (0t x) tr^rolJ U€d laddn -O :{g x) -J -B }o ra,^4ou :(t x) lcprq Jo ocelns rouur :ruJN 3utano11 y ouotuiowqal-dsrios ouotuuiuilal oa1aw146p-a];iayj

gl=

aeaae$leur{qJue{eJNnV uJotsoy' 'a^U "l'g 230 NuytsiaVol. 6, No. 2 (1988) Pedicelstsual\y I -2 mm long; hairs up to 3 mm long. F/owers bisexualor rarely female,white to paleyellow, sometimes pinkish in bud,not circumscissileor circumscissile2,3 mm above ovary portion, hairy insideat the throat and sometimeswith a few hairson baseof seoals but otherwiseglabrous inside, the longesthairs in throat reflexedand c. 1.5mm long; tube 8'14 mm long. Ovary-portion offloral tube 2 3 x c. 1.2mm, glabrousor with patent to antrorse hairs0.1-0.5 mm 1ong.Style-portion offloral tube 6-ll mm long, 1-1.5 mm diam.at summit, with long patentto widely antrorsehairs 4-6 mm long, but no short hairs,in the proximal half and antrorsehairs 0.5-1.5 mm long above.Sepals ovate or rarely narrowlyovate, 3-5.5 mm long,with a few antrorsehairs, the longesthairs 2 3 mmlong.Stamens greatly exceeding sepals;filament 5-7mm long;anther 0.7-1.4x 0.25-0.6mm; slitssemi-lateral to lateralafter dehiscence.S/y/e exserted by 6-10 mm. (Figures49, 50.)

Distribution. (Figure55.) Extends from the Darling Rangenear Perth to FitzgeraldRiver National Park.

Flowering period. August-September.

Afiinities. Closest to Pimelea spectabilis and P. rara.

Notes.The two subspeciesrecognised here are very distinct and could be regarded as separate species.Treating them as subspeciesresults in Pimeba lehmanniana being perhapsthe only speciesof the genusto have both early-circumscissileand non-circumscissilemembers. Subspeciesrank waschosen for the two taxabecause the morphologicaldifferences between them are lessthan thoseseparating all the other paiN of the taxa recognisedhere as species in sect.Heterolaena. Since they do not occur sympatdcally,it is not possibleto deteimine whetherthey arecapable of hybridisingin nature.Further studies,examining other aspects of their biology, suchas anatomy,biochemistry and cytology, might demonstratebetter which taxonomicrank is most appropriate.

Key to Subspecies

l. Floral tubecircumscissile; ovary-portion hairy...... 32a. subsp. lehmanniana 1. Floral tubenot circumscissile;ovary,portion glabrous...... 32b. subsp. nervosa

32a.subsp. lehmanniana

Shrub, 0.3-1.2m high. Leavesusually slightly recurved.Peduncle up to 10 mm long. Involucral bracts4 or very rarely6 but sometimeswith a pair of subsessilebract,like leaves immediatelybelow, ovate or broadlyov ate.InJlorescenrc ffndulous. F/owersbisexual or rarely female,white to pale yellow, circumscissile2 3 mm aboveovary-poftion of flo ral tube. Ovary- portion offlorulture with patentto antrorsehairs 0.1-0.5 mmlong. Anthers 0.7-1.2x 0.3-0.6 mm. (Figure49.)

Specimensemmined. WESTERN AUSTRALIA (selectedfrom over55 seen):Stirling Range, A.M. Ashby 2016 (AD, PERTH); betweenBremer Bay and Gordon Inlet, M.G. Corrick 7691(MEL); Middle Mt Banen,C.,4. Gardner 9159 (PERTH); West Mt Barren,A.S. Georse 6957 (PERTH);Jerramungup, F. LullJitz 3517 (PERTH);E .nd of PorongurupRange, k Newbey 1843 (PERTH); Mt Drummond,K. Newbey 2690 (PERTH); 7 km E of Mount Barker,P Wilson 3355 (AD); 2 mi [3 km] from Albany towardsDenmark, 12 Oct. 1968, J.W. Wrigley (CBG).

Distribution.(Figure 55.) Extends from LakeMuir (34028'S. I 16039'E) eastro EastMt Baren (33055'S, 120001'E). Fi 9) 8-I988! 'I00t8 anu Iq p4u0s dar qsorJurorJ u^\pJO (B x) uouets :(6 7B 'osotuaulErroteu O xJ ra,rol] J :(f x) lrerq Jo aJefns r3uur 'g :uais Suuel\o[I-y dsqnsouotuuDwqal Da]at ld, 0S arn3rl

'uoxet oql aceld,{lolturJop 01 olBnbep?ursr uotdursop oq} pu? ocuelsueur lrls suourrcodsodfl ,{ueeq ol J?oddelou op aleql osnecequnrqnp ue{xou? s3 pelsrTsr lnq .rc^ solcadsqnssrql Jo ru,{uou,4s ?,{lqsqoJd sr slarqDpqda7oaut ouDluuDwqalDalauld .sarcN 'raqu0^oN-lsn8ny .poltad Satauolg 'sllrq '[oA€13 .n1!qDH Jo se8u?Jur uouo Jo/pu?pu?s u pep]ocad '0,{U rta sgrcBtDlou^qJuerlgrsnv urolseid I g 232 NuytsiaVol. 6, No. 2 (1988) 32b.subsp. nervosa (Meissner) Rye, comb.et stat nov. (Figure 50.)

Pimelealehmanniana var. ne^)osa Meissner in Lehm.,PI. Preiss2: 270(1848).fyp€. south westernAustralia, 1843-1844, J. Drummondcoll. 3, n. 284ex parte(iso: NY).

S,4rub,usually 0.3 0.6 m high. Lea\)esflat or nearly so. Peduncleup to 17 mm long. Involucralbracts 4 or 6,ovate.Inflorescence usually erect, rarely pendulous. F/owers bisexual, white but often pinkish in bud, not circumscissile.Ovary'portion of Jlorul tube glallrous. Anthers| 1.4x 0.25-0.4mm. Specimensexamined. WESTERN AUSTRALIA (selectedfrom over 25 seen):Gleneagle- Jirrahdaleroad, T.E.H. Aplin 1222(PERTH); Preston River, 1896,G.H. Berthoud\MEL); Capel'Donneybrookroad, R.l Cranfield908 (PERTH);between Marradong and Pinjarra, l8 Nov. 1904,A. Morrison (PERTH);Gooseberry Hill, 22 Oct. 1970,A Monison (BRI. CANB); BlackwoodRiver, t 883,Mcllard (MEL); 2 mi [3 km] E of Dwellingap,K. Newbev 2627 (PERTH)I 3 mi [5 km] from Darkan towardsCollie, M'E. Phillips 684003(CBG, PERTH): Jarrahdale,Nov. 1916,Sroward (PERTH).

Distribution.(Figure 55.) Extends from GooseberryHill (3105'S, I1600'E) southto near Donneybrook133010' S, I l50l0' E).

Habitat. Occtrson the DarlingScarp and Range and in nearbyhilly areas,recorded in gravel.

Floweringperiod. SeptemberNovember.

33. Pimeleasessilis Rye, sp. nov. (Figure51.) ellipticis AflinrsP. IehmanniaLiaeMeissner sed differt foliis arcte sessilibus, involucri bracteis ad subcircularibuset tubo florali fauceintus nudo Natiglal Typus:0.'l km S of Kalbarri - Ajana"tss road on track 1o Meanarrafiill' Kalbani i'ii['w.rtii" e"itiaiia,za sept. 5,N. Hovte527 (holo: PERTH; iso: CANB' MEL)' bloadly Relatedto P. lehmannianaMeissner but differs in the closelysessile leaves, the throat of the aripii. to urLort.ircular involucralbracrs and the absenceof hairsinside the floral tube. Sftrub,erect, usually 0.15-0.4 m high,single stemmed at groundlevel. Sterrs deep red'brown or yellowishnear each inflorescence, becoming dark brownfurthe,r from apex.Ieaues usually paient,sessile, elliptic to almostcircular, almost concolorous to definitelydiscolorous, medium greenio dark bluishgreen on adaxialsurface, paler on abaxialsurface' variable in size,the iargerleaves 8-15 x 6-9 mm and the smallerleaves 2 8 x 1-6mm, somewhat2-lobed and stem-claspingat base,the marginrecwved, broadly obtuse but mucronulateat extremeapex' peduncteusially 5-25mm long.Involucral bracts 4, greenwith reddishor yellowishareas, broadly elliptic to almost circular, 7'15 x 6-14 mm' appressed-hairyinside' not ciliate' In/toriscenie erect.Pedicels 1.5-2 mm long; hairs up to 2.5 mm long Flowersbisexual or ra'relyfemale, white or cream,not circumscissile,glabrous inside; tube 8-1I mm long' Ouary- J 2 0 4 wrtion of florat tube c. 2 x 0.1 mm, usuallywith reflexed-to-relrotse minute hairs 0 (l mm longand sparselong hairs.Sty/e-portion ofJloral tube6.5-8.5 mm long,c. 0.7 mm diam I at rumriit, with long palent hairs 2 3.5 mm long but no small hairsin proximalhalf or for ll mostof styleportion, with antrorsehairs c. 1 mm longin distalhalf or only at summit-Sepals narrowly ovaie, 3-5 mm long, with hain l'3 mm long. Stamensexceeding sepals; filament 0. 2.5-4mm longt anther 0.7-1x c. 0.4 mm; slits semilateralafter dehiscenceSty/e exserted by 3-4.5mm. P; 'ro^ru '^,,i{H .y ,Iled uosrqrrnw Jo N [rDI gt] ru gZ le}s?o:) :HIUAd) gl9 tJDltoD 'l[H IAN 'g 1 I?uorleNrrr?qp) ?rJ?ueol{ol :(HJXgd) DlrDtg 1gDup.tDg .v .) '.sur?ld JJournJ 'pU 60L 14 e,rellntrtr''(HJUEd) 0gIl DtloI y U Bu?fv uo r EqlB)Ir[o{ lurl g.0] g.0 .yJ ,seuuensejIu :GJXAd) 191q3no2 T U'ro^rd uosrqrrnN :(DflJ) 0 tg9 3u1uun1 g .Vg6I oqJ prB,rol ppor e^\ellnl^{-uolpler00Jo S lur)t 6l 1rn/.S :GJUiId) talJng H.l4 'l3O '?lleg '.(HIdAdl 'surpld .g,EB6l IIDpolg g ,A'I%I ldos ?,^ro[nW:(HJUgd) srir ag '3nv .W.y ,Zt IZ'ureqle)IuorJ g'0]rur .:(yAn)prrryf :(HJdAd) 'punos[rU)I Z/t 6I,e{oou€pu?I 928 sMaquv J e8rooC EurX :VI.IVNJSOY NXAJSAIA paulwDxasuawoads

1.q86_l3rV lZ u-"qlp) )"'tu,rtlDlDg g \q plua.Jrderlerrlpld q.Jrt uror1urrerC .r9l yl urru?r..1:t0l\)rrttou.. .. ^\ol:quollJlurr.Jrol1Ur C .;:rJi,.ire:l g:ru:r,Bur.jmo1 y ,tlt,,a,oaiaw4 ig atia\

rtz aeaaeaelour{qfusrlBrtsnv uretso/A 'a,{d .I g 234 Nuvtsiavol.6, No.2 (1988) A. S. George1503 (PERTH); NW CoastalHwy, 9 mi [4.5 km] N of MurchisonRiver, .,4.S George7878 (PERTH);4 mi [6.5 km] SE of TamalaStation,,4.,S. George 9573 (PERTH); NW CoastalHwy, c. 42 km N of MurchisonRiver, EN.S. Jackson3727 (AD); c. 20 km by road N of Whelarra,E.N.S. Jackson3168 (AD); Shark Bay, 1877,F. Mueller (MEL); MurchisonRiver Gorge,27 Sept. 1962,M.E. Phillips ((36L 13 mi [2] km] by road SE of Kalbarri, R. Z Smith 661365(MEL); MurchisonRiver, 6 Sept. 1949,N.H. Specft(UWA); Hawks Head Lookout, Kalbarri National Park, A.S. Weston6922 (PERTH).

Distribution. (Figure55.) Extendsfrom TamalaStation (26044' S, 113043'E) south-eastto Yandanooka{29019' S. I15"34'E).

Habitat. Occursin sand in shrublands.

F lowering period. August-October

Derivation of name. Sessilis(L.) - sessile,referring to the leaves.

Affinities. Closestto Pimelealehmanniana, also similar to P. Ieucantha.

Notes. Of 11 collectionsexamined in PERTH. one wasfrom a femaleDlant and the remainder from bisexualplants.

34. Pimelearara Rye, Nuytsia 5: 9-10 (1984).- Pimelealehmanniana var. Iigustrinoides Benth.,Fl. Austral. 6: 9 (18131.Type: SwanRiver (WesternAustralia), date unknown,I Drummond, coll. I (lecto,fide Rye loc. cit.: K; isolecto:K, NY).

Shrub,erccI or decumbent,0.2-0.35 m high,single- or multi-stemmedat gound level,with few branchesabove. S/ems somewhat reddish near each inflorescence, becoming dark brown to black further from apex.kaves antrone to patent;petiole c. I mm long; lamina concolorous, dull bluishgreen, elliptic or obovate,l5'30 x c. 8 mm, flat, obtuseor almostacute. Peduncle 2-20mm long.Int)olucrul bracts 4, broadlyovate, 9-20 x c. l0 mm; outer bractsglabrous; innerbracts sparsely hairy inside,otlen with a few cilia,the cilia 0.5-1mmlong. Inflorescence erecI.Pedicels c. 0.6 mm long, with hairs 1-2 mm long. Flowers bisexual,white, not circumscissile,glabrous inside or with a few hairsc. 0.5 mm long in a ring at throat of tube; tube c. 9 mm lon9. Ovary-portionof lloral tube c. 3 x I mm, coveredby reflexedhairs c. 0.5 mm long. Style-portionof Jloral tubec.6 mm long,c. 1 mm diam. at summit,reflexed hairy at base,with a band of patent hairs 2.5-3mm long, usuallymixed with short hairs similarto thoseon the ovary-portionor longerand morespreading and often with antrorse hairs c. I mm long at summit.Sepals ovate,3-4 mm long, the outsidesparsely covered by hairs0.5- I mm long,glabrous inside. Stamens greatly exceeding sepals; filament 4-5 mm long; anther I .2-l.5 x c. 0.5mm;slits semi-lateralafter dehiscence.Sty/e exerted by up to 7 mm. (Figure- Rye 1984:9.) "Smith's Specirnensexamined. WESTERN AUSTRALIA: betweenParkerville and Mill lGlen Forrestl,Jan. 1904, W.V. Fitzgerald(NSW); Parkerville, W.A. Ross435 (PERTH).

Distribution.(Figure 56.) Extends from near Parkerville(3 1053' S. I 16008'E) south to near the Perth Observatory.

Habitat. Ocsrs on the Dading Range,recorded in lateritic soil in .

Floweringperiod. December-January. 101x)uoursls ,( :0x) ra,{orr .r :(, x) }cerq ro"""r.;919J"1:#?#fi,81iffi,:#il##:,#,H3J#?i:fi11S

'DtDuDl'd ,ouDlauDwqal ol osolc osl€ sdBqred oapuV ol tsosolc^lqegord .saau!l[{V r?ulse,roN'616r q apeu uooq auheq'"n'.?$T,,:1:1'l'"1,"t#1',i,rT;ii"jffifitJ3:3 puB .suoUJelloJ ellhle{l€d Jo ^lrurct^ aql uoJJ o,4\} ur,roul oelqi,tluo a:e,trs:aq1 ,{fuace: Flu{l poliulsal ^lecqdp$oo8 pue elEl,{lo^ eq o1s:uodde seroadi srq.L.sruojs uouottasuoJ .atrX .I eeocseelouIql u?rlellsnv ulalsoy'd B Vol 6' No 2 (1988) 2J6 Nuytsia Nores.The summerflowering time ol Pimelearara is unusualand may partially account for the small numberof collectionsof the spectes.

35. Pimeleaspectabilis Lindley, Sketch Veg. SwanR. 4l (1840).- Heterolaenaspectabilis (Lindley)Fischer & C. Meyer,Index Sem.Hort. Petrop.10: 48 (1845).- Banksiaspectabilis (Lindley)Kuntze, Revis. Gen. Pl. 2: 583 \1891\.Type: south westem Australia, 1839, I Drummond coll. I (lectohere designated: CGE); southwestern Australia, date unknown, J. Mangles(syn: CGE).

S,4rub,erect, usually 0.5-2 mm high,single stemmed at groundlevel. S/ems pale yellowish greenor brown neareach inflorescence, becoming dark red-brown to blackfurther from apex. fuavesantrorse to patent;petiole 0.5 L5 mm long;lamina discolorous, medium to dark green and often bluish on adaxialsurface, paler green on abaxialsurface, very narrowly elliptic, ll-42 x 2-7 mm or often broaderin leavesshortly below each inflorescence,flat, acute, mucronate.Peduncle 2 15 mm long. Involucral bracts4 or 6, greenand partially reddish or largelyreddish, ovate, 16-30 x 9-20mm, sometimespartially hairy inside;outermost bracts not ciliatel inner bracts sometimesciliate, the cilia 2'3 mm long. InJlorescenceerect or pendulous.Pedicels L5 2 mm long; hairs I 2( 4) mm long P/owersbisexual or very rarely female,white, pale pink or pale yellow, not circumscissile;tube 13-19mm long' glabrous inside.Ovary p:ortion of /loral tube 2'3.5x c. 0.7 mm, with minuteantrorse hairs c. 0.1 mm long.Style-portion offloral tube l0'15 mm long, l.l'1.5 mm diam.at summit,with a mixture of long'paienthairi 4-8mm longand minute hairs 0.1-0.2('0.3) mm longin proximalhalf and with antrorsehairs 0.5-l mm long in distal half. Sepalsnarrowly ovate or ovate,4-6 mm long, with short hairs and a few long hairs outside,the longesthairs 1.53 mm long, appressedhairy throughoutor at baseinside, the hain up to 1 mm long Sramerzsexceeding sepals,often greatly; filament 4-7 mm long;anther 1.2'1.5 x c. 0-2mmi slitssemi-lateral to lateral after dehiscence.Sry/e exserted by 7-10 mm. Bunjong. (Figure 52 )

Srycimensexamined WESTERN AUSTRALIA (selectedfrom over 90 seen):36 mi [57 km] N of Pembertonon road to Nannup, lC Anway 564 (AD, MEL, PERTH); Llbany' A. Ashby 23 (PERTH); near Augusta,A.M. Ashby 2695 (AD); SW of Shannonon road to Northcliffe, A.M. Ashby 3104 (AD, PERTH); near Mt Dale, M.G. Corrick 7838 (MEL); 34 mi peg 154.5kml, BrooktonHwy, H. Demarz 443 (PERTH);Notthcliffe, A.R. Fairall 859 (PERTH):Nornalup Inlet, C.A. Gardner 13026(PERTH); Thumb PeakRange,,4.S George7116 (PERTH);Shannon River, c. 25 km due E of Northcliffe, T.A. Hallidav 283 (AD, CANB, PERTH); East Mt Barren, T.B. Muir 4165 (MEL); Albanv Hwv, 35 mi [56 kml S of Perth,S Paust 1069(PERTH); 2 mi [3 km] E of CanningDam, 16 Oct. 1962,M Pftilrps(CBG)] Metricup, R.D Royce4685 ((PERTH); Fitzgerald River NationalPark, R D Royie 9259 (PERTH);upper Helena Vallev, I Seabrook398 (CANB, PERTH); between Cowaramupand MargaretRiver, D. J. E. Whibley 5045 (AD, PERTH); Denmark,C Z White 5365,(BRI); 4.5 mi [7 km] from Denmarktowards Mount Barker, 13 Oct. 1968,-/. Wrigley lAD, CBG).

Distribution.(Figure 57.) Extends from Mundaring(31054' S, 116010' E) andpossibly further north in the Darling Rangesouth to near Jarrahdale(c. 32050'S and I16013'E) and from CapeNaturaliste (33032'S, 115001'E) south-east to Albany(35002'S, I17053'E) Also occurs in FitzgeraldRiver National Park near Thumb Peak134'02'S, 119043'E) and on EastMt Barren(33055' S, t20001'E).

Habhat.In theDarlingRange and extreme south-west of WesternAustralia' recorded mainly in sandwith gravelor lateritic rocks,in Jarrahforest, sometimes with Marri or Karri trees as well as the Jarrah.In FitzgeraldRiver National Park, occurson the upperrocky slopes and summitsof hills. recordedin low shrublandwith outcroppingquartzite. 'l1'g) pup ,y) zlDwa1 rlorJ t6tn UtJilnTJ lJ tl6E'ut H u^\€.rc(/ x) ra^\ouC :(t x) tJerq]oacElns rouur J:(ix)J€01 B:etsts qse{ ur se se^Dol rll^\ 3tsSuuoMol] V DqluDrnaloaptudtgernaf

B

'eiu?J ssrcaoseql Jo seursrlxe0q1loJ polsrt solEts Jal3BJ?qJ 0r{1Jo suoneurquos ,\,\oqs seoJ? olerpouletururo4 suournodsluur0l-6 fq ^llensnpeuosxe el,(ts eqt pu8 sl?dasoqt Supoecxa '3uot 'el?qt3 fll?ar8 suetuEts ruur8-9 sJreq lsa8uol oql qtl,r sto/[oU,|aollo,{ ol?d ,{llBnsnslcplq Jouuroql qlr,{\socuorserouur snoFpu:d o^pq'lled tJ^rd pleJa8zlrdur fllercadso 'e8uer lpuotleN oql Jo tftd ulels?a,qlnosoql u.rolJsueursods turug,/ ,{qpoUosxa e1f,1s aq1 pue qedes aql Supaarxafluoqs sueruels'8uolruu 9-t ^lpnsn slleq lso8uoleql qtrr,rslo^\olj )lurdol altq,r'slc?rq oql uo erln tnoqll,{\ socue3soJolJur8up?aJds ol lcoJoe^eq ol pua} 'ol?pq?rJBt l?Jrnlo^ur 'o8ueJ ol 3uu?punl,^\trruo4 saroadsoql Joll"d uJequouoql tuo{ suaut3edssatoN 'DuDtuuDruqal D4autld ot $esol) sa!Iu!l[{V

'reque^oN'requeldas^lur?W po ad SuuaMoll 'a,{d 'I LE|, :e.cp0eleur^qlueqelsnv uraise/d g NuytsiaVol 6, No 2 (1988) 23S More study,including field work, is neededto determinewhether infraspecific taxa sholld t"i"*e.i;a ir ttris siecies.Two varietal namesgiven in the list of nomina dubia,P. s*ctabilis iir. ai{nrt gentn. aid P. spectabilisvat. rerschaffeltii (Morren) Meissner,may be applicable to the southem variant of P spectabilis.

36. Pimelealeucantha Diels in Diels & E. Pritzel, Bot. Jahrb.Svst. 35: 393 (1904\.Tvpe: GreenoughRiver, WesternAustralia, l0 Sept. 1901,L. Diels 4238(iso: PERTH). "near Pimelearosea var. calocephalaMeissner in Lehm.,Pl. Preiss.l: 602-603(18451. Type: Lake Keiermulu" lleedeiville area],Perth, Western Australia, 4 Oct. 1839,L. Preiss126'7 lecto here designated:LD; isolecto:MEL, NY).

Shrub, erect,0.4'2m high,single stemmed at groundlevel. S/ems pale brown or ted'brown neareach inflorescence, becoming dark grey to blackfurther from apex.Leaves antrorse to patenqpetiole 0.5-1.2 mm long;lamina slightly to distinctlydiscolorous' medium green then Lecomingbluish greenon adaxialsuriace, paler on abaxialsurface' narrowly ovate-elliptic or veryrirely ovaiebut oftenappearing linear when dried and laterally revolute '8'2'l x | 5'6.5 mm, with marginsrecurved at first and becoming(at leastwhen dried)revolute, obtuse or acuie.Peduncle l-6 mm long. Involuual brocts4 or rarely 6, mediumgreen or yellowish greenand commonlypink-tinged, ovate, L4'24x 6-t4 mm; outer bractsand medialbracts (whenpresent) glabrous or with fewer cilia than inner pair of bracts;inner bractsglabrous or subglabrousinside, sometimes ciliare, the cilia up to 4 mm long. Inllorescenceerecl to pendulous.Pedicels I'1.5 mm long; hairs up to 3 mm long. Flowersbisexua| pale yellow br cr"urn,not circumscissile,glabrous inside; tube 12 l8 mm long. Ovary'ponionof Jloral tube 2'2.5 x 0.7 I mm, with minute antrorseto patent hairs 0.1'0.3mm long, sometimes with a few long hairs at summit.Style'portion of floral tube 9-15mmlong, c. I mm diam. at summit,with a mixture of long patenthairs 3-4.5mm long and minute hairs 0.2 0.4mm long in proximalhalf, with shorterantrorse hairs mostly 0.5-1 mm long in distalpart.,Sepals ovaie oi narrowly ovate, 3 5.5 mm long,with sholt hairs and a few long hairs,the longest hairs I 2.5 mm long.,Stanens usually shorter than sepals;filament 1.5-2.5mm long;anther 0.9-l.3 x c. 0.3 mm; slits semilateralto lateralafter dehiscence.Sry/e exserted by 2.5-4mm (Figure53.)

Specimensexamined. WESTERN AUSTRALIA belectedflom ov,er35 seen):S of RedBluff, A.U. ,qtnOy 1825(AD); MurchisonHouse Station, J.S. Beard 6882(PERTH); Perrv Rd' E of Yanchep,J. Havel 94 (PERTH); 8 kmSof Eneabba,R. Hnatiuk 7'71518(PERTH): (CANB, LakeBanganup area , B.R.Maslin 1351(PERTH);Subiam J7-Ocl'1908'A. Mornson EDIN; 5;it8kml Wof Indarra,K. Newbev2162 (PERTH!-!:-*i-l? t kmlfrom Walkawav towardi Strawberri,15 Sept.1968, M.E. Phillips lAD, BP'I' CBG);Watheroo National Park' R.D. Royce 9569'(FERTil); KingswavRd, S of Wanneroo,H. Salasoo4129 (NSw)'

Distribution. (Figure56.) Exrends from Kalbarri National Park (27040'S, 114015' E) south to nearLake Banganup(32010' S, 115049'E).

Habitat. kt Kalbarri National Park, occursin sandwith exposedor outcroppingsandstone or limestone.Elsewhere generally recorded in deepsand

Floweringperiod. Mainly August October.

AfJinities. Closest to Pimelea spectabilis,P ciliata and P. avonensis. '9.tt2e .uorrrolor --J{ oauueu -r'.,,, adfi Fq peluosoJda.l qsa{ lloJJ .(01 ueruels-C xt romrlj :ip 9qr l€uJleultsrsaa u^iEto U 18 x, lciiqi:lur 1cr'aciryns _rauur tg :iits auurmod_v uo^DDpwH .t9 eJnztJ

B

'slueu?lrJ uelu?ls JeUoqsaql ur puB oplsur,{JrBq Auroq fluouruoc slc?Jq reuu oql u'str?Jq larcnloaurrotrno aleFc-uou oql u sleJJtplnqa^d D]D!I!Jii ol pol?lou s*ssqt,auuax.{x.€86r.rdosrr.qp4snvu,o'#3#.'#ry.'^;;3#J,fFr"..J}?3 'mqrJor^elqs4uerrrelJ urnururBls 'srtB![n le sBold sntrulenburnreld snquorJelur sreperq uou snqtlouetxasnqrlBlcnloaur srepeJq ueJJIp pos etd aD]olll2A slulJJV (.tS ernEld '^ou 'ds'efU slsuouo^?Balaurld .r€ o"a'9apl0u,{qJ uelellsnv ureNa^l 'e,{U .I g 240 NuYtsiaVol 6. No 2 (1988) Shrub, erect.usually 0.4 1 m high, single'stemmedat groundlevel. Stems red-brown near eachinfiorescence, beioming mediumgrey further from apex.Leaves patent oI sometimes antrorse;petiole 0.2-l mm long;lamina discolorous, medium to darkgreen and often becoming dark bluiih greenon adaxialsurface, paler green on abaxialsurface, naffowly ovate-elliptic or linear,5-2-4 x 0.5'3mm, with marginsrecurved and becoming (at least when dried) revolute' the apexinconspicuously mucronulate and fec'$ve.d.Peduncle 2-22 mm long.Involucral bracts 4 or 6, greenor'yellowiih greenand oftenred-tinged, 7'13 x 3.5-9mm; outer bractsnarrowly ouat. io ouate,glubrous; inner bracts (and medial bracts when present) ovaie or broadlyovate, often appressed-hairyinside, very rarely with a few cilia, the cilia up to 1.7 mm long' Infloreicenceerecl. Pedicels c. I mm long;hairs up to 1.5mm long F/owersbisexual, white ai maturity but often pink in bud, not circumscissile,glabrous inside: tube 8 12 mm long' Ovaryporiion ofJloral tube2'2.5 x c. 1.1mm. with shortpatent haim,0.2-0 4 mm long,often with a'few long fiairs towardssummit. Style'portion of floral tube 6 9 mm long, 0 7 I mm diam.at summit,with a mixtureof long patenthairs 1.5-2t3) mm longand shortpatent hairs0.1,0.3(-0.5) mm long in proximalpart or throughout,the distalpart oftenwith antrorse hairs0.5- I mm long.Sepals ovate, 2-4 mm long,with short hairsand a few longerhairs I l 5 mm long.Stamens shorier than to slightlyexceeding sepals;filament 1.5 2.5 mm long;anther 0.6-1 .1 i c. 0.2mm; slits semi lateral to lateralafter dehiscence. Style exserted by l 5 4 mm'

Specimensexamined WESTERN AUSTRALIA {selectedfrom over55 seen):Wilroy, '4 M (PERTH); Ashby 1806{AD, PERTH); betweenCoorow and Arrino, W.E. Blackall 2611 Tardun,26 eug. il+S, /a Cbbnd (AD:):Kalannie,CF. Daules282 (PERTH);Dalwallinu, A.L. F;iratl l04l (PERTH);Korrelocking, C.A. Gardner681(PERTH); Mt Singleton,C A' Gardner2211{PERTH); Wongan Hills, E.H. hing 83 (AD): Entranceto FowlersGully' WonganHills, K.F. Kennealty-1369 (MEL, PERTH); Cowcowing,M Koch ll'78 (MEL, PERTH); z mi t: t

Distibution.(Figure 57.) Extends ftom Wilroy (28038'S, I15038'EJ south-east to Moorine Rock(31019'S, il9o03'E). with isolatedrecords from near York (31'53'S,116046'E) and DongolockingReserve (34004' S, 117041'E)

Habitat. Recordedmainly in sandor sandysoils, oiten in open woodlandsor shrublands.

F lowering period. July'October.

Defivation of name.The specificepithet refers to the Avon BotanicalDistrict, in which the speciesoccuts.

Allinities. Closestto Pimeleaciliata. Also closeto P. brevifolia,showing greatest similarity to subsp.modesta, and P. brachyphylla.

38.Pimelea brevistyla Rye, Nuytsia 5: I 4 (1984).Zype: Glenburn Rd, Glen Forrest,Western Australia,31'55' S, 116d06'E, 6 Oct. 1983,N. Cohen1002 (holo: PERTH; iso: CANB, K, MEL. NSw).

Sftnrb,erect. usually 0.3-1.3 m high,single stemmed at groundIevel Sremsred'brown near eachinflorescence, becoming grey iurther from apex.leales usuallywidely antrone to patent; petiole0.5- 1.5 mm long;lamina concolorous or slightlydiscolorous'-medium green when young, becomingdark bluishgreen, usually nafiowly ovate,7'28 x.2'3 5 mm, adaxiallyconcave or nai, tupJring to a u.t:y nairow but obtuseapex, often slightly mucronulate,the margin recurved.piduncte 0.5,5 mm long.Involucral bracts2 of 4, usuallypale green and partly bl&s!^a/q 'dsqns a lS0rlrlerol ,/.{Js?^r./gd pue V nau polelos, o a slsuauolo d pue 'dsqns ./9 olK)st^atqoalawq Jo uotnqusrc gS oJnBrJ f slllqonadsDapwd Jo uounqutsrq otngrj

I I t-l , t\L t! .ne - b i l**o ) f I ! *. w t ! YJ \' \ 1V g a-aJ r I e ) \ a \t: 'l _\_i dk\i f{N l \\ \ \ \ lL .ni l- Y',i'^' \dilr'

slllssas pue oso^rau 'f 'dsqns f d a olltunnal d puB ouDluuowqald' a ,uoluuowqai Dr oalaald ttounqulslc 'd:qn\ I Jo 95 oln8rC ouotuuDwqalDdlrl./U Jo "i\\\\ uollnqul\lC SSarnSrl I I l 'T /h r i rf , \\\i a a \it\\ ^ r't(ry.i@,Fiin€ i \ -_1f^idquns 1 i I

f n: d tra \ \r tts v t v 1\ v ti\ 1f,\ ,1Nl ftx\a- \\\ \ Nl_.r Nil.,J r,*'i-- 1r3',i"-. rtz .e"Jeceleul{ql usrleJlmv uJolselA'e^U 'I'g 242 NuytsiaVol. 6, No.2 (t9881 yellowish,7-20 x 4-13mm, acuteor acuminate;outer bracts (when present) ovate or broadly ovate,glabrous or with a few cilia, often separatedby a distinctinternode from the inner pair of bracts;inner bractsbroadly ovate-elliptic, much thinner than the leaves,appressed hairy inside,glabrous or with a few hairsnear baseoutside, densely ciliate, the largestcilia 1.5-3mm long,the cilia aroundapex up to I mm long.Inflorescence erecl. Pedicels c. I mm long;hairs 0.1-0.2 mm long.Flowers bi:sexual, not circumscissile;tube 8- l5 mm long,glabrous inside.Ovary-portion offloral tube brown, 2 4 x c. 0.9 mm, appearingglabrous but usually sparselycovered by minute reflexedto patenthairs 0.1-0.2 mrn long.Style-portion offloral tube 6-12mm long, c. I mm diam. at summit, glabrousinside; proximal half brown, with minutehairs similar to thoseon the ovary-portionbut 0.1,0.3mm longand with patenthairs 2.5'5.5mm long at leastin distal part; distal half white, denselycovered by antrorsehairs 0.2-l mm long.Sepals white, ovate, 3,6 mm long, the outsidewith a similarindumentum to that of the distalpart of floral tubebut usuallywith a few largerhairs 1-2mm long,the insidemore sparsely appressed hairy or glabrous,the hairsinside up to 0.4mmlong. Stamens subsessileat throat of floral tube;filament c. 0.25 mm long; anther l-1.8 x c. 0.7 mm; slits strictly adaxial.,t/y/e not or scarcelyexserted. (Figures 59 and Rye 1984:3.)

Distribution. (Figure58.) Occursboth in the Darling Rangenear Perth and in the central wheatbeltfrom Wubin to Hyden. The two areasare separatedby over 130km.

Floweringperiod. August-October.

AJfnities. A very distinctspecies, somewhat intermediate between sect. Calyptrostegia and sect.Heterolaena but probablyclosest to Pimeleaciliata of the latter section.It is atypical of sect.Heterolaena in havingsubsessile, strictly intro$e anthen,with the connectivelaterally exceedingthe cellsslightly, and in having cilia around the apex of the involucral bracts.

Nores.There are two allopatricsubspecies, occurring in the Darling Rangeand wheatbelt respectively.The latter subspecieshas smallerleaves, involucral bracts and flowersthan the former,perhaps as a responseto its lesshumid environment.A moresignificant difference, perhaps,is the tendencyfor the long hairsof the floral tube to be differentlypositioned in the two taxa. Key to Subspecies

l. Involucralbracts 7-11 mm long.Floral tube 8ll mm long. Sepals3-4 mm long...... 38a.slbsp. minor 1. Involucralbracts 12'20 mm long.Floral tube 11-15mm long. Sepals4.5-6 mm long...... 38b.subsp.brevistyla

(Figure 38a.subsp minor Rye, subsp.nov. 59.) i I Differt a P brcvistylaRye subsp. brevistyla invofucri bracteis floribus antheris brevioribus. I Typus:Great Eastern Hwy, 4.1km W of HinesHill, WesternAustralia, 31033' S, 118004' \ E,27 Atg. 1983,N. Cohen1025 (holo: PERTH; iso:CANB, K, MEL, NSW). .4 o Differs from P. breyistylaRye subsp.brevistyla in the shofier involucralbracts, flowers and anthers. S (I Slirab, usually0.3-0.8 m h:igh.Leaves 7- 19 mm long or rarely shorter.Involucral bracts 4', 7-11x 4-9mm. Floral tube8l1mm long.Style-portion of floraltube 6 8 mm long,with a band of long hairs extendingfrom the base(or very near the base)upward, the longest D, hairs 2.5-3.5mm long. Sepak 3-4 mm long. Anthers l-7.2 mm long. to 's rsee-qlnos(a.8r0er r .r.0o0r) urqn^\ 'o+.'r sr,,$i9tJ3Ji,;i'(u;?."J, :it:il;;:;rZ 'urporror^r o JoA\NN urr 0s .J :rral^rr .9.068 .r"q"r"jtl*1firtj;1t5 {d!s!r1,44.r'f /1a,{?s r 1;tq-n[j6; !.1.1.7^,!!".ycl.v'nprttes.Jo I fTI's'w 6L rl rLu11 :r9s3l sloi"td1pJ ".14i :!D_s9t 'ze6rlxo z 'u!qn^e "i1ur l:lj, :l lTl_lsd! .f .n .orquu1:dnaa F s t,iii s-r i .ul,ir*,ti1-1iirijuaj _!,gJt0t6 lqd ol uro4r.ur r :u xiar ooor ,raaig ;y tl ltu :(Ht53dtayw.t lqn,tl -3.N]yIla f | 'wr;rltllB.v.L6Bt dos'irer4rrii;;':q|tni tigzqjox'w LLZqnx W :(HJ.uad qs :lHluad),ul.pe!€i{,:(1gr{]Z .ur8petunl zr}r z i; L$7.1o:r""if"i"{rxlf 989KaIDg J ? l :vrrvurSriv Nuaf's?r[- .paulutDxasuau*rads !3:'i#f":1'j*,f:3liij,'"Jlj;J:XIHJ:"tild'itg;(g,l,IliXYl:y:fi:#,e#H$'#Uij:,#jll"djiijil"i

Enz oEecroelaul,{qlu€rlellsnv uJelsalA.a,{U .1-{ NuytsiaVol. 6. No.2 (1988) "red Habitat. The habitatof the type locality was describedas sandyciay over latertte.on a flat roadverge". No othercoilections give habitat details but two of the localitiesare possibly granlteoutcrops. Floweringperiod. AugustOctober. Derivation of name.Minor (L.) - lesser,referdng to the smallersize of the leaves,bracts and flowers. 38b. subsp.brevistyla

Slzrub,usually 0.5 1.3mhigh. Leaves l4-28mmlong. 1n v olucral bracts l2 20 x 8-13mm' Floral tube l1-15 mm long.Style-portion of floral tube 8-12mm long' glabrousor with very short hairsin the proximal0 , mm, with a band of long hairsabove, the longesthairs 3-55 mm long.SeWk 4.5'6mm long.Anthers 1.5'1.8mm long.(Figure - Rye 1984:3.) "Smith\ Specimensexamined. WESTERN AUSTRALIA: Mill" [Glen Forrest],Sept l90l, L. oi"f U n. Pireel (PERTH);Karragullen, ,4.R. Fairall3l9A (PERTH);Serpentine Falls, J.W. Green361 (PERTH); Serpentine, Sept. 1922, G E. Penin IMEL);Darling Range,Sept l90l, E. Pritzel (PERTII); Glen Fonest, B.L Rye 82009(PERTFI); Helena Vallev, J Seabrook 584(PERTH).

Dist/ibution.(Figure 5 8.) Restricted to the DarlingRange. Extends from GIenForrest (31054' S, 116006'E) southto SerpentineFalls (32022' S, 116001'E).

Habitat.lJsrally occr:rson hillsideswith claysoil andgranite rocks. Also recordedin gravelly sand. Floweringperiod. August'October.

39. Pimeleaciliata Rye, Nuytsia 5: 6-9 (1984).Type: Glenburn Rd, Glen Forest, Western Australia.31055'S. Ii6006'E,6 Oct. 1983,N. Cohen1001 (holo: PERTH; iso: CANB, K, MEL. NSW). Shrub, erccl,usually 0.5-1m high, single'stemmedat ground level,many branchedand oflen bushyabove. Stelns red-browi or orangebrown near each inflorescence, becoming dark gx.y tu.ttr.t ftot "pex. lzaves usually widely antrorseto patent; petiole0 2'1 mm long; Iamina ti"olorou., mediumgreen to dark bluish greenon adaxialsurface, pale green-.onabaxial surface,usually ovate-to narrowly obovateor narrowlyoblong (often appearing linear when margini becomerevolute on drying),usually 5-22 x 1.5-7m-m, tape ng to base'the ma^rgin-s recuivedor somewhatrevolute; a$i acute,-.rcronulate,often recutved. Peduncle l'10('15\ rnmlong.Invotucral brrcts 4 or 6, usuallygxe€n with a pinkish base,sometimes largely reddish, ouui. oiUto"at' onate,8-13 x 5-i 0 mm, Citiate,otherwise glabrous or with hairsat baseand iarely also at centreinside; cilia up to 2.5 mm long.InJlorescence erect' Pedicels l'2 mm long; hairs 1-3.5mm long.Floweri bisexualor very rarely female,white or pale pink, not circ-umscissile,glabrous iniide; tube 8-14 mrnlong. Ovary-pottionofJloral tube with numerous trairsd.j-O.Z tn. long, very rarelywith a few long hairsin distal pafl. Style-portion fatent'of patent florat tube 5.5-12mm long,c. 1.5mm diam. at summit;proximal half with hairs /.i,5 .,n long nrurly always=mixedwith at leasta few minute hairssimilar to thoseon the ovary-poftio;but O.t-0.: im longi distal half coveredby very fine antrorsehain mostly 0.3-i rirm long. Sepalsovate or nariowly ovate,2.5-5.5 mm long, the outsidewith similar hairsto those-onthe distal part of the floral tube mixedwith a few largehairs, the longest hairs l-2.5mm long.Sramens slightly to greatlyexceeding sepals;filament 3 25-55 mm long; anther 0.75 1.5 x c-.0.3 mm; sliis semilateralto lateralafter dehiscence.S/y/e exserted by 3-6 mm. (Figure60.) fl spaourr,/r'dsqns uqlr.r uorrerrc^ oqrlnq posruaocor ,.,",.,r.0i";ttliljr:;#rXtT:ff?# 'scuauoAo .saglug[y d pue Dasotoapuq ol1saso11 'laqrueco(-lsn8nv pouadSuuauolE pu€re^r{ rare'retr.tr r'ouor qrnos uoop"ra J",{1r3XT:.,Jd$**ii+;:l.[111 I}',fr;::ff

qsa{ urorJu^\?.rc (gl x) uJruEts y).1:A^oU.):ltrl lrB.lq C:l0l ]o blouDDalaald 09 atn8rJ I

,?oc?e?lau^qJusll€rtsnv urats3/A o,{U l g 246 NuytsiaVol. 6, No.2 (1988) Key to Subspecies

l. Leavesrecurved at apex.Flowers white or very rarely palepink. Floral tube 8-12 mm long...... 39a. srbsp.ciliata 1. Leavesnot or scarcelyrecurved at apex. Flowerspale pink. Floral tube 12-14mm 1ong...... -....39b.subsp. longituba 39a.subsp. ciliata Leaves:laminanarrowly oblongto ellipticor broadestabove or below the middlewithin the samelength/width ratios, usually 5,22 x 1.57 mm, recuryedat apex.Involucral bracts: largestcilia 2-2.5mmlong. Flowerswhite or very rarelypale pink. F/oral nbe8l2 mm long; style-portion5.5-9 mm long. S?pals3-5.5 mm long. Srazensshortly to geatly exce€dingsepals. (Figure60.)

Specimensexamined WESTERN AUSTRALIA (selectedfrom over 110seen): Manjimup, H.J. Anderson28 (PERTH);42 mi [68 km] Perthto Toodyay,EM. Canning2808 (CBG)\ PickeringBrook, 13 Nov. 1928,E Del/ (PERTH);Helena Valley,I{ Demarz1l24 (C,\NB\: Bickley Reservoir,l{ Demarx 6199(PERT[I); Wagin, T.E. Gmrge 191 (MEL); Mount Barker, T.E. George561(MEL); Wooroloo,14 Koch 1424MELJ'Manjimup, M. Koch 2455(MEL); Kelmscott,I I Sept.1897, ,4. Morrison (BRI,CANB); Darlington,6 Oct. 1910,A. Moffison (BRI);2 mi [3 km] E of Kojonup, K. Newbey 1303(PERTH); 5 mi [8 km] W of Darkan, M.E. Phillips 4005 (AD, CBG);near Lesmurdie,I Pulley 1346(CBG); Glen Forrest,_8.t. Rye 82010(PERTH); Mundaring Weir, 1936,I Sco// (NSW);Rocky Pool, Sepr.1929, A. Steffanoni(ADW); Stirtng Range,F.W. lrye l54 (PERTH);Manjimup ro Nannup,r.l,E Whibley3242 (AD, NT); nearPorongurup Range, P G. Wilson3369 (AD, CBG, PERTH); Collie River, P.G. Wilson 3761 (PERTH).

Distibution. (Figure63.) Extends from north of Bindoon (c. 31050'S, I16010'E) south to Manjimup(34015'S, 116009'E) and south-east to nearthe PorongurupRange (c. 34044'S, I17055'E). Habitat. Recofied from healrysoils in lateritic or granitic areas,usually on hills.

Floweringperiod. August-November.

Noles.A variant in the Manjimup areais atypicalin havingthe long hairsof the floral tube extendingdown onto the ovary-portionand often havingfew or no short hairsmixed with the long hairs in the lower part of the style-portion.

39b. subsp.longituba Rye, subsp.nov.

Differt a P ciliata Piyestbsp. ciliata apicefoliorum magiserecta, ciliis involucd bractearum brevioribus,tubo florali longiore.

?nyprzs:Jindong, S of Busselton,Western Australia, 20 Oct. 1950,R.D. Royce 3402:holo'. PERTH; iso: CANB, MEL.

Differs from P. ciliata Piye sabsp.ciliata in the more erect leaf apex, the shorter cilia on the involucralbracts and the longerfloral tube. I Leaves:larnna narrowly obovateto narrowly ovate,usually 8-l3 x I .5-5mm, not or scarcely recurvedat apex.Involucral bracts:largesl cilia 1.5-2mm long. Flowerspale pink. Floral t tube 12-14mmlong; style-portion8.5-12 mm long.Sepals 2.5-3.5 mm long.Srazens shortly I exceedingsepals. c 5l nry:Y3-ryIjgrroqs,'nqars,oqnr,,"o;'il";J,'#lHl'#^|"Tr;'#ti"1,tLf,?JllT,:E'aqnl lBrolJreuoqs'rnoloo ra,nott. {rn_d_ daep isi,rpel reuos.rrrJoJ qrmora'rci:ouroj paqslnaunsrpe9 ue) -s1rps:611geq .a "p:i "q ,iii[ Dasotoa1aw14 ri1 oiottp puasul auurnmoaaso.r le qeq ,{qaelri:edoi_ecf or raioouie!1 qiiq,riql uo qanoqlt?.or4?dru,{s seunauros:{--_y-"9j"qlp .4 eq ,{eu pu?roqloaoi osolr Jnm o o"so, pui nqnliiuolldsqn ololllcnapw14 .sa1o1,J 'oqnl lerolJEuol eql 01AuureJor ,odrd _ ,Auol_ .awnu l.-i) snqry (-l snhuoT 1[ouotlnauag .Jequecoq-Joqopo .pat4 Buuauolg .slros ^o,(?Jcul stnJJoflq?qoJd ...Ios,{ur?ol poJ,, w. pepj(y)el .fijlqoH 'lfl.-,:0q't\1.'F:Zt'gE) ale8raquryol pxp(g.S6oslt 'S ,taluerg or qrnos(g lsuo .t90tg) ,zo,st ld ,6i,d; ,il.i;,iri; "i"t ,p'*r"a i.sq ,r*,g) .uoltnqltslo '@yl 'rls''u/rou)lun .dnAurJIeI 're^ru. aIr/,/A elep :(HIUAd .OV) y.u:)9._la|!!a4?r'o rorear_nr1glg.lryly^o3 :1111u3fu Jr,rjy-A-a-'.fid"ii ua nd '.q',{elulrg-r€eu :X .;eii 477lirifi :lqryf?'90sz0t .IAru 0!86. ,^eg i(.lsr/\fl"rr{d oqce-:aoig:11797si aaruJ oqduEooc:vrlviifsn-v nfti;ia1,\ .paarwDxasuauuads

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Flov 40. R. Br.,Prodr. 360 \1810). - Heterolaenarosea (R. Br.)C. Meyer,Bull. Cl. Phys.-Math.Acad. Imp. Sci. Saint-Petersbovg 4:73 (1845).- knksia rosea\R. Br.\ Ajlit Kuntze,Revis. Gen. Pl. 2: 583(1891). ,l,pe. KingGeorge Sound, Western Australia, Dec. Pime 1801.R. Brown{BM). picke a

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63. Distribution of Pmelea ciliata srbsp. ciliata L and P. c/iaJa subsp. longituba a .

n-ri'*\ J rti

a t sl .\ f"r-PE f-T I i l6unby'YfI ]T a I I .11 ri+,ft ta 'i-l ll \ \^ I l Ffgure65. Disrributionof Pimelealerruginea.

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".rr-$ Ll I co 'i-\ill\F t . tl sl I f t-i" FI( Figure 66. Distribution ol Pimelea hispida. Art 'otDlllJ.d pue oasotD4aug ot 6eso13.salnuglfV ',tueruqog_tsntnv .pouadBuuaMol{

uo$r'or Jo sad^r reqlo ro orru?ra ro^oo"r.,y;;lo$Jliot?:::i"ffitliltJffi1f,i;il 's (a,6sorr r ,zto6z) p?oH JJrrJ ruo{ }s?m .il rttfit"lJ,B;j,i;it,?}'."r,li'3 iY},Zi;;;ir:;

iDBJ ,ldgl KalBuA4./n.t'g96l .1rO .ur,rnea.l '1JgJl ',41'l'896l 9l ade3 :111i ^alzqt4 lro .{ueqtV'puElpBaq 'S'V "sI €t 1:oy ;1n9:tHlUlld.gNV)r I8f6 uo$a^ ^rots rosqo :(HJUAd ,SNVJ) i1u1od 'ocuereosg 9g7g,f.rotg y rets,rqsordJo S ru{ 0J:G{JUqJLr0z8 a'q 7.s gii,,r-l d or :tfriaat Ltt$nDd s.^Eg at?"g -dnrooru:(HJuad) zLs KaqMaN y.opnny j9 giir-fij:9 _.\_rsw) .s.J 'pugJcr-r adBJ :(Hrxsd!) 'kesSi,t{ a LsZr4nN 'I'Y 'uoopul s00rlapuo't .sw qroir,giii; sN ;"i i; iH:irsdr?l r6s-r \nliDuH €t€.I uDr :(HJUSd aBji,SS.3V JoiA 6t tiE lel"r dq*r.-C*tniriaij,rirro"cri,r,^u*ri-."i ?,92:l_:!f:i?^Y,?,'i9jT_s 'H!!lTa '"ruBiods.g-Jo r?eu:(HrusJ s7671nuwoe' ;/;r :tHJUgd'CVl0t66ltalqtg A\UDI S.0.c:fiffUaa 'f?g oqaersl:(HJd3'& g60tztlua1 OV)OZgTraiuoT l{.fi -D.W,,reg 77 r^g Js*oj ion*li,*ui-,irrr.liriin t^v pridqlf i.llgtg LngLy4tlo) reura:g:tSNVCt ZeOf )?ptqig .J.N;reg ."iOb.j o;1 \e^tnJrurp6jo i(Hruaa) p,r"s :15jlYgg].:^0r9.j:loo!_B_-e'w'lurod ^puBS:(uoes ,11lrua !11 ii br .pauwoxa{ss[ s:i 09I r0^ouo+ peicolas)VIlVuISnv NxaJSs,,11 suawltid-g

a1&secusJsrqep_roue rBrol€r01r'relerruos srqs:uo, e.0 1"t;;iTlfl;frf"o'1r[l"lfi:;1; ,{uq81;s,qensn saaara;5 au-oi ruru ie rz i ","q rsaauojaqr .i.,izq duoTIi.lT:llJ-tl:9:,pue pazs-unrpaur lrpaerxaqlt/r\ ,3uol .318{o rxru,,s.2 s7rda5.r:eolutslp ui auolri: ut 3.g.i.0 ut€uolruur,.0_t.0 j :jj:.1jrj:.11":^q]T..xedleutxord ,lururns sreq-rroqspue dirol Lur ri ,1g ir,'"[ lurl?o-buol Jo arnlxtrx € r.ltr,rl te.rrrerprxu 1.y_1.g,Buo1urul iqiljriolilo uotuodaf g rnorl8norql .auoj 0l_9., ro uedlasrp u1-s4eq i'tro1 qrrm uai.1"o ,, g.0I.0 sreqrsrorrue :]:^1ii.Y.,ji.yg(.1.xg,7.taqntpto11tlouoruod.,{tia)'alrssrcsunrrn 'Ienxesrg auolurui61_1iq:niipriuisno.rqela lou'lurd daapol elpd saMold.auoluruli_|7 916,ridq iauoy.iiiu suolurtu (t-)z-s 0 erlc:mo:qep eirmieqlo 'alrirpc rfensn !f:1.-9!!!!l!.].??.1^" r_rxelxea:y?tu-t/4 .ruul :ate,ro .Tl:g-r::9'rno{dnorqr lE srrpq.^{aJ e qll,r seulla^rxos 0 I_Sx tr1_9 qpeoiqX jjensn d}apot f Brrredpue .7 .aLibl {ud lud l uaarS{11ensn s rsotq iitjryoru1 ui I'br on alrunpadxode lE- pa^mml puB elelnuolcnut .suri.reur pjmncoj irudg.9_g:1 .}JEyns [ll,n * ql g,o, I!_":yj:3ry]!r",q,\torreuol cndlla [prx?q?u-o ueql -nJ-,'n, 1n,"npn uo uoa6ro{r?p qcnu ? 'snoroloosn^ra eurruel .Buo1urur 'Eunol :Buolurru I- 1.0-,16eiqelortod:luoieA nliensn iaaiz1 srl8qaql uoq,r,{rr?q ueryo 7lg.g ,le^ol }€qaouos o1u,norq_p'ar daap sut tS,.e^oq?osuep puepaqcu€rq"fueu punoJ8tE poruruels-el8uis ,{pnsn ,,larq,u g:1-g.g;1cen,qruqg'

'ro^ew'J jY,l,i8 ??:?:2?:tt:"d({(s'8r)€rr_it![[?r"q,%1931d.;,"qJju'tly^1ff],ojDssnrap _ .(0ISi) .rpo:j ,.r{.\ \'rg.E) DuaqorataH g9g Djossnrapoaput{

^!L]:{ly1"".rd :opq)a^lplDttlqDr ap,.H I.t.Z6LI rro ,,_,, ,"unnno,TT#.fft,,,i; aruurrdsgle patcrlor ,{tlEnpEl..ur,lrne.I nii":iii' .ueC.sr^od .azlun) DauBruDlDleruDg- 'o .ra.lrog ttSitt"EgiiZ:ra r'r[qe'rl g0grl 9'r i :i i-oN".:1i,qL1""urunrr"l "atau4d .J, I9? eeroBaelau,{qJuerl€Isnv ulelsa/ .0,{U .I.g NuytsiaVol. 6, No. 2 (1988) 252

A

(x 4); C-flower (x 7 5); D stamen(x 10)' FilAlre6l. Pimeleahisprda. A-flowerilg stemlB innel surfaceof bract D{awn from A.M- AshbY)682

(1810).- Heterotaenahispida (R'Br ') C Meyel Brll' 42.Pimelea hispida R. Br.,Prodr. 360 -_Banksia li. i,iiil,Mffi:'ilJ.lrnp.d.i. saint-peiersboure +:13tt8.4st. hispida.tR.Br.) i#J. il;;;.'c;". ir.'Ij sa: tgglt.rype, KinE Georee Sound. wesrern Ausrralia. Dec. 1801,.Ii. Brown (BM). red-brownnear Shrub,ercct,0.4'l5 m high,single-stemmed at gxound- level' S/ems usually grev'brownfurther from apexLeaves usuallv antrorse; .";h';li";;;;;,;6;fis'd".I -aiscotoio"i, palergreen il;ft; iamina mediumgreen-on adaxial surface and a ;;iil 1' recu ed'obtuse 5;;d;"i il;,?[ipii" o. n.utrv* ,g'zg x z'g mm,tat or withmarsins I

aqry .1\ SZ-Z lDJol[lo uotUod-tlD^Ouotl]od.^J?^o o^oqB uu t,Z fJl€nsnalrssrcsunclr3 ::prsur .i,roqe- a! :3::l :|u_ !.0.1.-2,ig"l snorq-e18 lurd pue rLEdllseq rqr ur lrq^\ ua1o u :0 'Jptsut i1"#"'ii,,1Tl#,Ts,tJi JI :JlErIJiiii.:ii,.^lidl;ff:l'#,' lou liEq-pJs$JddElifi [ ;rfi 1:t,5*.t!;9,ff ,r"fi 'apyqno ,{llprupdsDplq JJlno :JprMuru 9.i-g.g sursLeu qsrmog'eXaqi sno:qep'py q,t x g-9 .ot" ', o.surEreuoql u'd peauallirririauipouoJ p;; ;;di ol ueeJolnq uaarE^loEJ?l spDtqp)rryo^ul 1rn.r;ur iuol urir'61or iln ,ro,r,roguririoy ww allunpad elplnuolrnu .o^Ecuor ,rdleui 9r-t to rlplnld? ,{l;lnurul i(lplx?pe to fef s€^Eal,t\eJ E florel I lo rulIl \fZ x Sz-6,ecuecss:ogu1eqt ,roloq ^lloeJp sa^eoloql ur ol€^o 't .ro:qdq1a ,{;,tro::eu ,{1pnsn ,urF:riur lruerixj uii qsuloJle,{:b qqpper uJalb::113"T:.:.^l]r:^1 'os 1nq {r,tnrpau ^lJeJuro snoJoloJuoJEUrrxel :aJuaJsaJouur ue moloq {1ne.r,i irnual ,i rIx etouad .xadi lY.::.Ti:tJ::-ll9t !'t-Z 0 :asrorluesa,rzaT uo.rg:eqsnJ u,rorq,^era I,"p 'I ii,"rfri:,il,i.r;'i?1:L1x:1 JffJ"*q##;;:fr !#:,:v6:i;ffi,;,r5;:ij,l"ffi:il" lINg)uMorg. .V .I0g coc ,?lle4snv ,punos .ls,ts l ure$alA a8roa0B;C|.N:friJ.(1061) ,6€ :St sror(r_(rs, oruuoj.ten -.1g1stle .:1T{..19_'l.tjr.g-'s'uoC"lpuA ? d) o[1ds1t1o11"*1j :iioirir5 1j \'rg')d)olDuot otaa$ofidtlD) -.(0lgl) g9g .rporf,,:g.gz1eue1 ealarurj.i| .oasot.d ,ojouDl ol relnursosle D4aw6 ol lsasolJ .sapluri[{y .requoro6J_laqureldag .pouadBntauol g 'slJrq pws l?tseoouo pu?steu po8Aolrot?,{1. ^lpuos?os uo p}prcDe.u.lulqDH '(g .9t I0gt I ol quou^uEqlv ,s 'qrnos-rsee.'s ,f zotE)IJou)I'S JJntg urorJpuE (a .t90l. t t ,z00zr)'iiidigt,{u€qlv 01 tsea \g ,LZoStI ,l.t,fg r) ^EgrqderaoaC uroa spullxg:i99 .*ihqutlg

rlreceod:(Hru^ad)vtn,,lovqy_.dl,ij9ryr""1p,[t]);f t"{"J?\ a:l9ly-uoo.rreq:(HJIEdt a{!;rt :ii;1l::\rfl.i,ibie i"^J,f: 'lroux I 't ,L[Ug:i'^u"q'r$;o .tad s E,g':1H1;sij 9J^2!?!^"N )r !Jnts:(rsrN),:aanry 2l 151u1uoq,4 :(oy) u.o_etrD['S'NE're^tu^]rorsJolINgruy4c pxuleq3-airqr-lriiusairr0;i;"iir;D'.;;; ,82€ i(ssit8uuunj.hts..8sqt .burp,5Eu;'o;'Jo :l:!:l !r"d 1167i ^{i,t"i;rij lr"oi:, :F+ua.,g--qYlq89ztEtsv w'v'e$:.8nv.re:u:1ssxj4vsti-itav:ir:s.j r,i.i1rr{oirj Jo A\ :(u30s99 r0^o {xo{ pepolagvITVUTSnV-N)idiis,ld .pawuoxasuau)lrads

porrasxa,1tr./s .rruaJsr_qep .urur rarJErErorEr ror'rrrpr-rruas surs if ,t;":'ii ,lYJ;ff,i":l :]rruJ'€t lueurellJ:qedosSurpoocxe s Aemp Kpeeu suatirls auoiiuiu slrgq H,?'l'i'li1i;irili:,'ji:l#'l#',:i:#j:t_S.Z lse3uolaql ..d&i:iEi!?i,3T'illil"iiHi",i!"ji:1"'l,tlJensn ffifi 11T:i1"1-y- s 0'J'0luelzd 'Jpq ;1eqlerulxo:d Jo ir;q ;r;,i ;eq prsrpur Buoruur pu"porotl?cs aroru purxordur 6uo[ rxru puLpllarlajiroi ,t_--:_Z pm S_] ludieOl:rert auoi ::llYy|:9q: Euolpoxru qlr.r,uuruns 1e uetpiuiu'g.1,i ;auo1 . q::.":q:qp,o peu,ilrim ualsb {^'{!:' f#,fi',f1***t#i"#*,*: :;,i:i"{,fW'"J;.T; ^J€^r:ti:11".rlT,r!q1t;pffi#il;'?I'$';:,g,l:I:fJo sJrMolJ.BuoJ ,i?i:;!"{iw^i::: lenxesrq ruut? ot dn sJtuq:Buolww 1-i gs1at,p, anolurl 1g,;iiarc,rrja"uoteqi;ei",trc.urbr'u oqlpunor' o""o,."rqjt ;:11?3:?::i;#" slt q reuul:ar"ryr oreiruac "q1 ,uojJ -pes$rddeT]::].fj:q:lT:t,ddBstcerq .aprslno ,warlo1lesiq'eqr lsery1e aprsu ^rr?q ralno sno:q€l7 -L lypeor{tuoluoo ^oitJ qllr ,{ll€nsnpue ueara.v lt'02_6,e1eno rrj lurd spDtqlonnloiillau6i &w ws fiii"pua.e]Elnuorrnu ot ,o,tu.-I oeecEeeJout,{qluerc4snv ulols.ad B NuytsiaVol. 6 No. 2 (1988) 254

C

(x (x10i' gr:;:"1i.IT!",fr,H,2::ri;ii:tixl'"i'lll;i:;'ijfiTi'o'i*'E5);c'nower 75); D stamen

patenthairs near I mm- with reflexedhairs 0.1-0.3 mm long.with a few longantrorse or i'iiilit. iiyii ilni iiiira tuue5-8 mri.long, c' I mmdiam at summit' with a mixture lfl-onn o"ibnt'fr"its and muchshorter hairs; lonE hairs concentrated or sometimesconfined i";;;;;ilJ';;;il2-:.i rnt ronein proximalhalf. often reduced to l-2 5 mm longin drstal iilfrii'r,ir, rrli".. 0.i ;; bd in proximalhalf and c. 0.5 mm tongin disralhalf. sepals ;;-r;;il;;;i; ot o".G. z.s'l.s inm lons.with. lons and shorthairs' the lons-hairs i Sro*ensexceeding sepals; filarient 3-4 5 mm long;anther1'5-1'8 x c' 0'2mm' ( i. i ii'toie. (Figure ,ii[-ri-i-fui.."f to lateralafteadehiscence. ,Style exserted by 4.5-5.5mm. 68.) I '(I,l!lg)u$otg y '1081'rec 'Ert84snY 'punos 'sr^ou'e4rmx 'u) urslsolA o8IoocEur\:adtJ fi6glj t89 :z Id uoo ('rg DlsluDg- '(9t81) :t Emoqsrotod-luusrcs 'dul 'pmv 'qt?W-s,{qd'lJ ryoti^alq'refeni 'U) t/ '(0181) 'rpord ''rg 'U Infl J ('rg 4{o/ratq 48a1so4dtp, - 6St B[oJ!arq Balaurldtt

'vr'001 ^v'p-u? r(I'f r./r{oJN,{q pr)uas.rdar reua?Lu qsrJ urorJ u^\erc {6 r ururelsc:(6 J,tl/.99r!i"3?6] ^llqtllslra^\ou.):19xllJErqJoJcelnslalno-B:LUJ5Suuel1ou.VDuotr^dq'dsqns olloIt^drq tuldau 69eJnfu4 c

'otD.t'd 'Dpldslq 'saltlugv o! sanlurJJeqlrr{ oqe ,(lqrssod Dapwld 01 lsesol] 'lsnEnv-qcl?w '^JBnJqed-Jequ$e( 'poltad osl? ,{lurcI I Suuauolg 'uollsloSo^pu8[10.$ '^BJc Jo $uEld c4suolcBJBqrJeqto Jo s{JEqJededqlrrn pal"tcoss?uolJo JeAo[os ^puesJo puBsu! 'urqd pls?oc eql uo s1?$Sul^l-,{ol ro suo$serdoppeEEolJele,!|. f[euosees n smxxo 'JulqDH '(f 's ^uBqlv 'S.t0oz€) .tsol.t .zoo9t)'uoqnqulsle 01ls?oJ oq1 punors (g,lSo9ll olluroql uro{ spuolxafzl eln8rC) 'HIUiId DM11!,+l'!, 'ro^rultors :0{IXAd) ar$oY y 'dnuE,{og:(HJUAd) 09ZZ 'aolpn'I 'CIV) 96ZI O 'c tgll ac,bv Crdr :(HJXAd 862,ptDrlslo ? H'dnuro{ooJJo lA url 0t :GIUAd lGqMaN 'IJ?ruuoO :fiUO uon ol\l '2061'qeg. 'ltocstulo)0rzl )1 JoS [ur{ 9ll lru 0l V iI :(lAl,l!llLSge t4tox J4l'uo4cunfuoldd :(HJdSd)91 aSnag S'H',tunqung :fiIUAd aDwa1 'e!Fuor[ :(HIUAd uasua$! 'd Jo'lulod S luq S0t] IUI 9Z b Z I{ 'pauruDxatLZ t) lc?lg :(uoesgt re^o uro4 pelcolas)VI'IVUISnV NUSJSII^I suaunads oerJeoeleur,(ql ueqeJlsnv urotsold '0,{U 'I g 256 NuytsiaVol.6, No.2 (1988) Undershrubor shrub,0.1-l m high, often multistemmedat groundlevel. ,Srezs reddish brown to pale yellowishbrown near eachinflorescence, becoming dark greyto black then mediumgrey to brown further from apex.Leaves anlrorse or rarely patent,sessile or with a petioleup to 0.3 mm long; lamina concolorousor neady so, usuallymedium green, narrowly elliptic or rarely elliptic, l-16 x 0.5-6mm, flat or adaxiallyconcave, acute or obtuse,often mv$onulate. Peduncle ll)5-20130) mmlong. Involucral bracts 4, similar in colour to leaves or moreyellowish, sometimes reddish at baseor aprex,narrowly ovateto broadlyelliptic or broadlyovate, 4-12x 3-9mm, glabrousinside, rarely ciliate towardsbase, the longestcilia 0.5-0.8mm long.1n/orcscence ercct. Pedicels c. 0.5mm long;hairs up to 2 mm long.Flowers bisexualor female,white or rarelypale yellow, not circumscissile,glabrous inside; tube 6-12 mm long. Ovary-portionof floral tube2-3 x 0.7,1mm, with shortor long hairsor a mixture of both; hairsantrorse, the longestones 0.3-2 mm long.Style-portion ofJloral tube 3-8mm long,0.6-l mm diam. at summit,either shortly hairy throughoutor with a mixture of long patentand short hairs in proximal part and shortly hairy above,the longesthairs up to 2 mm long.Sepals ovate to broadlyovate,elliptic, 2-4 mm long,with shorthairs similar to those on distalpart of floral tube and longerhairs up to 1.3mm long.Stam€ns shorter than sepals; filament 0.3-1.5mm long; anther 0.6-l.l x 0.2-0.4mm; slits semilateralto almostadaxial after dehiscence.Sty/e exserted by up to 2 mm or very rarely not exserted.(Figures 69, 70.)

Distribution. (Figure73.) Extendsfrom nearWubin south to Albany and south-eastto Israelite Bav.

F lowerin g period. July-October.

Affnities. Close to .

A

C

rrAlrasubsp. modesta. A- flowetinqslem; B flower(x 9i; C- stamen(x l2). Drawnfrom nyrt:di|"l"{r!* 'raqopo-^Ft'pouadSmtauolg 'spu?lqnJqsur poploJoUalruz:8 .ro olJolel qlt^{ uouo'fBIc salurtauos Jo sllos,{puBs ur srlr.JJO'1u!qDH 'S.Z0lrSt) '(g 'S c) ,{394lla?rsl r?ou ol ls?epu? (A ,Zgotzl 'S.1,€o[€ .€Sol.ll ,tu?qlv01 qlnos (A .82081I ,90ott)ec?r|) oI?'I Iuo{ spuelxgfEl ernEld)uo!fiqltlslQ '(HJUAd) '}, 'poEEBd g1g1Ka4{!ry y }I tr:(quad 9tl 7snr2'5'ad:oqlsue^Bx Jo l[ [uI{ Otl lur SZ:(HIUAd) 60tI {aq(\aaNy'dnunrqlool U{ :(HJUAJ iaqkaN 'aJEJCe}l?.I g i(-Igl^ 'g'J'puerDsl Lrct ) JoA 'o8u?u luq t] lul I 66Z14nW ed?3Jo g uDt0l :('Igw) 0L8ttlnw 8,J sultrns' ou) #nls :GJdgd) 880t9/ 'I'V 'u4o,rlel4{ tu)l :(HJUAd) Dupto) 'X, 'uoJlEg1I apprW \n!fiuH 'CV) trurod'N'NJo t[ 'peorZ f I Lt6 tr :HJdAd gtO€nuuo] a:ue:edsg-ed:oqlsus^BUuo Sulssoro le^ld Sunol Jo IINNrq0e c:(HIUAd'(v) SZBZDuuoI N'N'f,8geIIaBrsI Jo ld tu){8 c:(HIUAd'CV) 069Zhuuoo NN lllH {tlrr oH JoN UDIt c :(HJuadl 690€ mlzotgg '11'dntflu4nau :(HIUSd) auuau g'dl].re8u} gS :(HIUgd '(Jy) {qqsv 'V 'oEuBU 9g6 W Jo [ru)l6Z] tur 8t h Errlrrls :(ums0, ro^ouro{ pepeles)\.I'I\.UJSnV NXAISAiA pauunxasuawnadg 169::n8r1) aJuf,Jstq3p relp lsolxp ol s1!FDqiuv eurJ'8uolurtx 'osJoJlu€flasolc ro lerx?pe'snorotunu lerolel-ruros 1,0-10 posseJdds ot,{eJ u€d lelsrpJo sr?q oqs:sseu)lclql uI unrpeu ol osl?oJ'Euol rurrr€ 0-10 sJmqetnuru qlr,r\ poxrrx uogo'Euol ruul Z'l sjreqEuol :e^oq€ sr"q Jeuoqspuo uoqJod-ol^lsJou?d l?urxord ol uorlrod-^rc^oJo edlElsrp ruo{ SurpuelxoJo uol od',{te^o 01peur.Juor srreq Suol ql\M aqni ptoll 'o4e^o.,(p?orq01 tlll'orreuslruq 'ozs 'qruqslotJryo^uJ'wldJ €-SI tl ) x 9l-gt€) u u.uoJrunl{nBJ ^[8nsn sr,lra1 qAIq tu I-910 ro qn.qsrapun '(lfj,\tr:u^sosl) aMxDW't 'u.rolj)lunol?p'erJ?rtsnv uJolsai['f?g acu8Jodsg:nAW :uf,sost) aMxDW 'u^{o{une}ep 'elle4snv '.(IgINtutsosl , 'EIIertsnV urelsod!'lolulumlleJcl^I natlpto U Z'u^\ou{un31Bp urelsel!\'re^ruu?8le)I :() :pelBu8rsopereq olr,el) ppt{pto j,t,'u,r\oulun at?p'€llerlsnv urolselA'ra^r{uopro] :adtJ (1681) :Z ueD'sr^au 'eztunx (qtueg xa'll3nl €89 ld 'l tr C) qpMxDtuDleluDg * (e/81)gl-ZI :9 l€rlsnv ld'qruog x|llonl{ qllakxowDapwq '(TAW:osr '.\:opql '9 'u,rou{unelep 'ell?4snv 'l?llmy ! tpttxow uals'a ['puv adeJ:adnJ'GLB ZI :9 Ig qlueg4lojm7uo nt mlotjotq oa1ary{ BlloJ.rarq dsqns Btt

.' " " nlsapow dsqns qtt a^oqPsJIEq oqs pu? uoluod-el^lsJo 'uoluod-^le^o ued Je/Aoluo sr?q uoqs pue sJleqluel?d tuol Jo oJnlxttuE qtl,{\ uo ^req fluoqs eqnl plotd clldllle ,{JpsolqJo clldlfia slc€lq Islsnlo^ul 'dsqns n11olltatq ett o^oqp sr?q uoqs pu€ uoruod-,{rzlo uo (sJIBquoqs ql!,|Apextu ua o) sJrBqSuol qlra eqnl lelold 018^0,{lp€ojqot ^l^\orl?u slc?Jql?lcnlo^ul salJadsqnsol ,{ay .sluqd pnxasrq usql uolxluoc ssel,{llueJBdd? lnq peeldseplt aJesluBld al€urec BoJBmelD elB'I oql ur seau"rraql Jsproquouuoc oql lE ap?l3rolut01 pual lnq sol?tsl3cld,41 Jleql uI Jo 'qlnos Irurlsrp^JeA eff selcedsqnsoqJ eql w. ottofiaatq dsqnspue quou oql uI SuuJnJso D$apolu dsqns'JUlBd?rBd flletlqdeJEosS eJz oseqJ posluBmojeJs setJodsqns o,^AJ saloN 'olU oeec?aeleur,{qJuerlerlsnv u.lolssld 'I g 258 NuytsiaVot.6 No.2 (1988) Noter. In the easternpart of the rangeof this subspecies,pafticularly betwe€n Esperance and IsraeliteBay, the flowershave largehairs on the ovary-portionof the floral tube but not on the style-portion.From Esperancewestwards, the largehairs of the floral tube tend to becomeprogressively higher in their position on the tube. In the northernmostDoDulations o,f subsp.brevdolia (in the easternpart of its range),the positionof the largehaiis on the floral tube approachesthat in subsp.rz odesta. Afew specimensfrom the easiernpart of the rangehave subsessileanthers with more or lessadaxial slits and the stvle not or scarcelv exserted.

m1l

Figure7l stamenlx ftf"1*.!{:,"!l*:!?^1;rf?}!1,?-,ii1fi,f.}?,1;,'tBi}"i^':3:B,"ilii"i,l}?:R"1,3},"iSilb"

44b.subsp. modesta (Meissner) Rye, comb. et stat.nov. (Figure7C.,

Pimeleamodesta Meissner in Lehm,,PL. preiss.2: 269.2i,0(1S4gj. Zype.. south_westem Australia,1843,1844, J. Drummondcoll. 3, n. 23g (iso:K, MEt, i{y).' '10 undershrubor shrub, mm l m high.Leaves often very variablein sizeon eachstem, thesmaller leaves 1-6 x 0.5-2mm, the larger leaves 6-14 x 1.5-6mm.Involu*al bracts ellipli; or broadlyelliptic. Floral tube with antrorseto patenthairs, those on ovary-portion0.1:0.3 mm long,with a mixtureof hairs1.2-2 mm longand much shorter hails b.i_0.3 mm tong in proximalpart of style-portionand hairs 0.4-f mm longabove. Anther slitssemi,latera'i afterdehiscence. '(HJXAd) .r.dr ,uoppew uos 1IAI 'H't'Ln6I ,qctedss?rD SI89 14 Jo l[S rut Z :(]At{) stlt/ ldes :HIU:Id) atY .7g ,,{eN 'pu 'cv) I Jo S 0t028 1I trJo ld,s {fl,roH :(aNyJ gLll ploqrlo a.r,lalul se{ols l" ls?oc rul Jo N €l.X LSLZ taqMaN y'ed:oqrsua,rag Jo A [tu)l t] :HJUAd) taqftaN 'dnroSuoJo?:{HIUAd) 'J.g,ZL6t.tdosIul Z ,feg Ztt IAN [rul S 9] Ilu uospN alrte?Isl t:(SNYJ)'a3uerodsa EI Jo N luq ZI] Itu 8 :(HJXAd VgB6aStoag :(HJUAd) aSnat) 'CV)S't, Jo AN [tu{ 6] Iur 9.9 \ZLg lW :(HJ{Ad DIrplEI 'ro^rU ptorJplo qtnour uDt gS .c S l,'uoqg '(Iyl 08€02 H Jo Jo N :(HlUSd tauuoe ur)t .c :(HIUAd) .g 'dnj8uH tl LZ rV N llrH {cL{.\oHJo N Z rc}t ttbpDlg 14 :(HIUAd) 'g'114's]0luuete>I g ru :\CI]|) .W.V 'lco1 886ZltDpolg Jo [ru{ Zt] 0Z Z,qqsi uuBJ :(uoos99 le^o ruoJJpolcolas) VI-IvUJSny NUAJSAIypauruoxa suawtc)dg

'ultx ( l/ arn8rj) t-g'l fq pallesxea/,&S $usJsrqep JeUeIerelEI rrxas sllJs :uur Z 0 c x g 0-t 0laqlu?:8uol uu [-g.l lueuBlrJ:sledosAurllenbo .,fur?q,Euo[ saurlourosro SurpeexxesuauDjs'3uo1 urru (Z-)g.l-l slr?q lso8uoJoql uruJ 'ale^o ,s:req t S.l slDdaslnoq8norgl sorxrloruospue Ed JBrurxotdu Buolturu Z-9.0 1ua1edBuol gtp\ us o'8uo[ ruur S 0-10 s]lBqesJollu€ ot lue]ed Uoqs qll,r,lturuns lB.ru€tp rxru '8uol .}rrutxns I,t.0 ulru E g Z aqu prc{lo uoulod-af$ l€ou slr?qAuol qllt seru4euros.a,roqe Suolru-ur t 0,1 0 sJreqesJoJlu€ ol luel?d oqs qlh\.es€q 1?snojqel8,ruur g.yl aqru ,elssrcsumorc l-g.0x.a1rq,r :eleure;ro lDlollfto uotuod-Kt ^O EuoJrrrru g,t oqnl leplsul snorqep lou IBnxosrqs.IaMolj. SuoJ rrflu sJreq:8uol uIlu sprlpad .peJenunsarcl)[ul .Buolurw 'el?qroS Z I 9.0'c S I-l elllo tso8uoloql uouo'aprsur^rpq-possalddE ^lletued uo o slc€rqtso{rrJouur :slrzrq tsouljouuru?qt,{Jr?q ssal lo snolqelE(lueseld ueq,,y\) spErq pu? slceJqlolno 'Jrld ,3tldrlo ,poJ Ippotu :uw L-g'Zx 8 g'? e flpeolq Jo el?Ao p?eldsepr^{orou qll,r sorunatxosJo 'g .?uo1w]dJ .po^rncoJ ul3relnqsppoJ ? qlr,{ uoe$ Jo, spotq lDnnloauJ t ol dn alzunpad pu? qeuorcnu xod? oql'sur8rEurelnlo^oJ qlr,|a,tuur x ,Auolqo-orldlllo 'ecEFns €-l 0 I,Z 01 t€eurl uo uoeJ8leled'"reJms uo uouo pu? snocn?I8 'snololoosrplerx?q? JBrx?p? ltep,(Jo^ lo uoala Bururel:3uol ruru 01 dn to ]uesq€oloqad :osloJ]u? lo sa^Dal x&E worl [ 0'ocuoosotouur luel"d JoqilnJ,{olEurnrporu Suruoroq qo?et?au u,roJq-pe.r sorullatxosto 'q3H {JBp'qruqstapun {c€lq suals'p^el puno$ 1? po{xurols-eleurs Ul I,l 0'lcaJa ^lpnsn ro qruqs

'(lgl,^rtr .u.S .JIo, :otjelosr:X :petEuErsopeloq opol) 6Zt rStgJ 'erT€llsnv .(l6gl) .ld .uaC .sr^ax .ezlun) puouaruO T uJalsen-q1nos:ad,{.; €S9:Z .lC ,.qluog .9f (quagJ o trtldtqrotq oleluog - (€tg l) I I :9 I"rlsny sll,iqd,tqcu.rqualaur;d

'ioqolco-lsn8nypouad SuuaMoli .puesar sJnxxo.1u!qDH ueuasroNpu? (g.0r0rzr 's.r.900t) erprearooJ olpu?pr pu' g 'S,1,000E) ,",i1itl'jfft$'9Jl13 e{EI ot ls?a-qlnos(g,g€o9t t ulqn,tlr?au rxo{ spualxa(.€/ erntrC).uounqulst1J

tuall 'l4 g 'ljJqn gN '.oouad '\ro1'Fo lHJuEd l8t 'op^xog AJo Joordllqq?u r? :(law) s ahl S'I g ogl t* 0s :(HJugd) pzjyd rBou:(cgJ) tdnl,aits 'ssorJ[.] \gL !r'ururu?J'S'W ,2961 ,uua^\octes w ureqtnosJo g lrul 6g]l{x E9'.lDgJlsduttrld tdos 6l Jol1A [ru)t :(HIUAd) taqntaN ,ururpuo1ufuroJ .poU o8roC '.1-IgI/llI tl lru I 9I9Z'tr?unx 17 JoS turf Sfl iu, Ot t.Ltr 4nI4l €rJ Jol[N,,\[ ru{ 08 :(HIUSd) 19€]!nW.9 g,a,rrasaXlcog urreJq oN:(HJ{gd) iuotp.tow'I)'N,SuttoLlo rln 6LZ|0L .u ,t .16 A\ [rx)tr] t.t:(HJUSJ I IgEI DuplDg'y3'SutItnqEIIngJ?ou :(Hluad 099 Dup.tug.u, ,auq€qrcI :(HJUgd) nDuae'Il'^,rH urolseg 8ed 'ur?p€tuntrAlI9Z-S lear3'[ru{ 068] ru ZtZ:fiJXAd) LW naftDgJ.g :(ueas0g re^oruo4 paloolos) vI-IvUJSnV NU;IJS1Lqpaututoxa suaunladg '0,4d 'I aeoJ€oelaur,{qlue[ulsnv urolsoiA B NultsiaVoL 6, No. 2 (1988)

Fisure ?2. Distributionol Pimelealanata

_t ,\^ ^l l' I *[tt4 l \ /l - a aa ! B! I I a F oa7 I I I

Figure 73. Distribution ol Pimelea brevifolia s\bsp. brerifolid a and P. brcvfolio stbsp moddta L

_t _/

PE

^ a"i".ro.fr Lt+rra. I t TAI 'i-f[I ]-J) I I I J Figure 74. Distribution ol Pimelea brachyphylla. 'pesruSoJereq plnoqs sorcods auou?qlerou roqleq^rJo ?xBt rrJrcods?4ur olur pepr^rpaq pqoqs o {qdtqrotq d Jeq]€r4/r' eurrulolap01 sluBldJo dnor8 srql euru?xeol pepoeusr ^a^Jns plerJ peJrelop v otrErpauralur oJ?ropurBrrroJ aql pue so^eolsnorn?[8 u?ql roqter uooJ8q1l,r ]nq luelJ?^ puocoseql ol 'lla,!t relruls ere ,reJE ^.ra^IUEUB^ lsru oql qc18u suournadsguros odroqlsuo^€U Jo 1s?A 'ouu atrnbll? 'sjr?q 'elBllrc-Euol Suolpus uoqs Jo emtrlu e sraroql eqnt Frolj oql uO eJBstcBrq reuul oql pue oJrluear? se^sal pu€ sljBrq EI sranaolJro8uol pue se^?olsnoJne[8 ro8uol qlr/r\ qnJqs Jo qruqsnpunpare u? srpug dn8unlu8uef Jo lsBo BoJE eql ol polculser$ tu?Il€^puocas oqJ 'e[rc {3el ueuo sl3?lq touur aql puBpeqlool ^[ecldoosoJrrur.{ll?nsn el" sa^?olreddn eq1 pu? slo?JqoqJ oqs ,{re^ ? sglxrlouosoJ? 'sJBq qcqi osr?ocruroJrun JaqtEJ e^eq ^ ensnpu" Iprus al? sJa,{ol}sU'suauroeds plo ur 'sa^?el uaer8{Jep .{Jan.otumoq tnq qso4ueq,[ uaoJ8umpeu al? qotq,{{ Sulp€eJdsuoqs sEq 'stuels olerlsordol luoqumcopSur^eq sourleuos lnq qnjqsropunpodeqs-euop e SururJoJ U 'lrqeq -eEu?J ,{["nsn Surp?oJdsa s?q slusrJ?^ asaql Jo auo selcadsaql Jo ued ujelssoaql ul sal"pouJalur aiuoserdoql JoJlou ojo,r 1l soroedsel?Ddes s? pesuSocer oq flp?er plnoo Jo 'edjoq$ue^?UJr qcrq^\slu?IJBA lcurlslp ^re o,r1 are oroql 01 utEBi[ tuo{'oEusJ sll Jo u?d urelso,r aqt q uox€l elqeuel ^ro^ e s Dlfqd{qlDiq Daputd 'peslru'?oxer^lluesod sV

'plerJoqi ur uo{xruocssal eq 01'^pnls sql ul ue{?uapun plorJpollunT eql ruo4'J€odd?osle pu? stu?ldpnxaslq ljo,r 'snolJoorpou,{3 u€ql suorlcolloJunuBqroq oql u luanbo{ ssoloft slu?ld optrlod ^p]loJun oq 01 sJeedd€sobads oqt 'aEuers1r ur ,{lopr,rpolce oJ uoeqe^?q suoruDedsapual sV saroN 'slsuauo^D'4 'saylug[V pue nlolnatq oapwld o! 6eso13

'Jeqopo-^lnt pouad Suuauolg

'saslnoJJJt€^ASuole Jo seIEJ 'surcld Jesupoprorol sormlo{xosspuBlqruqs Jo spuElpoo.{{€lleur ur IEIJlo sdoJqno Iaeueqde'eseql 'olualq '^BIo'lo^E.r8 'pues f{ooJ uo Josuorl?ulquroJ pue oIu?J? ur pepJoxan'loUqDH '(g 's rBg 'S,6lo€€ ,os,,€zt,s€o[€) olrTaBrslol lsee(g.lz0lll r) ur8Bl[ fteu uorJ spuetxgftr oJnBIC)uotlnqv$!(l '3^U 'J r9z e?ec?a€lo!u.{qluPIlsDsnV ulalser\A g NuytsiaVol. 6, No. 2 (1988) 2. THECANTHESWikstrom

ThecanthesW rkstrom,Kongl. Vetensk. Acad. Handl. 271 (1818). - Pimeleaa' Thecanthes Wikstrom)Endl., Gen. Pl. 331(1837). - CalypttostegiaA. CalyptridiumEndL,nom. illegit. a. ThecanthesWikstrom) Endl., Gen. Pl. Suppl.4:62 (1848\.- Pimeleasecr. Thecanthes (Wikstrom)Meissner in DC.,Prodr. 14: 496 (1857\. - Pimeleasubgen. Thecanthes (W*sIrom) Gilg in A. Engler& K. Prantl,Nat. Pflanzenfam.l1l, 6a: 24 (1894\.- Banksiasec! ThicanthesWikstrom) Kuntzein T. Post& Kuntze,Lex. Gen.Phan. 59 (1903).Type: T. punicea8.. Br.)Wikstrom 0ecto, fide S. Threlfall,Brunonia 5: 118(1983)). Annualherbs (possibly rarely biennials), usually erect, single'stemmed at ground level, usuallybranched above, often reddish or maroonat baseor on majorbranches, hermaphrcdite, glabrous.^ttems sometimes with a stringybark, fibrous beneath and hollow at thecentre when dried;branches often arising in pairsfrom oppositeleaf axils; nodes not protrudingabaxially beyondthe petiole.I-eaves opposite or subopposite,decussate, shortly petiolate ' Inflorescence a headlike condensedraceme, terminal, erec| rec€ptacleconcave; involucral bracts 4, appearingto form a continuationof the receptacle;pedic€ls dorsiventrally compressed, articulateat apex,the apicalscar left whenfruit is shedbeing circular or elliplic' Flowers whiteto deepred, completely glabrous or rarely(not in WesternAustralia) with simplehairs on baseof floral tube.Florul tubeof 2 portionssunounding the ovaryand style respectively; ovary-portionfusiform; style-portion nanowly cylindric,very slenderat base,broadest at summit,the gradualexpansion occurring mainly in thedistal half, circumscissileabove ovary in fruit. Sbpots4, the outer pair overlappinginner pair in bud, widely spreadingin flower (in WestemAustralia). Corolla lobes absent. Stamens 2, insertedat summitof floral tube oppositeouter sepals.Ovary 2-carpellateat first, effectivelyllocular at maturityi ovule l. Sryie arising laterally just below apex of ovary, filiform; stigmasmall, papilloseFruil indehiscent,dry, enclosedin ovary-portionof floral tube. A genusof 5 species,extending from the Philippinesto northern Australia, 3 species occurringin WesternAustralia. Notes.The name Thecanthes is deivf.dfrom the Greekwords theea lenvebw, sg.c\ and anthos (flower),presumably referring to thesac-like concave receptacle and attached bracts enclosing the flowers. Key to Species

l.Flowerswhite or sometimespale pink, with pinkishanthers. Floral tube9-11 mm long.Stamen filaments longer than sepa1s...... 1. T concreta l.Flowersdeep red, with brightyellow anthers. Floral tube 4'9 mm long. Stamenfilaments shorter than sepals. 2.Peduncle(6)20-85 mm long.Involucral bracts broadly ovate to depressed ovate,apiculate. Sepals 2.5-5 mm long.Anthers 1-1.4 mm long...... 2. T. punicea 2.Peduncle1-7 mm long. Involucralbracts narrowly ovateor ovate, acute.Sepals 1.5-2.5 mm long.Anthers 0.4-0.8 mm long...... 3.T. sanguinea l. Thecanthesconcreta (F. Muell.)Rye, comb. nov. PimeleaconcretaF.Muell.,Fragm.5:73-74(18651.-Banksiaconcrcta(F.Muell.)Kuntze, Revis.Gen. Pl. 2: 583 (18911.Type: Camden Harbour, Western Australia, date unknown, lS Roeftolo: MEL). PimeleabreviubaFawcett in Forbes,A NaturalisttWanderings 516 (1885). Zype: Mt Sobale, Samoro,Timor, 28 Apr.-3May 1883,H.O. Forbes3828 (holo: BM). :,;';ii:i.w,5iff js',*yj t"::a:t::u!'ltnlr ul tu.{rid{jqH?ir";';?,T,?^f * !:1,3,x.fTi:rui'"^ffi?Kt,::,uea,;'';t-.,;;r";;'.;i;i,;, .Auoler s s ol,ql d' 6-e"'e-e s t" uerp utruS.l_/ 0 uru q/ i1:jl{ln $*i*tfil'##[{{ff3d;&ff:ar"i{,,x}ift ;,?id;i#;vf

*Ti$i!1!11i'itT,:'T;'"ti'J,1T."J'H1,.',11,'3'tru;:U:;iil;f:u,roJq,\rollo^ al€d osl{u3qlo ,dn le llliXil*J;i* t$?rd*i$;:*i'd,lxjl*:,T,r-+*liqx"J,'drt##$'its1i]]v;f ,*itrt J?,l,gi,sLi;i?ilEg4"3i:l'",'.5!5,fi,,ii,"?"1"_5:ilfl:iq"iiliJ:.JT:iffiu,,j,ffi:.#e;i;11b,i

17

V t9a apaaBiploul{qI uellprtsnV ulalso/d,a,{U NuytsiaVol 6 No 2 (1988) 264 Frowerspt:rewhiteorsometimesi;l5ltrr:i.ff#l ##i;1,]^r:,|\:3fh",H:3'iJi1i; ilfiI 3r;:;::;i"T?"IiliTiTii*il'"tli,iirlllr;l'r',:ili1ii,liLlill; f*' u'o*iio uucl r +s x r 3r emm 3"1J;1il,T:?rYi:i':,'lf;l[3'J.;;;;';;;;uii S of settlement'Kalumburu Mission' ZE I{ ftSoecimensu,:ifu,:*$:+'tali$iirsl$tr{q{qft*61'": emmined.WEsTERl'l- AUSTRALIA: i,Fllt{ru:i:i'i','ty;iyff^l*;,l;:r'x*tl+:{str?liH:.s."T'*ir'*:ll: B:t'JfiiiftT"sff+/#lu'n.ll**lun:u:*',ff"',1l',#?f JIl;'"" (CANB,';;;;;;r;t""s78,80')occrrs.i''h'--I-":llll::3lfDNA, NT). 3l',llJ#i;J5L:il""jd'J."i,X'Tut',i':',i.'1',h"4:illrullntii{ifllilffl'+t:;t;illl;';.'."ff,iif ff $:f;""# EiTff:"tff l.9:i ll',d:,itl,iil11,i: rry;::l:,#':l;,'; speriesg':3;ft inIndonesia' ff19:',iil1 shown i'"ili,T+'J#:i':JJ'"j"li:il ffiit#il.s:*::1,**: (1 e60)' il"ffi:i't'8"d11'J'#il ilili'r'" i i s3at and Ding Hou areas with Eucalyptus woodlands' Habitat. I'.ecofied mainly in latedtic FloweringP ertod. JanuarY-June' punicea' AfJinities.Closest to Thecanthes #ffi:'iu'fi ##,'a[';.t#"f,:i"n;"j,ff:m##ili,t;t'"*ff;+ *#t$r'-ra:X"lt;i*r:r'r:l*i;.'3'i;i:?"'# Il';;;;';'" ;'*' pimeteapunicesvar qr,r/,q1 ni*"xxii",jffii$ffi*,r "yff",f"ij;"tr#,$jfil,!*."!] [*i-q#+i;:il:'$ ,"'H,l+ffi *#.tfil"fi;;;;rv;t1-011e:s:::i:il,*itii*l'lg*U;l;*:,1,t;lll,fl *rili":l:#*r-u*f*l[q-$A?',;i',t^g*,,1:-;ll$t$illft #lpl;$n{#;iltri'"t{"trffi #i{r+**,r;*;**i**,i{l*rl1r.,#'*iftffi{tr+j*t*;;""'.l"k',r,r,I*,}ff

:il:*i::r--'*-tt.itt#*llitilt'""ry,;;i$lilifto act mm long,narrowlY obtuse lm$n #{r*f#d;l#[ll#l*?l lgll"**:"ru':"t' [-

ffi.f#x{iiskx,#ilryi::,?iltffii1.sifl,ffi}'##:i;ffi ##d1*,iy**iiuairffil?{:{:iW^{p;**'*;nFt

uauaauw-z wotJuti\Brq x)uoruels .J 1, r :(0lx) ra^{ou,g :ulars Bu ""^r,.;r*t,W:;f J;fr:},i,}lry1p6rf

s9z aeo.eoqaulqJ .a,{N"l uplle4snv uolsalA I Nuytsia Vol 6 No 2 (1988) 266 200seen):.9 mi km]S of Bagheloj, NoRTHERNTERRITORY (selected from over [15.5 (LD' G.';;;;;;"iL i74oiiRr, csc, Msr, NSW,PERrH); Yirrkala, R't spechte05 ADW. BRI. CANB, MEL, NT). the Northern Botanical Dstribution. (Figures77, 81.)Confined (in Westem Australia)to Gulf-Lake Argvle area' ffifi;;;";dtc";;-ir;l;;.j itorn th.'Fitnov River areato the Cambridge Territorv ;j;h ;furd from Kalumburu Mission' Also occursin Northern ground.where specifiedthe locks Habitat. F(ecflidedin sandor sandyclay, often in lockf havebeen recorded as basaltor sandstone.rarely wlth gravel'

Floweringperiod. Flowersrecorded all year, especiallyApril-July'

Affinities. Closest to Thecanthesconcreta.

f' "i-a vh a. oI. oa G or C\ t \ t \$ a

lig*. 19. w.rt"tn Australiandistribution of Thecanthessongutnea' 'sesJnoJJolat\ qllr\ polBrcossBseurlaruos ,,{elc ur,{[el?l lo sJlos,{pu€sut pepJoced.pjlqoH Hi,##,"J::ilsi';nfi"xxr$i+d##j:f;f{*,;::l*$Jrui,,.,.,,, :(sNVr,l*t)#;*,,?Y+J3i#',#f,;{il.ff;f1:;T,?1,qi##,}fd,+#jl 3;,xg.i:,ff:,ti'Jtr&llll#i,iil."."",i'#i1i,;d:?'1fl^gii!K{ff K"ili"J]I^ r,#ft$Hf,jft1i.ffi ;'f'"1i"-"iitril*Fpidi,1ilffi r jlitfti'il:?lit;n,',i'l.i*itr, fr;'#:n{g#;fi}}1fi.l*i #l,ti,flffidlirr.,,"i:.ff [;;ffi',-",,frl{fillllf; j,',,#iliilf fi!1ii,l*i,3il:,,'ffilfrri#i,#*;;,.,;'**f ffi:,[:1f t',:,il{ffi ffi1{',t,;,.d-ffi*#*,:,tr,itrJi*#t*i*t jd^fi,ili"fjfr* ff:lffiif# j*'Jfi a*l:e #rgdilii*'#rii+lrii-::1,'.xiff #iTffJ:1":r,i:i":i..,#ti*ru.1l;r;"**iu.,*x"ffi#*nftr$:tff?,,,,ir :i*tffi+i:fili:s;:':,:ail;:;t;il,,l_'?,:?3i,Hf yH#ii',ilnit:!;,#:ji,*,ii (.9/ arn8rJ .^ou .quoc .o,td .l) (.[enr{ Batrrn8wssaqluBraql .t #*x'''r,**'la-ifrru#*l,i,**i.t*fiffi[ffi :lo]ipr-9r.r?"r3rouse^{ rr pup,.orru,t, o^r i,irElJl',X*li?::"1*{ii""€ord_,qc€e-od^ru,{s B.p€_u!?ruoJ re.qsed,{r oqr esn€caq ioiror,"r "qi',n u;,il:t;',isfi lY,\J"""::i1'.y t[];ijin#g#i,jrft# WBruorJ pr^,\ofloq oeoaeaplou,{qlusrleltsnv ulotsal\ .a,{U,l.g NuvtsiaVol' 6 No 2 {1988) 268 Flowering period. February-August'

Aftrnities.Closest to Thecanthespunicea' tend 10 have smallerflowers and Noles. Specimensfrom Northern f erritory,and Queensland ,tt.ii-i"""f r*rt bractsgreen on both surfaces

concrcta Figure 80. Distributionol Thecanthes

punicea' F,gr" 8l Dt.t,rb"*" of Thecanthes

e4' Ft*." sr: D#tb"tt. "f Tltecanthessangui ,{rs?os? ,{F?au ,'i:::#fdli;?:A:,;J:-;;:#ttt;ft ?:.1 :: ry"s:p touen'DuaDplapH pu.- ^r::'-"i:r{ry-#;fr3j6r,1,.1{.lp"i:$:l*}jli##fi }i},,i.T,#6*[ fu'#i'""j$,;l:}jif,,*]#r :.",:y"'i,"i;q,i" ;.;;;;;;;q sorrrnupuoJsrperq',rns fiffi#J;",:f i:$,Hf*1ii:1"ffi^:,1"il11*iiHffiTJllel1mvaqinei'* ;n,.#illffi',?:i:#ij'# j,11i,,Yiil,r;_ii6:ffi i#:f:ilii:fl'i'fr:';itf:li *i'jtr*,Tfffr{*i*#ili.iffi iiritrft,*d,+i#iTiifr itrfr;ti'}*i# Iti1rt**fifrffT"Hl[T{+;T",1,1.:ifr ';#,"dJ:ffiili",?"jtrr,}i*# tr;Illitfltr*,r:"*'*H,,t,w*,t* :1:[frT]:r#ffi11ft'#5#'slitx'ifi $fi:'J ffii#;lE1{;i5;;j$8fi;ryiqfrri,Ji"'i,*;g,;fr '*,+"-i-,f#fr;fl*i;ft,*;*li'#qfJ,,:i,,ffi }iltii*,,iif,; *f,-ff#:;;#'rl*t,?,rdlllidtlf;#,i*ll"xHtrffi #jsfrt .t*-*;f#*i*+,**'f:d#*;yliii***#Jl*ll'ffiq$p 'auqdDe :l':?i1"t;'iff3j'i::#1":i?lil:f'j"'.'.::"p esoq,^'Dlwa ouszut4 Eurpnpur '€Joua8 :il1ff:'Jr#,i:,8'l3i:ig3ll' #ou',ir,.,e,";*,:is;^,-dlj ;;;;iilJ",liiH;l ;ff :i'#'"TjB?xTJi3;x'",1;l;.1: il;1ff1"r,Y,",*T1^.,111".1_".u^1 e!r+ '(0861) qcns 'oBeceoBJotu[qJ 'r:r--' lrrvrrv] "'v+ Pduuoj eAEu ueJ il",#'s3]i:Xffi,1i11'#:i,:fl:i,y:i1t;i:ueJpjfnV-er ::rii:ry.;'h:i:ff 1#,1 ;xfit Jo liti:3leql s? urore?oc?oerau,rq.r. roeroue3 ""l},tl::tffTl;':'ljiY-":":1T?.u se1p"re,ii;op jloql ueql lelsoJE .t rgl'v /ruuLudul]"J?oes s?JU3JeJJIp .suoseorH?ii,:J::H:flo HY*,i""filuaJEre ":e 1i':,1:.1lxT*ffi;;fi;";;il,i"#o-,,Tji1"l3lnHflcrrls J'il]3ql *'";:::';$i?o.rfrroJ por?rsuror ti] J,?';:'#"r"o riiT,3l,',#Y,lY:"11y:-'113-:-Y.r.Jnp; ;:!i':i:"""' igfi;lifernpsqe :Ti:i|:jl,i eiri'rsir.} aqr3;}i,::ou:l#,;J;*:':llf,i:,:yl,lg.*,:*r::6"'ii';:q,i;:,i;j!:T'il[1i1lj#ju00q o^?q uorsr^er ff ffi !::,11 fr,ilfi crurouoxBirripp ,riroii#i.in a,,"i"iiii "i{Hrl};r-""r'"^ uoJssnrsro

o€aceeeJolulqfu€llel]snv ololso/t\ !e,{U,l.g NuytsiaVol 6 No 2 (1988) 2tn ilil;if:i,,i'.',"JJJl"Hl';';z';"-^:;2r}::n:#::h:!T!:Ulljiitilil'#iiil11triiilfi ;;;rtii-qr*l j";,tl*$li*6#*::tl"mlli:""i:',:1ig'"Yi'i'"* sepals,no corolla lobes,two or ol ",ftTfX*i'rH,',-;lt'*,11ilf,:"'"'Jl33tT':,::I;,lfr"'"il::"j';ifTi,i3f'Tln'i::U! H[*r.i'f;i,'',#j"#*::iil**lfe,li"",rffi n,'s"#'ff ft'*'ff'T#: not bccurring in WesternAustraln' ,xi!1i{'iiFl*:il#:riil"},:i,tfi*miffiffi:i::ffjlf l{x;jdf l:t :f,J:lf J[x'.",'#3iiil:Tl:T;]ffi";':'4,i ffir,H?.{ :l'll!Tfii.'.f;:1,113'li!LT"'"'ru;.;';;t-;'iift ,i]:dj:,':}Tfii,j:,,t!,fi,,:;}H{ llji':,,*"#*,ff 'ff 1i[fif$,;it*:i-*t':'**'i-' :li:'*t":Jilfriti:11tT'i*Tl';;J**iir,'-",r''n'vp"r*T.ie."r"pfi;;',fit1'1 il;o,^;i",6-pime,einae-,ry;n::l;:3:::*;1.,:'*llTffsueeJstt,,r,ru1t"+,,glulp;rj*r*,,,,,;,11i,xit** lgl;Uillffi:l1"fi:*d::ii!:*iili ffiifii{'ili{T#*:iilTili,','fl X*:t?TrfHiTf#:,*[?ii,r'ti",iiinuin,nn to* *':j: Futurestudies r hope ?:,?iljXil}.J.ff[:f]i'r:T]fd*",fft ji'J1t'.3,."; neeior tu'tire' sttdv ati Pimetea tehmanniana' Ki:#:'"?^:I::#i,'ffi;;;;tesi?irrri*#i:i:{t^{;r*,uw:rx*"'.:Tii'clli!:t1?::i*iv:'i:'"tr :t'#lst#li"i,?it$-lrf*i+$:f distinctspecies or assubspecles l l*+ri:l+l*lg;;*+rylr;; whethertheir.treatment here as separate speqes in southemand east".n a*uuuu io-dti"ttine *il1f';$#il*l;n:trj*?iilY;$':lx':H"fiii":,1'l3i"iil;il?ll131""1:i'"?'Hs#:'lf; the infragenedcclassification presented nere' InadditiontothePmtcrantla;d*;xilJfJi.trf;ri?il?"Fit"':t#'i 5tJ;ifiiffi i.''iio'vuna :ffi:fifi,3H':.ftr"iliilit"il';;;i;iiandrhecanthe',e*49:t;"i'r;;;fi :"'-ldiii:i r'#tft--'fftT${il'""'in,",:,':sHrm4iip;1;*'{13"i?r3fi two.occurin NorthernTerritory by oneor bothpapers or tnese"ta*axa'a['-a'-vti-*nuttO.' in mvforthcomms uioit. ott'.r,ln'*rt"rn -uinl#d-'i*i:lq*J5t"r-lllldescribed treatmentof the familY ThYme l,{r+yiffi;b{;##$,:d#-'##ii,if?tf,:tilitrftfd#',l*{iiir:., ll*i#';? ",!:r:::*'-3-oq'.ii-1"' ',ultn'o'uii"tsi[',"liiilid,:J:;;'t!,il,,1"'" "0"" ,Tl"eTl"isi$"",nTiil"*tr[H,,ff;'lif-?J'Jjs;,:,il,;a:#,i:iii;affi::::::#u:l:i"i!ff,:!:,,13ffi1,* ffrtry""r.Br',,1*f1**i#i1:T;#f,Ti#,*,i,:'{i,"j sl;ii,ib#fi}!#i,#,'r'j# txifl{ {,w K;3w:}y:"il,:#"tr,E1fglr#?iii#i\ {& lf;?,w \; ;zir;H!;i", #i:_=# :*;T#.ffiFJr#ilii::,'li:i,1!yf,,u91.',"d4,uJ,i11or;\"fi1i',,1,'J3lfr1,:#,;a,l:1w:*Xirii{i; iilifl:{i*J*i'ilt'*r"1f #ffitr}$ili:rfut"a{# Hi*;tfh'{TLd;*Lijt',,i*'r 'Yl'l?J'#l!#,lp:#;ffij:"{i"i,f",:,r,",,i,"H;,jfi:ff"i"#i1li,',ilii"q *i?j:lL"q';li'jl+llfi,idil,'i:,,,ff:3il?,y"".tij,? ,1,##_$f,ii;,:t,'f *lilt,m;n*nif lJ;g,,',:#U4:::,1;g

snoh d roJsB ad,{r'12}gl) g rzt-l euourod .uuv .senb 6 t8l ll xe[xa eJrcwa1,iDuDDlnb"' ,r"n o100*"p '"ffi o"'"liiL"J',il$g;:i;;'il#;#l"J,"Ii,fil:.*il1;i,JTJf,U'-'""iliJ*:11"#,f:':;:!:;,Hr;::;n# r(.c uopor'cored,{l oN.(rs8r) zos ,iFo":pJ:#H:j:,lJ,ir!,Nf,:{;f&r,3!i;i:t;:Ji;;::";:,:#;:y;}.,::;E 'ola3lfl ,ui";rrhfitf;i,3;eix;,ir,y!,',fi1iirjl?;jf,;#it#\ff.:,,"iffr*^uru;iiiij;i#,fre^ Dvr|qu! oataa] or;).tl 'u^\o.o*narsp'erl'rsnv .(€l8t)zl:9 urorsa^rqlnosrad,{J T'snvu ,rrr"d}3"tr};ff1::;t:*tfiwiy*l

ororlrqu!Daputdrepwrsqrparq(r/8r)ue*::llJ;,i,",snvrr{qruasxa uun,"o,,oiiii\i"i"i sulllodssrue ,{lluocdctv .rssrr./d qns.u,{s iy' ord,(ZSgl)I :,.ras. oH.^.U,.oure( xe.uoH, 'Dttol!1roul ;#:::;^yf:"rLfi te^opqdaxo.Drru il qns .u,{s ord ,(/Sgl) .Jpold..JC SIS:}l urreussrar{ xe uunJDlloltuoull oapatd ueqruaspup ,ts8l-rrc 1et8r) (/s8r co ,russrrr^i.ttssrt u.a".rql"u1;"ift.!r"J#iofinir;:n;t|ifi:J i$l:i;;:il):f;,;:;##u.i:,lH,,ffili?),ixiiiiJ#ifl,;1,is";i1,1?",,T:'\\:I+,i:tr;i:il;i\];#i;iij 'uatxDq.Je^ .u,{s ,(/9gl) on)zuqut A qns old : i.JC .uunJ /0, lpord ur taussler\lxe atxoq oapau 10981)609 :e lpg l.roH tddns.uopnol stulj[opapwl.t 'Dloqdato,tru oapw{ qns.u,{sord.(rsgl) :92 'roussrol .Ileny| .J 0S€ eeeln\'l l xo saploaputld D1pp/uaqrsv sxBI u?rru4sny uralsa/!\ ol palddBflqsqord ?pnN suluroN IL' a?orsaElau^qluprJellsnv uJalsold .a,(U ..J.g NuytsiaVol 6 No 2 (1988) 2'72 Xtr"Y:":T&\T,*:,yif:i:,X:fiil"i11.?!;"1'3f,i.:1"',TJ'i":'J.?tr'":x'*#itih*i,y;l* {;,?,ii\#"#" mrcrofichers nor'clear enough ior a Llefinileidenlilicalion fr;i:;':!#:{#,tr}ii"fi'ffi$"ti";H"Jl;"""",11:LtS:,1ir:Ti$.3,''xr"ii;,KfJ'.*'i:1,:!;;r';;,1 Gilbefl PimeteaDleissii \ar. gilbertii Meissnerin DC., Prodr. 141500 (1857).fypei Westem Australia, date unknowft. Type noi locatedoi G-DC microfiche. date PimeleasDectabilis var../rs/drs Benth., Fl. Austral.6: 9 (18?3)-Typer.King George Sound, western Australia, inkio*i- McLeon. Type not located at K and there is no illustration WeslernAusl ralia' Pimclea\ulphutco \a- . macrccephal|Ber,th .lt A u'trd l. b: l4 I | 87J I fypc Blackuood R;ver' dareunLn

I am very gratefulto ProfessorK.H. Rechingerfor nrovi{ir,r.gthe Latin diagnoses,Miss S.n. ilitO],i. irr. rustrations,Mr N. Cohenand Miss A.J.G. Wilsonfor technicalassistance and membenof the technicaland professionalstaff of the western Austlalian Herbarium, i".tal"g b, lfV. Creenfor commentson a draft of the manuscriptl particularlyappreciate paper' ttretime"spent by Mr P.G. Wilson checkingthe nome-nclatureand settingout of this iu* uir"ino.Utla to staff of the following herbariafor the loan of specimens:AD, ADW' irrl, eili, CANB, cBG, cGE, DNA, K, LD, MEL, Nsril, NT, NY and UWA This research was'partiallyfunded by two grantsfrom the AustralianBiological Resources Study' References B:ll$:i.31i3l3l: ;*,""liHiifJst:fff'.;.3liid#;.(|s'"i"lnf"':#s',#xl,Xili':'1"ff:iarium Research Notes3: 3?58. "rJ13111.;*$nfJifill€iit;;yii"',i"f.?:tT:ffii".::*fd:ili)rJohn,on &co : London l B:l#5fii,".tlti'Liidr"a"riitiiiii"i,i,rlti*1"tu1r*tf#:,*x'*..ri#,1?_1,,:,,iH.,*i.ills:rll3f'1"i1* giiiit'dJilf,1p,ffi**fi?*ff#g*ll,magrd,hil:'5xi"r"iirftluf;''# "*n:l*;35:391'39'7. A*if*m:ffi'T,1?.H"'51?ii,##'f;l3ll'.H"lirti"€r""runsderrhvmeraeaceenBibriorh Bor. "",i'jallir!ii. &t llibution a l,etudecaryorosique erparynorosique dequerques Thymeleaceae. Rev. cen. 66:ll0 161. -CeneraPlanlarurn Pan 4 rBeck:-Vienna) enoticnii.S.t . rl837r. ' Eliifj;"l.t"ll"rlli;"'ilrTi'ii:l'l(i&"i8i":,li#'fil*",'iJid"-Fro\re$andPranrs ofweirern Ausrraria t"STtt.llfib"ii\. "PoisonousPlants of Australia."2nd edn (Ansus &.Robertson: sv-dnev ) . - l:Sf::f;ff1ii2fflf:g.fg.",ff,l5fi.i:ii,5|3:.:fil.'Jf;"l;#,.: f:ff*rJ.l%?tfti#ill,,*",.on, ..rndex perropolitanus ,,r.Y.1lFiiT.'* ".r"r, c.A. (1845). Decimusseminum, quae Horrus Botanicus Inperiaris prinrer:penh.r ".r,'"".iF.Kiidlf'ili.'.".ii""ii'J]i,"'?,"tllT,]li'{,-1,*|i*qccidenrali(." rcovernmenr fi}ifiijlE#tiffftr#"*-,:ifl :'"flft]TI,",r+ ij'uu:jphtr TqlriTlffi mi Z9Z raun)I r8 .ut ISI'Z'I .9 I qpldq? s9l rersJoj ? tastof o!p!uE, I0( r4un) t.nrnhl J) Dud$auof .z )un){ fiJ|sv.Wl opunqyoi[ 961 tztunx (Wqyl) DauBruD{ a4un) (ltonhl .d) Z9Z .g ,r/; 9SI a4un) fllonw ) oplJun s9z tzrun)N lfiqen uoADtJ 6SZ raun) {..r9 \) Dtto[tMra 691 Jzun)J t.qluaa)D tttdttpDrq Jztun) fJB .\t DatuaBJb 661 .X) 9tr ?nm) { rg qpJisn8u, lzlunX l.ntnAl .!) srDtlJowuD ISI Z9T -... ez]un)fisluDqodtJ.|l,es o71m) {uollsll^l, srqllrJrll- .tJos 09r Jzrun) t9I {Jauu.ec xa 1o5y siuegi ralazj! rres t(e ezrun]{ | lpttj) slrpDtslDw .)xes t9l eauny. I lpual DudololdtH .;Jes ISI atluny r1pua1a?o11wi7 l:rls I9T .ztun) I r.ltueg)otwo1or74i1 .ttx 9tl eztun\A t lpull stpDlslrotl) ) str dzlua.r, (lpld3.) ouaotqfqd .q wnlp!4dtrlrJ e I9I'Z'I'Z'I'I ' I rolsloc c ? relslod ,rqrqg IIZ Dussror\l xo .llanlN .J sapoapwld IIZ .nen'i..CouppJuaqrsv If I ^lPqS,{rV eruequv* sroq'murosreoloqloraqerooc":i:!.f$iyillIll :li:#.:ili,'#'filt Jor.Fe:sauo8elpc ayauaEe.rg.q aqt JoJsrajtJl .snuaEflffH.:',ffi ro sJequnupj* ,roq,nnk*r! JuJESeqr ExslJrJrJedsBJJur pu? sercads eql Jo a.rqaq,{late.redai pr,r,i ,r" lrnrira,r*ls Jql J^oqE soue.relai "*qj puiiu"roi"i"*r"au sJuo8aleJ srueul?ort:i.1y:.i:l{g_:r.tntur 14uapr.brlrog r"i.r"q-idr"q";:;;; rn?u eql Jo uels eur rog sradurnuaa!.j urelsod\ lsrrersJ ui r!'p"r"r,p-, ar? erTersny u Suumrro tou ?xet pu? scrctr ur uaara-a:e iru{uou.,{s Jn prprnuo, sa{xeu IIV aBaJBoBIaudqJol xapul \suora^ISuo) reueqrourBers:pueroqBuruproue^unuaerour(qrlr,ror.r.litl;r:1,1"1,",p.",:f.,;l^"i.0, ^i;,lliiii,Tlit:if;t",,Hti?rfr,,-,.li"#illji*'.fdl3lfi*+"#,.,";bi*#n#if$if*,,-l,ffii;^.

qamqupd pJec .rrrlarr,(qJ rofl iou soloN snuotoqr uo,,l .r .lI aeaJ@elaur{qJ Jo v oq}u sorD"r;r{Sruuli,?1"; ff:1T;ffil{fi#:1H:i"Sr'$B5i,Hl[X:ffigg;6f,##i:?:J*T,"JH:i'f3#gli[:j:iYf'"",ff tt *q"t*1I;"f"f,"#',yf'*:Y39g1Y tJ:t""?.:?11.",*l::sNoluoso-u'Ioirrr'(r.s6r) vt'ff"uo"t"u,," itu&5il"i3:'"flq:i,tluturu,:wr*kb'n1l:y!ffi1dffi,li"tyj4h,1lm.j*sfuh#i1"ir#giaq1,:.:ri jii:"dm",i,i,j#lnrdfuff#f";b{Lff i 'd*l uli'l#rx prcv qler{'e(r{drJ .$tups,alrr./rr :'r"slrqal.so^,"sou$rfffi:J[:i}f$il!!S'iir&:![t'f-g.x":J,:iffi'"ftll+;fffr"i:lf",i';;#fins aunrlncorp roqn uoaun"",lrJ{r1",?l;1i;iiqtffj- ,i:i"^ j"""U"*ll": ,.qoyj6 *,unooi,a.ri:qr !,iiii lii j#"i# ;'S;lffi{i,fi l$TB3;',tr.''ffi llg, #},:f iff5,:*#jr:trfljtr*f*,"H,i{iii'ls#1iil'frfj"T";;i'}ryfi 1,ylf19.E".X..j,iilq-H:g: #,} ,jf iffi?aq"5-;1.i,[*ililiffiffi:qt j{!:i ."re,l?titeni 'azlunt '(eerreee.wtqr)oapuu pus (0"""slr^w) ,r,,?,,',rroursoln}cnrs lero[ Jo ;9fff"^g\#gij:tiS#)'l"r]81["#,";

o?eieeelou,{qJ uerletlsnv u.tolsojlA,ofu .I g (1988) 214 NuytsiaVol. 6, No.2 holrovdii lF. Muell.l Kuntze 226 imbr[cata (R. Br.) Kuntze 178 lehmanniana \Meissnetl K\ntze 228 longilloru lR. Br.l Kuntze t94 naxwelliilF.lvluell. ex Ben(h.rKunlze 257 micrcceDhala(R. Br.) Kuntze t57 /?eD'ost iMeissner) Kuntze t99 *tefaea (Meissner)Kunaze 184 rrvsodes (Hook.) Kuntze 222 ,/er:rsli {Meissner)Kuntze 197 punbea lR. Br.) Kuntze 264 roiea (R. Br.) Kuntze 248 sanguinealF. Muell.) Kuntze 267 serovllifolialR. Br.) Kuntze r52 spectablrrlLindley) Kuntze 236 ipiculisetu lF. Muell.l Kuntze t60 sil/phu-rcd(Meissner) Kuntze 203 Jy/vesalir(R. Br.) Kuntze 190 tilrcrolia (Meissner)Kuntze 2t2 trichostachya (Lindley) Kuntze 168 vilft/e/a (Meissner)Kuntze 187

Catypttostegia C. Meyer 132,I42, t75 a. frecdnti?cs(Wikstrom) Endl. 132,262 b. Hololaena EndL 17s A. Calypttidiun Endl 175 R. Chotistachys(Endl.) Endl. I5t', L Calyptridiun C. Meyer 175 ll. Malisrachys(Endl.l C. Meyer t 32,167 lll. Epelage lEndl.J C. Meyel I32.t67 anzusrifolia(R. Br.) C. Meyer t99 arientio \R. Br.l C. MeYer 169 brbvifolia(R. Br.) C. Meyer 255 c/t/yirbides (Meissner)walp. 152 drummondii Tur.z. 219 Ii ro l9l 'hypiricina%aci llo lHcft.)Endl. \C'Jnn.ex Hook.l C Meyer r42,t75 /drata (R. Br.) Endl. 253 lehmanniana (MeissneriEndl. 228 Ionsilloru lR. Bt.l Endl. t94 maiiocephala (Hook.)w alp. 271 menkeana lLehrn. ex Meissner) Endl 2t6 Br.l Endl. 157 miqocephala \R. 169 nryliafi i,tra {Meissner)Endl. ,dra (Graham) Endl. 271 nervosd (Meissner)Walp. 199 ,rcirrri (Meissner)Endl. 197 ,rrrtea (R. Br.) Endl 264 suaveolens(Meissner) Endl. 211 sulprrlea (Meissner)walp 203 sv/tesrrislR. Br.JC. Meyer t90 ,lrcrorid (Merssner)Endl 212 tric hostachya iLindley) WalP. 168 vr71ilefd(Meissner) walp 187 villosa T]uJcz. t96

Cookia L Gnl,elin r 42,l5t *griTid (Forster& C. Forster)J. GmeIn I42,t5 I

*DaphneL. 269 *GnidiaL. t3I

GymnococcqFischer & C. Meyer t3t.t42,151 L Melanococca C. Meyer 151

'dtupacea lLabll.l Fischer & C. Meyer 142,t51 e9e tnz'ztz'rnz )1aJ\\ef Dqnlrlaq ,tze ,z'ba rIU_loutu d\qn\ Itz rr r ei,,tlsr^sq.dsqns Ita e(U el(tsr^Jlq q)uag .qluegdlorlstguo.re^,aJ,uDJqUdu.)eA 09a'892t9a'sgz'\tz - 09z8st'Lsz's9z aAUUaussrel{) etsopout .dsqns tga'sszqra srloJhaJq.dsqns 89tt,I .lg .u 0^1-a.l ea'092'692'89t 99a0,z cbr .qluag erloJr^alq vtl e lqd,,lrlcerq .J6ler\{ uz .uua) elo[rq* 192.9ta'rn('6t z,6tz Sta.sr t Jaussrohlx. plirji) altg qsuauo,ri (,1,1_0Lf69189t DSOuac t9t €,t 9tr Ilenl^l I .tg re| .U eatua8ts 661 Jeussrahltoutw .Je,\ 66t Jeusstawlofow .re/\ 661 JauSStel/I1tpUouurtv p .Je J\ tL.a.i0; JOUSSIaWSrArsanlrr .J€,\ a)a00Z'66t S6I tfI ttI .U .lpnw rg ertoJqsn8ue 41.:ttt'sLt'nLI'aLt .l .n^ a tut trr lpuolltola.l llantr^l sueqcouue 0I a 602,802,Lje. s1a t, I'6t I '11rn11uopnol eurllD e"ourini!e su rtl 3 qluJB .Dasqns 69eSaa rsr,rtr 'qluJB llpufi DulDlolfrtd su ttt l ipnlr r f4.raisrJorjj rcesqns . . Z9Z'E' I qtll ag runpu tdt I D) De\gn\ 6 rtr BIr.) ru'torr\llll j raqtuo,it6 . uaiqns a9z'r€t'ztl dltD Dapardnl .ueaqn rou.sr.ry1 Ria9za saa,tzz t str cfr.r,I ruor"linr iaq)uoio,11 na, 8a(a9I't s t'6rt'Rb rrt r pri ii r.c,, a,{; s,{qcersodif5 roas 19trf I 'qtl]eB ealourd.lcos 01a69Z ZZZ'fbI rbt .cr ccl t.lpul) s,{r/Jrlslrrv .tJos ot. 6e.'sr|i, i i ; i i; i ;; ;.cII JIU { /JnI) pr8Jtsoljer\l.tJrs 6a; rAU SOIJLIIUet lH,IJOS ar?ltaLzzaza gotitt:tit.ert,stt $t 6 atI ,"u,,'.ap"r"1dr?l$ t,113ir1,3#il;,.:,8 .f reusslaw . :; 694-.SZ,a_89I' L9t t s I t, r.$ t.9t t.t € t .qluag oa pttt ! dn g lJas fJpug) a8ellEdl .tcos tst rtl .q)uegDtwDtDtt|tcl .tre\ qja,.bea Ilrnn rprr11\ttl tot?tb43 .ttx z 8az'azazaa'1Lt 8et (sI f,t.stt.ttI qtuo8.- J U'I9LI lra(rr,{ 3t eriaisollil(Je.) .r.ros ISt dltD Daputdotnv Da\ lJu9stew t9l aD4tuDtJrN.<_b stl teu.stehl ao ttld,{@ 89ttgt ttI reusstaw aottultquJ .E 891'I9I'I€I 1pu1 a8opdj \ 891'I9I'IT I .lpultpu.l {potepn , 9I I'I' I st|JblsuotlJ .p ua'ttr Ipng DuaolollttLt .J .92'I'I Ipug DuaqotalaH .g 'tpua .i 6'I \utorts\tt.IpuA J saqjuDrati Dalaluldnl e 0ta 69a'ZtI.tt/.0t1 6,I I€1 .los rauugec xa ,r, queg ealolurd Itl lc?I eueleqd* III ''I euuoss€d* Z"'"1'Zf I .zJJnI SUAJSaqtua aaa'z a€I .zalnJ. ggalsorrDrrl 9ta'taa'zatI .J 8ra ro,{et1l ? rorlcslC l^elpurl) sllqopads D^rhl .U, e9e .) J I.'l0 ,rso./ IIZ ._-. JJ^JW I.rB.U Dpldsltl tse tataUl,p twe\et1;t ttiostapuatl .J .l) ro ohl (.rg Dtbssnap Saataa'z 'atI ra,.ew Jg retlxsrfDuaop@laH 9LZ aeacEaulou^qfuprrcllsnv .e,{d . uretsoid I.B NuytsiaVol. 6, No. 2 (1988) 276 145, 19 1'193,193 calcicola Rye 14 5, ) 38-24D42.244'24 7-)4.9- 25.1 ciliata Rye 245-246.2)U subsp.ciliata 246.247'250 . subsb.tongituba RYe r .t39,t 44,r55,t56,t 57:158,168 chvala l2Dlll. t5) c/rydoides Meissnef 262 concrctaF . Mrell I 3 7.I 4 7,2 07, 208,209'2 I 0 dacens Rye 2t 0.211.2r2,219 subsD.cracens 210,211'219 subsp. elabra RYe 271 qinita L]JldleY 167,173,270 *curviflora-iivii-. R. Br' 173 iiiiino (F Muell ex MeissnerThrelfall 173 "i.kiiii,t i fr. Muell ex MeissnertBenrh 225 *derom Domin 251 decussataR. Br' 271 var. diosmoefolia [-odd. ex Me$sner 271 var.fiore'rubrc ?kfia1re er Jacques 271 v ar.'rcquiera no ?L€maireex Jacques 271 diosm4efolia L./Jd'd. 271 distilcriTirr4 F. Mue[ I 46,20J.2r9,2t9.220 22.t--2-7-l. drummondii (Turcz.) RYe l4! 1)J *druDaceaLabitl 151 *elnchantha F . M\tell 169 *elorBata ThrelJall I 46.I88.189,189.t 90 t9^3- erecta Rye t90 evlei F. Muell. L t 45.248,2 49 250.251,2-7t femrsinea Inbin. tJ/ *filiformis Hook. f. 203 '*flava/lata auct. non R Br' r52 R. Br. I 32. I 47, t 98,201,202 2 o3 ) 2D 7l floribunda Meissner t 36.t 38, I4 4, t sL t s). t 5q.lqll9!.i60^ forrestiana F. Muell 3 144,t4'ir. | )u. | )'.t;; ^l';an' F P;irel 130,138.134.I4 fl"iiai-iirF. itliiii. a rateex J. Blacktcarolin 142'15I *"midiaForster & C. Forster)wlrd' t93 -{aci[iflorudacilr'flora aucn. non Hook. l9l Hook. I 46.117'17'l '17I 'l8l Eaniticola Rve ihaemostachva 224-226 F. Muell I 43. I 52,154'154'l58 haloDhila Rve 271 hendersonii Graham 152 *hewardiana 151' Meissner t 32.t 44.249,250.252,252) 55 hisDidaR. Br' 1)J (R. & E rntzeL iw. lanata Br') Diels I 36,I 3 8, I 41, I 44,225,226'22'l F. Muell. holroydii 1J2.141,146.1 75.17 7.118,185 'I80 2-7^l imbricataR. Br. I tt2 f. boxteri lCunn ex Meissner) Benth 182 f. fiacillima (Meissner)Beflth. 183 l. biliseru Bf.nth. 182,271 vir. 6axteri Cunn. ex Meissner 271 var. c/irita (Lindley) DoDin 182 var. g/alrala-gruci Meissner 182 v at. ll ima Meissner r81,182,1 83, 184 var- lmDllcata I 79.180,18t 'r82 var. maior MeissnerlRYe 184.271 var naia lGraham|C.H. Ostenfeld '.183,1^8-4:18:..-:l var. oilisera{Benth., Dels & E Prilzel / lo,i 80.ts t . / 8 'var. ld1,l64tl dJ oe-iraeaMeissnet RYe 187 var. vi iferu Meissner) Domin 271 intemedia hort. ex Gmham * 142 laefigata Gaat1trrer 4.1 4 4. 1 4 5. 2 I 5,253'254.2^60 tanata-R. Br. I 32, 1 3 *latifolia 2/U R. Br. t44.228.230,2 32.2J4.2J5) 37) 70 lehmanniana Meissner 229'230'241 subsD.lghmannuna 2 30'231,232'241 subsi. nervosa (Meissrcr) RYe 234 v^r. Iigust noides B€'[th. '231 var . ieiocePhab Diel.s var. nen osa Meissner 145.234,237,238'241 leucantha Diels IIZ .qtlleg Loz'vo('Eoz'9,r.6t r.ht I qDqdaronou .re^ 98t 'gt'sgr'|It'oqi eii.rbt jeussroN e3Jnqdlns .IZ 'qrurg r^U illsrlrqL Br4rtlridns ilJussrJr\parottui Ijei 9IZ \116 .wyijt 6t( lrTrlrd f T' urusro4 io rroaryaut nn 8ta'glz'StZ .dsqns 617'8rz',Lrz9tI tt r suoloo^rns I l a 0az'rlz 9t r a,{d e^eu .dsqns ltulstJhl \uJloa\ens t9t .rl.tuJ€.rr .l n9r'tgr't9fB9r a^u(oroor^r lFnn Duqlrndr, s9t'n91t9t.89t .)"r:orf,iT5l:J;l 19t taI'091',a9t.t e r.rr r .Ienlt t9t Is r .C sjo8rlncrds ua 8€a .,"rr.,"^,u".,ro Ita Ste ^1.qluegrurfflr"Irn"t?#,. |1a,'tnz SF('gtz gtz (z.r,t suDlslp . re| 0ta." a,t.ztt ,{elput.I 6vl '9t slilqercads Itz'tt t'ztz'sb r r ,au.stowr,rrqff#11,ri{:l |9tfs/'zsr :dsqns r9I ts t'Est,asr . ertoJrtl(dres .Et o^U srleluoprraro BsIs9 I'r9 fzs|ff I I t dsqns Ita * Itz ,""r""^,*"u'o,n;i;h)grl?ffi )ausstew DloqdacoplJeA (tt'.t,5,:!tagta 91...srz. Lbzsr. I (..s,t.t t I 'lg I v. t.c vL(ut( rrl d easol 9tl reussrol .J .nanw ,9a l.xe,_u€nrs'qluog T , r3j*?a'.',^ ,9e xe .nanlt .goQo11a.aq.nt ISI'Z'I'I'I atz uraurcr (rorsroc c d,"r";xt HJ;i#;l ztt'861'L6t'96t'9tr lous$ow ?rJ.rrqlr3.Je,\ IIZ 9.(Zrza'tz('zzz 0z( s,t.( 6tt.1{t.att u*",* fH,.s}ol??i# l\ I f8l ,,."^oTTni,J,i,xxl )arsslaw Dao4ad+ 9tI .llonl^i.J 80e'L0z',90z's0z'tr l suellrcruad* s8l ,81 -- ney,,qrD!ar!1!^;1,9:;tiit.o lJBlluqf turus\rJ'1l)rrtl/i dsqris_ t9l .rg .U ell,{qdoDo* eta'ua ""ua .ro, Ita .euoeq *" .lJotl Ijl,,'il##,!, 661 xa nuolBradtriu t9I r.usst0l^l 'SO&a,1 IleJl)lqI erIJBueoau- atz . IIE''81 reusstrW DrI olirqq8 Je ^ 0tl 8Sa ,.rnffiW:lfl s9l .lpnl.jll JaussrewDlsapotu Z8I .J Dtljuoltsd .re^ Jeussbw tofotu .JeA 8SI . Ila JausstewDll olt tlDu!I re^ JeussrewDtDloa)uq .)e 091 .l \ 8Sr IJeIB f rr.lel T flanl,rt oqDl|.)e\, I lz'89t'l9I Jeu\ga DtD8uoldJe^ 09r'lsI elpudrao.rJrru.dsqns IIZ'291'69I'L9I llpJFrqj t:tJEtBf xr aeJ f 1:n11 jr erqe6.oqi.. .thIZ9I rtt.rb r .lg .U 0ta tLI'(LI'S tt I .1an;41epqclecor.rru IIZ roussrol4lxo .g eqlue.rciur 9IZ roussrow ,rqrp .ru^ laussrow xt .wtqs7 buoaluaa 0(z ._!ly_esx.o [ont! trlllaixow 0za'€0a Icslg I fqluog) eBolsorceu_ qoqdaJohDw Ita t6t'961 .qtueg\(nH 96I'€61 Drtotrtot.re^ 96I'96t't6l _ plour8uol d5qn\ '6€ r^U fllrnl roJ,{e.dsqns 66rt6l9 I'tt t.ztr tr J).tg .U 01atqz ztt erolJr8uol IIZ qlus eIoJ|4I* JJlrssrJl \, .uun-; o1o1lnii1 SII'ft1 J .d.ili." 9tl llpJiorqj{ IooH ra.uunjr EurJurd,{Li llrqe.Jeuulsn8rl* LLZ aEoceaslotu/tqJu€rJelisnv ulolse/!\,r,{X ..I.g NuytsiaVol. 6, No. 2 (1988) 218 svlvestrisR. Br. I I I, t45,190,191,192,193,194)71 - (F 2U6 vat. aeruginosa . Muell.)Benth. 199 t€rdt Scott 199 var. /otgitfvra Scott 164 rresioideJS. Moore I 6,219 Meissner I 46,2 I 0,212,213,2 tinctoria t 34.I J 7.I 4L l 44,I 45,168,112,17 4 trichostachyaLindley 271 versholFeltiiMofien 170 vesrimtr{eissner 185 auct.non Meissner vilrr&ra I 46.I 78. | 81,181,188,I 89. I 90 villifera Meissner 196 ,irlos4(f urcz.)Meissner /JJ viridulaLirdb, 272 weippugianaPaxlol I ThecanthesWil(strom I 31J 39,140,14,262,269,270

(F. 262,263,266,268 concrcta Muell.)RYe 70 Dunic€(R. Br.l Wikstrom I 34.262.263.264266,267,268,2 ianguinea(F. Muell ) RYe 262,265,266,261,268 *Wikst.oemiaEndl. 131,269 131 *indicaC. Meyet

Publicationdate of Nufsia Volume6, Number1: 23 December1987

L, DUFFIELD,Gov€rnm€nt Printori Westem aGtralia 58831/3/8a-750-L/450? GABRY