Lower Liassic Brachiopods from the Steinplatte-Kammerköhralm Area Near Waidring (Northern Calcareous Alps, Salzburg) 965-982 ©Geol

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Autor(en)/Author(s): Siblik M.

Artikel/Article: Lower Liassic Brachiopods from the Steinplatte-Kammerköhralm Area near Waidring (Northern Calcareous Alps, Salzburg) 965-982 ©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at

Gedenkband zum 100. Todestag von Dionys Stur Redaktion: Harald Lobitzer & Albert Daurer

Jb. Geol. B.-A. ISSN 0016-7800 Band 136 Heft 4 S.965-982 Wien, Dezember 1993

Lower Liassic Brachiopods from the Steinplatte-Kammerköhralm Area near Waidring (Northern Calcareous Alps, Salzburg)

By MilOS SIBLiK*)

With 10 Text-Figures and 2 Plates

Salzburg Nördliche Kalkalpen Steinplalle Lias Osterreichische Kartei: 50.000 Brachiopoden Bläller 91,92 Taxonomie

Contents

Zusammenfassung 965 Abstract...... 965 1. Introduction 965 2. Descriptions 966 Acknowledgements 977 References ...... 982

Brachiopoden aus dem Unterlias des Gebietes Steinplatte - Kammerköhralm bei Waidring (Nördliche Kalkalpen, Salzburg) Zusammenfassung 19 Brachiopoden-Taxa aus dem Unterlias (Marmorea-Zone) werden von drei Lokalitäten der Steinplatte beschrieben.

Abstract The Lower Liassic (Marmorea Zone) of the Steinplatte area has yielded 19 brachiopod taxa that are studied in the present paper.

1. Introduction

The present paper deals with the Lower Liassic brachio- a Locality ST-1 pod fauna coming from the Kammerköhralpe, NE of Stein- leilleria pertorata (PIETIE). platte (1896 m) near Waidring, Salzburg. The study is a Locality ST-2 based on my own collection made in 1991 and completed Calcirhynchia (?) plicatissima (QUENST.), Cirpa planitrans (OR- by several specimens found there by Dr. H. LOBITZERand MaS), Prionorhynchia (?) polyptycha (OPPEL), Furcirhynchia Dr. M. RAKUSin 1990. emmrichi (OPPEL), Cuneirhynchia retusitrons (OPPEL), "Rhyn- The Liassic of the Steinplatte area was studied in detail choella" traasi OPPEL, "Rh." aft. prana OPPEL, Liospiriterina al- by HAHN (1910) who mentioned 3 brachiopod species pina (OPPEL), L. obtusa (OPPEL), Liospiriferina sp., Lobothyris from the Lower Liassic variegated limestone ("Bunte ovatissimaetormis (BÖCKH), Linguithyris aspasia (MENEGH.), Ammonitenkalke") of Kammerköhralpe: Rhynchonella cf. leilleria pertorata (PIETIE), Z. mutabilis (OPPEL), Z. alpina traasi OPPEL, Rhynchonella plicatissima QUENSTEDTand Pygope (GEYER),Z. stapia (OPPEL), Z. aft. apenninica (ZITI.). nimbata OPPEL. a Locality ST-3 The new material described in the present paper comes Calcirhynchia (?) plicatissima (QUENST.), "Rhynchonella" aft. from 3 localities (for the localities see Text-Fig. 1), where paolii CAN., "Rhynchonella" sp., Linguithyris aspasia (ME- the following species were found: NEGH.).

*) Author's address: Dr. MILOSSIBLIK, Geologicallnstitute, Czech Academy of Science, Rozvojova 135, (;Z-16500 Praha 6, Czech Republic.

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/ I )

The accompanying ammonite fauna determined by M. 1943 Rhynchonella plicatissima Qu. - VIGH, p. 339, PI. 26, RAKUS(1993, this volume) indicates the Marmorea Zone. The Figs.11-16. 7 1957 Rhynchonella plicatissima QUENSTEDT - MAHEL, p. 182, preservation of brachiopods is relatively good, some spe- PI. 8, Figs. 13-14. cimens are fragmentary or unsuitable for sectioning due to 1970 "Rhynchonella" plicatissima (QUENSTEDT)- GAETANI, p. 381, the recrystallization of the inner parts of shell. If compared Text-Fig. 9. to the younger Hierlatz brachiopods, the Steinplatte bra- non 1954 Rhynchonella plicatissima QUENSTEDT- CONTI, p. 197, PI. 9, chiopod fauna seems to be slightly different, lacking in Fig. 12 (= Calcirhynchia rectemarginata (VECCHIA) fide GAE- numerous Terebratulidae and strongly ribbed rhynchonel- TANI, 1970). Mat e ria I: 11 specimens preserved as partly decorti- lids of the "alberti" and "guembeli" types. cated and fragmentary internal moulds. The dimensions All figured specimens are deposited in the collection of (in mm) of the better preserved specimens are as fol- the Geologische Bundesanstalt (Museum) in Vienna. lows:

Length Width Thickness 15.2 17.3 10.8 714.5 716.0 10.2 2. Descriptions 14.0 16.5 9.0 sectioned 713.8 16.0 7.8 PI. 1, Fig. 4 13.7 14.9 9.3 Order: Rhynchonellida KUHN, 1949 12.7 7 8.1 Superfamily: Rhynchonellacea GRAY, 1848 11.0 13.6 7.0 sectioned Family: Wellerellidae lIKHAREv In t ern a Ich a r act e r s: Lateral umbonal cavities sub- in RZHONSNITSKAYA, 1956 trigonal in cross-section. Dental lamellae strong and Genus: Calcirhynchia BUCKMAN, 1917 long, slightly divergent dorsally and anteriorly. Pedicle collar distinguished in 1 specimen sectioned. Double deltidial plates ascertainable. Hinge teeth expanded Calcirhynchia (?) plicatissima (QUENSTEDT, 1852) dorsally, crenulated and inserted into large, crenulated (PI. 1, Fig. 4; Text-Fig. 2) sockets. Squat denticula situated laterally. Hinge pla- 1852 Terebratulaplicatissima- QUENSTEDT,p. 451, PI. 36, Fig. 3. tes subhorizontal, at the distal ends with dorsally deve- 1889 Rhynchonella plicatissima QUENST. - GEYER, p. 57, PI. 6, loped crural bases. Neither true median septum nor Figs. 33-36, PI. 7, Figs. 1-7 (cum syn.). septalium observed. Crura prefalciform, extending 71889 Rhynchonella latifrons STUR m.s. - GEYER, p. 54, PI. 6, Fig. 29 slightly into pedicle valve. only. Rem ark s: This species and its allies belong undoubted- 1893 Rhynchonella plicatissima, QUENST. - FUCINI, p. 295, PI. 4, Figs.1-2. ly to the most difficult of all rhynchonellids which occur 1909 Rhynchonella plicatissima QUENST.sp. - TRAUTH, p. 64, PI. 2, in the Lower Liassic. There exists a series of small to Fig. 1. medium-sized, highly variable forms for which a great 1926 Rhynchonella plicatissima, Qu. - PETERHANS, p. 363, PI. 2, number of available names was used. Moreover, indi- Figs.5-8. stinct figures and insufficient original descriptions ren- 1932 Rhynchonella plicatissima QUENSTEDT - RENZ, p. 55 (cum syn.). der the taxonomy rather difficult. Our specimens agree 1936 Rhynchonella plicatissima FA QUENSTEDT- JOLY, p. 151 (cum very well externally with those figured and described in syn.). detail by GEYER(1889). The only difference in external 1937 Rhynchonella plicatissima QUENST.- ORMOS, p. 32. features is that the Hierlatz forms have, on the average,

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... ~ 0.3 0.5 0.6 \ 0.75 \ 2 mm, 0.9 1.1

1.2

» 4 2.7 f ~ 2.8 f ~ 3.2 ( "\ 3.7

( ) 3.95

I 4.1

Text-Fig. 2. Calcirhynchia (?) plicalissima (QuENSTEDT). Serial transverse sections through the posterior part of shell. Originallength of specimen 14.0 mm. ST 2. Magnified.

slightly more convex pedicle valves. The species is un- Genus: Cirpa 01 GREGORIO, 1930 doubtedly very close to "Rhynchonella" latifrons GEYER, 1889 that was distinguished chiefly by the more promin- Cirpa planifrons (ORMOS, 1937) ent beak and by the elliptic outline in anterior view. ÄGER (PI. 1, Figs.1-3; Text-Fig. 3) (1962, p. 86) with doubt referred "plicatissima" to Calcir- hynchia and mentioned close relation of serial sections of 1937 Rhynchonella planifrons nov.sp. - aRMOS, p. 35, 41, PI. 1, Calcirhynchia to those of Cirpa. GAETANI'Ssections (1970, Fig.19. ? 1964 Rhynchonella rimala OPP. - SACCHI VIALL!, p. 13, PI. 2, Fig. 7. Text-Fig. 9) of a topotypical specimen of "plicatissima" from Ofterdingen in Germany that revealed long dorsal Mat e ria I: 5 internal moulds with fragments of shell. The septum throw doubt on the attribution of "plicatissima" to figured specimens have the following dimensions: 13.7 Calcirhynchia. In the present paper, the question mark is x 15.6 x 9.9 mm (PI. 1, Fig. 1), ?11.5 x 13.0 x 8.8 mm used in connection with the generic name Calcirhynchia as (PI. 1, Fig. 2) and 11.4 x 12.4 x 8.8 mm (PI. 1, Fig. 3). the status of Calcirhynchia has still not been resolved sa- In tern a Ich a ra c t e rs: Could not be completely ascer- tisfactorily and there have not been gained more recent tained owing to the partial recrystallization of both spe- data on the internal structures of "plicatissima" and other cimens sectioned, but serial sections revealed the in- allocated species. ternal characters typical of Cirpa: pedicle collar, double Dis tri but ion: Stein platte - loc. 2 and 3 (Hierlatz, Bar- deltidial plates, short dental lamellae, fused hinge barastollen in Hinterholz, Viis, Germany, Switzerland, plates subhorizontally situated, neither septalium nor Belgium, Italy, Greece, CSFR, Hungary). median septum developed, and raduliform crura. Upper Hettangian to Upper Sinemurian-Lotharingian Rem ark s : The specimens display externally the charac- (according to ÄLMERAS, 1964). ?Middle Liassic of teristic features of Cirpa: rectangular outline of shell in Greece. anterior view and flattening of its anterior end. Number

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Text-Fig.3. Cirpa planifrons (ORMOS). Three transverse sections showing subparallel hinge plates and absent median septum. Original length of specimen c: 11.0 mm. ST 2. Magnified. (

J of the ribs confined to the fold is 4 (in 1 specimen)

or 5 (in 4 specimens). I This species is unoubt- I edly very close to Cirpa {ronlo (QUENSTEDT, 1871) which can be distinguished by its stronger ribs and by dorsally divergent dentallamellae. Lateral umbonal ca- producing common cynocephalous variants. vities subtrigonal in cross-section. Pedicle collar not Dis tri but ion: ORMOS (1937) recorded her new species ascertained. Hinge teeth strong, expanded dorsally and as coming from the Oxynoliceras oxynolum horizon of the crenulated. Stout denticula developed laterally. Sockets Lower Liassic succession in the Northern Bakony. Our large, crenulated, with better developed outer socket specimens derive from Stein platte -Ioc. 2. ridges. Hinge plates narrow, clearly demarcated from the inner socket ridges. Septalium narrow and deep. Median septum splitting early away from septalial plates. Crura raduliform but somewhat flattened and Genus: Prionorhynchia BUCKMAN, 1917 blade-like anteriorly, curving into cavity of pedicle valve. Prionorhynchia(?) polyptycha (OPPEL, 1861) Rem ark s: Our specimens have 5-9 sharp costae anteriorly on the brachial valves (3 or 5 in the fold), and 1861 Rhynchonella polyplycha OPP.- OPPEL,p. 544, PI. 12, Fig. 4. 1889 Rhynchonella polyplycha OPP.- GEYER,p. 51, PI. 6, Figs. 15-17. 40-50 fine capillae posteriorly. The representatives of 1911 Rhynchonella Greppini OPP.m.f. polyptycha OPP.- HAHN,p. 559, this genus are very distinctive, but only local and rela- PI. 21, Fig. 1. tively uncommon in the Alpine Liassic. Nearly all Liassic 1937 Rhynchonella polyptycha OPP.- ORMOS,p. 29. Furcirhynchia forms showing combination of costae and Mat e ria I: 3 fragmentary specimens. The best one capillae have been recorded as Rhynchonella {urcillala in the measures ?14.5 x 15.3 X8.8 mm. older literature. The authorship of this species has been attributed to THEODORIbut the name was first published Rem ark s: The material closely resembles the speci- by VON BUCH (1835, p. 43). His original material derived mens figured by OPPEL (1861) and GEYER (1889) from Hierlatz and shows the only difference - straight anterior from the Upper Liassic (see also AGER, 1958, p. 72). OP- PEL'S (1861) chief criterion for his new Lower Liassic margin (in the dorsal view). The specimen figured by species "Emmrichi" lay in the finer capillation of the pos- HAHN (1911, PI. 21, esp. Fig. 1c) is well comparable to terior parts of valves. This did not seem to be sufficient the Stein platte material. The species itself could be dis- ground for separating this form from and tinguished from the similar "Rhynchonella" {raasi OPPEL by "{urcillala" the lower and wider plication and by weaker ribs. Any- "emmrichi" has been usually included into the synonymy way, "Rhynchonella" {raasi and "Rhynchonella" greppini OPPEL of "{urcillata". The present author would, however, favour the separating of both species on the grounds that may produce variants that can be distinguished from the seems to have a more pronounced and upright species under consideration only with difficulties. "emmrichi" beak, and a slightly depressed outline in lateral view, Dis tri but ion: Stein platte - loc. 2 (Hierlatz, Achensee- with the two valves surfaces almost parallel. The strati- hof, Germany, CSFR, Hungary, ?Bulgaria). graphical difference between both species is also considered significant. Dis tri but ion: Stein platte - loc. 2 (Hierlatz). Sinemur- ian. Family: Rhynchone/lidae GRAY, 1848 Genus: Furcirhynchia BUCKMAN, 1917 Genus: Cuneirhynchia BUCKMAN, 1917

Furcirhynchia emmrichi (OPPEL, 1861) Cuneirhynchia retusifrons (OPPEL, 1861) (PI. 1, Fig. 9; Text-Fig. 4) (PI. 1, Fig. 10) 1861 Rhynchonella EmmrichiOpP. - OPPEL,p. 542, PI. 12, Fig. 1. 1861 Rhynchonella retusifrons OPP.- OPPEL,p. 544, PI. 12, Fig. 5. 1889 Rhynchonella furcillata THEOD.- GEYER,p. 60, PI. 7, Figs.? 1889 Rhynchonella retusifrons OPP.- GEYER,p. 62, PI. 7, Figs. 8-12. 16,17. 1932 Rhynchonella retusifrons OPP.- RENZ,p. 51 (cum syn.). 1936 Rhynchonella retusifrons A. OPPEL- JOLY,p. 152. Mat e ria I: 2 specimens with both valves and 1 incom- 1957 Rhynchonella retusifrons OPPEL- MAHEL,p. 183, PI. 8, Fig. 12. plete brachial valve. Dimensions: 20.4 x26.4 x 14.3 mm 1966 Cuneirhynchia retusifrons (OPPEL)- PEVNY, p. 273, PI. 11, (figured) and ?16.0 x21.5 x 10.8 mm (sectioned). Fig.2. In t ern a Ich a ra c t e r s: Delthyrial cavity characteristic- Mat e ria I: 2 partly damaged specimens, the figured one ally quadrate in cross-section, between subparallel or with dimensions ?8.2 x 9.5 x 6.8 mm.

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CJ'9

Text-Fig. 4. Furcirhynchia emmrichi (OPPEL). Measured from dorsal umbo. Crura persisted to 3.6 mm. Originallength ?16.0 mm. ST 2. Magnified.

Rem ark s: The specimens correspond well to the OP- Rem ark s: The specimen agrees very well externally with PEL'S original figures and description and to the that figured by OPPEL, though the umbones cannot be thorough description of GEYER(1889), and differ only by compared since it has been damaged in our specimen. their more inflated posterior parts of pedicle valves. The planareas are shallow, not sharply limited. Branch- "Rhynchonella" cartieri OPPEL, 1861 was thought by GEYER ing of the ribs occurring according to GEYERin the pos- to be closely related to "retusitrons" but it could be dif- terior parts of the valves, is well recognizable in the ferentiated by its relatively thicker shape and by its Stein platte specimen. On the contrary, the uniting of stronger and longer ribs. The type species of Cuneirhynch- two faint posterior ribblets into one anterior rib that was ia - Middle Liassic C. da/masi (DUMORTIER)- may develop observed occasionally in the Hierlatz material, is not specimens similar to "retusitrons" but these differ, present in the Steinplatte specimen. Such uniting oc- however, from "retusifrons" in having a nearly flat pedicle curs, however, in one (young) specimen from Steinplatte valve. (PI. 1, Fig. 8) that is included, with some hesitation, into Dis tri but ion: Steinplatte, loc. 2 (Hierlatz, Germany - the species under consideration. The catch-all genus Hindelang, Italy, Belgium, Hungary, CSFR). Rhynchonella is used here as practically nothing is known Hettangian to Upper Sinemurian (Lotharingian). Middle about the internal characters of "traasi" (except for three Liassic of Greece. not much telling sections through the shell publiShed by GEYER,1889). Dis tri but ion: Steinplatte - loc. 2 (Hierlatz, Germany - Genus: Rhynchonella FISCHER, 1809; s.1. Hindelang, CSFR, Hungary, Italy). Sinemurian-Domerian (according to ALMERAS,1964). "Rhynchonella" 'raasi OPPEL, 1861 (PI. 1, Figs. 7-?8) 1861 Rhynchonella FraasiOpP. - OPPEL, p. 543, PI. 12, Fig. 3. "Rhynchonella" aft. prona OPPEL, 1861 1889 p. 52, PI. 6, Figs. 18-24. RhynchonellaFraasiOpP.-GEYER, (PI. 1, Fig. 5) 1893 Rhynchonella Fraasi OPPEL- BÖSE, p. 643. ? 1907 Rhynchonella Fraasi, OPP. - DAl PIAZ, p. 31, PI. 2, Fig. 7. aff. 1861 Rhynchonella prona OPP.- OPPEL, p. 547, PI. 13, Fig. 7. 1937 Rhynchonella fraasi OPPEL - ORMOS, p. 30. 1886 Rhynchonella prona OPPEL - ROTHPLETZ, p. 37, PI. 12, 1956 RhynconellafraasiOpPEL-SELLI, p. 9, PI. 1, Fig. 5. Fig.35. 1964 "Rhynchonella" fraasiOpPEL - PEVNY, p. 164. aff. 1889 Rhynchonella prona OPPEL - GEYER, p. 68, PI. 7, Figs. 24-25. Mat e ria I: One incomplete internal mould with dimen- 1911 Rhynchonella sp.aff. prona OPP. - HAHN, p. 547, PI. 20, sions ?11.0 x 12.0 x 8.4 mm and another 2 fragments Fig.5. probably belonging here. 1966 Rhynchonella prona OPPEL - PEVNY, p. 277.

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Mat e ria I: 1 slightly damaged specimen with the dimen- ing a flatter pedicle valve. According to AGER(1967) Squa- sions ?12.5 x 15.1 x 8.1 mm. mirhynchia is a rare genus and "squamiplex" has not been Rem ark s: General shape and type of ribbing of the spe- reported from the Alpine Liassic with certainty so far. cimen from Stein platte remind one of some West Euro- Only with a further material enabling also the study of pean species, e.g. Stolmorhynchia bouchardi (DAVIDSON, internal characters, it would be possible to make a spe- 1852), Piarorhynchia ex gr. juvenis (QuENSTEDT,1852) etc. It cific determination of this specimen under considera- could be, however, distinguished on the grounds of its tion. weaker convexity of the brachial valve, by the lower Dis tri but ion: Steinplatte -Ioc. 3. plication and by the different outline, respectively. On the other side, our specimen is quite well comparable to those figured by ROTHPLETl(1886) and HAHN(1911) as "aff. prona"OpPEL. They differfrom true "prona", the origin- Order: Spiriferida WAAGEN, 1883 al of which is rather peculiar (OPPEL,1861, PI. 13, Fig. 7) Superfamily: Spiriferinacea DAVIDSON, 1884 in having another form of the plication and a flatter bra- Family: Spiriferinidae DAVIDSON, 1884 chial valve. Further comparison is for present made dif- Genus: Liospiriferina ROUSSELLE, 1977 ficult owing to the scarcity of the material and to the damage of the posterior part of our specimen. (OPPEL, 1861) Dis tri but ion: Steinplatte -Ioc. 2 (Praghorn im Steiner- Liospiriferina alpina nen Meer, Germany - Reichenbachquelle near Pfronten, (PI. 2, Fig. 7) CSFR). 1861 Spiriferina a/pinaOPP.- OPPEL, p. 541, PI. 11, Fig. 5. Lower Liassic (for "aff. prona"). According to ALMERAS, 1889 Spiriferina a/pina OPP. - GEYER, p. 71, PI. 8, Figs. 4-8 (cum (1964) "Rhynchonella" prona ranges from Lower Sinemurian syn.). to Upper Sinemurian (Lotharingian). 1894 Spiriferina a/pina OPP. - FucINI, p. 34, PI. 6, Fig. 10 (cum syn.). 1907 Spiriferina a/pina, OPP. - DAL PIAZ, p. 11, PI. 1, Fig. 1 (cum syn.). 1937 Spiriferina a/pina OPP.- ORMOS, p. 15 (cum syn.). "Rhynchonella" aft. paoIiiCANAvARI, 1880 1943 Spiriferina a/pina OPP.- VIGH, p. 349, Text-Fig. 14. (PI. 2, Fig. 9) ? 1954 Spiriferina a/pina OPPEL- CONTI, p. 194, PI. 9, Figs. 5-9. 1956 Spiriferina a/pinaOPPEL - SELLI, p. 14. aft. 1880 Rhynchone//aPao/ii, novo form. - CANAVARI, p. 69, PI. 1, 1964 Spiriferina a/pinaOPPEL - SIBLlK, p. 158, PI. 7, Fig. 2. Fig.1. 1966 Spiriferina a/pinaOPPEL - SIBLlK, p. 143, PI. 1, Fig. 3. ? 1880a Rhynchonella variabi/is SCHL. sp. var. - CANAVARI,p. 31, 1969 Spiriferina a/pinaOPPEL - DELANCE, p. 9, PI. A, Figs. 4,6. PI. 4, Fig. 11 only. 1990 Spiriferina a/pinaa/pina OPPEL - TCHOUMATCHENCO,p. 6, Text- ? 1897 RhynchonellaPao/ii CANAVARI-BOSE, p. 191, PI. 14, Fig. 3 Figs. 2-3, PI. 3, Figs. 4-7, PI. 4, Figs. 1-5. only. Mat e ria I: 1 well-preserved internal mould of the dimen- Mat e ria I: 1 slightly damaged specimen measuring 8.9 sions 12.6 x 12.9 x 8.8 mm. x 8.3 x 5.6 mm. Rem ark s: The specimen is placed under this variable Rem ark s: The specimen shows considerable resemb- species because of its outline, straight and long hinge lances to the smooth variants of "Rhynchonella" paolii as line, and its flat brachial valve. It differs from OPPEL'S figured by BÖSE,1897 and to the specimen figured by original specimen in the narrower cardinal area, in the CANAVARI,1880a under the name Rhynchonella variabi/is lower curved beak and in the development of weak sul- (this latter specimen was ascribed to "Rhynchonella"pao/ii cation of the pedicle valve resulting in the uniplication of already by GEYER,1889, p. 67). It differs from them, the anterior commissure. These characters were ascer- however, apart from the smaller dimensions and narrow- tained also later in larger material e.g. by GEYER(1889, er outline in having a lower plication and a massive beak. PI. 8, Fig. 8 - curved beak and uniplication of the an- The typical "Rhynchonella" paolii is semiplicate. terior commissure), by DALPIAl (1907,PI. 1, Fig. 1 - nar- Dis tri but ion: Steinplatte -Ioc. 3. rower cardinal area) etc. Considerable external resemb- "Rhynchonella" paolii originates from Pliensbachian but it lances to our specimen could be also found in Spiriferina was described by GEYER,1889 also from the Lower Pichleri NEUMAYR,1879 (differing in outline and in much Liassic of Hierlatz. shorter hinge line) and in the Middle Liassic Spiriferina can- tianensis CANAVARI,1881 (straight beak, no sulcus and no anterior uniplication). A very similar, but uncomplete "Rhynchonella" sp. specimen was described by RENl (1932, p. 15, PI. 1, Fig. 8)from the Upper Liassic of Greece as Spiriferina Pich- (PI. 1, Fig. 6) leri mut. Spanish representatives of "alpina" as figured by Mat e ria I: 1 slightly damaged specimen of the dimen- DELANCE(1969) and by ROUSSELLE(1977) possess short sions 10.6 x 14.6 x 9.2 mm. hinge lines. The latter author put surprisingly "alpina"into Rem ark s: Despite relatively very good preservation, it is the synonymy of Liospiriferina rostrata (SCHLOTH.).However, not possible to assign the specimen with certainty to a I adhere to a moderately conservative concept of "alpina" particular taxon, because it shows external features and keep both these species separate. common to several species. The shape of shell and Dis tri but ion: Steinplatte - loc. 2 (Hierlatz, Seehof- character of ribs suggest Calcirhynchia (?) plicatissima graben/Achensee, Breitenberg/Attersee; Germany - (QUENSTEDT)which has, however, a slightly different type Hindelang, France, Switzerland, Belgium, Spain, Italy, of plication and a more convex brachial valve. Interest- CSFR, Hungary, Bulgaria, Roumania, Turkey, Greece, ing is a strong resemblance to some English variants of ?Morocco). Squamirhynchia squamiplex (QUENSTEDT,1871), esp. to that Sinemurian-Domerian (according to ALMERAS,1964), on PI. 12, Fig. 7 by AGER(1967), which only differs in hav- ?Toarcian.

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Liospiriferina obtusa (OPPEL, 1861) (PI. 2, Fig. 6)

1861 Spiriferina ob/usa OPP. - OPPEL, p. 542, PI. 11, Fig. 8. 1889 Spiriferina ob/usa OPP. - GEYER, p. 75, PI. 8, Figs. 13-15, PI. 9, Figs. 1-5 (cum syn.). () 1907 Spiriferina ob/usa, OPP. - DAL PIAZ, p. 17 (cum syn.). 0.3 ? 1932 Spiriferina ob/usa OPPEL var. - RENZ, p. 11 (cum syn.). 1937 Spiriferina ob/usa OPP. - ORMOS, p. 17 (cum syn.). 1943 Spiriferina ob/usa OPP. - VIGH, p. 351 , PI. 27, Figs. 33-35. 1956 Spiriferina ob/usa OPPEL- SELLI, p. 17. .J 2.8 1957 Spiriferina ob/usa OPPEL- MAHEL, p. 187, PI. 8, Figs. 6-10. '- 1964 Spiriferina ob/usa OPPEL- SIBLlK, p. 159. 1967 Spiriferina ob/usa OPP. sensu GEYER - SACCHI VIALLI & CAN- ) l 3.1 TALUPPI, p. 89, PI. 13, Fig. 8. 1.~ Mat e ria I: 1 slightly damaged specimen with both valves ( ) 4.5 (dimensions 18.2 x 18.9 x 13.8 mm), 1 pedicle and 1 brachial valve. \...... / L-/ 4.8 ~] A e mar k s : The specimens agree in all observed external I j features with the Hierlatz forms as described in detail 2.~ and figured by GEYER(1889). He discussed also very si- l J 5.5 milar species Liospiriferina angulata (OPPEL, 1861). The dif- Text-Fig.5. ferences are probably not granted specific rank and it is Lobo/hyris ovatissimaeformis (BÖCKH). possible that L. obtusa is the same species as L. angulata. Incomplete series of transverse sections, due to the bad preservation. But proof of identity will require a revisional study of the Length of specimen 19.0 mm. ST 2. Magnified. Hierlatz material. Another species, which has to be considered here, is Middle Liassic Liospiriferina sicula (GEM- Mat e ria I: 3 partly damaged specimens. The sectioned MELLARO,1874). It is distinctive in its almost equi-dimen- one measures 19.0 x 15.5 x 9.4 mm. sional form and in wider and deeper ventral sulcus. In t ern a Ich a ract e rs: The complete absence of dental Judging from the illustration of L. obtusa by GEYER, 1889, lamellae and of a median septum leave no doubt that the PI. 9, Fig. 1 there is every reason to believe that "obtusa" specimen belongs to the Terebratulidae. Internal details and "sicula"should be considered closely allied. are basically the same as those made known by GAETANI Dis tri but ion: Steinplatte - loc. 2 (Hierlatz, Germany - (1970). Our specimen showed lateral insertion of teeth, Hindelang, Switzerland, Italy, CSFA, Hungary, very wide sockets with flat bottoms, clearly separated ?Greece). from strongly bent hinge plates, and a short loop. Lower Sinemurian - Upper Sinemurian (Lotharingian) - Aemarks: BÖCKH (1874) distinguished his new species ?Domerian (according to ALMERAS, 1964). from Terebratula ovatissima QUENSTEDT,1858 above all by its smaller size and narrower beak. The material from Stein- platte corresponds well to BÖCKH'S description and re- Liospiriferina sp. sembles considerably the specimen figured by him on (PI. 2, Fig. 5) PI. 1, Fig. 12. Externally very similar "Terebratula"juvavica Mat e ria I: 1 damaged internal mould of the dimensions GEYER, 1889 from Hierlatz is well distinguishable from 14.7 x 13.5 x 10.1 mm. "ovatissimaeformis"by its sharp beak ridges. A e mar k s: The specimen resembles some variants of Dis tri but ion: Steinplatte - loc. 2 (Hungary, Italy). Het- Liospiriferina rostrata (ZIETEN, 1830) but differs from them in tangian-?Lower Sinemurian. having lesser convexity of brachial valve, lower beak and shorter hinge line. In regard to this last character, the specimen is comparable to Liospiriferina terebratuloides Family: Pygopidae MUlA-WOOD, 1965. (SEGUENZA, 1883) from the Upper Liassic that is, Genus: Linguithyris BUCKMAN, 1917 however, much bigger and lacks ventral sulcation. The definite specific attribution of the specimen is, for the present, made difficult owing to the scarcity of Linguithyris aspasia (MENEGHINI, 1853) material. (PI. 2, Figs. 1,8, Text-Fig. 6) Dis tri but ion: Steinplatte -Ioc. 2. 1853 Terebra/ulaAspasia - MENEGHINI, p. 13 (non vidi). 1880 Terebra/ulaAspasia - UHLIG, p. 274 (cum syn.). 1887 Terebra/ula (Pygope) Aspasia MENEGHINI - DE STEFANI, p. 43, Order: Terebratulida WAAGEN, 1883 PI. 1, Figs. 6-9. Superfamily: Terebratulacea GRAY, 1840 1889 Terebra/ufa Aspasia MENEGH. - GEYER, p. 14, PI. 2, Figs. 13-15. Family: Terebratulidae GRAY, 1840 1932 Terebra/ula (Pygope) Aspasia MENEGHINI - RENZ, p. 28, PI. 2, Genus: Lobothyris BUCKMAN, 1917 Figs. 3,5 (cum syn.). 1943 Glosso/hyris aspasia MGH. - VIGH, p. 331 (cum var. major ZITT., minor ZITT., dila/a/a CAN., comparabile CAN., var.(?), n.var.), Lobothyris ovatissimaeformis (BÖCKH, 1874) Text-Figs. 9,10a, PI. 25, Figs. 20-27, PI. 26, Figs. 1-2. (Text-Fig. 5) 1953 Pygope aspasia MENEGHINI- RossI RONCHETTI& BRENA, p. 10, PI. 10, Fig. 3. 1874 Terebra/ula ovatissimaeformis n.sp. - BOCKH, p. 141, P1.1, 1959 Propygope aspasia MENEGHINI- AGER, p. 1024, PI. 128, Fig. 6. Figs.11-14. 1964 Propygope? aspasia (MENEGHINI)- SIBLlK, p. 163, PI. 7, Fig. 3. 1970 Lobo/hyris ovatissimaeformis (BOCKH) - GAETANI, p. 384, PI. 30, 1967 Nuclea/a aspasia (MGH.) - SACCHI VIALLI & CANTALUPPI, p. 100, Figs. 2-6, Text-Fig. 10 (cum syn.). PI. 14, Figs. 13-15, Text-Fig. 24.

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000.3 0.6 0.8

~--.-#, ~"..// ~~ ".aß ~1.4 ~ 1.3 ~1.8 ~1.6 ~J

\.. J

Text-Fig. 6. Linguithyris aspasia (MENEGHINI). Originallength C. 14.0 mm. ST 2. Magnified.

Mat e ria I : 6 partly damaged specimens with both valves does not warrant a study of the taxonomic relations be- and 4 single valves. The figured specimens measure tween these two species. According to VOROS in PRO- 13.9 x 17.9 x 8.8 mm (PI. 2, Fig. 1) and 8.8 x 12.0 x SOROVSKAYA& VOROS (1988), ZITTEL (1869) should be 7.1 mm (PI. 2, Fig. 8). considered the author of "aspasia" as he described it for In t ern a Ich a ra c t e r s : There are no dentallamellae and the first time. pedicle collar present. Cardinal process unclearly sug- Dis tri but ion: Steinplatte - loc. 2 and 3 (Hierlatz, gested (?). Short hinge teeth inserting laterally into Breitenberg/Attersee, Aurikelwand and Ob. Brunntal/ sockets that are well separated from the thin hinge Untersberg, Schafberg, Enzesfeld, Germany - Fagstein, plates. Dorsal median septum absent. Short loop about Switzerland, CSFR, Hungary, Italy, Spain, Greece, Tur- 0.3 length of valve developed. key, ?Tunis). Rem ark s: Linguithyris aspasia together with its "varieties" The species ranges from the Hettangian to the Upper and with a series of allied sulcate species are character- Liassic and seems to be nearly useless from the strati- istic elements of the Tethyan brachiopod fauna. "Aspasia" graphical point of view. Its long-ranging stratigraphical is no more attributed to Propygope BITTNER,1890 because distribution was mentioned already by UHLIG, 1880, p. this genus develops a strong dorsal septum which is 275. missing in the species under consideration. It seems. that there is a continuous range of variation between "aspasia" and "Terebratula" nimbata OPPEL, 1861. Our two Superfamily: Zeilleriacea ALLAN, 1940 narrower specimens with only shallow sulcation approach externally to the latter species and could be eas- Family: Zeilleriidae ALLAN, 1940 ily determined as "nimbata". The material at disposal Genus: Zeilleria BAYLE, 1878

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Zeilleria perforata (PIETTE, 1856) Septalium large. Median septum persisting for about (PI. 2, Fig. 4, Text-Fig. 7) one third of the length of shell. All details concerning loop not known due to bad preservation. 1856 Terebratu/aper/orata- PIETTE,p. 206, PI. 10, Fig. 1. ? 1909 Watdheimia (leit/eria) per/orata PTT.sp. - TRAUTH,p. 71, Rem ark s: The species was very thoroughly described PI. 2, Fig. 11 (cum syn.). by DELANGE(1974) and nearly nothing can be added to it. 1936 lei/teria per/orata E. PIETTE- JOLY,p. 157 (cum syn.). The specimens figured by him are, on the average, wider 1970 lei/teria perforata (PIETTE)- GAETANI,p. 387, PI. 30, Figs. 7-15 (cum syn.). if compared with our material. The outlines of Steinplat- 1974 leit/eria (leilleria) per/orata (PIETTE)- DELANCE,p. 75, PI. 1, te specimens approach to those of PIETTE'S specimen Figs. 1-7, Text-Figs. 6-1 to 6-9 (cum syn.). (the longer one) or of GAETANI'SFigs. 11 and 14 on PI. 30. non 1967 Lobothyris perforata (PIETTE)- SACCHIVIALLI& CANTALUPPI, TULUWEIT(1965) referred "perfarata"to his new genus Kera- p. 94, PI. 14, Figs. 9-10, Text-Fig. 19. tathyris, but this is not thought to be likely. Moreover, the Mat e ria I: 2 internal moulds with shell remains. The diagnosis of the new genus is vague and fits equally well dimensions: 19.8 x 14.1 x 11.3 mm (sectioned) and with lei/feria. Keratathyris was later reasonably included in 19.4 x 14.0 x 9.1 mm (figured). the synonymy of lei/feria by DELANGE(1974). In t ern a Ich ara c t e rs: Pedicle collar and clear cardinal Dis tri but ion: Steinplatte - loc. 1 and 2 (Breitenberg/ process were not observed. Dental lamellae rather Attersee, Germany - Pfonsjoch, Italy, England, France, short, dorsally convergent and then subparallel. Low Belgium, Switzerland, CSFR, Hungary, ?Crimea, denticula developed laterally. Hinge teeth without ?Caucasus). crenulation, in free contact with large sockets. Outer Hettangian to lowermost Lotharingian - Obtusum Zone socket ridges better developed than the inner ones. (according to DELANGE,1974).

~ ~ {1 V

4.~ 4.8~

Text-Fig. 7. leilleria perfora/a (PIETTE). Median septum persisted as a low ridge to a point 7.1. Originallength of specimen 19.8 mm. ST 2. Magnified.

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Zeilleria mutabilis (OPPEL, 1861) Rem ark s: Our specimens correspond well to those fi- (PI. 2, Fig. 3, Text-Figs. 8, 9) gured by OPPEL (1861) and GEYER (1889) and show the same variability which was discussed in detail by the 1861 Terebratuta mutabitis OPP. (WALDHEIMlA) - OPPEL, p. 538, latter author. Zeil/eria mutabilis is by far the most common PI. 10, Fig. 7. 1889 Wa/dheimia mutabilis OPP. - GEYER, p. 18, PI. 2, Figs. 31-36, and most varied of all Lower to Middle Liassic zeillerids PI. 3, Figs. 1-7 (cum syn.). from Alpine domain. A great number of different names 1893 Watdheimia mutabitis OPPEL- BÖSE, p. 639. have been given to the "mutabilis" group of zeillerids that 1895 Wa/dheimia mutabitis OPPEL- FUCINI, p. 84, PI. 7, Figs. 29-31. should be taken into consideration during the determi- 1907 Wa/dheimia mutabitisOpP. - DAL PIAZ, p. 48, PI. 3, Fig. 7. nation. However, the present material does not warrant 1911 Wa/dheimia mutabi/is OPP. - HAHN, p. 557, PI. 20, Fig. 8. a further discussion of the complex synonymies in- 1937 Wa/dheimia mutabilis OPP. - ORMOS, p. 21. 1943 Wa/dheimia mutabitis OPP. - VIGH, p. 327, Text-Figs. 7 left, 8c, volved. ANTOSTGHENKO(1973) erected a new genus Spi- PI. 25, Figs. 9-10. nulothyris (with S. patilensis sp.n. from the Sinemurian of 1956 lei/teria mutabi/is (OPPEL) - SELL!, p. 21, PI. 2, Fig. 1. Caucasus as the type-species) and referred "mutabilis"to 1964 leilleria mutabi/is (OPPEL) - SIBL!K, p. 165, Text-Fig. 2. it. As one of the most important features distinguishing 1966 leilleria mutabi/is (OPPEL) - PEVNY, p. 280. new genus from Zeil/eriathe absence of the inner cardinal 1967 leit/eria mutabi/is (OPP.) - SACCHI VIALL! & CANTALUPPI, p. 108, plates was reported. DELANGE(1974, p. 31) in the com- Text-Fig. 29, PI. 15, Fig. 10. prehensive monograph on West European zeillerids Mat e ria I: 8 more or less fragmentary internal moulds. negates, however, the possibility of discerning between The better preserved specimens measure: 17.5 x 14.3 inner and outer parts of zeillerid cardinal plates. A more x 8.8 mm, 17.4 x 16.3 x 8.5 mm, 17.0 x 7 x 9.2 mm, detailed study is required to support ANTOSTGHENKO'S 716.7 x 14.8 x 8.6 mm (figured), 13.6 x 12.3 x opinion before definite taxonomic conclusions can be 6.8 mm. drawn. In the present paper the traditional affiliation of In t ern a Ich a r act e rs: Dentallamellae subparallel and "mutabilis"to "Zeil/eria" is still used. very short. Cardinal process not ascertained. Hinge plates thick and narrow, poorly separated from the in- Distribution: Stein platte - loc. 2 (Hierlatz, Aurikel- ner socket ridges. Septalium broad, but shallow. Me- wand/Untersberg, Kramsach bei Rattenberg, Germany dian septum strong and extending less than a third of - Hindelang, Switzerland, CSFR, Hungary, Italy, Turkey, the length of valve. Long loop with undulated ascending 7Crimea, 7Caucasus). branches. Sinemurian to Domerian (according to ALMERAS, 1964).

2.W \C:Q7 2.3

J J q

~ ~.3~.5 3.5 ~1

\ 3mm )

11.2 12.6 ~ ~ Text-Fig. 8. leilleria mutabi/is (OPPEL). Due to the bad preservation, only discontinous series of transverse sections has been made. Originallength 16.8 mm. ST 2. Magnified.

@~ 0.1 0.3 0.5

(J C:>

~.2 2mm

Text-Fig. 9. leilleria mutabitis (OPPEL). Sections through another specimen showing a large septalium. Originallength 12.0 mm. ST 2. Magnified.

Zeilleria stapia (OPPEL, 1861) Zeilleria alpina (GEYER, 1889) (PI. 2, Fig. 2, Text-Fig. 10) (PI. 2, Fig. 10)

1861 Terebratuta stapia OPP. (Watdheimia) - OPPEL, p. 539, PI. 11, 1889 Waldheimia alpina nov.sp. - GEYER, p. 29, PI. 3, Figs. 33-38. Fig.2. 1937 Waldheimia alpina GEYER- ORMOS, p. 24. 1889 Waldheimia stapia OPP. - GEYER, p. 16, PI. 2, Figs. 25-30. 1943 Waldheimia alpina GEY. - VIGH, p. 324, Text-Fig. Sc, PI. 25, 1936 leilieriastapiaA. OPPEl-JOlY, p. 157. Fig.4. 1943 Waldheimia stapia OPP. - VIGH, p. 327. 1966 leilleria alpina (GEYER)- PEVNY, p. 281. 1964 leitleria stapia OPP. - SACCHI VIAlLl, p. 19, PI. 2, Fig. 11. Mat e ria I: 3 more or less damaged specimens. The Mat e ria I: 5 internal moulds. The best preserved ones figured one measures 10.1 x 10.2 x 5.4 mm. measure 20.5 x 14.0 x 10.0 mm and 16.5 x 14.1 x Rem ark s : The material shows all the characters given by 9.7 mm (figured). GEYER and differs in having a very shallow sulcation in In t ern a Ich a ra c t e r s: Delthyrial cavity subquadrate in the brachial valves, and a rounded cardinal margin cross-section, separated from semicircular umbonal which is observable in the figured specimen only. The cavities by short, ventrally divergent dental lamellae. same rounded character of the cardinal margin was Strong teeth inserted into deep sockets. Denticula not ascertained also in the Hungarian material (VIGH, 1943, observed. Hinge plates practically horizontal, fused or Text-Fig. 5c). From the mutual morphological "pas- forming very shallow septalium. Median septum extend- sages" among "a/pina", lei/feria enge/hardti (OPPEL), Z. ewa/di ing for about one third of the length of valve. Loop about (OPPEL) and Z. mutabi/is (OPPEL) is it clear that the conven- 2/3 length of valve, with undulated ascending branch- tional species concept within this group needs recon- es. sideration and that the variation could well be intraspe- cific. Hungarian lei/feria baconica comp/anata (BÖCKH, 1874) Rem ark s: The specimens agree in all observed external seems to belong to the same morphological group. features with those described in detail and figured by Dis tri but ion: Stein platte - loc. 2 (Hierlatz, CSFR, GEYER (1889) who discussed the great variability of the Hungary, Italy, ?Crimea). species and its "passages" to lei/feria mutabi/is. Thicker Hettangian to Upper Sinemurian-Lotharingian (accord- specimens resemble in some respect lei/feria venusta ing to ALMERAS, 1964). (UHLIG, 1880) but differ from it in the absence of planareas. It is probable that "stapia" is the same species as "mutabi/is", but proof of identity will require a revisional Zeilleria aft. apenninica (ZITTEL, 1869) study of the Hierlatz material. aff. 1869 Terebratula (Waldheimia) Apenninica. ZITT. - ZITTEL, p. 127, PI. 14, Fig. 9. Distribution: Steinplatte - loco 2 (Hierlatz, Belgium, aff. 1889 Waldheimia Apenninica V. ZITTEL - GEYER, p. 33, PI. 4, CSFR, Hungary, Italy, ?Crimea). Figs.8-12. Sinemurian. aff. 1892 Waldheimia Apenninica ZITT.- PARONA, p. 49, PI. 2, Fig27.

\J f~W~ 3.5 2.7 2.5 2.2

} (1 ~\/ ( ) r

3.7

~ r-\ \ / \ /

Text-Fig. 10. leilleria stapia (OPPEL). Length of specimen? 17.0 mm. ST 3. Magnified.

Mat e ria I: 2 incomplete internal moulds. The better pre- Dis tri but ion: Steinplatte -Ioc. 2. served one measures 11.2 x ?11.5 x 7.8 mm. lei/feria apenninica comes from the Middle Liassic but was Remarks: The specimens are quite well comparable to reported by GEYER, 1889 from the Lower Liassic of Hier- those described and figured by GEYERand PARONAbut latz, too. differ from them, however, in having subtrigonal outlines with the maxium width situated near the anterior margin Acknowledgements of shell. A median septum persisting for about one third of the length of valve, and short dentallamellae are as- I am particularly indebted to Dr. Harald LOBITZER (Geologische Bun- certainable on our material. lei/feria ampezzana (BÖSE & desanstalt, Vienna) who kindly offered me the possibility to participate in the biostratigraphical research of the Steinplatte area, and gave me ad- SCHLOSSER, 1900) is distinguishable from our speci- vice on field work and assistance with collecting. The financial support mens by its subpentagonal outline with strongly emargi- by the Geologische Bundesanstalt in Vienna that made my field work nate anterior margin. possible is heartily acknowledged.

[ Plate 1 J

Fig. 1: Cirpa planifrons (ORMOS). Locality sr 3. Collection of the Geologische Bundesanstalt Wien (Museum). GBA no. 1992/6/1. x2. Fig. 2: Cirpa planifrons (ORMOS). Locality sr 2. GBA no. 1992/6/2. x2. Fig. 3: Cirpa planifrons (ORMOS). Locality sr 3. GBA no. 1992/6/3. x2. Fig. 4: Calcirhynchia (?) plicalissima (QUENSTEDT). Locality sr 2. Unusually flat specimen. GBA no. 1992/6/4. x2. Fig. 5: "Rhynchonella"aff. pronaOpPEl. Locality sr 2. GBA no. 1992/6/5. x2. Fig. 6: "Rhynchonella"sp .. Locality sr 3. GBA no. 1992/6/6. x2. Fig. 7: "Rhynchonella" fraasiOpPEL. Locality sr 2. GBA no. 1992/6/7. x2. Fig. 8: ? "Rhynchonella" fraasi OPPEL. Locality sr 2. Young specimen (?). GBA no. 1992/6/8. X3. Fig. 9: Furcirhynchia emmrichi (OPPEL). Locality sr 2, GBA no. 1992/6/9. x 1.5. Fig. 10: Cuneirhynchia retusifrons (OPPEL). Locality sr 2. GBA no. 1992/6/10. x2.5.

All specimens were coated with ammonium chloride before photographing. Photographs (Pis. 1 and 2) by Mr. J. BROZEK(Prague).

( Plate 2 J

Fig. 1: Linguithyris aspasia (MENEGHINI). Locality ST 2. GBA no. 1992/6/11. x 2. Fig. 2: Zeilleria stapia (OPPEL). Locality ST 2. GBA no. 1992/6/12. x 2. Fig. 3: Zeilleria mutabilis (OPPEL). Locality ST 2. GBA no. 1992/6/13. x 2. Fig. 4: Zeilleria perforata (PIETTE). Locality ST 1. GBA no. 1992/6/14. x 2. Fig. 5: Liospiriferina sp .. Locality ST 2. GBA no. 1992/6/15. x 2. Fig. 6: Liospiriferina obtusa (OPPEL). Locality ST 2. GBA no. 1992/6/16. x 1.5. Fig. 7: Liospiriferina alpina (OPPEL). Locality ST 2. GBA no. 1992/6/17. x 2. Fig. 8: Linguithyris aspasia (MENEGHINI). Locality ST 2. GBA no. 1992/6/18. x 2. Fig. 9: "Rhynchonella" aff. paolii CANAVARI. Locality ST 3. GBA no. 1992/6/19. x 3. Fig.10: Zeilleria alpina (GEYER). Locality ST 2. GBA no. 1992/6/20. x 2.5.

References

AGER, D.V. (1956-1967): A Monograph of the British Liassic aRMOS, E. (1937): Die Brachyopoden-Fauna der unteren Lias in Rhynchonellidae. - I (1956), I-XXVI, 1-50, Pis. 1-4, 11(1958), Kekhegy (Bakonyerwald). - Abh. min. geol. Inst. Tisza Univ., 9, 51-84, Pls.5-7, III (1962), 85-136, Pis. 8-11, IV (1967), 1-45, PI. 1, Debrecen. 137-172, Pis. 12-13,London. PETERHANS, E. (1926): Revision des Brachiopodes Iiasiques du AGER, D.V. (1959): Lower Jurassic brachiopods from Turkey. - Grammont, des Tours d' Ai, du Pissot et de Rossiniere figures Journ. Paleont., 33/6,1018-1028, Pis. 128-9, Tulsa. dans I'ouvrage de M.H. Haas. - Mem. Soc. Vaud. Sei. nat., 2/6, ALMERAS, y. (1964): Brachiopodes du Lias et du Dogger. - Doc. 353-383, Pis. 1-2, Lausanne. Labor. Geol. Fac. Sei., 5, 1-161, Lyon. PEVNY, J. (1964): On brachiopods from Northern Little Carpa- ANTOSTCHENKO,ZA (1973): On Spinulothyris ANTOSTCHENKOgen. thians. - Geol. prace, Zpravy, 33,157-172, Pis. 4-6, Bratislava novo (Brachiopoda). - Bull. Moscow Soc. Nat. Geol., 48/4, (in Slovak). 109-116, PI. 1, Moscow (in Russ.). PEVNY, J. (1966): Liassic Brachiopoda from the Mountains Cach- ticke pohrie West Slovakia. - Geol. sbornik, 17/2, 271-282, BÖCKH, J. (1874): Die geologischen Verhältnisse des südlichen Pis. 11-12, Bratislava. Teiles des Bakony. II. - Mitt. Jb. ung. geol. Anst., 3/1,1-180, PIETTE, E. (1856): Notice sur les gres d'Aiglemont et de Rimogne. Pis. 1-7, Budapest. - Bull. Soc. geol. France, 2. ser., 13, 188-207, PI. 10, Paris. BÖSE, E. (1893): Die Fauna der liasischen Brachiopodenschichten PROSOROVSKAYA,E.L. & VÖRÖS, A. (1988): Pliensbachian, Bajocian bei Hindelang (Algäu). - Jb. k.k. geol. Reichsanst., 42 (1892), and Callovian Brachiopoda. -In: Evolution ofthe Northern Mar- 627-650, Pis. 14-15, Wien. gin of Tethys, vol. 1., Mem. Soc. geol. France, N.S. 154, 61-70, BÖSE, E. 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