Guide to British Dung : APHODIINI

D: Distribution area (see map) Months most likely to find Widespread species Months possible to find Rare species Months not usually found

ACROSSUS ACROSSUSAGOLIINUS

depressus luridus rufipes lapponum nemoralis 6‐9 mm D: 1‐9 6‐9 mm D: 2‐9 9‐13 mm D: 1‐9 4‐6 mm D: 1‐4, 9 4‐6 mm D: 1‐2 All habitats Well drained soils All habitats Upland habitats Upland, shaded habitats J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D

AGRILINUS AGRILINUSAPHODIUS APHODIUS BODILOIDES

ater constans fimetarius foetidus pedellus ictericus 4‐6 mm D: 1‐9 5‐6 mm D‐ 4‐9 5‐8 mm D: 1‐9 5‐8 mm D: 1‐9 5‐8 mm D: 1‐9 3‐6 mm D: 1‐9 All habitats Prefers shaded habitats All habitats Grasslands All habitats Well drained soils J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D

BODILOPSIS BODILOPSISCALAMOSTERNUS CHILOTHORAX CHILOTHORAX CHILOTHORAX

rufa sordida granarius conspurcatus distinctus paykulli 5‐7 mm D: 1‐9 5‐8 mm D: 1‐9 3‐5 mm D: 3‐9 4‐5 mm D: 2‐8 3‐6 mm D: 2‐8 3‐5 mm D: 1‐8 All habitats Well drained soils Grasslands Shaded habitats, Prefers sandy soils Well drained soils sometimes dunes in North J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D

COLOBOPTERUS ESYMUS ESYMUS EUHEPTAULACUS EUHEPTAULACUS EUORODALUS

erraticus merdarius pusillus sus villosus coenosus 6‐9 mm D: 1‐9 4‐5 mm D: 1‐9 3‐5 mm D: 1‐9 4‐5 mm D: 6‐8 3‐5 mm D: All 3‐5 mm D: 6‐8 Grasslands Grasslands Grasslands Well drained soils Well drained soils Well drained soils J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D HEPTAULACUS LIMARUS LIOTHORAX

testudinarius lividus zenkeri niger plagiatus 3‐4 mm D: 7 3‐6 mm D: 5‐7 4‐6 mm D: 3‐8 4‐5 mm D: 7 4‐5 mm D: 1‐9 Sandy soils Well drained soils Shaded habitats Poolside debris Coastal, organic debris

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MELINOPTERUS MELINOPTERUS MELINOPTERUSNIMBUS NIMBUS

consputus prodromus sphacelatus contaminatus obliteratus 3‐6 mm D‐ 6‐8 4‐7 mm D: 1‐9 4‐6 mm D: 1‐9 5‐7 mm D: 1‐9 4‐7 mm D: 1‐8 Well drained soils All habitats All habitats All habitats Shaded habitats J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D

OTOPHORUS OXYOMOUS PHALACRONOTHUS PLAGIOGONUS PLANOLINUS PLANOLINUS

haemorrhoidalis sylvestris quadrimaculatus arenarius borealis fasciatus 3‐6 mm D: 2‐7 3‐4 mm D: 3‐8 3‐4 mm D: 3‐8 3‐5 mm D: 3,6,8 3‐6 mm D: 1‐9 3‐5 mm D: 1‐4 Grasslands Organic debris & Well drained soils Well drained soils Shaded habitats & Upland, often Manure heaps dunes in North shaded habitats

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RHODAPHODIUS SIGORUS TEUCHESTES VIOLINUS

foetens porcus fossor sticticus 6‐9 mm D: 1‐9 4‐6 mm D: 2‐9 6‐12 mm D: 1‐9 4‐6 mm D: 2‐9 Grasslands Grasslands Grasslands Shaded habitats

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