An invertebrate survey of Penn Common (concentrating on the wet habitats, especially the springs and flushes) to cover those species and groups that help evaluate its biological importance and help to inform its habitat management under the Higher Level Scheme agreement.

Peter Boardman, Ian Cheeseborough, Nigel Jones, Richard Wright

November 2011 Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

INDEX

i. Summary …………………………………………………………………………….. 3

1. Rationale …………………………………………………………………………………………. 4

2. Methodology…………………………………………………………………………………… 5

3. Results – overview………………………………………………………………………….. 5

4. Results – analysis of invertebrate communities………………………………. 9

5. Results – summary of ISIS analysis…………………………………………………… 10

6. Management Suggestions……………………………………………………………….. 11

7. Discussion……………………………………………………………………………………….. 13

ii. References………………………………………………………………………………. 15

iii. Acknowledgements…………………………………………………………………. 15

Appendix 1 – Full survey data for Penn Common golf course ……………………. 16

Appendix 2 – ISIS outputs for Penn Common …………………………………………….. 31

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Summary

 An invertebrate survey of Penn Common, near Wolverhampton, recorded 457 species of invertebrates during 2011.

 Of these, the vast majority were common species with only 7 species recorded of national or regional importance. These were the flies; Spania nigra, Syntormon macula, Sphaerophoria taeniata, and Herina frondescentiae. The weevil Trachyphloeus asperatus, the spider-hunting wasp Anoplius caviventris, and the solitary wasp Tiphia minuta.

 24 species were noted as new to Staffordshire (V.C. 39).

 Data was put through ISIS software to identify species assemblages for the site. As a whole site, significant species assemblage types (SATs) were identified for scrub edge, rich flower resource, and bark & sapwood decay assemblages. The most significant broad assemblage type (BATs) noted was that for flowing water suggesting the wetland habitats are important at the site. Heathland communities were not significant across the whole site data.

 When data was split down into habitat areas the species assemblage types produced significant results for scrub-heath and moorland species assemblage amongst the dry acid grassland habitats, and the bark and sapwood decay assemblages amongst the aculeate habitats.

 The calcareous flush was found to be of little significance currently.

 Penn Common was found to be regionally important for its aculeate hymenoptera assemblage, though much of this is associated with dead-wood rather than heathland species.

 Management suggestions were made to increase the suitability of habitat for heathland hymenoptera, amongst other habitats across the site.

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

1. Rationale

1.1 An invertebrate survey was carried out on Penn Common (SO898945), near Wolverhampton, to identify the most important invertebrate communities at the site and examine how they compare to other similar sites in the area (Staffordshire and West Midlands primarily but also east Shropshire and north Worcestershire).

1.2 Outputs were to examine how the invertebrate communities can be conserved and, where possible, enhanced (under management through the Higher Level Stewardship (HLS) agreement).

1.3 Penn Common has historically been recognised as an important site in terms of nature conservation with historical botanical records suggesting important habitats (particularly base rich flushes and heathland) for invertebrates at the site. It was noted that grazing had recently ceased but more up to date information regarding habitat quality and suitability for invertebrate assemblages was lacking, hence the need to assess the site through survey work.

1.4 The site hosts Penn Common Golf Course, where golf has been played since 1893 (see map 1), and is intersected roughly down the centre on a north / south plane by a minor road. It sits to the south of Penn, a suburb of the major conurbation of Wolverhampton, and to the west of Sedgley, and is located in Staffordshire (V.C. 39).

Map 1 – Penn Common Golf Course (www.maps.google.co.uk.)

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

2. Methodology

2.1 The survey methodology followed that laid out in ‘Surveying terrestrial and freshwater invertebrates for conservation evaluation’ (Drake et al, 2007) and results were analysed using the ISIS software, developed by Natural England, to assess invertebrate communities at the site.

2.2 The surveyors used appropriate methodologies for their particular invertebrate interests; Pete Boardman & Nigel Jones (diptera – timed sweeping over flushes, damp and marginal habitats, plus netting individual species of interest, water traps, limited Malaise trapping), Ian Cheeseborough (aculeate hymenoptera – netting individual species and observation of nesting aggregations and dead wood habitats, water traps, limited Malaise trapping), Richard Wright (coleoptera – pitfall trapping and sweeping / beating heather and other vegetation, hand searching).

2.3 Where possible other invertebrate and insect orders were recorded when encountered. These included the Lepidoptera which were being surveyed by another contractor so little attention was given to the group during our site visits.

2.4 Visits were timed to assess the peak phenology of the groups studied.

3. Results - overview

3.1 Survey visits were undertaken in April, May, June, July, September and October by the surveyors, after an initial visit in early April accompanied by the course manager, Paul Mills from Penn Common Golf Course.

3.2 Survey work during 2011 was in some ways hampered by the very dry conditions that occurred through the year. A hot, dry spring was followed by a generally dull, dry summer and the dry weather continued on into autumn. Several of the wet or damp areas where survey work was focused were subject to desiccation and at times insects were hard to come across. Nevertheless a total of 457 species were recorded at the site.

3.3 The table below details the breakdown of invertebrate orders recorded;

Order No of Species Diptera (true flies) 220 Coleoptera (beetles) 82 Hymenoptera (bees, wasps and ants) 73 Aranae & Opiliones (spiders and harvestman) 33 Hemiptera (true bugs) 21 Lepidoptera (butterflies and moths) 15 Diplopoda (millipedes) 4 Odonata (damselflies and dragonflies) 4 Isopoda (woodlice) 3 Orthoptera (grasshoppers and crickets) 2 Total - 457 Table 1 – summary of invertebrate orders recorded at Penn Common during the 2011 survey

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

3.4 Of the 457 species recorded few were noted to be of conservation importance except for the following 7 species;

3.5 Diptera (Spaniidae) Spania nigra Meigen, 1830 Liverwort Snipefly. This species is listed as Notable but being small is often overlooked. Larvae leaf mine species of liverworts and Pellia neesia has been noted to host the fly (Stubbs & Drake, 2001). It has a widespread distribution within the United Kingdom (www.data.nbn.org.uk) and appears here at its first Staffordshire site (Jukes, 2005). It has been recorded in Shropshire from dingle woodland near Ironbridge Gorge and at three sites around the Long Mynd in south Shropshire (Shropshire Ecological Data Network pers. comm.)

3.6 Diptera (Dolichopodidae) Syntormon macula Parent, 1927. This uncommon long-legged fly is often associated with dingle woodlands or damp places and has a mostly south-western distribution. It was originally designated a Red Data Book species but is now recognised as more commonly occurring, so was given the lower risk (near threatened) status to reflect this (Falk & Crossley, 2005). It appears here at its first Staffordshire site (Jukes, 2005). It is known from three dingle woodlands in Shropshire (Boardman, 2005 & Shropshire Ecological Data Network pers. comm.).

3.7 Diptera (Syrphidae) Sphaerophoria taeniata (Meigen, 1822). This hoverfly has a mostly southern distribution with Kent across to Somerset seeing the majority of records. It is uncommon outside of this range (www.hoverfly.org.uk), and it appears to be heading north and is recorded here at its first Staffordshire location (Jukes, 2005).

3.8 Diptera (Ulidiidae) Herina frondescentiae (Linnaeus, 1758). This non-tephritid picture-wing fly has a widespread distribution nationally but can be considered local in the Midlands. It is associated with damp grasslands and appears new to Staffordshire at this site (Jukes, 2005).

Herina frondescentiae (photo by Nigel Jones)

3.9 Coleoptera (Curculionidae) Trachyphloeus asperatus Boheman, 1843. This weevil is associated with lowland calcareous grassland and has a widespread distribution in England, with mainly a maritime distribution in Wales. It is classed as Notable A, and is the first Staffordshire record since 1907 (Slawson & Brown, 2009).

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

3.10 Hymenoptera (Pompilidae) Anoplius caviventris (Aurivillius, 1907). This spider-hunting wasp has a mostly southern distribution (www.data.nbn.org) and is classed as Nationally Notable B by Falk (1991) but ‘very rare’ by Michael Archer in his more recent review. It appears here as new to Staffordshire (Webb, Bloxham and Slawson, 2005). It is known from one site in Worcestershire (Trevis, 2004) and a single site in Shropshire (Ian Cheeseborough pers. comm. & Shropshire Ecological Data Network pers. comm.).

3.11 Hymenoptera (Tiphiidae) Tiphia minuta Vander Linden, 1827. This solitary wasp is a parasite of dung beetles (Scarabaeidae) and is considered to be regionally scarce (Jukes, 2004).

3.12 As well as these important species several others were noted as new to Staffordshire when checked against current checklists;

3.13 Diptera;  craneflies (Limoniidae); Dicranomyia affinis, Eloeophila submarmorata, Erioptera squalida.  long-legged flies (Dolichopodidae); Hercostomus parvilamellatus, Teuchophorus calcaratus, T. monacanthus, T. nigricosta, T. simplex,  snail-killing flies (Sciomyzidae); Pherbellia schoenherri, Tetranocera fuscinervis.  flat-footed flies (Platypezidae); Agathomyia antennata  druid flies (Clusiidae); Clusiodes gentilis  stalk flies (Psilidae); Chamaepsila rosae  shore flies (Ephydridae); Ditrichophora calceata  big-headed flies (Pipunculidae); Jassidophaga villosa  lauxaniid flies (Lauxaniidae); Minettia fasciata  parasitic flies (Tachinidae); Lydella stabulans, Vibrissina debilitata  dung flies (Scathophagidae); Nanna flavipes  anthomyid flies (Anthomyiidae); Anthomyia procellaris  muscid flies (Muscidae); Coenosia infantula

3.14 Hemiptera;  leatherbugs (Coreidae); Coreus marginatus (Dock Bug)

3.15 Coleoptera;  ground beetle (Carabidae); Agonum emarginatum

3.16 Hymenoptera;  solitary wasp (Sphecidae); Psenulus concolor

3.17 As well as assessing the site as a whole, it was broken down to the main areas of interest as detailed in the tender document, to begin to assess the invertebrate communities present at the site.

3.18 They were as follows (see map 2);  1 – damp grassland, pond etc at SO89709483  2 – damp, rushy grassland with scrub at SO89629479  3 – species-rich dry grassland at SO895294444

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

 4 – acid grassland / heather at SO89459440  5 – wet woodland / flush with calcareous influence at SO89599442  6 – acid grassland / heather at SO89799476  7 – damp woodland at SO90309478  8 – marsh / wet woodland and wet grassland at SO90379448  8a* – open scrub / woodland with bracken at SO903944  9 – woodland flush / pond / open ditch at SO90259456  9a* –cleared patch with dead wood at SO90269452  10 – flower-rich clearing at SO90269452  11 – damp woodland at SO89639441  12 – eastern footpath / sheltered hedgerow at SO90349456

3.19 Other records away from these areas were noted as ‘general site records’ and given a site centroid grid reference of SO898945.

3.20 Areas 8a and 9a were separated from areas 8 and 9 respectively to take into account good aculeate hymenoptera habitat including dead wood, which would otherwise potentially biased species assemblage analysis in the larger parent areas.

3.21 All records are detailed in Appendix of this report within Table 2.

Map two – compartments survey during Penn Common survey 2011

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

4. Results - Analysis of Communities

4.1 Data was put into the ISIS spreadsheet to identify Species Assembly Types (SATs) and Broad Assemblage Types (BATs) that may exist at Penn Common. It should be noted that approximately 30 species could not be used in the ISIS analysis as they registered an “error”. These species are ones excluded from ISIS as they are ones where sufficient information is known about them, are vagrants, or their habitats do not fit into the classification.

4.2 All SATs and BATs tables used in this analysis are produced in the Appendix of this report.

4.3 Whole site data - ISIS recognised 11 SATs of which three (scrub edge, rich flower resource and bark & sapwood decay) suggested strong species assemblies (favourable condition in CSM). Less strong SATs were those from the dung, open short sward, shingle banks, seepage, reed-fen and pools, scrub-heath and moorland, bare sand and chalk, and heartwood decay assemblies.

4.4 This suggests that the site as a whole has more value for its scrub / woodland edge, associated dead wood, and floral resources than for its heathland or wetland features. This may represent a decline in the state of the habitat but this can only be suggested, rather than proven, without previous evidence of invertebrate survey work at the site and is based upon the perception that features such as the calcareous flush and heathland were perhaps once in better condition due to the historical botanical records that exist.

4.5 Whole site data - 8 BATs were identified, of which the flowing water broad assembly was noted as favourable (in terms of CSM) and achieved the highest rarity score (154). The grassland and scrub matrix achieved the highest BAT species richness score of 154. These figures suggest that within the site as a whole the grassland and scrub matrix and the flushes are key features. Other BATs identified were; unshaded early successional mosaic, permanent wet mire, wood decay, shaded field and ground layer, mineral marsh and open water, arboreal canopy.

4.6 Calcareous flush at SO89599442 (area 5) - ISIS only recognised a single SAT from this data, a single species of the shingle bank assembly.

4.7 Calcareous flush at SO89599442 (area 5) - 5 BATs were identified; grassland and scrub matrix, flowing water, mineral marsh and open water, shaded field and ground layer, and permanent wet mire. No features of either SATs or BATs reached favourable levels and this suggests that this flush is of limited value for invertebrates in its current condition, with few specialists identified. It should be noted that part of this area of the site has only recently been cleared of scrub.

4.8 Wet habitats in the east of the site (areas 8 and 9 – not including 8a and 9a) - ISIS recognised 8 SATs from this area but none of them were strong communities. They were; seepage, open short sward, scrub edge, reed-fen and pools, bark and sapwood decay, rich flower resource, scrub-heath and moorland, bare sand and chalk. 8 BATs were recognised with the flowing water broad assembly noted as favourable.

4.9 Wet habitats in the east of the site (areas 8 and 9 – not including 8a and 9a) - BATs recognised were; grassland and scrub matrix, permanent wet mire, flowing water, mineral marsh and open water, shaded field and ground layer, unshaded early successional mosaic, wood decay and arboreal canopy.

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

4.10 This analysis suggests no strong species assemblages occur within this wetland area, though it does recognise that flowing water is important within the broader assemblage of species and is a key feature of the site.

4.11 Scrubby, flower rich and drier habitats (areas 8a, 9a and 10) - ISIS showed 5 SATs within these areas (mostly targeted for aculeate hymenoptera) and one species assembly type stands out as favourable – the bark and sapwood decay. We had already noted that there was a higher than expected list of dead wood aculeate hymenoptera, considering tree cover is limited, and few old or veteran trees occur at the site, so it was pleasing to see ISIS agreeing with our own interpretation. ISIS suggests 26 species within the bark and sapwood decay assembly. Other SATs listed were; rich flower resource, dung, scrub edge and open short sward.

4.12 Scrubby, flower rich and drier habitats (areas 8a, 9a and 10) - 5 BATs were identified with three standing out; unshaded early successional mosaic, wood decay and grassland and scrub matrix. The two remaining and less strongly represented BATs were arboreal canopy and mineral marsh and open water.

4.13 Dry acid grassland and heather (areas 3, 4 and 6) - ISIS listed 4 SATs within these areas (mostly targeted for coleoptera) and the scrub heath and moorland assembly was noted as favourable with 10 species within the assembly type. The remaining three SATs were bare sand and chalk, scrub edge, and open short sward.

4.14 Dry acid grassland and heather (areas 3, 4 and 6) - 7 BATs were identified with the grassland and scrub matrix assembly standing out far above all others listed. The others were; unshaded early successional mosaic, mineral marsh and open water, shaded field and ground layer, permanent wet mire, flowing water. This suggests that this area of the site on the western side is the most important area of acid grassland and scrubby heath, but bare ground and short sward assemblies are poorly represented.

4.15 Damp grassland (areas 1 and 2) - ISIS was unable to identify any SATs for this habitat.

4.16 Damp grassland (areas 1 and 2) - 5 BATs. Of these the strongest broad assembly was amongst grassland and scrub matrix species. The permanent wet mire broad assembly was not overly strong but nevertheless worthy of mention whilst the mineral marsh and open water, shaded field and ground layer, and flowing water broad assemblies were poorly represented. This data suggests that damp grassland habitats at the site of the least productivity for species assemblages on the site.

5. Results – Summary of ISIS analysis

5.1 Analysis of the species and broad species assemblages of Penn Common using ISIS suggests that the scrub matrix assembly, with associated dead wood, flowing water, dry acid grassland and flower rich grassland are the key current habitat types at the site for invertebrates.

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

5.2 Heathland containing good quality heather and bare open ground seems in too short a supply to host important invertebrate assemblages at present levels. Aculeate hymenoptera assemblages are more reliant upon dead wood than traditionally suitable heathland habitat at the site.

5.3 The calcareous flush in the east of the site is currently not important in terms of species assemblages; through it does host three broad wetland assemblages and a scrub matrix assemblage. This suggests that as the flush scrubbed up it became less important for wetland species, though with recent and continued habitat management may revert to hold more characteristic wetland assemblages of note in the future.

5.4 Other wet habitats at the site were not viewed as strongly important in terms of individual species assemblages but flowing water as a broad assembly was noted as important.

5.5 Damp grassland at the north of the site is of little importance for species assemblages, which may reflect the lack of grazing at the site, or simply the dry weather from 2011.

6. Management Suggestions

6.1 Current management of habitats needs to be continued and increased to maintain a mosaic of shady wet, open wet, flower-rich (low growing and tussocky) grassland, and heathland habitats with heather and bare ground. For example the aculeate hymenoptera at Penn Common need nesting opportunities (sandy bare ground or dead wood), and foraging/hunting grounds in proximity to these nesting sites. Keeping the mix of bare ground, short flower-rich grassland, rank patches of vegetation, scrub and woodland is ideal.

6.2 Wet areas and flushes;

 Probably too many of the wet areas are, or have been until recently, scrubbed up to the detriment of wetland diptera and other groups. These areas should be rotationally cut as has been done in area 5.

 The main passage of water from north of south along the eastern boundary of the site (area 8) is canalised and the grass is mown right up to the edge. Though there is probably not too much that can be done about the canalization given the proximity to the fairway, some grading down the stream banks (to make them more gently sloped) and a more relaxed approach to grass cutting may provide more marginal habitat in this area.

 The pond at the bottom of this ditch is becoming choked with vegetation and would benefit from some clearance.

6.3 Heathland;

 The current managed heather areas on the site are just too small to be of much use as heathland pockets, and are often just a couple of straggly heather plants in the middle of a fairway. Areas should be expanded and could be seeded with heather seeds from a local source.

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

 More bare ground would benefit nesting aggregations of bees. Bare ground is often seen as an erosion problem or unsightly on golf courses and seeded with a grass mixture at the first opportunity. Some decent sized scrapes on marginal areas of the fairways (as large as practicably possible) would provide potentially a great deal of bare ground across the course. This sort of approach has recently been undertaken at Highgate Common for the rare solitary bee Andrena nigrispina. The scrapes were observed on a recent site visit there by two of our surveyors. Several of these bee scrapes are needed around the site to encourage nesting aggregations providing horizontal and vertical surfaces into which bees and wasps can burrow. These can cover a variety of aspects, but are probably the single most important habitat management tool for heathland bees.

 Solitary bees Andrena barbilabris and Crossocerus quadrimaculatus were observed using the bunkers for nesting but only around the unraked sides, therefore whilst bunkers are a key aspect for the golfers at the site, parts of them need not be regulary raked to maintain more breeding habitat for solitary bees. This may be a matter of education of golfers rather than management per se.

6.4 Grassland;

 The amount of acid grassland and species rich grassland should be increased at the site. This could be done by relaxing mowing regimes on the fairways. Instead of a standard lozenge shape of a golf hole, the fairways could be altered so that more of an hour glass shape exists filtering golfers down the middle of the fairway in areas less commonly used on each hole (for instance between the tee and where the average drive from that tee lands).

 Flower rich areas need cutting and raking, with the arisings removed to restrict nutrient input. Cutting of flowering areas needs to be timed so that they are cut after flowering (not during) and with areas left for invertebrates to over-winter in seed heads or tussocks.

 Where thick areas of bracken persist, some rotational cutting to open up the structure would add another dimension to the habitat.

6.5 Dead wood;

 ISIS analysis identified dead wood as a key feature of aculeate hymenoptera at the site, therefore as well as ensuring existing amounts of dead wood are retained, more dead wood should be left available for aculeate hymenoptera and coleoptera to utilise.

 Standing dead wood is more valuable than fallen dead wood. Often standing dead wood is removed as it is deemed to be deleterious to health and safety, however reducing the height of dead wood by removing large branches or the upper half of the tree whilst leaving the trunk still standing to maintain safety is still leaving a valuable resource in place.

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 Standing dead wood could be further improved by drilling small holes into it with a battery powered drill and bit. This method has worked well at sites in Shropshire where it has been adopted and used by a variety of dead wood nesting bees and wasps (Cheeseborough pers. comm.).

 The existing dead wood left on the ground is very quickly covered with vegetation making it unsuitable quickly unsuitable. Fallen dead wood should therefore be kept clear of vegetation or kept higher up of the ground if this is a problem.

7. Discussion

7.1 Penn Common was not considered to be of high conservation status in its current state. The vast majority of species encountered were common in terms of nature conservation interest, though some can be considered heathland, or wetland specialists.

7.2 The strongest species assemblages represented those of scrub edge, flower-rich grassland, and dead wood assemblages of aculeate hymenoptera. Had heathland habitat been in better condition the species assemblages would have reflected this with higher scores in the scrub-heath and bare sand assemblages.

7.3 The calcareous flush was found to be of less importance for wetland species than had been hoped based upon historical botanical records.

7.4 One surprise was the large amount of dead wood aculeate hymenoptera found, considering the lack of established veteran trees.

7.5 Clearly the impact of a very popular local amenity such as the Golf Course undoubtedly has had an effect upon the habitat, though this is not to say the impact of more considered habitat management could improve the existing situation under the Higher Level Scheme.

7.6 It should be noted that our results were affected by the lack of rain through the 2011 season, which led to the drying out of most of the “wet” areas on the site by the end of the 2011 recording season, and clearly had a detrimental effect on the numbers of wetland diptera and coleoptera sampled.

7.7 In terms of the aculeate hymenoptera recorded at the site, the species list achieved during the survey compares favourably with other Staffordshire heathlands, and Penn Common could perhaps be listed as fourth best Staffordshire heathland site (behind Highgate Common, Gentleshaw Common and Bevins Birches) (Jukes, 2004) on its aculeate list.

7.8 It is noted that the better West Midlands heathland sites have decent amounts of bare or sparsely vegetated ground, heather dominated areas and flower-rich heathy verge (Falk, 2010). Penn Common has some flower-rich areas but not enough heather dominated areas and too little bare or sparsely vegetated ground to host the level of aculeate hymenoptera that might improve its standing within the West Midlands league table.

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7.9 As already mentioned the amount of dead wood fauna was unexpected and this seems to be boosting the site aculeate list. This is perhaps masking the lack of heathland specialists found during this survey. Although above sites like Pelsall Common in the league table, Penn Common does however have fewer species of national or regional interest than many other of the key West Midlands heathland sites, with only one very rare species recorded (Anoplius caviventris) and two regionally scarce species (Tiphia minuta and Psenulus concolor*) (Note – Psenulus concolor is viewed as common in Shropshire (Cheeseborough pers.comm.) but is absent from the Staffordshire checklist (Webb, Bloxham & Slawson, 2011), and is also absent from Steven Falk’s West Midlands region review of bee and wasp assemblages (Falk, 2010)). Both Anoplius caviventris and Psenulus concolor appear to be new to Staffordshire. It should be noted that Anoplius caviventris was listed as a general site record during our survey and as such was not included in the ISIS analysis for any individual compartment, though was part of the whole site analysis.

7.10 Most coleoptera noted were common species with only three heathland specialists recorded (Bradycellus ruficollis, Lochmaea suturalis and Micrelus ericae). This again illustrates that good quality heathland habitat is currently at a premium on Penn Common. Approximately 27 species of beetle recorded were wetland specialists, though all were considered common.

7.11 Wetland diptera were well represented but again most species were considered common. As stated earlier the subsequent drying out of the wet areas, as the 2011 recording season progressed, did have a deleterious effect upon our survey and towards the end of the season there were few wetland flies around, particularly craneflies.

7.12 It would be hoped that further habitat management work could improve the wetland areas across the site. ISIS analysis of the species assemblages of the calcareous flush showed how species associated with scrub currently dominate the wetland assemblages. The continued removal of scrub here should begin to show a change in the assemblages over time, as the vegetation responds to more available light.

7.13 Elsewhere, on the eastern side of the site, the flowing water species assemblages were identified by ISIS as of importance and these should at least be maintained.

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References;

Boardman, P.J. (2005); The Red Data Book Invertebrates of Shropshire, a compilation and review of data. Unpublished report for Shropshire Council.

Drake, C.M, Lott, D.A., Alexander, K.N.A, & Webb, J. (2007); Surveying terrestrial and freshwater invertebrates for conservation evaluation. Natural England. Sheffield.

Jukes, A. (2004); An evaluation of the invertebrate fauna of Highgate Common in relation to other sites within Staffordshire and the West Midlands. Unpublished report for Natural England.

Jukes, A. (2005); A provisional atlas of the True Flies (including hoverflies) (Diptera) of Staffordshire. Staffordshire Ecological Record (web version)

Falk, S.J. (1991); A review of the scarce and threatened bees, wasps and ants of Great Britain. JNCC. Peterborough.

Falk, S.J. (2010); The bee and wasp assemblages (Hymenoptera: Aculeata) of some key heathland sites in the West Midlands region. British Journal of Entomology and Natural History. Vol. 23 p77-97. BENHS. Dinton.

Falk, S.J. & Crossley, R. (2005); A review of the scarce and threatened flies of Great Britain. JNCC. Peterborough.

Slawson, C., & Brown, E. (2009); A provisional atlas of the Beetles (Coleoptera) of Staffordshire. Staffordshire Ecological Record (web version).

Stubbs, A.E., & Drake, C.M., (2001); British Soldierflies and their allies: an illustrated guide to their identification and ecology. BENHS. Dinton Pastures.

Trevis, G. (2004); Aculeate Hymenoptera in Worcestershire. Worcestershire Record No.16 p27-28

Webb, J., Bloxham, M. & Slawson C. (2005); A provisional atlas of the Bees and Wasps (Hymenoptera) of Staffordshire. Staffordshire Ecological Record (web version).

Websites www.data.nbn.org.uk – The NBN Gateway www.hoverfly.org.uk – The Hoverfly Recording Scheme www.staffs-ecology.org.uk – Staffordshire Ecological Record

Acknowledgements

The authors would like to thank Paul Mills at Penn Common Golf Course for access to the course and showing us around on our first visit, and Jon Webb at Natural England for supplying ISIS. We would also like to thank for golfers who were playing the course during our visits and fully expect that our attendance was blamed for more than one bad shot or lost ball.

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

APPENDIX 1 – Full survey data for Penn Common Golf Course

Order Taxon Compartment Grid ref Date Determiner Araneae Alopecosa pulverulenta 3 SO89529444 2011 Richard Wright Araneae Alopecosa pulverulenta 4 SO89459440 2011 Richard Wright Araneae Antistea elegans 5 SO89599442 2011 Richard Wright Araneae Antistea elegans 8 SO90379448 2011 Richard Wright Araneae Bathyphantes gracilis 6 SO89799476 2011 Richard Wright Araneae Centromerita bicolor 2 SO89629479 2011 Richard Wright Araneae Centromerita bicolor 6 SO89799476 2011 Richard Wright Araneae Centromerita bicolor 8 SO90379448 2011 Richard Wright Araneae Drassyllus pusillus 3 SO89529444 2011 Richard Wright Araneae Drassyllus pusillus 7 SO90309478 2011 Richard Wright Araneae Erigone atra 1 SO89709483 2011 Richard Wright Araneae Erigone dentipalpis 3 SO89529444 2011 Richard Wright Araneae Erigone dentipalpis 4 SO89459440 2011 Richard Wright Araneae Gongylidiellum vivum 5 SO89599442 2011 Richard Wright Araneae Gongylidiellum vivum 8 SO90379448 2011 Richard Wright Araneae Lepthyphantes tenuis 2 SO89629479 2011 Richard Wright Araneae Lepthyphantes tenuis 3 SO89529444 2011 Richard Wright Araneae Lepthyphantes tenuis 4 SO89459440 2011 Richard Wright Araneae Lepthyphantes tenuis 8 SO90379448 2011 Richard Wright Araneae Leptorhoptrum robustum 4 SO89459440 2011 Richard Wright Araneae Leptorhoptrum robustum 6 SO89799476 2011 Richard Wright Araneae Meta mengei 1 SO89709483 2011 Richard Wright Araneae Meta segmentata 2 SO89629479 2011 Richard Wright Araneae Micaria pulicaria 3 SO89529444 2011 Richard Wright Araneae Micaria pulicaria 4 SO89459440 2011 Richard Wright Araneae Micaria pulicaria 8 SO90379448 2011 Richard Wright Araneae Micrargus herbigradus 7 SO90309478 2011 Richard Wright Araneae Neottiura bimaculata 4 SO89459440 2011 Richard Wright Araneae Oedothorax gibbosus 1 SO89709483 2011 Richard Wright Araneae Oedothorax gibbosus 2 SO89629479 2011 Richard Wright Araneae Oedothorax gibbosus 6 SO89799476 2011 Richard Wright Araneae Oedothorax gibbosus 8 SO90379448 2011 Richard Wright Araneae Oedothorax gibbosus 9 SO90259456 2011 Richard Wright Araneae Oedothorax retusus 1 SO89709483 2011 Richard Wright Araneae Oedothorax retusus 5 SO89599442 2011 Richard Wright Araneae Pachygnatha clercki 1 SO89709483 2011 Richard Wright Araneae Pachygnatha clercki 2 SO89629479 2011 Richard Wright Araneae Pachygnatha clercki 7 SO90309478 2011 Richard Wright Araneae Pachygnatha clercki 9 SO90259456 2011 Richard Wright Araneae Pachygnatha degeeri 1 SO89709483 2011 Richard Wright Araneae Pachygnatha degeeri 2 SO89629479 2011 Richard Wright Araneae Pachygnatha degeeri 3 SO89529444 2011 Richard Wright Araneae Pachygnatha degeeri 4 SO89459440 2011 Richard Wright Araneae Pardosa nigriceps 3 SO89529444 2011 Richard Wright Araneae Pardosa nigriceps 6 SO89799476 2011 Richard Wright Araneae Pardosa palustris 4 SO89459440 2011 Richard Wright Araneae Pardosa prativaga 1 SO89709483 2011 Richard Wright Araneae Pardosa prativaga 3 SO89529444 2011 Richard Wright Araneae Pardosa prativaga 4 SO89459440 2011 Richard Wright Araneae Pardosa prativaga 6 SO89799476 2011 Richard Wright Araneae Pardosa prativaga 7 SO90309478 2011 Richard Wright Araneae Pardosa prativaga 9 SO90259456 2011 Richard Wright Araneae Pardosa pullata 1 SO89709483 2011 Richard Wright Araneae Pardosa pullata 2 SO89629479 2011 Richard Wright Araneae Pardosa pullata 4 SO89459440 2011 Richard Wright

Page 16

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Araneae Pardosa pullata 6 SO89799476 2011 Richard Wright Araneae Pardosa pullata 8 SO90379448 2011 Richard Wright Araneae Pardosa pullata 9 SO90259456 2011 Richard Wright Araneae Pirata hygrophilus 9 SO90259456 2011 Richard Wright Araneae Pirata piraticus 1 SO89709483 2011 Richard Wright Araneae Pirata piraticus 5 SO89599442 2011 Richard Wright Araneae Pirata piraticus 8 SO90379448 2011 Richard Wright Araneae Pocadicnemis pumila 5 SO89599442 2011 Richard Wright Araneae Pocadicnemis pumila 6 SO89799476 2011 Richard Wright Araneae Robertus lividus 4 SO89459440 2011 Richard Wright Araneae Robertus lividus 6 SO89799476 2011 Richard Wright Araneae Walckenaeria vigilax 5 SO89599442 2011 Richard Wright Araneae Walckenaeria vigilax 9 SO90259456 2011 Richard Wright Araneae Xysticus cristatus 3 SO89529444 2011 Richard Wright Araneae Xysticus cristatus 4 SO89459440 2011 Richard Wright Araneae Xysticus cristatus 6 SO89799476 2011 Richard Wright Coleoptera Acupalpus dubius 1 SO89709483 2011 Richard Wright Coleoptera Acupalpus dubius 2 SO89629479 2011 Richard Wright Coleoptera Acupalpus dubius 5 SO89599442 2011 Richard Wright Coleoptera Acupalpus dubius 6 SO89799476 2011 Richard Wright Coleoptera Acupalpus dubius 7 SO90309478 2011 Richard Wright Coleoptera Acupalpus dubius 8 SO90379448 2011 Richard Wright Coleoptera Acupalpus dubius 9 SO90259456 2011 Richard Wright Coleoptera Adalia bipunctata site record SO898945 20/05/2011 Nigel Jones Coleoptera Adalia bipunctata 9 SO90259456 29/06/2011 Pete Boardman Coleoptera Agonum emarginatum 5 SO89599442 2011 Richard Wright Coleoptera Agonum emarginatum 8 SO90379448 2011 Richard Wright Coleoptera Agonum emarginatum 9 SO90259456 2011 Richard Wright Coleoptera Agonum fuliginosum 5 SO89599442 2011 Richard Wright Coleoptera Agonum fuliginosum 7 SO90309478 2011 Richard Wright Coleoptera Agonum muelleri 8 SO90379448 2011 Richard Wright Coleoptera Agonum viduum 5 SO89599442 2011 Richard Wright Coleoptera Agonum viduum 8 SO90379448 2011 Richard Wright Coleoptera Agriotes acuminatus 3 SO89529444 2011 Richard Wright Coleoptera Agriotes lineatus 2 SO89629479 2011 Richard Wright Coleoptera Agriotes pallidulus 3 SO89529444 2011 Richard Wright Coleoptera Agriotes sputator 4 SO89459440 2011 Richard Wright Coleoptera Amara communis 3 SO89529444 2011 Richard Wright Coleoptera Amara communis 4 SO89459440 2011 Richard Wright Coleoptera Amara communis 6 SO89799476 2011 Richard Wright Coleoptera Amara lunicollis 3 SO89529444 2011 Richard Wright Coleoptera Amara lunicollis 4 SO89459440 2011 Richard Wright Coleoptera Amara lunicollis 6 SO89799476 2011 Richard Wright Coleoptera Amara plebeja 1 SO89709483 2011 Richard Wright Coleoptera Amara plebeja 2 SO89629479 2011 Richard Wright Coleoptera Amara plebeja 5 SO89599442 2011 Richard Wright Coleoptera Amara plebeja 8 SO90379448 2011 Richard Wright Coleoptera Amara plebeja 9 SO90259456 2011 Richard Wright Coleoptera Anacaena globulus 1 SO89709483 2011 Richard Wright Coleoptera Anacaena globulus 5 SO89599442 2011 Richard Wright Coleoptera Anacaena globulus 6 SO89799476 2011 Richard Wright Coleoptera Anacaena globulus 8 SO90379448 2011 Richard Wright Coleoptera Anacaena globulus 9 SO90259456 2011 Richard Wright Coleoptera Anotylus rugosus 1 SO89709483 2011 Richard Wright Coleoptera Anotylus rugosus 2 SO89629479 2011 Richard Wright Coleoptera Anotylus rugosus 3 SO89529444 2011 Richard Wright Coleoptera Anotylus rugosus 4 SO89459440 2011 Richard Wright Coleoptera Anotylus rugosus 6 SO89799476 2011 Richard Wright Coleoptera Anotylus rugosus 7 SO90309478 2011 Richard Wright Page 17

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Coleoptera Anotylus rugosus 9 SO90259456 2011 Richard Wright Coleoptera Anthobium unicolor 2 SO89629479 2011 Richard Wright Coleoptera Anthobium unicolor 7 SO90309478 2011 Richard Wright Coleoptera Aphodius prodromus 8 SO90379448 2011 Richard Wright Coleoptera Aphthona euphorbiae 3 SO89529444 2011 Richard Wright Coleoptera Apion frumentarium 5 SO89599442 20/04/2011 Pete Boardman Coleoptera Apion frumentarium 7 SO90309478 20/04/2011 Pete Boardman Coleoptera Athous haemorrhoidalis 7 SO90309478 2011 Richard Wright Coleoptera Barypeithes pellucidus 2 SO89629479 2011 Richard Wright Coleoptera Barypeithes pellucidus 3 SO89529444 2011 Richard Wright Coleoptera Barypeithes pellucidus 4 SO89459440 2011 Richard Wright Coleoptera Barypeithes pellucidus 7 SO90309478 2011 Richard Wright Coleoptera Bembidion decorum 5 SO89599442 2011 Richard Wright Coleoptera Bembidion guttula 5 SO89599442 2011 Richard Wright Coleoptera Bembidion guttula 6 SO89799476 2011 Richard Wright Coleoptera Bembidion lampros 3 SO89529444 2011 Richard Wright Coleoptera Bembidion lampros 4 SO89459440 2011 Richard Wright Coleoptera Bembidion lampros 9 SO90259456 2011 Richard Wright Coleoptera Bembidion mannerheimi 1 SO89709483 2011 Richard Wright Coleoptera Bembidion mannerheimi 2 SO89629479 2011 Richard Wright Coleoptera Bembidion mannerheimi 6 SO89799476 2011 Richard Wright Coleoptera Bembidion mannerheimi 7 SO90309478 2011 Richard Wright Coleoptera Bembidion mannerheimi 8 SO90379448 2011 Richard Wright Coleoptera Bembidion mannerheimi 9 SO90259456 2011 Richard Wright Coleoptera Bembidion obtusum 1 SO89709483 2011 Richard Wright Coleoptera Bembidion obtusum 3 SO89529444 2011 Richard Wright Coleoptera Bembidion obtusum 4 SO89459440 2011 Richard Wright Coleoptera Bembidion obtusum 6 SO89799476 2011 Richard Wright Coleoptera Bembidion obtusum 9 SO90259456 2011 Richard Wright Coleoptera Brachygluta haematica 4 SO89459440 2011 Richard Wright Coleoptera Brachygluta haematica 8 SO90379448 2011 Richard Wright Coleoptera Bradycellus ruficollis 4 SO89459440 2011 Richard Wright Coleoptera Bradycellus ruficollis 6 SO89799476 2011 Richard Wright Coleoptera Bradycellus verbasci 4 SO89459440 2011 Richard Wright Coleoptera Calvia quattuordecimguttata 7 SO90029443 20/04/2011 Pete Boardman Coleoptera Calvia quattuordecimguttata site record SO898945 20/05/2011 Nigel Jones Coleoptera Cantharis nigra 3 SO89529444 2011 Richard Wright Coleoptera Cassida rubiginosa 3 SO89529444 2011 Richard Wright Coleoptera Ceratapion gibbirostre 3 SO89529444 2011 Richard Wright Coleoptera Chaetocnema concinna 3 SO89529444 2011 Richard Wright Coleoptera Chaetocnema concinna 4 SO89459440 2011 Richard Wright Coleoptera Chaetocnema hortensis 3 SO89529444 2011 Richard Wright Coleoptera Chilocorus renipustulatus site record SO898945 20/05/2011 Nigel Jones Coleoptera Clivina fossor 8 SO90379448 2011 Richard Wright Coleoptera Clytus arietis 12 SO90349456 18/04/2011 Ian Cheeseborough Coleoptera Clytus arietis 8 SO90379448 20/05/2011 Nigel Jones Coleoptera Coccidula rufa 1 SO89709483 2011 Richard Wright Coleoptera Coccidula rufa 5 SO89599442 2011 Richard Wright Coleoptera Coccinella septempunctata 4 SO89459440 2011 Richard Wright Coleoptera Coccinella septempunctata site record SO899945 18/04/2011 Ian Cheeseborough Coleoptera Coccinella septempunctata 7 SO90029443 20/04/2011 Pete Boardman Coleoptera Coccinella septempunctata 7 SO90309478 20/04/2011 Pete Boardman Coleoptera Coccinella septempunctata site record SO898945 20/05/2011 Nigel Jones Coleoptera Corticarina minuta 3 SO89529444 2011 Richard Wright Coleoptera Cortinicara gibbosa 4 SO89459440 2011 Richard Wright Coleoptera Cortinicara gibbosa 6 SO89799476 2011 Richard Wright Coleoptera Cortinicara gibbosa 7 SO90309478 2011 Richard Wright Coleoptera Dryops ernesti 1 SO89709483 2011 Richard Wright Coleoptera Dryops ernesti 5 SO89599442 2011 Richard Wright Page 18

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Coleoptera Dryops ernesti 8 SO90379448 2011 Richard Wright Coleoptera Dryops ernesti 9 SO90259456 2011 Richard Wright Coleoptera Eutrichapion ervi 3 SO89529444 2011 Richard Wright Coleoptera Eutrichapion viciae 3 SO89529444 2011 Richard Wright Coleoptera Exochomus quadripustulatus 7 SO90309478 2011 Richard Wright Coleoptera Gabrius breviventer 5 SO89599442 2011 Richard Wright Coleoptera Gabrius breviventer 8 SO90379448 2011 Richard Wright Coleoptera Gymnetron pascuorum 3 SO89529444 2011 Richard Wright Coleoptera Gymnetron pascuorum 4 SO89459440 2011 Richard Wright Coleoptera Harmonia axyridis site record SO898945 20/05/2011 Nigel Jones Coleoptera Lathrobium geminum 5 SO89599442 2011 Richard Wright Coleoptera Lathrobium geminum 8 SO90379448 2011 Richard Wright Coleoptera Leiosoma deflexum 3 SO89529444 2011 Richard Wright Coleoptera Lesteva sicula 5 SO89599442 2011 Richard Wright Coleoptera Lesteva sicula 8 SO90379448 2011 Richard Wright Coleoptera Lochmaea suturalis 4 SO89459440 2011 Richard Wright Coleoptera Lochmaea suturalis 6 SO89799476 2011 Richard Wright Coleoptera Malthinus flaveolus 8 SO90379448 29/06/2011 Pete Boardman Coleoptera Mecinus pyraster 3 SO89529444 2011 Richard Wright Coleoptera Micrelus ericae 4 SO89459440 2011 Richard Wright Coleoptera Micrelus ericae 6 SO89799476 2011 Richard Wright Coleoptera Nargus velox 4 SO89459440 2011 Richard Wright Coleoptera Nargus velox 8 SO90379448 2011 Richard Wright Coleoptera Nargus velox 9 SO90259456 2011 Richard Wright Coleoptera Nebria brevicollis 3 SO89529444 2011 Richard Wright Coleoptera Nebria brevicollis 4 SO89459440 2011 Richard Wright Coleoptera Nebria brevicollis 9 SO90259456 2011 Richard Wright Coleoptera Neocrepidodera ferruginea 9 SO90259456 2011 Richard Wright Coleoptera Neocrepidodera transversa 3 SO89529444 2011 Richard Wright Coleoptera Notaris acridulus 5 SO89599442 2011 Richard Wright Coleoptera Notaris acridulus 8 SO90379448 2011 Richard Wright Coleoptera Ochthebius minimus 9 SO90259456 2011 Richard Wright Coleoptera Oedemera nobilis 9a SO90269452 20/05/2011 Ian Cheeseborough Coleoptera Olibrus aeneus 5 SO89599442 2011 Richard Wright Coleoptera Oxystoma craccae 3 SO89529444 2011 Richard Wright Coleoptera Paradromius linearis 3 SO89529444 2011 Richard Wright Coleoptera Perapion curtirostre 1 SO89709483 2011 Richard Wright Coleoptera Perapion curtirostre 3 SO89529444 2011 Richard Wright Coleoptera Perapion curtirostre 9 SO90259456 2011 Richard Wright Coleoptera Perapion hydrolapathi 1 SO89709483 2011 Richard Wright Coleoptera Philonthus cognatus 1 SO89709483 2011 Richard Wright Coleoptera Philonthus cognatus 2 SO89629479 2011 Richard Wright Coleoptera Philonthus cognatus 3 SO89529444 2011 Richard Wright Coleoptera Philonthus cognatus 4 SO89459440 2011 Richard Wright Coleoptera Philonthus cognatus 8 SO90379448 2011 Richard Wright Coleoptera Phyllobius pomaceus 9 SO90259456 20/04/2011 Pete Boardman Coleoptera Phyllobius roboretanus 7 SO90309478 2011 Richard Wright Coleoptera Phyllotreta tetrastigma 5 SO89599442 2011 Richard Wright Coleoptera Phyllotreta tetrastigma 6 SO89799476 2011 Richard Wright Coleoptera Phyllotreta tetrastigma 8 SO90379448 2011 Richard Wright Coleoptera Poecilus versicolor 3 SO89529444 2011 Richard Wright Coleoptera Propylea quattuordecimpunctata 7 SO90309478 2011 Richard Wright Coleoptera Propylea quattuordecimpunctata site record SO899945 18/04/2011 Ian Cheeseborough Coleoptera Propylea quattuordecimpunctata 8a SO903945 28/07/2011 Ian Cheeseborough Coleoptera Protapion assimile 3 SO89529444 2011 Richard Wright Coleoptera Protapion fulvipes 3 SO89529444 2011 Richard Wright Coleoptera Protapion trifolii 3 SO89529444 2011 Richard Wright Coleoptera Psyllobora vigintiduopunctata 3 SO89529444 2011 Richard Wright Coleoptera Pterostichus diligens 5 SO89599442 2011 Richard Wright Page 19

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Coleoptera Pterostichus diligens 9 SO90259456 2011 Richard Wright Coleoptera Pterostichus madidus 3 SO89529444 2011 Richard Wright Coleoptera Pterostichus melanarius 1 SO89709483 2011 Richard Wright Coleoptera Pterostichus melanarius 2 SO89629479 2011 Richard Wright Coleoptera Pterostichus melanarius 8 SO90379448 2011 Richard Wright Coleoptera Pterostichus melanarius 9 SO90259456 2011 Richard Wright Coleoptera Pterostichus nigrita 5 SO89599442 2011 Richard Wright Coleoptera Pterostichus nigrita 7 SO90309478 2011 Richard Wright Coleoptera Pterostichus nigrita 8 SO90379448 2011 Richard Wright Coleoptera Pterostichus nigrita 9 SO90259456 2011 Richard Wright Coleoptera Pterostichus strenuus 1 SO89709483 2011 Richard Wright Coleoptera Pterostichus strenuus 2 SO89629479 2011 Richard Wright Coleoptera Pterostichus strenuus 4 SO89459440 2011 Richard Wright Coleoptera Pterostichus strenuus 5 SO89599442 2011 Richard Wright Coleoptera Pterostichus strenuus 6 SO89799476 2011 Richard Wright Coleoptera Pterostichus strenuus 7 SO90309478 2011 Richard Wright Coleoptera Pterostichus strenuus 8 SO90379448 2011 Richard Wright Coleoptera Pterostichus strenuus 9 SO90259456 2011 Richard Wright Coleoptera Pterostichus vernalis 5 SO89599442 2011 Richard Wright Coleoptera Pterostichus vernalis 8 SO90379448 2011 Richard Wright Coleoptera Pterostichus vernalis 9 SO90259456 2011 Richard Wright Coleoptera Quedius curtipennis 1 SO89709483 2011 Richard Wright Coleoptera Quedius curtipennis 2 SO89629479 2011 Richard Wright Coleoptera Quedius curtipennis 3 SO89529444 2011 Richard Wright Coleoptera Quedius curtipennis 6 SO89799476 2011 Richard Wright Coleoptera Quedius curtipennis 7 SO90309478 2011 Richard Wright Coleoptera Quedius fuliginosus 4 SO89459440 2011 Richard Wright Coleoptera Quedius fuliginosus 9 SO90259456 2011 Richard Wright Coleoptera Quedius maurorufus 5 SO89599442 2011 Richard Wright Coleoptera Quedius maurorufus 6 SO89799476 2011 Richard Wright Coleoptera Quedius maurorufus 9 SO90259456 2011 Richard Wright Coleoptera Quedius molochinus 7 SO90309478 2011 Richard Wright Coleoptera Rhagonycha fulva 3 SO89529444 2011 Richard Wright Coleoptera Rhagonycha limbata 7 SO90309478 2011 Richard Wright Coleoptera Rhyzobius litura 1 SO89709483 2011 Richard Wright Coleoptera Rhyzobius litura 2 SO89629479 2011 Richard Wright Coleoptera Rhyzobius litura 3 SO89529444 2011 Richard Wright Coleoptera Rhyzobius litura 5 SO89599442 2011 Richard Wright Coleoptera Rugilus rufipes 2 SO89629479 2011 Richard Wright Coleoptera Rutpela maculata 9a SO90269452 20/06/2011 Ian Cheeseborough Coleoptera Sitona cambricus 2 SO89629479 2011 Richard Wright Coleoptera Sitona lineatus 3 SO89529444 2011 Richard Wright Coleoptera Sitona lineatus 9 SO90259456 2011 Richard Wright Coleoptera Sphaeroderma testaceum 3 SO89529444 2011 Richard Wright Coleoptera Sphaeroderma testaceum 8 SO90379448 2011 Richard Wright Coleoptera Stenus clavicornis 1 SO89709483 2011 Richard Wright Coleoptera Stenus clavicornis 2 SO89629479 2011 Richard Wright Coleoptera Stenus clavicornis 5 SO89599442 2011 Richard Wright Coleoptera Stenus clavicornis 6 SO89799476 2011 Richard Wright Coleoptera Stenus clavicornis 7 SO90309478 2011 Richard Wright Coleoptera Stenus clavicornis 8 SO90379448 2011 Richard Wright Coleoptera Stenus clavicornis 9 SO90259456 2011 Richard Wright Coleoptera Stenus flavipes 2 SO89629479 2011 Richard Wright Coleoptera Stenus flavipes 5 SO89599442 2011 Richard Wright Coleoptera Stenus providus 5 SO89599442 2011 Richard Wright Coleoptera Sunius propinquus 3 SO89529444 2011 Richard Wright Coleoptera Sunius propinquus 6 SO89799476 2011 Richard Wright Coleoptera Tachinus rufipes 2 SO89629479 2011 Richard Wright Coleoptera Tachinus rufipes 3 SO89529444 2011 Richard Wright Page 20

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Coleoptera Tachinus rufipes 4 SO89459440 2011 Richard Wright Coleoptera Tachinus rufipes 7 SO90309478 2011 Richard Wright Coleoptera Tachyporus chrysomelinus 1 SO89709483 2011 Richard Wright Coleoptera Tachyporus chrysomelinus 3 SO89529444 2011 Richard Wright Coleoptera Tachyporus chrysomelinus 6 SO89799476 2011 Richard Wright Coleoptera Tachyporus dispar 5 SO89599442 2011 Richard Wright Coleoptera Tachyporus dispar 8 SO90379448 2011 Richard Wright Coleoptera Tachyporus dispar 9 SO90259456 2011 Richard Wright Coleoptera Tachyporus hypnorum 1 SO89709483 2011 Richard Wright Coleoptera Tachyporus hypnorum 2 SO89629479 2011 Richard Wright Coleoptera Tachyporus hypnorum 3 SO89529444 2011 Richard Wright Coleoptera Tachyporus hypnorum 4 SO89459440 2011 Richard Wright Coleoptera Tachyporus hypnorum 5 SO89599442 2011 Richard Wright Coleoptera Tachyporus hypnorum 6 SO89799476 2011 Richard Wright Coleoptera Tachyporus hypnorum 7 SO90309478 2011 Richard Wright Coleoptera Tachyporus hypnorum 9 SO90259456 2011 Richard Wright Coleoptera Tachyporus solutus 2 SO89629479 2011 Richard Wright Coleoptera Tachyporus solutus 7 SO90309478 2011 Richard Wright Coleoptera Tachyporus solutus 9 SO90259456 2011 Richard Wright Coleoptera Trachyphloeus asperatus 4 SO89459440 2011 Richard Wright Coleoptera Trachyphloeus asperatus 6 SO89799476 2011 Richard Wright Coleoptera Trixagus dermestoides 3 SO89529444 2011 Richard Wright Coleoptera Trixagus dermestoides 4 SO89459440 2011 Richard Wright Coleoptera Trixagus dermestoides 6 SO89799476 2011 Richard Wright Coleoptera Xantholinus linearis 3 SO89529444 2011 Richard Wright Coleoptera Xantholinus longiventris 2 SO89629479 2011 Richard Wright Coleoptera Xantholinus longiventris 8 SO90379448 2011 Richard Wright Diplopoda Cylindroiulus punctatus 6 SO89799476 2011 Richard Wright Diplopoda Cylindroiulus punctatus 7 SO90309478 2011 Richard Wright Diplopoda Ophyiulus pilosus 5 SO89599442 2011 Richard Wright Diplopoda Ophyiulus pilosus 6 SO89799476 2011 Richard Wright Diplopoda Ophyiulus pilosus 7 SO90309478 2011 Richard Wright Diplopoda Ophyiulus pilosus 8 SO90379448 2011 Richard Wright Diplopoda Polydesmus angustus 1 SO89709483 2011 Richard Wright Diplopoda Polydesmus angustus 2 SO89629479 2011 Richard Wright Diplopoda Polydesmus angustus 4 SO89459440 2011 Richard Wright Diplopoda Polydesmus angustus 5 SO89599442 2011 Richard Wright Diplopoda Polydesmus angustus 8 SO90379448 2011 Richard Wright Diplopoda Tachypodoiulus niger 1 SO89709483 2011 Richard Wright Diplopoda Tachypodoiulus niger 2 SO89629479 2011 Richard Wright Diplopoda Tachypodoiulus niger 4 SO89459440 2011 Richard Wright Diplopoda Tachypodoiulus niger 5 SO89599442 2011 Richard Wright Diplopoda Tachypodoiulus niger 6 SO89799476 2011 Richard Wright Diplopoda Tachypodoiulus niger 7 SO90309478 2011 Richard Wright Diplopoda Tachypodoiulus niger 8 SO90379448 2011 Richard Wright Diplopoda Tachypodoiulus niger 9 SO90259456 2011 Richard Wright Diptera Agathomyia antennata 8 SO90379448 20/05/2011 Nigel Jones Diptera Anomoia purmunda site record SO898945 28/07/2011 Nigel Jones Diptera Anthomyia procellaris 9 SO90259456 20/05/2011 Nigel Jones Diptera Argyra argentina 8 SO90379448 28/07/2011 Nigel Jones Diptera Beris chalybata 5 SO89599442 20/05/2011 Nigel Jones Diptera Beris chalybata 7 SO90309478 20/05/2011 Nigel Jones Diptera Beris morrisii 8 SO90379448 28/07/2011 Nigel Jones Diptera Beris vallata 8 SO90379448 28/07/2011 Nigel Jones Diptera Bibio johannis 1 SO89709483 18/04/2011 Nigel Jones Diptera Bibio johannis 5 SO89599442 18/04/2011 Nigel Jones Diptera Bibio marci site record SO898945 18/04/2011 Nigel Jones Diptera Bibio marci 7 SO90029443 20/04/2011 Pete Boardman Diptera Neolimnomyia nemoralis 5 SO89599442 20/05/2011 Pete Boardman Page 21

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Diptera Neolimnomyia nemoralis 8 SO90379448 20/05/2011 Pete Boardman Diptera Neolimnomyia nemoralis 9 SO90259456 29/06/2011 Pete Boardman Diptera Campsicnemus curvipes 8 SO90379448 18/04/2011 Nigel Jones Diptera Campsicnemus curvipes 8 SO90379448 20/05/2011 Nigel Jones Diptera Campsicnemus curvipes 8 SO90379448 28/07/2011 Nigel Jones Diptera Chaetostomella cylindrica 1 SO89709483 20/05/2011 Nigel Jones Diptera Chaetostomella cylindrica site record SO898945 28/07/2011 Nigel Jones Diptera Chamaepsila rosae 5 SO89599442 20/05/2011 Nigel Jones Diptera Chamaepsila rosae 8 SO90379448 20/05/2011 Nigel Jones Diptera Cheilosia albitarsis 9 SO90259456 20/05/2011 Nigel Jones Diptera Cheilosia illustrata 9a SO90269452 20/06/2011 Ian Cheeseborough Diptera Cheilosia illustrata site record SO898945 28/07/2011 Nigel Jones Diptera Cheilotrichia cinerascens 5 SO89599442 20/04/2011 Pete Boardman Diptera Cheilotrichia cinerascens 7 SO90029443 20/04/2011 Pete Boardman Diptera Cheilotrichia cinerascens 9 SO90259456 20/04/2011 Pete Boardman Diptera Cheilotrichia cinerascens 7 SO90309478 20/05/2011 Pete Boardman Diptera Chelifera precatoria 5 SO89599442 20/05/2011 Nigel Jones Diptera Chrysogaster solstitialis site record SO898945 28/07/2011 Nigel Jones Diptera Chrysopilus cristatus 5 SO89599442 20/05/2011 Nigel Jones Diptera Chrysopilus cristatus 8 SO90379448 20/05/2011 Nigel Jones Diptera Chrysopilus cristatus 9 SO90259456 29/06/2011 Pete Boardman Diptera Chrysopilus cristatus 9 SO90259456 29/06/2011 Pete Boardman Diptera Chrysotoxum festivum site record SO898945 28/07/2011 Nigel Jones Diptera Chrysotus gramineus site record SO898945 28/07/2011 Nigel Jones Diptera Cleigastra apicalis 9 SO90259456 18/04/2011 Nigel Jones Diptera Clusiodes albimanus 8 SO90379448 28/07/2011 Nigel Jones Diptera Clusiodes gentilis 8 SO90379448 20/05/2011 Nigel Jones Diptera Coenia palustris 8 SO90379448 28/07/2011 Nigel Jones Diptera Coenosia femoratus site record SO898945 28/07/2011 Nigel Jones Diptera Coenosia infantula site record SO898945 28/07/2011 Nigel Jones Diptera Coenosia tigrina site record SO898945 20/05/2011 Nigel Jones Diptera Coenosia tigrina 9 SO90259456 28/07/2011 Nigel Jones Diptera Coenosia tigrina site record SO898945 28/07/2011 Nigel Jones Diptera Coenosia tigrina site record SO898945 28/07/2011 Nigel Jones Diptera Conops quadrifasciatus site record SO898945 28/07/2011 Nigel Jones Diptera Dasysyrphus venustus 8 SO90379448 20/05/2011 Nigel Jones Diptera Dicranomyia affinis 5 SO89599442 20/05/2011 Pete Boardman Diptera Dicranomyia chorea 5 SO89599442 20/04/2011 Pete Boardman Diptera Dicranomyia chorea 7 SO90029443 20/04/2011 Pete Boardman Diptera Dicranomyia chorea 7 SO90309478 20/05/2011 Pete Boardman Diptera Dicranota pavida 5 SO89599442 20/05/2011 Pete Boardman Diptera Dicranota subtilis 5 SO89599442 20/05/2011 Pete Boardman Diptera Dioctria atricapilla 8 SO90379448 20/05/2011 Nigel Jones Diptera Dioctria baumhaueri 8 SO90379448 29/06/2011 Pete Boardman Diptera Ditrichophora calceata 8 SO90379448 28/07/2011 Nigel Jones Diptera Dolichocephala irrorata 8 SO90379448 18/04/2011 Nigel Jones Diptera Dolichopus brevipennis 2 SO89629479 20/05/2011 Nigel Jones Diptera Dolichopus brevipennis 5 SO89599442 20/05/2011 Nigel Jones Diptera Dolichopus brevipennis 9 SO90259456 20/05/2011 Nigel Jones Diptera Dolichopus discifer 8 SO90379448 28/07/2011 Nigel Jones Diptera Dolichopus picipes 5 SO89599442 20/05/2011 Nigel Jones Diptera Dolichopus plumipes 5 SO89599442 20/05/2011 Nigel Jones Diptera Dolichopus plumipes 8 SO90379448 20/05/2011 Nigel Jones Diptera Dolichopus plumipes 9 SO90259456 20/05/2011 Nigel Jones Diptera Dolichopus plumipes 8 SO90379448 28/07/2011 Nigel Jones Diptera Dolichopus plumipes 9 SO90259456 28/07/2011 Nigel Jones Diptera Dolichopus popularis 5 SO89599442 20/05/2011 Nigel Jones Diptera Dolichopus popularis 8 SO90379448 28/07/2011 Nigel Jones Diptera Dolichopus trivialis 5 SO89599442 20/05/2011 Nigel Jones Page 22

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Diptera Dolichopus trivialis 8 SO90379448 28/07/2011 Nigel Jones Diptera Dolichopus trivialis 9 SO90259456 28/07/2011 Nigel Jones Diptera Dolichopus ungulatus 5 SO89599442 20/05/2011 Nigel Jones Diptera Dolichopus ungulatus 8 SO90379448 28/07/2011 Nigel Jones Diptera Elgiva cucularia 1 SO89709483 20/05/2011 Nigel Jones Diptera Elgiva solicita 1 SO89709483 20/05/2011 Nigel Jones Diptera Eloeophila maculata 5 SO89599442 20/05/2011 Pete Boardman Diptera Eloeophila submarmorata 5 SO89599442 20/05/2011 Nigel Jones Diptera Eloeophila submarmorata 5 SO89599442 20/05/2011 Pete Boardman Diptera Empis aestiva 1 SO89709483 18/04/2011 Nigel Jones Diptera Empis livida 7 SO90309478 28/07/2011 Nigel Jones Diptera Empis livida site record SO898945 28/07/2011 Nigel Jones Diptera Empis nigripes 9 SO90259456 18/04/2011 Nigel Jones Diptera Empis nigripes 5 SO89599442 20/05/2011 Nigel Jones Diptera Empis nigripes 8 SO90379448 20/05/2011 Nigel Jones Diptera Empis praevia 5 SO89599442 20/05/2011 Nigel Jones Diptera Empis stercorea 5 SO89599442 20/05/2011 Nigel Jones Diptera Empis tessellata site record SO898945 20/05/2011 Nigel Jones Diptera Epistrophe eligans 8 SO90379448 20/04/2011 Pete Boardman Diptera Epistrophe eligans site record SO898945 20/05/2011 Nigel Jones Diptera Epistrophe grossulariae 8 SO90379448 28/07/2011 Nigel Jones Diptera Episyrphus balteatus site record SO898945 20/05/2011 Nigel Jones Diptera Episyrphus balteatus 9 SO90259456 29/06/2011 Pete Boardman Diptera Episyrphus balteatus 9 SO90259456 29/06/2011 Pete Boardman Diptera Episyrphus balteatus 1 SO89709483 15/09/2011 Pete Boardman Diptera Episyrphus balteatus 9 SO90259456 15/09/2011 Pete Boardman Diptera Erioconopa trivialis 9 SO90259456 18/04/2011 Pete Boardman Diptera Erioconopa trivialis 9 SO90259456 18/04/2011 Pete Boardman Diptera Erioconopa trivialis 5 SO89599442 20/04/2011 Pete Boardman Diptera Erioconopa trivialis 8 SO90379448 20/05/2011 Pete Boardman Diptera Erioptera fusculenta 9 SO90259456 29/06/2011 Pete Boardman Diptera Erioptera squalida 9 SO90259456 29/06/2011 Pete Boardman Diptera Eriothrix rufomaculata site record SO898945 28/07/2011 Nigel Jones Diptera Eristalis arbustorum site record SO898945 20/05/2011 Nigel Jones Diptera Eristalis intricarius 9 SO90259456 20/04/2011 Pete Boardman Diptera Eristalis pertinax 8 SO90379448 18/04/2011 Nigel Jones Diptera Eristalis pertinax 9 SO90259456 20/04/2011 Pete Boardman Diptera Eristalis tenax site record SO898945 20/05/2011 Nigel Jones Diptera Eristalis tenax 8 SO90379448 28/07/2011 Nigel Jones Diptera Eristalis tenax 1 SO89709483 15/09/2011 Pete Boardman Diptera Eristalis tenax 9 SO90259456 15/09/2011 Pete Boardman Diptera Eudasyphora cyanella site record SO898945 28/07/2011 Nigel Jones Diptera Eumerus funeralis site record SO898945 20/06/2011 Nigel Jones Diptera Eupeodes latifasciatus 1 SO89709483 20/05/2011 Nigel Jones Diptera Eupeodes luniger site record SO898945 20/06/2011 Nigel Jones Diptera Fannia armata 5 SO89599442 20/05/2011 Nigel Jones Diptera Fannia armata 5 SO89599442 20/05/2011 Nigel Jones Diptera Fannia lustrator 8 SO90379448 20/05/2011 Nigel Jones Diptera Fannia serena 5 SO89599442 20/05/2011 Nigel Jones Diptera Fannia similis 5 SO89599442 20/05/2011 Nigel Jones Diptera Geomyza tripunctata 1 SO89709483 18/04/2011 Nigel Jones Diptera Gloma fuscipennis 5 SO89599442 20/05/2011 Nigel Jones Diptera Gloma fuscipennis 8 SO90379448 20/05/2011 Nigel Jones Diptera Helina impuncta 5 SO89599442 20/05/2011 Nigel Jones Diptera Helina impuncta 8 SO90379448 28/07/2011 Nigel Jones Diptera Helina reversio 5 SO89599442 28/07/2011 Nigel Jones Diptera Helina reversio 9 SO90259456 28/07/2011 Nigel Jones Diptera Helophilus hybridus 1 SO89709483 28/07/2011 Nigel Jones Diptera Helophilus pendulus 8 SO90379448 18/04/2011 Nigel Jones Page 23

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Diptera Helophilus pendulus 8 SO90379448 20/05/2011 Nigel Jones Diptera Helophilus pendulus site record SO898945 20/05/2011 Nigel Jones Diptera Helophilus pendulus 9 SO90259456 29/06/2011 Pete Boardman Diptera Hercostomus aerosus 8 SO90379448 28/07/2011 Nigel Jones Diptera Hercostomus cupreus 1 SO89709483 20/05/2011 Nigel Jones Diptera Hercostomus cupreus 5 SO89599442 20/05/2011 Nigel Jones Diptera Hercostomus germanus 8 SO90379448 28/07/2011 Nigel Jones Diptera Hercostomus parvilamellatus 5 SO89599442 20/05/2011 Nigel Jones Diptera Herina frondescentiae 9 SO90259456 20/05/2011 Nigel Jones Diptera Herina frondescentiae site record SO898945 28/07/2011 Nigel Jones Diptera Hilara curtisi 8 SO90379448 20/05/2011 Nigel Jones Diptera Hilara litorea 8 SO90379448 28/07/2011 Nigel Jones Diptera Hilara lurida 8 SO90379448 28/07/2011 Nigel Jones Diptera Hybos femoratus 8 SO90379448 28/07/2011 Nigel Jones Diptera Hydrellia maura 5 SO89599442 20/05/2011 Nigel Jones Diptera Hydrellia maura 9 SO90259456 20/05/2011 Nigel Jones Diptera Hydrellia maura 9 SO90259456 20/05/2011 Nigel Jones Diptera Hydrellia maura 8 SO90379448 28/07/2011 Nigel Jones Diptera Hydrellia maura 9 SO90259456 28/07/2011 Nigel Jones Diptera Hydromya dorsalis 8 SO90379448 18/04/2011 Nigel Jones Diptera Hydromya dorsalis 9 SO90259456 18/04/2011 Nigel Jones Diptera Hydromya dorsalis 5 SO89599442 20/04/2011 Pete Boardman Diptera Hydromya dorsalis 9 SO90259456 28/07/2011 Nigel Jones Diptera Hydrotaea cyrtoneurina 5 SO89599442 20/05/2011 Nigel Jones Diptera Hydrotaea cyrtoneurina 8 SO90379448 28/07/2011 Nigel Jones Diptera Ilisia maculata 8 SO90379448 20/05/2011 Nigel Jones Diptera Ilisia maculata 9 SO90259456 20/05/2011 Pete Boardman Diptera Jassidophaga villosa 9 SO90259456 18/04/2011 Nigel Jones Diptera Lauxania cylindricornis site record SO898945 20/05/2011 Nigel Jones Diptera Leptogaster cylindrica site record SO901944 20/06/2011 Ian Cheeseborough Diptera Leucophora obtusa 8a SO903945 18/04/2011 Nigel Jones Diptera Limnellia quadrata 9 SO90259456 18/04/2011 Nigel Jones Diptera Limnia paludicloa 9 SO90259456 28/07/2011 Nigel Jones Diptera Limnia unguicornis 8 SO90379448 20/05/2011 Nigel Jones Diptera Limonia flavipes 5 SO89599442 20/05/2011 Pete Boardman Diptera Limonia macrostigma 8 SO90379448 20/05/2011 Pete Boardman Diptera Limonia nubeculosa 8 SO90379448 20/05/2011 Pete Boardman Diptera Limonia nubeculosa 9 SO90259456 15/09/2011 Pete Boardman Diptera Limonia phragmitidis 8 SO90379448 20/05/2011 Pete Boardman Diptera Lonchaea chorea 8 SO90379448 20/05/2011 Nigel Jones Diptera Lonchoptera lutea 5 SO89599442 18/04/2011 Nigel Jones Diptera Lonchoptera lutea 7 SO90309478 18/04/2011 Nigel Jones Diptera Loxocera albiseta 8 SO90379448 28/07/2011 Nigel Jones Diptera Loxocera albiseta 9 SO90259456 28/07/2011 Nigel Jones Diptera Lydella stabulans site record SO898945 28/07/2011 Nigel Jones Diptera Machimus cingulatus site record SO898945 28/07/2011 Nigel Jones Diptera Meigenia mutabilis 8 SO90379448 28/07/2011 Nigel Jones Diptera Meiosimyza decipiens site record SO898945 28/07/2011 Nigel Jones Diptera Meiosimyza rorida site record SO898945 28/07/2011 Nigel Jones Diptera Melanogaster hirtella 9 SO90259456 20/05/2011 Nigel Jones Diptera Melanostoma mellinum 9 SO90259456 28/07/2011 Nigel Jones Diptera Melanostoma scalare 5 SO89599442 18/04/2011 Nigel Jones Diptera Melanostoma scalare 5 SO89599442 20/05/2011 Nigel Jones Diptera Melanostoma scalare 8 SO90379448 20/05/2011 Nigel Jones Diptera Melanostoma scalare 5 SO89599442 28/07/2011 Nigel Jones Diptera Melanostoma scalare 8 SO90379448 28/07/2011 Nigel Jones Diptera Melanostoma scalare 9 SO90259456 28/07/2011 Nigel Jones Diptera Merodon equestris 8 SO90379448 20/05/2011 Nigel Jones Diptera Mesembrina meridiana 9 SO90259456 15/09/2011 Pete Boardman Page 24

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Diptera Microchrysa flavicornis 8 SO90379448 28/07/2011 Nigel Jones Diptera Microchrysa polita 8 SO90379448 20/05/2011 Nigel Jones Diptera Microphor holosericeus 5 SO89599442 20/05/2011 Nigel Jones Diptera Minettia fasciata site record SO898945 28/07/2011 Nigel Jones Diptera Minettia inusta 8 SO90379448 28/07/2011 Nigel Jones Diptera Minettia longipennis site record SO898945 20/05/2011 Nigel Jones Diptera Minettia longipennis site record SO898945 28/07/2011 Nigel Jones Diptera Molophilus cinereifrons 5 SO89599442 20/05/2011 Pete Boardman Diptera Molophilus obscurus 1 SO89709483 20/05/2011 Pete Boardman Diptera Molophilus obscurus 9 SO90259456 29/06/2011 Pete Boardman Diptera Morellia hortorum 8 SO90379448 28/07/2011 Nigel Jones Diptera Muscina levida 8 SO90379448 28/07/2011 Nigel Jones Diptera Myathropa florea site record SO898945 20/05/2011 Nigel Jones Diptera Nanna fasciata 2 SO89629479 18/04/2011 Nigel Jones Diptera Nanna flavipes 2 SO89629479 18/04/2011 Nigel Jones Diptera Neoascia podagrica 5 SO89599442 18/04/2011 Nigel Jones Diptera Neoascia podagrica 5 SO89599442 28/07/2011 Nigel Jones Diptera Nephrotoma appendiculata 8 SO90379448 20/05/2011 Pete Boardman Diptera Nephrotoma appendiculata 9 SO90259456 29/06/2011 Pete Boardman Diptera Nephrotoma quadrifaria 9 SO90259456 29/06/2011 Pete Boardman Diptera Neria cibaria 8 SO90379448 20/05/2011 Nigel Jones Diptera Neuroctena anilis 5 SO89599442 20/05/2011 Nigel Jones Diptera Notiphila maculata 9 SO90259456 28/07/2011 Nigel Jones Diptera Ocydromia glabricula 5 SO89599442 20/05/2011 Nigel Jones Diptera Oedalea stigmatella 8 SO90379448 20/05/2011 Nigel Jones Diptera Opomyza germinationis 5 SO89599442 20/05/2011 Nigel Jones Diptera Opomyza petrei 8 SO90379448 28/07/2011 Nigel Jones Diptera Ormosia lineata 11 SO89649435 20/04/2011 Pete Boardman Diptera Ormosia lineata 5 SO89599442 20/05/2011 Pete Boardman Diptera Ormosia lineata 7 SO90309478 20/05/2011 Pete Boardman Diptera Oxycera rara 9 SO90259456 29/06/2011 Pete Boardman Diptera Pachygaster leachii 8 SO90379448 28/07/2011 Nigel Jones Diptera Palloptera scutellata 1 SO89709483 18/04/2011 Nigel Jones Diptera Palloptera scutellata 5 SO89599442 18/04/2011 Nigel Jones Diptera Palloptera scutellata 7 SO90309478 18/04/2011 Nigel Jones Diptera Palloptera scutellata 9 SO90259456 18/04/2011 Nigel Jones Diptera Parydra coarctata 5 SO89599442 20/05/2011 Nigel Jones Diptera Parydra coarctata 9 SO90259456 28/07/2011 Nigel Jones Diptera Pelatachina tibialis site record SO898945 28/07/2011 Nigel Jones Diptera Phaonia errans 8 SO90379448 28/07/2011 Nigel Jones Diptera Phaonia fuscata site record SO898945 28/07/2011 Nigel Jones Diptera Phaonia subventa 8 SO90379448 28/07/2011 Nigel Jones Diptera Phasia obesa site record SO898945 20/05/2011 Nigel Jones Diptera Phasia pusilla 5 SO89599442 20/05/2011 Nigel Jones Diptera Pherbellia schoenherri site record SO898945 28/07/2011 Nigel Jones Diptera Pilaria discicollis 9 SO90259456 29/06/2011 Pete Boardman Diptera Platycheirus clypeatus 9 SO90259456 28/07/2011 Nigel Jones Diptera Platycheirus granditarsus 9 SO90259456 28/07/2011 Nigel Jones Diptera Platycheirus manicatus 9 SO90259456 20/05/2011 Nigel Jones Diptera Platycheirus peltatus 9 SO90259456 28/07/2011 Nigel Jones Diptera Platycheirus tarsalis 9 SO90259456 20/05/2011 Nigel Jones Diptera Platycheirus tarsalis 5 SO89599442 28/07/2011 Nigel Jones Diptera Platypalpus longicornis 8 SO90379448 20/05/2011 Nigel Jones Diptera Poecilobothrus nobilitatus 9 SO90259456 29/06/2011 Pete Boardman Diptera Poecilobothrus nobilitatus 8 SO90379448 28/07/2011 Nigel Jones Diptera Portevinia maculata 8 SO90379448 20/05/2011 Nigel Jones Diptera Psila fimetaria site record SO898945 20/05/2011 Nigel Jones Diptera Ptychoptera albimana 5 SO89599442 20/04/2011 Pete Boardman Diptera Ptychoptera albimana 8 SO90379448 20/04/2011 Pete Boardman Page 25

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Diptera Ptychoptera albimana 5 SO89599442 20/05/2011 Pete Boardman Diptera Ptychoptera albimana 8 SO90379448 20/05/2011 Pete Boardman Diptera Rhagio lineola 8 SO90379448 28/07/2011 Nigel Jones Diptera Rhagio scolopaceus 8 SO90379448 20/05/2011 Nigel Jones Diptera Rhamphomyia albipennis 1 SO89709483 18/04/2011 Nigel Jones Diptera Rhamphomyia albohirta 5 SO89599442 18/04/2011 Nigel Jones Diptera Rhamphomyia albohirta 8 SO90379448 20/05/2011 Nigel Jones Diptera Rhamphomyia nigripennis 5 SO89599442 20/05/2011 Nigel Jones Diptera Rhamphomyia subcinerascens 8 SO90379448 18/04/2011 Nigel Jones Diptera Rhamphomyia subcinerascens 9 SO90259456 18/04/2011 Nigel Jones Diptera Rhamphomyia sulcata 7 SO90309478 18/04/2011 Nigel Jones Diptera Rhamphomyia umbripennis 5 SO89599442 18/04/2011 Nigel Jones Diptera Rhamphomyia umbripennis 7 SO90309478 18/04/2011 Nigel Jones Diptera Rhaphium brevicorne 8 SO90379448 28/07/2011 Nigel Jones Diptera Rhaphium caliginosum 8 SO90379448 18/04/2011 Nigel Jones Diptera Rhingia campestris 7 SO90029443 20/04/2011 Pete Boardman Diptera Rhingia campestris 8 SO90379448 20/05/2011 Nigel Jones Diptera Sapromyza quadricincta 8 SO90379448 20/05/2011 Nigel Jones Diptera Sapromyza sexpunctata site record SO898945 28/07/2011 Nigel Jones Diptera Scaeva pyrastri 8a SO903945 28/07/2011 Ian Cheeseborough Diptera Scatella tenuicosta 9 SO90259456 28/07/2011 Nigel Jones Diptera Scathophaga furcata site record SO898945 28/07/2011 Nigel Jones Diptera Scathophaga lutaria 8 SO90379448 18/04/2011 Nigel Jones Diptera Scathophaga stercoraria site record SO898945 20/05/2011 Nigel Jones Diptera Sepedon sphegea 1 SO89709483 18/04/2011 Nigel Jones Diptera Sepedon sphegea 1 SO89709483 20/04/2011 Pete Boardman Diptera Sepsis fulgens site record SO898945 28/07/2011 Nigel Jones Diptera Sepsis punctum site record SO898945 28/07/2011 Nigel Jones Diptera Sicus ferrugineus site record SO898945 20/06/2011 Nigel Jones Diptera Siphona geniculata 8 SO90379448 28/07/2011 Nigel Jones Diptera Spania nigra 5 SO89599442 20/05/2011 Nigel Jones Diptera Sphaerophoria interrupta site record SO898945 28/07/2011 Nigel Jones Diptera Sphaerophoria scripta 9 SO90259456 15/09/2011 Pete Boardman Diptera Sphaerophoria taeniata 2 SO89629479 28/07/2011 Nigel Jones Diptera Sphegina clunipes 8 SO90379448 20/05/2011 Nigel Jones Diptera Sybistroma obscurellum 8 SO90379448 28/07/2011 Nigel Jones Diptera Sympycnus desoutteri 5 SO89599442 20/05/2011 Nigel Jones Diptera Sympycnus desoutteri 9 SO90259456 20/05/2011 Nigel Jones Diptera Syntormon macula 8 SO90379448 18/04/2011 Nigel Jones Diptera Syntormon macula 8 SO90379448 20/05/2011 Nigel Jones Diptera Syritta pipiens site record SO898945 20/05/2011 Nigel Jones Diptera Syrphus vitripennis site record SO898945 28/07/2011 Nigel Jones Diptera Tachina fera 9a SO90269452 20/05/2011 Ian Cheeseborough Diptera Tachina fera 9 SO90259456 15/09/2011 Pete Boardman Diptera Tephritis cometa 9 SO90259456 20/05/2011 Nigel Jones Diptera Terellia ruficauda 8 SO90379448 28/07/2011 Nigel Jones Diptera Terellia ruficauda 9 SO90259456 28/07/2011 Nigel Jones Diptera Tetanocera elata site record SO898945 28/07/2011 Nigel Jones Diptera Tetanocera ferruginea site record SO898945 28/07/2011 Nigel Jones Diptera Tetanocera fuscinervis 8 SO90379448 20/05/2011 Nigel Jones Diptera Teuchophorus calcaratus 8 SO90379448 28/07/2011 Nigel Jones Diptera Teuchophorus monacanthus 9 SO90259456 20/05/2011 Nigel Jones Diptera Teuchophorus monacanthus 8 SO90379448 28/07/2011 Nigel Jones Diptera Teuchophorus nigricosta 8 SO90379448 28/07/2011 Nigel Jones Diptera Teuchophorus simplex 8 SO90379448 28/07/2011 Nigel Jones Diptera Themira annulipes site record SO898945 28/07/2011 Nigel Jones Diptera Thricops diaphana 8 SO90379448 28/07/2011 Nigel Jones Diptera Tipula fascipennis 9 SO90259456 29/06/2011 Pete Boardman Diptera Tipula lateralis 11 SO89649435 20/04/2011 Pete Boardman Page 26

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Diptera Tipula lateralis 9 SO90259456 29/06/2011 Pete Boardman Diptera Tipula paludosa 1 SO89709483 15/09/2011 Pete Boardman Diptera Tipula paludosa 9 SO90259456 15/09/2011 Pete Boardman Diptera Tipula paludosa 9 SO90259456 15/09/2011 Pete Boardman Diptera Tipula paludosa site record SO898945 15/09/2011 Pete Boardman Diptera Tipula submarmorata site record SO89399444 20/05/2011 Pete Boardman Diptera Tipula vittata 9 SO90259456 18/04/2011 Pete Boardman Diptera Tipula vittata 5 SO89599442 20/04/2011 Pete Boardman Diptera Tipula vittata 7 SO90309478 20/04/2011 Pete Boardman Diptera Tipula vittata 9 SO90259456 20/04/2011 Pete Boardman Diptera Tipula vittata 9 SO90259456 20/04/2011 Pete Boardman Diptera Trichopsomyia flavitarsis 9 SO90259456 20/05/2011 Nigel Jones Diptera Tricyphona immaculata 5 SO89599442 20/04/2011 Pete Boardman Diptera Tricyphona immaculata 5 SO89599442 20/05/2011 Pete Boardman Diptera Tricyphona immaculata 8 SO90379448 20/05/2011 Pete Boardman Diptera Urophora cardui site record SO898945 28/07/2011 Nigel Jones Diptera Urophora jaceana 8 SO90379448 29/06/2011 Pete Boardman Diptera Urophora jaceana site record SO898945 28/07/2011 Nigel Jones Diptera Vibrissina debilitata site record SO898945 28/07/2011 Nigel Jones Diptera Volucella bombylans site record SO898945 20/05/2011 Nigel Jones Diptera Xyphosia miliaria site record SO898945 28/07/2011 Nigel Jones Hemiptera Acanthosoma haemorrhoidale 1 SO89709483 15/09/2011 Pete Boardman Hemiptera Capsus ater 3 SO89529444 2011 Richard Wright Hemiptera Coreus marginatus 9a SO90269452 20/06/2011 Ian Cheeseborough Hemiptera Deraeocoris lutescens 7 SO90309478 2011 Richard Wright Hemiptera Elasmostethus interstinctus site record SO898945 20/05/2011 Nigel Jones Hemiptera Himacerus mirmicoides 4 SO89459440 2011 Richard Wright Hemiptera Ischnodemus sabuleti 5 SO89599442 2011 Richard Wright Hemiptera Leptopterna dolabrata 2 SO89629479 2011 Richard Wright Hemiptera Leptopterna dolabrata 3 SO89529444 2011 Richard Wright Hemiptera Liocoris tripustulatus 7 SO90309478 2011 Richard Wright Hemiptera Nabis rugosus 2 SO89629479 2011 Richard Wright Hemiptera Nabis rugosus 3 SO89529444 2011 Richard Wright Hemiptera Nabis rugosus 4 SO89459440 2011 Richard Wright Hemiptera Nabis rugosus 6 SO89799476 2011 Richard Wright Hemiptera Nabis rugosus 7 SO90309478 2011 Richard Wright Hemiptera Nabis rugosus 8 SO90379448 2011 Richard Wright Hemiptera Notostira elongata 3 SO89529444 2011 Richard Wright Hemiptera Orthotylus ericetorum 4 SO89459440 2011 Richard Wright Hemiptera Orthotylus ericetorum 6 SO89799476 2011 Richard Wright Hemiptera Palomena prasina 7 SO90309478 2011 Richard Wright Hemiptera Palomena prasina site record SO899945 18/04/2011 Ian Cheeseborough Hemiptera Palomena prasina 9a SO90269452 20/05/2011 Ian Cheeseborough Hemiptera Piezodorus lituratus site record SO902948 18/04/2011 Ian Cheeseborough Hemiptera Pithanus maerkelii 9 SO90259456 29/06/2011 Pete Boardman Hemiptera Scolopostethus decoratus 4 SO89459440 2011 Richard Wright Hemiptera Scolopostethus decoratus 6 SO89799476 2011 Richard Wright Hemiptera Stenodema calcarata 3 SO89529444 2011 Richard Wright Hemiptera Stenotus binotatus 3 SO89529444 2011 Richard Wright Hemiptera Stygnocoris pedestris 3 SO89529444 2011 Richard Wright Hemiptera Stygnocoris rusticus 4 SO89459440 2011 Richard Wright Hemiptera Stygnocoris rusticus 8 SO90379448 2011 Richard Wright Hemiptera Stygnocoris rusticus 9 SO90259456 2011 Richard Wright Hemiptera Velia caprai 5 SO89599442 2011 Richard Wright Hymenoptera Ancistrocerus gazella 8a SO903944 28/07/2011 Ian Cheeseborough Hymenoptera Andrena barbilabris 11 SO89649435 18/04/2011 Ian Cheeseborough Hymenoptera Andrena bicolor site record SO899948 18/04/2011 Ian Cheeseborough Hymenoptera Andrena bicolor 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Andrena chrysosceles 12 SO90349456 18/04/2011 Ian Cheeseborough Page 27

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Hymenoptera Andrena cineraria 11 SO89649435 18/04/2011 Ian Cheeseborough Hymenoptera Andrena cineraria site record SO899948 18/04/2011 Ian Cheeseborough Hymenoptera Andrena dorsata 8a SO903944 28/07/2011 Ian Cheeseborough Hymenoptera Andrena fucata 8a SO903944 20/05/2011 Ian Cheeseborough Hymenoptera Andrena fulva 11 SO89649435 18/04/2011 Ian Cheeseborough Hymenoptera Andrena fulva 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Andrena fulva site record SO899945 18/04/2011 Ian Cheeseborough Hymenoptera Andrena haemorrhoa 11 SO89649435 18/04/2011 Ian Cheeseborough Hymenoptera Andrena haemorrhoa 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Andrena haemorrhoa site record SO899945 18/04/2011 Ian Cheeseborough Hymenoptera Andrena haemorrhoa site record SO901944 18/04/2011 Ian Cheeseborough Hymenoptera Andrena haemorrhoa site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Andrena nigroaenea 7 SO90309478 18/04/2011 Ian Cheeseborough Hymenoptera Andrena nigroaenea site record SO901944 18/04/2011 Ian Cheeseborough Hymenoptera Andrena scotica 8a SO903945 18/04/2011 Ian Cheeseborough Hymenoptera Andrena scotica site record SO899945 18/04/2011 Ian Cheeseborough Hymenoptera Andrena scotica site record SO901944 18/04/2011 Ian Cheeseborough Hymenoptera Andrena subopaca 8a SO903944 18/04/2011 Ian Cheeseborough Hymenoptera Andrena wilkella 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Andrena wilkella 8a SO903944 28/07/2011 Ian Cheeseborough Hymenoptera Anoplius caviventris site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Anoplius nigerrimus 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Anthidium manicatum 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Anthophora furcata 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Anthophora furcata 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Argogorytes mystaceus 8a SO903944 20/05/2011 Ian Cheeseborough Hymenoptera Bombus hortorum site record SO899943 29/06/2011 Ian Cheeseborough Hymenoptera Bombus hypnorum 9a SO90269452 20/05/2011 Ian Cheeseborough Hymenoptera Bombus hypnorum site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Bombus hypnorum site record SO898945 20/05/2011 Nigel Jones Hymenoptera Bombus hypnorum 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Bombus lapidarius site record SO901944 18/04/2011 Ian Cheeseborough Hymenoptera Bombus lapidarius site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Bombus lapidarius 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Bombus lapidarius 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Bombus lucorum 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Bombus lucorum 9a SO90269452 20/05/2011 Ian Cheeseborough Hymenoptera Bombus lucorum 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Bombus lucorum 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Bombus pascuorum 7 SO90309478 18/04/2011 Ian Cheeseborough Hymenoptera Bombus pascuorum site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Bombus pascuorum 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Bombus pascuorum 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Bombus pratorum 9a SO90269452 20/05/2011 Ian Cheeseborough Hymenoptera Bombus pratorum site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Bombus pratorum 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Bombus sylvestris 9a SO90269452 20/05/2011 Ian Cheeseborough Hymenoptera Bombus sylvestris site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Bombus terrestris 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Bombus terrestris site record SO899945 18/04/2011 Ian Cheeseborough Hymenoptera Bombus terrestris 9a SO90269452 20/05/2011 Ian Cheeseborough Hymenoptera Bombus terrestris 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Bombus vestalis site record SO899945 18/04/2011 Ian Cheeseborough Hymenoptera Crossocerus annulipes 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Crossocerus capitosus 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Crossocerus cetratus 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Crossocerus megacephalus 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Crossocerus nigritus 8a SO903944 20/05/2011 Ian Cheeseborough Hymenoptera Crossocerus podagricus 9a SO90269452 28/07/2011 Ian Cheeseborough Page 28

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Hymenoptera Crossocerus quadrimaculatus 8 SO90379448 20/06/2011 Ian Cheeseborough Hymenoptera Dolichovespula norwegica 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Dolichovespula sylvestris 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Ectemnius cavifrons 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Ectemnius continuus 8a SO903944 20/05/2011 Ian Cheeseborough Hymenoptera Ectemnius continuus 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Ectemnius continuus 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Halictus rubicundus 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Halictus tumulorum 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Hylaeus communis 9 SO90259456 28/07/2011 Nigel Jones Hymenoptera Hylaeus communis 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Lasioglossum calceatum 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Lasioglossum leucopus 8a SO903945 28/07/2011 Ian Cheeseborough Hymenoptera Lasioglossum villosulum 8a SO903945 18/04/2011 Ian Cheeseborough Hymenoptera Lasius flavus 7 SO90309478 18/04/2011 Ian Cheeseborough Hymenoptera Lasius niger site record SO899948 18/04/2011 Ian Cheeseborough Hymenoptera Lindenius albilabris site record SO899943 29/06/2011 Ian Cheeseborough Hymenoptera Lindenius albilabris 8a SO903944 28/07/2011 Ian Cheeseborough Hymenoptera Megachile ligniseca 8a SO903945 29/06/2011 Ian Cheeseborough Hymenoptera Megachile willughbiella 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Megachile willughbiella 10 SO89639441 28/07/2011 Ian Cheeseborough Hymenoptera Megachile willughbiella 8a SO903944 28/07/2011 Ian Cheeseborough Hymenoptera Mimumesa dahlbomi 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Myrmosa atra site record SO899943 29/06/2011 Ian Cheeseborough Hymenoptera Nomada fabriciana 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Nomada flava site record SO901944 18/04/2011 Ian Cheeseborough Hymenoptera Nomada flava/panzeri 7 SO90309478 18/04/2011 Ian Cheeseborough Hymenoptera Nomada goodeniana 7 SO90309478 18/04/2011 Ian Cheeseborough Hymenoptera Nomada lathburiana 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Nomada ruficornis 11 SO89649435 18/04/2011 Ian Cheeseborough Hymenoptera Nomada ruficornis 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Osmia caerulescens 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Osmia rufa 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Osmia rufa site record SO899945 18/04/2011 Ian Cheeseborough Hymenoptera Osmia rufa site record SO901944 20/06/2011 Ian Cheeseborough Hymenoptera Passaloecus gracilis 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Passaloecus singularis 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Pemphredon lugubris 8 SO90379448 20/05/2011 Nigel Jones Hymenoptera Pemphredon lugubris 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Psenulus concolor 8a SO903944 20/05/2011 Ian Cheeseborough Hymenoptera Psenulus pallipes 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Rhopalum clavipes 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Rhopalum clavipes site record SO898945 28/07/2011 Nigel Jones Hymenoptera Sphecodes ephippius 12 SO90349456 18/04/2011 Ian Cheeseborough Hymenoptera Stigmus solskyi 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Tiphia minuta 8a SO903945 29/06/2011 Ian Cheeseborough Hymenoptera Trichrysis cyanea 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Trypoxylon attenuatum 9a SO90269452 20/05/2011 Ian Cheeseborough Hymenoptera Trypoxylon attenuatum 9a SO90269452 20/06/2011 Ian Cheeseborough Hymenoptera Trypoxylon attenuatum 8a SO903945 29/06/2011 Ian Cheeseborough Hymenoptera Trypoxylon clavicerum 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Vespula germanica 7 SO90309478 18/04/2011 Ian Cheeseborough Hymenoptera Vespula germanica 9a SO90269452 28/07/2011 Ian Cheeseborough Hymenoptera Vespula rufa site record SO898945 20/05/2011 Ian Cheeseborough Hymenoptera Vespula rufa 8a SO903944 29/06/2011 Ian Cheeseborough Hymenoptera Vespula vulgaris 11 SO89649435 18/04/2011 Ian Cheeseborough Hymenoptera Vespula vulgaris 8a SO903945 29/06/2011 Ian Cheeseborough Isopoda Oniscus asellus 7 SO90309478 2011 Richard Wright Isopoda Philoscia muscorum 1 SO89709483 2011 Richard Wright Page 29

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Isopoda Philoscia muscorum 2 SO89629479 2011 Richard Wright Isopoda Philoscia muscorum 3 SO89529444 2011 Richard Wright Isopoda Philoscia muscorum 4 SO89459440 2011 Richard Wright Isopoda Philoscia muscorum 6 SO89799476 2011 Richard Wright Isopoda Philoscia muscorum 8 SO90379448 2011 Richard Wright Isopoda Philoscia muscorum 9 SO90259456 2011 Richard Wright Isopoda Trichoniscus pusillus 1 SO89709483 2011 Richard Wright Isopoda Trichoniscus pusillus 2 SO89629479 2011 Richard Wright Isopoda Trichoniscus pusillus 6 SO89799476 2011 Richard Wright Isopoda Trichoniscus pusillus 9 SO90259456 2011 Richard Wright Lepidoptera Aglais urticae 8 SO90379448 20/04/2011 Pete Boardman Lepidoptera Aglais urticae 9a SO90269452 20/05/2011 Ian Cheeseborough Lepidoptera Aglais urticae 9 SO90259456 29/06/2011 Pete Boardman Lepidoptera Anthocharis cardamines site record SO902948 18/04/2011 Ian Cheeseborough Lepidoptera Anthocharis cardamines 8 SO90379448 20/04/2011 Pete Boardman Lepidoptera Anthocharis cardamines 9 SO90259456 20/04/2011 Pete Boardman Lepidoptera Aphantopus hyperantus 9a SO90269452 20/06/2011 Ian Cheeseborough Lepidoptera Aphantopus hyperantus 9 SO90259456 29/06/2011 Pete Boardman Lepidoptera Aphantopus hyperantus 8a SO903945 28/07/2011 Ian Cheeseborough Lepidoptera Callophrys rubi 9a SO90269452 20/05/2011 Ian Cheeseborough Lepidoptera Coenonympha pamphilus 9a SO90269452 20/05/2011 Ian Cheeseborough Lepidoptera Coenonympha pamphilus 9a SO90269452 20/06/2011 Ian Cheeseborough Lepidoptera Inachis io 8 SO90379448 20/04/2011 Pete Boardman Lepidoptera Inachis io 9a SO90269452 20/06/2011 Ian Cheeseborough Lepidoptera Inachis io 8a SO903944 28/07/2011 Ian Cheeseborough Lepidoptera Lycaena phlaeas 9a SO90269452 28/07/2011 Ian Cheeseborough Lepidoptera Maniola jurtina 9 SO90259456 29/06/2011 Pete Boardman Lepidoptera Ochlodes venata 9a SO90269452 20/06/2011 Ian Cheeseborough Lepidoptera Ochlodes venata 9 SO90259456 29/06/2011 Pete Boardman Lepidoptera Pararge aegeria 7 SO90309478 20/04/2011 Pete Boardman Lepidoptera Pararge aegeria 9a SO90269452 20/05/2011 Ian Cheeseborough Lepidoptera Pararge aegeria 9 SO90259456 29/06/2011 Pete Boardman Lepidoptera Pararge aegeria 9 SO90259456 15/09/2011 Pete Boardman Lepidoptera Pieris napi 8 SO90379448 20/04/2011 Pete Boardman Lepidoptera Pieris napi 9a SO90269452 20/05/2011 Ian Cheeseborough Lepidoptera Pieris rapae 9a SO90269452 20/06/2011 Ian Cheeseborough Lepidoptera Pieris rapae 9a SO90269452 28/07/2011 Ian Cheeseborough Lepidoptera Polygonia c-album 8 SO90379448 20/04/2011 Pete Boardman Lepidoptera Pyronia tithonus 9a SO90269452 28/07/2011 Ian Cheeseborough Lepidoptera Stigmella aurella 7 SO90029443 20/04/2011 Pete Boardman Lepidoptera Thymelicus sylvestris 9a SO90269452 20/06/2011 Ian Cheeseborough Odonata Coenagrion puella 9a SO90269452 20/05/2011 Ian Cheeseborough Odonata Coenagrion puella 9a SO90269452 20/06/2011 Ian Cheeseborough Odonata Ischnura elegans 9 SO90259456 29/06/2011 Pete Boardman Odonata Pyrrhosoma nymphula 9a SO90269452 20/05/2011 Ian Cheeseborough Odonata Sympetrum striolatum 1 SO89709483 15/09/2011 Pete Boardman Opiliones Mitopus morio 2 SO89629479 2011 Richard Wright Opiliones Nemastoma bimaculatum 2 SO89629479 2011 Richard Wright Opiliones Nemastoma bimaculatum 5 SO89599442 2011 Richard Wright Opiliones Nemastoma bimaculatum 7 SO90309478 2011 Richard Wright Opiliones Nemastoma bimaculatum 9 SO90259456 2011 Richard Wright Opiliones Paroligolophus agrestis 1 SO89709483 2011 Richard Wright Opiliones Paroligolophus agrestis 3 SO89529444 2011 Richard Wright Opiliones Paroligolophus agrestis 9 SO90259456 2011 Richard Wright Opiliones Rilaena triangularis 1 SO89709483 2011 Richard Wright Opiliones Rilaena triangularis 3 SO89529444 2011 Richard Wright Opiliones Rilaena triangularis 4 SO89459440 2011 Richard Wright Orthoptera Omocestus viridulus 9a SO90269452 20/06/2011 Ian Cheeseborough Orthoptera Tetrix subulata 5 SO89599442 2011 Richard Wright Page 30

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

Orthoptera Tetrix subulata 8 SO90379448 2011 Richard Wright Orthoptera Tetrix subulata 9 SO90259456 2011 Richard Wright Orthoptera Tetrix subulata 5 SO89599442 20/04/2011 Pete Boardman

Appendix 2 – ISIS outputs for Penn Common

WHOLE SITE

SAT % of national species Related BAT rarity code SAT name No. spp. Condition pool score F001 scrub edge 19 fav 11 F002 rich flower resource 15 fav 6 A212 bark & sapwood decay 23 fav 5 131 F006 dung 3 3 F112 open short sward 4 2 120 W111 shingle banks 1 2 154 W126 seepage 1 2 154 W314 reedfen and pools 2 2 135 F003 scrub-heath & moorland 5 1 F111 bare sand & chalk 5 1 120 A211 heartwood decay 1 1 131

BAT Representation Rarity code BAT name (1-100) score Condition BAT species richness IEC F2 grassland & scrub matrix 34 115 154 unshaded early successional F1 mosaic 11 120 51 W3 permanent wet mire 10 135 44 W1 flowing water 8 154 fav 35 A2 wood decay 7 131 30 0 F3 shaded field & ground layer 6 130 29 W2 mineral marsh & open water 5 112 25 A1 arboreal canopy 3 13

CALCAREOUS FLUSH

SAT % of national species Related BAT rarity code SAT name No. spp. Condition pool score W111 shingle banks 1 2 132

BAT Representation Rarity code BAT name (1-100) score Condition BAT species richness IEC F2 grassland & scrub matrix 22 127 22 W1 flowing water 19 132 19 W2 mineral marsh & open water 18 122 18 F3 shaded field & ground layer 11 11 W3 permanent wet mire 10 10

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

WET HABITATS IN EAST OF SITE

SAT % of national species Related BAT rarity code SAT name No. spp. Condition pool score W126 seepage 2 4 177 F112 open short sward 3 2 F001 scrub edge 2 1 W314 reedfen and pools 1 1 121 A212 bark & sapwood decay 3 1 F002 rich flower resource 1 0 F003 scrub-heath & moorland 1 0 F111 bare sand & chalk 1 0

BAT Representation Rarity code BAT name (1-100) score Condition BAT species richness IEC F2 grassland & scrub matrix 29 116 73 W3 permanent wet mire 16 121 40 W1 flowing water 12 177 fav 31 W2 mineral marsh & open water 11 118 28 F3 shaded field & ground layer 6 15 unshaded early successional F1 mosaic 3 8 A2 wood decay 3 8 0 A1 arboreal canopy 1 3

DRY ACID GRASSLAND

SAT % of national species Related BAT rarity code SAT name No. spp. Condition pool score F003 scrub-heath & moorland 10 fav 3 F111 bare sand & chalk 3 1 F001 scrub edge 1 1 F112 open short sward 1 1

BAT Representation Rarity code BAT name (1-100) score Condition BAT species richness IEC F2 grassland & scrub matrix 63 116 92 unshaded early successional F1 mosaic 5 8 W2 mineral marsh & open water 4 6 F3 shaded field & ground layer 3 4 W3 permanent wet mire 3 4 W1 flowing water 1 1

DAMP GRASSLAND

SAT % of national species Related BAT rarity code SAT name No. spp. Condition pool score

BAT Representation Rarity code BAT name (1-100) score Condition BAT species richness IEC F2 grassland & scrub matrix 47 114 43 W3 permanent wet mire 16 15 W2 mineral marsh & open water 4 4 F3 shaded field & ground layer 2 2 W1 flowing water 2 2

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Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

ACULEATE HABITATS

SAT % of national species Related BAT rarity code SAT name No. spp. Condition pool score A212 bark & sapwood decay 26 fav 5 119 F002 rich flower resource 8 3 F006 dung 3 3 F001 scrub edge 5 3 F112 open short sward 2 1 104

BAT Representation Rarity code BAT name (1-100) score Condition BAT species richness IEC unshaded early successional F1 mosaic 29 104 28 A2 wood decay 28 119 27 0 F2 grassland & scrub matrix 21 100 20 A1 arboreal canopy 2 2 W2 mineral marsh & open water 2 2

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