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Pliocene, Southern Spain) Contr. Tert. Quatern. Geol. 36(1-4) 41-44 1 fig- Leiden, December 1999 A new species of Cyllene (Cyllenina) (Mollusca, Mesogastropoda) from the Arenas de Huelva Formation (Pliocene, Southern Spain) Bernard M. Landau International Health Centres Albufeira, Portugal and Robert Marquet Antwerp, Belgium Landau, Bernard M. & Marquet, Robert. A new species of Cyllene (Cyllenina) (Mollusca, Mesogastropoda) from the Arenas de Huelva Formation (Pliocene, southern Spain). — Contr. Tert. Quatern. Geol., 36(1-4): 41-44,1 fig. Leiden, December 1999. A new species of the nassariid subgenusCyllene (Cyllenina) Bellardi, 1882, is described from the Arenas de Huelva Formation (Pliocene, of Santa Catalina,near the of del in from strata at Velerin Zanclean) village Lucena Puerto southern Spain and coeval (Antena), Estepona, near It is the record of the from in Lucena Malaga. first genus these units. The occurrence of other Nassariidae the fauna is commented upon. Key words — Zanclean, Pliocene, Spain, Mollusca, Nassariidae, new species. B.M. Landau, International Health Centres, Avenida Infante de Henrique 7, Areias Sao Joao, P-8200 Albufeira, Portugal; R. Marquet, Constitutiestraat 50, B-2060 Antwerp, Belgium Caecum has been described the last Contents during twenty-four et al., years (de Porta 1993). The the Salamanca team paper published by covering Introduction p. 41 the Nassariidae(Gonzalez Delgado, 1989) does not men- 42 Systematic palaeontology p. tion the of the in the following species family, present Acknowledgements 44 p. authors' private collections: Nassarius bollenensis References 44 p. (Tournouer, 1874), N. tersus (Bellardi, 1882), N. tumidus (von Eichwald, 1830), N. bonellii (Sismonda, 1847) and Cyclope neritea (Linne, 1758). There are also a number of Introduction smaller nassariids, which are probably endemicand unde- scribed, the most interesting of which is Cyllene (Cyl- Although the occurrence of Pliocene marine fossil molluscs lenina) lucenensis sp. nov., described herein. from southwest Spain has been documented for over a Strata exposed in the Santa Catalina outcrops are of century, taxonomic work on the rich and well-preserved Zanclean (Early Pliocene) age. The new species is re- fauna did not start until the early 1980s. stricted to the 'grey sands' in localities 4 and 5 of Landau Since In that Landau commented the distinction then numerous papers have been published by the (1984). paper, on University of Salamanca covering the systematics and between these fine grey sands, which only appear in bore- taxonomy of the commoner species found in these deposits holes in the vicinity of Santa Catalina and the coarser (Andres, 1983, 1985, 1987, 1989; Gonzalez Delgado, 'yellow sands' from Bonares and Lucena del Puerto, named 1985, 1986, 1988, 1989, 1993). Unfortunately, material the Arenas de Huelva Formation by Civis et al. (1987). outside the Salamanca collections has not been consulted Cyllene (C.) lucenensis is also known from the conglomer- and the more problematic or endemic species have been ates ofVelerin (Antena), Estepona (near Malaga, Spain). ignored. Indeed, apart from the six new species described The Santa Catalina environment was infralittoralwith than those around the by one of us (Landau, 1984), just a single new taxon of warmer waters existing southwestern 42 coast of Spain today (Andres, 1986). The 'grey sands' have - J. Martinell Collection, University of Barcelona, and not been worked on since their correlationto the Arenas Department of Stratigraphy and Palaeontology, Faculty of de Huelva Formation is unclear, although they probably Geology). represent a slightly different depositional environment, possibly at somewhat greater depth. Superfamily Buccinacea Rafinesque, 1815 Family Nassariidae Iredale, 1916 SYSTEMATICPALAEONTOLOGY Subfamily Nassariinae Iredale, 1916 Genus Cyllene Gray in Griffiths & Pidgeon, 1834 Abbreviations — To denote the repositories of specimens Subgenus Cyllenina Bellardi, 1882 referred to in the text the following abbreviations are used: IRScNB - Institut royal des Sciences naturelles de Belgique, Type species — Buccinum ancillariaeformis de Grateloup, Brussels (1ST = Invertébrés du Secondaire et Tertiaire), UB 1834, by subsquent designation of Peyrot (1925). 1. of C. Santa 4 of Lucena Fig. Representatives Cyllene (Cyllenina): 1-3 are (C.) lucenensis sp. nov., Catalina, locality Landau (1984), near del Puerto, Huelva, Spain; Arenas de Huelva Formation (Zanclean, Early Pliocene), 1 - IRScNB IST 6386 (holotype), x 2; 2 - IRScNB IST 6387 - 25; - view, 3 - IST (paratype), a protoconch, x b, x 2; c protoconch, adapical x 28; IRScNB 6388 (paratype), lateral view of 4 apertural lip, x 6; is C. (C.) ancillariaeformis (de Grateloup, 1834), IRScNB IST 6389, Montegibbio, near Modena (northern Italy); Late Miocene (Tortonian), x 2.5. lucenensis Stratum — Arenas de Huelva Formation Cyllene (Cyllenina) sp. nov. typicum (Zanclean, Fig. 1/1-3 Early Pliocene). Material studied — Holotype is IRScNB 1ST 6386, Derivatio — nominis Named after the village of Lucena del paratypes are IRScNB 1ST 6387-IST 6388, and UB 01135- Puerto. 01136, from the type locality; 30 additionalspecimens in the Locus — typicus Locality 4 (Landau 1984), Santa Catalina, B.M. Landau Collection from the type locality and from near Lucena del Puerto (Huelva, southern Spain). Santa Catalina Locality 5 of Landau (1984); 5 specimens in 43 the and 98 of whorl and distinct anal the R. Marquet Collection from type locality; juncture body outer lip forming a flared below, smooth within. specimens from Velerin (Antena), Estepona (near Malaga) in groove. Outer lip thin, slightly — is in the the B.M. Landau Collection, from Zanclean deposits (no Remarks The present species not uncommon sands' Santa Catalina. Some adult shells more formation name available yet). 'grey at are but constant Diagnosis — A small to medium-sizedCyllene (Cyllenina) globular with a more strongly developed callus, of thick callus in other shell characteristics. It also occurs commonly in the characterised by the presence completely covering the body whorl and spire, obscuring all surface conglomerates of Velerin (Antena), Estepona, near Malaga; tend be than those from the ornament except for elongated nodules on the periphery of these specimens to slightly larger the the body whorl. type locality. With its extensive callus formation, species attribution — The could form evolution ofthe olivid Generic and subgeneric new species a parallel to species genus undoubtedly belongs to the subgenus Cyllenina Bellardi Baryspira P. Fischer, 1883. and ofcallus formation in this 1882, which is distinguished from Cyllene (s. str.) in having The thickness extent new it from a thickened parietal pad below the suture. In some specimens species immediately distinguish congeners. Cyllene C. sismondae this thick parietal callus pad is more developed expanding (Cyllenina) irregularis (Bellardi, 1882), (C.) of the and the and C. (C.) (Bellardi, 1882) over the upperpart outer lip extending up to (Bellardi, 1882) subumbilicata Cavallo & This feature is in the new than from the Italian Pliocene, figured by Repetto apex. more developed species all have axial ribs and the the in any other member of the subgenus. Thiele (1931) (1992) same general shape as but not covered callus. Another mentioned as shell characters of the genus Cyllene the new Spanish species, are by teeth the inside of the Italian Pliocene C. protoconch with three whorls, on outer species, (C.) paulucciana (d'Ancona, and columellawith several The number which has also been recorded from the Iberian apertural lip a plicae. 1864), has of protoconch whorls should, however, only be used in Pliocene near Malaga (Vera-Pelaez et al., 1995), a quite and almost smooth, with no axial defining species and not genera. Cyllene (C.) lucenensis differentshell, elongated lacks the apertural teeth and columellar plicae, but these ribs. Nassariidae. (de Grateloup, characters vary strongly among Cyllene (Cyllenina) ancillariaeformis found in the Late Miocene of Description — Shell small to medium sized, glandiform, 1834) (Fig. 1/4) (Tortonian) the Badenian of adult height ranging from 9.5 to 20.7 mm, maximum Italy (Bellardi, 1882), Hungaria (Strausz, and the Helvetian of the Basin diameter 9.0 mm. Juvenile shells ranging in size between 7 1966) Aquitaine (France; with distinct and is closest the and 9 mm have a convex spire sutures a Peyrot, 1927) to new Spanish species, having below the and well- barrel-shaped body whorl. The protoconch, composed of two axial ribs with a blunt knob suture a This callus in the Miocene and a half, tumid whorls with a deep suture, is high spired developed parietal pad. parietal does extend the whorl and and smooth. Protoconch and teleoconch are not clearly species not, however, over body the shell to in the delimited.The teleoconch starts with the appearanceof axial envelop any significant degree, as new ornament, while the shell becomes less smooth. The first species. The is in the teleoconch whorl is concave with about twelve strong axial genus Cyllene widespread Neogene ribs faunas, in the of the Paratethys, ribs. The second is flat with the upper part of the axial European occurring Neogene the Mediterraneanand the Basin Fifteen becoming obsolete. The third teleoconch whorl is concave Aquitaine (France). recorded from West Africa with much weaker ribs, which all but disappear on the body extant species have been to The closest is whorl. Spiral ornament is also present in the juvenile forms, Japan. species
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