Geographic distribution of khapra , granarium, by location as recorded in six publications

Location EPPO2016 Australia2016 Myers2012 SPIRes2009 French2005 Banks1977 5 Present, no details Afghanistan Afghanistan Afghanistan Present, flourishes certain areas Algeria 5 Present, widespread Algeria Algeria Algeria Present, flourishes certain areas Present, restricted distribution Angola 5 Absent, unreliable record Angola Angola Absent, doubtful record or intercepted only Argentina 1 Argentina 5 Absent, intercepted only Australia Australia Absent, intercepted only Absence, eradicated Austria 3 Absent, intercepted only Austria Absent, fomerly present Bahrain 2 Bahrain Bahrain Present, restricted distribution Bangladesh 5 Present, no details Bangladesh Bangladesh Bangladesh or intercepted only Belgium 3 Absent, pest no longer present Belgium Absent, fomerly present Benin 2 Benin Benin Bolivia 1 Bolivia Botswana 1 Botswana Bulgaria 3 Absent, intercepted only Bulgaria (i1940) Absent, intercepted only Burkina Faso 5 Present, no details Burkina Burkina Faso Burkina Faso Present, flourishes certain areas Burundi 1 Burundi 2 Canada Canada Cambodia 1 Cambodia Cameroon 1 Cameroon Central African Republic 1 Central Afr. Rep. Chad 1 Chad China 5 Absent, unreliable record China China Absent, doubtful record Uncertain Comoros 1 Comoros Present, restricted distribution Cote d'Ivoire 5 Absent, unreliable record Ivory Coast Ivory Coast Absent, doubtful record or intercepted only Croatia 3 Absent, intercepted only Croatia Absent, intercepted only Cyprus 6 Present, widespread Cyprus Cyprus Cyprus Cyprus Present, flourishes certain areas Czech Republic 3 Absent, intercepted only Czech Republic Absent, intercepted only Denmark 4 Absent, pest no longer present Denmark Denmark Absent, fomerly present Djibouti 1 Djibouti 6 Present, no details Egypt Egypt Egypt Egypt Present, flourishes certain areas El Salvador 1 El Salvador England 3 England England (i1916) Absent, fomerly present Equatorial Equatorial Guinea 1 Guinea Eritrea 1 Eritrea Ethiopia 4 Ethiopia Ethiopia Ethiopia Absent Finland 1 Finland France 2 France France Gabon 1 Gabon Gambia 5 Absent, unreliable record Gambia Gambia Absent, doubtful record Present, restricted distribution or intercepted only Germany 4 Absent, pest eradicated Germany Germany (i) Absent, eradicated Ghana 2 Ghana Ghana Guatemala 1 Guatemala Present, restricted distribution Guinea 5 Absent, unreliable record Guinea Guinea Absent, doubtful record or intercepted only Guinea Bissau, Guinea Bissau, Rep. 1 Rep. Hong Kong 2 Hong Kong Unknown Hungary 3 Absent, pest eradicated Hungary Absent, eradicated 6 Present, no details India India India India Present, flourishes certain areas Indonesia 4 Absent, pest no longer present Indonesia Indonesia Absent, fomerly present 6 Present, no details Iran Iran Iran Iran Present, flourishes certain areas Iraq 6 Present, no details Iraq Iraq Iraq Iraq Present, flourishes certain areas Present, restricted distribution Ireland 4 Absent, pest no longer present Ireland Absent, fomerly present or intercepted only Israel 6 Present, restricted distribution Israel Israel Israel Israel Present, flourishes certain areas Italy 4 Absent, intercepted only Italy Absent, intercepted only Uncertain Italy (Sicily) 1 Absent, doubtful record Present, restricted distribution Jamaica 3 Jaimaica Jamaica or intercepted only Present, restricted distribution Japan 5 Absent, pest eradicated Japan Japan (i) Absent, eradicated or intercepted only Jordan 2 Jordan Jordan Kazakhstan 1 Kazakhstan Kenya 4 Absent, pest no longer present Kenya Kenya Absent, fomerly present Korea, Republic 5 Present, no details Korea Korea Korea, Republic of Uncertain 2 Kuwait Kuwait Kyrgyzstan 1 Kyrgyzstan Laos 2 Laos Laos Lebanon 5 Present, no details Lebanon Lebanon Lebanon Lebanon Lesotho 1 Lesotho Liberia 2 Liberia Liberia Libya 5 Present, no details Libya Libya Libya Uncertain Luxembourg 3 Absent, pest no longer present Luxembourg Absent, fomerly present Madagascar 4 Madagascar Madagascar Madagascar unknown Malaya 1 Malaya Malaysia (peninsular) 1 Absent, doubtful record Malaysia 5 Absent, pest eradicated Malaysia Malaysia Absent, eradicated Absent (E), disagree (W) Malawi 1 Malawi Mali 6 Present, no details Mali Mali Mali Mali Present, flourishes certain areas Present, restricted distribution Mauritania 5 Present, no details Mauritania Mauritania Mauritania or intercepted only Mauritius 2 Mauritius unknown Mexico 4 Absent, pest no longer present Mexico Absent, fomerly present Absence, eradicated 5 Present, widespread Morocco Morocco Morocco Present, flourishes certain areas Mozambique 5 Absent, unreliable record Mozambique Mozambique Absent, doubtful record Uncertain Myanmar (formerly Burma) 5 Present, no details Myanmar Myanmar Myanmar Disagree Namibia 1 Namibia Netherlands 3 Absent, pest no longer present Netherlands Absent, fomerly present New Guinea 1 Absent New Zealand 3 Absent, pest no longer present New Zealand Absent, fomerly present Niger 5 Present, no details Niger Niger Niger Present, flourishes certain areas Nigeria 6 Present, restricted distribution Nigeria Nigeria Nigeria Nigeria Present, flourishes certain areas 1 Oman Pakistan 6 Present, no details Pakistan Pakistan Pakistan Pakistan Present, flourishes certain areas Philippines 5 Absent, unreliable record Philippines Philippines Absent, doubtful record Uncertain Poland 3 Absent, intercepted only Poland (i) Absent, intercepted only Portugal 4 Absent, intercepted only Portugal Absent, intercepted only Unknown 3 Qatar Qatar Qatar Present, restricted distribution Reunion Island 4 Absent, unreliable record Reunion Is. Absent, doubtful record or intercepted only Russia 4 Absent, pest no longer present Russia Absent, fomerly present Disagree Rwanda 1 Rwanda Saudi Arabia 5 Present, no details Saudi Arabia Saudi Arabia Saudi Arabia Saudi Arabia Senegal 6 Present, no details Senegal Senegal Senegal Senegal Present, flourishes certain areas Serbia and Montenegro 3 Absent, unreliable record Serbia Absent, doubtful record Present, restricted distribution Sierra Leone 5 Absent, pest no longer present Sierra Leone Sierra Leone Absent, fomerly present or intercepted only Singapore 2 Singapore Singapore unknown Slovakia 3 Absent, intercepted only Slovakia Absent, intercepted only Somalia 4 Present, no details Somali Republic Somalia Absent, eradicated South Africa 4 Absent, pest eradicated South Africa Absent, eradicated Absence, eradicated Spain 5 Present, restricted distribution Spain Spain Spain Uncertain (formerly Ceylon) 6 Present, no details Sri Lanka Sri Lanka Sri Lanka Sri Lanka Disagree Sudan 5 Present, no details Sudan Sudan Sudan Sudan Sweden 3 Absent, pest no longer present Sweden Absent, fomerly present Swaziland 1 Swaziland Switzerland 3 Present, restricted distribution Switzerland Switzerland 6 Present, widespread Syria Syria Syria Syria Uncertain Taiwan 5 Absent, pest no longer present Taiwan Taiwan Taiwan Uncertain Tajikistan 1 Tajikistan Tanzania 6 Absent, pest no longer present Tanzania Tanzania Tanzania Absent, fomerly present Absence, eradicated Thailand 5 Absent, unreliable record Thailand Thailand Absent, doubtful record Disagree Togo 1 Togo Tunisia 6 Present, widespread Tunisia Tunisia Tunisia Tunisia Present, flourishes certain areas Turkey 6 Present, restricted distribution Turkey Turkey Turkey Turkey Present, flourishes certain areas Uganda 1 Uganda Ukraine 2 Ukraine Ukraine (Odessa) Present, restricted distribution United Kingdom 2 Absent, pest no longer present or intercepted only 3 UAE UAE UAE United States of America 4 Absent, pest eradicated USA (i1946) Absent, eradicated Absence, eradicated Uruguay 2 Absent, confirmed by survey Uruguay Uzbekistan 2 Uzbekistan Uzebekistan Venezuela 4 Absent, pest no longer present Venezuela Venezuela Absent, fomerly present Vietnam 2 Vietnam Vietnam Yemen 5 Present, restricted distribution Yemen Yemen Yemen Yemen Yugoslavia 2 Yugoslavia Uncertain Zaire (formerly Congo) 1 Zaire Zambia 6 Present, no details Zambia Zamibia Zambia Zambia Uncertain Zanzibar 2 Zanzibar Uncertain Zimbabwe (formerly Rhodesia) 5 Present, widespread Zimbabwe Zimbabwe Zimbabwe Uncertain

References:

Australia2016: Department of Agriculture and Water Resources, (DAWR) 2017. Khapra beetle countries last revised 2016: http://www.agriculture.gov.au/import/before/pests/khapra_beetle There are more than 70 countries known to be infested with khapra beetle throughout Asia, the Middle East, Africa and Europe

Banks1977: Banks, H. J. 1977. Distribution and establishment of Trogoderma granarium Everts (Coleoptera: ): climatic and other influences. Journal of Stored Products Research 13(4): 183-202.

EPPO2016: European and Mediterranean Plant Protection Organization (EPPO) 2017. EPPO Global Data Base. Trogoderma granarium. Available at: https://gd.eppo.int/taxon/TROGGA https://gd.eppo.int/taxon/TROGGA/distribution

French2005: French, S. and R.C. Venette. 2005. Mini Risk Assessment, Khapra Beetle, Trogoderma granarium (Everts) (Coleoptera: Dermestidae). USDA–APHIS–PPQ–Cooperative Agriculture Pest Survey–Pest Risk Assessment (PRA); 22 pages.

Myers2012: Myers, S.W. and Hagstrum, D.W. 2012. Quarantine. p. 297–304. In: Hagstrum DW, Phillips TW, Cuperus G, editors. Stored Product Protection. Kansas State University, Kansas, USA.

SPIRes2009: Hagstrum, D. W., and Bh. Subramanyam. 2009. Stored-Product Resource. AACC International, St. Paul, Minnesota.