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Folia Conchyliologica N° 5 2 , Décembre 201 9 Folia conchyliologica N° 5 2 , décembre 201 9 Drymaeus ( Mesembrinus ) iris (Piaget, 1914), Colombia, Calí. Photo G. Kattan. Sommaire Abraham S.H. Breure & Francisco J. Borrero - A review of Stenostylus Pilsbry , 1898 and Drymaeus Albers, 18 5 0 (Mollusca: Gastropoda: Orthalicoidea: Bulimulidae) from Colombia , with description of new species . 1 - 79 Boussenac , décembre 2019 - ISSN 2107 - 7010 Folia Conchyliologica, N° 5 1 , novembre 2019 Ligne éditoriale Soumission d’articles La priorité est donnée aux articles traitant de Les manuscrits sont rédigés au format texte malacologie continentale, autrement dit de la ( Word , OpenOffice, etc.) dans la langue souhaitée faune terrestre et dulcicole. Les articles traitant par l’auteur. Ils sont accompagnés d’un résumé et de la faune saumâtre ou marine sont acceptés de mots clés lorsque la longueur de l’article le afin de répondre à une demande croissante de justifie ; des résumés dans d’autres langues sont lec teurs intéressés par l’ensemble de la mala - aussi admis. cologie. Ils sont envoyés par voie postale ou par mail aux Sur le plan géographique, il n’est donné aucune rédacteurs : limitation, les faunes des contrées les plus lointaines ayant ici autant leur place que les Philippe Feray : [email protected] faunes de nos jardins. Alain Bertrand : abela 11 @wanadoo.fr Les articles peuvent concerner aussi bi en des Ils seront soumis à deux relecteurs du comité de observations biologiques ou naturalistes, des lecture et éventuellement à des référents exté - inventaires faunistiques, que des travaux effec - rieurs. tués sur les collections anciennes ou sur les personnes qui les ont constituées. Recommandations Les articles d’ethno - malacologie, d’archéo - Il n'y a pas de recommandation particulière dans malacologie ou de paléontologi e sont également la rédaction des articles : nous nous chargeons acceptés . de la mise en forme en accord avec vos souhaits. Dans la m e sure du possible, inspirez vous de la L es descriptions de taxons nouveaux pour la mise en forme des numéros les pl u s récents. science, ainsi que les nouvelles combinaisons doivent répondre aux exigences formulées par le N éanmoins, il est souhatable de partir d’un Code International de Nomenclature zoologique. article de la revue comme modèle, notamment Si besoin nous consulter. pour la mise en forme des citations bibliographiques et la présentation des figures. Des exemplaires papier sont déposés à Lyon (Musée des Confluences et Société Linnéenne de Nous vous demandons d'utiliser la nomen - Lyon) et à Mâcon (siège de Cernuelle)pour clature latine et de ne pas utiliser les no ms assurer une pérennité des documents publiés. vernaculaires sauf exception comme dans une perspective historique par exemple. Directeur de la publication Comité de lecture Henri Nicolas BÜSCHER Cédric AUDIBERT Laurent CHARLES Rédacteurs S y lvain CLANZIG Michaël DIERKENS Alain BERTRAND Sandrine HEUSSER Jacques MOUTHON Philippe FERAY Jean Michel PACAUD Daniel PAVON Christophe PERRIER Quentin WACKENHEIM Antoine WAGNER Francisco WELTER - SCHULTES Coordonnées - contact Alain Bertrand, Abéla, 09320 Bousse nac [email protected] Folia Conchyliologica 52 décembre 20 19 A review of Stenostylus Pilsbry, 1898 and Drymaeus Albers, 18 5 0 (Mollusca: Gastropoda: Orthalicoidea: Bulimulidae) from Colombia , with description of new species Abraham S.H. Breure 1 , 2, 3 & Francisco J. Borrero 4 1 Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B - 1000 Brussels (Belgium) 2 Natural History Museum, Invertebrate Division, Department of Life Sciences, Cromwell Road, London SW7 5BD (U.K.) 3 Naturalis Biodiversity Center, P.O. Box 9517, Leiden (the Nether lands) 4 USDA Na tional Malacology Laboratory, The Academy of Natural Sciences, Philadelphia, PA, 19103 ( U.S.A. ) Corresponding author : ashbreure@ proton mail.com http://zoobank.org:pub:4778E4BF - 86D9 - 4 956 - B1D6 - 08CF32CCF6BA Abstract - The land snails of the genera Drymaeus Albers, 18 5 0 and Stenostylus Pilsbry, 1898, both belonging to the family Bulimulidae, and occurring within northwestern South America are revised and notes on their distribution a re given. 78 species of Drymaeus and t wo of Stenostylus are herein confirmed from Colombia, and are illustrated for comparison. Six new (sub) species are described: Dry maeus ( Drymaeus ) denticulus , D . ( D . ) duplexannulus , D . ( D . ) felix restrepoensis , D . ( D . ) iniurius , D . ( D . ) intermissus , D . ( D . ) luciensis . Drymaeus flexuosus megas Pilsbry, 1944 is now upgraded to species level. A lectotype is designated for Drymaeus roseatus montanus Pilsbry, 1901. The following nominal taxa are herein synonymi s ed: Bulimus antioquiensis L. Pfeiffer, 1855 = B. baranguillanus L. Pfeiffer, 1853; Bulimus hachensis Reeve , 1850 = B. virgo Lea, 1838 = B. columbianus Lea, 1838; Drym aeus eversus alata Piaget, 1914 = Drymaeus eversus subula Piaget, 1914 = Bulimus violaceus Mousson, 1 873 = B. confluens L. Pfeiffer, 1855; Drymaeus cantatus medinanus Pilsbry, 1935 = D. tusagasuganus Pilsbry , 1935 = Bulimulus ( Drymaeus ) plicatoliratus da Costa, 1898 = Bulimus convexus L. Pfeiffer, 1855; Drymaeus fallax chicoensis Breure, 1977 = Bulimus fa llax L. Pfeiffer, 1853; Bulimus trivittatus Mousson, 1869 = B. felix L. Pfeiffer, 1862; Bulimus andicola L. Pfeiffer, 1847 = B. multili neatus Say, 1825; Bulimulus ( Drymaeus ) comis Preston, 1907 = Bulimus pealianus Lea, 1838; Drymaeus incognita da Costa, 1 907 = D. bellus da Costa, 1906 = D. blandi Pilsbry, 1898 = Bulimulus ( Drymaeus ) smithii da Costa, 1898 . For the following species, precise localities are given for the first time: Drymaeus ( Drymaeus ) angusta da Costa, 1906, D. ( D. ) auris (L. Pfeiffer, 1 866), D. ( D. ) baranguillanus (L. Pfeiffer 1853), D. ( D. ) cognatus Pilsbry, 1901, D. ( D. ) geometricus (L. Pfeiffer 1846), D. ( D. ) inclinatus (L. Pfeiffer 1862), D. ( D. ) spadiceus da Costa, 1906, D. ( Mesembrinus ) koppelli (G.B. Sowerby III, 1892), D . ( M .) mu liebris (Reeve 1849) . Newly recorded for the Colombian malacofauna are the following five taxa: Drymaeus ( Drymaeus ) fordii Pilsbry, 1898, D. ( D. ) glaucostomus (Albers, 1852), D. ( D. ) vols us Fulton , 1 907 , D. ( Mesembrinus ) interruptus (Preston, 1909) . Th e following 27 taxa are excluded from the Colombian fauna as we consider them based on erroneous or doubtful records : Stenostylus meleagris (L. Pfeiffer, 1853), Drymaeus ( Drymaeus ) attenuatus ( L. Pfei f fer, 1853) , D . ( D . ) chimborasensis (Reeve, 1848) , D . ( D . ) edmuelleri (Albers, 1854) , D . ( D . ) linostoma (d’Orbigny, 1835) , D. ( D. ) membielinus (Crosse, 1867), D . ( D . ) phryne (L. Pfeiffer, 1863) , D . ( D . ) poecilus (d’Orbigny, 1835) , D . ( D . ) protractus (L. Pfeiffer, 1855) , D . ( D . ) rugistriatus Haas, 1952 , D . ( D . ) strigatus (Sowerby, 1833) , D . ( D . ) subinterruptus (L. Pfeiffer, 1853) , D . ( D . ) trigonostomus (Jonas, 1844) , D . ( Mesembrinus ) cactivorus (Broderip, 1832) , D . ( M . ) deshayesi (L. Pfeiffer, 1845) , D . ( M . ) dubius (L. Pfeiffer, 1853) , D . ( M . ) flavidus ( Menke, 18 29) , D . ( M . ) granadensis (L. Pfeiffer, 1848) , D . ( M . ) liliaceus (Férussac, 1821) , D . ( M . ) loxanus (Higgins, 1872) , D . ( M . ) manupictus (Reeve, 1848) , D . ( M . ) multifasciatus (Lamarck, 1822) , D . ( M . ) nitidus (Broderip, 1832) , D . ( M . ) pertristis Pilsbry, 1898 , D . ( M . ) pervariabilis (L. Pfeiffer, 1853) , D . ( M . ) studeri (L. Pfeiffer, 1847) , D . ( M . ) translucens (Broderip, 1832) . Key words - Northwestern South America, Ecuador, Panama, Venezuela, Stylommatophora, taxonomy, distribution . Introduction The la nd snail superfamily Orthalicoidea is a dominant element within the Neotropical malacofauna. Its taxonomy at the family level has seen several conflicting views, based on shell morphology and anatomy (Breure 1979, Schileyko 1999). Recently, Breure et al . ( 2010) and Breure & Romero (2012), presented phylogenetic studies covering representatives of the whole superfamily. Although some relationships need further study, the overall conclusion is that at present , seven families can be recognised within the Ortha licoidea. One of these families is the Bulimulidae, occurring in South and Central America, adjacent areas with in the U.S.A., and the Caribbean . In this paper we deal with the genera Stenostylus Pilsbry, 1898, and Drymaeus Albers, 18 5 0 of Colombia . W hile t he former is a small genus with only a handful of species, the latter is one of the most species - rich gen era of macro land snails in Colombia and also in the overall northern South American malacofauna. In the recent catalogue by Linares & Vera (2012) th re e Stenostylus respectively 99 Drymaeus species are listed; their list is, however, largely - 1 - Folia Conchyliologica 52 décembre 20 19 based on literature and museum records from electronic databases, so the specimens were not critically examined and evaluated. A few considerations motivated thi s study. First, many nominal taxa of Drymaeus have been described from the region, but most of these species are known only from the type material where the locality data is often very imprecise, such as a country, or often using places names now restricte d to areas far from their namesakes, making it difficult to ascertain where these species actually lived or may live today. Further more , many species are known from few or single specimens so any intraspecific variability cannot be evaluated. A final motiv at ion was that of producing a document that would allow
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