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Svensk Mykologisk Tidskrift Volym 38 · Nummer 2 · 2017 Svensk Mykologisk Tidskrift �������������������7 Svensk Mykologisk Tidskrift Volym 38 · nummer 2 · 2017 Svensk Mykologisk Tidskrift 1J@C%RV`:` 1R1$:`7 www.svampar.se 0VJ@7@QCQ$1@0 Sveriges Mykologiska Förening 0VJ]%GC1HV`:`Q`1$1J:C:` 1@C:`IVR0:I]R Föreningen verkar för :J@J7 J1J$QH.IVR0VJ@ QH.JQ`RV%`Q]V1@ R VJ G?`V @?JJVRQI QI 0V`1$V 0:I]:` QH. 1J `VV80VJ% @QIIV`IVR`7`:J%IIV` 0:I]:``QCC1J: %`VJ ]V`B`QH.?$:00V`1$V7@QCQ$1@:DV`VJ1J$8 R@7RR:0J: %`VJQH.:0:I]]CQH@J1J$QH.- : `%@ 1QJV` 1CC`V``: :`V`1JJ]BD7.VI1R: J: %]] `?R:JRV1@Q$QH.I:`@@V`%JRV`1:@ - 11180:I]:`8V80VJV`.BCC$VJQI- :$:JRV:0$?CC:JRVC:$:` CVI@:] 1 D80VJ ``:I ?CC IVR G1R`:$ R:@QJ :@ V`IVCC:JCQ@:C:0:I]`V`VJ1J$:`QH. ``BJ0Q`V=: .Q` I1JJV`QJR8 0:I]1J `VV`:RV1C:JRV %JRV`C?: R:@QJ :@ %]]`?.BCCIVRI7@QCQ$1@:`V`- $:`1$`:JJC?JRV` R VJ :I0V`@:J IVR I7@QCQ$1@ `Q`@J1J$ QH. Redaktion 0V VJ@:]8 HVR:@ V`QH.:J0:`1$% $10:`V 1@:VCIQJ VRCVI@:]V`.BCCV$VJQI1J?J1J$:0IVRCVIR JK=: :J :0 VJ]B`V`VJ1J$VJG:J@$1`Q 0HLJJKN00<= 5388-7733 =0:I]:`8V VRCVI:0 VJ` 7 [ 7`V`IVRCVII:`GQ::10V`1$V Jan Nilsson [ 7`V`IVRCVII:`GQ::% :J`V`0V`1$V IVGV`$ [ 7 `V` %RV`:JRVIVRCVII:`GQ::1 ;JJ<H6= 0V`1$^6 _ [ 7 `V``=^=0_ =$8V %GH`1]``QI:G`Q:R:`V1VCHQIV84:7IVJ Äldre nummer :00VJ@7@QCQ$1@0^ `Q` ^69 _H:JGVI:RVG7H`VR1 H:`RG7 ILHE0INH=<=_`1JJ]BVJAEQI@:JGV ?CC: 01Q%`1VG.Q]: 11180:I]:`8VQ` QQ%` ``BJD8 G:J@:HHQ%J7 Previous issues Q` 0VJ@ 7@QCQ$1@ 0 ^ILHE0INH=<=_:`V:0:1C:GCVQJ:AE1 :;<=76 H:JGVQ`RV`VR``QID8 ;:AL:D076E6 Sveriges Mykologiska Förening :=V VJ@:] GVGQ`$%J10V`1 V ;6 GVGQ`$ 11180:I]:`8V Omslagsbild 7]`11B`V `Q Q ?0C1J$$1H@ 1CC60:< VR IVRG1CRVJ:0$C1 V`GC?H@0:I]^Coprinellus micaceus 8_DQ Q ?` :$V ]BG 7`VCVQH.`VR:@ 1QJ$`: %CV`:`8 I detta nummer nr 2 2017 *_77`J`7 C*'3 2 QCG1% H:CC1 % , papegojskivling, 58 *?$VJ0:I]:`1J`V`VR :J funnen i Sverige 2020 patrick Björck & Aleksandar Tudzarovski Jan-Olof Tedebrand 7 Q]`QC:0:I]:`1@BJV5 1 V :@BJVRV`R V`]B SMF $:0V V: % 79 HI0.RV% Sven-Åke Hanson I7@$ @% %:`0 22 :H :`1% V].VJ 11, en hypogeisk riska, Jan Nilsson & Mikael Jeppson funnen i Sverige Elna Hultqvist & Mikael Jeppson ? V .7 I: i Sverige Jörgen Jeppson & Mikael Jeppson *'377`J` 46 *? R0:I]`7R1QI`R $ Q`:9 =? 1;`=?^? : 9V R_ ?8A%QJV`. :$$0:I]:` Thomas Grönlund C1 QH7GV:$`VHarmaja. Svenskt namn saknas. Läs mer i 0K SVENSK MYKOLOGISK TIDSKRIFT 38:2 (2017) 1 *_77`J`7 0VJ@7@QCQ$1@0 ^_7–6 2017 QCG1%H:CC1 %, papegojskivling, funnen i Sverige 4J-W;U-W<Y7W`3036*WJ ;,- QCG1% H:CC1 % `%R1R8 0R1H%V .VQ`:J%J%%:C]VH1VQ`QCG1%. In October 2016 the :% .Q`011 VR:1J V`V`%J$%CQH:C1 71J:CI@V]1J$51VRVJ8L:]1CVQ`1- H.1] .V7 %IGCVR1J Q:]VH%C1:`:$:`1H80HQCQ%`5<V:JRH.:`:H V`1Q` .V GQR1V1JR%HVR:JV6 VJRVRI:H`QR:JRI1H`QIQ`].QCQ$1H:CV6:I1J:5CV:R1J$ Q .V HQJHC%1QJ .: 1 1::]VH1V1J;5$VJ%QCG1%8:1:]`Q011QJ:CC71RVJ- :QCG1%]1:H1J%K5< 34K<A1V`VVJ Q<:C:G1J ]:1J`_<1`V0V:CVR .: .V1VR1.HVH C7I: H.VR: _``QI .V.QCQ 7]VQ`8]1:H1J% deposited in GenBank. Moreover, a GenBank sequence of the American species 8H:CC1 %5Q`1$1J:CC7RVH`1GVRG74VH@ :Agari- H%H:CC1 %5 %`JVRQ% QGV1RVJ:C8Q`].QCQ$1H:CR: :Q`Q%`1V`VHQI]:`VR 1RVH`1]1J .VC1 V`: %`VQ`GQ .8]1:H1J% and 8H:CC1 % (Peck) Watling and 1JR1H: VRH8V .%HQJ1RV`8]1:H1J%:7Q%J$V`7JQJ7IQ`8H:CC1 % 1:V`V]Q` VR`6Q]V8 ?R$ lar intensivt färgade. Riktigt unga svampar var I mitten av oktober 2016 besökte vi en lokal där ärggröna, övergående i mörkrött och bleknande i vi tidigare hittat många intressanta arter, bland tegel till terracotta på utvuxna exemplar. Skivor- annat Pholiota squarrosoides (kryddtofsskiv- na gav ett blågrått intryck på unga exemplar och ling) och Leucoagaricus meleagris. Lokalen är övergick i ockra på spormogna fruktkroppar. ett före detta grustag strax utanför Malmköping Färgskalan skiftade mycket på de exemplar vi i Södermanland som idag, efter att täkten upp- hittade men alla tillhörde samma art. Svampen \ liknade inget som någon av oss tidigare sett och \ väckte förstås vårt intresse. En stunds pillande \ och dissekerande kombinerat med litteraturstud- [! ier ledde oss in på släktet Bolbitius (guldskivling- löv- och barrved, framförallt gran, asp, björk och ar). Fruktkropparna var något bräckliga, foten tall men med en stor andel ädellöv som lind, ek var ihålig och såväl habitus i stort som växtsättet och alm. Flislagrets tjocklek varierar och är < 2 på substratet gjorde att svampen var en uppen- – 50 cm, men är på enstaka platser mer än meter- "%& djupt. Lokalen är alltså ett typexempel på en bio- Bolbitius enligt gängse litteratur sakna arter med top där den människoskapade livsmiljön är stadd den färgskala vi såg. Vi fann dock en från Tjeck- i ständig förändring då nytt timmer kontinuerligt ien nybeskriven art, B. psittacinus, som verkade \ " vara en perfekt matchning mot vårt fynd. \ #$\ K$5I:@``$ cirka femtio kvadratmeter i ett drygt hundratal Som ung är svampens hatt konisk och blir i takt exemplar. Fruktkropparna var hos unga exemp- med fruktkroppens mognad välvd till nästin- 2 SVENSK MYKOLOGISK TIDSKRIFT 38:2 (2017) *_77`J`7 till plan, oftast med en liten, ibland lätt rynkad har även som äldre en grön anstrykning eller umbo, ofta även med strimmig, men aldrig fårad framstår som helt olivgröna. Doft och smak hos hattkant. Hatten är särskilt hos unga fruktkrop- fruktkropparna är svag, inte distinkt, men upp- par något klibbig i väta. Skivorna är vidvuxna, fattas något obehaglig och påminner om jord- på vissa exemplar med en liten nedlöpande an- källare eller inte helt fräscha rotfrukter, rätt lik tydan, relativt täta. Skivfärgen är blågrå, ibland doften hos Amanita citrina*+ nästan turkos hos unga exemplar för att sedan bli gråvita och så småningom ockrafärgade. På K$5I1@``$ äldre exemplar är skiveggen aningen ljusare än Sporer: 6,4 – 9 × 4,1 – 5,5 μm (medel 7,7 × 4,8 skivorna. På ett fåtal äldre exemplar syns anty- μm), elliptiska, något mandelformade, med slät, dan till anastomoser mellan skivorna, men det är /);/) inget konsekvent karaktärsdrag. Hattens bredd Rostfärgade till orangegula i KOH (3%). varierar hos de utvuxna exemplaren mellan 25 Basidier: fyrsporiga, med enstaka tvåsporiga, och 80 mm. Den är i regel också lätt ojämn, med ca 15,5 – 21 x 8 –/>?) ibland något vågig hattkant. Foten är 30 – 60 Cheilocystidier: (15–) 19 – 43 x 7,5 –//) mm hög, 3 – 8 mm bred, ihålig. Fotens hålighet mestadels utriforma, sällan svullna, med förlängd har hos unga exemplar oftast en tydligt blå ton. @>) ' Pleurocystidier: saknas. \ $ Caulocystidier: />>$/>) med en blå anstrykning. Den är något oregelbun- Pileipellis: globosa element, 29 – 33 x 18 – AB?) det cylindrisk med en lätt förtjockad bas där vitt Pileocystidier: saknas. mycelludd, ibland även där med blå anstrykning, Söljor: förekommer hos hyferna i tramat. oftast förekommer. Hattens färg varierar extremt mycket både med utvecklingsstadiet och mellan A7?%RV$ olika fruktkroppar. Som ung är färgen blågrön ITS-regionen för nukleärt ribosomalt DNA hos till grön och övergår i mörkröd, ibland nästin- vår kollekt sekvenserades vid Alvalab i Spanien till svart på ännu inte spormogna exemplar som $ ! inte har fullt utvecklad hatt. I takt med att hatten en sekvens bestående av 662 baspar. Tillgäng- öppnar sig förändras också färgskalan från rött liga och representativa ITS-sekvenser av Bol- till orange, ockra och gult. Många fruktkroppar bitius J" tillsammans med vår nya sekvens. Som utgrupp användes Laccaria laccata (laxskivling) och Lactarius rufus (pepparriska). Alignmenten med efterföljande maximum likelihood- och boot- strapanalys gjordes i mjukvaran MEGA 7. Det fylogenetiska trädet med bootstrapvärden pre- / Den fylogenetiska analysen visar att vår kollekt från Sörmland, tillsammans med den tjeckiska typkollekten av B. psittacinus och den enda i Genbank tillgängliga sekvensen av B. callistus, D E `7CQ$: `?RV 01:` : 0B` `7JR bildar en helt egen klad med hundraprocentigt (`V _ ``BJ :CI@V]1J$5 V :IV`1@:J@ `7JR :0 bootstrapstöd. QCG1% H:CC1 % ^6 _ QH. .QCQ 7]VJ :0 B. ]1:H1J% ^6D _ .:` 1RVJ: :0VJV`8 ; `:]0?`RVJ:J$V01RJQRV`J:86 V`0=:` - Ny för Sverige J:IJ:J$VRVJ:J:C7V`:RVV@0VJVJ1RRJ%IIV` De tre kollekter vars ITS-sekvenser här jämförs, i GenBank. dvs typkollekten av Bolbitius psittacinus från SVENSK MYKOLOGISK TIDSKRIFT 38:2 (2017) 3 *_77`J`7 Fig. 2. QCG1%H:CC1 %^?]:]V$=?_D @`Q]]:`]B1VJ:`V:JR ?@ 85- köping, 2016-10-13, leg. & det. P. Björck & A. Tudzarovski (herb. GB). Foto p. Björck. Tjeckien, kollekten av B. callistus (som B. cal- En ovanlig art som är liten, kan undgå upptäckt listeus) från Nordamerika och vårt svenska fynd, under lång tid. De tre europeiska fyndens geo- bör alltså tillhöra samma art. Namnet B. callis- [ tus har prioritet, eftersom det går tillbaka till en kanske för en undanskymd tillvaro i biotoper beskrivning gjord av den amerikanske mykolo- som sällan besöks, såsom kärrliknande och gen Peck redan 1874 (som Agaricus callistus). våta miljöer, som de Peck (1874) anger i den Den tjeckiska beskrivningen av B. psittacinus ursprungliga beskrivningen av arten. I Nord- publicerades 2007. I samband med att A. callis- amerika förekommer arten i svämskogar, vid tus\Bolbitius av Watling kärr och vattenhål, liksom också på vedrester i (1977), råkade han felaktigt använda epitetet människoskapade miljöer enligt Watling & Mil- callisteus, dvs med ett extra e jämfört med origi- ler (1987).
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