2012 MBL Annual Report 2012 63 Non Profit ORG

2 0 1 2 M B L A n n u a l R e p ort 2012 Top Highlights

• Harvard biologist and long-time MBL Trustee Joan Ruderman appointed President and Director of the MBL

• Michael Sheetz, James Spudich, and Ron Vale, who discovered the motor protein kinesin at the MBL in 1984, were awarded the Lasker Award for Basic Medical Research

• MBL Catalyst Campaign was completed, surpassing goal of $125 million with $136 million raised as of December 31, 2012

• $835,889 in scholarships awarded to 252 highly qualified students

• MBL educational programs trained 562 students from 345 institutions in 52 countries

The MBL Annual Report is published by the Office of Communications Marine Biological Laboratory. Although the MBL greatest possible care has been taken in the 7 MBL Street preparation of this record, the MBL recognizes Woods Hole, MA 02543 the possibility of omissions or inaccuracies. If [email protected] any are noted, please accept our apology and advise us of any corrections to be made. Contents

2 Message from the Chairman and the President Visit us online! 3 Research & Education Highlights To view the comprehensive report—including 12 MBL at a Glance investigator, faculty, and course lists—please visit:

13 Financials mbl.edu/annualreport2012

16 Gifts

57 Leadership Message from THE Chairman and the President Research and Education Highlights

2012 was a year of transitions. Notably, the MBL While working toward the affiliation, the MBL continued to make welcomed its 14th Director and President Joan V. Ruderman, its mark in our three major program areas: education, visiting formerly MBL Trustee and Marion V. Nelson Professor of Cell research, and year-round research. The photo essay on pages 3-11 Biology at Harvard Medical School. We concluded the Catalyst provides a summary of the year’s highlights, including the Lasker Campaign, which raised more than $136 million dollars Award for Basic Medical Research for work done in part at the since 2006. MBL by three scientists with strong MBL ties.

The lab also charted a course to form a landmark affiliation with We also worked to resolve ongoing deficits and cash shortfalls. the University of Chicago, involving months of discussion and Full details of this year’s financials can be found on pages 14-15. input from many communities. This new link will contribute to both institutions’ missions of research and education, and will 2012 was both challenging and revitalizing. We are tremendously help strengthen the MBL’s financial foundation. grateful to our faculty, staff, trustees, overseers, funders, and other supporters for not only standing by the MBL, but also for helping MBL and the University of Chicago have a long history together position the MBL to fulfill its potential to impact the biological dating to their earliest days. The University is a renowned sciences in far-reaching ways. research institution with a great record of complementing the strengths and missions of its other affiliates. The affiliation strengthens both institutions’ missions of leadership and innovation in scientific research and education. It also John W. Rowe, MD promises to foster areas of research and training in areas Chair, MBL Board of Trustees ranging from molecular mechanisms of basic cellular processes to global-scale ecosystem analysis. The affiliation also offers opportunities to build collaborative research and educational programs in areas including neuroscience, evolutionary and developmental biology, regenerative biology, cell biology, Joan V. Ruderman microbial sciences, molecular engineering, ecosystems science, MBL President and Director and global environmental change.

2 MBL Annual REport 2012 Research and Education Highlights

The MBL sparks biological discovery by providing a charged scientific environment and attracting the world’s leading researchers. MBL science has helped reveal the mechanisms of cellular processes and their roles in disease, the processes that sustain life, and the genetic ancestry of life itself. It has also promoted a cross-fertilization of ideas, nurturing numerous scientific fields that continue to advance our understanding of human and environmental systems.

3 MBL Annual REport 2011 MBL Annual REport 2012 3 REsearch and Education highlights

Exploring the Hidden Realm of Microbes Beneath the Ocean Floor

Using remotely operated vehicles, Bay Paul Center Associate Scientist Julie Huber and her colleagues sampled the world’s deepest hydrothermal vents in the Mid-Cayman Rise and discovered high-temperature microbial communities thriving in the hydrogen- rich fluids. Also in 2012 Huber and colleagues from several institutions received a $2.25 million award from the Gordon and Betty Moore Foundation to undertake novel experiments—some at the bottom of the ocean—to understand the microbes and viruses in the subseafloor rocks of Axial Seamount, a deep-sea volcano in the Pacific Ocean.

Harvard Biologist Joan Ruderman Named MBL President and Director

Joan Ruderman was named President and Director of the MBL on November 3, 2012. Ruderman was previously the Marion V. Nelson Professor of Cell Biology at Harvard Medical School. She also served as Senior Science Advisor at the Radcliffe Institute for Advanced Study at Harvard University. Ruderman is best known for her work on fertilization and on the molecular mechanisms that regulate mitosis, the last part of the cell division cycle. Ruderman has a long affiliation with the MBL, having served on the Board of Trustees and its Executive Committee and as Speaker of the MBL Corporation. Ruderman is an alumna of the MBL Embryology course and a former MBL course faculty member and visiting scientist.

4 MBL Annual REport 2012 REsearch and Education highlights

Vale, Sheetz, and Spudich Receive Lasker Award for Basic Medical Research

Scientists working at the MBL in the 1980s invented ways to study how “cargo” is moved within cells and, as a result, discovered a new biological motor protein, kinesin. In 2012, they received the Lasker Award for Basic Medical Research for that work. Michael Sheetz of Columbia University; James Spudich of Stanford University School of Medicine; and Ron Vale of University of California, San Francisco/Howard Hughes Medical Institute

Why Are Our Salt Marshes Falling Apart?

Northeast U.S. coastal salt marshes are more vulnerable to nitrogen pollution than previously believed, according to a study published in Nature by MBL Ecosystems and MBL Whitman Center shared the Center Senior Scientist Linda Deegan and colleagues working in New England’s largest award, which is among the most respected coastal marsh at Plum Island, MA. The researchers found that by adding an amount of science prizes in the world, and often nitrogen that is about the same as concentrations from septic and fertilizer runoff to the anticipates the Nobel Prize for its recipients. water entering salt marsh creeks over eight years, they stimulated the decomposition The team’s fundamental discoveries have in creek banks and induced slumping and marsh loss. The long-term study suggests spurred research on new treatments for that salt marshes in New England and perhaps other locations are less likely to act as cardiac problems, neurological disorders, “sponges” that take up large quantities of nitrogen indefinitely and more likely to be and cancer. themselves damaged by nitrogen runoff to the coast.

MBL Annual REport 2012 5 REsearch and Education highlights

Neuroscientist Joins Bell Center An Evolutionary Surprise

Jennifer Morgan, formally on the faculty at The evolutionary origin of the vertebrate brain is a mystery, because nothing anatomically like University of Texas-Austin, was appointed an it has been found in invertebrates. Yet a clue to Assistant Scientist in the Eugene Bell Center for its origins was discovered by Whitman Center Regenerative Biology and Tissue Engineering. Investigators Ariel Pani, Chris Lowe, and their With funding from the NIH, Morgan seeks to colleagues. The team discovered some of the understand the mechanisms of neurodegeneration genetic processes that regulate vertebrate brain and regeneration in the vertebrate central nervous development in, surprisingly, the acorn worm, a brainless, burrowing marine invertebrate whose system. She is currently using the sea lamprey as lineage diverged from vertebrates more than a model organism for spinal cord regeneration 500 million years ago. In vertebrates, Pani says, following injury. She has discovered that the these signaling centers are major components protein synuclein accumulates after injury and of a gene expression program “that sets up the modulates neuronal survival such that reducing foundation of how the brain develops.” synuclein accumulation may be a useful mechanism to increase regeneration after spinal cord injury. Her work holds promise for the development of novel therapeutic approaches for affected patients.

Christopher Neill Named Director of Ecosystems Center

Christopher Neill, MBL Senior Scientist and the Phyllis and Charles M. Rosenthal Director of the Brown-MBL Partnership, was named Director of the Ecosystems Center. For more than two decades Neill has studied how deforestation in the Brazilian Amazon affects soil fertility, water quality, and stream habitats. In addition to a long-standing interest in local applied environmental research, Neill also has considerable experience in communicating science to the public and directs the Hands-on Environmental Laboratory for the MBL’s Logan Science Journalism Program.

6 MBL Annual REport 2012 REsearch and Education highlights

Nerves Control Iridescence in Squid’s Joint Brown University/MBL Graduate Remarkable “Electric Skin” Program Awards Four PhDs

The mystery behind how squid control their skin’s iridescence, an activity responsible for the animal’s rainbow of color and extraordinary camouflage capability, was uncovered by Sensory Physiology and Behavior Program Assistant Research Scientists Paloma Gonzalez Bellido, Trevor Wardill, and their colleagues. The researchers reported that nerves in the animal’s skin control the spectrum of shimmering hues, as well as Having Serious Fun in the MBL’s their speed of change. The work marked Physiology Course the first demonstration of neural control of iridescence in an invertebrate species. How Several former directors of the MBL squid choose and hold particular skin colors to Physiology Course detailed their winning camouflage themselves remains unknown and Four Brown-MBL Program Ph.D. students, formula for instilling in students the is particularly interesting because the animals Priya Dutta, Yuko Hasegawa, Shelby Riskin, and passion for and ability to conduct “real are completely colorblind. Susanna Theroux, defended their dissertations research,” in Science. The article presented in 2012, bringing the total number of program the overwhelmingly positive feedback from Ph.D. graduates to date to ten. Graduate a poll of 2004-2010 course alumni; and students in this joint program draw on the remarkable list of research papers and the expertise of faculty mentors at Brown meeting abstracts that developed out of University and the MBL and often conduct Physiology course projects. The course is one research in the MBL’s laboratories in Woods of 25 courses the MBL offers for advanced, Hole or at field sites in Alaska, Antarctica, the laboratory-based research training in fields Amazon, Africa, North America, Panama, and such as cellular physiology, embryology, other locales. neurobiology, and microbiology.

MBL Annual REport 2012 7 REsearch and Education highlights

“Safe” Levels Of Arsenic In Drinking Water New Program in Sensory Physiology Have Negative Health Effects on Pregnant/ and Behavior Launched Lactating Mothers And Offspring The Program in Sensory Physiology and A study led by Joshua Hamilton, Bay Paul Behavior was formed to capitalize on the Center Senior Scientist and MBL Chief Academic MBL’s many strengths in sensory biology— and Scientific Officer and his colleagues at the especially vision, balance, and hearing Dartmouth Superfund Research Program on Toxic research. Building on the growth and success Metals reported that exposure to arsenic in drinking of existing Marine Resources laboratories—as water deemed safe in the U.S. has negative health well as long-time strengths in sensory biology effects on pregnant mice and their offspring. The and neuroscience in the Whitman Center, results add to the body of knowledge the team is Grass Laboratory, and summer courses—a developing pertaining to arsenic sources, its effects central goal of the program is to synergize at the cellular level, the ways in which it affects the the many MBL sensory biology/neuroscience health of mammals like mice and, ultimately, how activities into a cohesive group and provide a it causes disease and harmful health outcomes for career path for young scientists. Senior humans. Scientist Roger Hanlon directs the program with Associate Director and Senior Scientist Stephen Highstein.

New Imaging Technique Spurs Interdisciplinary Collaboration

Tomomi Tani, an Associate Scientist in the Cellular Dynamics Program, perfected a technique using fluorescence polarization and a novel imaging system he developed that allows one to image single molecules inside living cells and determine their three-dimensional orientation. He applied the technique in a new collaboration with Cellular Dynamics Program Senior Scientist Rudolf Oldenbourg, an expert in polarization microscopy, and Amy Gladfelter, a Whitman Investigator from Dartmouth College who is an expert on septins, key proteins involved in cell division. This project has led to a successful NIH collaborative research grant and the recruitment of Assistant Research Scientist Shalin Mehta.

8 MBL Annual REport 2012 REsearch and Education highlights

Human microbiome project Relies on MBL Innovations, Insights

Bay Paul Center scientists Susan Huse, Hilary Morrison, and Mitchell Sogin were among more than 200 scientists who reported the results of the NIH’s Human Microbiome Project, an effort to define the boundaries of bacterial variation found in healthy human beings. The project adopted several advanced genetic sequencing and analysis methods, some of which were originally developed by the MBL. The project discovered that the human body contains a few highly abundant bacterial types plus many more low-abundance types (the so-called “rare biosphere,” a phenomenon first discovered in ocean samples by Sogin and his MBL colleagues). Marine Bacteria Distribution “Bipolar”

A research team led by Bay Paul Center Associate Scientist Linda Amaral Zettler and Director Mitchell Sogin found “bipolar” species of bacteria that occur in the Arctic and Antarctic, but nowhere else. The findings show that bacteria follow biogeographical distribution patterns more typical of macroscopic organisms and their survival may be vulnerable to impacts of environmental stressors, climate warming, and degradation in habitat quality that affect biodiversity of animal and plant species.

MBL Renews Diversity Efforts

The leaders of the six scientific and educational organizations in Woods Hole, including the MBL, renewed their commitment to building a diverse and inclusive scientific community by signing an updated Woods Hole Diversity Initiative Memorandum of Understanding (MOU). The MOU, first signed in 2004, reaffirms the institutional commitments to work together to attract and retain a more diverse workforce, one that represents the changing demographics of the nation and the international community.

9 MBL Annual REport 2011 MBL Annual REport 2012 9 REsearch and Education highlights

Draft of National Climate Assessment Released

The most comprehensive assessment of climate change effects for the United States ever produced was coordinated by Ecosystems Center Distinguished Scientist Jerry Melillo in his role as chair of the National Climate Assessment and Development Advisory Committee. Melillo organized more than 240 authors who wrote thirty chapters that covered the effects of climate change across multiple sectors including water resources, energy, agriculture, human health and rural communities. The National Climate Assessment, which operates under the U.S. Global Change Research Program, also contains region-by-region accounts of what climate change is happening and what the effects of those changes will mean for regional ecosystems and economies. A final version of the Assessment has passed review by the National Academy of Sciences and the public and will be released in early 2014.

Encyclopedia of Life Reaches Milestone; Provides Global Access Research Opportunities for Undergraduates The Encyclopedia of Life (EOL) surged past one million pages of content with the During the summer of 2012, nine Brown addition of hundreds of thousands of images undergraduate students participated in MBL and specimen data from the Smithsonian research programs, working on topics that Institution’s National Museum of Natural varied from the microbial communities in the History. EOL continued to fulfill its mission aquarium pet trade, to the effect of nitrogen on to provide “global access to knowledge about coastal salt marshes, to the makeup of the plant life on Earth” with the launch of EOL China communities in suburban Boston house lots. in Beijing. This version included Chinese Additionally, nine students from the MBL’s content, new layouts to accommodate Chinese Semester in Environmental Science Program characters, and the ability to search EOL using Class of 2011 completed internships with individual Chinese characters. EOL content was program faculty. also successfully replicated between the Library of Alexandria in Egypt and MBL, a prerequisite step for developing worldwide EOL nodes.

10 MBL Annual REport 2012 REsearch and Education highlights

American Society FOR Cell Biology Presents Award to the Physiology Course

The American Society for Cell Biology presented an award to the Physiology course “for its tremendous impact on developing the classical and modern eras of cell biology.” The award was given in honor of the course’s 120th anniversary.

Distinguished Scientist Joins the MBL’s Cellular Dynamics Program

Jonathan Gitlin joined the MBL as a Senior Scientist in the Cellular Dynamics Program. For more than 35 years, Gitlin’s career has combined pediatric medicine, research, administration, and teaching at such renowned institutions as Boston Children’s Hospital, Washington University School of Medicine, and Vanderbilt Children’s Hospital. Gitlin’s MBL Receives Gift of T. H. Morgan’s Nobel Prize Medal and Diploma research concentrates on birth defects related to genetics and nutrition. His laboratory Three granddaughters of Thomas Hunt Morgan, the late MBL scientist, trustee, and pioneer of uses zebrafish as a model to study embryonic modern genetics, donated his Nobel Prize medal and brilliantly colored diploma to the MBL development. Gitlin holds a joint appointment Archives. In 1933, Morgan received the first Nobel Prize ever awarded in the field of genetics of senior research scientist in the Bell Center for his “discoveries concerning the role played by the chromosome in heredity.” Morgan first and is currently serving as its director. came to the MBL as a student in 1889 and returned nearly every summer for the next half- century to conduct research.

MBL Annual REport 2012 11 MBL AT A glance* The Marine Biological Laboratory (MBL) is dedicated to scientific discovery and improving the human condition through research and education in biology, biomedicine, and environmental science. Founded in Woods Hole, Massachusetts in 1888, the MBL is a private, nonprofit institution and an affiliate of the University of Chicago.

LEADERSHIP and education. The affiliation builds training in fields such as cellular SCIENTIFIC CONFERENCES Chairman of the Board of Trustees: on shared values and historical ties physiology, embryology, neurobiology, The MBL annually hosts scientific John W. Rowe between Chicago and the MBL, which and microbiology, including (in 2012) meetings, conferences, and President and Director: was led by University of Chicago 582 students from 384 institutions and departmental retreats, accommodating Joan V. Ruderman faculty members for the first four 48 countries; and 888 faculty, staff, more than 2,600 participants from Speaker of the MBL Society: decades of its existence. and lecturers from 282 institutions and around the world. Colleen M. Cavanaugh 20 countries. MAJOR PROGRAMS & RESEARCH AREAS RESOURCES SCIENTISTS AND AFFILIATED STAFF Ecosystems Center: Ecosystems Educational programs also include The MBLWHOI Library houses one The MBL has approximately 300 year- processes, biogeochemistry, and undergraduate education and of the world’s foremost print and round employees, about half of which climate change independent research, graduate electronic literature collections in the are scientists and science support staff. training, the Brown-MBL Graduate biological, biomedical, ecological, and The staff is joined each year by more Bay Paul Center for Comparative Program in Biological and oceanographic sciences; and conducts than 300 visiting scientists, summer Molecular Biology & Evolution: Environmental Sciences, postdoctoral digitization and informatics projects. staff, and research associates from Microbial ecology; genetics and training, and community continuing hundreds of institutions around the metagenomics; molecular evolution education programs. The W. M. Keck Ecological and world, as well as a large number of Evolutionary Genetics Facility is a high- faculty and students participating in Bell Center for Regenerative Biology OTHER PARTNERSHIPS throughput DNA template production MBL courses. & Tissue Engineering: The Brown-MBL Partnership fosters and sequencing facility in the MBL’s Cell, developmental and regenerative research collaborations between Josephine Bay Paul Center. AWARD-WINNING SCIENCE biology; tissue engineering and scientists from the MBL and Brown Among the scientists with a significant regeneration University, and includes undergraduate The Marine Resources Department is an affiliation with the MBL (scientists, training, a joint graduate program, advanced facility for maintaining, course faculty and students) are 55 Cellular Dynamics Program: graduate training grants, and culturing, and providing aquatic Nobel Prize winners (since 1929); 118 Microscopy, imaging and cellular collaborative research programs. organisms essential to biological, Howard Hughes Medical Institute dynamics; cellular physiology and biomedical, and ecological research. investigators, early career scientists, biocurrents; biosensors The Woods Hole Consortium is a research international researchers, and and educational alliance between the National Xenopus Resource: breeds and professors (since 1960); 202 Members Program in Sensory Physiology & MBL, Woods Hole Oceanographic maintains Xenopus (frog) genetic stocks; of the National Academy of Sciences Behavior: Sensory biology (especially Institution, and Woods Hole Research provides training in Xenopus husbandry, (since 1960); and 178 Members of the vision, balance, hearing); cephalopod Center. cell biology, imaging, genetics, American Academy of Arts and Sciences camouflage; animal behavior transgenesis, and genomics. (since 1960). Encyclopedia of Life: The MBL is one Whitman Center for Visiting of 13 Global Partner institutions in FINANCES AFFILIATION WITH THE UNIVERSITY Research: Collaborative research by the Encyclopedia of Life, a website 2013 Operating Budget: $40.7 million OF CHICAGO leading scientists from around the dedicated to providing online access (52% government grants and contracts; The MBL became an affiliate of the world to knowledge about the 1.9 million 20% nonfederal grants and contracts; University of Chicago on July 1, named species of animals, plants, and 5% contributions; 7% endowment 2013. This affiliation enhances both EDUCATIONAL PROGRAMS other life forms on Earth. The EOL’s earnings; 16% other) institutions’ missions of leadership MBL Courses: 25 courses providing Biodiversity Informatics Group is based Endowment: $69 million (as of 12/31/12) and innovation in scientific research advanced, laboratory-based research at the MBL. Net Assets: $110 million (as of 12/31/12)

*as of August 2013 12 MBL Annual REport 2012 Financials Report of the Treasurer

2012 continued the trend of challenging financial times for the MBL. While the balance sheet shows an increase in net assets from $107.3 million at the end of 2011 to $109.7 million on December 31, 2012, the increase is due entirely to investment market gains, a $2 million gain on the sale of some long-held private stock, and a gain of $1.2 million from the sale of surplus real estate. The results from operations show a deficit of nearly $6 million for the year. Government grant revenue declined in 2012 due to the winding down of the federal stimulus funding received during the prior few years. MBL leadership and the Board of Trustees spent most of 2012 assessing the financial situation and identifying actions to mitigate the operational cash shortfalls that have plagued the MBL in recent years.

On the positive side, 2012 saw the completion of the Catalyst Capital Campaign, which raised a record amount of funding for the MBL. This comprehensive campaign provided sup- port for MBL programs, operations, physical plant, and endowment. The value of the MBL’s endowment rose from $54.9 million at the end of 2006 to $69.4 million at the end of 2012. The physical plant is in very good condition and the severe storms that impacted much of the East Coast during the year left the MBL largely unscathed.

FUTURE EXPECTATIONS

It has become clear that the growth initiatives of the past several years have strengthened the scientific staff of the MBL and improved the physical infrastructure on campus, but that the business model has not kept pace. By the end of 2012, MBL leadership and the Board had begun to explore the potential of affiliating with a strong research university partner in order to stabilize the financial position of the MBL, and also to open new opportunities The financial statements of the Marine Biological Laboratory for global scientific and educational initiatives. Following several months of discussions, an for the fiscal year ending December 31, 2012, were audited by Grant Thornton LLP. affiliation was formed with University of Chicago on July 1, 2013. With the success of the Catalyst Campaign, the highly acclaimed research and educational programs currently in Complete financial statements are available on request from place, and the opportunities afforded by the affiliation, the MBL’s future looks bright. Financial Services Chief Financial Officer MBL 7 MBL Street Paul R. Dupee, Treasurer Woods Hole, MA 02543

14 MBL Annual REport 2012 Statements of Financial Position December 31, 2012 and 2011 Statement of Activities and Changes in Net Assets Year ended December 31, 2012

Temporarily Permanently Assets 2012 2011 Unrestricted Restricted Restricted Total Operating support and revenues: Government grants $ 20,544,313 $ - $ - $ 20,544,313 Cash and cash equivalents $ 1,332,031 $ 1,990,094 Non-Federal grants and contracts 3,982,418 900,000 - 4,882,418 Cash held for others 430,483 298,619 Laboratory rentals 1,451,710 - - 1,451,710 Accounts receivable, net of allowance for Tuition, net 718,267 - - 718,267 doubtful accounts of $35,000 in 2012 and 2011 697,520 825,062 Fees for conferences and services 4,365,749 - - 4,365,749 Contributions receivable, net 3,228,301 7,327,936 Contributions 2,120,415 1,842,329 1,372,503 5,335,247 Receivables due for costs incurred on grants, contracts 3,872,180 4,881,482 Investment income (loss) (7,225) 6,268 - (957) Inventory and prepaid expenses 517,660 637,773 Investment earnings used for operations 596,625 2,653,024 - 3,249,649 Investments, at fair value 72,103,586 63,630,089 Other revenue 744,717 - - 744,717 Charitable remainder trusts 554,089 251,319 Present value adjustment to annuities (15,568) 251 - (15,317) Change in donor intent 4,245,789 254,211 (4,500,000) - Plant assets, net 68,772,203 71,530,516 Net assets released from restrictions 5,297,809 (5,352,730) 54,921 - Other assets 919,143 877,960 Total operating support and revenues 44,045,019 303,353 (3,072,576) 41,275,796 Total assets $ 152,427,196 $ 152,250,850 Expenses: Research 27,696,077 - - 27,696,077 Liabilities and Net Assets Visiting research 3,930,669 - - 3,930,669 Education 9,744,455 - - 9,744,455 Accounts payable and accrued expenses $ 3,125,863 $ 3,382,042 Conferences, housing and dining 3,823,871 - - 3,823,871 Future program resources 430,483 298,619 Other expenses 2,032,313 - - 2,032,313

Deferred income 1,985,976 2,059,426 Total expenses 47,227,385 - - 47,227,385 Annuities and unitrusts payable 516,119 504,026 Advances on contracts - 507,267 Change in net assets before Line of credit - 250,000 non-operating activities (3,182,366) 303,353 (3,072,576) (5,951,589) Interest rate swap 6,220,676 6,302,139 Non-operating revenue (expense): Liability for post-retirement benefits - 439,147 Change in value of interest rate swap 81,463 - - 81,463 Long-term debt 30,415,000 31,185,000 Post retirement - related charges other than periodic pension costs 488,445 - - 488,445 Total liabilities 42,694,117 44,927,666 Gain on sale of property 1,222,052 - - 1,222,052 Contributions for plant 13,246 1,000 - 14,246

Commitments and contingencies Change in net assets from non-operating activities 1,805,206 1,000 - 1,806,206 Net assets: Unrestricted 40,438,285 38,950,341 Investment income and gains (losses): Net investment income and gains 2,887,666 6,917,261 - 9,804,927 Temporarily restricted 18,105,918 14,111,391 Less investment earnings used for operations (596,625) (2,653,024) - (3,249,649) Permanently restricted 51,188,876 54,261,452 Deficiencies in historical values 574,063 (574,063) - -

Total net assets 109,733,079 107,323,184 Excess investment income 2,865,104 3,690,174 - 6,555,278

Total liabilities and net assets $ 152,427,196 $ 152,250,850 Change in net assets 1,487,944 3,994,527 (3,072,576) 2,409,895 Net assets, December 31, 2011 38,950,341 14,111,391 54,261,452 107,323,184

Net assets, December 31, 2012 $40,438,285 $18,105,918 $51,188,876 $109,733,079

MBL Annual REport 2012 15 GIFTS

16 MBL Annual REport 2012 Report of the development committee

What a year! In 2012 we raised $13,511,207 in research foundation and private support, and completed the most ambitious fundraising campaign that the MBL has ever undertaken—the $125 million Catalyst Campaign. Thanks to our trustees, overseers, devoted scientists, alumni, friends, foundations, and staff, I am thrilled to report that we not only met, but exceeded our campaign goal, raising more than $136 million between June 2006 and December 2012 for MBL programs! The number itself is remarkable, but the fact that we surpassed our goal during a challenging economy can only be described as amazing. Thank you to everyone who made this achievement possible.

In 2013, with the 125th anniversary of the MBL, we look to honor the past and celebrate the future of this amazing institution. For 125 years, the MBL has been at the forefront of biological discovery. Today, we are poised to continue making extraordinairy discoveries and significant advancements.

Throughout our long history, the MBL has relied on the strength of its community to lead the way forward. The success of the Catalyst Campaign is a wonderful example of this important partnership. On behalf of the Development Committee, the Catalyst Campaign Executive Committee, MBL students, scientists and staff, I offer my deepest thanks to the individuals, companies and foundations listed here.

— Jeffrey Pierce, Chair

MBL Annual REport 2012 17 CATALYST CAMPAIGN DONORS The individuals, foundations, and corporations listed here gave generously and contributed to the success of the Catalyst Campaign between June 1, 2006 and December 31, 2012. We are most grateful for their support.

$15,000,000+ $500,000 - $1,000,000 Sally Cross Howard Hughes Medical Institute Anonymous (1) Arthur Vining Davis Foundations Porter W. Anderson, Jr. Gerald D. and Ruth L. Fischbach $10,000,000 - $15,000,000 Davis Educational Foundation The Friendship Fund Massachusetts Life Sciences Center The Charles Evans Foundation Genentech Foundation John W. and Valerie Rowe The Grass Foundation Sibyl R. Golden Martin and Audrey Gruss Gordon Gund $5,000,000 - $10,000,000 Robert and Margot Haselkorn Harken Foundation Millicent Bell Susan and David Hibbitt J. Woodland Hastings Estate of Laura Hunter Colwin William and Lauren Huyett The William Randolph Hearst Foundation Dart Neuroscience LLC Shinya and Sylvia Inoué Kurt J. and Rhoda S. Isselbacher John D. and Catherine T. MacArthur Foundation Futaba Koike George Frederick Jewett Foundation Alfred P. Sloan Foundation The Gruss Lipper Family Foundation Barbara Woll Jones Estate of Margery J. Milne Jane C. Kaltenbach-Townsend $1,000,000 - $5,000,000 The David and Lucile Packard Foundation H.F. and Marguerite Lenfest Anonymous (2) Alice Shure Walter E. and Shirley A. Massey The Bay and Paul Foundations William Speck and Evelyn Lipper Nikon Instruments, Inc. Frederick and Christine N. Bay Saul Pannell and Sally Currier Burroughs Wellcome Fund $100,000 - $500,000 The Pfizer Foundation Paul R. Dupee, Jr. Anonymous (2) Jean W. Pierce The Ellison Medical Foundation The American Society for Cell Biology Jeffrey and Margaret Pierce Hermann Foundation, Inc. David and Sandra Bakalar Robert A. Prendergast W.M. Keck Foundation Bruce Beal and Francis Cunningham Walter J. and Marjorie B. Salmon George W. and Harmon H. B. Logan Joan R. Berman Cecily C. Selby Ambrose K. and Lili Monell Norman and Diane Bernstein Simons Foundation Gordon and Betty Moore Foundation Gary G. Borisy and Sally M. Casper Society for Neuroscience George and Kathy Putnam Margaret C. and Francis P. Bowles Albert J. Stunkard and Margaret S. Maurin Patricia and Charles Robertson Ed and Amy Brakeman Christopher M. and Susanna B. Weld Charles and Phyllis Rosenthal Estate of David Bulloch Wellcome Trust The G. Unger Vetlesen Foundation Frank M. and Julie S. Child John S. and Judith H. Willis The Clowes Fund, Inc. The Winkler Family Foundation Molly N. Cornell Alfred M. and Joyce V. Zeien Martha W. and William C. Cox Thomas and Geraldine Crane

18 MBL Annual REport 2012 Campaign Executive Committee Porter Anderson Charles Robertson Sally Currier Charles M. Rosenthal Jeffrey Pierce, Catalyst Campaign Chair Richard B. DeWolfe Christopher Weld John W. Rowe, Chairman of the Board Paul R. Dupee Gary G. Borisy, President and Director Margot Haselkorn Pamela Clapp Hinkle, Director of Development and External Relations

$50,000 - $100,000 The Seth Sprague Education and Charitable Raymond E. Stephens Anonymous (1) Foundation William H. Telfer, Ph.D Alison and Robert Ament Ann Sutherland and Raymond Keller Elaine Troll American Federation for Aging Research Gerard L. and Mary Swope Byron and Joyce Waksman American Society for Biochemistry and Gerald and Ann Weissmann Joan Wheeler Molecular Biology Carl Zeiss MicroImaging, Inc. Alice S. Whittemore and Joseph B. Keller Anderson-Rogers Foundation, Inc. World Health Organization Beech Tree Trust $25,000 - $50,000 Joshua and Lisa Bernstein Anonymous (2) $10,000 - $24,999 Anonymous (4) Alexis Borisy and Lia Meisinger Estate of Edward A. Adelberg Louise Adler Cabot Family Charitable Trust American Association for the Advancement Constance M. and Dean C. Allard Margaret Clowes of Science Peter B. and Margaret T. Armstrong Mary and Jonathan Conrad American Society for Reproductive Medicine Elizabeth Heald Arthur The Nature Conservancy Center for Molecular Biology David Baltimore and Alice Huang Elizabeth M. Davis David and Patty Cohen Patricia Ann Barlow John E. and Judith F. Dowling The Company of Biologists Limited John and Lisa Batter Saint Michael and All Angels Espiscopal Church Richard B. and Marcia H. DeWolfe Bauer Center for Genomics Research John E. Fetzer Institute, Inc. The Dorr Foundation George and Harriet Berkowitz Patrick Gage and Irina Wilson The Eppley Foundation for Research, Inc. George H. Billings Marianne and Eric Billings Gibson Associates, Inc. Peter and Ginny Foreman Gail Boettiger Estate of William T. Golden Bruce and Barbara Furie Richard G. Kessel Thomas F. and Virginia C. Gregg Cardinal Brook Trust Laurie J. Landeau and Robert J. Maze Caroline Harwood and E. Peter Greenberg Eloise E. Clark Hans Laufer M Howard and Frances F. Jacobson James M. and Ruth C. Clark Cobalt Technologies Inc. Merck & Company Inc. Kenneth and Nelda Johns Elsimar M. Coutinho Ronald P. and Karen E. O’Hanley Stephen M. and Cynthia R. Krane Bernice Cramer and Paul Friedman Keith R. Porter Endowment for Cell Biology Inga Mahler William Townsend Porter Foundation The Andrew W. Mellon Foundation Alexander T. Daignault Julia S. Rankin Marianne F. Milkman Joel P. and Ruth H. Davis Linda A. Deegan and Christopher Neill Elise and Stanley Rose Richard S. and Susan G. Morse Diarmaid and Margaret M. Douglas-Hamilton Vincent J. Ryan and Carla E. Meyer Edward and Linda Owens Phillip B. Dunham and Gudrun Bjarnarson The Schooner Foundation Elaine Pierson-Mastroianni Douglas R. and Virginia M. Shanklin Marius A. Robinson Gordon W. and Joyce M. Ellis James and Mary Sharp Edward D. and Nancy M. Salmon Federation of American Society for Experimental Biology Martha S. Ferguson Society for Developmental Biology, Inc. Shawmut Design and Construction Seiichiro Fujimoto

MBL Annual REport 2012 19 CAtalyst Campaign Donors

Charles M. Ganson Pfizer Inc $5,000 - $10,000 David C. and Patricia S. Gadsby Sallie A. Giffen Robert W. and Carroll C.D. Pierce Anonymous (3) James N. and Nancy Galloway Robert D. and Anne E. Goldman Thomas D. and Patricia S. Pollard Garland E. Allen Craig and Nancy Gibson Susan P. and Thomas Goux Presbyterian Church of Chestnut Hill Douglas F. and Sarah H. Allison Prosser Gifford Mary Greer James L. and Helene M. Anderson Alfred L. and Joan H. Goldberg Linda L. and Stephen A. Greyser Harris and Jeanne Ripps Associates of Cape Cod Barbara L. Goldsmith Foundation Hilde Rosenthal Austin Foundation, Inc Timothy H. and Mary Helen M. Goldsmith Howard H. Hiatt Priscilla F. Roslansky Moïse H. Goldstein, Jr. John and Olivann Hobbie Edward S. and Susie W. Rowland Robert G. and Harriet Baker Robert and Trudy Gottesman Hub International New England Joan V. and Gerald S. Ruderman William H. and Paula Beers Mary W. Graham Carmela J. Huettner Kerry S. Bloom and Elaine Yeh Philip Grant Hugh E. and Frances M. Huxley Andrew E. Sabin Gloria S. Borgese Katherine Graubard and William H. Calvin Santa Monica Seafood Max M. Burger Paul R. Gross Hans and Elizabeth Jannasch Paul R. Schloerb Butler’s Hole Fund Audrey M. Schneiderman Joshua and Melissa Hamilton Freda Kaminer Harriet F. Segal R. Andrew and Barbara Cameron Hamilton Thorne Inc. Mayer and Morris Kaplan Sholley Foundation, Inc. Malcolm O. and Jeanne Campbell Susan Harding Hawkins A. Sidney and Alice Knowles Esther Simon Charitable Trust James F. Case The William H. Harris Foundation Tsoi Kobus & Associates, Inc. Society of General Physiologists Richard L. and Alice M. Chappell William A. Haskins Masakazu Konishi Mitchell L. Sogin and Laurel Miller Paul N. Chervin Diane E. Heck Hans and Donna Kornberg Melvin and Evelyn Spiegel The Church of the Messiah John G. Hildebrand and Gail D. Burd Dusan Stefoski and Craig Savage Estate of Octavia C. Clement James Hildreth Lassalle Fund Marjorie E. Steinberg Alexander W. Clowes and Susan E. Llewellya W. Hillis and Paul Colinvaux Anna and Tom Lawson Alfred and Dorothy A. Stracher Detweiler Timothy T. Hilton Paul B. Lazarow Ann E. Stuart and John W. Moore Mary Clutter Pamela C. and Gregory J. Hinkle Ethan and Lisa Lerner John F. Swope William and Susan Cobb John and Olivann Hobbie Monique Liuzzi Sautter Marion Cohen Joseph Hoffman and Elena Citkowitz Werner R. and Birgit R. Loewenstein Technic, Inc. Catherine B. Cramer Howard and Sybil Holtzer Laszlo Lorand Edward and Louise Tsoi Stephen D. and Beth G. Crocker Henry Hornblower Fund, Inc. Mervyn J. Turner The Charles Evans Hughes Memorial Dave Madden and Jennifer Segal Madden Eric H. Davidson Foundation, Inc. John R. and Catherine B. Mandile USNC-IBRO Committee Leyla deToledo-Morrell David and Miriam J. Mauzerall Joseph and Sarah Dowling Gary and Susan Jacobson Nawrie Meigs-Brown and David Brown Ronald D. Vale and Karen Dell Hugh and Beverly Ducklow Mary D. Janney Jerry and Lalise Melillo John J. and Frederica W. Valois Robert A. and Diane B. Jaye Matthew Meselson and Jeanne Barbara E. Ehrlich Guillemin-Meselson Waksman Foundation for Microbiology Herman N. Eisen and Natalie Aronson Morris J. and Shirley Karnovsky Leonard and Eve Warren Stephen and Lois Eisen Allan Keith The New York Times Company Washington & Jefferson College Doris W. Epstein Malka Keynan Foundation Earl H. Weidner Dan Kiehart and Patricia Lee Saylor R. Dana Ono and Anne Wagner Annette L. Williamson and A. Verdi Farman-Farmaian Edward A. and Kathryn F. Kravitz Michael McColm Michael Fenlon and Linda Sallop The Saul & Gitta Kurlat Charitable Foundation Arthur B. Pardee and Ann B. Goodman Dyann and Peter K. Wirth The Flatley Foundation Bertha Person Barbara Wu and Eric Larson Pamela and David Follett Homer W. and Mary C. Lane George and Pam Friedman Marnie and John Langford

20 MBL Annual REport 2012 CAtalyst Campaign Donors

Leonard and Ruth Ann Laster Alan R. and Phyllis J. Silver David S. and Nancy C. Babin Laurence P. and Katharine B. Cloud Deena and Alan Laties Society for the Study of Reproduction Katherine Bachman and Robert Fleming Jewel Plummer Cobb Jennifer Letitia Jennifer Sorenti and Ralph Burns Shawky Z. Badawy Adam E. Cohen Joseph Levine and Steven A. Cadwell James A. and Annamma Spudich John M. and Carol T. Baitsell Carolyn Cohen William W. and Noelle Locke Alan B. and Sala Steinbach Hope Baker Sarah Cohen Richard Lounsbery Foundation The Judy and Michael Steinhardt Talbot Baker R. John and Joan M. Collier William K. and Winnie C. Mackey Foundation David Baltimore and Alice S. Huang Rita R. and Jack H. Colwell John R. and Catherine B. Mandile Carol G. and Joseph T. Stewart Barbo’s Furniture Community Counselling Service Edward Barker Harry F. Conner and Carol E. Scott-Conner March of Dimes Foundation Carolyn and John Stimpson The Barno - Montgomery Family Trust Charles and Beatrice Cooper Michael T. and Jean M. Martin Gregg Strumwasser, Esq. Susan R. and Daniel T. Barry Roberta Corliss Joe L. Martinez and Kimberly Donata O. and William M. Sugden David C. Beebe Jeffrey T. Corwin Smith-Martinez Foundation for Sustainability and Jerry A. Bell and Mary Ann Stepp William G. and Sally Coughlin R. K. Mellon Family Foundation Innovation Michael V.L. and R. Suzanne Bennett Joseph T. Coyle, M.D. Jason and Jennifer Meyers Andrew and Ursula Szent-Györgyi Celeste A. Berg and Mark Kot Marilyn E. and J. Sterling Crandall Gerrish H. Milliken Foundation Robert O. and Jean Bigelow Charles M. Crane and Wendy Breuer Ralph and Muriel Mitchell Edwin and Heather Taylor Jonathan and Elizabeth Billings Michael J. Cronin Timothy Mitchison and Christine James G. and Mary Elizabeth Townsel William C. Black Gorham & Joan Cross Foundation Marie Field Thomas P. and Jane W. Bleck Dorothy I. Crossley Merle Mizell The Irving Weinstein Foundation, Inc. Neil G. Bluhm Anthony J. and Diane M. Cutaia Kathrin R. Winkler and Angus Campbell The Bodman Founation Hiroko Naka Nancy Wolf and Aric Greenfield David A. and Marla Bodznick Giuseppe and Mirella V. D’Alessio Toshio and Kyoko Narahashi Kendall B. and Karin A. Bohr George Q. Daley The National Organization for Barbara K. Zeisler Thomas C. and Elizabeth D. Bolton Devra Lee Davis Hearing Research Joshua J. Zimmerberg and Elayne Bornslaeger-Bednar and Michael S. Joseph P. Day Peter A. Nickerson Teresa L.Z. Jones Bednar Robert C. and Ellen L. deGroof Ronald Zweig and Christina Rawley Robert W. Brackenbury and Linda Parysek Martha Bridge Denckla OceanSolution Eleanor D Bronson Susan Dillmann Donald D. and Linda W. Brown Jonathan O’Herron $1,000 - $5,000 Allison Doupe David Bryant and Donna Nimec The Doverspike Family Fund James L. Olds and Robin E. Buckley Anonymous (11) Elisabeth M. Buck Arthur B. and Roberdeau C. DuBois Michael R. Paine Conservation Trust Donald A. and Barbara R. Abt Darryl A. and Janet Buckingham David and Sherri Durica Parkinson’s Disease Foundation Blaise Aguera Arcas and Adrienne L. Fairhall The Bufftree Foundation AHA Consulting Engineers, Inc. Thoru and Judith Pederson John E. and Sally S. Burris Nina Stromgren Allen Cynthia D. Eaton Frank Press Amy K. Butler Michael and Marie Angelini George P. and Sally Edmonds Robert and Lynne Angerer Robert Eimstad Michael and Diane Rabinowitz Ronald Calabrese and Christine Cozzens James and Cynthia Angstadt Paul Elias and Marie Lossky Peter W. and Elizabeth Renaghan Graciela C. Candelas Kiyoshi Aoki David and Lois Epel Reynolds Family Foundation Cape Cod Five Cents Savings Bank Aquatic Habitats, Inc. David A. Epstein Monica Riley D. Bret Carlson Bryan and Donna Arling Karen Ann Epstein John and Joan B. Ripple Frank C. and Sheila G. Carotenuto Clay M. Armstrong and Clara Anne Etgen Center for Reproductive Rights Jack and Rosalyn Rosenbluth Franzini-Armstrong Evolved Machines, Inc. Joseph A. Cerro James Ross Samuel C. and Elizabeth F. Armstrong Chenbei Chang Carol Arnosti and Andreas P. Teske Associates Fairbanks Foundation, Inc. Sallie Chisholm Amy Segal and Andrew Blum Farooq Azam Falmouth Toyota Scion Michael P. Sheetz Ann P. and James M. Cleary David and Doris L. Fausch

MBL Annual REport 2012 21 CACAtalysttalyst C Campaiampaiggnn D Donorsonors

Daniel E. Feldman Gary G. and Nancy Hayward John R. and Andrea O. Lakian Douglas A. Melton and Gail O’Keefe Max and Martha Fink Marian B. and Edmund Hazzard George M. and Sylvia T. Langford Martin Mendelson Rachel D. Fink Joseph Heitman Catherine C. and John Lastavica Merck Sharp & Dohme Mark C. and Martha Fishman Gordon E. Hering Holly A. Lavoie Irwin and Ruth Merkatz Flagship Roofing & Sheetmetal Co., Inc. Patricia D’Amore and Ira Herman Edward R. and Gloria A. Leadbetter Melanie and Klein Merriman Stephen E. Follansbee Stephen and Gabrielle Highstein Joel A. and Gail Leavitt Mid-American Water of Wauconda, Inc. Jean-François and Rachel Formela Richard W. and Susan D. Hill John Lechner and Mary Higgins Stanley Mierzwa John P. Fowler Charles A. and Young Hobbie Matt K. Lee Ricardo and Mela Miledi Thomas O. Fox Paul and Patricia Hogan Marian E. LeFevre David A. and Virginia I. Miller Lawrence C. Fritz Kenneth Holden and Frances C. McGuire Millie Lerner Thomas J. and Susan R. Miller John J. and Avery T. Funkhouser Kelly R. Holzworth Jack and Francine Levin Millicent and Eugene Bell Foundation Ruth E. Fye Charles S. and Marianita Hopkinson John E. and Natasha C. Lisman Mills Family Charitable Foundation Alan F. Horwitz Rodolfo R. and Gillian K. Llinás Hideo Mohri Harold and Ruth S. Gainer Robert Howarth and Roxanne Marino James E. and Caroline S. Lloyd Ronald B. Moir and Toni Czekanski William Galardi Ronald R. Hoy and Margaret C. Nelson Irving M. and Huguette London James F. and Lisa M. Mooney Joseph G. Gall and Diane M. Dwyer Lindsey Humes Loomis, Sayles & Company Richard D. Mooney Frank P. and Joyce Gallagher Tim and Mary Hunt Michele Lorand and Paul Ringel Sean and Tracy Moore Branko J. Gerovac Charles and Berit Hunter Joanna E. Lowell Robert and Maura Morey Marion R. Ginsberg Hunter College Victoria H. and Francis C. Lowell James T. Morris Helen W. Gjessing Thomas J. and Nicole Hyne Howard W. Lowy H. Edward Muendel Michele Gallati Glasgow The Lubin Family Foundation Saylo and Elizabeth Munemitsu Daniel E. Goldberg David Isenberg and Paula Blumenthal Anna Lysakowski Joan M. Murray Mary L. Goldman and Debra Weiner Kurt J. and Rhoda S. Isselbacher Mary U. Musacchia and James E. Faber Paul K. Goldsmith Robert E. and Marjorie J. Mainer Margaret Ann Goldstein Laurinda A. Jaffe Mary Malloy Knute J. Nadelhoffer and Barbara Billings Bruce and Mary Goodman Anthony A. James Phillip B. and Yvonne R. Maples Harold Nash Gary Gorbsky Ernest G. and Pauline B. Jaworski James Marcello Enrico Nasi and Maria E. Gomez Joel S. Gordon The Joby Foundation Inc. Julian B. and Priscilla K. Marsh Philip and Sima Needleman Peter Gottlieb Robert K. Josephson Martha’s Vineyard Savings Bank Phil Nehro The Julie Gould Fund Sally S. and Ramsey E. Joslin Charitable Fund John M. and Linda C. Nelson Michael J. and Rebecca H. Greenberg Lowell V. and Ann J. Martin Jeanne M. Nerbonne Newton H. and Evelyn Gresser Darcy B. Kelley W.B. Mason Neurobiotex Inc. Alexandre Grigorovitch R. Emmet and Gail Kenney Dale and Edward J. Mathias New England Biolabs, Inc. Gustav Grosch and Jane Kulesza Alexey L. Khodjakov Peter Matthiessen Santo V. Nicosia Larry Grossberg David Kicklighter Edwin McCleskey Bryan D. and Elaine K. Noe Barbara Grossman David Kleinfeld William M. and Elaine D. McDermott Dana T. Nojima Carlos Gual James J. Knierim Richard B. McElvein Estate of John W. Norman Mark Koide Dianne McFarlane Normandie Foundation Valerie A. and Robert R. Hall Samuel S. and Sumi M. Koide McKinsey & Company, Inc. Catherine N. Norton Zach W. Hall and Julia A. Giacobassi David B. Kriser Foundation, Inc. Jane A. McLaughlin Jean Halvorson Kiyoshi Kusano William N. and Marianne W. Mebane Penelope S. Hare Alan M. and B.J. Kuzirian Shuya Meguro Kenneth Olden Glenn W. Harrington Dennis E. Meiss Rudolf and Nannette D. Oldenbourg John P. and Mary E. Harrington Ezra and Aimlee D. Laderman Heidi Gifford Melas-Kyriazi George D. and Bernice Pappas William Harrison Kevin L. and Sallie D. Lake Richard P. and Katherine W. Mellon The Parasol Tahoe Community Foundation

22 MBL Annual REport 2012 CAtalyst Campaign Donors

John W. and Cecilia H. Pehle Robert Rubenstein and Marie Raftery Margaret W. Taft Leslie J. and Elizabeth M. Wilson Robert H. and Pamela D. Pelletreau John D. and Jean M. Rummel Anne C. and Robert Talley John and Patty Wineman Nancy Pendleton Michael G. Ryan and Linda A. Joyce G. Pran Talwar Wise Living Polly Penhale Carol Ann Ryder Martha E. Tarafa Jonathan B. and Beatrice Wittenberg The Pennyghael Foundation D. Lansing and M. Margaret Taylor W. Redwood and Mary Wright Ralph F. Peo Foundation, Inc. James Sailer and Cass Conrad Emil D. and Eleanor H. Tietje Frederick S. Peters Joseph W. and Jean M. Sanger John Tochko and Christina Myles-Tochko Linda and William Zammer Foundation Fund Stanley A. and Virginia V. Peterson Noriyuki Satoh Victor K. and Julia Tolkan Michael J. and Naomi Zigmond Victor Petkauskos Anne W. Sawyer Laurence C. Torsher Steven and Dorothy Zottoli Robert B. and Evelyn M. Pharr Howard K. and Ethel L. Schachman Noah and Janet B. Totten Frederick H. Pierce Mari Takasu Scheiffele and Peter Scheiffele David M. and Jeanne D. Travis Other Gifts Harmony Glazebrook Pittet Jay Schippers Kenyon S. and Joan Tweedell Anonymous (45) William K. Plunkett Herbert Schuel Thomas H. Aal Jeanne S. Poindexter Scott M. Schulte Volker and Mary Ulbrich Robert F. Aarstad Harvey B. and Bette S. Pollard Laura Segal and Matthew Petrie Unitarian Universalist Fellowship Mark Abad The Polly Hill Arboretum Sol Sepsenwol of Falmouth Mark and Joan Ann Abbett Robert Greenberg and Carol Pooser Harriet S. and Howard E. Shapiro Jeanine A. Ursitti Jayne Abbott Joan Abbott Jahn Pothier Gaius and Eleanore Shaver Laurence and Catherine Abbott John S. Price Kate Lake Shaw Toby J. Velte John Aber Promega Corporation Alexandra E. Shea Gilbert Verney Foundation Ronald F. Abercrombie Gordon and Claire Prussian Guojun Sheng Estate of Claude A. Villee, Jr. Irwin Abrams Ronald J. Przybylski David and Marilyn Sheprow Susan Volman Robin W. Ackroyd Keith Purpura L. David Sibley Leslie B. Vosshall James A. and Candy Adams Felicity A. Putnam Arthur and Frances Silverstein Josephine Adams Bonnie W. Simon Carol A. and Stephen P. Wagner Kristina M. Adams Waldorf James P. and Joan M. Quigley Roger D. and Carol J. Sloboda Warburg Pincus Nii A. Addy Greenfield and Patrica Sluder Gary Ward and Zail Berry David E. Adelberg Irving W. Rabb Stephen J. Smith and JoAnn Buchanan Richard H. and Doris Waring Lea H. Adelstein Edward Rastetter and Karen Hendrickson John and Evelyn Steele Jane E. Warshaw Abdulbaki Agbas Regional Technology Development Center Andreas C. Stemmer Clare M. Waterman Goran Agren of Cape Cod, Inc. William K. and Jane B. Stephenson Annemarie Weber David C. and Jean E. Ahearn Richard and Bonnie Reiss Paul A. and Pamela T. Steudler Janis C. Weeks and William M. Roberts Helen Ahearn William S. and Catherine A. Reznikoff Edward and Anne Stimpson Frances Weiffenbach Kamran Ahmed The Rhad Trust Steven F. Stoddard Edward and Dia Weil Jonas Ahnfelt-Ronne Fred and Gloria Rhoades Elijah W. Stommel and Jasmin Bihler The Adam J. Weissman Foundation Marc Airhart Emre R. Aksay Richard H. and Margaret Rhoads Jerome and Catherine Strauss Andrew Weissmann Merryl Alber Paul and Susan Rhodes Antony O. W. Stretton John and Barbara Weyand Daniel and Nancy Albert Philip S. and Anne Robertson Boris Striepen Clifton R. Wharton MaryAnn Albertine David and Nan Robinson Sugar, Friedberg & Felsenthal Byron and Anita Wien David F. Albertini The Rockefeller University Press Mutsuyuki and Shirley M. Sugimori Eric Wieschaus and Trudi Schupbach Beverly Alexander Lawrence C. and Victoria H. Rome William C. and Karen A. Summers Roland L. and Nancy Wigley Douglas G. and Kathleen Alexander Joel L. Rosenbaum and Connie Drysdale Raquel and Maurice Sussman Clare M. Wilber Marselle O. Alexander-Ozinskas Paul and Melissa Ross Kelly J. Suter Clayton A. and Arlene Wiley Dianne M. and Thomas V. Allen William N. Ross and Nechama Lasser-Ross Helen and Albert Wilson Donald G. and Judith Allen

MBL Annual REport 2012 23 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

Rosalind J. Allen Spencer L. Baird and Judy P. Spencer Donald and Virginia M. Belanger Eric Blum Mark and Mary Anne Alliegro Clare V. Baker Marlene and George Belfort Joanne Blum Peggy and Fred Alsup Nicholas J. Baker Brittany J. Belin Scott Blum Francisco Alvarado Arax Balakian Curtis Bell Edward M. Blumenthal David S. and Beverly K. Ament Joseph Baldino David M. Bell Thomas Blumenthal Penny S. Amy Thomas O. Baldwin and Miriam M. Ziegler Albert Belman Amy Lynn Boardman and Robert Purcell Collins Anchor Print & Design Ganesh Balgi Andrew Belmonte Richard S. Bockman C. Ronald Anderson Emily P. Balskus Elia Ben-Ari Susan and Hank Bode Ruth Anderson Michael J. Baltzley Jim and Jessica Benjamin Brenda J. Bodian W. Banks Anderson Rolf and Edda Bandle Thomas L. and Mary Jo Benjamin Elinor W. Bodian Andor Technology Molly Bang Kenneth Bento Susan E. Boehme William DeWitt Andrus, Jr. Matthew R. Banghart C. John and Elizabeth B. Berg David Boettiger Agnes Lee Ang James D. Bangs Christopher Berger Andrew W. Bollen Lynne and Robert Angerer Banner Environmental Services, Inc. Jane A. Berger and Roger O. Gittines Lonegan Bonczek and Mary Makela Scott L. Applebaum Gary Banta Joanne E. Berger-Sweeney Edward Michael Bonder Erin and Barry Aprison Jianxin Bao Everett and Anne Bergman Mark R. Boothby Ricardo C. Araneda Madelyn M. Baran Dominique C. Bergmann Elizabeth A. Booth-Gauthier Irina Arkhipova Robert Barker and Marla Schay Baker William H. Bergstrom David Borden Alexandra Armstrong James Barlow Gerald Bergtrom Andrew and JeanneMarie Borgese Ellen P. and Richard Armstrong John E and Helen V. Barnes John and Barbara Berkenfield Richard T. Born Gwyneth Armstrong Michael J. Barnes Selim Berker and Carol Trauner Lawrence and Sandy Bornstein William F. Arndt, Jr. Nancy Barnett Ari Berkowitz Chiara Boschetti Elizabeth I. Arnold Barnstable Intermediate School Edward R. and Doreen A. Berling Sarah W. Bottjer Mike and Elana Arnold Harold and Marion Barr Joel C. Berman Joyce Bottum Senthil Arumugam Ernest Barreto Judith and Michael Berman Willliam and Linda Bowden Shinobu Asano Dennis Barrett Joan M. Bernhard Jennifer L. Bowen Michael S. Ascher Elizabeth Barrows Elaine K. Bernstein Gwynn Akin Bowers Robert F. Ashman Diana C. Bartelt David Berreby Henry Bowers Francis T. Ashton Nelson R. Barton Viviana M. Berthoud Nancy J. Bowers Duncan P. and Dorothy D. Aspinwall Alexandra L. Basolo Neil Bettez and Gretchen Gettel Courtney M. Bowker Vijay H. Aswani Lawrence E. Bathgate Olive C. Beverly Barbara C. and John F. Boyer Jelle Atema Barbara-Anne Battelle and James P. Alligood Francisco Bezanilla Peter and Joan Boyer Mary S. Atkisson and Joel White George H. Batterson Maja I. Bialecka-Fornal Anne Weinberger Bracy Barbara Atwood William Battin Stanley J. Bielen Albert and Deborah Bradley June and Joseph Atwood Norman Bauman George H. and Wendy Bigelow Scott T. Brady Nathaniel B. Atwood and Susan A. Parkes David H. Beach Michael Bihari Thomas and Janet Brady James W. Atz Cheryl Beano Jay B. Bikoff Margaret Brandon W. Lena Austin Nancy Bearss and Jennifer Lindsay Allen and Joan Bildner Brigitte F. Brandriff Andrea R. Avila Michael A. Becker Maks and Naomi Birnbach Aaron Bravman John J. and Lorraine S. Aziz John Beckerle Ethel E. Bishop Stanley G. Braxton Jonathan Beckwith Stephen H. and C. Lynn Bishop Elizabeth Breen Serapio M. Baca Mary Beckwith Shirley Bissen Catherine A. Brennan Robert L. Bacallao Matthew and Judy Beecher Dieter and Gigi Blennemann Phyllis S. Brenner Beth Bacha Deborah Y. Beers-Jones Dawn M. Blitz John Brereton Rebecca K. Baer Paul Begovac Avis Blomberg David N. Breslauer Michele and Frank Bahr Elizabeth Kujawinski Behn Gail A. Blout Charles and Helen Bresnahan Ronald C. Baird Edward Joseph Behrman Blue Spruce Technologies Inc. Elizabeth Allen Brett

24 MBL Annual REport 2012 CAtalyst Campaign Donors

Giovanna Breu George and Yara T. Cadwalader Alvarus Chan Jean Cohen John and Renee Breznak Yuping A. Cai Jane Chance Joel and Helen Cohen Anthony Briana Arthur and Booje Calfee Albert Chang Maynard M. and Doris V. Cohen Lawson W. Brigham Patrick J. and Mary R. Calie Donald C. Chang Rochelle S. Cohen and Rex Sexton Kenneth H. Brink John L. and Virginia C. Callahan Yueh Chang and Jeffery Watson Saul Cohen Bill Brinkley Linda M. Callahan The Chapin School Seymour S. Cohen Andrew Brittingham David and Janet Campbell Louise K. Charkoudian George Cohn Peter Brodfuehrer James W. Campbell Richard and Patricia G. Chartrand Robert Colasacco Burt V. Bronk Leah J. Campbell John and Sandra Checklick Barbara A. Colburn Rebecca Bronson Patricia Campbell Lu Chen Jonathan J. Cole Austin E. Brooks Ronald and Vivian Campbell Clarissa M. Cheney Annette W. and John R. Coleman Phil and Shirley Brooks Francesco Canganella Lisa Y. Chiang Katherine Coleman Laura C. Broughton Julie C. Canman Chi-Bin Chien Rebecca Coleman John and Elaine F. Brouillard Thomas Cannold Children’s School of Science, Inc. Scott L. Collins Jennifer Byrnes Brower Cynthia S. Cappello Florin V. Chirila Stephen D. Collins Barbara Brown Diana Caracino Matthew and Sarah Chisholm Beth Colt and P.K. Simonds M. Kathryn S. and Thomas A. Brown Zoe and Andrew Cardon Carol Chittenden Marc D. Coltrera and Anne L. Buchinski Stephen C. and Patricia S. Brown Kevin R. Carman A. Kent Christensen Committee for the Conservation of Melinda Browne Joanne B. Carnevale John A. Christian Horseshoe Crabs Robert H. Broyles Robert J. and Louise A. Carney Keith Christianson Elizabeth B. Conant John L. Brubacher Carol Laikin Carpenter Scott G. and Penelope P. Chrysler Susan B. Conaty-Buck Judy A. Bruguiere Edward J. Carpenter and Gerdi Weidner Magdalena B. Chrzanowska-Wodnicka Concord Christian Academy Jerí Bryant Stephen R. and Susan G. Carpenter Isabel G. Cintron-Garcia Priscilla N. and Alan Conger Gurdon S. Buck Catherine Emily Carr Ellen S. and George B. Citron Sean D. Conner Peter M. Buck Margaret W. and Samuel B. Carr Thomas and Rachel Claflin Peter and Susan Connolly Daniel H. Buckley Robert H. and Myra S. Carrier Benton C. Clark Lou-Anne Conroy Ann C. Bucklin and Peter Wiebe Ashley R. Carter George A. Clark Martha Constantine-Paton and H. Robert Horvitz John E. Buffington Robert and Evelyn Carvalho Maria E. Clark Catherine E. Cook David A. Bumcrot Lilliam Casillas-Martinez Tucker Clark Joseph A. and Betty Ann Cook Susan M. Burden-Gulley Geraldine J. Casper Vera S. Clark Ruth Cooke David R. Burgess and Rita Hogan Robert G. Cassens Charles G. Clarke Nathaniel S. and Catherine E. Coolidge Deborah M. Burgess Joseph D. Cassidy David Clarke Elizabeth Cooney Richard R. Burgess Lynne U. Cassimeris John M. and Theresa M. Clarkin Edwin L. Cooper Alan H. Burghauser Edward Castaneda Philippa Claude Frederick Cooper and Karen Gordon Ingrid Burke and William Lauenroth Maria Castello Janet Clay Gail Cooper Joseph W. Burke J. Douglas Caston Leonard Cleary Margaret S. Cooper Patrick D. and Patricia A. Burke Margaret Catley-Carlson Susan D. Clements-Rabatin Margery and Neil Cooper Arnold H. and Ruth B. Burrough James Catterall and Jean Schweis Hollis T. Cline Alice Cooperstein Elizabeth H. Burrows Robert and Penny Catzen Graeme Cloutte Marie Cordner Harold L. and Joan N. Burstyn Mary P. Cavanaugh William T. Clusin Sallie Riggs Corgan Shirley Businski Michael C. Cerio Margaret H. and Edward W. Coburn Anita Corliss William and Geraldine Butman Anthony Cerrone Eric P. and Christine Cody Bruce and Teresa Corliss Bruce E. and Patrice L. Buxton Alfred B. Chaet Mort and Susan Cohan John O. and Yuemei G. Corliss Karen Buzby Jungwon Chai Bradley and Katie Cohen Robert A. Cornell Robert David Bynum Remy P. Chait Bruce S. Cohen Trinka S. Coster Vetria L. Byrd Susan Chamberlain Elizabeth Ann Cohen Laura F. Cotlin Natalie Chambers Maryanne and Isidore Cohen Paul Cotran

MBL Annual REport 2012 25 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

Sandra Cottingham Florence H. Davidson and Leonard Kreidermacher Laneta Dorflinger Lincoln and Ruth Ekstrom Ernest F. and Keiko Couch Graham C. Davis Barbara J. Douglass Kate Eldred Yves Courtois Gregory K. Davis David L. Douglass Carlyn Ellms Pamela A. Coutchie Roger S. and Marcia Davis Dustin L. Dovala Encore of Falmouth Newcomers Patrica Cowan Rowland H. Davis David D. Dow Alvin and Hsiao Eng Vincent Cowling and Charlene R. Lewis Nigel W. and Leila Daw Karen G. Dowell Mary Engel and Nolan Hester John Crabbe Syrel Dawson Daniel Doyle Gabriella J. Engelhardt Farnham Susan W. and Nessly C. Craig Marisol de Jesus-Berrios Peter Drake Paul T. Englund and Christine S. Schneyer Melvin C. and Margery L. Crain Robilee F. Deane Julie Drawbridge Helen Mittlacher Erickson Michael and Catherine Crane Joseph A. DeGiorgis Joseph and Esther Dresner Henry Erle John D. Crawford Robert L. and Marianne S. DeHaan Barbara T. Driggers Ali Eroglu Karen Crawford Nahid S. DeLanney Charles and Kimberley M. Driscoll Barbara B. Espey Sally A. Crawford Nicholas and Elena Delbanco Cindy Driscoll Juan S. Espinosa Chris Crockett James M. Denegre Suzanne Droban Gordon C. Estabrooks Jeffrey G. Croll John and Beverly G. Densler Dorothy L. Drummey Marilynn E. Etzler John B. Cromie Katy A. Depew Arthur Dudden Thomas E. Evans Christopher S. Cronan Charles Derby Henry and Marilyn P. Dudley Robert R. and Ann Everett Helen M. Crossley Abbey E. DeRocker Mark and Lorraine S. Duewiger Arnold G. and Paula B. Eversole W. Peter Crossley David J. DeRosier and Esta B. Markin Maureen J. Dugan James F. and Ann C. Crow Richard J. DeSa Donald Dugger Donald and Jo Faber John Henry Crowe Wolf-Dietrich Dettbarn Quan-Yang Duh and Ann Comer Suzanne Fachon-Kalweit Bernard G. and Karen W. Crowell Stephan M. Deutsch Catherine Asleson Dundon Debra Ann and James M. Fadool Jeff Crowell Marie A. DiBerardino Kathleen Dunlap Patricia M. Failla Prince S. Crowell, III Charles A. and Molly A. DiCecca Allan and Linda Dunn Alexandra S. Fairfield Paul Cruikshank Suzanne M. DiChiara Ivy Dunn Peter Falco John Cunniff, M.D. Byram Dickes Nigel Stuart Dunn-Coleman Nancy L. Falk Lynn P. and Courtland B. Cunningham Michael H. Dickinson Christopher and Portia Durbin Falmouth Newcomers Club Mary Ellen M. Cunningham Paul and Martha Diehl Rita and Paul E. Dussault Jerome and Barbara Fanger James F. Cupo Virginia A. and Richard C. Dierker Charles Dwane Alan Scott Fanning Alice M. Curry Kenneth Dill Richard and Kathi Dwelle Elizabeth M. Farnham Robert A. and Donna Curtis Ellen and Fred DiMeco Anne W. Dyer Helen C. Farrington Richard D. and Virginia M. Cutler Bruce A. Diner Rhonda Dzakpasu John W. Farrington Vincent E. Dionne George Feher Ellen Roter Dirksen Brian F. Eames Peter S. Feibleman Yuri A. Dabaghian Karen L. Divelbiss Early Learning of Medfield, Inc. Andrew and Amy Feinberg Sandra L. Dabora Helen and Joseph Dixon Eastside Christian Chrurch Marta Feldmesser Veronica Dagostino Thelma D. and Jonathan S. Dixon Clifton H. Eaton and Nilde Diefenbach Marla B. Feller Stephen C. Dahl Thuy A. Doan Hoyt and Deborah Ecker Benjamin Felzer Leslie M. Dalmacio Dirk Dobbelaere Laurel A. Eckhardt Caraciolo J. Fernandes Sandra L. Damato Fred C. Dobbs Carol Bernstein Eckstein Philip and Barbara Ferro Amber L. Dance Howard T. Dodd Kenneth T. and Kathryn H. Edds Barry William Festoff Janet B. Daniels Liam Dolan Marcia Edwards Joseph R. Fetcho John and Gwen Daniels Michael F. Dolan David A. Egloff Mark Fichman and Ruth Fauman-Fichman William C. Daniels Pavel Dolezal Frank Egloff Stephen A. and Denise P. Fiedler Gregory Alan Dasch Ellen P. Donaldson Frank Egloff, II Don and Joyce Filiault Andrea d’Avella Peter J. Donaldson Judith S. Eisen Caleb E. Finch John R. and Roberta A. David Tabitha Doniach George T. Eisenhoffer, Jr. Stuart Findlay

26 MBL Annual REport 2012 CAtalyst Campaign Donors

Edward J. and Debra L. Fine Stephan and Anne Franzino Maureen A. Gannon Fred Golden Gerald Fine and Victoria LeFevre Nancy and Joseph V. Fraser David E. and Andrea Garber Heidi E. Golden Alan and Joan Finkelstein Chris and Cathleen Frederickson Amaya Garcia Costa Jerome and Gloria Golden Eric B. Finkelstein Ted and Harriet Fredman John A. Garcia Joan R. Golden Jacques C. Finlay and Sarah E. Hobbie Michael A. Freed Jose E. Garcia-Arraras Michael R. and Ellen Goldenberg Cyril V. Finnegan, Jr. Kenneth Freedman and Steven Weiss Frederic Gardner and Sherley G. Smith Robert Golder Bruce and Virginia Finnie Anne Freeman Michael R. and Mary Garfield David J. Goldhamer First United Methodist Church of Chatham Arthur and Linda Freeman Sallyann Garner Charles R. Goldman Robert S. Fischer Christopher J. Freeman Eugene and Evelyn Garnett Jerome and Eileen Goldman Lawrence K. and Atsuko Fish Hadley Mack French Patricia E. Garrett and Oliver H. Woshinsky Jonathan and Nicole Goldman Michael J. and Joann Fishbein Krystyna Frenkel Rita A. and Leslie Garrick Rina Goldman Frederick S. and Ingeborg K. Fisher Hugo D. and Anita R. Freudenthal Paul A. and Kim Garris Michael P. Goldring Laura S. Fishler Paul J. and Patricia S. Freyheit Patricia Garrity Joshua Goldstein Harvey M. and Susan Fishman Friedberg Properties & Associates Jennifer H. Garvin Kenneth H. Goldstein Ruth Alice Fitz David F. Friedman Mark Gasarch Thelma Goldstein Kevin W. Fitzpatrick Edward and Susan Friedman C. Vernon Gaw Ethan Goldstine Robert and Winifred Fitzpatrick Larry Jay Friedman Stephen E. Gellis Josef Goldufsky Thomas R. Flanagan Marvin and Rivka Friedman Alan and Kate Gelperin Gabrielle B. Goode John W. Fleeger Ronny Frishman Elizabeth L. George Charlotte Goodhue John Fleischman Matthew Frosch Mary George Vicki L. Goodman Roger Fleischman Robert A. and Jessica R. Frosch Susan Gerbi-McIlwain Sarah C. Goodrich Karl W. Flessa and Mari N. Jensen William Frost and Lise S. Eliot Gabriele Gerlach Holly V. Goodson-Hildreth Mark and Ellen Floyd Anne E. Fry Robin Gerlach Charlotte O. Goodwin Mary Pat Flynn Barry J. Fry James L. and Margaret F. German Arthur C. and Ilse E. Gorbach John W. Folino, Jr. Brian D. Fry Thomas Gewecke Katherine Gordon Andrew A. Forbes Paul A. Fuchs Hugh H. Gibbons Marion K. Gordon Byron D. Ford Ichiro Fujita Wallace and Anne Gibbons Martin Gorovsky John H. Ford Yoshiko Fujita Suzanne M. Gibson Michael Gottlieb Darhl Lois Foreman Shigeki Fujiwara Cameron E. and Margaret J. Gifford Esther M. Goudsmit Warren Forma Howard J. Fuller Gilbane Building Company Laura Gough and Henry Lamousin Franklin W. Fort William Fuqua Cole Gilbert Douglas B. Gould John Forte John Furfey William and Joanne Gilbrook Muriel Gould Alvin K. Fossner Edwin J. Furshpan and Shelagh A. Foreman Donald Giller Robert M. and Marianne Gould Barbara A. Foster Martha J. Gillis Robert A. Goyer David R. Foster Theresa Gaasterland Bruce P. and Gael Gilmore Bernice Grafstein James A. Foster The Mordecai & Elinor Gabriel Foundation Ann M. Ginsberg Louise and Jim Graham Sarah Q. Foster Michael and Bridget Gabriel Marguerite Anne Girling Roger K. Graham Stenio P. Fragoso Mordecai L. and Elinor R. Gabriel Barbara B. Glade Henry J. Grass, M.D. Charles D. Frame Barbara P. Gaffron Amy S. Gladfelter Judith P. and J. Frederick Grassle Ivy Frances Arthur G. Gaines David L. Glanzman Paul A. Gray Peter J. Franco Timothy R. Gainey MaryLou Glazer Philip and Carol Greco Elizabeth Frank Marzena M. Gajecka James A. Glazier Adrian Green Frank R. and Agnes Franklin Lisa Gallagher and Gregory Ruffa Richard H. Glew Deborah A. and Anthony D. Green Peter Franklin Paul E. Gallant Ann I. Glusker Edith S. Green Carie M. Frantz Mark A. Gallo Don Richard Goe Philip and Barbara E. Green Ben FrantzDale Gloria Gallo-Cromie Ethan M. Goldberg Stefan J. Green Alfred Franzblau Nicki Gamble Catherine Morrison Golden Lewis J. Greene

MBL Annual REport 2012 27 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

Frederic and Jocelyn Greenman Patricia C. and Frederick J. Hancox Philip L. Hertzler Mary C. Hopkinson Julius Sidney Greenstein Carla Cerami Hand Ellen Hertzmark Kenneth W. Horch Debra S. Grega Benjamin and Judith G. Handelman Sharon E. Hesterlee Erich F. Horgan Janet M. Gregg Roger T. and Arlene Hanlon Catherine E. Hibbitt and Tim Rockwell Anita B. Horn Leslie G. and James K. Gregg-Jolly Anne M. Hanneken Karen M. Hibbitt Edward G. Horn Huson and Edith Gregory Debra Tumbula Hansen Joseph and Barbara Hichar Melvyn Horwith Roger L. Greif Frank and Barbara Hanson Caitlin E. Hicks Sally Hoskins Edwin R. Griff Clifford V. Harding, Jr. Jay Hiller Richard A. and Susan M. Houghton Jeanne B. and B. H. Griffith Clifford V. Harding and Mina K. Chung David M. and Ann Hillis James A. and Marie G. Houle Patrick A. Griffith Marie Hardwick Ralph Hillman Robert C. Howard James A. Grifo Linda and Gerry Hare Howard D. Hinkle Christine L. Howe Warren M. Grill Maureen and Arthur Harlow-Hawks Rodney Hinkle and Kirstin A. Moritz Zdenka Hradecna Laura Grosch and Herb Jackson Kristen M. Harris and Max Snodderly Larry Hinzman Ru-Chih C. Huang Barbara S. and Leslie F. Gross Mary Harris Arlene A. Hirano Mary B. Hubbard Jerome Gross Stephen Coplan Harrison Stephen Hirschhorn Franz Huber Robert Grossfeld Peter Hartline and Rebecca Kucera Akira Hishinuma Robert Huber Albert and Sondra Grossman James and Martha Hartmann Sarah E. Hitchcock-DeGregori Jennifer M. Huettner David M. Grossman Buzz and Janis Harvey David Hobbie Linda A. Hufnagel Howard I. Grossman Janet M. Harvey Erik and Alison Hobbie Jeffrey and Karen A. Hughes Lawrence I. and Esta S. Grossman Pamela D. Harvey Lawrence and Layne Hobbie Edward M. Hulburt Jeanne Guillemin-Meselson Audrey E. Haschemeyer Ann Snow Hobbs Joan S. Hunt Jay M. Gulledge Valerie J. Haskins Harry Hochheiser Patricia A. Hunt Charuni A. Gunaratne William A. and Jeannine M. Hatch Nina W. and Andrew C. Hocker Alice Hunter Gregg Gundersen Sally Hauck Ruth A. Hodges Colleen D. Hunter M. Joslyn Gurley Dorothy Haugaard Edward S. Hodgson, Jr. W. Bruce and Suzanne I. Hunter Gilbert F. Gwilliam John Hay Tracy M. Hodgson Charles E. Huntington Nancy J. Gwynne Raymond L. Hayes, Jr. Eileen E. Hofmann Deborah Hursh and Mark Mortin UnJa L. Hayes Deborah A. Hogan Ralph and Joyce Hurvitz Janet Haas David S. Hays Cheryl Hogue Jerard Hurwitz Matthew Hackenberg Rhonda L. Hazell Mark Hollander Jodie L. Hurwitz Antoine F.O. Hadamard Jane M. Heald Robert and Patricia Hollett Tim and Mary Husted E. Janet Hager Judith M. Healy Holliston High School Deborah Hyland Jill K. Hahn Edward S. and Mary G. Heard George Holman Kerwin E. Hyland, Jr. Stephen L. and Ann S. Hajduk Alice B. Hearn Max and Gabby Holmes Thomas R. Hynes Kasturi Haldar Gina and Edmund A. Hebert Richard Holt Julie Hysom John Haldenstein Jeffrey C. and Lois S. Hecht Robert G. Holt Ronald J. Haley William Allen Held Raymond W. Holton Michael J. Iadarola John E. Hall Edward E. Heller Jane Holtz Jayne M. Iafrate and Barbara Stephens Sidney Halperen Simone Helluy Seymour and Victoria E. Holtzman Lynne Ierardi-Curto Marnie E. Halpern James R. Henriksen Timothy E. Holy Antje Ihlefeld Keri A. Halsema John H. Henson George G. Holz, IV Ichio F. and Naoko Ii Halsey Family Foundation Peter K. and Margaret H. Hepler Christina Holzapfel Robert F. Illgen Lisa M. Halvorson James J. Herlan Frederick C. Homan Kachi Illoh Tom and Lisa Hammett Michael and Donna Herlihy Uwe Homberg Nicholas A. and Laura M. Ingoglia Jin-Kwan Han Caterina M. Hernandez Jon Honea Felix and Yolanda E. Inigo Arnold Hanawalt Marco A. Herrera-Valdez Betsy and John Honey Institute for Sensory Research Scott B. Herrick Lewis and Carol Honig Integralis Inc. Boston Theodore T. and Ethel G. Herskovits Charles Hoopes Richard and Nancy Intres

28 MBL Annual REport 2012 CAtalyst Campaign Donors

Allen Isaacson Harry S. Kahn Louis M. and Brenda Kerr Kenneth Korach Sara Isenberg James W. and Ann P. Kalat Nasim Khan Ellyn V. Korzun Haruhiko Itagaki Doris Kalisman Lynda A. Kiefer Barry Evan Kosofsky Stephen K. and Patricia W. Itaya Caramai N. Kamei Thomas L. Kieft Edward M. Kosower Minami Ito Yoshie Kamei Kathleen A. Killian Michael and Jennifer Kosowsky Michael T. Ivy Fred I. Kamemoto Jennifer Kimbell Birgit Kovacs Irving and Selma Kamil Betsy King George H. Kowallis Jon W. Jacklet Ritsu Kamiya Camille M. King C. Edward Kowalski Kenneth Jackson Marianna M. Kane David and Camilla King Keith G. Kozminski Michael and Gail Jackson Edna S. Kaneshiro Donald W. King Timothy W. Kraft Christina Jacobi Joan T. and John W. Kanwisher Gilbert and Katharine King Mike and Beth Kramer Allison E. Jacobs Benjamin Kaplan and Rita Farber Jennifer M. King Matthew D. Krasowski Beverly and Paul Jacobson James L. Kaplan Richard H. and Reta King Geri Kreitzer Dieter Jaeger Matthew Kaplan Robert C. and Virginia King James N. Kremer Richard and Kathleen Jaeger Richard Kaplan Robert P. King John and Susan Krezoski Lionel Jaffe Frank Karger John M. and Louise Kingsbury Elinor Kritzman Daniel E. James Richard F. Karger Kevin M. Kingsland Andrew M. Kropinski and Peggy A. Pritchard Edward and Jeannine James Gloria Kariotis Jane Kinkle Jason Krumholz and Emily Manz Eric R. James David M. Karl Otto Kinne J. Gijs Kuenen Gregory J. Jennings Melanie Karp George W. Kipphut Virginia Kuykendall Mari N. Jensen Sally Karush Robert V. Kirchen Daniel A. Jimenez Karen E. Kasza David Louis Kirchman Kate R. Labrador Sigrid Schenk Joachimsthaler David C. Katz Mark D. Kirk Leo E. and Joan LaChance A. Kristine Johnson Marcella Katz Leonard B. Kirschner Alta S. and Bela Lahner Benjamin R. Johnson Michael G. Kauffman Marc W. and Phyllis Kirschner Monina R. Lahoz Carl F. Johnson Seth N. Kauppinen Jessica Kissinger David Lambert Carl H. Johnson Gordon I. Kaye Elizabeth R. Kittrie Robert P. and Ginny Lambrecht Carolyn L. Johnson Murat Kaynar Peter N. Kivy and Joan Pearlman Yolanda C. Landrau Hansen Johnson Jon Kayyem Norman R. Klath Michael J. Landzberg James E. and Kathryn M. Johnson Edward L. and Edna Kean David L. Klein Nancy Lang Nancy A. Johnson M. Whitney Keen Donald and Sandra Klein Meryl and Melvin Langbort Barbara Johnson-Wint Michael J. Kell Ophir D. Klein James and Elinor Langer Daniel and Jean Johnston Hartmut E. and Karla H. Keller Paul Kligfield Deborah A. Lans Dean and Mary Johnston Thomas C. S. and Laura R. Keller George W. Kling Jeffry B. Lansman Elizabeth Ann Jonas and Thomas D. Eisen Hans Kellner and Ruth Gross Michele M. Klingbeil Louis S. and Camilla B. Larrey Andrew C. Jones Doreen R. Kelly Peter A. Klock Melinda Larsen and Michael Gerdes Benjamin F. Jones and Juliet Sorensen Robert E. Kelly Alan Klotz Gail Larsen-MacPhee James M. and Olaive Jones Samuel W. Kelsey Paul W. and Virginia Knaplund Keith LaScalea Margaret H. and DeWitt C. Jones James and Paula Kemler Alan K. Knapp Howard S. Lasher Marlene S. Jones Robert G. Kemp Charles Knauss Stephen and Debi Laster James D. Jontes Richard E. and Cynthia Kendall Collin Knauss Susan Laster and Terry Little Marilyn J. Jordan Margaret A. Kenna, M.D. Paul M. and Carol L. Knopf Bruce B. Latham Erik M. Jorgensen Katherine Kennedy Robert E. Knowlton Joyce Whitehead Lathbury Susan Joslin Thomas C. and Mary Ann Kenny Robert A. and Cynthia Knudson Richard and Adrienne Latimer Richard S. Kent William Kobyljanec James Laundre and Bonnie Kwiatkowski Kathryn Kachavos Patricia E. Keoughan Susan Kohn Jennifer LaVail Leonard K. and Sheila F. Kaczmarek Lee Kerkhof Dana J. Kolibachuk A.W.D. and Fay LaVigne Gar Kaganowich Don and Marlys Kerr Deborah and Walther Kondig Sylvia Law

MBL Annual REport 2012 29 CAtalyst Campaign Donors

Mary E. Lawrence Scott Lindell and Alison Leschen Eduardo R. and Nancy P. Macagno Aric Martinez Marie M. Lawrie Jon Martin Lindstrom Raymond MacDonald Inez Martinez Jeffrey Lazarus and Phyllis Cohen Jairam Lingappa Sally MacIntyre Magdalena Martinez-Canamero Corinne Le Bovit Benedikt Linskens Neil and Barbara MacKenzie E. Mary Martini Berton J. and Barbara E. Leach Anne M. Linton Margaret M. MacLeish Joseph C. and Connie Martyna Marie-Claire Lebeau Raymond J. Lipicky Peter MacLeish Takeshi Maruo Janine M. LeBlanc-Straceski Jennifer Lippincott-Schwartz Jane and Daniel MacNeil Karla E. Marz Denis R. Lecavalier Eve and Ted Lippold Phyllis M. MacNeil Timothy C. Marzullo Elizabeth E. LeClair Stephen J. Lipson Katherine MacQueen Trina Mascott Joan Lederman James C. Lisak Alix Madigan Massachusetts Marine Educators Cindy Lee Joan W. Lisak John J. Magnuson Anna Mastroianni and Greg Shaw Kyeng Gea Lee Barbara C. Little Gisele Magnusson Sandra K. Masur Sue Ying Lee Little Harbor Foundation Vinay S. Mahajan Natalie N. Mather Susan E. Leeman Madeleine Littman Patrick J. and Lorraine Mahoney Walter Matherly Vanessa LeFevre Jylene M. Livengood Richard Mains Rita W. Mathews Diane Rielinger Legault Robert and Laura Livingstone Guido Majno and Isabelle Joris-Majno Allen Wray Mathies, Jr. Robert Owen Lehman William and Joan M. Lloyd Stephen E. Malawista Paul Matsudaira William J. Lehman Phillip S. Lobel Craig C. and Darlene M. Malbon Joan Mattson George Leipzig Edward and Jennifer Lobenhofer Robert P. Malchow Alex S. Mauss Tanya Leise Edward Locke Antoniette M. Maldonado-Devincci Dorothea J. Mautner Ardean Leith Charles F. and Keran Lockhart Nancy R. Malkiel Barbara Jo May Russ Lemcke Edgar Lockwood and Claire Cohen Matthew Mallow and Ellen Chesler Gregory S. May Ellen K. LeMosy Robert A. Loest Kevin Maloney and Jean Nolan Anne E. McBride David P. Lenzi Frank A. and Mary W. Loewus Richard S. and Sylvia A. Manalis Frances V. McCann Murray Mark Lesk Ann M. Lohof Frank T. and Ose Manheim John W. and Judy McCarter Charles Leverone Barbara M. Lom Sridhar Mani Robert McClary Janet Miller Levie Frank J. and Kathryn L. Longo Charles K. and Susanne J. Mann Megan N. McClean Myron Levine Paulette Loomis Diann and Thomas Mann Ashleigh McCord Richard B. and Jane M. Levine Barbara T. Loring Douglas G. Mann and Jane G. Schweitzer Kathryn E. McCormick Thomas Levine Janet Loring Uri Manor Derek J. and Nancy McDonald Francoise Levinthal Susan Loucks Mary Ann Mans Donald and Laurie McEnery Herbert and Karen B. Levitan Robert L. and Gwyneth E. Loud Peter and Anna Manso Matthew McFarlane and Jennifer Nadeau Ann L. Lezberg Alane Love Bernard and Isabelle Manuel Grace McGill Steven William L’Hernault Warner E. Love Tianyi Mao Paul McGonigle Elizabeth and Paige L’Hommedieu Thomas E. Lovejoy Diane B. and Joseph S. Maranchie David and Cynthia McGrath Chenghui Li Richard C. Lovering Robin Marantz Henig A. David McGuire Jade Li Janet S. Loynes Ronald and Barbara Marcks Joseph and Darlene McGuire Wei Li Roger C. and Mary Ann Lucas Eugenia Marcus Gary McHatton Bruce T. Liang Kaitlyn Lucey Eve Marder Robert McIntire Min-Ken Liao Richard H. C. Lucius David Margoliash Robert B. McKane Richard T. Libby Louise M. Luckenbill Deborah Margules Margaret H. McKibben Gene E. Likens William B. Ludington Dawn Morin Marick W. Paul and Karen McKinney James S. and Alice J. Liljestrand Bjoern Ch Ludwar Andrew C. Marinucci Mary W. McKoan Bruss R. Lima Chris M. Luecke David and Jessica Mark Welch Arthur and Jo Ann McLendon James and Cynthia Limberakis Zella Luria Junko Munakata Marr Katherine D. McMahon Shin-Yi Lin Lawrence Lynch Corp. Mars Educational Tours Cornelia Hanna McMurtrie Richard W. and Madeleine A. Linck Craig Lyons Charles E. Martin, Jr. Samuel McMurtrie, Jr. Nancy Kane Lind Gary E. Lyons Jack Martinelli and Nancy Couts Michael and Lyn McNaught Lennart and Ann C. Lindberg W. Berry Lyons Adria E. Martinez and Juan Morin James and Joanne McSherry Katherine May Lyser

30 MBL Annual REport 2012 CAtalyst Campaign Donors

Bruce L. Mehler Florence E. Mixer Robert J. and Jeanne A. Naiman Victor and Ruth S. Nussenzweig Jay and Phyllis Mehlman Hazime Mizoguchi Angus Nairn and Marina Picciotto Mary Jane Nutter Jonathan and Jane Meigs Rebecca Moffitt Kenneth E. Najarian Rebecca W. Meigs William A. Mohler Mia Nakachi Ana Lia Obaid Sarah Meigs Ruel and Alice Mohnkern Hiromichi Tsuda Narahara Steven Oberbauer Alexander Meigs-Rives Sukhbir S. Mokha Nashua Christian Academy Michael and Carol Oberdorfer Anne D. Melamet Jean and John Monahan Lewis and Phyllis Nassikas John and Marion O’Brien Arleen T. Mellon Yiannis Monovoukas National Wild Turkey Federation, Inc. Richard O’Brien DeForest Mellon, Jr. Lawrence A. Monte John E. and Ethel H. Naugle Clifford T. and Hope H. O’Connell Lynn Mendelman Charles H. and Mary Louise Montgomery David E. Naylor James J. and Patricia O’Connor Everett Mendelsohn Melita H. Moore Pius N. Nde Joe and Patty O’Connor Allen F. Mensinger Stephen A. and Marjorie H. Moore Ruth H. Neff Tim and Mercedes O’Connor Irene H. Mentzel Steve S. Moore Cheryl and Nabil Nehme William and Sarah Odenkirk Wilhelm and Nonie Merck Alexis MH Moran Anne Nelson Lynda Oehme Wesley J. and Emmy C. Merritt Marius R. Moran Pamela Nelson and Christopher Olmsted Daniel B. Oerther Carmen Merryman Mary Ann Moran Ralph F. Nelson Susan Offner Vivagean V. Merz Nancy Mordente Victoria L. Nelson Daniel O’Grady Blanche Meslin Jennifer R. Morgan Iris Nelson-Schwartz Shinpei and Catherine B. Ohki Anne Messer Michael M. Morgan Jeniel E. Nett Dennis Ojima and Jill Baron Shanta Messerli Yasuhiro Morita Jill Neubauer John and Karen O’Leary Robert T.P. and Barbara Metcalf Hideaki Moriyama Kristine Neville Stephanie Oleksyk Richard and Phyllis Meyers James Morrell Ann Little Newbury Thomas J. Oleksyk Steven Miano Hilary and Archie Morrison Robert F. Newkirk Stephanie A. Olexa Roger and Barbara Michaels Naomi S. Morrissette Dianne K. Newman Timothy N. Oliver Sara Michie James C. Morrone John N. and Kathleen K. Newman Janice and Albert Olszowka Leonard A. and Stephanie J. Miele Betty L. Morse Nicholas N. Newman Emily Fowler Omura Jill A. Mikucki M. Patricia Morse Carol Newton Jeffery and Valerie Oppenheim Nancy S. and Richard Milburn Stephen S. and Marilyn G. Morse David J. Newton Norman K. Orida Milhench Supply Company Isabelle Mort David L. and Linda D. Newton Iris M. Ornberg Helene Miller Thomas C. and Elizabeth D. Moseley William E. Newton James L. and Nora Oschman Holly Miller Henry A. Moses Brian D. Nibbelink Ann-Beth Ostroff Lynn Miller Day O. and Kathie C. Mount Teresa A. Nick, Ph.D Kimberley O’Sullivan Rita K. Miller William R. Mowrey Kenneth W. and Ann W. Nickerson Renee Bennett O’Sullivan Rolf Mount Miller Walter J. Mozgala Christof Niehrs Yuko Ota Aaron Mills and Linda Blum Ann-Kristin Mueller Christina Niemeyer and Robert W. Zeller George O’Toole John W. Mills Christoph Mueller Eliot Nierman Lorrence J. and Beverly C. Otter Margaret A. Mills-Michael Thomas A. Mulholland Anna E. Nimeskern Timothy Otter Brenda Milner Sc.D. Paul and Mary Mulloy Rae Nishi James G. Ottos Leonard E. Mindich Multitemp Mechanical Corp. Lee A. Niswander Peter J. Ouellette Martin M. Miner Stephen H. Munroe Scott W. Nixon Robert Ouimette T. Richardson Miner and Bobbie Loop Patricia N. Murata Amy Nobu Paul A. Overbeek Edwin A. Mirand Robert E. Murphy Drew M. Noden Scott F. Owen Steven D. Mirsky Seanan B. Murphy Edmund F. and Ann Nolan Brett A. and Kathy Oxberry Thomas Misgeld Stephanie A. Murphy Phyllis Norris Carol and Daniel Mishell Xavier Musacchia North Star Financial Services Corp. Mary G. Pacifici George and Denise Missall Frank B. Myers Thomas and Virginia Noyes Daniel Page and Kirsi M. Savusalo Charles A. and Betty Mitchell Wallace H. Myers Harriet Nurse Joanne and Alan Page Peter and Olga Mitchell Donald L. Mykles Adam Nussenbaum and Shari Abramowitz Kathryn Paine William M. Mitchell Eleanor M. Nace David Nussenbaum David J. Pajerowski

MBL Annual REport 2012 31 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

Sarah L. Pallas and Paul S. Katz Debra Peters Carl D. Prota Judith C. Rhodes David R. and Anastasia Palmer Niles and Mary Peterson Richard Purple Lockwood W. and Mary Rianhard Kelli L. Palmer Olivia R. Petrasch Dale Purves and Shannon Ravenel Andrea Ricca Richard and Barbara Palson Carey R. and Ryan J. Petrie Christian Putnam Marian C. Rice Yude Pan Jane Petrie Mary Elizabeth Rice Harish C. and Kamala Pant Ronald J. and Dawn G. Pfohl Helen Quinby Mary Esther Rice Nicholas and Carmela Pantazis Dwight E. and Katharine Phillips Neil K. Quinn Robert V. Rice Costa and May Papastephanou William G. Phillips Richard Quinn John G. Richardson Suzanne Paradis Flynn Picardal Frederick R. and Kathryn G. Rickles Anastasios Parafestas Albert T. and Lorraine Piccirilli Esther L. Racoosin Conly L. Rieder Rosevelt L. Pardy Norman J. Pieniazek George Radcliffe Austen F. Riggs David and Lois Parker, Jr. Louis Pierro Paul Raeburn and Elizabeth DeVita- Diane M. Ring Jaqui D. Parker Carl B. Pilcher Raeburn Elizabeth Ripple Sophie S. Parker Sara C. Pimental Keen A. Rafferty, Jr. John and Jeanine Ripple KiTani A. Parker-Johnson Jerome Pine James W. Raich and Ann Russell Paul B. Rizzoli Carolyn L. Parmenter Ricardo Pineda Wilfrid Rall Alison A. Robb Charles and Patricia P. Parmenter Leslie and Donna Pinsof Jeffrey L. Ram David and Rosemary Roberts James L. Parmentier and Elizabeth Fowler Anthony and Wendy S. Pires David J. Ramsey Jean Roberts Barbara Jane Parpart Helen and David Piwnica-Worms Khela Ransier Thomas H. and Kimberley H. Roberts Jane L. Parpart Stephen and Robin Plattman Shirley Raps Robertson Trust Carolyn Partan Sharon Plautz Nancy Ratner Rex R. Robison Leonard M. Passano David E. and Jeanette Pleasure Dianne Margaret Rausch Jennifer Rocca Angel M. Pastor-Loro and Rosa R. de la Cruz Samuel J. Pleasure, M.D. Luciana Rava, M.D. Jonathan Rockoff Paul H. Patterson George H. and Louann S. Plough Fred J. and Catherine E. Ravens John R. Roderick Thomas Patterson and Lucy Bartlett Mark Pochapin Jennifer L. Raymond Dana and Alison Rodin Kimberly S. Paul and Charles Thomas Tanya D. Podchiyska Peter Raymond Ramona M. Rodriguiz Richard Payne and Deborah Siegal Jessica K. Polka Samuel O. and Holly Raymond Snezna P. Rogelj John B. and Ruth E. Pearce Daniel A. and Linda K. Pollen Neal E. Ready Laura M. Rogers David and Rose Pearson Nancy R. Pollis Willis B. and Joan Reals Warren Rogers and Linda St. Jacques Rogers Kay Pechilis Lawrence Pomeroy Geoffrey Ream Richard E. Rohr Russell H. Peck William F. Pomputius, III Matthew Reardon Nichlas Romano Alejandro Peinado David Poppick Alfred G. and Sarah C. Redfield Michael Romero Pelican Cove ROMEO’S Rachel S. Poretsky Graham Redinger Brian K. Romias Cara Pellegrini Charles Porter Renee and Michael Redman Christian Roop Joyce S. Pendery Mary E. Porter and Thomas Sewell Hays Bernard B. Rees Matthew F. Rose Nick Peng Pamela Porter Ann N. Reese Wendy E. and Edward A. Rose Susan M. Pennington Anton Post Jean F. and Ronald R. Regal Jody Rosenblatt Elizabeth Pennisi Aubrey and Margareta B. Pothier Kathleen M. Regan and Jeffrey Leshin Clare Rosenfield Alberto E. Pereda Michael J. Povelones Vincent Rehder Manuel Rosenfield Ruth Perez Anahid E. Powell Marcus Reidenberg Allen Rosenspire John B. and Barbara M. Peri Sabrina and Bradford Powell Susanna K. Remold Atholie K. Rosett Francine Perler Maureen K. Powers Michael J. Renner John D. and Martha A. Ross Richard D. Perron Catherine Prendergast Lisa Repko Virginia F. Ross Charles Perrow Edna Pressler Kristin Lester Revill Helen Ross-Sangster Barbara C. Perry Anne T. Pressman Randall W. Reyer Cordelia Roth Melissa J. Perry Donald L. Price Carol C. Reynolds Lois Rothburd Matthew A. Person and Jill Erickson David J. and Merry L. Prior Prudence B. Reynolds Rudi J. and Hanna Rottenfusser Robert M. and Kristine M. Reynolds Pierre E. Rouviere Virginia R. Reynolds Frans and Marga Rowaan

32 MBL Annual REport 2012 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

Lew and Esther Rowland Nina Chandramani Saxena Terrence J. Sejnowski Rolly L. Simpson Tania J. Rozario Tina M. Saxowsky Christina M. Sekaer John Sinard Uldis Roze Ruth L. Saz Paul Selvin Maxine F. Singer Alan A. Rozycki Thomas and Judith S. Sbarra Deborah G. Senft Dorothy P. Sinha Gualberto Ruano Shirley W. Scaife Hannah Senft Stephanie M. Sit Laurens N. Ruben William and Kathleen Scala Joseph P. Senft Nawaphon Sittisawassakul Daniel and Nancy Rubenstein Mary L. Scalera Owen J. and Mildred B. Sexton Edward R. Siuda Shirley Rubinow-Friedman Mary M. Scanlan Laura and Jamey Shachoy Kathleen K. and Robert Siwicki Parke A. Rublee Deborah G. and John F. Scanlon Deana Shackelford Watts Raymond A. Sjodin Lucinda A Rudin Patrick Schaefer Shanti Shahani Philip J. Skehan, Jr. Rivka Rudner Raffael Schaffrath Gary A. Shangold Stephen Skillman David Rudnick Rolf Schauder Herbert and Rose A. Shanker Wendy Skinner Roland R. Rueckert Jeffrey Schell Carolyn and Joseph Shannon Rosanne Skirble William A. and Andrea Rugh Ellen R. Schellhause Courtney H. Shannon George J. Skladany Smil and Marion Ruhman Artur Scherf Natalie and Mike Shapiro Jean and Bruce Skrine Linda L. Runft Suzaynn F. Schick Joan Sharp James Slock Alice M. Rushforth Mary E. Schiffman Victor E. and Angela V. Shashoua Ruth C. Slocum Norman B. Rushforth Ralph Schiller Edward I. Shaw Roy M. Smeal Ann L. Russell Joshua Schimel Jerry W. Shay Cynthia C. Smith Karen R.M. Russell Hans L. and Luise Schlesinger Padraic J. Shay Frederick E. and Marguerite A. Smith Naomi Rutenberg Helen Schlesinger John Shea and Sallyann Garner George and Joyce Smith John G. Rutherford, Jr. Jeffery A. Schloss Margaret Shea Gregory A. Smith Ueli Rutishauser Cathleen Schmidt Daniel and Debbie Shearer Kirby D. and Linda Smith Ginny L. Ryan Buresh Marc F. Schmidt Martin I. Sheldon Lindsay Byron Smith Charlotte Ryan Thomas M. and Susan Schmidt E. Frances Shepherd Marva Eugenia Smith David Ryan Ian C. Schneider Gordon M. Shepherd Paul F. and Mary L. Smith Dorothy C. Ryder Joseph H. Schneider Alan and Stephanie Sher Peter J.S. Smith Jan S. Ryerse Anne M. Schneiderman Denise R. Shereff Ruth T. Smith Schnitzer Family Foundation Irwin W. Sherman Stephen and Mary Jane Smith Ali and Markrid Saab John F. and Diane S. Schoenfelder David R. Sherwood Una Smith Matthew and Kelly Saal School of the Holy Child Ujwal P. Sheth Roxanna S. and Ronald J. Smolowitz Theodora Saal David Schoppik Celia E. Shiau Snow and Jones Inc. Sacks Family Foundation Ethan Schrank Timothy and Kathleen Short Thomas F. and Anne W. Snyder Mary Beth Saffo Robert F. Schroeder Richard and Elizabeth Shriner Samuel J. Sober Richard Sailor and Mary Johnston Andrew Schutrumpf Carl N. Shuster Jill A. Sohm Wendy C. Salmon Jessica C. Schwartz Enid K. Sichel Karam F.A. Soliman Brian M. Salzberg Lawrence M. Schwartz and Carol Bigelow James M. Sidie Lilianna Solnica-Krezel David R. Samols Melvin E. and Elizabeth Schwartz Herbert and Sydell Siegel Pamela Soltis Robert N. Sandberg Emily Schwartz-Clark Richard M. Siegel Linda Sommer Christopher Sanders Ralph T. Schwarz Hansruedi Siegrist Alan M. Sooho William B. Sandler Elisabeth C. and Gary Schwarzman Thomas Sikina Martha M. and Albert L. Sorenson Steven B. Sands Hilda Scire Bertram R. and Dorothy Silver Carmen S. Soto Joshua R. Sanes Barinetta Scott Barry & Judy Silverman Foundation, Inc. David B. Space and Linda A. Jarvis Jean-Louis G. Sankale Elouise C. Scott Roy L. Silverstein and Jacquelyn A. Helen Spaulding Robert J. Saqueton Harold H. and Kristin T. Sears Joseph-Silverstein Louise M. Specht William Sass John J. Seder and Frances R. Plough Charles Simenstad David and Mona Spector Satterlee Stephens Burke & Burke Jerome and Bernice Segal Eric J. Simon John W. and Cathleen Speer Morton T. Saunders Robert and Carole Seidler John A. and Michele B. Simourian Edward A. and Barbara C. Spiegel Kenneth E. Sawin Mary Ann Seifert Joyce M. Simpson Debra and Donald Spinelli

MBL Annual REport 2012 33 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

John E. Spiro Josephine Strauss Saul Teichberg Tom Tullius Nadja Spitzer Paul Strebel Sally Temple Louis C. and Lee R. Turner Sven and Anna Spoerri Theresa V. Strong Mari DeCosta Terman David and Nancy P. Twichell Donald Spoon Daniel E. Strongin Alejandro Terrazas Robert Tyler Allan C. Spradling Raymond G. Stross Catherine D. Thaler Michael and Frances W. Tytell Nelson Spruston Brad Stumcke Roger Elliot Thannerthies Helene E. Spurrier Sylvia O. Suadicani Roselle L. Tharion Hiroshi Ueno St. Andrews Foundation Nobuo Suga The Schmitt Family Charitable Foundaton Eleanor S. and Kevin Uhlinger St. Barnabas Memorial Church Charles H. and Donna J. Sullivan Kristin Tholke Zoltan Ungvari Sheila M. Stackhouse David T. Sullivan Suzanne M. Thomas United Nations Association of Greater Boston Walter F. Stafford, III John and Barbara Sullivan William R. Thomas Vivek K. Unni Joel P. Stafstrom Lawrence R. and Nancy L. Sullivan Holly Thompson Smitha B. Uthaman Wallace and Pamela Stark Walter Sullivan Richard and Kathy Thompson Charles Stasey Eric T. and Holly Sundquist Wesley J. Thompson and Mary Ann Rankin Ivan and Virginia Valiela Robin L. Staub Kathy A. Suprenant and William L. Dentler W. Nicholas and Joan I. Thorndike Pauline Valois David A. Stauffer Timothy A. Sutton Mary and James Thorne Alice H. van Buren Barbara A. Stay Dorothy N. Swanson Robert F. Thorne Matthijs van der Meer Norman S. Stearns and Irma Mann Dorman J. Swartz Carl L. Thurman, II Miranda Van Gelder Tim Stearns Hyla C. Sweet Marjorie Till Kensal and Barbara van Holde Robert E. Steele Harold E. Sweetman Lewis G. and Elaine Tilney Andrew and Catherine Van Praagh Lucy Steere Lawrence Sweetman Jay A. Tischfield Daria N. Van Tyne John J. Stegeman Susan Swigor Natascia Tiso David L. Van Vactor, Jr. Michael Stein Alexander O. Sy Cathy Tobin Joseph W. Vanable Sidney and Sue Stein Barbara A. Sylvia Alice Todd Dallas A. Vanorny Claude Steinberg Ben G. Szaro Laurie Tompkins Garret VanWart Eleanor K. Steinberg Robert and Sylvia Szulkin Michael Toner Anthony Vattay Jay Steinberg Aron and Irene Szulman Mary F. Toomey Sylvia Vatuk and George Rosen Jeffery Steinberg Leana and Joby Topper Kevin T. Vaughan Julie Steinberg and Richard Engerman Tabor Academy Kevin G. Townsel Robert Veeder Joan A. and Thomas A. Steitz Masanori Takahashi Amy Townsend-Small Pauline T. Velasco Thomas A. and Joan A. Steitz Mary J. Talbot Walter V. and Catherine Trainor Mark A. Velleca Sherry E. Stekloff Sidney L. and Signhild Tamm William E. and Belle Traver Antonio Ventosa StemCell Technologies Inc. Josephine P.G. Tan Ann H. and Mark F. Trax Joseph and Lorraine Viglione Ann Stern Kevin S.W. Tan Mead Treadwell Greg J. Villareal Eleanor Stern Frank and Loretta Tang Steven N. and Joanne Treistman Dorothy Villee Judith G. Stetson Tomomi Tani Barbara S. Trela Angela C. Vincent George D. Stetten Marvin L. and Betsy C. Tanzer Barbara S. Trelstad Vincent Associates Karl Stetter Alexis Tapanes-Castillo Walter H. Troffkin Charlotte M. Vines Craig W. Stevens Steve C. Tarrant Natalie Trousof Maria Luisa Virata Charles N. Stewart Peter Tassia and Maija Lutz Keith A. Trujillo Arthur D. and Joanne R. Voorhis James M. and Joly Stewart Belle K. Taylor Philip N. Truluck Ellen Vopicka Randall R. Stewart Charles W. Taylor Tsuchiura First High School Marc Stieglitz Richard S. Taylor Tsuchiyra High School Ralph and Judith Wadleigh Norman L. and Lisa Stockbridge Richard Sears Taylor Christopher W. Tubbs Patricia Wadsworth Jeffrey Stoddard Betty Taymor Greg and Kay Tuber Daniel Wagenaar Alexandra G. Stone Richard B. Tayrien James Tucker Irving and Vida Wagner Gary Stormo Edwin H. and Priscilla S. Tebbetts Jane Tucker Gregory Wagoner Adam Stracher Technical Video, Ltd. Steven Tucker Thomas Wakim

34 MBL Annual REport 2012 CACAtalysttalyst CCampaiampaiggnn DDonorsonors

Maryalice Walker Mark J. West and Maryam Golchin George Wittenberg Colby J. Zaph Robin A. Wallace Jane A. Westfall Nancy Woitkoski Veronika I. Zarnitsyna Emily C. Walsh Nancy and Peter Weston Jason S. Wolfe Arturo R. Zavala Mary J. Walsh Wayde M. Weston Ralph and Gretka Y. Wolfe Ricardo Zayas Kerry Walton Michael Wheeler Ann S. Wolff Zazzle Peggy L. Walton and William Walton Paul S. and Barbara B. Wheeler Robert G. Wolk M. Jade Zee Nancy L. Wanek Mabel Y. Whelpley Lily L. Wong Steven Zeichner Brant G. Wang Elizabeth A. Whitchurch Michael and Mary Wood Linda A. and Erik R. Zettler Jing W. Wang Gerry and Gina White Michelle A. Wood Jiawei Zhang Yihong Wang John White and Sonia Witte Teresa Woodruff and Thomas O’Halloran LingLi Zhang Lawrence J. and Katherine S. Wangh Stephen and Anna White George M. and Katharine Woodwell Ming Zhou Eric R. Ward Tanya B. White Nancy Woolford Yi Zhou Frederick Ward Barbara M. Whitehead Sarah Woolley Qianlai Zhuang Margery Ward Ben B. Whitlock John A. Worcester Bunnie Rose Zigman Samuel and Anne Ward Stuart H. Whitlock and Rachel Richwine Worldwide Planning Services, LLC Richard E. Zigmond Joseph T. Warden Chris L. Whittle Susan Wray Kenneth H. and Patricia E. Zimble Falk Warnecke Konrad Wiese Kathryn M. Wrubel Bruce and Elizabeth I. Zimmerli Deborah M. Warner Susan Wigley and Jay Burnett Carl Wu Andrew W. and Lenore N. Zimmerman Estelle P. Warner Athula H. Wikramanayake Chao-Ting Wu Arthur Zimmerman Dorothy Wass Linda E. Wilbrecht David D. Wu Andrew and Susan Zimmermann John Waterbury and Vicky Cullen Harry W. Wilcox Zeba B. Wunderlich Jeffrey A. and Johanna A. Zinn Talbot H. Waterman Martin Wild James Allen Wyban Alexander and Judith S. Ziss Coreen A. Waters Ann S. and Lynn H. Wilke Richard M. Wylie Ludek Zurek James D. Watson John J. Wille, Jr. Charles R. and Ellen G. Wyttenbach Andrew and Ruth Zweifler Priscilla J. Way Benjamin Douglas Williams Kate K. Webster Carrie A. Williams Duane and Kathleen Yaindl William K. Webster Edward Williams Xian-Cheng Yang John T. and Jeanne T. Weeks Mathew Williams Yuriko Yano Harrison Wehner Patricia C. Williams Charles Yanofsky Aguan Daniel Wei Milton E. and Susan G. Williamson Patrice O Yarbough James Wei Darcy B. and Diane H. Wilson Norman R. Yetman George Weinbaum Dorothy L. Wilson George and Miriam Yevick Melvin and Flora Weinberg Gerrard Wilson Anne D. Yoder Alfred M. and Naida D. Weisberg Louis A. Wilson Isabel and Richard Yoder Alice Sara Weiss Nancy Wilson Abigail Young Herbert M. Weiss Steven J. Wilson Wise and Lily Young Howard D. Weiss T. Hastings and Dorothy M. Wilson Rafael M. Yuste Leon P. and Ellen Weiss Fred H. Wilt Anne-Marie C. Yvon Lisa Weissmann and Debra Shapiro Ellen Winchell Joshua S. Weitz Carolyn P. Winn Lucia F. Zacchi Jeffrey M. Welch Donn and Fluffy Winner Morton and Nurit Zachter Louis Wells and Ellen Seidensticker Helene D. Winstanley Marilyn G. Zacks Phyllis R. Wendorff Ashley Wires Verónica Zaga-Clavellina Robert H. Werner H. Kern Wisner Allison L. Zajac Ronald D. Wesley Louise Witonsky Harold H. Zakon and M. Lynne McAnelly Gary M. Wessel Stephan N. Witt William and Linda Zammer Janiene West Beatrice and Jonathan B. Wittenberg

MBL Annual REport 2012 35 2012 MAJOR GIFTS

The MBL gratefully acknowledges the generous The Charles Evans Foundation contributions and pledges of the individuals, $500,000 to establish the endowed Charles foundations, and companies who supported our Evans Research Development Fund. programs and operations in 2012. John and Valerie Rowe $1,000,000-$3,000,000 $500,000 in support of the Strategic Planning Fund and Annual Fund. Gordon and Betty Moore Foundation $2,258,548 grant to investigate microbial Joan R. Berman ecosystems at the Axial Seamount, a deep sea $295,577 bequest intention. volcano on the North Pacific; Julie Huber, Principal Investigator. Arthur Vining Davis Foundations $150,000 in support of the Semester in $100,000-$999,999 Environmental Science Program.

The G. Unger Vetlesen Foundation David and Susan Hibbitt $850,000 for support of the Bay $119,185 in support of the Strategic Planning Paul Center and Bay Paul Faculty. Fund and Annual Fund.

Anonymous Robert A. Prendergast $640,000 in support of the Swope Elevator $110,050 in support of the MBL General project and the Annual Fund. Endowment and the Annual Fund.

Dart Neuroscience LLC The William Randolph Hearst Foundation $500,000 to provide lab space for visiting $100,000 in support of the William Randolph investigators. Hearst Scholarship Fund.

36 MBL Annual REport 2012 SPECIAL GIFTS $5,000-$9,999 Prosser Gifford William and Susan Cobb Robert D. and Anne E. Goldman $10,000-$99,999 David and Patty Cohen Peter Gottlieb Anonymous (3) Mary W. Graham William H. and Margaret J. Greer The American Society for Cell Biology March of Dimes Foundation Larry Grossberg Peter B. and Margaret T. Armstrong Donald and Patricia Mintmire Carlos Gual Beech Tree Trust Monica Riley Robert and Margot Haselkorn Burroughs Wellcome Fund John and Joan B. Ripple Carmela J. Huettner Sally Cross Edward D. and Nancy M. Salmon Mark Koide The Ellison Medical Foundation Harriet F. Segal John R. and Andrea O. Lakian Saint Michael and All Angels Espiscopal Church Douglas R. and Virginia M. Shanklin Hans Laufer Peter and Ginny Foreman Michael P. Sheetz Michele Lorand and Paul Ringel Seiichiro Fujimoto Society for the Study of Reproduction Walter E. and Shirley A. Massey Howard Hughes Medical Institute James A. and Annamma Spudich William N. and Marianne W. Mebane George Frederick Jewett Foundation The Judy and Michael Steinhardt Foundation Matthew Meselson and Jeanne Guillemin-Meselson Marianne F. Milkman Ronald D. Vale and Karen Dell John Mills Nikon Instruments, Inc. Gerald and Ann Weissmann Hideo Mohri Elaine Pierson-Mastroianni Dyann and Peter K. Wirth Harold Nash William Townsend Porter Foundation Ms. Barbara K. Zeisler Bryan D. and Elaine K. Noe Joan V. and Gerald S. Ruderman Nancy Northup Society for Developmental Biology, Inc. Sidney K. Pierce, Jr. William Speck and Evelyn Lipper $1,000-$4,999 William S. and Catherine A. Reznikoff USNC-IBRO Committee Porter W. Anderson, Jr. Marius A. Robinson Wellcome Trust James and Cynthia Angstadt Lawrence C. and Victoria H. Rome John S. and Judith H. Willis Robert W. and Jean Ashton Charles M. and Phyllis Rosenthal Shawky Z. Badawy John D. and Jean M. Rummel David and Sandra Bakalar Sara Schupf Frederick and Christine N. Bay Society of General Physiologists Millicent Bell Marjorie E. Steinberg Gloria S. Borgese Andrew and Ursula Szent-Györgyi Gary G. Borisy and Sally M. Casper G. Pran Talwar Paul N. Chervin Waksman Foundation for Microbiology Marion Cohen Washington & Jefferson College The Company of Biologists Limited Alfred M. and Naida D. Weisberg David and Karen Davis Clifton R. Wharton, Jr. Stephen and Lois Eisen Joshua J. Zimmerberg and Teresa L.Z. Jones David C. and Patricia S. Gadsby

MBL Annual REport 2012 37 Special Gifts

Other Gifts Peter J. Donaldson Jessica Kissinger Anonymous (4) Henry Erle Peter N. Kivy and Joan Pearlman Emre R. Aksay Anne Etgen Charles Knauss David F. Albertini Andrew and Amy Feinberg Collin Knauss Marselle O. Alexander-Ozinskas First United Methodist Church of Chatham William Kobyljanec Ricardo C. Araneda Laura S. Fishler Elinor Kritzman Bryan and Donna Arling Andrew A. Forbes George M. and Sylvia T. Langford Carol Arnosti and Andreas P. Teske Warren Forma Leonard and Ruth Ann Laster Harold and Marion Barr Sarah Q. Foster Stephen and Debi Laster Susan R. and Daniel T. Barry Christopher J. Freeman Holly A. Lavoie Viviana M. Berthoud Ronny Frishman Marie-Claire Lebeau Jay B. Bikoff Harold and Ruth S. Gainer Laszlo Lorand Joanne Blum Sallyann Garner Richard H. C. Lucius Brenda J. Bodian Mark Gasarch Bjoern Ch Ludwar Margaret C. and Francis P. Bowles Wallace and Anne Gibbons Vinay S. Mahajan Aaron Bravman Bruce and Mary Goodman Joe L. Martinez and Kimberly Smith-Martinez John Brereton Muriel Gould Takeshi Maruo Giovanna Breu Bernice Grafstein Anna Mastroianni and Greg Shaw Andrew Brittingham Henry J. Grass, M.D. Alex S. Mauss Melinda Browne Paul A. Gray Bruce L. Mehler Elizabeth H. Burrows Gustav Grosch and Jane Kulesza Jerry and Lalise Melillo Elaine Cahn Stephen L. and Ann S. Hajduk Irwin and Ruth Merkatz Margaret Catley-Carlson John Haldenstein Steven Miano James Catterall and Jean Schweis Peter Hartline and Rebecca Kucera Mid-American Water of Wauconda, Inc. Robert and Penny Catzen Joseph Heitman David A. and Virginia I. Miller The Chee Pearlman Company, Inc. Ellen Hertzmark Steve S. Moore David Clarke John and Olivann Hobbie Yasuhiro Morita Bradley and Katie Cohen Richard Holt James Morrell Sarah Cohen James A. and Marie G. Houle Ann-Kristin Mueller Frederick Cooper and Karen Gordon Patricia A. Hunt Anne Nelson Catherine B. Cramer David Isenberg and Paula Blumenthal Teresa A. Nick, Ph.D John and Gwen Daniels Allison E. Jacobs John and Karen O’Leary William C. Daniels Hansen Johnson Daniel Page and Kirsi M. Savusalo Paul J. and Maria S. De Weer Freda Kaminer Elizabeth Pennisi Robilee Frederick Deane James L. Kaplan Charles Perrow Michael H. Dickinson Richard Kaplan Bertha Person Dirk Dobbelaere Frank Karger Jean W. Pierce Howard T. Dodd Richard F. Karger Leslie and Donna Pinsof Pavel Dolezal Darcy B. Kelley Harmony Glazebrook Pittet

38 MBL Annual REport 2012 Special Gifts

Plant Operations & Maintenance Staff - MBL Daniel Wagenaar GIFTS TO NAMED FELLOWSHIPS Mark Pochapin Leonard and Eve Warren Rachel S. Poretsky Harrison Wehner AND SCHOLARSHIPS Gordon and Claire Prussian Edward and Dia Weil Keith Purpura Robert H. Werner Scholarships and fellowships enable the MBL to Helen Quinby Michael Wheeler stay competitive by attracting and supporting top Geoffrey Ream Ellen Winchell researchers and students. We gratefully acknowledge Fred and Gloria Rhoades John and Patty Wineman donors to the following funds: Michael G. Ryan and Linda A. Joyce Carolyn P. Winn Matthew and Kelly Saal Ann S. Wolff AFAR Aging Research Program Fellowship Theodora Saal Steven and Dorothy Zottoli The Ellison Medical Foundation Sacks Family Foundation Joseph W. and Jean M. Sanger GIFTS IN KIND Robert Day Allen Fellowship Artur Scherf Constance M. and Dean C. Allard Joseph W. and Jean M. Sanger Shanti Shahani Fine Science Tools Carolyn and Joseph Shannon Gordon Gund Kate Lake Shaw Pamela D. Jacobsen American Society for Cell Biology Summer Irwin W. Sherman Maison Villatte Research Awards Timothy and Kathleen Short Christopher Neill and Linda A. Deegan The American Society for Cell Biology L. David Sibley Barbara Morgan Roberts St. Andrews Foundation William D. Cohen Endowed Scholarship Fund Eleanor K. Steinberg David Borden Jay Steinberg David and Patty Cohen Jeffery Steinberg In 2012, the MBL continued to benefit from Marion Cohen Julie Steinberg and Richard Engerman major multi-year gifts pledged in previous Sarah Cohen StemCell Technologies Inc. years from the following: Randall R. Stewart Masanori Takahashi Cornell Research Development Award Anonymous (2) Betty Taymor Molly N. Cornell The Bay and Paul Foundations Hiroshi Ueno Volker and Mary Ulbrich Millicent Bell Bernard Davis Scholarship Fund Smitha B. Uthaman Burroughs Wellcome Fund Porter W. Anderson, Jr. John J. and Frederica W. Valois The Ellison Medical Foundation Matthijs van der Meer The Grass Foundation Eugene Floyd DuBois Memorial Fund Miranda Van Gelder Howard Hughes Medical Institute Harmony Glazebrook Pittet Daria N. Van Tyne George and Harmon Logan Gordon and Betty Moore Foundation Embryology Fund George and Kathy Putnam The Company of Biologists Limited March of Dimes Foundation Society for Developmental Biology, Inc.

MBL Annual REport 2012 39 Gifts to Named Fellowships and Scholarships

The Charles Evans Research Development IBRO Scholarships for Advanced Neural Systems and Behavior Course Fund Neuroscience Courses Scholarship Fund Alice Shure/The Charles Evans Foundation USNC-IBRO Committee Anonymous Emre R. Aksay The Fischbach Research Awards Holger and Friederun Jannasch Scholarship James and Cynthia Angstadt Simons Foundation in Microbial Diversity Jay B. Bikoff Carol Arnosti and Andreas P. Teske Michael H. Dickinson Gerald D. and Ruth L. Fischbach Scholarship Bjoern Ch Ludwar Fund Benjamin Kaminer Scholarship Alex S. Mauss Gerald D. and Ruth L. Fischbach Freda Kaminer Bruce L. Mehler Joseph W. and Jean M. Sanger Teresa A. Nick, Ph.D Frontiers in Reproduction Keith Purpura Scholarship Fund Edwin S. Linton Scholarship Fund Randall R. Stewart Burroughs Wellcome Fund Washington & Jefferson College Masanori Takahashi Joseph Heitman Smitha B. Uthaman Patricia A. Hunt Logan Science Journalism Program Matthijs van der Meer Holly A. Lavoie George and Harmon Logan Daniel Wagenaar Society for the Study of Reproduction Giovanna Breu Janis C. Weeks and William M. Roberts Ronny Frishman Albert and Ellen Grass Faculty Grant Howard Hughes Medical Institute Nikon Research Awards Program Elizabeth Pennisi Nikon Instruments, Inc. The Grass Foundation Miranda Van Gelder Waksman Foundation for Microbiology Semester in Environmental Science Fund Daniel S. and Edith T. Grosch Scholarship Marselle O. Alexander-Ozinskas Fund S.O. Mast Endowed Scholarship Bryan and Donna Arling Gustav Grosch and Jane Kulesza Fred and Gloria Rhoades John Brereton Elizabeth H. Burrows Aline D. Gross Scholarship Mastroianni-Segal-Koide Scholarship Fund Bradley and Katie Cohen Freda Kaminer Elaine Pierson-Mastroianni John and Gwen Daniels Alfred M. and Naida D. Weisberg William C. Daniels James A. and Faith Miller Fellowship Fund Arthur Vining Davis Foundations Hartline MacNichol Research Award David A. and Virginia I. Miller Stephen and Lois Eisen Peter Hartline and Rebecca Kucera Andrew and Amy Feinberg Mountain Memorial Fund Andrew A. Forbes William Randolph Hearst Scholarship Constance M. and Dean C. Allard Peter and Ginny Foreman The William Randolph Hearst Foundation Brenda J. Bodian

40 MBL Annual REport 2012 Gifts to Names Fellowships and Scholarships

Sarah Q. Foster Christopher J. Freeman Society of General Physiologists Scholarships GIFTS TO MBL ANNUAL FUND Sallyann Garner Society of General Physiologists Mark Gasarch Through their annual unrestricted gifts, annual Bruce and Mary Goodman The Evelyn and Melvin Spiegel Fellowship fund donors enable the MBL to cover costs not met Peter Gottlieb Fund by grants or fees, seed key strategic initiatives, and Ellen Hertzmark Joseph W. and Jean M. Sanger provide important bridge funding. Allison E. Jacobs Hansen Johnson Burr and Susie Steinbach Research Awards The Chairman’s Circle James L. Kaplan Volker and Mary Ulbrich $25,000 or more Richard Kaplan Frank Karger Eva Szent-Györgyi Scholarship Millicent Bell Richard F. Karger Freda Kaminer Paul R. Dupee, Jr. Charles Knauss Joseph W. and Jean M. Sanger Robert and Margot Haselkorn Collin Knauss Andrew and Ursula Szent-Györgyi David and Susan Hibbitt William Kobyljanec William and Lauren Huyett Steven Miano William Townsend Porter Scholarship George and Harmon Logan Mid-American Water of Wauconda, Inc. William Townsend Porter Foundation Saul Pannell and Sally Currier John Mills Jeffrey and Margaret Pierce Donald and Patricia Mintmire J.P. and Madeleine Trinkaus Endowed George and Kathy Putnam Steve S. Moore Scholarship Patricia and Charles Robertson James Morrell Joseph W. and Jean M. Sanger Elise and Stanley Rose John and Karen O’Leary Ann Sutherland and Raymond Keller Charles M. and Phyllis Rosenthal Leslie and Donna Pinsof John and Valerie Rowe Rachel S. Poretsky Stanley W. Watson Education Fund Walter J. and Marjorie B. Salmon Gordon and Claire Prussian Anonymous Matthew and Peggy Winkler Geoffrey Ream Sacks Family Foundation Walter L. Wilson Endowed Scholarship Fund Sara Schupf Paul N. Chervin The Director’s Circle Carolyn and Joseph Shannon $10,000 - $24,999 Timothy and Kathleen Short Young Scholars Program St. Andrews Foundation David and Karen Davis Anonymous (4) The Judy and Michael Steinhardt Foundation Alison and Robert Ament Edward and Dia Weil Porter W. Anderson, Jr. Michael Wheeler Bruce A. Beal and Francis Cunningham John and Patty Wineman Ann S. Wolff

MBL Annual REport 2012 41 Gifts to the MBL Annual Fund

Norman and Diane Bernstein Sibyl R. Golden Saul and Gitta Kurlat Gary G. Borisy and Sally M. Casper Thomas F. and Virginia C. Gregg Deena and Alan Laties Mary and Jonathan Conrad Caroline Harwood and E. Peter Greenberg Anna L. and Tom Lawson Martha W. and William C. Cox Stephen M. and Cynthia R. Krane Joseph Levine and Steven A. Cadwell John E. and Judith F. Dowling Laurie J. Landeau and Robert J. Maze William W. and Noelle Locke Patrick Gage and Irina Wilson H.F. and Marguerite Lenfest David and Miriam J. Mauzerall Craig and Nancy Gibson Monique Liuzzi Sautter Nawrie Meigs-Brown and David Brown Kenneth and Nelda Johns John R. and Catherine B. Mandile Jonathan O’Herron Ethan and Lisa Lerner Jean W. Pierce R. Dana Ono and Anne Wagner Walter E. and Shirley A. Massey Dusan Stefoski and Craig Savage Arthur B. Pardee and Ann B. Goodman Richard S. and Susan G. Morse Bonnie W. and Stephen A. Simon Thomas D. and Patricia S. Pollard Edward and Linda Owens Carolyn and John Stimpson Harris and Jeanne Ripps Robert A. Prendergast John J. and Frederica W. Valois Cecily Cannan Selby Vincent J. Ryan and Carla E. Meyer Earl H. Weidner Jennifer Sorenti and Ralph Burns James and Mary Sharp Ann E. Stuart and John W. Moore William Speck and Evelyn Lipper The Lillie Society Gerard L. and Mary Swope The Seth Sprague Education & Charitable $2,500 - $4,999 John F. Swope Foundation Leonard and Eve Warren Christopher M. and Susanna B. Weld David and Sandra Bakalar Gerald and Ann Weissmann Eloise E. Clark Annette Williamson and Michael McColm The President’s Club James M. and Ruth C. Clark Kathrin R. Winkler and Angus Campbell $5,000 - $9,999 Bernice Cramer and Paul Friedman Nancy Wolf and Aric Greenfield Sally Cross Barbara Wu and Eric Larson Don Aukamp Diarmaid and Margaret M. Douglas-Hamilton Ronald Zweig and Christina Rawley David Baltimore and Alice S. Huang Joseph and Sarah Dowling John and Lisa Batter Jean-François and Rachel Formela Joshua and Lisa Bernstein Charles M. Ganson The Whitman Society George H. Billings Catherine Morrison Golden $1,000 - $2,499 Marianne and Eric Billings Philip Grant ______Francis P. and Margaret C. Bowles Mary Greer Life Members Ed and Amy Brakeman Howard H. Hiatt Frank M. and Julie S. Child Richard B. and Marcia H. DeWolfe Pamela C. and Gregory J. Hinkle Margaret Clowes Michael Fenlon and Linda Sallop Shinya and Sylvia Inoué J. Woodland Hastings Gerald D. and Ruth L. Fischbach Gary and Susan Jacobson William Jeffery and Meredith Yeager-Jeffery Pamela and David Follett Barbara Woll Jones Hans Laufer ______

42 MBL Annual REport 2012 Gifts to the MBL Annual Fund

Louise Adler Susan P. and Thomas Goux Ambrose K. and Lili Monell Blaise Aguera Arcas and Adrienne L. Fairhall Katherine Graubard and William H. Calvin James F. and Lisa M. Mooney Garland E. Allen Linda L. and Stephen A. Greyser Hiroko Naka James L. and Helene M. Anderson Joshua and Melissa Hamilton Toshio and Kyoko Narahashi Peter B. and Margaret T. Armstrong Susan Harding Hawkins Alice Netter Samuel C. and Elizabeth F. Armstrong Synnova Hayes Peter A. Nickerson Elizabeth Heald Arthur Diane E. Heck James L. Olds and Robin E. Buckley Robert G. and Harriet Baker John G. Hildebrand and Gail D. Burd Thoru and Judith Pederson Patricia Ann Barlow Timothy T. Hilton Bertha Person Jackson Barno John and Olivann Hobbie Robert W. and Carroll C. D. Pierce Susan R. and Daniel T. Barry Joseph Hoffman and Elena Citkowitz Frank Press Max M. Burger Kenneth Holden and Frances C. McGuire John S. Price R. Andrew and Barbara Cameron Howard and Sybil Holtzer Keith Purpura Colleen M. Cavanaugh and Philip M. Gschwend Carmela J. Huettner Michael and Diane Rabinowitz Mary Clutter Hugh E. and Frances M. Huxley Edward Rastetter and Karen Hendrickson Adam E. Cohen Freda Kaminer Peter W. and Elizabeth Renaghan Rita R. and Jack H. Colwell Morris J. and Shirley Karnovsky Marius A. Robinson Roberta Corliss A. Sidney and Alice Knowles Jack and Rosalyn Rosenbluth Molly N. Cornell Hans and Donna Kornberg Joan V. and Gerald S. Ruderman Thomas S. and Geraldine Crane Edward A. and Kathryn F. Kravitz Edward D. and Nancy M. Salmon Stephen D. and Beth G. Crocker Homer W. and Mary C. Lane Audrey M. Schneiderman Eric H. Davidson Marnie and John Langford Osamu and Akemi Shimomura Paul J. and Maria S. De Weer Leonard and Ruth Ann Laster Melvin and Evelyn Spiegel Leyla deToledo-Morrell Burton J. and Ann K. Lee Corinne Steel The Doverspike Family Fund John E. and Natasha C. Lisman Marjorie E. Steinberg Philip B. Dunham and Gudrun Bjarnarson Werner R. and Birgit R. Loewenstein Raymond E. Stephens Barbara E. Ehrlich Laszlo Lorand Alfred and Dorothy A. Stracher Herman N. Eisen and Natalie Aronson Michele Lorand and Paul Ringel Andrew and Ursula Szent-Györgyi A. Verdi Farman-Farmaian William K. and Winnie C. Mackey Edwin and Heather Taylor Martha S. Ferguson Heidi Gifford Melas-Kyriazi Edward and Louise Tsoi David C. and Patricia S. Gadsby Jason and Jennifer Meyers Ronald D. Vale and Karen Dell James N. and Nancy Galloway Thomas J. and Susan R. Miller John and Barbara Weyand Sallie A. Giffen Gerrish H. Milliken Elaine Yeh and Kerry S. Bloom Alfred L. and Joan H. Goldberg Timothy Mitchison and Christine Marie Field Alfred M. and Joyce V. Zeien Timothy H. and Mary Helen M. Goldsmith Merle Mizell

MBL Annual REport 2012 43 Gifts to the MBL Annual Fund

$500 - $999 $100 - $499 Vera S. Clark Gloria S. Borgese Anonymous (3) Ann P. and James M. Cleary Patrick D. and Patricia A. Burke Donald A. and Barbara R. Abt Laurence P. and Katharine B. Cloud Alfred B. Chaet Nina Stromgren Allen Maryanne and Isidore Cohen Sallie Chisholm Michael and Marie Angelini R. John and Joan M. Collier Carolyn Cohen Duncan P. and Dorothy D. Aspinwall Marc D. Coltrera and Anne L. Buchinski Beth Colt and P.K. Simonds David S. and Nancy C. Babin Nathaniel S. and Catherine E. Coolidge Harry F. Conner and Carol E. Scott-Conner Spencer L. Baird and Judy P. Spencer Jeffrey T. Corwin Robert C. and Ellen L. deGroof Hope Baker Patrica Cowan Susan Dillmann James D. Bangs Marilyn E. and J. Sterling Crandall David and Doris L. Fausch John E and Helen V. Barnes Charles M. Crane and Wendy Breuer Flagship Roofing & Sheetmetal Co., Inc. Harold and Marion Barr Karen Crawford Ruth E. Fye Cheryl Beano Joan and Jerry Cross Harold and Ruth S. Gainer David C. Beebe Prince S. Crowell, III Joseph G. Gall and Diane M. Dwyer Deborah Y. Beers-Jones Richard D. and Virginia M. Cutler William A. Haskins Edward Joseph Behrman Giuseppe and Mirella V. D’Alessio Gary G. and Nancy Hayward Marlene and George Belfort Elizabeth M. Davis Gordon E. Hering Thomas L. and Mary Jo Benjamin Joel P. and Ruth H. Davis Robert K. Josephson Francisco Bezanilla Nigel W. and Leila Daw Kiyoshi Kusano Stephen H. and C. Lynn Bishop Joseph P. Day Catherine C. and John Lastavica Thomas P. and Jane W. Bleck Robert L. and Marianne S. DeHaan Robert E. and Marjorie J. Mainer David A. and Marla Bodznick Martha Bridge Denckla Edwin McCleskey Thomas C. and Elizabeth D. Bolton Charles A. and Molly A. DiCecca Abigail Norman Thomas and Janet Brady Ellen P. Donaldson Rudolf and Nannette D. Oldenbourg Eleanor D Bronson Julie Drawbridge Ronald J. Przybylski Jennifer Byrnes Brower Arthur B. and Roberdeau C. DuBois William S. and Catherine A. Reznikoff M. Kathryn S. and Thomas A. Brown Kathleen Dunlap Howard K. and Ethel L. Schachman Judy A. Bruguiere Carol Bernstein Eckstein Douglas R. and Virginia M. Shanklin Harold L. and Joan N. Burstyn Frank Egloff, II Roger D. and Carol J. Sloboda Amy K. Butler Lincoln and Ruth Ekstrom David and Mona Spector Graciela C. Candelas Paul Elias and Marie Lossky John and Evelyn Steele Thomas Cannold Gordon W. and Joyce M. Ellis Alan B. and Sala Steinbach Margaret W. and Samuel B. Carr David and Lois Epel Edward and Anne Stimpson John and Sandra Checklick Doris W. Epstein Gary Stormo Children’s School of Science, Inc. Thomas E. Evans Margaret W. Taft John A. Christian Robert R. and Ann Everett David M. and Jeanne D. Travis Ellen S. and George B. Citron Patricia M. Failla Richard and Dorothy Verney Thomas and Rachel Claflin Robert and Joanne Fallon Frances Weiffenbach

44 MBL Annual REport 2012 Gifts to the MBL Annual Fund

Elizabeth M. Farnham Robert and Taska Hener Vanessa LeFevre Marla B. Feller Peter K. and Margaret H. Hepler Robert Owen Lehman Max and Martha Fink Catherine E. Hibbitt and Tim Rockwell David P. Lenzi Rachel D. Fink Richard W. and Susan D. Hill Jack and Francine Levin Alan and Joan Finkelstein Seymour and Victoria E. Holtzman Francoise Levinthal Bruce and Virginia Finnie Erich F. Horgan James S. and Alice J. Liljestrand Michael J. and Joann Fishbein Edward G. Horn Lennart and Ann C. Lindberg Harvey M. and Susan Fishman Ronald R. Hoy and Margaret C. Nelson Raymond J. Lipicky Elizabeth Frank Nicholas A. and Laura M. Ingoglia Frank J. and Kathryn L. Longo Arthur and Linda Freeman Laurinda A. Jaffe Victoria H. and Francis C. Lowell Hadley Mack French Ernest G. and Pauline B. Jaworski Janet S. Loynes John J. and Avery T. Funkhouser Elizabeth Ann Jonas and Thomas D. Eisen Louise M. Luckenbill Barbara P. Gaffron Sally S. and Ramsey E. Joslin Robert P. Malchow Frank P. and Joyce Gallagher Yoshie Kamei James Marcello Michael R. and Mary Garfield Michael G. Kauffman Dawn Morin Marick Stephen E. Gellis M. Whitney Keen Julian B. and Priscilla K. Marsh Elizabeth L. George Katherine Kennedy Lowell V. and Ann J. Martin Thomas Gewecke Thomas C. and Mary Ann Kenny Jack Martinelli and Nancy Couts Cameron E. and Margaret J. Gifford Louis M. and Brenda Kerr Timothy C. Marzullo Barbara B. Glade Nasim Khan Richard B. McElvein Amy S. Gladfelter Alexey L. Khodjakov Robert B. McKane Mary L. Goldman and Debra Weiner Min Kyung Kim Jane A. McLaughlin Charlotte O. Goodwin Betsy King Martin Mendelson Frederic and Jocelyn Greenman Gilbert and Katharine King Wilhelm and Nonie Merck Huson and Edith Gregory Richard H. and Reta King Melanie and Klein Merriman Newton H. and Evelyn Gresser John M. and Louise Kingsbury Nancy S. and Richard Milburn Alexandre Grigorovitch Peter N. Kivy and Joan Pearlman Ricardo and Mela Miledi Albert and Sondra Grossman Norman R. Klath David A. and Virginia I. Miller Gregg Gundersen Paul W. and Virginia Knaplund Holly Miller Gilbert F. Gwilliam Edward M. Kosower T. Richardson Miner and Bobbie Loop John F. and Sandra Halsey John and Susan Krezoski Robert and Maura Morey Lisa M. Halvorson Andrew M. Kropinski and Peggy A. Pritchard James T. Morris Tom and Lisa Hammett Alan M. and B. J. Kuzirian Day O. and Kathie C. Mount Clifford V. Harding and Mina K. Chung Kevin L. and Sallie D. Lake Christina J. Myles-Tochko Glenn W. Harrington George M. and Sylvia T. Langford Angus Nairn and Marina Picciotto Dorothy Haugaard Louis S. and Camilla B. Larrey Enrico Nasi and Maria E. Gomez Marian B. and Edmund Hazzard Howard S. Lasher Phil Nehro Edward E. Heller Joyce Whitehead Lathbury Ann Little Newbury Simone Helluy Marian E. LeFevre David L. and Linda D. Newton

MBL Annual REport 2012 45 Gifts to the MBL Annual Fund

Peter M. and Virginia Nicholas Thomas J. Schnitzer Kensal and Barbara van Holde Santo V. Nicosia Herbert Schuel Carol A. and Stephen P. Wagner Lee A. Niswander Harold H. and Kristin T. Sears Brant G. Wang Ana Lia Obaid Robert and Carole Seidler James D. Watson William and Sarah Odenkirk Deborah G. Senft Annemarie Weber Janice and Albert Olszowka Harriet S. and Howard E. Shapiro John T. and Jeanne T. Weeks Jeffery and Valerie Oppenheim Gaius and Eleanore Shaver Ellen Weiss Lorrence J. and Beverly C. Otter Kate Lake Shaw Robert H. Werner Costa and May Papastephanou James M. Sidie Gary M. Wessel David and Lois Parker, Jr. Alan R. and Phyllis J. Silver Paul S. and Barbara B. Wheeler James L. Parmentier and Elizabeth Fowler Arthur and Frances Silverstein Tanya B. White Robert H. and Pamela D. Pelletreau Eric J. Simon Roland L. and Nancy Wigley Matthew A. Person and Jill Erickson Rolly L. Simpson Clare M. Wilber Carey R. and Ryan J. Petrie Maxine F. Singer Clayton A. and Arlene Wiley Helen and David Piwnica-Worms Stephen Skillman Gerrard Wilson David E. and Jeanette Pleasure Roxanna S. and Ronald J. Smolowitz Jonathan B. and Beatrice Wittenberg George H. and Louann S. Plough John W. and Cathleen Speer Charles R. and Ellen G. Wyttenbach Jeanne S. Poindexter Lucy Steere Linda A. and Erik R. Zettler Harvey B. and Bette S. Pollard William K. and Jane B. Stephenson Michael J. and Naomi Zigmond Aubrey and Margareta B. Pothier Judith G. Stetson Bruce and Elizabeth I. Zimmerli Donald L. Price Elijah W. Stommel and Jasmin Bihler Donald J. Zinn James P. and Joan M. Quigley Alexandra G. Stone Frederick R. and Kathryn G. Rickles Mutsuyuki and Shirley M. Sugimori Up to $100 Philip S. and Anne Robertson William C. and Karen A. Summers Anonymous (1) Dana and Alison Rodin Aron and Irene Szulman Robin W. Ackroyd Lawrence C. and Victoria H. Rome Marvin L. and Betsy C. Tanzer Peggy and Fred Alsup John D. and Martha A. Ross Martha E. Tarafa Ruth Anderson William N. Ross and Nechama Lasser-Ross Peter Tassia and Maija Lutz Mike and Elana Arnold Rudi J. and Hanna Rottenfusser William R. Thomas Shinobu Asano William A. and Andrea Rugh W. Nicholas and Joan I. Thorndike Barnstable Intermediate School Alice M. Rushforth Mary and James Thorne Dennis Barrett Norman B. Rushforth Emil D. and Eleanor H. Tietje Kenneth Bento Robert N. Sandberg Noah and Janet B. Totten Jane A. Berger and Roger O. Gittines Christopher Sanders James G. and Mary Elizabeth Townsel Olive C. Beverly Noriyuki Satoh Natalie Trousof Dieter and Gigi Blennemann Anne W. Sawyer Tsuchiyra High School Courtney M. Bowker Jay Schippers James Tucker Barbara C. and John F. Boyer Paul R. Schloerb Kenyon S. and Joan Tweedell Charles and Helen Bresnahan Thomas M. and Susan Schmidt Volker and Mary Ulbrich Jerí Bryant

46 MBL Annual REport 2012 Gifts to the MBL Annual Fund

Ann C. Bucklin and Peter Wiebe Kenneth H. Goldstein Shanta Messerli Ronald and Vivian Campbell Sarah C. Goodrich Margaret A. Mills-Michael Joanne B. Carnevale Louise and Jim Graham Ruel and Alice Mohnkern Robert H. and Myra S. Carrier Edwin R. Griff Stephen A. and Marjorie H. Moore John M. and Theresa M. Clarkin Lawrence I. and Esta S. Grossman Isabelle Mort Graeme Cloutte Matthew Hackenberg Thomas C. and Elizabeth D. Moseley Mort and Susan Cohan Ronald J. Haley Christoph Mueller Elizabeth Ann Cohen Maureen and Arthur Harlow-Hawks Thomas A. Mulholland Seymour S. Cohen Pamela D. Harvey Paul and Mary Mulloy Concord Christian Academy David S. Hays Iris Nelson-Schwartz Priscilla N. and Alan Conger Rodney Hinkle and Kirstin A. Moritz Jill Neubauer Peter and Susan Connolly Lawrence and Layne Hobbie Eliot Nierman Lou-Anne Conroy Ann Snow Hobbs Shinpei and Catherine B. Ohki Melvin C. and Margery L. Crain Holliston High School Dennis Ojima and Jill Baron Jeffrey G. Croll Betsy and John Honey Harish C. and Kamala Pant Dorothy I. Crossley Richard A. and Susan M. Houghton Charles and Patricia P. Parmenter Syrel Dawson Julie Hysom Leonard M. Passano Joseph and Esther Dresner Leonard K. and Sheila F. Kaczmarek Cara Pellegrini Dorothy L. Drummey Harry S. Kahn Joyce S. Pendery Quan-Yang Duh and Ann Comer Edna S. Kaneshiro Nancy Pendleton Allan and Linda Dunn Robert G. Kemp Barbara C. Perry Rita and Paul E. Dussault Michael and Jennifer Kosowsky William G. Phillips Hoyt and Deborah Ecker Michael J. Landzberg Sara C. Pimental Frank Egloff Susan E. Leeman Stephen and Robin Plattman Carlyn Ellms Herbert and Karen B. Levitan Jessica K. Polka Ruth Alice Fitz Elizabeth and Paige L’Hommedieu Daniel A. and Linda K. Pollen John W. Fleeger William and Joan M. Lloyd Catherine Prendergast John H. Ford Lawrence Lynch Corporation Anne T. Pressman Franklin W. Fort Margaret M. MacLeish Fred J. and Catherine E. Ravens Alvin K. Fossner Jane and Daniel MacNeil Willis B. and Joan Reals Peter Franklin Patrick J. and Lorraine Mahoney Carol C. Reynolds Nancy and Joseph V. Fraser Frank T. and Ose Manheim Mary Elizabeth Rice Paul J. and Patricia S. Freyheit Charles K. and Susanne J. Mann Robert V. Rice Larry Jay Friedman Diane B. and Joseph S. Maranchie Alison A. Robb William Fuqua Joseph C. and Connie Martyna Jonathan Rockoff Michael and Bridget Gabriel Joseph and Darlene McGuire Warren Rogers and Linda St. Jacques-Rogers Patricia E. Garrett and Oliver H. Woshinsky Gary McHatton Helen Ross-Sangster William and Joanne Gilbrook Cornelia Hanna McMurtrie Daniel and Nancy Rubenstein Ann I. Glusker Samuel McMurtrie, Jr. Richard Sailor and Mary Johnston

MBL Annual REport 2012 47 Gifts to the MBL Annual Fund, cont.

Thomas and Judith S. Sbarra 2012 MBL GALA PATRONS OF THE CONCERT Mary L. Scalera Gail Cooper Ralph Schiller The MBL thanks Margot and Robert Haselkorn Sallie and Michael Corgan John F. and Diane S. Schoenfelder for their generosity in sponsoring the 2012 Gala Gloria Cormack School of the Holy Child concert, in which the renowned Tokyo String Prince Crowell Paul Selvin Martha Bridge Denckla Hannah Senft Quartet made their final MBL performance before Dick and Cathy Fay Joan Sharp retiring from the international stage. We are also Padraic J. Shay deeply grateful to the following sponsors, hosts, Pat Ferguson Gordon M. Shepherd patrons, and many others who made the 2012 MBL Prosser Gifford Stephanie M. Sit Gala a great success, raising more than $43,000 Ginny and Tom Gregg James Slock for the MBL Annual Fund. Janis and Buzz Harvey Sven and Anna Spoerri Olivann and John Hobbie Charles Stasey CONCERT SPONSORS Barbara Woll Jones John J. Stegeman Margot and Robert Haselkorn Birgit and Werner Loewenstein Ann Stern Winnie and Bill Mackey Daniel E. Strongin Lalise and Jerry Melillo Brad Stumcke RECEPTION SPONSORS Jeanne Guillemin and Matthew Meselson Alexander O. Sy Associates of Cape Cod Ann Stuart and John Moore Tabor Academy Diane and Norman Bernstein Frank and Loretta Tang Elise and Stan Rose Ann Goodman and Arthur Pardee Barbara H. and William A. Toomey Shawmut Design and Construction Frank Press William E. and Belle Traver John Price Barbara S. Trela RECEPTION HOSTS PrintSynergy Solutions Barbara S. Trelstad Gary Borisy and Sally Casper Willis Reals Michael and Frances W. Tytell Patricia and Paul Hogan Jack and Valerie Rowe Irving and Vida Wagner Marjorie and Walter Salmon Mary J. Walsh Hub International New England Karen and Tom Tierney Harriet Segal Kerry Walton Cecily Selby James Wei Dyann and Peter Wirth Marilyn Sheprow Phyllis R. Wendorff Gerry and Mary Swope Donn and Fluffy Winner GALA FRIENDS Ashley Wires Barbo’s Furniture Richard Taylor Louise Witonsky Flagship Roofing & Sheetmetal Co., Inc. Vincent Associates Charles Yanofsky Patrick Gage and Irina Wilson Ann and Gerry Weissmann Abigail Young Lisa and Ethan Lerner Ronald Zweig and Christina Rawley Morton and Nurit Zachter Harmon and George Logan Harold H. Zakon and M. Lynne McAnelly Saul Pannell and Sally Currier Kenneth H. and Patricia E. Zimble Alexander and Judith S. Ziss Tsoi Kobus & Associates, Inc. W.B. Mason

48 MBL Annual REport 2012 GIFTS TO THE MBL Edward M. Blumenthal David D. Dow ALUMNI FUND Andrew W. Bollen David and Sherri Durica Edward Michael Bonder Laurel A. Eckhardt Henry Bowers Marcia Edwards MBL alumni and faculty from 42 states and 6 Brigitte F. Brandriff Robert Eimstad countries made gifts to the Alumni Fund to support David N. Breslauer Judith S. Eisen the 2012 MBL discovery courses. Burt V. Bronk Paul T. Englund and Christine S. Schneyer Alumni Fund Donors Donald D. and Linda W. Brown Ali Eroglu Susan M. Burden-Gulley Arnold G. and Paula B. Eversole Robert David Bynum Suzanne Fachon-Kalweit Anonymous (9) David and Janet Campbell Nancy L. Falk Laurence and Catherine Abbott Patricia Campbell Alan Scott Fanning Penny S. Amy Cynthia S. Cappello George Feher William DeWitt Andrus, Jr. Ashley R. Carter Caleb E. Finch Robert and Lynne Angerer Robert G. Cassens Karl W. Flessa and Mari N. Jensen Erin and Barry Aprison Chenbei Chang James A. Foster William F. Arndt, Jr. Lisa Y. Chiang Kenneth Freedman and Steven Weiss Elizabeth I. Arnold Magdalena B. Chrzanowska-Wodnicka Hugo D. and Anita R. Freudenthal Senthil Arumugam George A. Clark William Frost and Lise S. Eliot Michael S. Ascher Annette W. and John R. Coleman Anne E. Fry Robert F. Ashman Elizabeth B. Conant Edwin J. Furshpan and Shelagh A. Foreman Serapio M. Baca Susan B. Conaty-Buck Arthur G. Gaines Katherine Bachman and Robert Fleming Edwin L. Cooper Maureen A. Gannon Michael J. Baltzley Jeffrey T. Corwin John A. Garcia Madelyn M. Baran Pamela A. Coutchie Jennifer H. Garvin Robert Barker and Marla Schay Baker John Cunniff, M.D. Susan Gerbi-McIlwain Nelson R. Barton Mary Ellen M. Cunningham Cameron E. and Margaret J. Gifford David H. Beach James F. Cupo Helen W. Gjessing Elizabeth Kujawinski Behn Sandra L. Dabora Michele Gallati Glasgow Jerry A. Bell and Mary Ann Stepp Charles Derby James A. Glazier William H. Bergstrom Bruce A. Diner Margaret Ann Goldstein Gerald Bergtrom Long Ding Gary Gorbsky Joel C. Berman Thuy A. Doan Joel S. Gordon Joan M. Bernhard Laneta Dorflinger Martin Gorovsky Maja I. Bialecka-Fornal Dustin L. Dovala Stefan J. Green Dawn M. Blitz

MBL Annual REport 2012 49 Gifts to the MBL Alumni Fund

Debra S. Grega David Kleinfeld Wolfram Moebius Leslie G. and James K. Gregg-Jolly Alan Klotz Ronald B. Moir and Toni Czekanski Charuni A. Gunaratne James J. Knierim Marius R. Moran Janet Haas Robert E. Knowlton Walter J. Mozgala E. Janet Hager Birgit Kovacs Kenneth E. Najarian John E. Hall George H. Kowallis David E. Naylor Kristen M. Harris and Max Snodderly Keith G. Kozminski Ruth H. Neff James R. Henriksen Deborah A. Lans Ralph F. Nelson Scott B. Herrick Jeffry B. Lansman Jeanne M. Nerbonne Sharon E. Hesterlee Melinda Larsen and Michael Gerdes Jeniel E. Nett Ralph Hillman Bruce B. Latham Dana T. Nojima Akira Hishinuma Matt K. Lee Michael and Carol Oberdorfer Sarah E. Hitchcock-DeGregori William J. Lehman Susan Offner Raymond W. Holton Ardean Leith Stephanie A. Olexa Timothy E. Holy Ellen K. LeMosy Brett A. and Kathy Oxberry Christine L. Howe Steven William L’Hernault Joanne and Alan Page Charles E. Huntington Anne M. Linton Kelli L. Palmer Deborah Hursh and Mark Mortin Jennifer Lippincott-Schwartz Melissa J. Perry Antje Ihlefeld Joanna E. Lowell Flynn Picardal Kachi Illoh Howard W. Lowy Louis Pierro Richard and Nancy Intres Gary E. Lyons Mark Plummer Jon W. Jacklet Stephen E. Malawista William K. Plunkett Laurinda A. Jaffe Phillip B. and Yvonne R. Maples William F. Pomputius, III Daniel E. James Junko Munakata Marr Richard Purple Nancy A. Johnson Sandra K. Masur Esther L. Racoosin Fred I. Kamemoto Allen Wray Mathies, Jr. George Radcliffe Karen E. Kasza Joan Mattson Wilfrid Rall Seth N. Kauppinen Anne E. McBride Jeffrey L. Ram Gordon I. Kaye Megan N. McClean Alfred G. and Sarah C. Redfield Murat Kaynar Kathryn E. McCormick Jean F. and Ronald R. Regal R. Emmet and Gail Kenney Matthew McFarlane and Jennifer Nadeau Marian C. Rice Lynda A. Kiefer Everett Mendelsohn Mary Esther Rice Jennifer M. King Sara Michie Austen F. Riggs David Louis Kirchman Lynn Miller Ramona M. Rodriguiz Marc W. and Phyllis Kirschner Edwin A. Mirand Richard E. Rohr

50 MBL Annual REport 2012 Gifts to the MBL Alumni Fund

Brian K. Romias Roger Elliot Thannerthies FALMOUTH FORUM Allen Rosenspire Holly Thompson ENDOWMENT FUND Pierre E. Rouviere Carl L. Thurman, II Uldis Roze Laurence C. Torsher The following donors contributed to the Falmouth Laurens N. Ruben Ann H. and Mark F. Trax Forum Endowment Fund in 2012. Jan S. Ryerse Jeanine A. Ursitti Wendy C. Salmon Anthony Vattay Anonymous (2) Robert J. Saqueton Antonio Ventosa David and Sandra Bakalar Suzaynn F. Schick Susan Volman Catherine B. Cramer Paul R. Schloerb Yihong Wang Prosser Gifford Joseph H. Schneider Gary Ward and Zail Berry Robert H. Werner David Schoppik Samuel and Anne Ward Ethan Schrank Joseph T. Warden Andrew Schutrumpf Jane E. Warshaw MEMORIAL AND TRIBUTE GIFTS Terrence J. Sejnowski Coreen A. Waters Deana Shackelford Watts Wayde M. Weston These donors have chosen to support the MBL as a Alan and Stephanie Sher Harold Bancroft White special way to remember or honor a relative or friend. Philip J. Skehan, Jr. John White and Sonia Witte George J. Skladany Eric Wieschaus and Trudi Schupbach Biology of Parasitism Endowed Fund Samuel J. Sober Athula H. Wikramanayake Irwin W. Sherman in memory of: Lilianna Solnica-Krezel Linda E. Wilbrecht Villa G. Sherman Alan M. Sooho Benjamin Douglas Williams John E. Spiro John S. and Judith H. Willis Thomas A. and Gloria S. Borgese Rare Book Joel P. Stafstrom Helene D. Winstanley and Journal Fund Barbara A. Stay Stephan N. Witt Gloria S. Borgese in memory of: Robert E. Steele George Wittenberg Thomas A. Borgese Joan A. and Thomas A. Steitz Robert G. Wolk Daniel E. Strongin Zeba B. Wunderlich William D. Cohen Endowed Scholarship Fund Charles H. and Donna J. Sullivan James Allen Wyban David S. Cohen, Marion Cohen, and Sarah David T. Sullivan Judith L. Yanowitz and Harry Hochheiser Cohen in memory of: Kelly J. Suter Patrice O Yarbough William D. Cohen Harold E. Sweetman Anne-Marie C. Yvon Ben G. Szaro Veronika I. Zarnitsyna Eugene Floyd DuBois Memorial Fund Alexis Tapanes-Castillo Steven Zeichner Harmony Glazebrook Pittet in memory of: D. Lansing and M. Margaret Taylor Jiawei Zhang Rebeckah D. Glazebrook Saul Teichberg Andrew W. and Lenore N. Zimmerman

MBL Annual REport 2012 51 memorial and Tribute GIFTS

The Harlyn Halvorson Fund Gottlieb, Richard Kaplan, Frank Karger, Richard F. Elizabeth M. Farnham Andrew and Ursula Szent-Györgyi in memory of: Karger, Mid-American Water of Wauconda, Inc., Alan M. Sooho Harlyn O. Halvorson John Mills, Leslie and Donna Pinsof, Gordon and Claire Prussian, St. Andrews Foundation, Edward Anne E. Goldman Tay Hayashi Lectureship in Cell Physiology and Dia Weil, John and Patty Wineman, and Ann David and Mona Spector Fund S. Wolff in memory of: Andrew and Ursula Szent-Györgyi in memory of: Harold E. Foreman Harlyn O. Halvorson Teru Hayashi Lisa M. Halvorson Sustainable Aquaculture Fund Robert Huettner Rare Book Endowed Fund Anonymous in honor of: Pamela Clapp Hinkle Carmela J. Huettner in memory of: Dorianne Mebane Rodney Hinkle and Kirstin A. Moritz John Alexander Hogg Robert Huettner Eva Szent-Györgyi Scholarship Fund John E. Hobbie Andrew and Ursula Szent-Györgyi in memory of: Lawrence and Layne Hobbie Library Fund Eva Szent-Györgyi Elaine Cahn, Paul J. and Maria S. De Weer, and Elizabeth A. Jonas Bertha Person in memory of: Whitman Center General Endowment Fund Kenneth H. Goldstein Leon P. Weiss Hans Laufer in memory of: Mona Gross, Alexander Keynan, Malcolm S. Eugene P. and Adelaide Kennedy Neural Science and Behavior Scholarship Fund Steinberg, Byron H. Waksman, and Leon Katherine Kennedy James D. Angstadt in honor of: P. Weiss Ronald L. Calabrese Marine Biological Laboratory Library Wolfgang O. Friesen Annual and Alumni Funds Ann I. Glusker William B. Kristan Gifts made in honor of: Michael H. Dickinson in honor of: Richard W. Linck Masakazu Konishi Ned Cabot Dana T. Nojima Ronald B. Moir and Toni Czekanski Rare Books Fund Luciane Marcus Anne Nelson in memory of: Ercole Canale-Parola Thomas Cannold Leonard Nelson Jane E. Warshaw Joe L. Martinez Semester in Environmental Science Maggie Carmody Ramona M. Rodriguiz Christopher J. Freeman in memory of: Joan Sharp Beth Chenicek Jeffrey Pierce Bradley and Katie Cohen, Peter and Ginny John Crane Tom and Lisa Hammett Foreman, Bruce and Mary Goodman, Peter Margaret M. MacLeish

52 MBL Annual REport 2012 Memorial and Tribute GIFTS

Warren Rogers Barbara Atwood Herman T. Epstein Luigi Mastroianni, Jr. Warren Rogers and Linda St. Elizabeth M. Farnham Alan R. and Phyllis J. Silver Santo V. Nicosia Jacques-Rogers Barbara P. Gaffron John J. and Frederica W. Valois Herbert Seymour Festoff Kathryn Mendelson Charles M. Rosenthal Barry William Festoff Martin Mendelson Howard S. Lasher Ann W. Aukamp Patrick Gage and Irina Wilson Jeanette Fossner Harold M. Mills, Jr. Joseph P. Senft Alvin K. Fossner Margaret A. Mills-Michael Hannah Senft Robert B. Barlow, Jr. Michael and Carol Oberdorfer David L. Garbers Frank Morrell Kate L. Shaw Carey R. and Ryan J. Petrie Thomas P. and Jane W. Bleck Kevin L. and Sallie D. Lake Evelyn Bestic Noah and Janet B. Totten Margaret C. L. Gildea Mark Mozgala Andrew G. Szent-Györgyi Frank Egloff, II Walter J. Mozgala Sarah E. Hitchcock-DeGregori Valentin and Elisabeth Braitenberg Cornelia Hanna McMurtrie Walter Heiligenberg William D. Russell-Hunter William Ward George Wittenberg Arnold G. and Paula B. Eversole Warren Rogers and Linda St. William D. Cohen Jacques-Rogers William and Sarah Odenkirk Gertrude N. Illoh Sheldon J. Segal Kachi Illoh Harold and Marion Barr Steven J. Zottoli Robert A. Cowan Jason and Jennifer Meyers Patrica Cowan Hekimcan Kaynar Malcolm S. Steinberg Murat Kaynar Shinobu Asano Christopher Dillmann Julie Drawbridge Gifts made in memory of: Susan Dillmann Salih Kirli Bruce and Virginia Finnie Murat Kaynar Gary Gorbsky Louise S. Armstrong Eric Christopher Dillmann Norman R. Klath Samuel C. and Elizabeth F. Susan Dillmann James W. Lash Michael and Jennifer Kosowsky Armstrong Joel S. Gordon Joyce Whitehead Lathbury Nancy Egloff Elizabeth and Paige L’Hommedieu Dr. Philip B. Armstrong Frank Egloff, II Anthony Liuzzi Costa and May Papastephanou Samuel C. and Elizabeth F. Monique Liuzzi Sautter Daniel and Nancy Rubenstein Armstrong Dora Epstein Helen Ross-Sangster Susan E. Leeman Joyce Lorand Robert N. Sandberg Laszlo Lorand Stephen Skillman Michele and Paul Lorand Marjorie E. Steinberg

MBL Annual REport 2012 53 Memorial and Tribute Gifts, cont.

Barbara S. Trelstad Martin K. White David and Lois Epel James Wei Tanya B. White Martha S. Ferguson Louise Witonsky Yvonne R. Fuortes Morton and Nurit Zachter Charles G. Wilber Bruce and Barbara Furie Clare M. Wilber Susan Gerbi-McIlwain Albert Szent-Györgyi Gilbert W. Glass Elizabeth I. Arnold Donald and Margery Zinn Sol H. Goodgal Jeffrey A. and Johanna A. Zinn Philip Grant Eva Szent-Györgyi J. Woodland Hastings Sarah E. Hitchcock-DeGregori Gertrude W. Hinsch New Century Society Joseph Hoffman and Elena Citkowitz John Hervey M Howard and Frances F. Jacobson Richard Purple The New Century Society recognizes and honors Mary D. Janney those who have made bequests and other deferred Daniel and Jean Johnston J. P. Trinkaus gifts arrangements that will benefit the MBL in Sally S. and Ramsey E. Joslin Lilianna Solnica-Krezel the future. We thank all the members of the New Jane C. Kaltenbach-Townsend Century Society for their generosity. Darcy B. Kelley Lee VanBremen Richard G. Kessel Judy A. Bruguiere Anonymous (6) Andrew M. Kropinski and Peggy A. Pritchard Peter B. and Margaret T. Armstrong Hans Laufer Walter S. Vincent Patricia Ann Barlow Paul B. Lazarow Douglas R. and Virginia M. Shanklin Joan R. Berman Edward R. and Gloria A. Leadbetter Margaret C. and Francis P. Bowles Marian E. LeFevre Byron H. Waksman Mario H. and Aida E. Burgos Howard W. Lowy Children’s School of Science, Inc. Ronald Calabrese and Christine Cozzens Gordon MacIver and Eleanor MacIver Maryanne and Isidore Cohen Graciela C. Candelas Robert E. and Marjorie J. Mainer Elizabeth M. Davis James F. Case Melanie and Klein Merriman Adolph S. Cavallo John W. Moore and Ann E. Stuart Leon P. Weiss Paul N. Chervin Arthur B. Pardee and Ann B. Goodman Cheryl Beano Julie S. and Frank M. Child Jeanne S. Poindexter Stephen and Robin Plattman Lawrence B. Cohen Thomas D. and Patricia S. Pollard Harris and Jeanne Ripps Seymour S. Cohen Robert A. Prendergast John E. and Judith F. Dowling Ronald J. Przybylski Lillian Wendorff Joseph and Sarah Dowling Michael and Diane Rabinowitz Phyllis R. Wendorff Paul R. Dupee, Jr. Monica Riley Barbara E. Ehrlich Michael G. Ryan and Linda A. Joyce

54 MBL Annual REport 2012 Cecily Cannan Selby ATCC Graftek Imaging Inc. Enid K. Sichel AutoMate Scientific, Inc. Grass Technologies / Astro-Med, Inc. Roger D. and Carol J. Sloboda Marjorie E. Steinberg BD Accuri Cytometers Hamamatsu Corporation Ann E. Stuart and John W. Moore BD Biosciences Hamilton Thorne, Inc. John Tochko and Christina Myles-Tochko Beckman/ Coulter, Inc. Flow Cytometry Harvard Apparatus, Inc. John J. and Frederica W. Valois Beckman / Coutler Inc. HEKA Instruments, Inc. Susan Volman Bioptechs Herzan, LLC Byron and Joyce Waksman Bio-Rad Laboratories Horiba Jobin Yvon Inc. Earl H. Weidner BioTek Instruments, Inc. Clare M. Wilber Bitplane Inc. Imagine Optics / Axiom Optics John S. and Judith H. Willis Boston Electronic Corporation Inteligent Imaging Innovations Brownlee Precision Company INTRACEL Limited Invitrogen Corp. / Life Technologies Cambridge Electronic Design Ltd. ISS Corporate Lending Program Carestream Health Carl Zeiss Microscopy, LLC Jackson Immunoresearch Laboratories, Inc. The quality and success of the MBL educational Cell Signaling Technology Inc. Journal of Histochemistry and Cytochemistry program is maintained through research equipment, Charles River Laboratories, Inc. reagents, and computers valued at $26 million Chroma Technology Corporation Keyence Corporation of America loaned by: Coherent Kramer Scientific Conoptics Inc. USA 1st Vision Inc. Coriell Instute for Medical Research Leica Microsystems Inc. Leica Microsystems Biosystems Division Abcam Dagan Corporation Leica Microsystems Confocal Systems North AD Instruments DataWave Technologies America Adams & List Associates Diagnostic Instruments, Inc. Leica Microsystems Nanotechnology Division Agilent Technologies, Inc. Digitimer Ltd. LI-COR Biosciences, Inc A-M Systems Drummond Scientific Co., Inc. Life Technologies AMG / Advanced Microscopy Group Lonza Walkersville Inc. Andor Technology Electron Microscopy Sciences Ludl Electronic Products Ltd. Applied Biosystems / Life Technologies Eppendorf North America Ludlum Measurements Applied Precision / GE Healthcare Luigs & Neumann Applied Scientific Instrumentation, Inc. FHC, Inc. Lumen Dynamics Group Aquatic Habitats Fine Science Tools Asylum Research Fisher Scientific Fluidity Software, Inc.

MBL Annual REport 2012 55 Corporate Lending Program, cont.

Mathworks QIMAGING Unisense A/S MatTek Corporation Quest Scientific Instruments Inc. MBF Bioscience Varian / Agilent Technologies Micro Video Instruments Inc. Rapp OptoElectronic GmbH Vector Laboratories Miltenyi Biotec Inc RedShirtImaging, LLC Veeco Instruments GmbH Millipore Corporation Research Precision Instruments Co., Inc. Vibratome / Leica Microsystems, Inc. Molecular Devices, Inc. Roche Diagnostics Corp. Viewpoint Life Sciences MyNeuroLab / Leica Microsystems Inc. Visiopharm A/S SAGE Publications Vortran Laser Technology, Inc. Nanodrop / Thermo Fisher Scientific Scientifica Ltd. VWR International Nanosight Ltd. SciMedia Narishige USA, Inc. ScopeLED, A DiCon Brand Warner Instruments National Instruments Seahorse Bioscience, Inc. WaveMetrics, Inc. Neuralynx Inc. Semrock, Inc. / a unit of IDEX Corporation Wescor, Inc. – an ELITech Group Company Nevtek SIMPore Inc. World Precision Instruments, Inc. Nexcelom Bioscience LLC Siskiyou Corporation Nikon, Inc. Solamere Technology Group Inc. Yokagawa Corporation of America Nikon Confocal Systems Soma Scientific Instruments, Inc. Noldus Information Technology Spectra Physics, A Division of Newport Corp. Stoelting Co. / ANY-maze Olink Bioscience Sutter Instrument Company Olympus America, Inc. – Scientific Equipment Group Technical Manufacturing Corporation Omega Optical, Inc. Ted Pella, Inc. The Jackson Laboratory Parker Hannifin Corp. Thermo Scientific Pearson / Brock Education Thorlabs Inc. PerkinElmer Life and Analytical Sciences TILL Photonics USA Photometrics Titertek Instruments, Inc. Photron USA, Inc. Tokai Hit Prairie Technologies, Inc. Toohey Company Prior Scientific, Inc. Tucker-Davis Technologies Prizmatix Ltd. Protech International, Inc.

56 MBL Annual REport 2012 Leadership

MBL Annual REport 2012 57 MBL LEadership

Chairman of the Board of Trustees John W. Rowe, Columbia University BOARD OF TRUSTEES DIRECTOR EMERITI Robert H. Ament, Esq., Ament Law Firm, LLC Gary G. Borisy Vice Chair of the Board of Trustees John E. Burris William I. Huyett, McKinsey & Co., Inc. Bruce A. Beal, The Beal Companies Millicent Bell, Boston, MA Paul R. Gross Speaker of the CORPORATION Rita R. Colwell, University of Maryland William T. Speck Colleen M. Cavanaugh, Harvard University Thomas S. Crane, Esq., Mintz, Levin, Cohn, Ferris, Glovsky PRESIDENT EMERITUS and Popeo, PC President and Director John E. Dowling Joan V. Ruderman, Marine Biological Laboratory Paul R. Dupee, Haddeo Partners, LLP Gerald R. Fink, Whitehead Institute, Cambridge, MA TRUSTEES EMERITI Treasurer of the CORPORATION L. Patrick Gage, enGage Biotech Consulting Paul R. Dupee, Haddeo Partners, LLP Norman Bernstein James N. Galloway, University of Virginia Margaret C. Bowles Clerk of the Corporation Caroline S. Harwood, University of Washington Alexander W. Clowes Robert H. Ament, Esq., Ament Law Firm, LLC Robert Haselkorn, University of Chicago Mary B. Conrad H. David Hibbitt, Bristol, RI Eric H. Davidson Chairman of the Science Council Gerald D. Fischbach David C. Gadsby, The Rockefeller University William I. Huyett, McKinsey & Co., Inc Ethan A. Lerner, Massachusetts General Hospital Robert D. Goldman Mary Greer George W. Logan, Charlottesville, VA Kurt J. Isselbacher Walter G. Massey, School of the Art Institute of Chicago M Howard Jacobson Ambrose K. Monell, Palm Beach, FL George M. Langford Richard S. Morse, Jr., Woods Hole, MA Burton J. Lee R. Dana Ono, VIMAC Ventures, LLC Irwin B. Levitan Saul J. Pannell, Wellington Management Company, LLC Paul A. Marks Jeffrey H. Pierce, Pierce Aluminum Co. Jean W. Pierce Robert A. Prendergast, Falmouth, MA Thomas D. Pollard George Putnam, New Generation Advisors, LLC Frank Press Patricia K. Robertson, Rockland, DE Vincent J. Ryan John W. Rowe, Columbia University Gerald Weissmann Christopher M. Weld Elise Rose, Falmouth, MA Annette L. Williamson Charles M. Rosenthal, First Manhattan Company Alfred M. Zeien Walter J. Salmon, Harvard Business School James A. Sharp, Carl Zeiss Microscopy, LLC Dyann F. Wirth, Harvard Global Health Institute

All lists as of December 31, 2012.

58 MBL Annual REport 2012 MBL LEadership

BOARD OF OVERSEERS Saul Kurlat, Cambridge, MA ASSOCIATES EXECUTIVE BOARD Donald R. Aukamp, Woods Hole, MA Deena G. Laties, Philadelphia, PA Thomas F. Gregg, President David Bakalar, Chestnut Hill, MA Anna L. Lawson, Daleville, VA Huson R. Gregory, Vice President Richard M. Barno, Weston, MA Joseph S. Levine, Concord, MA Michael Fenlon, Secretary Timothy A. Barrows, Matrix Partners Richard Lipkin, Commerce Health Ventures Robert C. King, Treasurer John F. Batter, WilmerHale William W. Locke, Cataumet, MA Alice M. Chappell* (July, 2013) George Berkowitz, Legal Sea Foods William K. Mackey, Mackey & Foster, PA Syrel Dawson Joshua B. Bernstein, Bernstein Management Corporation Dale Mathias, Washington, DC Joe A. DeGiorgis George H. Billings, Falmouth, MA Thomas McMurray, Jackson, WY Michael J. Fishbein Marianne Billings, Potomac, MD Nawrie Meigs-Brown, Woods Hole, MA Erich F. Horgan Margaret C. Bowles, Lyme, NH Ambrose K. Monell, G. Unger Vetlesen Foundation Barbara W. Jones Ed Brakeman, Bain Capital LLC H. Edward Muendel, Stanton Chase International Hans Kornberg Beth Colt, Woods Hole Inn Edward P. Owens, Norwell, MA Ruth Ann Laster Bernice Cramer, Newton, MA Jean W. Pierce, Boca Grande, FL Jeffrey O. Oppenheim Catherine B. Cramer, New York, NY Robert W. Pierce, Pierce Aluminum Co. Marjorie B. Salmon Thomas S. Crane, Mintz, Levin, Cohn, Ferris, Glovsky, John Ripple, Virdante Pharmaceuticals & Popeo, PC Robert Samuelson, Monitor Group LLC ex-officio members Sally Cross, Falmouth, MA Cecily C. Selby, New York, NY John W. Rowe, Chair of the Board of Trustees, MBL Diarmaid Douglas-Hamilton, Hamilton Thorne Inc. Bonnie W. Simon, Washington, DC Joan V. Ruderman, President amd Director, MBL Joseph L. Dowling, The Rhode Island Eye Institute Carolyn Stimpson, Wachusett Mountain Pamela Clapp Hinkle, Director of Development and Richard Ellis, New York, NY Ann E. Sutherland, University of Virginia External Relations, MBL Pamela L. Follett, Lewis Rhodes Lab Gerard L. Swope, Woods Hole, MA Jean-François Formela, Atlas Ventures John F. Swope, Sheehan, Phinney, Bass and Green Craig B. Gibson, Lawrence General Hospital Karen Tierney, Wellesley, MA Prosser Gifford, Woods Hole, MA Victor K. Tolkan, Penzance Management LLC Catherine M. Golden, New York, NY Gerald Weissmann, New York University Medical Center Mary Greer, Cambridge, MA Nancy G. Wolf, University Hospitals of Cleveland Gary Jacobson, Westwood Pembroke Health System Barbara Wu, Wilmette, IL Thomas P. Jalkut, Nutter McClennen & Fish LLP James H. Wyche, Howard University Mary D. Janney, Washington, DC* (October, 2012) William Zammer, Cape Cod Restaurants Barbara W. Jones, Barbara Woll Jones Designer/Builder Ronald Zweig, Woods Hole, MA

MBL Annual REport 2012 59 MBL LEadership

SCIENTIFIC LEADERSHIP

The Josephine Bay Paul Center for Comparative Whitman Center Molecular Biology and Evolution Robert Goldman, Director Mitchell Sogin, Director, Senior Scientist Mark Alliegro, Senior Scientist Cellular Dynamics Program Gary Borisy, Senior Scientist Joshua Hamilton, Acting Director Joshua W. Hamilton, Senior Scientist Shinya Inoué, Distinguished Scientist Anton Post, Senior Scientist Osamu Shimomura, Distinguished Scientist David Mark Welch, Associate Scientist Gary Borisy, Senior Scientist Linda Amaral Zettler, Assistant Scientist Jonathan Gitlin, Senior Scientist Irina Arkhipova, Assistant Scientist Rudolf Oldenbourg, Senior Scientist Julie Huber, Assistant Scientist Michael Shribak, Associate Scientist Sheri Simmons, Assistant Scientist Tomomi Tani, Associate Scientist

The Ecosystems Center Program in Sensory Physiology and Behavior Hugh Ducklow, Director**, Senior Scientist Roger Hanlon, Senior Scientist Christopher Neill, Director**, Senior Scientist Stephen Highstein, Senior Scientist John Hobbie, Distinguished Scientist Alan Kuzirian, Associate Scientist Jerry Melillo, Distinguished Scientist Zoe Cardon, Senior Scientist Center for Library and Informatics Linda Deegan, Senior Scientist Nathan Wilson, Director, Associate Research Scientist Anne Giblin, Senior Scientist David Patterson, Senior Scientist Edward Rastetter, Senior Scientist Gaius Shaver, Senior Scientist SENIOR STAFF Joseph Vallino, Associate Scientist Gary G. Borisy, President and Director** Jianwu Tang, Assistant Scientist Richard Cutler, Director Facilities, Services and Projects Susan Goux, Director of Human Resources Eugene Bell Center for Regenerative Biology and Joshua Hamilton, Chief Academic and Scientific Officer Tissue Engineering Pamela Clapp Hinkle, Director of Development and Gary G. Borisy, Acting Director External Relations Marko Horb, Associate Scientist Maureen Nunez, Chief Financial Officer** Jennifer Morgan, Assistant Scientist** William Reznikoff, Director of Education Joel Smith, Assistant Scientist Joan Ruderman, President and Director** Mark Messerli, Assistant Scientist

*deceased **Staff who joined or left the MBL during 2012.

60 MBL Annual REport 2012 PHOTOS

Cover and Table of Contents: The green alga Chlamydomonas reinhardtii, which has Channelrhodopsin proteins important in optogenetic research. (Laura Hilton, courtesy of Lynne Quarmby, Simon Fraser University) Page 3: Nerves in red can be easily traced among the distinctive chromatophores and iridophores that they innervate. (Wardill, Gonzalez- Bellido, Crook & Hanlon, Proceedings of the Royal Society B: Biological Sciences) Page 4: Sampling fluids for microbiology and geochemistry at Von Damm Vent field. (Chris German/Woods Hole Oceanographic Institution); Joan Ruderman (Bachrach Studios) Page 5: Longfin squid (Tom Kleindinst); After eight years of nutrient enrichment, the edge of this salt marsh at Plum Island Estuary has fragmented and turf has slumped into the channel. (Christopher Neill) Page 6: Cross-section of a lamprey spinal cord stained with toluidine blue. (Paul Oliphint, Jennifer Morgan); Chris Neill (Tom Kleindinst); Fluorescent staining with each square shows where different genes are turned on in acorn worm embryos. Multiple colors in a single image show the activities of different genes in the same embryo.(A. Pani) Page 7: Physiology course (Tom Kleindinst); Confocal imaging of squid skin reveals relative morphology of chromatophores with their radial muscles (large brown chromatophore in center) and an underlying iridescence iridophore (Paloma Gonzalez-Bellido and Trevor Wardill); Brown-MBL students at Plum Island LTER. (David Rand) Page 8: Water faucet (dreamstime.com); A single hair cell from a frog ear (J. Meyers) Page 9: Intestinal section from a gnotobiotic mouse model inoculated with selected bacterial species found in the human gut. (Yuko Hasegawa/MBL); The waters of the Antarctic Peninsula, one of several sites where water samples were collected for MBL studies of bacterial distributions (Hugh Ducklow) Page 10: Tree frog (Encyclopedia of Life); L to R Semester in Environmental Science alumni Hansen Johnson, Fiona Jevon, and Andrew Miano climb Alaska’s Brooks Range while on a break from their work as summer research interns at the Toolik Lake Field Station. (MBL) Page 11: The diploma for T.H. Morgan’s 1933 Nobel Prize in Physiology or Medicine, which is now in the MBL Archives. (Diana Kenney) Page 13: Woods Hole village (Woods Hole Oceanographic Institution) Page 14: Coronal section of an entire mouse brain, stained to delineate anatomical boundaries in many brain regions. (Allen Institute for Brain Science) Page 15: MBL on Eel Pond (Tom Kleindinst) Page 16: Jeffrey Pierce (Tom Kleindinst) Page 36: Pycnopodia helianthoides (Casey Debehhan/Census of Marine Life) Page 57: Daffodils on Lillie front lawn (Pam Wilmot) Page 62: MBL student (Tom Kleindinst)

MBL Annual REport 2012 61 The Marine Biological Laboratory does not discriminate in employment or in access to any of its activities or programs or take any retaliatory action on the basis of race, color, religion, sex, sexual orientation, national origin, ancestry, age, participation in a discrimination-related complaint, physical or mental disability, military status or genetic predisposition. In addition, the MBL is committed to the prevention and elimination of sexual harassment, as well as other forms of unlawful harassment, in the workplace. Through training programs and disseminated information, MBL strives to educate its employees, students, faculty, and visitors on these important issues.

62 MBL Annual REport 2012 MBL Annual REport 2012 63 Non Profit ORG. U.S. Postage PAID Brockton, MA Permit No. 402 Marine Biological Laboratory 7 MBL Street Woods Hole, MA 02543 www.mbl.edu

64 MBL Annual REport 2012 Printed with recycled materials