SALAMANDRA 53(2) 267–278 15 May 2017 StatusISSN of Rhinella0036–3375 beebei Toads, tall mountains and taxonomy: the Rhinella granulosa group (Amphibia: Anura: Bufonidae) on both sides of the Andes John C. Murphy1,7, Teddy Angarita Sierra2,3, J. Roger Downie4 & Michael J. Jowers5,6 1) Science and Education, Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, USA 2) Yoluka ONG, Fundación de Investigación Biodiversidad y Conservación, Bogotá, Colombia 3) Grupo de investigación Biogeografía Histórica y Cladística Profunda, Laboratorio de anfibios, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia 4) School of Life Sciences, Graham Kerr Building, University of Glasgow, Glasgow G12 8QQ, Scotland, UK 5) CIBIO/InBIO (Centro de Investigação em Biodiversidade e Recursos Genéticos), Universidade do Porto, Campus Agrario De Vairão, 4485-661, Vairão, Portugal 6) National Institute of Ecology, 1210, Geumgang-ro, Maseo-myeon, Seocheon-gun, Chungcheongnam-do, 33657, Korea Corresponding authors: Michael J. Jowers, e-mail:
[email protected],
[email protected] Manuscript received: 3 December 2015 Accepted: 19 April 2016 by Stefan Lötters Abstract. A toad in the Rhinella granulosa group has been recognized as present on Trinidad since 1933. In 1965, the Trini- dadian population was described as a subspecies of Bufo granulosus, B. g. beebei. It has its type locality on the island and was eventually raised to species status as B. beebei (Beebe’s toad). Recently Beebe’s toad was synonymized with Rhinella humboldti, a species with a type locality in the Magdalena Valley of western Colombia. The Magdalena Valley is separated from the Orinoco Basin by the Eastern and Merida Cordilleras.