DOCSLIB.ORG

Szwedo J. 2006. First Fossil Record of Cedusini in the Eocene Baltic

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Szwedo J. 2006. First Fossil Record of Cedusini in the Eocene Baltic Russian Entomol. J. 15(3): 327–333 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2006 First fossil record of Cedusini in the Eocene Baltic amber with notes on the tribe (Hemiptera: Fulgoromorpha: Derbidae) Ïåðâàÿ èñêîïàåìàÿ íàõîäêà Cedusini â ýîöåíîâîì áàëòèéñêîì ÿíòàðå ñ çàìå÷àíèÿìè î òðèáå (Hemiptera: Fulgoromorpha: Derbidae) Jacek Szwedo ß. Øâåäî Department of Systematics and Zoogeography, Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, PL 00- 679 Warszawa, Poland. E-mail: [email protected] Отделение систематики и зоогеографии, Музей и институт зоологии ПАН, Вильча 64, PL 00-679 Варшава, Польша. KEY WORDS: Hemiptera, Derbidae, Cedusini, taxonomy, new genera, new species, new combinations, Eocene, Baltic amber, fossil insects. КЛЮЧЕВЫЕ СЛОВА: Hemiptera, Derbidae, Cedusini, таксономия, новые роды, новый вид, новые комбинации, эоцен, балтийский янтарь, ископаемые насекомые. ABSTRACT. New extinct genus and species — Emelja- Introduction novedusa gentarna gen.n. et sp.n., of the planthopper family Derbidae, tribe Cedusini, from the middle Eocene The planthopper family Derbidae houses a large Baltic amber is described. New extant genera of Cedusini number of species. It is one of the largest and most are established: Hauptenia gen.n., Muiredusa gen.n., morphologically differentiated family among Fulgoroi- Produsa gen.n. New combinations are given: Hauptenia dea. Fennah [1952] and Emeljanov [1995] elaborated magnifica (Yang & Wu, 1993), comb.n.; H. jacula (Yang classification and evolution of Derbidae, but the distri- & Wu, 1993), comb.n.; H. fellea (Yang & Wu, 1993), bution of some characters still remains controversial comb.n., H. glutinosa (Yang & Wu, 1993), comb.n., H. and the available data are not sufficient to substantiate idonea (Yang & Wu, 1993), comb.n., Muiredusa brun- nea (Muir, 1914), comb.n., M. littorea (Yeh & Yang, the phylogeny of the group. Numerous taxonomic prob- 1993), comb.n., M. ignota (Yeh & Wu, 1993), comb.n., lems within the group remain, and Derbidae, as well as Produsa concava (Yang & Wu, 1993), comb.n., and P. their subunits, are sometimes believed to be paraphyle- cubica (Yang & Wu, 1993), comb.n. Key to genera placed tic [Emeljanov & Fletcher, 2004]. The higher classifica- in Cedusini is given and placement of the fossil genus tion of extant Derbidae has recently been presented by and its relationships within the tribe are discussed. Emeljanov [1995], who presented a more detailed sys- tem based on external morphological characters. He РЕЗЮМЕ. Новый ископаемый род и вид цикадо- divided Derbidae into three subfamilies and 18 tribes. вых семейства Derbidae, трибы Cedusini — The subfamily Cedusinae comprises the tribes Ipsnoli- Emeljanovedusa gentarna gen.n. et sp.n. описан из ni, Goneokarellini, Vinatini, Cedusini, Phrygiini, and re- среднего эоцена балтийского янтаря. Установлены cently [Emeljanov & Fletcher, 2004] the tribe Breddinio- новые роды для современных Cedusini: Hauptenia lini Fennah, 1950, formerly placed in Achilidae [Fennah, gen.n., Muiredusa gen.n., Produsa gen.n. Предложе- 1950; Emeljanov, 1991, 1992b] but lately transferred here. ны новые комбинации: Hauptenia magnifica (Yang & Subfamily Derbinae comprise the tribes Cedochreini, Wu, 1993), comb.n.; H. jacula (Yang & Wu, 1993), Dawnarioidini, Derbini, Cenchreini and Nicertini, while comb.n.; H. fellea (Yang & Wu, 1993), comb.n., H. subfamily Otiocerinae include the tribes: Kamendakini, glutinosa (Yang & Wu, 1993), comb.n., H. idonea (Yang Rhotanini, Otiocerini, Patarini, Neocyclocarini, Phenici- et Wu, 1993), comb.n., Muiredusa brunnea (Muir, 1914), ni, Zoriaidini, Sikaianini and recently described [Ba- comb.n., M. littorea (Yeh & Yang, 1993), comb.n., M. naszkiewicz & Szwedo, 2005] tribe Aquaeliciini. Accor- ignota (Yeh & Wu, 1993), comb.n., Produsa concava ding to Emeljanov [1995], the taxa placed in the tribe (Yang & Wu, 1993), comb.n. и P. cubica (Yang & Wu, Mysidiini, proposed by Broomfield [1985], are to be 1993), comb.n. Дана определительная таблица родов, placed within Derbini. помещённых в трибу Cedusini, и обсуждены поло- The Eocene Baltic amber inclusions of Derbidae жение и родственные отношения ископаемого рода comprise only Positrona shcherbakovi Emeljanov, 1994, в пределах трибы. of the tribe Otiocerini [Emeljanov, 1994] and Lugeilan- 328 Jacek Szwedo gor elektrokleistis Szwedo, 2005 [Szwedo, 2005] of the sometimes convex, concave, or subparallel to the outer tribe Otiocerini are known. Unnamed representatives of ventral margin, [Flynn & Kramer, 1983; Kramer, 1983]. the tribe Cedusini mentioned by Szwedo [2002] are de- One species, Cedusa wolcotti Muir, 1924 seems not to scribed below. From Oligocene/Miocene resins of the be congeneric with the other American species ascribed New World more taxa are described. Derbidae inclu- to Cedusa, as it is the only member of the genus with sions described from Dominican amber are: Cedusa the posterior prolongation of the pygofer and is very credula Emeljanov & Shcherbakov, 2000, of the tribe unique possessing a ventral plate [Caldwell & Mar- Cedusini and Dysimia imprudens Emeljanov & torell, 1951, Flynn & Kramer, 1983]. It seems that the Shcherbakov, 2000 of the tribe Derbini [Emeljanov & genus Cedusa is a paraphyletic unit, and needs to be Shcherbakov, 2000]. Inclusions recorded in Mexican revised, as well as other taxa placed in Cedusini. amber are: Cedusa baylissae Szwedo & Ross, 2003 of the tribe Cedusini [Szwedo & Ross, 2003] and Co- The following key could be used for delimitation of pallinges chiapasensis Szwedo, 2004 of the tribe Cen- genus-level taxa ascribed to Cedusini. chreini [Szwedo, 2004]. A few other unnamed Derbidae 1.Tegmen with joined claval veins Pcu+A1 reaching commis- are known from Oligocene/Miocene Dominican amber sural margin of tegmen, reaching vein A2. Tegmen with — specimens figured in Poinar & Poinar [1999] and branches of stem of the vein M forming anterior comb2 –.Tegmen with joined claval veins Pcu+A reaching claval Eocene Bitterfeld amber [Szwedo, 2002]. Mysidioides 1 migdisovae Emeljanov, 2002, of the tribe Otiocerini has suture, reaching vein CuP, near apex of clavus. Tegmen with branches of stem of the vein M forming posterior been described by Emeljanov [2002] from Miocene stra- comb.............................................................................7 ta of Stavropol’ (Northern Caucasus Mts.) The record 2.Wing with vein ScRA1 very short, reaching anterior margin of Derbidae from Upper Triassic strata of Brazil [Pinto, near wing coupling apparatus......................................3 1956] is based on misinterpreted material [Emeljanov, –.Wing with vein ScRA1 long, reaching anterior margin near 1994; Szwedo, 2002; Szwedo et al., 2004]. This fossil its apex..................................... Emeljanovedusa gen.n. known as Sanctipaulus mendesi Pinto, 1956, has been 3.Transversely oblique carina across the gena distinct from transferred to Trichoptera, undetermined family, by the subantennal process, as long as basal width of the Martins-Neto et al. [2003]. process. Antennal pedicel about as long as wide, slightly flattened. Genital styles asymmetrical, New World spe- cies......................................................... Cedusa Fowler Taxonomy type species: Cedusa funesta Fowler, 1904: 103; by subse- quent designation by Muir 1913: 35. –.Genital styles symmetrical, Old world species...........4 The tribe Cedusini could be characterized as fol- 4.Transversely oblique carina across the gena between the lows: head and tegmina without sensory pits, apical subantennal process and lateral carina of frons not longer segment of rostrum distinctly longer than wide, female than half of the basal width of the process, confluent with with abdominal sternite VII not fused with abdominal the process as a broad flange. Tegmen with veins Sc+R fork tergite VIII. The last feature is of particular interest as, basad of claval veins junction, fork of vein CuA at level fusion of the sternite VII with tergite VIII is believed to claval veins junction. Antennal pedicel about twice as long be synapomorphy for all Derbidae, with exception of as wide. African and Palaearctic species........................ Cedusini and Derbini [Emeljanov 1992a]. According to ................................................................ Malenia Haupt Emeljanov [1995] the tribe Cedusini comprises three type species: Lamenia bosnica Horváth, 1907: 322; by original designation by Haupt 1929: 194. genera: Cedusa Fowler, 1904, Melusa Emeljanov, 1995, –.Transversely oblique carina across the gena between sub- and Eocenchrea Muir, 1913. The taxonomic status of antennal process and lateral carina of frons indistinct. Anal the genus Malenia Haupt, 1929 (synonym of Cedusa or tube not distinctly elongated, about twice as long as wide, distinct genus) remained controversial for a long time. usually with apex not reaching level of apex of genital Fennah [1961] discriminated those two genera on the styles. Oriental species................................................5 basis of venation features. Yang & Wu [1993] separat- 5.Genital styles slender, dorsobasal projection basad. Hind ed Taiwanese species ascribed to genera Cedusa and leg tibiotarsal spinal formula 6 : 5 : 5...........................6 Malenia on the basis of hind leg femorotarsal formula –.Genital styles short and stout, dorsobasal projection and structure of the male genital block. All African distad. Hind leg tibiotarsal spinal formula 7 : 6 : 5. Anal style of male turned ventrad or nearly so, slightly
Load more

Recommended publications
  • A New Species of Planthopper in the Genus Agoo Bahder & Bartlett
    Zootaxa 4853 (2): 254–264 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2020 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4853.2.6 http://zoobank.org/urn:lsid:zoobank.org:pub:949733F1-D47A-44B4-8315-29EA3BB0A85F A new species of planthopper in the genus Agoo Bahder & Bartlett (Hemiptera: Fulgoroidea: Derbidae) from coconut palm (Cocos nucifera L.) in Jamaica BRIAN W. BAHDER1, DE-FEN MOU2, CHARLES R. BARTLETT3, ERICKA E. HELMICK4, ASSUNTA BERTACCINI5 & WAYNE MYRIE6 1University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. �[email protected]; https://orcid.org/0000-0002-1118-4832 2University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. �[email protected]; https://orcid.org/0000-0003-4620-727X 3University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716-2160, USA. � [email protected]; https://orcid.org/0000-0001-9428-7337 4University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. �[email protected]; https://orcid.org/0000-0002-5153-0891 5University of Bologna, Department of Plant Pathology; Viale Fanin 42, 40127, Bologna, Italy. �[email protected]; https://orcid.org/0000-0002-5650-1512 6Coconut Industry Board, 18 Waterloo Road, Kingston 10, Jamaica W. I. �[email protected]; https://orcid.org/0000-0001-8969-1204 Abstract A new species of the genus Agoo Bahder & Bartlett, Agoo beani sp.
    [Show full text]
  • Accepted Version
    This document is the accepted manuscript version of the following article: Davranoglou, L. R., Mortimer, B., Taylor, G. K., & Malenovský, I. (2019). On the morphology and possible function of two putative vibroacoustic mechanisms in derbid planthoppers (Hemiptera: Fulgoromorpha: Derbidae). Arthropod Structure and Development, 52, 100880 (15 pp.). https://doi.org/10.1016/j.asd.2019.100880 This manuscript version is made available under the CC-BY-NC-ND 4.0 license http://creativecommons.org/licenses/by-nc-nd/4.0/ 1 On the morphology and possible function of two putative vibroacoustic 2 mechanisms in derbid planthoppers (Hemiptera: Fulgoromorpha: 3 Derbidae) 4 Leonidas-Romanos Davranogloua*, Beth Mortimera, Graham K. Taylora, Igor 5 Malenovskýb 6 7 *e-mail: [email protected] 8 aDepartment of Zoology, University of Oxford, Oxford OX1 3SZ, U.K. 9 bDepartment of Botany and Zoology, Faculty of Science, Masaryk University, Kotlářská 2, Brno, CZ-611 37, 10 Czech Republic 11 12 Abstract 13 A mechanism involving interaction of the metathoracic wing and third abdominal segment of 14 derbid planthoppers was first discovered over a century ago, and interpreted as a stridulatory 15 organ for sound production. Although referred to occasionally in later taxonomic works, the 16 detailed morphology, systematic distribution, and behavioural significance of this structure 17 have remained unknown, and its proposed use in sound production has never been 18 corroborated. Here we examine the distribution and morphology of the supposed stridulatory 19 organ of Derbidae and the recently-described vibratory mechanism of planthoppers – the 20 snapping organ, across 168 species covering the entire taxonomic spectrum of the family.
    [Show full text]
  • Taxonomia De Fulgoroidea No Brasil (Insecta: Hemiptera: Auchenorrhyncha), Com Ênfase Em Dictyopharidae
    MARCELO DA SILVA BAPTISTA TAXONOMIA DE FULGOROIDEA NO BRASIL (INSECTA: HEMIPTERA: AUCHENORRHYNCHA), COM ÊNFASE EM DICTYOPHARIDAE Tese apresentada à Universidade Federal de Viçosa, como parte das exigências do Programa de Pós- Graduação em Entomologia, para obtenção do título de Doctor Scientiae. VIÇOSA MINAS GERAIS – BRASIL 2006 Ficha catalográfica preparada pela Seção de Catalogação e Classificação da Biblioteca Central da UFV T Baptista, Marcelo da Silva, 1971- B222t Taxonomia de Fulgoroidea no Brasil ( Insecta: 2006 Hemiptera: Auchenorrhyncha ), com ênfase em Dictyopharidae / Marcelo da Silva Baptista. – Viçosa : UFV, 2006. x, 274f. : il. (algumas col.) ; 29cm. Texto em português e inglês. Orientador: Paulo Sérgio Fiuza Ferreira. Tese (doutorado) - Universidade Federal de Viçosa. Inclui bibliografia. 1. Fulgoroidea - Identificação. 2. Fulgoroidea - Catálogos e coleções. 3. Fulgoroidea - Distribuição geográfica. 4. Hemíptera. I. Universidade Federal de Viçosa. II.Título. CDD 22.ed. 595.754 MARCELO DA SILVA BAPTISTA TAXONOMIA DE FULGOROIDEA NO BRASIL (INSECTA: HEMIPTERA: AUCHENORRHYNCHA), COM ÊNFASE EM DICTYOPHARIDAE Tese apresentada à Universidade Federal de Viçosa, como parte das exigências do Programa de Pós- Graduação em Entomologia, para obtenção do título de Doctor Scientiae. APROVADA: 16 de setembro de 2006 Prof. Elidiomar Ribeiro da Silva Prof. José Eduardo Serrão (Co-Orientador) (Co-Orientador) Prof. Lucio Antonio de Oliveira Campos Profa. Milene Faria Vieira Prof. Paulo Sérgio Fiuza Ferreira (Orientador) Aos meus pais, Aldo Baptista e Sidnéa da Silva Baptista, que sempre fizeram de tudo para que eu alcançasse e conquistasse mais essa etapa na vida. À minha filha Victoria Nolding Barros Baptista, que sonho estar sempre ao meu lado. Amo vocês. ii AGRADECIMENTOS Em primeiro lugar gostaria de agradecer a cinco pessoas, que me concederem o privilégio de desenvolver a este trabalho no Laboratório de Insetos Aquáticos na Universidade Federal do Estado do Rio de Janeiro.
    [Show full text]
  • Hemiptera: Auchenorrhyncha: Fulgoroidea: Derbidae) with a New Species from Costa Rica and New Country Records
    Zootaxa 4718 (4): 521–535 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2020 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4718.4.6 http://zoobank.org/urn:lsid:zoobank.org:pub:E315343C-3418-4F43-BB85-61C0E8AACC81 Revised status of Omolicna subgenus Agoo (Hemiptera: Auchenorrhyncha: Fulgoroidea: Derbidae) with a new species from Costa Rica and new country records BRIAN W. BAHDER1, CHARLES R. BARTLETT2, ERICKA E. HELMICK3, EDWIN A. BARRANTES BARRANTES4, MARCO A. ZUMBADO ECHAVARRIA5, ERICA M. GOSS6 & MARINA S. ASCUNCE7 1University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. E-mail: [email protected] 2University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716-2160, USA E-mail: [email protected] 3University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. E-mail: [email protected] 4Universidad of Costa Rica—Sede San Ramón, Departmento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica. E-mail: [email protected] 5Universidad of Costa Rica—Sede San Ramón, Departmento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica. E-mail: [email protected] 6University of Florida, Department of Plant Pathology; 1453 Fifield Hall, Hull Rd., Gainesville, FL 32611-0680, USA E-mail: [email protected] 7University of Florida, Department of Plant Pathology; 1453 Fifield Hall, Hull Rd., Gainesville, FL 32611-0680, USA E-mail: [email protected] Abstract An ongoing survey for novel phytoplasmas and viruses that affect palms (Arecaceae) and their potential vectors is being conducted in Costa Rica.
    [Show full text]
  • Hemiptera): New Phylogenetic Evidence
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Cicadina = Beiträge zur Zikadenkunde Jahr/Year: 2011 Band/Volume: 12 Autor(en)/Author(s): Hamilton K.G. Andrew Artikel/Article: Making sense of Fulgoroidea (Hemiptera): new phylogenetic evidence. 57-79 Cicadina 12: 57-79 (2011)©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at 57 Making sense of Fulgoroidea (Hemiptera): new phylogenetic evidence K.G. Andrew Hamilton1 Abstract: Antennal sensilla in more than 450 genera o f all 21 putative families o f Fulgoroidea provide characters that correlate well with other characters, including those of nymphal wax filaments and adult behaviour. Together, these characters indicate that Issidae, Lophopidae and Tropiduchidae are polyphyletic taxa, and suggest that Achilidae, Caliscelidae, Cixiidae and Dictyopharidae are paraphyletic. Two strongly supported monophyletic lineages comprise six plesiomorphic taxa (Achilidae+ Cixiidae+ Achilixiidae+ Derbidae+ Kinnaridae+ Meenoplidae) and five apomorphic taxa (Eurybrachidae+ Gengidae+ Lophopidae tribes Acarnini+ Colpopterini+ Elasmoscelini). Tetdgometridae share the most highly modified antennal type with Ahomocnemiellinae (of Caliscelidae). Weaker evidence suggests that Trypetomorphini (of Tropiduchidae) together with Augilini (of Lophopidae) is the basal lineage of Delphacidae, that Fulgoridae together with Dictyopharidae and Hiraciini (of Tropiduchidae) form a fifth monophyly, and that the remainder of Fulgoroidea are related to Issidae and to Nogodinidae. The most characteristic antennal sensilla type (known since 1890) is shared by seven putative families (Acanaloniidae, Flatidae, Hypochthonellidae, Ricaniidae, Issidae, Lophopidae and Tropiduchidae). The unexpected relationships of Issidae to various other such families support a 1977 hypothesis by Fennah that derive an “issid” wing type in Mithjmna Stal from that of Nogodinidae.
    [Show full text]
  • Hemiptera: Fulgoroidea: Derbidae) from Coquito Palms (Astrocaryum Alatum) in Costa Rica
    Zootaxa 4779 (3): 409–418 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2020 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4779.3.8 http://zoobank.org/urn:lsid:zoobank.org:pub:8EA647FD-FCB7-47F4-9629-4E746499694E A new species of planthopper in the genus Agoo (Hemiptera: Fulgoroidea: Derbidae) from coquito palms (Astrocaryum alatum) in Costa Rica BRIAN W. BAHDER1, MARCO A. ZUMBADO ECHAVARRIA2, EDWIN A. BARRANTES BARRANTES3, GERNOT KUNZ4, ERICKA E. HELMICK5 & CHARLES R. BARTLETT6 1University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. E-mail: [email protected] �[email protected]; https://orcid.org/0000-0002-1118-4832 2Universidad de Costa Rica—Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste car- retera hacia San Pedro, San Ramón, Alajuela, Costa Rica. E-mail: [email protected] �[email protected]; https://orcid.org/0000-0002-2591-7662 3Universidad de Costa Rica—Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste car- retera hacia San Pedro, San Ramón, Alajuela, Costa Rica. E-mail: [email protected] �[email protected]; https://orcid.org/0000-0001-9565-2105 4Karl-Franzens-Universität, Universitätsplatz 2, Zoologie, 8010 Graz, Austria. E-mail: [email protected] �[email protected]; https://orcid.org/0000-0001-7858-0402 5University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA.
    [Show full text]
  • The First Mesozoic Derbidae (Homoptera: Fulgoroidea) from Cretaceous Burmese Amber
    Russian Entomol. J. 29(3): 237–246 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2020 The first Mesozoic Derbidae (Homoptera: Fulgoroidea) from Cretaceous Burmese amber Ïåðâûå ìåçîçîéñêèå Derbidae (Homoptera: Fulgoroidea) èç ìåëîâîãî áèðìàíñêîãî ÿíòàðÿ A.F. Emeljanov1, D.E. Shcherbakov2 À.Ô. Åìåëüÿíîâ1, Ä.Å. Ùåðáàêîâ2 1 Zoological Institute, Russian Academy of Sciences, Universitetskaya Emb. 1, St.-Petersburg 199034, Russia. E-mail: [email protected]. 2 Borissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya St. 123, Moscow, 117647, Russia. E-mail: [email protected]. 1 Зоологический институт РАН, Университетская наб. 1, Санкт-Петербург 199034, Россия. 2 Палеонтологический институт им. А.А. Борисяка РАН, ул. Профсоюзная 123, Москва 117647, Россия. KEY WORDS: planthoppers, Achilidae, sensory pits, phylogeny, fossil, Cenozoic. КЛЮЧЕВЫЕ СЛОВА: носатки, Achilidae, сенсорные ямки, филогения, ископаемые, кайнозой. ABSTRACT. Derbachile hochae gen. et sp.n., D. переднего крыла, узкое лицо, уплощённые усики и aschei sp.n. and Achiderbe obrienae gen. et sp.n. from подусиковый киль. Эта триба отнесена к примитив- mid-Cretaceous Burmese amber described in Derbachil- ному подсемейству, название которого, согласно ini trib.n. are the first Mesozoic and the earliest fossil приоритету, изменено на Breddiniolinae Fennah, 1950, Derbidae. D. hochae sp.n., with 2.6 mm tegmina, is one stat.n. (= Cedusinae Emeljanov, 1992, syn.n.). of the smallest derbids known. Like their modern rela- tives, Burmite derbids could feed on monocots and Derbidae, with more than 160 described genera and woody dicots, and their nymphs could dwell in rotting almost 1,700 species, is the third most diverse planthop- wood and plant debris. Transformation of achilid-like per family after Cixiidae and Delphacidae [Bourgoin, ancestors into typical derbids is discussed.
    [Show full text]
  • Derbid Planthoppers (Hemiptera: Fulgoroidea: Derbidae) Associated with Coconut and Oil Palm in Brazil
    Neotrop Entomol https://doi.org/10.1007/s13744-020-00788-5 SYSTEMATICS, MORPHOLOGY AND PHYSIOLOGY Derbid Planthoppers (Hemiptera: Fulgoroidea: Derbidae) Associated with Coconut and Oil Palm in Brazil 1,2 3 4 5 6 7 8 MDOLLET ,EGFIDELIS ,EDOS PASSOS ,FDA SILVA ,HPABERLENC ,DASCHURT ,BBAHDER , 4 9 LC DINIZ ,CRBARTLETT 1Cirad, Umr Ipme Cirad/Ird/UM, Campus International, de Baillarguet, Montpellier Cedex 5, France 2Embrapa Cenargen, Brasilia, DF, Brasil 3Embrapa Cerrados, Brasilia, DF, Brasil 4Embrapa Tabuleiros Costeiros, Aracaju, SE, Brasil 5Instituto Federal do Amapá, Macapá, AM, Brasil 6Cirad, Umr Bgpi, Montpellier, France 7Embrapa Roraima, Boa Vista, RR, Brasil 8Univ of Florida, Fort Lauderdale, FL, USA 9Univ of Delaware, Newark, DE, USA Keywords Abstract Fulgoromorpha, Auchenorrhyncha, We present surveys of derbid planthoppers associated with coconut Cenchreini, phytoplasmas, lethal yellowing (Cocos nucifera L.) and oil palm (Elaeis guineensis Jacq.) collected in type syndromes, vectors Northeastern (Sergipe) and North (Pará and Roraima) Brazil. The surveys Correspondence were intended to contribute to our knowledge of possible vectors of M Dollet, Embrapa Cenargen, Brasilia, DF, Brasil; [email protected] phytoplasmas or other phloem-restricted plant pathogens. Eight derbid taxa were found, two in the subfamily Cedusinae, tribe Cedusini (Cedusa Edited by Takumasa Kondo – CORPOICA yipara Kramer and C. yowza Kramer) and six in the subfamily Derbinae, Received 9 March 2020 and accepted 25 tribe Cenchreini: Herpis sp., Persis pugnax Stål, Omolicna anastomosa May 2020 (Caldwell), O. nigripennis (Caldwell), and two new species in the genus Agoo Bahder & Bartlett are described here. Genus-level features between * The Author(s) 2020 Omolicna and Agoo are discussed and a key to the species of Agoo is provided.
    [Show full text]
  • Insects of Guam-Ii
    INSECTS OF GUAM-II BERNICE P. BISHOP MUSEUM BULLETIN 189 HONOLULU, HAWAII PUBLISHED BY TH:t Mus:tuM · 1946 Issued DECEMBER20, 1946 FOREWORD "Insects of Guam-II" consists of papers completed after publication of tlie first volume in 1942. About 470 species and varieties are here discussed, of which 94 are described as new. The material is comprised mainly of col­ lections made in Guam by D. T. Fullaway in 1911, and by 0. H. Swezey and R. L. Usinger in 1936. Some groups have not yet been reported on, the most important being the Microlepidoptera, which includes about 75 to 80 species. It seems desirable, however, to print the manuscripts on hand now, rather than to delay until the remainder of the material has been submitted for publication. Appreciation is hereby expressed to those who have participated in the production of this second bulletin on Guam insects. -0. H. SwitzitY. I i J CONTENTS PAGJI Foreword ............................................................................................................................................ Map of Guam ....................................................................................................................................iii ORTHOPTERA AND REI,A'.l'ED ORDERS Orthoptera and related orders of Guam, by 0. H. Swezey.................................................. 3 IsoPTERA Isoptera of Guam, by S. F. Light .............., ............. ,..... ,............................................................ 9 HETEROP'.l'ERA Heteroptera of Guam, by R. L. Usinger .....................,.....................................................
    [Show full text]
  • Making Sense of Fulgoroidea (Hemiptera): New Phylogenetic Evidence K.G
    Cicadina 12: 57-79 (2011) 57 Making sense of Fulgoroidea (Hemiptera): new phylogenetic evidence K.G. Andrew Hamilton1 Abstract: Antennal sensilla in more than 450 genera of all 21 putative families of Fulgoroidea provide characters that correlate well with other characters, including those of nymphal wax filaments and adult behaviour. Together, these characters indicate that Issidae, Lophopidae and Tropiduchidae are polyphyletic taxa, and suggest that Achilidae, Caliscelidae, Cixiidae and Dictyopharidae are paraphyletic. Two strongly supported monophyletic lineages comprise six plesiomorphic taxa (Achilidae+ Cixiidae+ Achilixiidae+ Derbidae+ Kinnaridae+ Meenoplidae) and five apomorphic taxa (Eurybrachidae+ Gengidae+ Lophopidae tribes Acarnini+ Colpopterini+ Elasmoscelini). Tettigometridae share the most highly modified antennal type with Ahomocnemiellinae (of Caliscelidae). Weaker evidence suggests that Trypetomorphini (of Tropiduchidae) together with Augilini (of Lophopidae) is the basal lineage of Delphacidae, that Fulgoridae together with Dictyopharidae and Hiraciini (of Tropiduchidae) form a fifth monophyly, and that the remainder of Fulgoroidea are related to Issidae and to Nogodinidae. The most characteristic antennal sensilla type (known since 1890) is shared by seven putative families (Acanaloniidae, Flatidae, Hypochthonellidae, Ricaniidae, Issidae, Lophopidae and Tropiduchidae). The unexpected relationships of Issidae to various other such families support a 1977 hypothesis by Fennah that derive an “issid” wing type in Mithymna Stål from that of Nogodinidae. It is proposed here that various characteristic wing venation types in Fulgoroidea arose in numerous independent lineages through a few simple genetic modifications that induce or suppress ramification of vein systems during development. Six lineages are identified that could represent more broadly defined families (Cixiidae, Delphacidae, Eurybrachidae, Fulgoridae, Issidae, Tettigometridae) with most other putative families reduced to subfamily status.
    [Show full text]
  • Planthoppers of Delaware (Hemiptera, Fulgoroidea), Excluding Delphacidae, with Species Incidence from Adjacent States
    A peer-reviewed open-access journal ZooKeysPlanthoppers 83: 1–42 (2011) of Delaware (Hemiptera, Fulgoroidea), excluding Delphacidae, with species... 1 doi: 10.3897/zookeys.83.1176 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research Planthoppers of Delaware (Hemiptera, Fulgoroidea), excluding Delphacidae, with species incidence from adjacent States Charles R. Bartlett1, Erin R. Adams1, Anthony T. Gonzon, Jr.2 1 Department of Entomology and Wildlife Ecology, 250 Townsend Hall, University of Delaware, Newark, De- laware, 19717-1303. U.S.A. 2 DE Natural Heritage and Endangered Species Program, DE Division of Fish and Wildlife, DNREC 4876 Hay Point Landing Road, Smyrna, Delaware, 19977. U.S.A. Corresponding author : Charles R. Bartlett ( [email protected] ) Academic editor: Michael Wilson | Received 8 November 2010 | Accepted 29 January 2011 | Published 25 February 2011 Citation: Bartlett CR, Adams ER, Gonzon AT Jr. (2011) Planthoppers of Delaware (Hemiptera, Fulgoroidea), excluding Delphacidae, with species incidence from adjacent States. ZooKeys 83 : 1 – 42 . doi: 10.3897/zookeys.83.1176 Abstract Th e number of species of planthoppers (excluding Delphacidae) known from Delaware is updated from 7 (in 4 families) to 62 species (in 9 families). Specimen abundance is tallied by county and seasonally by two week intervals. Th e Chao1 abundance estimator suggests that the true fauna may be 74 species, al- though species incidence tallied from adjacent states (MD, NJ, PA and DC) suggests that a total fauna of approximately 100 species may be possible. An artifi cial key is presented to genus and select species with photos of most included taxa. Keywords Auchenorrhyncha, Fulgoromorpha, Acanaloniidae, Achilidae, Caliscelidae, Cixiidae, Derbidae, Dicty- opharidae, Flatidae, Issidae, species inventory, generic key Introduction Th e distribution of planthoppers (Hemiptera: Auchenorrhyncha: Fulgoroidea) in the eastern United States was most recently summarized by Wilson and McPherson (1980a).
    [Show full text]
  • A New Species of Planthopper in the Genus Agoo Bahder & Bartlett (Hemiptera: Fulgoroidea: Derbidae) from Coconut Palm (Cocos Nucifera L.) in Jamaica
    Zootaxa 4853 (2): 254–264 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2020 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4853.2.6 http://zoobank.org/urn:lsid:zoobank.org:pub:949733F1-D47A-44B4-8315-29EA3BB0A85F A new species of planthopper in the genus Agoo Bahder & Bartlett (Hemiptera: Fulgoroidea: Derbidae) from coconut palm (Cocos nucifera L.) in Jamaica BRIAN W. BAHDER1, DE-FEN MOU2, CHARLES R. BARTLETT3, ERICKA E. HELMICK4, ASSUNTA BERTACCINI5 & WAYNE MYRIE6 1University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. [email protected]; https://orcid.org/0000-0002-1118-4832 2University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. [email protected]; https://orcid.org/0000-0003-4620-727X 3University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716-2160, USA. [email protected]; https://orcid.org/0000-0001-9428-7337 4University of Florida, Department of Entomology and Nematology—Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314-7719, USA. [email protected]; https://orcid.org/0000-0002-5153-0891 5University of Bologna, Department of Plant Pathology; Viale Fanin 42, 40127, Bologna, Italy. [email protected]; https://orcid.org/0000-0002-5650-1512 6Coconut Industry Board, 18 Waterloo Road, Kingston 10, Jamaica W. I. [email protected]; https://orcid.org/0000-0001-8969-1204 Abstract A new species of the genus Agoo Bahder & Bartlett, Agoo beani sp.
    [Show full text]