Early Land Plants Today: Index of Liverworts & Hornworts 2006-2008

EarlyLand Plants Today: Indexof Liverworts &Hornworts 2006-2008

Matt von KONRATa*,Lars SÖDERSTRÖMb ,Anders HAGBORGa , Marshall R.CROSBYc &John J.ENGELa

a Departmentof Botany,The Field Museum,1400 SouthLake ShoreDrive, Chicago,IL60605-2496,U.S.A.

b Departmentof Biology,NorwegianUniversity of Scienceand Technology, N-7491,Trondheim,Norway

c MissouriBotanicalGarden,P.O.Box299,St.Louis,MO63166-0166 U.S.A.

(Received 19 August 2009,accepted 21October2009)

Abstract –Awidelyaccessible working list of knownplantspeciesisconsidered the number one targetof the GlobalStrategyforPlantConservation. Anindexof published namesof liverworts and hornworts between 2006 and 2008isprovided aspart of acontinued effort in working towardproducing apreliminary world checklist of theseplants.Included in the list arethosenamesoverlooked byearlierindices.The list includesover20highertaxa, 246speciesorinfraspecificvalid names,and three fossils.

Liverworts /hornworts /index/nomenclature

INTRODUCTION

Recently,therehasbeen amajorinternationallycoordinated effort to synthesizethe vast amountof nomenclatural,taxonomicaland globaldistribu- tionaldataforliverworts (Marchantiophyta)and hornworts (Anthocerotophyta) (e.g.,Söderström etal., 2008;von Konrat etal., 2008,2009). Theseendeavours are criticalinproviding the foundation todevelop aworking checklist of liverworts and hornworts worldwide. Thisissignificantbecausethe Convention on BiologicalDiversity (CBD)adopted the GlobalStrategyforPlantConservation (GSPC)whichsetthe targetdateof2010tocompleteawidelyaccessible working list of all knownplantspecies(Lughadha, 2004). The criticalneed forthistarget tobeachieved and the implicationsforconservation,taxonomy,and botanical sciencewererecentlyoutlined byPaton etal. (2008). Thisindexfollows on from the resourcefulindicesof recentyears:Index of Hepatics(Crosby&Engel,2006)and the seriesIndexof Bryophytes2001-2004 (Crosby&Magill,2005) and Indexof Bryophytes2005(Crosby&Magill,2006).

*Correspondenceand reprints:[email protected] 4M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Inthisindexweattempttolist citationsfornamespublished forliverworts and hornworts during the period January 1,2006 through December31,2008. Names thatwereoverlooked byearlierindicesarealsoincluded. Abibliographycontains completecitationsforthe placesof publication of the names.Although basionyms fornewcombinationsarelisted aswell asthe originalpublication,thesearenot referenced in the bibliography.The formatgenerallyfollows thatof previous indices.The type country of anewspeciesand infraspeciesisprovided. Forfossils westatethe geologicalperiod. The type Taxon isalsoprovided fornewnamesof highertaxa.All names,including basionymsand replaced namesarearranged alphabetically.Asamatterof referenceautonymsareincluded onlyforthe period in whichthe presentindexcovers,providing ausefulresourceindicating publication dateatwhichthe autonymwasfirst created. Although namesthatwe judge tobeinvalidlypublished areincluded in the list,wehaveattempted notto include datathatcould beinterpreted asvalidating them. Therefore,the invalidly published namesarenotaccepted bythe authors of thisindex,and theyarenot thereforepublished underthe ViennaCode of BotanicalNomenclature(McNeill etal. 2006). Asin previous indices,no disposition isprovided toanaccepted name ortothe taxonomy. The valid numberof published namesincludesover20highertaxa, 246speciesorinfraspecificnames,and three fossils.Noteworthyisthe influence of molecularinvestigationswithhigherclassification and taxonomy.Inthe past severalyears,the application of molecularphylogeneticstothe unraveling of liverwort phylogenyhasgenerated newinsights intothe evolutionary history of the groupand revolutionized liverwort classification (Crandall-Stotler etal., 2009b). Thisisevidentwiththe useofmolecularmarkers forall but one of the 24highertaxa(abovethe rank of genus)described. Recently,Crandall-Stotler etal. (2009a, 2009b)provided atreatisesummarizing historicaland contemporary liverwort classification. Incontrast,moleculardatain conjunction withmorphologicalcharacters wasused forless than15taxaatthe rank of species.

METHODS

Authoritiesand citation abbreviationsfollowthe on-line version of Authors of PlantNamesatthe RoyalBotanicGardens,KewWebsite (www.ipni.org ),whichisanupdated version of Brummitt &Powell (1992). For nomenclaturalelements wefollowthe ViennaCode of BotanicalNomenclature (McNeill etal., 2006). The following outlinesthe formatfornamespublished during the period: Name,Author(s),Citation [journal/publication,volume,page number], Year, Type country [subdivision notesin some cases]. The full referenceforaname isprovided in the index.Atthe end of each name,referraltothe full citation forthe journal/book article in whichthe name appeared isin the form: SEE Engel (2007). Entriesmayhaveanannotation in the formofaNOTE nearthe end. Theseannotationstypicallyidentifyinvalid or illegitimatenamesand mayprovide abrief description of the problem withthe name. EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-20085

The name of a newTaxon published during the period takesthe form: Allisoniellanigra var. acutiloba J.J.Engel, Novon 17:313.2007.NewZealand [Stewart Island].S EE Engel (2007). Its article from the Bibliography: ENGEL J.J.,2007.—Studiesof NewZealand Hepaticae. 20-38. Amiscellaneaof newtaxaand combinations. Novon 17:310-314.

A newcombination published during the period takesthe form: Austrolejeuneaconchophylla (Grolle) Pócs, J.HattoriBot.Lab. 99:186.2006. (Basionym: Nephelolejeuneaconchophylla Grolle, J.HattoriBot.Lab. 48: 164. 1980). S EE Pócs(2006). Its article from the Bibliography: PÓCS T.,2006.—Contributionstothe bryofloraof Australia, II.Onthe Australasian“Tuyamaelloideae” (Lejeuneaceae),withthe description of Austrolejeuneaoccidentalis . Journalofthe Hattoribotanicallaboratory 99: 185-196.

Basionymsthathavebeen used toestablishnewcombinationsduring the index period arealsolisted. Forexample, Nephelolejeuneaconchophylla Grolle, J.HattoriBot.Lab. 48:164. 1980.O THER USE: Austrolejeuneaconchophylla (Grolle) Pócs,2006.

Anasterisk,*,preceding anentry indicatesthatthe name hasbeen interpreted by us ascontrary toMcNeill etal. 2006 and appliestobothinvalid and illegitimate names.Anannotation in the formofaNOTE indicatesthe problem. Forexample, * Lopholejeuneanongstoinii AjitP.Singh et V.Nath, Phytotaxonomy 6:32.2006 [2007].N OTE:nom. inval.,holotype notdesignated. India[Meghalaya]. S EE Singh &Nath(2007c).

Adagger,†,preceding anentry indicatesaname forafossil. Forexample, † Schizolepidellabirkenmajeri Ociepa, PolishPolarResearch 28:34. 2007. Jurassic.S EE Ociepa(2007).

Incaseswherenewinfraspecifictaxahavebeen developed,weprecede eachentry withthe species,authority,and yeartoprovide historicalcontext.Forexample, Allisoniellanigra (Rodway)R.M.Schust.,1971[1972]. Allisoniellanigra var. acutiloba J.J.Engel, Novon 17:313.2007.NewZealand [Stewart Island].S EE Engel (2007a).

THE INDEX

Adelanthus crossii Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:404. 1885. O THER USE: Pseudomarsupidiumcrossii (Spruce) J.J.Engel,2007. Adelanthus decipiens (Hook.) Mitt.,1863 [1864]. Adelanthus decipiens subsp. aureocinctus R.M.Schust., Phytologia 39:250.1978. O THER USE: Pseudomarsupidiumaureocinctum (R.M.Schust.) J.J.Engel, 2007. Allisoniellanigra (Rodway)R.M.Schust.,1971[1972]. Allisoniellanigra var. acutiloba J.J.Engel, Novon 17:313.2007.NewZealand [Stewart Island].S EE Engel (2007a). 6M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Allisoniellanigra var. nigra , Novon 17:313.2007.S EE Engel (2007a). Amblyolejeuneafulfordiae Ast, Rev.Bryol. Lichénol. 17:25. 1948. O THER USE: Lejeuneafulfordiae (Ast)R.L.Zhu,2008. Amphicephaloziaafricana Vá≈ a et Wigginton, J.Bryol. 30:55. 2008. Rwanda.S EE Vá≈ a&Wigginton (2008). Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17: 311. 2007.B ASIONYM: Cephalolobus hodgsoniae R.M.Schust., Rev.Bryol. Lichénol. 34:251. 1966.S EE Engel (2007a). Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel, Novon 17:311. 2007.B ASIONYM: Cephaloziascabrella C.Massal., NuovoGiorn. Bot.Ital. 17:233.1885. S EE Engel (2007a). Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17:311. 2007.B ASIONYM: Cephalolobus sphenoloboides R.M.Schust., Rev.Bryol. Lichénol. 34:251. 1966.S EE Engel (2007a). Aneuracoronopus DeNot. ex Steph., Hedwigia 32:19. 1893.O THER USE: Lobatiriccardiacoronopus (DeNot. ex Steph.) Furuki,2006. * Aneuramerrillana Steph. ex Furuki, J.HattoriBot.Lab. 100:94. 2006.N OTE: nom. inval. Art.34.1(c),prosyn. (cf. Riccardianagasakiensis (Steph.) S.Hatt.). S EE Furuki (2006b). Aneurapolyclada Mitt. ex Thurn, Timehri 5:222.1886.O THER USE: Riccardia polyclada (Mitt. ex Thurn) Hässel, Bryologist 109:34. 2006. Anthoceroscoriaceus Steph., Spec.Hepat. (Stephani) 5:991. 1916.O THER USE: Phaeomegaceroscoriaceus (Steph.) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, 2007. Anthocerosendiviaefolius Mont., Voy.Pôle Sud 1:211. 1845. O THER USE: Nothocerosendiviaefolius (Mont.) J.Haseg. ex J.C.Villarreal,Hässel et N.Salazar,2007. Anthocerosevanidus Steph., Spec.Hepat. (Stephani) 5:990.1916.O THER USE: Phaeocerosevanidus (Steph.) Cargill et Fuhrer,2008. Anthocerosgiganteus Lehm. et Lindenb., Nov.Stirp. Pug. 4:25. 1832.O THER USE: Nothocerosgiganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar,2007. Anthoceroshallii Austin, Bull. TorreyBot.Club 6:26.1875. O THER USE: Paraphymatoceroshallii (Austin) Hässel,2006. Anthocerosinflatus Steph., Spec.Hepat. (Stephani) 5:990.1916.O THER USE: Phaeocerosinflatus (Steph.) Cargill et Fuhrer,2008. Anthocerosminutus Mitt., J.Proc.Linn. Soc.,Bot. 16:195. 1877 [1878].O THER USE: Paraphymatocerosminutus (Mitt.) Hässel,2006. Anthocerosphymatodes M.Howe, Bull. TorreyBot.Club 25:12.1898. O THER USE: Phymatocerosphymatodes (M.Howe) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, 2007. Aphanolejeuneaexigua A.Evans,1911. Aphanolejeuneaexigua var. africana Pócs, Cryptog. Bryol. Lichénol.5:247.1984. O THER USE: Cololejeuneamicroscopica var. africana (Pócs)Pócs et Bern.- Lück.,2008. Aphanolejeuneagracilis Ast, Rev.Bryol. Lichénol. 16:21. 1947.O THER USE: Cololejeuneagracilis (Ast)Pócs,2008. Aphanolejeuneasintenisii Steph., Spec.Hepat. (Stephani) 5:861. 1916.O THER USE: Cololejeuneasintenisii (Steph.) Pócs,2008. Aphanolejeuneasubsphaeroidea R.M.Schust., Phytologia 39:431. 1978. O THER USE: Cololejeuneasubsphaeroidea (R.M.Schust.) Pócs,2008. EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-20087

Arachniopsisindica S.C.Srivast. et P.K.Verma, Natl. Acad. Sci. Lett. 27:270.2004. O THER USE: Telaraneaindica (S.C.Srivast. et P.K.Verma)A.E.D.Daniels et P.Daniel,2007.S EE Srivastava&Verma(2006). Archilejeuneahelenae Pócs et Gyarmati, Cryptog. Bryol. 27:104. 2006. Madagascar.S EE Pócs&Sass-Gyarmati(2006). Austrolejeuneaconchophylla (Grolle) Pócs, J.HattoriBot.Lab. 99:186.2006. B ASIONYM: Nephelolejeuneaconchophylla Grolle, J.HattoriBot.Lab. 48: 164. 1980.S EE Pócs(2006a). Austrolejeuneahamata (Grolle) Pócs, J.HattoriBot.Lab. 99:186.2006. B ASIONYM: Nephelolejeuneahamata Grolle, J.HattoriBot.Lab. 48:167. 1980.S EE Pócs(2006a). Austrolejeuneaoccidentalis Pócs, J.HattoriBot.Lab. 99:187.2006.Australia [WesternAustralia].S EE Pócs(2006a). Austrolejeuneapapillosa (Glenny)Pócs, J.HattoriBot.Lab. 99:186.2006. B ASIONYM: Nephelolejeuneapapillosa Glenny, NewZealand J.Bot. 34: 195. 1996.S EE Pócs(2006a). Austrolejeuneatalinayi (S.W.Arnell) Pócs, J.HattoriBot.Lab. 99:187.2006. B ASIONYM: Harpalejeuneatalinayi S.W.Arnell, SvenskBot.Tidskr. 50: 309. 1956.S EE Pócs(2006a).

Bazzania sect. Exemptae J.J.Engel, J.HattoriBot.Lab. 99:200.2006.T YPE: Bazzaniaexempta J.J.Engel, J.HattoriBot.Lab. 99:197.2006.S EE Engel (2006a). Bazzania sect. Glaucobazzania J.J.Engel, Novon 17:310.2007.T YPE: Bazzania tayloriana (Mitt.) Kuntze, Revis.Gen. Pl. 2:832.1891. S EE Engel (2007a). Bazzania sect. Protobazzania J.J.Engel, J.HattoriBot.Lab. 99:203.2006.T YPE: Bazzaniainvoluta (Mont.) Trevis, Mem. Reale Ist.Lombardo Sci.,Ser.3, Cl. Sci. Mat. 4:414. 1877.S EE Engel (2006a). Bazzaniacitharodes Meagher, NovaHedwigia 86:481. 2008. Australia [Queensland].S EE Meagher(2008). Bazzaniacorbieri (Steph.) Meagher, NovaHedwigia 86:483.2008. B ASIONYM: Mastigobryumcorbieri Steph., J.&Proc.Roy.Soc.NewSouthWales 48: 122.1914. S EE Meagher(2008). Bazzaniaengelii Glenny, Fieldiana, Bot., n.s.47:176.2008. NewZealand [South Island].S EE Glenny&Bartlett (2008). Bazzaniaexempta J.J.Engel, J.HattoriBot.Lab. 99:197.2006.NewZealand [NorthIsland].S EE Engel (2006a). Bazzaniaokaritana Meagher et Glenny, J.Bryol. 29:60.2007.NewZealand [SouthIsland].S EE Meagher&Glenny(2007). Bazzaniarimosa Meagher, NovaHedwigia 86:489. 2008. PapuaNewGuinea.S EE Meagher(2008). Bazzaniascalaris Meagher, Telopea 11(3):247.2006.PapuaNewGuinea.S EE Meagher(2006). Bazzaniazonulata Meagher, NovaHedwigia 86:491. 2008. Australia [Queensland].S EE Meagher(2008). Blasiidae He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27.2006. B ASED ON:Blasiales(R.M.Schust.) Stotler et Crand.-Stotl. T YPE: Blasia L., Sp. Pl.:1138. 1753.S EE He-Nygrén etal. (2006). Blepharostomataceae W.Frey et M.Stech, NovaHedwigia 87:263.2008. B ASED ON:Pseudolepicoleaceaesubfam. BlepharostomoideaeGrolle, J.Bryol. 7: 205. 1972.T YPE: Blepharostoma (Dumort.) Dumort., Recueil Observ. Jungerm. 18. 1835. S EE Frey&Stech(2008). 8M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Calyculariaceae He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27. 2006.T YPE: Calycularia Mitt., J.Proc.Linn. Soc.,Bot. 5:122.1861. S EE He-Nygrén etal. (2006). Calyculariineae He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27. 2006.B ASED ON:CalyculariaceaeHe-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27.2006.T YPE: Calycularia Mitt., J.Proc.Linn. Soc., Bot. 5:122.1861. S EE He-Nygrén etal. (2006). Calypogeiakhasiana AjitP.Singh et V.Nath, Taiwania 52:320.2007.T YPE:India [Meghalaya].S EE Singh &Nath(2007a). Castanoclobos J.J.Engel et Glenny, Novon 17:424. 2007.T YPE: Leiomitrajulacea Hatcher ex J.J.Engel, Novon 9:26,1999. S EE Engel &Glenny(2007). Castanoclobosjulaceus (Hatcher ex J.J.Engel) J.J.Engel et Glenny, Novon 17:426. 2007.B ASIONYM: Leiomitrajulacea Hatcher ex J.J.Engel, Novon 9:26. 1999. S EE Engel &Glenny(2007). Cephalojonesiaincuba Grolle et Vanden Berghen,1970. Cephalojonesiaincuba subsp. incuba, J.HattoriBot.Lab. 100:37.2006.S EE Burghardt etal. (2006). Cephalojonesiaincuba subsp. mexicana Burghardt,Gradst. et Vá≈ a, J.Hattori Bot.Lab. 100:37.2006.Mexico[Michoacán].S EE Burghardt etal. (2006). Cephalolobus hodgsoniae R.M.Schust., Rev.Bryol. Lichénol. 34:251. 1966. O THER USE: Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel,2007. Cephalolobus sphenoloboides R.M.Schust., Rev.Bryol. Lichénol. 34:251. 1966. O THER USE: Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel,2007. Cephalozia subgen. Eocephalozia J.J.Engel, Novon 17:312.2007.T YPE: Cephaloziabadia (Gottsche) Steph., Bull. Herb.Boissier ,sér.28:483. 1908(Spec.Hepat. (Stephani) 3:313.1908). S EE Engel (2007a). Cephaloziaaustrigena R.M.Schust. ex J.J.Engel, Novon 17:312.2007.New Zealand [SouthIsland].S EE Engel (2007a). Cephaloziapachygyna R.M.Schust. ex J.J.Engel, Novon 17:313.2007.New Zealand [SouthIsland].S EE Engel (2007a). Cephaloziascabrella C.Massal., NuovoGiorn. Bot.Ital. 17:233.1885. O THER USE: Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel,2007. Cephaloziaschusteri S.K.Singh et D.K.Singh, Lindbergia 32:1. 2007.India [HimachalPradesh].S EE Singh &Singh (2007a). Cephaloziaschusteriana J.J.Engel, Novon 17:313.2007.NewZealand [South Island].S EE Engel (2007a). Cephaloziellaarctica Bryhn et Douin, 1913. Cephaloziellaarctica var. subantarctica R.M.Schust., NovaHedwigia 22:197. 1972.O THER USE: Cephaloziellavarians var. subantarctica (R.M.Schust.) R.M.Schust. ex J.J.Engel,2007. Cephaloziellapseudocrassigyna R.M.Schust. ex J.J.Engel, Novon 17:313.2007. NewZealand [SouthIsland].S EE Engel (2007a). Cephaloziellaspinigera (Lindb.) Jørg.,1894. Cephaloziellaspinigera f. striatula (C.E.O.Jensen) Damsh., Lindbergia 28:124. 2003.B ASIONYM: Cephaloziastriatula C.E.O.Jensen, Rev.Bryol. 31:25. 1904. S EE Damsholt(2003). Cephaloziellavarians (Gottsche) Steph.,1905. Cephaloziellavarians var. subantarctica (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17:314. 2007.B ASIONYM: Cephaloziellaarctica var. subantarctica R.M.Schust., NovaHedwigia 22:197.1972.S EE Engel (2007a). EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-20089

Ceratolejeuneaatlantica Alvarenga et Ilk.-Borg., NovaHedwigia 86:238. 2008. Brazil [Alagoas].S EE Ilkiu-Borges(2008). Cheilolejeuneabeyrichii (Lindenb.) M.E.Reiner, NovaHedwigia 83:474. 2006. B ASIONYM: Lejeuneabeyrichii Lindenb., Syn. Hepat. 3:371. 1845. S EE Reiner-Drehwald (2006). Cheilolejeuneacompacta (Steph.) M.E.Reiner, NovaHedwigia 83:474. 2006. B ASIONYM: Hygrolejeuneacompacta Steph., Spec.Hepat. (Stephani) 5: 532.1914. S EE Reiner-Drehwald (2006). Cheilolejeuneaeximia (Ast et Tixier)R.L.Zhu et M.L.So,Beih. NovaHedwigia 121:114. 2001. B ASIONYM: Pycnolejeuneaeximia Ast et Tixier, Rev.Bryol. Lichénol. 31:31. 1962.Vietnam. S EE Zhu&So(2001). Cheilolejeunealacerata C.Bastos et Gradst., J.Bryol. 28:133.2006.Brazil [Bahia]. S EE Bastos&Gradstein (2006). Cheilolejeuneaneblinensis Ilk.-Borg. et Gradst., NovaHedwigia 87:522.2008. Venezuela[Amazonas].S EE Ilkiu-Borges&Gradstein (2008). Cheilolejeunearupestris C.Bastos et Gradst., J.Bryol. 28:136.2006.Brazil [Bahia]. S EE Bastos&Gradstein (2006). Chiloscyphus sect. Lophocolea (Dumort.) Hentschel, Pl. Syst.Evol. 262:129. 2006. B ASIONYM: Jungermannia sect. Lophocolea Dumort., Syll. Jungerm. Europ.:59. 1831. T YPE: Jungermanniabidentata L.S EE Hentschel etal. (2006). Chiloscyphus anisolobus J.J.Engel et Glenny, Bryologist 111:118. 2008. New Zealand [SouthIsland].S EE Engel &Glenny(2008a). Chiloscyphus dissitifolius (Herzog) Hentschel et Heinrichs, Taxon 56:1139. 2007. B ASIONYM: Pachyglossadissitifolia Herzog, Rev.Bryol. Lichénol. 21:259. 1952.S EE Hentschel etal. (2007a). Chiloscyphus fissus (Mitt.) Hentschel et Heinrichs, Taxon 56:1139. 2007. B ASIONYM: Herpocladiumfissum Mitt., J.Proc.Linn. Soc.,Bot.Botany 15:69. 1876.S EE Hentschel etal. (2007a). Chiloscyphus grolleanus (Vá≈ a)Hentschel et Heinrichs, Taxon 56:1139. 2007. B ASIONYM: Pachyglossagrolleana Vá≈ a, Cryptog. Bryol. 26:86.2005. S EE Hentschel etal. (2007a). Chiloscyphus inflatus Steph.,1907. Chiloscyphus inflatus var. fragilissimus N.Kitag., ActaPhytotax.Geobot. 30:32. 1979. O THER USE: Heteroscyphus inflatus var. fragilissimus (N.Kitag.) M.J.Lai et R.L.Zhu, Ann. Bot.Fenn. 45:334. 2008. Chiloscyphus spegazzinianus (C.Massal.) Hentschel et Heinrichs, Taxon 56:1139. 2007.B ASIONYM: Lophocoleaspegazziniana C.Massal., NuovoGiorn. Bot.Ital. 17:255. 1885. S EE Hentschel etal. (2007a). Chiloscyphus tenacifolius (Hook.f. et Taylor)Hentschel et Heinrichs, Taxon 56: 1139. 2007.B ASIONYM: Jungermanniatenacifolia Hook.f. et Taylor, Hooker, Bot.Antarct.Voy.I(Fl. Antarct.1) “1844” 1:40.1844. S EE Hentschel etal. (2007a). Chiloscyphus yunnanensis C.Gao et Y.H.Wu, ActaBot.Yunnan. 28:119. 2006. China[Yunnan].S EE Gao&Wu(2006). Cololejeuneaarfakiana Pócs et J.Eggers, PolishBot.J. 51:155. 2006.Indonesia [NewGuinea, West Irian].S EE Pócs&Eggers (2006). Cololejeuneadauphinii R.L.Zhu, J.Bryol. 28:277.2006.N OTE:nom. nov.pro Cololejeuneatixieri M.I.Morales et G.Dauphin, Trop. Bryol. 14:133.1998, nom. illeg. Art.53.1,non Cololejeuneatixieri Onr., Bull. Jard. Bot.Natl. Belgique 59:436.1989. S EE Zhu(2006). 10M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Cololejeuneadianae Wigginton, J.Bryol. 28:364. 2006.StHelena.S EE Wigginton (2006). Cololejeuneaeustacei Pócs in Müller&Pócs, J.Bryol. 29:83.2007.Equatorial Guinea[Bioko Island].S EE Müller&Pócs(2007). Cololejeuneagracilis (Ast)Pócs in Dauphin etal., Cryptog. Bryol. 29:233.2008. B ASIONYM: Aphanolejeuneagracilis Ast, Rev.Bryol. Lichénol. 16:21. 1947.S EE Dauphin etal. (2008). Cololejeuneagradsteinii M.J.Lai et R.L.Zhu, Ann. Bot.Fenn. 45:334. 2008. N OTE: nom. nov.pro Cololejeuneapusilla Tixier, Nat.Hist.Bull. SiamSoc. 23: 548. 1970,nom. illeg. Art.53.1,non Cololejeuneapusilla Steph., Hedwigia 34:251. 1895. S EE Lai etal. (2008). Cololejeuneagrossestyla Wigginton, J.Bryol. 28:369. 2006.StHelena.S EE Wigginton (2006). Cololejeunealichenyae R.D.Porley,N.G.Hodgetts et M.Wigginton in Wigginton et al., J.Bryol. 29:7.2007.Malawi. S EE Wigginton etal. (2007). Cololejeuneametzgeriopsis (K.I.Goebel) Gradst.,R.Wilson,Ilk.-Borg. et Heinrichs, J.Linn. Soc.,Bot. 151:306.2006.B ASIONYM: Lejeunea metzgeriopsis K.I.Goebel,Flora72:2.1889. S EE Gradstein etal. (2006). Cololejeuneamicroscopica (Taylor)Schiffn.,1893. Cololejeuneamicroscopica var. africana (Pócs)Pócs et Bern.-Lück. InDauphin et al., Cryptog. Bryol. 29:234. 2008. B ASIONYM: Aphanolejeuneaexigua var. africana Pócs, Cryptog. Bryol. Lichénol. 5:247.1984. S EE Dauphin etal. (2008). Cololejeuneamicroscopica var. microscopica, Cryptog. Bryol. 29:234. 2008. S EE Dauphin etal. (2008). Cololejeuneapanamensis G.Dauphin et Pócs in Dauphin etal., Trop. Bryol. 27:76. 2006.Panama.S EE Dauphin etal. (2006). Cololejeuneaplanissima ( Mitt.) Abeyw.,1959. Cololejeuneaplanissima var. chagosensis Pócs in Seaward etal., J.Bryol. 28:14. 2006.ChagosArchipelago. S EE Seaward etal. (2006). Cololejeuneaplanissima var. planissima , J.Bryol. 28:14. 2006.S EE Seaward etal. (2006). Cololejeuneasanctae-helenae Wigginton, J.Bryol. 28:366.2006.StHelena.S EE Wigginton (2006). Cololejeuneasintenisii (Steph.) Pócs in Dauphin etal., Cryptog. Bryol. 29:235. 2008. B ASIONYM: Aphanolejeuneasintenisii Steph., Spec.Hepat. (Stephani) 5:861. 1916.S EE Dauphin etal. (2008). Cololejeuneasubsphaeroidea (R.M.Schust.) Pócs in Dauphin etal., Cryptog. Bryol. 29:235. 2008. B ASIONYM: Aphanolejeuneasubsphaeroidea R.M.Schust., Phytologia 39:431. 1978. S EE Dauphin etal. (2008). Cololejeuneatixieriana M.Dey,D.Singh et D.K.Singh, Taiwania 53:258. 2008. India[Darjeeling].S EE Dey etal. (2008a). Cololejeuneaturbinifera (Spruce) León-Yánez,Gradst. et Wegner, Hepát.y Antoc.del Ecuador,Catál.:33.2006.B ASIONYM: Lejeuneaturbinifera Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:295. 1884. S EE León- Yánez etal. (2006a). Coluracalderae Pócs, J.Bryol. 29:84. 2007.EquatorialGuinea[Bioko Island]. S EE Müller&Pócs(2007). Coluravitiensis Pócs et J.Eggers, PolishBot.J. 52:88. 2007.Fiji. S EE Pócs& Eggers (2007). Cylindrocoleabrasiliensis D.P.Costa, N.D.Santos et Vá≈ a, Bryologist 111:2008: 667.Brazil [Rio de Janeiro].S EE Costa etal. (2008). EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200811

Dendrocerotidae R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:240. 2007.T YPE: Dendroceros Nees, Syn. Hep. 4:579. 1846.S EE Duff etal. (2007). Dinckleriafruticella (Hook.f. et Taylor)J.J.Engel et Heinrichs, Cryptog. Bryol. 29: 194. 2008. B ASIONYM: Jungermanniafruticella Hook.f. et Taylor, London J.Bot. 3:565. 1844. S EE Engel &Heinrichs(2008). Dinckleriapleurata (Hook.f. et Taylor)J.J.Engel et Heinrichs, Cryptog. Bryol. 29: 194. 2008. B ASIONYM: Jungermanniapleurata Hook.f. et Taylor, London J.Bot. 3:372.1844. S EE Engel &Heinrichs(2008). Diplasiolejeuneaandringitrae Schäf.-Verw., Cryptog. Bryol. 27:447.2006. Madagascar.S EE Pócs&Schäfer-Verwimp (2006). Diplasiolejeuneaeggersii Pócs, Bryologist 109:408. 2006.CostaRica.S EE Pócs (2006b). Diplasiolejeuneaingekarolae Schäf.-Verw., Herzogia 19:239. 2006.Indonesia [Sumatra].S EE Schäfer-Verwimp (2006). Diplasiolejeuneaornata Pócs et Schäf.-Verw., Cryptog. Bryol. 27:440.2006. Madagascar.S EE Pócs&Schäfer-Verwimp (2006). Diplasiolejeunearanomafanae Pócs, Cryptog. Bryol. 27:444. 2006.Madagascar. S EE Pócs&Schäfer-Verwimp (2006). Diplophylloideae W.Frey et M.Stech, NovaHedwigia 87:262.2008. B ASED ON: DiplophyllaceaePotemkin, Ann. Bot.Fenn. 36:281. 1999. T YPE: Diplophyllum (Dumort.) Dumort., Recueil Observ.Jungerm. 15. 1835. Scapaniaceae . S EE Frey&Stech(2008). Drepanolejeunealongifolia AjitP.Singh,V.Nath et A.K.Asthana , Bryologist 111: 2008:477.India[Meghalaya].S EE Singh etal. (2008). Drepanolejeuneamawtmiana AjitP.Singh et V.Nath, Hepat.KhasiJaintiaHills: East.Himalayas 251. 2007.India[Meghalaya].S EE Singh &Nath(2007b). Dumortieraceae D.G.Long, Edinburgh J.Bot. 63:260.2006.T YPE: Dumortiera Nees, NovaActaPhys.Med. Acad. Caes.Leop. Carol. Nat.Cur. 12:410. 1824. S EE Long (2006).

Exormotheca bischlerae Furuki et Higuchi, Cryptog. Bryol. 27:98. 2006. “ bischleri .” China[Sichuan].S EE Furuki &Higuchi (2006).

Fossombroniaaltilamellosa G.A.M.Scott et D.C.Pike ex Cargill in P.M. McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[Victoria].N OTE: based on Fossombroniaaltilamellosa G.A.M.Scott et D.C.Pike,1987, nom. inval. Art.37.2(based on morethanone gathering). S EE McCarthy (2003). Fossombroniaauricolor G.A.M.Scott et D.C.Pike ex Cargill in P.M.McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[SouthAustralia].N OTE: based on Fossombroniaauricolor G.A.M.Scott et D.C.Pike,1987,nom. inval. Art.37.2(based on morethanone gathering). S EE McCarthy (2003). Fossombronialophoscypha Hässel, Beih. NovaHedwigia 131:16.2007.Chile [Valparaíso].S EE Hässel de Menéndez&Villagrán(2007). Fossombroniamagnaspora G.A.M.Scott et D.C.Pike ex Cargill in P.M.McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[Victoria].N OTE:based on Fossombroniamagnaspora G.A.M.Scott et D.C.Pike,1987,nom. inval. Art.37.2(based on morethanone gathering). S EE McCarthy(2003). *Fossombroniamaritima G.A.M.Scott et D.C.Pike ex Cargill in P.M.McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[Victoria].N OTE:based on Fossombroniamaritima G.A.M.Scott et D.C.Pike,1987,nom. inval. 12M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Art.37.2(based on morethanone gathering);nom. illeg. Art.53.1,non Fossombroniamaritima (Paton) Paton,1994. S EE McCarthy(2003). Fossombroniapurpureospora G.A.M.Scott et D.C.Pike ex Cargill in P.M.McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[South Australia].N OTE:based on Fossombroniapurpureospora G.A.M.Scott et D.C.Pike,1987,nom. inval. Art.37.2(based on morethanone gathering). S EE McCarthy(2003). Fossombroniarupestris G.A.M.Scott et D.C.Pike ex Cargill in P.M.McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[SouthAustralia].N OTE: based on Fossombroniarupestris G.A.M.Scott et D.C.Pike,1987,nom. inval. Art.37.2(based on morethanone gathering). S EE McCarthy (2003). Fossombroniatruncata G.A.M.Scott et D.C.Pike ex Cargill in P.M.McCarthy, Fl. AustraliaSuppl. 21:114. 2003.Australia[Victoria].N OTE:based on Fossombroniamagnaspora G.A.M.Scott et D.C.Pike 1987,nom. inval. Art.37.2(based on morethanone gathering). S EE McCarthy(2003). Fossombroniavalparaisiana Hässel, Beih. NovaHedwigia 131:14. 2007.Chile [Valparaiso].S EE Hässel de Menéndez&Villagrán(2007). Frullaniadulimensis Uribe, Cryptog. Bryol. 27:309. 2006.Colombia[Tolima]. S EE Uribe(2006). Frullaniapran-nathii M.Dey et D.K.Singh, J.Jap. Bot. 83:281. 2008. India [Darjeeling].S EE Dey&Singh (2008). Frullaniapseudoalstonii Tsudo et J.Haseg., Bryol. Res. 9:44. 2006.Japan [Kyushu].S EE Tsudo &Hasegawa(2006). Frullaniatamarisci (L.) Dumort.,1835. Frullaniatamarisci var. azorica J.-P.Frahm, Trop. Bryol. 27:102.2006.Portugal [Azores].S EE Frahm (2006). Frullaniaudarii V.Nath et AjitP.Singh, Curr.Sci. 91:744. 2006.India [Meghalaya].S EE Nath&Singh (2006). Frullaniavivipara Pócs, Fieldiana, Bot. ,n.s.47:151. 2008. Fiji. S EE Pócs(2008a). Frullaniawairua von Konrat et Braggins, NewZealand J.Bot. 43:886.2005. New Zealand [NorthIsland].S EE von Konrat(2005).

Gottscheliagrollei D.G.Long et Vá≈ a, J.Bryol. 29:167.2007.China[Yunnan]. S EE Long &Vá ≈ a(2007). Grimaldiafragrans (Balb.) Corda ex Nees ,1838. Grimaldiafragrans var. brevipes Kaal., Forh. Vidensk.-Selsk. Kristiania 1898:3. 1898. O THER USE: Manniafragrans var. brevipes (Kaal.) Damsh., Ill. Fl. Nord. Liverw.Hornw. :717.2002.

Hamatostrepta Vá≈ a et D.G.Long, Fieldiana, Bot. ,n.s.47:134. 2008. T YPE: Hamatostreptaconcinna Vá≈ a et D.G.Long, Fieldiana, Bot., n.s.47:134. 2008. S EE Vá≈ a&Long (2008). Hamatostreptaconcinna Vá≈ a et D.G.Long, Fieldiana, Bot. ,n.s.47:134. 2008. Myanmar[Kachin].S EE Vá≈ a&Long (2008). Haplomitriidae Stotler et Crand.-Stotl., Taxon 57:290.2008. T YPE: Haplomitrium Nees.S EE Stotler&Crandall-Stotler(2008). Harpalejeuneatalinayi S.W.Arnell, SvenskBot.Tidskr. 50:309. 1956.O THER USE: Austrolejeuneatalinayi (S.W.Arnell) Pócs,2006. Herbertus acanthelius Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:344. 1885. O THER USE: Herbertus juniperoideus subsp. acanthelius (Spruce) Feldberg et Heinrichs,2006. EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200813

Herbertus bivittatus Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:343.1885. O THER USE: Herbertus juniperoideus subsp. bivittatus (Spruce) Feldberg et Heinrichs,2006. Herbertus buchii Juslén, Ann. Bot.Fenn. 43:416.2006.Russia[Amur].S EE Juslén (2006). Herbertus juniperoideus (Sw.) Grolle,1961. Herbertus juniperoideus subsp. acanthelius (Spruce) Feldberg et Heinrichs, J.Linn. Soc.,Bot. 151:326.2006.B ASIONYM: Herbertus acanthelius Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:344. 1885. S EE Feldberg& Heinrichs(2006). Herbertus juniperoideus subsp. bivittatus (Spruce) Feldberg et Heinrichs, J.Linn. Soc.,Bot. 151:328. 2006.B ASIONYM: Herbertus bivittatus Spruce, Trans. &Proc.Bot.Soc.Edinburgh 15:343.1885. S EE Feldberg&Heinrichs (2006). Herbertus juniperoideus subsp. juniperoideus, J.Linn. Soc.,Bot. 151:326.2006. S EE Feldberg&Heinrichs(2006). Herbertus juniperoideus subsp. pensilis (Taylor)Feldberg et Heinrichs, J.Linn. Soc.,Bot. 151:329. 2006.B ASIONYM: Sendtnerapensilis Taylor, London J.Bot. 5:372.1846.S EE Feldberg&Heinrichs(2006). Herbertus mauritianus N.G.Hodgetts, J.Bryol. 30:247.2008. Mauritius.S EE Hodgetts (2008). Herbertus pocsii N.G.Hodgetts, J.Bryol. 30:249. 2008. Tanzania.S EE Hodgetts (2008). Herbertus spicatus N.G.Hodgetts, J.Bryol. 30:244. 2008. Tanzania.S EE Hodgetts (2008). Herpocladiumfissum Mitt., J.Proc.Linn. Soc.,Bot. 15:69. 1876.O THER USE: Chiloscyphus fissus (Mitt.) Hentschel et Heinrichs,2007. Heteroscyphus inflatus (Steph.) S.C.Srivast. et A.Srivast.,1989 [1991]. Heteroscyphus inflatus var. fragilissimus (N.Kitag.) M.J.Lai et R.L.Zhu, Ann. Bot. Fenn. 45:334. 2008. B ASIONYM: Chiloscyphus inflatus var. fragilissimus N.Kitag., ActaPhytotax.Geobot. 30:32.1979. S EE Lai etal. (2008). Heteroscyphus polychaetus (Spruce) Hentschel et Heinrichs, Taxon 56:1139. 2007. B ASIONYM: Lophocoleapolychaeta Spruce, Trans.&Proc.Bot.Soc. Edinburgh 15:436.1885. S EE Hentschel etal. (2007a). Hyalolepidoziamicrophylla R.M.Schust. ex J.J.Engel, Novon 17:310.2007.New Zealand [SouthIsland].S EE Engel (2007a). Hygrolejeuneacompacta Steph., Spec.Hepat. (Stephani) 5:532.1914. O THER USE: Cheilolejeuneacompacta (Steph.) M.E.Reiner,2006. Hypolophozia (R.M.Schust.) Bakalin, Monogr.ObrabotkaRoda Lophozia: 38. 2006.B ASIONYM: Lophozia subgen. Hypolophozia R.M.Schust., Rev. Bryol. Lichénol. 34:276.1966.T YPE: Lophoziasubinflata (R.M.Schust.) R.M.Schust., Rev.Bryol. Lichénol. 34:276.1966. Hypolophoziadusenii (Steph.) Bakalin, Monogr.ObrabotkaRoda Lophozia: 39. 2006.B ASIONYM: Lophoziadusenii Steph., Bih. Kongl. SvenskaVetensk.- Akad. Handl. 26(III, 6):24. 1900.S EE Bakalin (2006).

* Isopachespumicicola (Berggr.) Bakalin, Monogr.ObrabotkaRoda Lophozia: 27. 2006.N OTE:nom. inval. Art.33.1,basionym Lophoziapumicicola Berggr. notfullycited S EE Bakalin (2006). Isopachespumicicola (Berggr.) Bakalin, Arctoa 17:162,2008. B ASIONYM: Lophoziapumicicola Berggr.1898. NewZealand Hepat. :21. S EE Bakalin (2008a). 14 M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Jamesoniellaceae He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27. 2006.T YPE: Jamesoniella (Spruce) Carrington in Lees, London Catal. Brit.Moss.Hep. (ed. 2):25. 1881). S EE He-Nygrén etal. (2006). Jamesoniellales W.Frey et M.Stech, NovaHedwigia 87:262.2008. T YPE: Jamesoniella (Spruce) Carrington in Lees, London Catal. Brit.Moss.Hep. (ed. 2)25. 1881. S EE Frey&Stech(2008). Jubulahutchinsiae (Hook.) Dumort.,1822. Jubulahutchinsiae subsp. australiae Pócs et A.Cairns, NovaHedwigia 86:232. 2008. Australia[Queensland].S EE Pócs&Cairns(2008). Jubulales W.Frey et M.Stech, NovaHedwigia 87:264. 2008. T YPE: Jubula Dumort., Comment.Bot. 112.1822.S EE Frey&Stech(2008). *Jungermanniaamakawana AjitP.Singh et V.Nath, Hepat.KhasiJaintiaHills: East.Himalayas 117.2007.India[Meghalaya].N OTE:nom. illeg. Art.53.1, non Jungermanniaamakawana Grolle, J.Jap. Bot. 41:230.1966.S EE Singh &Nath(2007b). *Jungermanniaerectii AjitP.Singh et V.Nath, Hepat.KhasiJaintiaHills:East. Himalayas 117.2007.India[Meghalaya].N OTE:nom. illeg. Art.53.1,non Jungermanniaerecta (Amakawa)Amakawa, J.HattoriBot.Lab. 22:13, 1960.S EE Singh &Nath(2007b). Jungermanniagongshanensis C.Gao et J.Sun in Sun&Duan, Bull. Bot.Res., Harbin 27:140.2007.N OTE:nom. nov.for Jungermanniabrevicaulis C.Gao et X.L.Bai, Philippine Scientist 38:124. 2001,nom. illeg. Art.53.1, non Jungermanniabrevicaulis Raddi, Jungermanniogr.Etrusca 20.1818. S EE Sun&Duan(2007). Jungermanniaindrodayana S.K.Singh et D.K.Singh, Cryptog. Bryol. 28:103.2007. India[HimachalPradesh].S EE Singh &Singh (2007). Jungermanniajenseniana Grolle, Österreichische Bot.Z. 111:190.1964. O THER USE: Solenostomajenseniana (Grolle) Bakalin,2003. Jungermanniajuniperoidea Sw., Prodr.(Swartz) :144. 1788. O THER USE: Herbertus juniperoideus subsp. juniperoideus,2006. Jungermanniamairangii AjitP.Singh et V.Nath, Hepat.KhasiJaintiaHills:East. Himalayas 120.2007.India[Meghalaya].S EE Singh &Nath(2007b). Jungermanniamulticarpa C.Gao et J.Sun in Sun&Duan, Bull. Bot.Res.,Harbin 27: 139. 2007.N OTE:nom. nov.pro Jungermanniapolycarpa C.Gao et X.L.Bai, Philippine Scientist 38:147.2001;nom. illeg. Art.53.1,non Jungermannia polycarpa Nees, Fl. Bras. 1:350.1833.S EE Sun&Duan(2007). Jungermannianilgiriensis A.Alam,Ad.Kumar et S.C.Srivast., Bull. Bot.Surv. India 49:220.2007.India[Tamil Nadu].S EE Alam etal. (2007). Jungermanniapyriflora Steph.,1917. Jungermanniapyriflora var. minutissima Amakawa, J.HattoriBot.Lab. 22:61. 1960.O THER USE: Solenostomapyriflorum var. minutissimum (Amakawa)Bakalin,2007. Jungermanniarufiflora Colenso, TransactionsTrans.&Proc.NewZealand Inst. 18:237.1886.O THER USE: Solenostomarufiflorum (Colenso) J.J.Engel, Novon 17:312.2007. Jungermanniatenacifolia Hook.f. et Taylor, Fl. Antarct. 1:40.1844. O THER USE: Chiloscyphus tenacifolius (Hook.f. et Taylor)Hentschel et Heinrichs, 2007. Jungermanniasparsofolia C.Gao et J.Sun in Sun&Duan, Bull. Bot.Res.,Harbin 27:139. 2007.N OTE:nom. nov.pro Jungermannialaxifolia C.Gao,2003. B LOCKING NAME: Jungermannialaxifolia Hook., Brit.Jungerm. Pl. 59. 1816.S EE Sun&Duan(2007). EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200815

Kurziahelophila R.M.Schust.,1980. Kurziahelophila var. flaccida R.M.Schust. ex J.J.Engel, Novon 17:310.2007.New Zealand [SouthIsland].S EE Engel (2007a). Kurziahelophila var. helophila , Novon 17:310.2007.S EE Engel (2007a).

Leiocoleacollaris (Nees)Schljakov,1976. *Leiocoleacollaris var. pumila Dragi† evi † et Velji† , Pregl. mahov.Crne Gore : 14 2006.N OTE:nom. inval. Art.32.1(d) (no description) and Art.36.1, apparentlybased on Jungermanniamuelleri f. pumila Nees,1836.S EE Dragi† evi † &Velji† (2006). Leiocoleaholmeniana (Inoue et Steere) Konstant., Arctoa 9:33.2000.B ASIONYM: Lophoziaholmeniana Inoue et Steere, J.HattoriBot.Lab. 44:285. 1978. S EE Konstantinova(2000). Leiocoleaturbinata (Raddi) H.Buch,1938. *Leiocoleaturbinata var. wilsoniana Dragi† evi † et Velji† , Pregl. mahov.Crne Gore :4. 2006.N OTE:nom. inval. Art 32.1(d),apparentlybased on Jungermanniawilsoniana Nees,1838. S EE Dragi† evi † &Velji† (2006). Leiocoleaussuriensis Bakalin, Arctoa 17:103,2008. RussianFarEast [Primorsky]. S EE Bakalin (2008b). Leiomitrajulacea Hatcher ex J.J.Engel, Novon 9:26.1999. O THER USE: Castanoclobosjulaceus (Hatcher ex J.J.Engel) J.J.Engel et Glenny,2007. Leiosporocerotopsida Stotler et Crand.-Stotl., Bryologist 108:19. 2005. T YPE: Leiosporoceros Hässel, J.HattoriBot.Lab. 64:82.1988. S EE Stotler& Crandall-Stotler(2005). Lejeuneabeyrichii Lindenb., Syn. Hep. 3:371. 1845. O THER USE: Cheilolejeunea beyrichii (Lindenb.) M.E.Reiner,2006. Lejeuneacordifissa Taylor, London J.Bot. 5:395. 1846.O THER USE: Lepidolejeuneacordifissa (Taylor)M.E.Reiner, NovaHedwigia 83:478. 2006. Lejeuneafulfordiae (Ast)R.L.Zhu, Syst.Bot. 33:617.2008. B ASIONYM: Amblyolejeuneafulfordiae Ast, Rev.Bryol. Lichénol. 17:25. 1948. S EE Zhu&Cheng (2008). Lejeuneahalei H.Rob.,1964. Lejeuneahalei var. africana Pócs, J.Bryol. 29:89. 2007.EquatorialGuinea[Bioko Island].S EE Müller&Pócs(2007). Lejeuneahalei var. halei, J.Bryol. 29:89. 2007.S EE Müller&Pócs(2007). Lejeuneajuruana Gradst. et M.E.Reiner, Syst.Bot. 32:488. 2007.B ASIONYM:nom. nov.pro Potamolejeuneauleana Steph., Spec.Hepat. (Stephani) 5:641. 1914. B LOCKING NAME: Lejeuneauleana (Steph.) Steph., Spec.Hepat. (Stephani) 5:675. 1915. S EE Gradstein &Reiner-Drehwald (2007). Lejeuneakashyapii M.Dey,D.K.Singh et D.Singh, J.Bryol. 30:126.2008. India [Sikkim].S EE Dey etal. (2008b). Lejeunealaii R.L.Zhu, J.Bryol. 30:173.2008. N OTE:nom. nov.pro Lejeunea ramulosa (Herzog) R.M.Schust., Beih. NovaHedwigia 9:133,nom. illeg. Art.53.1,non Lejeunearamulosa Spruce, Trans.Proc.Bot.Soc. Edinburgh 15:274. 1884. S EE Wang &Zhu(2008). Lejeuneamehrana M.Dey,D.K.Singh et D.Singh, J.Bryol. 30:128. 2008. India [Sikkim].S EE Dey etal. (2008b). Lejeuneametzgeriopsis K.I.Goebel, Flora 72:2.1889. O THER USE: Cololejeunea metzgeriopsis (K.I.Goebel) Gradst.,R.Wilson,Ilk.-Borg. et Heinrichs,2006. Lejeuneaperpapillosa M.E.Reiner et K.C.Pôrto, NovaHedwigia 85:542.2007. Brazil [Pernambuco].S EE Reiner-Drehwald &Pôrto(2007). 16M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Lejeuneasanctae-helenae Wigginton, J.Bryol. 29:12.2007.St.Helena.S EE Wigginton (2007). Lejeuneasullivantii Gottsche, Mexik. Leverm.:196.1863.O THER USE: Lepidolejeuneasullivantii (Gottsche) M.E.Reiner,2006. Lejeuneatalamancensis M.E.Reiner et Schäf.-Verw., NovaHedwigia 87:414. 2008. CostaRica.S EE Reiner-Drehwald &Schäfer-Verwimp (2008). Lejeuneatopoensis Gradst. et M.E.Reiner, Syst.Bot. 32:488. 2007.Ecuador.S EE Gradstein &Reiner-Drehwald (2007). Lejeuneaturbinifera Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:295. 1884. O THER USE: Cololejeuneaturbinifera (Spruce) León-Yánez,Gradst. et Wegner,2006. Lejeuneaxiphophylla (Herzog) M.E.Reiner, NovaHedwigia 87:409. 2008. B ASIONYM: Taxilejeuneaxiphophylla Herzog, SvenskBot.Tidskr. 46:99. 1952.S EE Reiner-Drehwald &Schäfer-Verwimp (2008). Lepidolejeuneacordifissa (Taylor)M.E.Reiner in Reiner-Drehwald &Schäfer- Verwimp, NovaHedwigia 83:478. 2006.B ASIONYM: Lejeuneacordifissa Taylor, London J.Bot. 5:395. 1846.S EE Reiner-Drehwald &Schäfer- Verwimp (2008). Lepidolejeuneasullivantii (Gottsche) M.E.Reiner in Reiner-Drehwald & Schäfer-Verwimp, NovaHedwigia 83:479. 2006.B ASIONYM: Lejeunea sullivantii Gottsche, Mexik. Leverm.:196.1863.S EE Reiner-Drehwald & Schäfer-Verwimp (2008). Lepidoziaazorica H.Buch et Perss., Commentat.Biol. 8:4. 1941. O THER USE: Telaraneaazorica (H.Buch et Perss.) Pócs ex Schumacker et Vá≈ a, 2005. Leptolejeuneaarunachalensis SudipaDas et D.K.Singh, J.Jap. Bot. 83:343.2008. India[ArunachalPradesh].S EE Das&Singh (2008). Leptophyllopsoideae J.J.Engel, NovaHedwigia 85:422.2007.T YPE: Leptophyllopsis R.M.Schust., J.HattoriBot.Lab. 26:269. 1963.S EE Engel (2007b). Leucolejeunea subg. Neoleucolejeunea R.M.Schust., J.HattoriBot.Lab. 100:375. 2006.T YPE: Leucolejeuneacaducifolia Gradst. et Schäf.-Verw.S EE Schuster(2006). Leucolejeunea subg. Omphalanthopsis (R.M.Schust.) R.M.Schust., J.HattoriBot. Lab. 100:382.2006.T YPE: Leucolejeuneadecurrens (Steph.) Mizut., J.HattoriBot.Lab. 40:443.1976. S EE Schuster(2006). Leucolejeunea subg. Pseudoleucolejeunea R.M.Schust., J.HattoriBot.Lab. 100: 376.2006.T YPE: Leucolejeuneasuprema Grolle et Piippo. S EE Schuster (2006). Lobatiriccardiacoronopus (DeNot.exSteph.) Furuki, J.HattoriBot.Lab. 100: 90.2006.B ASIONYM: Aneuracoronopus DeNot. ex Steph.,1893.S EE Furuki (2006b). Lobatiriccardiayunnanensis Furuki et D.G.Long, J.Bryol. 29:162.2007.China. S EE Furuki &Long (2007). Lophocoleabrookwoodiana Paton et Sheahan, J.Bryol. 28:163.2006.United Kingdom [England].S EE Paton &Sheahan(2006). * Lophocoleagemmiparous A.Evans ex W.T.Doyle et Stotler, Madroño 53:135. 2006.N OTE:nom. inval. Art.32.1(d);Doyle &Stotler(2006)erronously used the name Lophocoleagemmiparous A.Evans,based on Chiloscyphus gemmiparus A.Evans.S EE Doyle &Stotler(2006). Lophocoleapolychaeta Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:436.1885. O THER USE: Heteroscyphus polychaetus (Spruce) Hentschel et Heinrichs, 2007. EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200817

Lophocoleaspegazziniana C.Massal., NuovoGiorn. Bot.Ital. 17:255. 1885. O THER USE: Chiloscyphus spegazzinianus (C.Massal.) Hentschel et Heinrichs,2007. Lophocoleales W.Frey et M.Stech, NovaHedwigia 87:264. 2008. T YPE: Lophocolea (Dumort.) Dumort., Recueil Observ.Jungerm. 17.1835. S EE Frey&Stech(2008). * Lopholejeuneainflata Steph. exR.L.Zhu et Gradst., Monogr.Syst.Bot.Missouri Bot.Gard. 74:86.2005. Dominica.N OTE:nom. invalArt.34.1(c);sub. Lopholejeuneanigricans (Lindenb.) Schiffn. S EE Zhu&Gradstein (2005). * Lopholejeuneakarstenii Steph. ex R.L.Zhu et Gradst., Monogr.Syst.Bot. MissouriBot.Gard. 74:86.2005. Indonesia[Java].N OTE:nom. inval. Art.34.1(c),sub Lopholejeuneaapplanata (Reinw.,Blume et Nees) Schiffn. S EE Zhu&Gradstein (2005). * Lopholejeuneanongstoinii AjitP.Singh et V.Nath, Phytotaxonomy 6:32.2006 [2007].India[Meghalaya].N OTE:nom. inval. Art.37.3.S EE Singh &Nath (2007c). Lopholejeuneavojtkoana Gyarmati, NovaHedwigia 87:480.2008. Fiji. S EE Sass- Gyarmati(2008). * Lophoziaandrogyna R.M.Schust.,Beih. NovaHedwigia 119:266.2002.Australia [Tasmania].N OTE:nom. inval. Art.36.1. S EE Schuster(2002). Lophoziadusenii Steph., Bihang Bih. Kongl. SvenskaVetensk.-Akad. Handl. 26:24. 1900.O THER USE: Hypolophoziadusenii (Steph.) Bakalin,2006. Lophoziamonoica (E.A.Hodgs.) J.J.Engel, Novon 17:311. 2007.B ASIONYM: Metahygrobiellamonoica E.A.Hodgs., Trans.Roy.Soc.NewZealand, Bot. 3:76.1965. S EE Engel (2007a). Lunulariales D.G.Long, Edinburgh J.Bot. 63:259. 2006.T YPE: Lunularia Adans., Fam. Pl. (Adanson) 2:15. 1763 S EE Long (2006).

Makinoiineae He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27. 2006.B ASED ON:MakinoaceaeNakai, Ord. Fam. Trib.Gen. Sect.nov.ed.: 201. 1943.T YPE: Makinoa Miyake, Hedwigia 38:202.1899. S EE He- Nygrén etal .(2006). Manniacontroversa (Meyl.) D.B.Schill, Edinburgh J.Bot. 65:45. 2008. B ASIONYM: Grimaldiacontroversa Meyl., Beitr.Kryptogamenfl. Schweiz 6:87.1924. S EE Schill etal .(2008). Manniacontroversa subsp. asiatica D.B.Schill et D.G.Long, Edinburgh J.Bot. 65: 45. 2008. China[Qinghai].S EE Schill etal .(2008). Manniacontroversa subsp. controversa , Edinburgh J.Bot. 65:45. 2008. S EE Schill etal .(2008). Manniafragrans (Balb.) Frye et L.Clark,1937. Manniafragrans var. brevipes (Kaal.) Damsh., Ill. Fl. Nord. Liverw.Hornw. : 717.2002.B ASIONYM: Grimaldiafragrans var. brevipes Kaal., Forh. Vidensk.-Selsk. Kristiania 1898:3.1898. S EE Damsholt(2002). Marsupellasprucei (Limpr.) Bernet,1888. Marsupellasprucei var. sprucei, Ill. Fl. Nord. Liverw.Hornw. :262,2002.S EE Damsholt(2002). Marsupellasprucei var. neglecta (Limpr.) Damsh., Ill. Fl. Nord. Liverw.Hornw. : 262.2002.B ASIONYM: Sarcocyphosneglectus Limpr., Forh. Vidensk.- Selsk. Kristiania 58:180.1880[1881].S EE Damsholt(2002). Marsupellasprucei var. ustulata (Spruce) Damsh., Ill. Fl. Nord. Liverw.Hornw. : 262.2002.B ASIONYM: Marsupellaustulata Spruce, Rev.Bryol. 8:100. 1881. S EE Damsholt(2002). 18 M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Marsupellaustulata Spruce, Rev.Bryol. 8:100.1881. O THER USE: Marsupella sprucei var. ustulata (Spruce) Damsh.,2002. Marsupidiumpapillosum J.J.Engel et Glenny, NovaHedwigia 87:289. 2008. New Zealand [SouthIsland].S EE Engel &Glenny(2008b). Marsupidiumperpusillum (Colenso) E.A.Hodgs.,1958. Marsupidiumperpusillum var. denticulatum J.J.Engel et Glenny, NovaHedwigia 87:284. 2008. NewZealand [SouthIsland].S EE Engel &Glenny(2008b). Marsupidiumperpusillum var. perpusillum , NovaHedwigia 87:284. 2008. Marsupidiumplagiochiloides J.J.Engel et Glenny, NovaHedwigia 87:284. 2008. NewZealand [SouthIsland].S EE Engel &Glenny(2008b). Mastigobryumcorbieri Steph., J.&Proc.Roy.Soc.NewSouthWales 48:122. 1914. O THER USE: Bazzaniacorbieri (Steph.) Meagher,2008. Mastigolejeuneaauriculata (Wilson) Schiffn.,1893. Mastigolejeuneaauriculata var. auriculata , Bull. Bot.Surv.India 49:231. 2007. S EE Daniels&Daniel (2007). Mastigolejeuneaauriculata var. ciliata (U.S.Awasthi et Udar)A.E.D.Daniels et P.Daniel, Bull. Bot.Surv.India 49:231. 2007.B ASIONYM: Mastigolejeunea humilis var. ciliata U.S.Awasthi et Udar, Proc.IndianAcad. Sci. Pl. Sci. 93:487.1984. S EE Daniels&Daniel (2007). Mastigolejeuneahumilis (Gottsche) Schiffn.,1893. Mastigolejeuneahumilis var. ciliata U.S.Awasthi et Udar, Proc.IndianAcad. Sci. Pl. Sci. 93:487.1984. O THER USE: Mastigolejeuneaauriculata var. ciliata (U.S.Awasthi et Udar)A.E.D.Daniels et P.Daniel,2007. Megaceroscanaliculatus (Pagán) A.J.Shaw et Renzaglia ex J.C.Villarreal et N.Salazar in Dauphin etal., Trop. Bryol. 27:82.2006.B ASIONYM: Dendroceroscanaliculatus Pagán, Bryologist 45:111. 1942.S EE Dauphin etal. (2006). Meinungeria Frank.Müll., Bryologist 110:494. 2007.T YPE: Meinungeriamouensis Frank.Müll. S EE Müller(2007). Meinungeriamouensis Frank.Müll., Bryologist 110:494. 2007.NewCaledonia. S EE Müller(2007). Metahygrobiellamonoica E.A.Hodgs., Trans.Roy.Soc.NewZealand,Bot. 3:76. 1965. O THER USE: Lophoziamonoica (E.A.Hodgs.) J.J.Engel, Novon 17:311. 2007. † Metzgeriothallus sharonae Hernick,Landing et Bartowski, Rev.Palaeobot. Palynol. 148:160.2008. UpperMiddle Devonian. S EE Hernick etal. (2008). Mniolomanovaezelandiae J.J.Engel, Cryptog. Bryol. 27:111. 2006.NewZealand [SouthIsland].S EE Engel (2006b). Mniolomastamatotonum M.A.M.Renner et E.A.Br., Fieldiana, Bot. ,n.s.47:173. 2008. Australia[Queensland].S EE Renner&Brown(2008). Moerckiajaponica Inoue, Bull. Natl. Sci. Mus.,Tokyo,B11:8. 1985. O THER USE: Sandeothallus japonicus (Inoue) Crand.-Stotl. et Stotler,2007. Moerckiaceae Stotler et Crand.-Stotl., Beih. NovaHedwigia 131:54. 2007.T YPE: Moerckia Gottsche, HepaticaeEuropaeExsiccatae :121. 1860.S EE Stotler &Crandall-Stotler(2007).

Nardiaminutifolia Furuki, Bryol. Res. 9:73.2006.Japan[Honshu].S EE Furuki (2006a). Neohodgsoniaceae D.G.Long, Edinburgh J.Bot. 63:258. 2006.T YPE: Neohodgsonia Perss., Bot.Not. 107:39. 1954. S EE Long (2006). Neohodgsoniales D.G.Long, Edinburgh J.Bot. 63:258. 2006.T YPE: Neohodgsonia Perss., Bot.Not. 107:39. 1954. S EE Long (2006). EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200819

Nephelolejeuneaconchophylla Grolle, J.HattoriBot.Lab. 48:164. 1980.O THER USE: Austrolejeuneaconchophylla (Grolle) Pócs,2006. Nephelolejeuneahamata Grolle, J.HattoriBot.Lab. 48:167.1980.O THER USE: Austrolejeuneahamata (Grolle) Pócs,2006. Nephelolejeuneapapillosa Glenny, NewZealand J.Bot. 34:195. 1996.O THER USE: Austrolejeuneapapillosa (Glenny)Pócs,2006. Noterocladales W.Frey et M.Stech, NovaHedwigia 87:262.2008. B ASED ON: Pelliaceaesubfam. NoterocladoideaeR.M.Schust., J.HattoriBot.Lab. 70: 145. 1991. T YPE: Noteroclada Taylor ex Hook.f. et Wilson, London J.Bot. 3:166.1844. S EE Frey&Stech(2008). Nothoceroscanaliculatus (Pagán) J.C.Villarreal,Hässel et N.Salazar, Bryologist 110:283.2007.B ASIONYM: Dendroceroscanaliculatus Pagán, Bryologist 45:111. 1942.S EE Villarreal etal. (2007). Nothocerosendiviaefolius (Mont.) J.Haseg. ex J.C.Villarreal,Hässel et N.Salazar, Bryologist 110:283.2007.B ASIONYM: Anthocerosendiviaefolius Mont., Voy.Pole Sud 1:211. 1845. S EE Villarreal etal. (2007). Nothocerosgiganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal,Hässel et N.Salazar, Bryologist 110:283.2007.B ASIONYM: Anthocerosgiganteus Lehm. et Lindenb., Nov.Stirp. Pug. 4:25. 1832.S EE Villarreal etal. (2007). Nothocerossuperbus J.C.Villarreal,Hässel et N.Salazar, Bryologist 110:280.2007. CostaRica.S EE Villarreal etal. (2007). Notothylatidae R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:240. 2007.T YPE: Notothylas Sull. ex A.Gray, AmericanJournalofScienceand Arts,ser.2,1:74. 1846.S EE Villarreal etal. (2007). † Notothylites Chitaley et Yawale, Botanique 9:111-118. 1980.Eocene-Oligocene. S EE Chitaley&Yawale (1980). † Notothylitesnirulai Chitaley et Yawale, Botanique 9:111-118. 1980.Eocene- Oligocene. S EE Chitaley&Yawale (1980). *Nowelliacurvifolia var. baueri Dragi† evi † et Velji† , Pregl. mahov.Crne Gore :19. 2006.MonteNegro. N OTE:nom. inval. Art.32.1(d),apparentlybased on Jungermanniabaueri Mart.,1817.S EE Dragi† evi † &Velji† (2006).

* Odontoschismaalpinum R.M.Schust., Beih. NovaHedwigia 119:83.2002. Venezuela[Mérida].N OTE:nom. inval.,no Latin description. S EE Schuster(2002). Odontoschismaengelii Gradst. et Burghardt, Fieldiana, Bot. ,n.s.47:194. 2008. Ecuador.S EE Gradstein &Burghardt(2008). Omphalanthus subg. Evansiolejeunea (Vanden Berghen) R.M.Schust., J.Hattori Bot.Lab. 100:395. 2006.T YPE: Omphalanthus roccatii (Gola) R.M.Schust., Beih. NovaHedwigia 9:96.1963.S EE Schuster(2006). Omphalanthus subg. Peltolejeunea (Spruce) R.M.Schust., J.HattoriBot.Lab. 100: 393.2006.T YPE: Omphalanthus ovalis (Lindenb. et Gottsche) Gradst., Proc.Kon. Ned. Akad. Wetensch. C 80:411. 1977.S EE Schuster(2006). Omphalanthus baracoensis Mustelier,M.E.Reiner et Gradst., J.Bryol. 29: 95. 2007.Cuba.S EE Reiner-Drehwald etal. (2007).

Pachyglossadissitifolia Herzog, Rev.Bryol. Lichénol. 21:259. 1952.O THER USE: Chiloscyphus dissitifolius (Herzog et Grolle) Hentschel et Heinrichs,2007. Pachyglossagrolleana Vá≈ a, Cryptog. Bryol. 26:86.2005. O THER USE: Chiloscyphus grolleanus (Vá≈ a)Hentschel et Heinrichs,2007. Paraphymatoceros Hässel, Phytologia 88:208. 2006.T YPE: Paraphymatoceros diadematus Hässel. S EE Hässel de Menéndez(2006a). 20 M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Paraphymatoceroscoriaceus (Steph.) Stotler, Fieldiana , Bot .,n.s.47:233.2008. B ASIONYM: Anthoceroscoriaceus Steph., Spec.Hepat. (Stephani) 5:991. 1916.S EE Crandall-Stotler etal. (2008). Paraphymatocerosdiadematus Hässel, Phytologia 88:209. 2006.Chile [Valparaíso].S EE Hässel de Menéndez(2006a). Paraphymatocerosfimbriatus (Gottsche) Stotler, Fieldiana, Bot. ,n.s.47:233. 2008. B ASIONYM: Anthocerosfimbriatus Gottsche, Ann. Sci. Nat.,Bot. , sér.51:187.1864. S EE Crandall-Stotler etal. (2008). Paraphymatoceroshallii (Austin) Hässel, Phytologia 88:209. 2006.B ASIONYM: Anthoceroshallii Austin, Bull. TorreyBot.Club 6:26.1875. S EE Hässel de Menéndez(2006a). Paraphymatoceroshirticalyx (Steph.) Stotler, Fieldiana, Bot. ,n.s.47:233.2008. B ASIONYM: Aspiromitus hirticalyx Steph., Spec.Hepat. (Stephani) 5:966. 1916.S EE Crandall-Stotler etal. (2008). Paraphymatocerosminutus (Mitt.) Hässel, Phytologia 88:210.2006.B ASIONYM: Anthocerosminutus Mitt., J.Linn. Soc.,Bot. 16:195. 1877.[1878].N OTE: erroneouslypublished as Phymatocerosminutus (cf. abstract). S EE Hässel de Menéndez(2006a). Paraphymatocerosskottsbergii (Steph.) Stotler, Fieldiana, Bot. ,n.s.47:234. 2008. B ASIONYM: Anthocerosskottsbergii Steph., Kongl. Svenska Vetenskapsakad. Handl.,n.s.46:90.1911. S EE Crandall-Stotler etal. (2008). * Pelliaalpicola (R.M.Schust.) Damsh., Lindbergia 31:98. 2007.N OTE:nom. inval. Art.37.7,based on Pelliaendiviifolia subsp. alpicola R.M.Schust., 1991,nom. inval. Art.37.7(no herbariumspecified). S EE Damsholt (2007). * Pelliaendiviifolia subsp. alpicola R.M.Schust., J.Hatt.Bot.Lab. 70:145. 1991. N OTE:nom. inval. Art.37.7(no herbariumspecified). O THER USE: Pellia alpicola (R.M.Schust.) Damsh.,2007. Pelliales He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27.2006. B ASED ON:PelliineaeR.M.Schust. ex Schljakov.T YPE: Pellia Raddi, Jungermanniogr.Etrusca:38. 1818. S EE He-Nygrén etal. (2006). Pelliidae He-Nygrén,Juslén,Ahonen,Glenny et Piippo, Cladistics 22:27.2006. T YPE: Pellia Raddi, Jungermanniogr.Etrusca:38. 1818. S EE He-Nygrén etal. (2006). Phaeocerosdonnellii (Austin) J.Haseg. ex Stotler et Crand.-Stotl., Bryologist 108: 24. 2005. B ASIONYM: Anthocerosdonnellii Austin, Bull. TorreyBot.Club 6:304. 1879. S EE Stotler etal. (2005). Phaeocerosengelii Cargill et Fuhrer, Fieldiana, Bot. ,n.s.47:248. 2008. Australia [Victoria].S EE Cargill &Fuhrer(2008). Phaeocerosevanidus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. ,n.s.47:245. 2008. B ASIONYM: Anthocerosevanidus Steph., Spec.Hepat. (Stephani) 5:990. 1916.S EE Cargill &Fuhrer(2008). Phaeocerosinflatus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. ,n.s.47:246.2008. B ASIONYM: Anthocerosinflatus Steph., Spec.Hepat. (Stephani) 5:990. 1916.S EE Cargill &Fuhrer(2008). Phaeocerosproskaueri Stotler,Crand.-Stotl. et W.T.Doyle, Fieldiana, Bot. ,n.s.47: 232.2008. U.S.A.[California].S EE Crandall-Stotler etal. (2008). Phaeolejeuneaamicorum (Hürl.) Pócs, Fieldiana, Bot. ,n.s.47:140.2008. B ASIONYM: Phaeolejeuneaetesseana subsp. amicorum Hürl.,Bauhinia9: 263.1991. S EE Pócs(2008b). Phaeolejeuneaetesseana (Steph.) Mizut.,1968. EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200821

Phaeolejeuneaetesseana subsp. amicorum Hürl., Bauhinia 9:263.1991. O THER USE: Phaeolejeuneaamicorum (Hürl.) Pócs,2008. Phaeomegaceros R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110: 241. 2007.T YPE: Phaeomegacerosfimbriatus (Gottsche. R.J.Duff, J.C.Villarreal,Cargill et Renzaglia, 2007.S EE Duff etal. (2007). Phaeomegaceroscoriaceus (Steph.) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:241. 2007.B ASIONYM: Anthoceroscoriaceus Steph., Spec. Hepat. (Stephani) 5:991. 1916.S EE Duff etal. (2007). Phaeomegacerosfimbriatus (Gottsche) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:241. 2007.B ASIONYM: Anthocerosfimbriatus Gottsche, Ann. Sci. Nat.,Bot. sér.51:187.1864. S EE Duff etal. (2007). Phaeomegaceroshirticalyx (Steph.) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:241. 2007.B ASIONYM: Aspiromitus hirticalyx Steph., Spec. Hepat. (Stephani) 5:966.1916.S EE Duff etal. (2007). Phaeomegacerosskottsbergii (Steph.) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:241. 2007.B ASIONYM: Anthocerosskottsbergii Steph., Kongl. SvenskaVetenskapsakad. Handl. 46:90.1911. S EE Duff et al. (2007). Phaeomegacerotoideae R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:241. 2007.T YPE: Phaeomegaceros R.J.Duff,J.C.Villarreal,Cargill et Renzaglia . , Bryologist 110:41. 2007.S EE Duff etal. (2007). Phymatocerosphymatodes (M.Howe) R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:240.2007.B ASIONYM: Anthocerosphymatodes M.Howe, Bull. TorreyBot.Club 25:12.1898. S EE Duff etal. (2007). Phymatocerotaceae R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110: 240.2007.T YPE: Phymatoceros Stotler,W.T.Doyle et Crand.-Stotl. S EE Duff etal. (2007). Phymatocerotales ord. R.J.Duff,J.C.Villarreal,Cargill et Renzaglia, Bryologist 110:240.2007.T YPE: Phymatoceros Stotler,W.T.Doyle et Crand.-Stotl. S EE Duff etal. (2007). Pictolejeuneapiconii Pócs, ActaBot.Hung. 49:110.2007.Venezuela[Bolívar]. S EE Pócs(2007). Plagiochilaartsii Pócs, J.HattoriBot.Lab. 100:334. 2006.Réunion. S EE Pócs (2006c). Plagiochilagymnocalycina (Lehm. et Lindenb.) Lindenb.,1839. Plagiochilagymnocalycina var. surinamensis (Molk. ex Sande Lac.) Heinrichs et D.S.Rycroft, J.HattoriBot.Lab. 100:139. 2006.B ASIONYM: Plagiochila surinamensis Molk. exSande Lac., Verh. Kon. Ned. Akad. Wetensch.,Afd. Natuurk.,Sect.2 ,5:103.1856[1857].S EE Heinrichs etal. (2006). Plagiochilahiroshiana Pócs, J.HattoriBot.Lab. 100:335. 2006.Tanzania.S EE Pócs(2006c). Plagiochilakiaeri Gottsche,1882. Plagiochilakiaeri var. myriocarpa (Pearson) Pócs, J.E.Afr.Nat.Hist. 96:38. 2007.B ASIONYM: Plagiochilamyriocarpa Pearson, Ark. Bot. 1924. S EE Pócs&Luke (2007). Plagiochilameghalayensis K.K.Rawat et S.C.Srivast., Geophytology 35:49. 2005. India[Meghalaya].S EE Rawat&Srivastava(2005). Plagiochilamyriocarpa Pearson, Ark. Bot. 1924. O THER USE: Plagiochilakiaeri var. myriocarpa (Pearson) Pócs,2007. Plagiochilanorvegica H.H.Blom et Holten, Lindbergia 14:8. 1988. O THER USE: Plagiochilaporelloides var. norvegica (H.H.Blom et Holten) Schumacker et Vá≈ a, 2005. 22 M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

Plagiochilapalangiensis S.C.Srivast.,K.K.Rawat etP.K.Verma , Natl. Acad. Sci. Lett. 29:267.2006.India[Tamil Nadu].S EE Srivastava etal. (2006). Plagiochilaporelloides (Torr. ex Nees)Lindenb.,1840. Plagiochilaporelloides var. norvegica (H.H.Blom et Holten) Schumacker et Vá≈ a, Identif. keys liverw.Hornw.Europe,2nd ed.:131. 2005. B ASIONYM: Plagiochilanorvegica H.H.Blom et Holten, Lindbergia 14:8. 1988. S EE Schumacker&Vá ≈ a(2005). Plectocoleainfusca Mitt., Trans.Linn. Soc.London,Bot. 3:196.1891. O THER USE: Solenostomainfuscum (Mitt.) Hentschel,2007. Plectocoleainfusca var. ovicalyx (Steph.) Bakalin, Arctoa 16:207.2007. B ASIONYM: Solenostomaovicalyx Steph., Spec.Hepat. (Stephani) 6:82. 1917.S EE Bakalin (2007). *Proskauera Heinrichs et J.J.Engel, Pl. Syst.Evol. 258:235. 2006.T YPE: Proskauerapleurata (Hook.f. et Taylor)Heinrichs et J.J.Engel, Pl. Syst. Evol. 258:235. 2006.S EE Heinrichs etal. (2006). N OTE:nom. illeg. Art. 52.1,type of Dinckleria Trevis1877 included. S EE Engel &Heinrichs (2008). Proskauerafruticella (Hook.f. et Taylor)Heinrichs et J.J.Engel, Pl. Syst.Evol. 258: 237.2006.B ASIONYM: Jungermanniafruticella Hook.f. et Taylor, London J.Bot. 3:565. 1844. S EE Heinrichs etal. (2006). O THER USE: Dinckleria fruticella (Hook.f. et Taylor)J.J.Engel et Heinrichs,Engel &Heinrichs, 2008. Proskauerapleurata (Hook.f. et Taylor)Heinrichs et J.J.Engel, Pl. Syst.Evol. 258: 235. 2006.B ASIONYM: Jungermanniapleurata Hook.f. et Taylor, London J.Bot. 3:372.1844. S EE Heinrichs etal. (2006). O THER USE: Dinckleria pleurata (Hook.f. et Taylor)Trevis.,Engel &Heinrichs,2008. Pseudolepicoleales W.Frey et M.Stech, NovaHedwigia 87:263.2008. T YPE: Pseudolepicolea Fulford et J.Taylor, NovaHedwigia 1:412.1960.S EE Frey&Stech(2008). Pseudomarsupidiumaureocinctum (R.M.Schust.) J.J.Engel, Novon 17:312.2007. B ASIONYM: Adelanthus decipiens subsp. aureocinctus R.M.Schust., Phytologia 39:250.1978. S EE Engel (2007a). Pseudomarsupidiumcrossii (Spruce) J.J.Engel, Novon 17:312.2007.B ASIONYM: Adelanthus crossii Spruce, Trans.&Proc.Bot.Soc.Edinburgh 15:404. 1885. S EE Engel (2007a). Pycnolejeunearetusa R.M.Schust., J.HattoriBot.Lab. 2006.100:402.PapuaNew Guinea.S EE Schuster(2006).

Radulafujitae Furuki, Bryol. Res. 9:143.2007.Japan[Nansei-shoto].S EE Furuki (2007). Radulapseudoscripta M.A.M.Renner, NewZealand J.Bot. 44:340.2006.New Zealand [NorthIsland].S EE Renner(2006). Riccardiamagnicellularis Furuki, J.HattoriBot.Lab. 100:93.2006.Philippines [NegrosIsland].S EE Furuki (2006b). Riccardiaphilippinensis Furuki, J.HattoriBot.Lab. 100:94. 2006.Philippines [Luzon Island].S EE Furuki (2006b). Riccardiapolyclada (Mitt. ex Thurn) Hässel, Bryologist 109:34. 2006.B ASIONYM: Aneurapolyclada Mitt. ex Thurn,1886.S EE Hässel de Menéndez(2006b). Ricciapoihaiana A.E.D.Daniels et P.Daniel, Bull. Bot.Surv.India 44:135. 2002 [2003].India[Tamil Nadu].S EE Daniels&Daniel (2002). Ricciavelimalaiana A.E.D.Daniels et P.Daniel, Bull. Bot.Surv.India 44:139. 2002 [2003].India[Tamil Nadu].S EE Daniels&Daniel (2002). EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200823

Saccogynidiumirregularospinum C.Gao,T.Cao et M.J.Lai, Bryologist 104:129. 2001. China[Xizang].S EE Gao&Cao(2001). Sandeothallus japonicus (Inoue) Crand.-Stotl. et Stotler in Stotler&Crandall- Stotler, Beih. NovaHedwigia 131:58. 2007.B ASIONYM: Moerckia japonica Inoue, Bull. Natl. Sci. Mus.,Tokyo,B11:8. 1985. S EE Stotler& Crandall-Stotler(2007). Sarcocyphosneglectus Limpr., Jahresber.Schles.Ges.Vaterl. Cult. 58:180.1880 [1881].O THER USE: Marsupellasprucei var. neglecta (Limpr.) Damsh., 2002. Scapanianemorea (L.) Grolle,1963. *Scapanianemorea var. purpurascens Dragi† evi † et Velji† , Pregl. mahov.Crne Gore :17.2006.MonteNegro. N OTE:nom. inval. Art.32.1(d) (no description),apparentlybased on Jungermannianemorosa var. purpurascens Hook.,1812.S EE Dragi† evi † &Velji† (2006). Scapaniaparaphyllia T.Cao,C.Gao,J.Sun et B.R.Zuo in Zuo etal., Acta Phytotax.Sin. 45:311. 2007.China[Zhejiang].S EE Zuo etal. (2007). Schiffneriayunnanensis C.Gao et W.Li, Bull. Bot.Res.,Harbin 26:390.2006. China[Yunnan].S EE Li etal. (2006). † Schizolepidellabirkenmajeri Ociepa, PolishPolarRes. 28:34. 2007.Jurassic. S EE Ociepa(2007). Sendtnerapensilis Taylor, London J.Bot. 5:372.1846.O THER USE: Herbertus juniperoideus subsp. pensilis (Taylor)Feldberg et Heinrichs,2006. Solenostomacryptogynum R.M.Schust. ex J.J.Engel, Novon 17:311. 2007.New Zealand [NorthIsland].S EE Engel (2007a). *Solenostomahasskarlianum (Nees)R.M.Schust., Beih. NovaHedwigia 119:378. 2002.N OTE:nom. inval. Art.33.4 (basionymnotcited),apparentlybased on Aliculariahaskarliana Nees,1844. S EE Schuster(2002). *Solenostomahodgsoniae (Grolle) J.J.Engel, Novon 17:312.2007.B ASIONYM: Jungermanniahodgsoniae Grolle, Bot.Mag. 78:83.1965. N OTE:nom. illeg. Art.52.1,incl. sp. prior.,the basionym Jungermanniahodgsoniae Grolle wasanom. nov.for Lethocoleagrandifolia Berggr.(non Jungermanniagrandifolia Hook.f. et Taylor,1844). S EE Engel (2007a). Solenostomainfuscum (Mitt.) Hentschel, Pl. Syst.Evol. 268:152.2007. “infusca” B ASIONYM: Plectocoleainfusca Mitt., Trans.Linn. Soc.London,Bot. 3: 196.1891. S EE Hentschel etal. (2007b). *Solenostomajenseniana (Grolle) Bakalin, Arctoa 12:89. 2003.B ASIONYM: Jungermanniajenseniana Grolle, Österr.Bot.Z. 111:190.1964. N OTE: nom. illeg. Art.52.1,incl. sp. prior.,the basionym Jungermannia jenseniana Grolle wasanom. nov.for Aploziapusilla C.E.O.Jensen (non Jungermanniapusilla L.,1753). S EE Bakalin (2003). *Solenostomaobliquifolium (Schiffn.) R.M.Schust., Beih. NovaHedwigia 119: 378. 2002.N OTE:nom. inval. Art.36.1 (no Latin description),apparently based on Nardiaobliquifolia Schiffn.,1898). S EE Schuster(2002). Solenostomaovicalyx Steph., Spec.Hepat. (Stephani) 6:82.1917.O THER USE: Plectocoleainfusca var. ovicalyx (Steph.) Bakalin,2007. Solenostomapyriflorum Steph.,1917. Solenostomapyriflorum var. minutissimum (Amakawa)Bakalin, Arctoa 16:208. 2007.B ASIONYM: Jungermanniapyriflora var. minutissima Amakawa, J.HattoriBot.Lab. 22:61. 1960.S EE Bakalin (2007). Solenostomarufiflorum (Colenso) J.J.Engel, Novon 17:312.2007.B ASIONYM: Jungermanniarufiflora Colenso, Trans.&Proc.NewZealand Inst. 18: 237.1886.S EE Engel (2007a). 24M.v.Konrat,L.Söderström,A.Hagborg,M.R.Crosby&J.J.Engel

*Solenostomatruncatum (Nees)R.M.Schust., Beih. NovaHedwigia 119:378. 2002.N OTE:nom. inval. Art.33.4 (basionymnotcited),apparentlybased on Jungermanniatruncata Nees,1830.S EE Schuster(2002). *Solenostomawattsianum (Steph.) R.M.Schust., Beih. NovaHedwigia 119:378. 2002.N OTE:nom. inval. Art.33.4 (basionymnotcited),apparentlybased on Jungermanniawattsiana Steph.,1917.S EE Schuster(2002). Sphenolobus langkyrdumii AjitP.Singh et V.Nath, Hepat.KhasiJaintiaHills: East.Himalayas 93.2007.India[Meghalaya].S EE Singh &Nath(2007b).

Taxilejeuneaxiphophylla Herzog, SvenskBot.Tidskr. 46:99. 1952.O THER USE: Lejeuneaxiphophylla (Herzog) M.E.Reiner,2008. Telaraneaazorica (H.Buch et Perss.) Pócs ex Schumacker et Vá≈ a, Identif. keys liverw. Hornw.Europe,2nd ed.:160.2005. B ASIONYM: Lepidoziaazorica H.Buch et Perss., Commentat.Biol. 8:4. 1941. S EE Schumacker&Vá ≈ a(2005). Telaraneabischleriana Pócs, ActaBot.Hung. 48:120.2006.Seychelles.S EE Pócs (2006d). Telaraneaindica (S.C.Srivast. et P.K.Verma)A.E.D.Daniels et P.Daniel, Bull. Bot.Surv.India 49:231. 2007.B ASIONYM: Arachniopsisindica S.C.Srivast. et P.K.Verma, Natl. Acad. Sci. Lett. 27:270.2004. S EE Daniels &Daniel (2007). Telaraneamaorensis Pócs, ActaBot.Hung. 48:124. 2006.Comores[Mayotte].S EE Pócs(2006d). Trabacelluloideae R.M.Schust., Beih. NovaHedwigia 119:41. 2002.B ASED ON: TrabacellulaceaeFulford, in Steyermark, ActaBot.Venez. 2:85,1967. T YPE: Trabacellula Fulford, ActaBot.Venez. 2:86.1967.Cephaloziaceae. SEE Schuster(2002). Treubiidae Stotler et Crand.-Stotl., Taxon 57:290.2008. T YPE: Treubia K.I.Goebel. S EE Stotler&Crandall-Stotler(2008). Trichocoleales W.Frey et Stech, NovaHedwigia 87:263.2008. T YPE: Trichocolea Dumort., Comment.Bot. 113.1822.S EE Frey&Stech(2008). Tritomariaexsecta (Schmidel ex Schrad.) Schiffn. ex Loeske,1909 Tritomariaexsecta subsp. novaezelandiae J.J.Engel, Bryologist 109:61. 2006.New Zealand [SouthIsland].S EE Engel (2006c). Trocholejeuneameghalayensis AjitP.Singh et V.Nath, J.Jap. Bot. 83:2.2008. India[Meghalaya].S EE Singh &Nath(2008). Tylimanthus andinopatagonicus M.Stech et W.Frey in Stech etal., NovaHedwigia 83:28. 2006.Chile [La Araucania].S EE Stech etal. (2006).

Zoopsisbicruris Glenny et E.A.Br. in Renner etal., J.Bryol. 28:332.2006.New Zealand [SouthIsland].S EE Renner etal. (2006). Zoopsismatawaia M.A.M.Renner in Renner etal., J.Bryol. 28:334. 2006.New Zealand [SouthIsland].S EE Renner etal. (2006).

Acknowledgments. Datawerecompiled and edited byAnders Hagborg,Lars Söderström,and Matt von Konrataspart of the “EarlyLand Plants TodayProject”,which waskindlysupported byGBIF (AwardNo. 2007/41) and the NSF funded project“Bringing Schuster,Standleyand Co. Intothe ThirdMillennium:The Field Museum’s180K Bryophyteand Lichen Conversion &Digitization Project”(Award#0749762). Wealso acknowledge the usefulresourcesTROPICOS and the RecentLiteratureseriesof The Bryologist compiled byWilliamR.Buck,BruceAllen,and Ronald A.Pursell. Wealso thank David Long (RoyalBotanicGarden,Edinburgh) forassistancewithsome literature and criticalfeedbackonthe manuscript.Finally,weexpress our gratitude toJörnHentschel (FriedrichSchillerUniversity of Jena)forhiscomments and suggestionson the manuscript. EarlyLand Plants Today:Indexof Liverworts &Hornworts 2006-200825

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