Buckinghamshire & Milton Keynes Environmental Records Centre

Tring Road  Halton  Aylesbury  Bucks  HP22 5PN Tel 01296 696012 Fax 01296 624519 www.bucksmkerc.org.uk

Craneflies and Winter Gnats (Diptera: ) recorded in Buckinghamshire

DRAFT 1. Author: Martin C. Harvey. Date: 15 March 2007 (with minor edits at 15 October 2008).

Based on data held on the MapMate database by BMERC (data retrieved 15 March 2007), and including all species covered by the national cranefly recording scheme (craneflies, , plus fold-winged craneflies, Ptychopteridae, and winter gnats, Trichoceridae). This is a very provisional document – the data has not yet been fully validated, and has not been checked with the national scheme. Records have been contributed by around 30 people, especially Ken Merrifield, Martin Harvey, Tony Marshall and Alan Stubbs.

1. Species list

There are approximately 550 records of hoverfly for Buckinghamshire on the database, comprising 90 species: 37 long-palped craneflies, 41 short-palped craneflies, 7 winter gnats and 5 fold-winged craneflies. There are 339 British species altogether.

Long-palped craneflies

Taxon Authority Records Last Status Taxon Authority Records Last Status Recorded Recorded Ctenophora Meigen, 1 2005 RDB1 Tipula livida van der 1 2001 Scarce ornata 1818 Wulp, 1858 Ctenophora (Fabricius, 1 2005 RDB2 Tipula lunata Linnaeus, 7 2006 flaveolata 1794 ) 1758 Ctenophora (Linnaeus, 7 2005 Scarce Tipula vernalis Meigen, 31 2006 pectinicornis 1758 ) 1804 Dictenidia (Linnaeus, 1 1984 Tipula Meigen, 8 1991 bimaculata 1761) varipennis 1818 Prionocera (Fabricius, 2 1981 Tipula confusa van der 3 2006 turcica 1787) Wulp, 1883 Nephrotoma (Pierre, 25 2006 Tipula Meigen, 12 1989 appendiculata 1919) marmorata 1818 Nephrotoma (Linnaeus, 3 1981 Tipula obsoleta Meigen, 3 2005 cornicina 1758) 1818 Nephrotoma (Linnaeus, 14 2006 Tipula pagana Meigen, 25 2006 flavescens 1758) 1818 Nephrotoma (Meigen, 14 1983 Tipula rufina Meigen, 7 1984 flavipalpis 1830) 1818 Nephrotoma (Westhoff, 1 1978 Tipula signata Staeger, 3 1978 guestfalica 1879) 1840 Nephrotoma (Meigen, 17 1997 Tipula staegeri Nielsen, 2 1982 quadrifaria 1804) 1922 Nephrotoma Edwards, 1 1978 Tipula oleracea Linnaeus, 21 2001 submaculosa 1928 1758 Tipula De Geer, 5 2005 Tipula paludosa Meigen, 50 2006 fulvipennis 1776 1830 Tipula luna Westhoff, 1 1988 Tipula czizeki de Jong, 1 1977 1879 1925 Tipula maxima Poda, 1761 17 2006 Tipula Alexander, 1 1972 Tipula vittata Meigen, 2 1982 subcunctans 1921 1804 Tipula hortorum Linnaeus, 1 1982 RDB3 Tipula Meigen, 7 2006 1758 flavolineata 1804 Tipula scripta Meigen, 15 1997 Tipula cava Riedel, 11 1998 1830 1913 Tipula lateralis Meigen, 5 1983 Tipula Meigen, 8 2006 1804 fascipennis 1818

The Buckinghamshire and Milton Keynes Environmental Records Centre is a service provided by Buckinghamshire County Council and Milton Keynes Council

Craneflies in Buckinghamshire

Short-palped craneflies

Taxon Authority Records Last Status Limonia (Schrank, 3 2006 Recorded phragmitidis 1781) Achyrolimonia (Loew, 1873) 3 2006 Limonia (Fabricius, 12 1995 decemmaculata tripunctata 1782) Austrolimnophila (Meigen, 20 2006 (Staeger, 3 1989 ochracea 1804) appendiculatus 1840) (Meigen, 8 1998 Molophilus de Meijere, 3 1974 cinerascens 1804) cinereifrons 1920 Cheilotrichia (Meigen, 1 1974 Scarce Molophilus (Meigen, 1 1999 imbuta 1818) griseus 1804) Dicranomyia (Meigen, 14 2006 Molophilus de Meijere, 1 2006 chorea 1818) occultus 1918 Dicranomyia (Meigen, 4 2006 Neolimnomyia (Meigen, 5 1983 mitis 1830) nemoralis 1818) Dicranomyia (Meigen, 1 1999 Neolimonia (Meigen, 5 2006 modesta 1818) dumetorum 1804) Dicranomyia (Meigen, 1 1980 Edwards, 2 1974 sericata 1830) albitibia 1921 Dicranota (Schummel, 2 1997 Ormosia lineata (Meigen, 1 1983 bimaculata 1829) 1804) Epiphragma (Linnaeus, 3 2006 Ormosia (Macquart, 2 1997 ocellare 1761) nodulosa 1826) (Walker, 1 1999 Phylidorea (Schummel, 2 2006 diuturna 1848) fulvonervosa 1829) Erioptera lutea Meigen, 1804 1 2006 Pilaria discicollis (Meigen, 1 1990 Euphylidorea (Meigen, 4 2006 1818) dispar 1818) Rhipidia authors, 5 1982 Euphylidorea (Meigen, 3 1983 duplicata misident. lineola 1804) Rhipidia Meigen, 1818 20 2007 Helius flavus (Walker, 2 2006 maculata 1856) Goetghebuer, 2 1999 maculata (Meigen, 3 1988 bifurcatus 1920 1804) (Meigen, 3 2006 Ilisia occoecata Edwards, 2 1974 stictica 1818) 1936 Thaumastoptera Mik, 1866 2 1990 Scarce Limonia flavipes (Fabricius, 6 1997 calceata 1787) Tricyphona (Meigen, 5 1982 Limonia (Schummel, 6 2006 immaculata 1804) nigropunctata 1829) Trimicra pilipes (Fabricius, 3 1974 Limonia Meigen, 1804 29 2006 1787) nubeculosa Ula sylvatica (Meigen, 3 1989 1818)

Winter gnats

Taxon Authority Records Last Status Trichocera Meigen, 1 1999 Recorded parva 1804 Trichocera Meigen, 20 2006 Trichocera (Linnaeus, 16 2006 annulata 1818 regelationis 1758) Trichocera (De Geer, 4 1985 Trichocera Edwards, 1 1983 hiemalis 1776) rufescens 1921 Trichocera Edwards, 5 1983 Trichocera (Harris, 11 2007 major 1921 saltator [1776])

Fold-winged craneflies

Taxon Authority Records Last Status Ptychoptera Meigen, 1 1988 Recorded lacustris 1830 Ptychoptera (Fabricius, 6 1981 Ptychoptera Tonnoir, 1 1989 albimana 1787) minuta 1919 Ptychoptera (Linnaeus, 5 1990 Ptychoptera Meigen, 1 1988 contaminata 1758) paludosa 1804

BMERC, March 2007 Craneflies in Buckinghamshire

2. Top 10 most frequently recorded species 3. Top 9 most widespread species

Species Records Species Tetrads Tipula paludosa 50 Tipula paludosa 25 Tipula vernalis 31 Tipula vernalis 22 Limonia nubeculosa 29 Nephrotoma appendiculata 21 Tipula pagana 25 Tipula oleracea 16 Nephrotoma appendiculata 25 Tipula maxima 13 Tipula oleracea 21 Limonia nubeculosa 13 Rhipidia maculata 20 Tipula pagana 10 Austrolimnophila ochracea 20 Austrolimnophila ochracea 10 Nephrotoma quadrifaria 17 Dicranomyia chorea 9 Tipula maxima 17

5. Recorded tetrads in Buckinghamshire

6

5

4

3

2

1

SP TL

9

8

7 SU TQ 5 6 7 8 9 1

For further information about the national cranefly recording scheme please contact: John Kramer, 31 Ash Tree Road, Oadby, Leicester LE2 5TE See also www.dipteristsforum.org.uk

Records can be sent in to the above scheme or to BMERC (we will pass them on): Buckinghamshire and Milton Keynes Environmental Records Centre email: [email protected] - web: www.bucksmkerc.org.uk phone: 01296 696012 - post: BMERC, Museum Resource Centre, Tring Road, Halton, Aylesbury, Buckinghamshire, HP22 5PN

BMERC, March 2007 Craneflies in Buckinghamshire

Identification resources

A number of draft keys have been produced by Alan Stubbs for the Cranefly Recording Scheme. Alan is working on a cranefly book that will be published by the British Entomological and Natural History Society in a year or two. Copies of these keys are available from BMERC for anyone who is recording craneflies. Alan would welcome any feedback from people using his keys.

Order Key Level Classification Date A4 pages 1 Key to families Family Tipuloidea families, + Undated 2 Trichoceroidea: Trichoceridae 2 Keys to Tipulidae Genus Tipulidae April 3 and 19(94?) Family Cylindrotomidae Species Cylindrotomidae April 1994 3 Test key for Tipula Species Tipulidae – Tipula April 1997 16 4 Keys to long-palped Species Tipulidae except Tipula Undated 4 craneflies other than Tipula 5 Test key – hairy-eyed Species Pediciidae Undated 6 craneflies: Pediciidae [6 – Key to Limoniidae Subfamilies Limoniidae missing] 7 Test key to (= Species Limoniidae – Chioneinae (plus March 2001 14 ) – key 1: open all other species with an open discal cell discal cell) 8 Test key to Chioneinae (= Species Limoniidae – Chioneinae July 2001 17 Eriopterini) – key 2: closed discal cell 9 Test key – Molophilus Species Limoniidae – Chioneinae – April 1994 5 Molophilus 10 Key to British species of Species Limoniidae – Chioneinae – [Freeman, 1 Skuse Tasiocera 1951) [9 – Key to Dactylolabinae (two Species Limoniidae – Dactylolabinae missing] species) 10 Test key to subfamily Species Limoniidae – Limnophilinae February 9 Limnophilinae 1997 11 Test key to subfamily Species Limoniidae – January 15 Limoniinae 1998 13 Test keys to Trichoceridae Species Trichoceridae and September 5 (wintergnats) and fold- Ptychopteridae (not part of 2002 winged craneflies Tipuloidea but included in the (Ptychopteridae) recording scheme)

Also available:

Item Classification Date A4 pages Test key: larvae and pupae in dead wood Some Tipulidae and some February 5 Limoniidae 1998 Wing diagrams for various species Various - 8 ‘Rough guides’ to: orange and yellow craneflies; black Various - 2 or blackish species; Genitalia illustrations for various species Tipulidae - 9

Other references:

Stubbs, A.E. 1992. Provisional atlas of the long-palped craneflies (Diptera: Tipulinae) of Britain and Ireland. Biological Records Centre, Monks Wood. Stubbs, A.E. 1993. Provisional atlas of the ptychopterid craneflies (Diptera: Ptychopteridae) of Britain and Ireland. Biological Records Centre, Monks Wood. [Includes key to Ptychopteridae, more detailed than item 13 above.]

BMERC, March 2007