DINOSAURS AMONG US INSIDE • Map of the Exhibition • Essential Questions • Teaching in the Exhibition • Come Prepared Checklist • Correlation to Standards • Glossary
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Journal of Paleontology
Journal of Paleontology http://journals.cambridge.org/JPA Additional services for Journal of Paleontology: Email alerts: Click here Subscriptions: Click here Commercial reprints: Click here Terms of use : Click here New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra, Diego Pol, Oliver W.M. Rauhut and Ignacio A. Cerda Journal of Paleontology / Volume 90 / Issue 03 / May 2016, pp 555 - 577 DOI: 10.1017/jpa.2016.24, Published online: 27 June 2016 Link to this article: http://journals.cambridge.org/abstract_S002233601600024X How to cite this article: Marcos G. Becerra, Diego Pol, Oliver W.M. Rauhut and Ignacio A. Cerda (2016). New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids. Journal of Paleontology, 90, pp 555-577 doi:10.1017/jpa.2016.24 Request Permissions : Click here Downloaded from http://journals.cambridge.org/JPA, IP address: 190.172.49.57 on 16 Aug 2016 Journal of Paleontology, 90(3), 2016, p. 555–577 Copyright © 2016, The Paleontological Society 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.24 New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra,1 Diego Pol,1 Oliver W.M. Rauhut,2 and Ignacio A. Cerda3 1CONICET- Museo Palaeontológico Egidio Feruglio, Fontana 140, Trelew, Chubut 9100, Argentina 〈[email protected]〉; 〈[email protected]〉 2SNSB, Bayerische Staatssammlung für Paläontologie und Geologie and Department of Earth and Environmental Sciences, LMU München, Richard-Wagner-Str. -
A Chinese Archaeopterygian, Protarchaeopteryx Gen. Nov
A Chinese archaeopterygian, Protarchaeopteryx gen. nov. by Qiang Ji and Shu’an Ji Geological Science and Technology (Di Zhi Ke Ji) Volume 238 1997 pp. 38-41 Translated By Will Downs Bilby Research Center Northern Arizona University January, 2001 Introduction* The discoveries of Confuciusornis (Hou and Zhou, 1995; Hou et al, 1995) and Sinornis (Ji and Ji, 1996) have profoundly stimulated ornithologists’ interest globally in the Beipiao region of western Liaoning Province. They have also regenerated optimism toward solving questions of avian origins. In December 1996, the Chinese Geological Museum collected a primitive bird specimen at Beipiao that is comparable to Archaeopteryx (Wellnhofer, 1992). The specimen was excavated from a marl 5.5 m above the sediments that produce Sinornithosaurus and 8-9 m below the sediments that produce Confuciusornis. This is the first documentation of an archaeopterygian outside Germany. As a result, this discovery not only establishes western Liaoning Province as a center of avian origins and evolution, it provides conclusive evidence for the theory that avian evolution occurred in four phases. Specimen description Class Aves Linnaeus, 1758 Subclass Sauriurae Haeckel, 1866 Order Archaeopterygiformes Furbringer, 1888 Family Archaeopterygidae Huxley, 1872 Genus Protarchaeopteryx gen. nov. Genus etymology: Acknowledges that the specimen possesses characters more primitive than those of Archaeopteryx. Diagnosis: A primitive archaeopterygian with claviform and unserrated dentition. Sternum is thin and flat, tail is long, and forelimb resembles Archaeopteryx in morphology with three talons, the second of which is enlarged. Ilium is large and elongated, pubes are robust and distally fused, hind limb is long and robust with digit I reduced and dorsally migrated to lie in opposition to digit III and forming a grasping apparatus. -
LETTER Doi:10.1038/Nature14423
LETTER doi:10.1038/nature14423 A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings Xing Xu1,2*, Xiaoting Zheng1,3*, Corwin Sullivan2, Xiaoli Wang1, Lida Xing4, Yan Wang1, Xiaomei Zhang3, Jingmai K. O’Connor2, Fucheng Zhang2 & Yanhong Pan5 The wings of birds and their closest theropod relatives share a ratios are 1.16 and 1.08, respectively, compared to 0.96 and 0.78 in uniform fundamental architecture, with pinnate flight feathers Epidendrosaurus and 0.79 and 0.66 in Epidexipteryx), an extremely as the key component1–3. Here we report a new scansoriopterygid short humeral deltopectoral crest, and a long rod-like bone articu- theropod, Yi qi gen. et sp. nov., based on a new specimen from the lating with the wrist. Middle–Upper Jurassic period Tiaojishan Formation of Hebei Key osteological features are as follows. STM 31-2 (Fig. 1) is inferred Province, China4. Yi is nested phylogenetically among winged ther- to be an adult on the basis of the closed neurocentral sutures of the opods but has large stiff filamentous feathers of an unusual type on visible vertebrae, although this is not a universal criterion for maturity both the forelimb and hindlimb. However, the filamentous feath- across archosaurian taxa12. Its body mass is estimated to be approxi- ers of Yi resemble pinnate feathers in bearing morphologically mately 380 g, using an empirical equation13. diverse melanosomes5. Most surprisingly, Yi has a long rod-like The skull and mandible are similar to those of other scansoriopter- bone extending from each wrist, and patches of membranous tissue ygids, and to a lesser degree to those of oviraptorosaurs and some basal preserved between the rod-like bones and the manual digits. -
New Heterodontosaurid Remains from the Cañadón Asfalto Formation: Cursoriality and the Functional Importance of the Pes in Small Heterodontosaurids
Journal of Paleontology, 90(3), 2016, p. 555–577 Copyright © 2016, The Paleontological Society 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.24 New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra,1 Diego Pol,1 Oliver W.M. Rauhut,2 and Ignacio A. Cerda3 1CONICET- Museo Palaeontológico Egidio Feruglio, Fontana 140, Trelew, Chubut 9100, Argentina 〈[email protected]〉; 〈[email protected]〉 2SNSB, Bayerische Staatssammlung für Paläontologie und Geologie and Department of Earth and Environmental Sciences, LMU München, Richard-Wagner-Str. 10, Munich 80333, Germany 〈[email protected]〉 3CONICET- Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, Museo Carlos Ameghino, Belgrano 1700, Paraje Pichi Ruca (predio Marabunta), Cipolletti, Río Negro, Argentina 〈[email protected]〉 Abstract.—New ornithischian remains reported here (MPEF-PV 3826) include two complete metatarsi with associated phalanges and caudal vertebrae, from the late Toarcian levels of the Cañadón Asfalto Formation. We conclude that these fossil remains represent a bipedal heterodontosaurid but lack diagnostic characters to identify them at the species level, although they probably represent remains of Manidens condorensis, known from the same locality. Histological features suggest a subadult ontogenetic stage for the individual. A cluster analysis based on pedal measurements identifies similarities of this specimen with heterodontosaurid taxa and the inclusion of the new material in a phylogenetic analysis with expanded character sampling on pedal remains confirms the described specimen as a heterodontosaurid. Finally, uncommon features of the digits (length proportions among nonungual phalanges of digit III, and claw features) are also quantitatively compared to several ornithischians, theropods, and birds, suggesting that this may represent a bipedal cursorial heterodontosaurid with gracile and grasping feet and long digits. -
The Dinosaurs Are Back!
EXHIBITION FOR HIRE THE DINOSAURS ARE BACK! THE DINOSAURS ARE BACK! In partnership with Emilio’s Creations, Universeum proudly presents the unique dinosaur exhibition “The dinosaurs are back”. The latest scientific discoveries are summarised in a spectacular visitor experience with 16 life-size dinosaurs that move and make noises. Ten of them are feathered in accordance with recent research. The exhibition for hire comprises eight different species set in a prehistoric forest landscape with trees and plants from that specific era. Included is the world’s largest known land predator, theSpinosaurus , at 14 metres long and five metres tall. Also on display are the four-metre tall and fully feathered Therizinosaurus and the legendary Tyrannosaurus rex, together with a feathered juvenile. Feathers The dinosaurs in the exhibition for hire The dinosaurs never really died out. They live The dinosaurs in the exhibition have been carefully on and we see and hear them every day. After selected to show a diverse range of species and all, birds are directly descended from some of highlight scientifically interesting points. the most ferocious and predatory dinosaurs, Sinosauropteryx (x2). The first genus of dinosaur discovered with recent research confirming the close outside of Avialae(birds) that is known to have been links between dinosaurs and birds. This feathered. point is clearly illustrated when visitors see the Tyrannosaurus rex (x2). As well as being famous, it is of great feathered dinosaurs in the exhibition. scientific interest. Researchers are speculating as to whether the adult lost the coat of feathers it had when young or Environment whether it retained all or part of it throughout its life. -
A New Raptorial Dinosaur with Exceptionally Long Feathering Provides Insights Into Dromaeosaurid flight Performance
ARTICLE Received 11 Apr 2014 | Accepted 11 Jun 2014 | Published 15 Jul 2014 DOI: 10.1038/ncomms5382 A new raptorial dinosaur with exceptionally long feathering provides insights into dromaeosaurid flight performance Gang Han1, Luis M. Chiappe2, Shu-An Ji1,3, Michael Habib4, Alan H. Turner5, Anusuya Chinsamy6, Xueling Liu1 & Lizhuo Han1 Microraptorines are a group of predatory dromaeosaurid theropod dinosaurs with aero- dynamic capacity. These close relatives of birds are essential for testing hypotheses explaining the origin and early evolution of avian flight. Here we describe a new ‘four-winged’ microraptorine, Changyuraptor yangi, from the Early Cretaceous Jehol Biota of China. With tail feathers that are nearly 30 cm long, roughly 30% the length of the skeleton, the new fossil possesses the longest known feathers for any non-avian dinosaur. Furthermore, it is the largest theropod with long, pennaceous feathers attached to the lower hind limbs (that is, ‘hindwings’). The lengthy feathered tail of the new fossil provides insight into the flight performance of microraptorines and how they may have maintained aerial competency at larger body sizes. We demonstrate how the low-aspect-ratio tail of the new fossil would have acted as a pitch control structure reducing descent speed and thus playing a key role in landing. 1 Paleontological Center, Bohai University, 19 Keji Road, New Shongshan District, Jinzhou, Liaoning Province 121013, China. 2 Dinosaur Institute, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, California 90007, USA. 3 Institute of Geology, Chinese Academy of Geological Sciences, 26 Baiwanzhuang Road, Beijing 100037, China. 4 University of Southern California, Health Sciences Campus, BMT 403, Mail Code 9112, Los Angeles, California 90089, USA. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Was Dinosaurian Physiology Inherited by Birds? Reconciling Slow Growth in Archaeopteryx
Was Dinosaurian Physiology Inherited by Birds? Reconciling Slow Growth in Archaeopteryx Gregory M. Erickson1,6*, Oliver W. M. Rauhut2, Zhonghe Zhou3, Alan H. Turner4,6, Brian D. Inouye1, Dongyu Hu5, Mark A. Norell6 1 Department of Biological Science, Florida State University, Tallahassee, Florida, United States of America, 2 Bayerische Staatssammlung fu¨r Pala¨ontologie und Geologie and Department of Earth and Environmental Sciences, LMU Munich, Mu¨nchen, Germany, 3 Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology & Paleoanthropology, Chinese Academy of Science, Beijing, China, 4 Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, United States of America, 5 Paleontological Institute, Shenyang Normal University, Shenyang, China, 6 Division of Paleontology, American Museum of Natural History, New York, New York, United States of America Abstract Background: Archaeopteryx is the oldest and most primitive known bird (Avialae). It is believed that the growth and energetic physiology of basalmost birds such as Archaeopteryx were inherited in their entirety from non-avialan dinosaurs. This hypothesis predicts that the long bones in these birds formed using rapidly growing, well-vascularized woven tissue typical of non-avialan dinosaurs. Methodology/Principal Findings: We report that Archaeopteryx long bones are composed of nearly avascular parallel- fibered bone. This is among the slowest growing osseous tissues and is common in ectothermic reptiles. These findings dispute the hypothesis that non-avialan dinosaur growth and physiology were inherited in totality by the first birds. Examining these findings in a phylogenetic context required intensive sampling of outgroup dinosaurs and basalmost birds. Our results demonstrate the presence of a scale-dependent maniraptoran histological continuum that Archaeopteryx and other basalmost birds follow. -
Perinate and Eggs of a Giant Caenagnathid Dinosaur from the Late Cretaceous of Central China
ARTICLE Received 29 Jul 2016 | Accepted 15 Feb 2017 | Published 9 May 2017 DOI: 10.1038/ncomms14952 OPEN Perinate and eggs of a giant caenagnathid dinosaur from the Late Cretaceous of central China Hanyong Pu1, Darla K. Zelenitsky2, Junchang Lu¨3, Philip J. Currie4, Kenneth Carpenter5,LiXu1, Eva B. Koppelhus4, Songhai Jia1, Le Xiao1, Huali Chuang1, Tianran Li1, Martin Kundra´t6 & Caizhi Shen3 The abundance of dinosaur eggs in Upper Cretaceous strata of Henan Province, China led to the collection and export of countless such fossils. One of these specimens, recently repatriated to China, is a partial clutch of large dinosaur eggs (Macroelongatoolithus) with a closely associated small theropod skeleton. Here we identify the specimen as an embryo and eggs of a new, large caenagnathid oviraptorosaur, Beibeilong sinensis. This specimen is the first known association between skeletal remains and eggs of caenagnathids. Caenagnathids and oviraptorids share similarities in their eggs and clutches, although the eggs of Beibeilong are significantly larger than those of oviraptorids and indicate an adult body size comparable to a gigantic caenagnathid. An abundance of Macroelongatoolithus eggs reported from Asia and North America contrasts with the dearth of giant caenagnathid skeletal remains. Regardless, the large caenagnathid-Macroelongatoolithus association revealed here suggests these dinosaurs were relatively common during the early Late Cretaceous. 1 Henan Geological Museum, Zhengzhou 450016, China. 2 Department of Geoscience, University of Calgary, Calgary, Alberta, Canada T2N 1N4. 3 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China. 4 Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada T6G 2E9. 5 Prehistoric Museum, Utah State University, 155 East Main Street, Price, Utah 84501, USA. -
A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi
www.nature.com/scientificreports OPEN A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Received: 12 October 2017 Accepted: 7 March 2018 comments on size variation among Published: xx xx xxxx oviraptorosaurs Yilun Yu1, Kebai Wang2, Shuqing Chen2, Corwin Sullivan3,4, Shuo Wang 5,6, Peiye Wang2 & Xing Xu7 The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi difers from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with signifcant posterior orientation; accessory trochanter low and confuent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior fange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae. Oviraptorosauria is a clade of maniraptoran theropod dinosaurs characterized by a short, high skull, long neck and short tail. -
Therizinosaurus Cheloniformis
3-EURO-SUPERSAURS THERIZINOSAURUS CHELONIFORMIS Series Supersaurs Appearances can be deceptive. Weighing five tons, up to ten metres tall and boas- ting sharp claws that were longer than those of any other land animal that has ever Face value ¤ 3 existed, the Therizinosaurus was not as dangerous as it looked. The subject of the Date of Issue 17 February 2021 sixth coin in the superlative Supersaurs series was actually a vegetarian and its claws Design K. Kuntner/ H. Andexlinger more suited to detaching vegetation than attacking the other animals that roamed Diameter 34 mm the planet some 70 million years ago. Quality Uncirculated Mintage 65,000 As a theropod, Therizinosaurus cheloniformis was not only distantly related to pre- Alloy Coloured metal dators such as Tyrannosaurus rex but also to present-day birds. Originally carnivor- Total Weight 16.00 g ous, over time the Therizinosaurus changed its diet and became a pure herbivore. Edge Plain During the late Cretaceous period, it lived in areas of the northern continent of Colour application, glow in the dark, Laurasia that now correspond to North America, China and Mongolia. Up to one no packaging. metre in length, flat, curved and pointed at the end, the dinosaur’s claws are re- sponsible for its nickname, ‘scythe lizard’. They were used primarily to reach high Recommended branches in trees as well as perhaps for self-defence. Well preserved remains of one initial issue price: ¤ 12.60 (incl. 20% VAT) of its close relatives have suggested that, rather surprisingly, the Therizinosaurus was also partially feathered. WITH COIN MOTIF COLOURED GLOW-IN-THE-DARK Therizinosaurus features all 12 of the extreme prehistoric animals in the Supersaurs EFFECT series in silhouette on its obverse, while a colour-printed Therizinosaurus is shown in its native prehistoric habitat next to a nest full of eggs on the coin’s glow-in-the- dark reverse. -
New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia
Bull. Natn. Sci. Mus., Tokyo, Ser. C, 30, pp. 95–130, December 22, 2004 New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia Junchang Lü1, Yukimitsu Tomida2, Yoichi Azuma3, Zhiming Dong4 and Yuong-Nam Lee5 1 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China 2 National Science Museum, 3–23–1 Hyakunincho, Shinjukuku, Tokyo 169–0073, Japan 3 Fukui Prefectural Dinosaur Museum, 51–11 Terao, Muroko, Katsuyama 911–8601, Japan 4 Institute of Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 5 Korea Institute of Geoscience and Mineral Resources, Geology & Geoinformation Division, 30 Gajeong-dong, Yuseong-gu, Daejeon 305–350, South Korea Abstract Nemegtia barsboldi gen. et sp. nov. here described is a new oviraptorid dinosaur from the Late Cretaceous (mid-Maastrichtian) Nemegt Formation of southwestern Mongolia. It differs from other oviraptorids in the skull having a well-developed crest, the anterior margin of which is nearly vertical, and the dorsal margin of the skull and the anterior margin of the crest form nearly 90°; the nasal process of the premaxilla being less exposed on the dorsal surface of the skull than those in other known oviraptorids; the length of the frontal being approximately one fourth that of the parietal along the midline of the skull. Phylogenetic analysis shows that Nemegtia barsboldi is more closely related to Citipati osmolskae than to any other oviraptorosaurs. Key words : Nemegt Basin, Mongolia, Nemegt Formation, Late Cretaceous, Oviraptorosauria, Nemegtia. dae, and Caudipterygidae (Barsbold, 1976; Stern- Introduction berg, 1940; Currie, 2000; Clark et al., 2001; Ji et Oviraptorosaurs are generally regarded as non- al., 1998; Zhou and Wang, 2000; Zhou et al., avian theropod dinosaurs (Osborn, 1924; Bars- 2000).