Mollusca, Pulmonata, Buliminoidea, Enidae
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TurkJZool 25(2001)427-429 ©TÜB‹TAK RediscoveryofSesteriagallandi Bourguignat,1884 (Mollusca,Pulmonata,Buliminoidea,Enidae) HartwigSCHÜTT Haydnstrasse50,D-40593Düsseldorf-Benrath. R›dvanfiEfiEN BiologyDepartment,ScienceandArtFaculty,DicleUniversity.TR-21280Diyarbak›r-TURKEY Received:04.10.2000 Abstract: Rediscoveryoftheland-snailSesteriagallandi Bourguignat,1884isreported.Theoriginaldescriptionwasbasedononly oneemptyshell,whichlaterwaslostandneverfoundagain,andtheexacttypelocalityinTurkeywasunknown.Nowwefoundthis speciesinMardinProvincenearthevillageofSariköyontheCizre-‹dil-Midyatroad20kmafter‹dilintheforest,inMay 2000. KeyWords: Mollusca,Pulmonata,Buliminoidea,Mardin,Taxonomicalstudy Sesteriagallandi Bourguignat,1884(Mollusca,Pulmonata,Buliminoidea,Enidae)'ün YenidenSaptanmas› Özet: Karasalyangozu Sesteriagallandi Bourguignat,1884yenidensaptanm›flt›r.Butürüntan›m›birtekboflkavk›üzerinde yap›lm›fl,dahasonrakaybolanbukavk›tekrarbulunamamifltir.‹lktan›m›nverildi¤içal›flmadalokalitetamolarakbelirtilmemifltir.Bu tür,Mayis2000de,MardiniliCizre-Idil-MidyatyoluüzerindeSariköyköyücivar›ndakiormanl›kalandayenidensaptanm›flt› r. AnahtarSözcükler: Mollusca,Pulmonata,Buliminoidea,Mardin,Taksonomikçal›flma Introduction CompletebibliographyofSesteria TheTurkishland-snail Sesteriagallandi was Thebibliographyofthegenus Sesteria hasbeen describedbyBourguignatin1884andhecreatedthe summarizedchronogicallyasfollows(3-12): monotypicgenusSesteria "Jeneconnais,decenouveau 1884Sesteriagallandi Bourguignat,Bull.Soc.Malac. genre,qu'uneespècerecueilliemorteaupieddes France,1:135-138,T.3F.1-5. arbustes,surlarivedroiteduTigre,entreMardinet Djezireh,lelongdelaroutedeDiarbekiràMossoul". 1887Sesteriagallandi,--Westerlund,Fauna,3:76. Thatmeanshehadonlyoneemptyshellofthisnew 1897Sesteriagallandi,--Westerlund,Synopsis,1:3. genusandspecies,whichwasfoundatthebottomof 1898 Sesteriagallandi ,--Kobelt,Zoogeographie,2: bushesabovetherightbankalongtheDicleNehri 76. (Tigris)betweenMardinandCizre,alongtheoldroad fromDiyarbakirtoMosulinIraq(1). 1901Sesteriagallandi,--Kobelt,Martini-Chemnitz,1. 13(2):707,T.105,F.14-16. AsBourguignatwrote,theroadconstructionengineer J.Gallandfoundonlyoneemptyshellandhegaveno 1901Sesteriagallandi,--Kobelt,Iconographie,(2)9 moredetailedinformationaboutthetypelocalitythan (3/4):36,T.254F.1637. thatmentioned.For116yearsafterthedescription,this 1903 Sesteriagallandi ,--Kobelt,Nachr.Bl.dtsch. uniquespeciesneverwasfoundagainbecauseitsexact malakozool.Ges.,35(7/8):118. localitywasunknownandthissingleshell,inthe 1931Sesteriagallandi,--Thiele,Handbuch,1:522. collectionofBourguignatinMuseumGenève, 1940Sesteriagallandi,--Forcart,Verh.Naturf.Ges. Switzerland,waslost(2). Basel,51(1):133. 427 RediscoveryofSesteriagallandi Bourguignat,1884(Mollusca,Pulmonata,Buliminoidea,Enidae) 1959 Sesteriagallandi ,--ZilchinWenz,Handb. characterforthegenus Sesteria.Theothercharacter Paläont.,6(2,1):184,Abb.636. mentionedbyBourguignatalsocannotbeaccepted,since 1996 Sesteriagallandi ,--Schütt,Landschneckender weknowthespecies Buliminus (Pene)brunneus (P. Türkei:129,Textfig. Hesse,1914)[synonym Buliminus(Pene)coluplicatus Gittenberger&Menkhorst,1991],whichshowsan 1998Sesteriagallandi,--Schileyko,Treatise,2:209, equallypronouncedendofthecolumellainitsaperture, Fig.258. buthasabulgiershell. Therefore,werecommendthecancellationofthe Material taxon Sesteria andthedesignationofthissoutheast AfterlongendeavoursinMay2000wefoundshellsof Anatolianspeciesas Buliminusgallandi (Bourguignat, Sesteriagallandi ,whichenableustospecifythetype 1884). locality.TheTurkishprovinceofMardinontheCizre-‹dil Thusfar,circumstanceshavenotallowedustofind -Midyatroad20kmafter‹dilinthevillageSariköy,near livinganimalsforanatomicaldissection,butthiswillbe theoldroadfromMardintoMosul,inaforestwith doneinthefuture.AtSariköywefoundsevenshells, great-leafyoaks(37º17'N41º37'E).Wealsofounditat whicharedepositedinSenckenbergMuseumFrankfurt, asecondlocationabout23kmwestofCizreontheway Germany,andinNaturhistorischesMuseumWien, to‹dil.Inthespacesaroundtheseplacesmusthavebeen Vienna,Austria,aswellasinthecollectionsofboth thelocalitywhereGallandfoundhistypicalshell. authors. Associatedmalacologicalfaunaatbothplacesfollows: DiscussionandResults Orculellapalatalis (Pilsbry,1922), Orculella sirianocoriensislibanotica (Tristram,1865), Buliminus Thedecisivediagnosticmarkofthegenus Sesteria Bourguignat,1884isthehollowandcorkscrew-like undulatedaxisofthecolumella:"l'axecolumellaire, Bull.Soc.malac.FranceJuin1884. PL.III. tubulairedanstoutesalongueur ,prend, àpartirde 5 2 l'avant-derniertour (pl.III,f.5)unetorsionsiforteetsi 1 accentuée,qu'ilressembleàuntire-bouchonà circonvolutionstrèsexcentriques ".Nearlynothingneeds tobeaddedtotheoriginaldescriptionbyBourguignat, apartfromthefactthatinourshellsthelowermostpart ofthecolumelladoesnotoverlaptheaperturetothe extentdepictedinhisfigure(Figure1).Also,ourshells aresomewhatshorterthanhis.Thehighestshellofours is21mm,whilehissingleshellwas"alt.25millim."Itis worthmentioningthattheborderoftheapertureatthe areaoftheangularlumpisespeciallythickened,andthis andtheupperinsertionoftheperistomeformasmall canal. Wecheckedthehollowandcorkskrew-likeundulated 3 4 axisofthecolumella"l'axecolumellaire, tubulairedans toutesalongueur",andcomparedthisdecisivediagnostic markofthegenuswithotherTurkishspeciesof Buliminus,anddeterminedthatallspecieshaveahollow 1-5.SesteriaGallandi.- axisaslongasthetotalshell:B.akkumensis Gittenberger &Menkhorst,1991, B.alepensis (L.Pfeiffer,1841), B. Maubertdel. Imp.Becquetfr.Paris. carneus (L.Pfeiffer,1846), B.labrosus (Olivier,1804). Figure1. ReproductionoftheoriginaldrawingofBourguignat, Therefore,thiscannotbeusedasarealdiagnostic 1884. 428 H.SCHÜTT,R.fiEfiEN alepensis (L.Pfeiffer,1841), Buliminusgallandi Acknowledgement (Bourguignat,1884), Eopolitaderbentina (O.Boettger, WethankMissBrigitteGraack,Wedemark,Germany, 1886), Assyriellaescheriana (Bourguignat1864), forthephotographs(Fig.2). Assyriellamardinensis (Kobelt,1900), Helixcheikliensis Zilch,1952. Figure2. Buliminusgallandi,specimenfrom Sariköyköyü,Mardin-Turkey. Scalebar=5mm,photo:B. Graack. References 1. Bourguignat,J.R.DescriptiondunouveaugenreSesteria.Bulletin 7. Kobelt,W.IconographiederLand-&Süsswasser-Mollusken,Neue delaSociétéMalacologiquedeFrance,1:135-138,pl.3.1884. Folge,9:1-86,Taf.241-276,1901. 2. Forcart,L.MonographiederTürkischenEnidae(Moll.,Pulm.). 8. Kobelt,W.UnsereKenntnissderFaunaEuropaea.Nachrichtenblatt VerhandlungenderNaturforschendenGesellschaftBasel,51(1): derDeutschenMalakozoologischenGesellschaft,35(7/8):113- 106-263.Tafel1-3.,1940. 121,1903. 3. Westerlund,C.A.FaunaderinderPalaearctischenRegion 9. Thiele,J.HandbuchdersystematischenWeichtierkunde.1:I- lebendenBinnenconchylien,3:Buliminus etc.:1-183,Register1- VI,1931Jena,778p. 15.Lund,1887. 10. Zilch,A.&Wenz,W.HandbuchderPalaeozoologie,6(2):I-XII, 4. Westerlund,C.A.Synopsismolluscorumextramarinorumregionis 1959Berlin,834p. palaearcticae.Fasc.1GeneraetspeciesextypisBulimietPupae. 11. Schütt,H.LandschneckenderTürkei.--ActaBiologicaBenrodis, Lund,1897. Supplement4,Solingen1996.497p. 5. Kobelt,W.StudienzurZoogeographie.2:DieFaunader 12. Schileyko,A.A.TreatiseonRecentTerrestialPulmonateMolluscs. MeridionalenSub-Region,I-X,1898,Wiesbaden,368p. Part2.Gastrocoptidaeetc.Ruthenica,Supplement2:Moscow, 6. Kobelt,W.DieFamilieBuliminidae.In:Martini&Chemnitz, 1998,128-261. SystematischesConchylien-Cabinet,1.13(2):FamilieBuliminidae: 397-1051,Taf.71-133.1901. 429.