Contributions to Zoology, 72 (2-3) 85-89 (2003)

SPB Academic Publishing bv, The Hague

New data on the Binkhorstia ubaghsii (Late ;

NE , SE )

Coole4 Barry+W.M. van Bakel John+W.M. Jagt René+H.B. Fraaije³ & Yvonne ¹, ²,

2 1Schepenhoek 235, NL-5403 GB Uden, the Netherlands; Natuurhistorisch Museum , deBosquet-

3 plein 6-7, P.O. Box 882, NL-6200 AW Maastricht; Oertijdmuseum de Groene Poort, Bosscheweg 80, NL-

4 5283 WB Boxtel, the Netherlands; Sl Maartenslaan 88, NL-6039 BM Stramproy, the Netherlands

Keywords:: Crustacea, type Maastrichtian, Binkhorstia, systematic placement

Abstract 1997; Jagt et al2000). Many new of and

new data for the largely western Tethyan Carcinere-

Well-preserved material ofBinkhorstia ubaghsii reveals some tidae have been described in the past two decades additional information that helps clarify the taxonomic affini- (Feldmann et al., 1999; Feldmann& Villamil, 2002; ties with the Torynommidae of this Late crab from Vega & Feldmann, 1991; Vega et al., 1995; Vega the Maastricht area of Belgium and Netherland et al., 1997; Vegaet al., 2001). Feldmann& Villamil

included five in the (2002) genera Carcineretidae,

namely Carcineretes Withers, 1922, Branchiocar- Introduction cinus Vega et al., 1995, Mascaranada Vega &

Feldmann, 1991, Ophthalmoplax Rathbun, 1935 and Carcinologists dealing with are often Woodbinax Stenzel, 1952. They suggested that Bink- faced with difficulties in classifying their material. horstia and Withersella should be retained in the With only a single or a handful of specimens avail- Cyclodorippidae Ortmann, 1892. than of able, more often not consisting only (par-

the tial) carapaces, dorsal carapace characters are

sole features to on. Classification of extant crabs go Results mostly cannot be applied, since ventral morphol-

ogy is here of prime importance (Guinot, 1977, The new material of Binkhorstia ubaghsii consists char- 1978). Having come to realize that carapace of three more or less complete specimens all col- of unrelated acters may overlap in representatives lected from the uppermost Nekum Member (Maas- in the decade have families, past paleontologists tricht the CBR-Romontbos Formation), at quarry the few in used cases which crabs are preserved in Eben Emael, NE Belgium. Here, B. ubaghsii is with well-preserved ventral parts with much suc- fairly common and is found in association with

cess (Karasawa, 2003, Guinot & Tavares, 2001). Paranecrocarcinus quadriscissus Noetling, 1881, Binkhorstia 1881, known Noetling, exclusively “Pseudoraninella” muelleri(van Binkhorst, 1857),

from Upper Maastrichtian deposits in the area type Homolopsis declinata Collins et al., 1995, Eumor- of the Maastrichtian (SE Netherlands, NE phocorystes sculptus (van Binkhorst, 1857), Pa- has been in different families by Belgium), placed guristes? florae Collins et al., 1995 and Lyreidina

various authors. Glaessner (1980) erected the fam- 1998. pyriformis Fraaye & van Bakel, Two spe- and included ily Torynommidae (Cyclodorippoidea), cimens most likely represent corpses; the third Withersella & Collins, 1972and Binkhorstia. Wright is probably a molt. The best preserved specimen Withersella was originally placed in the Carcinere- [NUMBER] has been prepared in detailboth dorsally ,

tidae and with Binkhorstia Beurlen, 1930, together and ventrally, and shows important details at the

again assigned to the Carcineretidae by later au- ventral side with all the abdominal segments, ster-

thors (Collins al., 1995; 1996; Wright, nites and et Fraaye, bases of limbs preserved. The coxae of

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and References the fifth pair of pereiopods are diminutive, situated laterally ofthe small and slender first abdo-

Beurlen K. 1930. minal segment, covered by the posterior carapace Vergleichende Stammesgeschichte. Methoden, Probleme unter besonderer The second shows both Grundlagen, margin. corpse chelipeds, Beriicksichtigung der hoheren Krebse. Fortschr. Geol. some disarticulated limb fragments, and a well- Paldont. 8: 317-586. preserved, broad sternum. The carapace shows the Binkhorst JT van. 1857. Neue Krebse aus der Maestrichter

rostrum, a feature that be Tuffkreide. Verh. Naturhist. Ver. 14: unique spatulated may preuss. Rheinl. Westf.

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Collins JSH, RHB & Jagt JWM. 1995. Late Creta- specimen, a presumed molt, lie on top of the cara- Fraaye ceous anomurans and brachyurans from the Maastrichtian pace, but with the left-hand cheliped preserved the type area. Acta palaeont. pol. 40: 165-210. identification as B. ubaghsii is indisputed. Pereio- Fddmann RM,Villamil T. 2002. A new carcineretid crab (Up- P3 pods P2, and P4 are and ex- ofColombia. Jour. Paleo. 76: VIS- partially exposed per Turanian,Cretaceous) tremely flattened, assumed to have served for 724.

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aus dem Senon . Ueber NoetlingF. 1881 einige Brachyuren von We thank Rudi W. Dortangs (Amstenrade) for taking photo- Mastricht [sic] und dem Tertiar Norddeutschlands. Z. dt. geol. graphs, Robert Pieters (Geel) for donation of material. Ges. 33: 357-371.

P'g. I. Binkhorstia ubaghsii (van Binkhorst, 1857); Upper Nekum Member (Maastricht formation, Late Maastrichtian) of CBR-

Romontbos of quarry, Eben Emael (NE Belgium). 1, Right cheliped. 2, Left cheliped the same specimen. 3, Dorsal view of this

both claws and limb Dorsal view. Ventral the delicate specimen, showing complete carapace, fragments. 4, 5, view, showing preser- vation.

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Downloaded from Brill.com10/04/2021 11:30:54AM via free access Contributions to Zoology, 72 (2-3) - 2003 89

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fdg. 2. Binkhorstia ubaghsii (van Binkhorst, 1857) and Binkhorstia euglypha Collins, Fraaye & Jagt, 1995. 1, Ventral view, with

details of the sternum ofB. ubaghsii. 2, Orbital view ofthis specimen, showing the unique spatulated rostrum. 3, Binkhorstia euglypha

& ventral view of k, the ofthe Member Collins, Fraaye Jagt, 1995, holotype (MAB, 1033) from upper part Meerssen (latest Maastricht

of the Blom Dorsal view formation) former quarry, Berg en Terblijt (SE Netherlands). 4, of the holotype. 5, Dorsal view of assumed molt, B. ubaghsii showing the left-hand and flattened , claw, pereiopods.

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