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A Break-Up of the Australian Gecko Genus Strophurus

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A Break-Up of the Australian Gecko Genus Strophurus 36 Australasian Journal of Herpetology Australasian Journal of Herpetology 34:36-56. ISSN 1836-5698 (Print) Published 20 July 2017. ISSN 1836-5779 (Online) A break-up of the Australian gecko genus Strophurus Fitzinger, 1843 sensu lato as currently recognized, from one to four genera, with two new subgenera defined, description of nine new species and two new subspecies. RAYMOND T. HOSER 488 Park Road, Park Orchards, Victoria, 3134, Australia. Phone: +61 3 9812 3322 Fax: 9812 3355 E-mail: snakeman (at) snakeman.com.au Received 15 January 2017, Accepted 20 May 2017, Published 20 July 2017. ABSTRACT The genus Strophurus Fitzinger, 1843 sensu lato has been the subject of numerous taxonomic reviews in recent years. With the exception of Wells and Wellington (1985), no recently publishing herpetologists have broken up the genus beyond that defined by Cogger (2014), which sums up the current position in Australian herpetology. Recognizing recent molecular work on the assemblage (e.g. Sadlier, Omeally and Shea (2005) or Nielsen et al. 2016), Strophurus is herein divided into four obvious and divergent genera. One is formally named for the first time. One of these genera is subdivided three ways into subgenera, two being named for the first time. In spite of four species being named in the past three years (two as subspecies, but elevated herein to species status on the basis of time of divergence), molecular evidence clearly shows numerous unnamed forms. To partially correct this situation, nine new easily diagnosed species and two new subspecies are formally named for the first time. A complete list of recognized and valid species, confirmed by published molecular data is also provided. Keywords: Taxonomy; lizards; Australia; Western Australia; Northern Territory; Queensland; Gecko; Strophurus; Eremiastrophrurus; Oedurella; new genus; Adelyndactylus; new subgenus; Graciledactylus; Parvusdactylus; new species; jackyae; dannybrowni; gedyei; chriswilliamsi; jenandersonae; alba; jamielindi; garystephensoni; sonnemanni; new subspecies; obscurum; minima. INTRODUCTION separate paper published at the same time as this (Hoser, The genus Strophurus Fitzinger, 1843 more recently 2017a) and is therefore ignored for the purposes of this applied to the so-called spiny tailed geckos, was effectively paper. ignored by most herpetologists until resurrected by Wells In terms of Strophurus sensu lato, while all species are and Wellington in their major work of 1985 (Wells and united by caudal mucous glands and associated ejection Wellington, 1985). mechanisms, and transversely enlarged (as opposed to Since then the genus has been widely recognized in rounded and paired) proximal subdigital lamellae, the Australian herpetology as being separate from other individual species form distinct morphologically distinct species within the genus Oedura Gray, 1842 sensu lato groups. with this group (Strophurus) including all the species with On this basis, Wells and Wellington (1985) resurrected the caudal mucous glands and associated ejection genus Oedurella Lönnberg and Andersson, 1913, for the mechanisms, and transversely enlarged (as opposed to divergent and small so-called “phasmid gecko”, originally rounded and paired) proximal subdigital lamellae. described as “Oedurella taeniata Lönnberg and Andersson, The rest of Oedura Gray, 1842 sensu lato is dealt with in a 1913”, and also created a new genus, Eremiastrophrurus Hoser 2017 - Australasian Journal of Herpetology 34:36-56. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 37 Wells and Wellington, 1985, for the divergent species coincided with an audit of Strophurus sensu lato (as described as “Diplodactylus elderi Stirling and Zeitz, 1893”. defined by Cogger (2014) and similar texts) by myself, with At the same time, they divided the two main populations of a view to dividing the genus along obvious morphological the species “Diplodactylus elderi” as then recognized into and phylogenetic lines. two. At the same time I audited the known species within the In spite of overwhelming evidence accumulating in favour group to see if there were any unrecognized groups in of the Wells and Wellington actions in the three decades need of either being formally recognized and/or formally since their publication of 1985, spearheaded by a gang of named for the first time. thieves known as the Wüster gang as described by Hoser Long aware of the illegal suppression of the works of Wells (2015a-f), not one single publishing herpetologist has used and Wellington by the Wüster gang and long aware of the either the Wells and Wellington genus name or recognized fact that valid and appropriate Wells and Wellington names the species Eremiastrophrurus manhoodi as originally for taxa were not being used, this paper already had a described by them. reason for publication on that basis alone. This attests to the power of threats of violence and other Also having spent more than 30 years active across forms of personal harm made on herpetologists by Australia catching and observing relevant species, I was members of the Wüster gang over the three decades since well aware that the recognized species diversity of the publication of Wells and Wellington (1985) and the group as listed in Cogger (2014) or the Wüster gang immense damage done to the science of herpetology by controlled “The Reptile Database” managed by Peter Uetz, this gang. listing just 19 species as at end 2016, was an In terms of overwhelming evidence in favour of the understatement of the total, so I decided to identify and taxonomy and nomenclature of Wells and Wellington with name obvious unnamed taxa, which is done in this paper. regards to the genus Eremiastrophrurus, or the formal MATERIALS AND METHODS recognition of the species Eremiastrophrurus manhoodi, The identification of the relevant genus and species groups one need look no further than the publication of Nielsen et was easily achieved by simple inspection of relevant al. (2016) at Fig. 1, page 486, which shows a significant specimens, live in the field, in museums and via images divergence at both genus and species levels for the sent to me by others with accurate locaility and/or other relevant taxa. Alternatively, one may look at the very data. In terms of species level groups, biological barriers animals themselves, or for those who prefer not to leave were identified by combining known locality data with the comfort of their own armchair, merely take a look at known geographical barriers, most of which have become page 355 of Brown (2014) and one sees pictures of both well known to myself in my various researches on other taxa, side by side at the bottom of the page reptile groups inhabiting the same regions. (Eremiastrophrurus elderi on the left and The formal naming exercises are a direct result of a review Eremiastrophrurus manhoodi to the right), with all their of the relevant literature to identify all previously named obvious differences, including those identified and spelt out groups at both species and genus level, including known by Wells and Wellington in 1985. synonyms and potentially available names according to the On this basis, the very valid and properly identified Wells rules of the International Code of Zoological Nomenclature and Wellington taxa are herein formally recognized and (Ride et al. 1999). their nomenclaturally available names are used in I should mention that any names coined in online non peer accordance with the rules of the International Code of reviewed or PRINO (peer reviewed in name only) journals Zoological Nomenclature (Ride et al. 1999). like Zootaxa are usually available under the current and Illegal names for these or other taxa coined by the Wüster relevant rules of the International Code of Zoological gang in online PRINO (peer reviewed in name only) Nomenclature as amended online and so are treated as journals they have hijacked, such as Zootaxa, should be valid and used when appropriate herein. ignored and not used under any circumstance. This assumes that the names are not junior synonyms of However, I recognize the ongoing threats and harassment earlier properly proposed names. of scientists who go against the terrorist like edicts and Available names are used as appropriate (in the paper demands of Wolfgang Wüster and his associated band of below) and where none was available the relevant entities thugs and thieves. are named according to the provisions of the International It is also a matter of public record that illegal actions of the Code of Zoological Nomenclature. Wüster gang and associates have caused deaths of a While the species, genera and subgenera diagnosed number of very competent herpetologists, including for herein are done so on the basis of their own physical example Luke Yeomans in the UK and Nathan Garrod in characters, it is important to note the guidance given by Queensland, Australia. relevant earlier publications (quoted herein), which in For defying the demands of the Wüster gang, myself and combination show that the taxonomic conclusions within my family have been subject of criminal attacks to both our this paper are not only logical, but are in fact a mere persons and property, as have many of my co-workers and statement of the obvious. it is the fear of such occurring that has been forcing many Divergence times of species or genus level groups are competent herpetologists to bow down to the illegal taken from the published literature as cited herein. demands of the Wüster gang and using the illegal How long it will take other herpetologists to adopt and use nomenclature (coined in breach of the International Code the taxonomy within this paper will not depend on the of Zoological Nomenclature) instead of the legal merits of what is published herein, so much as how willing nomenclature. they are to brave the hatred and harassment from a group The publication of the paper by Nielsen et al. (2016) known as the Wüster gang, who will seek to do all they can Hoser 2017 - Australasian Journal of Herpetology 34:36-56.
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