DOCSLIB.ORG

The Genus Anoectangium (Pottiaceae, Bryophyta) in Russia Ðîä Anoectangium (Pottiaceae, Bryophyta) Â Ðîññèè

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
The Genus Anoectangium (Pottiaceae, Bryophyta) in Russia Ðîä Anoectangium (Pottiaceae, Bryophyta)  Ðîññèè Arctoa (2009) 18: 167-176 THE GENUS ANOECTANGIUM (POTTIACEAE, BRYOPHYTA) IN RUSSIA ÐÎÄ ANOECTANGIUM (POTTIACEAE, BRYOPHYTA)  ÐÎÑÑÈÈ ELENA A. IGNATOVA1 ÅËÅÍÀ À. ÈÃÍÀÒÎÂÀ1 Abstract Three species of Anoectangium are currently known in Russia: A. aestivum (Hedw.) Mitt., A. handelii Schiffn. and A. straheyanum Mitt., the latter species is reported for the first time for the country. Anoectangium thomsonii Mitt. is ex- cluded from the bryoflora of Russia. Anoectangium amurense Broth. and A. contortum Broth., both described from the Amurskaya Province of the Russian Far East, are synonymized with A. stracheyanum. Anoectangium aestivum is a rare species known from northwestern European Russia and the Caucasus; all its records from Asia were based on erroneously identified specimens. Anoectangium stracheyanum oc- curs in the East of Asiatic Russia and is locally abundant in Primorsky and Khabarovsk Territories, spreading westwards almost to Baikal Lake in Irkutsk Province. Anoectangium handelii was found only once on Olkhon Island at Baikal Lake. Key for identification, descriptions and illustrations of the species are provided. Ðåçþìå Ðîä Anoectangium ïðåäñòàâëåí â Ðîññèè òðåìÿ âèäàìè: A. aestivum (Hedw.) Mitt., A. handelii Schiffn. è A. straheyanum Mitt., ïîñëåäíèé âèä ïðèâîäèòñÿ äëÿ ñòðàíû âïåðâûå. Anoectangium thomsonii Mitt. èñêëþ÷åí èç ôëîðû ìõîâ Ðîññèè. Anoectangium amurense Broth. è A. contortum Broth., êîòîðûå áûëè îïèñàíû èç Àìóðñêîé îáëàñòè, îòíåñåíû â ñèíîíèìû A. stracheyanum. Anoectangium aestivum ðåäêèé âèä, èçâåñòíûé èç íåìíîãèõ ìåñòîíàõîæäåíèé íà ñåâåðî- çàïàäå åâðîïåéñêîé Ðîññèè è íà Êàâêàçå; âñå óêàçàíèÿ íà íàõîæäåíèå ýòîãî âèäà â àçèàòñêîé ÷àñòè Ðîññèè áûëè îøèáî÷íûìè. Anoectangium stracheyanum âñòðå÷àåòñÿ íà Ðîññèéñêîì Äàëüíåì Âîñòîêå, ìåñòàìè îí äîâîëüíî îáû÷åí â Ïðèìîðñêîì è Õàáàðîâñêîì êðàÿõ; ñàìîå çàïàäíîå ìåñòîíàõîæäåíèå â Èðêóòñêîé îáëàñòè. Anoectangium handelii áûë ñîáðàí òîëüêî îäèí ðàç íà î. Îëüõîí íà Áàéêàëå. Ïðèâîäèòñÿ êëþ÷ äëÿ îïðåäåëåíèÿ âèäîâ, îïèñàíèÿ è èëëþñòðàöèè. KEYWORDS: mosses, Anoectangium, Pottiaceae, taxonomy, Russia, phytogeography In the recent check-list of mosses of East Eu- Subsequent study revealed a number of mis- rope and North Asia (Ignatov, Afonina, Ignatova takes in species identification and also ambigous et al., 2006) three species of Anoectangium application of some names. So, the expanded re- Schwägr. were listed for the territory of Russia: A. vision of collections from Russia was underta- aestivum (Hedw.) Mitt. (reported from northwest- ken, based on herbarium material of MW, MHA, ern European Russia, Caucasus and East Siberia), LE, VLA, IRK, KPABG, H, S and some smaller A. handelii Schiffn. (one locality in Baikal Lake collections. Their study resulted in the following area) and A. thomsonii Mitt. (Russian Far East). taxonomy of the genus in Russia. 1 Department of Geobotany, Biological Faculty, Moscow State University, Moscow 119991 Russia Ðîññèÿ 119991, Ìîñêâà, Ìîñêîâñêèé óíèâåðñèòåò, Áèîëîãè÷åñêèé ôàêóëüòåò, êàô. ãåîáîòàíèêè. E-mail: [email protected] 168 E. A. IGNATOVA ANOECTANGIUM Schwägr., Sp. Musc. Leaves ovate-lanceolate to oblong-lanceolate, Frond. Suppl. 1(1): 33. 1811. >0.6 mm long; laminal cells densely papil- Type species: A. compactum Schwägr. (= A. lose; gemmae absent ................................. 2 aesticum) 2.Leaves incurved when dry, distantly ar- Plants in dense or loose tufts, yellow-green ranged, stem exposed at places; leaves ovate- in upper part, brownish below. Stems 0.5-4 cm, lanceolate, 0.8-1.6 mm long .... A. aestivum weakly branching, usually moderately tomentose, Leaves contorted and slightly or occasional- rounded-triangular in transverse section, with ly strongly spirally twisted when dry, dense- rather strong central strand, weak sclerodermis, ly arranged, stem not exposed; leaves ovate- hyalodermis absent. Leaves distantly or densely lanceolate to oblong lanceolate, (1.0-)1.5-2.5 arranged, straight or contorted and twisted when mm long ........................... A. stracheyanum dry, erect-spreading when wet, uneven in size Anoectangium aestivum (Hedw.) Mitt., J. along the stem, forming alternating zones of Linn. Soc., Bot. 12: 175. 1869. Gymnostomum smaller and larger leaves, ovate, ovate-lanceolate aestivum Hedw., Sp. Musc. Frond. 32. 2 f. 47. or oblong-lanceolate, with not differentiated or 1801. Figs. 1, 5. differentiated base, occasionally constricted above Plants medium-sized, in moderately dense soft base, acute or acuminate, strongly keeled, nar- tufts or cusions, yellowish-green in upper part, rowly grooved along costa ventrally; margins brownish below. Stems 1-4 cm, erect. Leaves dis- plane, entire or slightly crenulate at leaf base, tantly arranged and exposing stem at places, in- sometimes uneven proximally; costa strongly curved and slightly curled when dry, erect-spread- projecting abaxially, ending few cells below apex, ing to patent when wet, ovate-lanceolate or short- percurrent or shortly excurrent, with only dorsal lanceolate, without or with the scarcely differenti- stereid band, dorsal and ventral epidermis, occa- ated base, narrowly acute or acuminate, keeled, sionally ventral epidermis absent and guide cells carinate, 0.8-1.4(-1.6) mm long, 0.25-0.3 mm exposed, ventral surface cells elongate, smooth, wide; margins plane, uneven, slightly papillose- dorsal surface cells elongate, smooth proximal- crenulate except for the leaf base; costa strong, ly, slightly to strongly papillose distally; lamina 40-75 µm wide at base, ending few cells below unistratose, distal and median laminal cells sub- apex or percurrent, papillose dorsally in distal 2/3; quadrate, densely papillose, papillae low and upper and median laminal cells rounded-quadrate, wide, bifid, more rarely loosely papillose, basal rounded-hexagonal or transversely widened, 7- laminal cells short rectangular, shorter to the mar- 10 5-10 µm, with moderately thickened walls, gins, smooth. Specialized asexual reproduction × opaque, densely covered by massive low bifid pa- rare, by multicellular gemmae on rizhoids in leaf pillae; basal laminal cells hyaline, with moderate- axils. Dioicous. Gametangia and sporophytes ter- ly thickened walls, subquadrate to short-rectan- minal on short lateral branches. Perichaetial gular, 10-20 8-12 µm, becoming more narrow to leaves convolute-sheathing, ovate-acuminate. × the margins; several cells at leaf apex smooth, Setae yellow-brownish. Urns yellow-brown, with forming short apiculus. Sporophytes unknown red rim, elliptic, smooth and glossy, exothecial from Russia. Inner perichaetial leaves up to 0.8- cells rectangular, thin-walled, annulus of two 1.0 0.3-0.4 mm, gradually acuminate. [Seta 7-10 rows of slightly vesiculose cells, peristome ab- × mm. Urn 0.5-1.0 mm long. Peristome absent. sent. Spores 12-16 µm, finely papillose. Opercu- Operculum with long oblique beak. Spores 12-16 lum with long oblique beak. KOH laminal reac- µm (from Smith, 2004; Zander & Eckel, 2007)]. tion yellow. Specimens examined: RUSSIA: Murmansk KEY FOR IDENTIFICATION OF SPECIES OF Province: Kutsa Nature Reserve, Jänisköngäs ra- ANOECTANGIUM IN RUSSIA pids, 10.VII.1934, Tuomikoski s.n. (LE); Kara- 1. Leaves ovate, very small, 0.3-0.4(-0.6) mm chaevo-Cherkessia: Teberda Nature Reserve, long; laminal cells loosely papillose; multi- Shumka Creek, 2750 m alt., Onipchenko #205/ cellular gemmae sometimes present in leaf 96; Khutyj Creek, 1600 m alt., Onipchenko #36/ axils........................................... A. handelii 98 (MW); Karachaevsk District, Shubaidai-chat The genus Anoectangium in Russia 169 Gorge, 2600 m alt., Onipchenko #137/94 (MW); upper part, brownish below. Stems 1-2 cm, erect Arkhyz, Sofiya Mt., 25.VIII.1997, 2635 m alt., or often curved downwards in upper part. Leaves Korotkov s.n. (MW). densely arranged and hiding stem, incurved and Differentiation. In most cases A. aestivum is slightly contorted when dry, often slightly secund, separated from A. stracheyanum by ovate-lan- sometimes slightly to strongly spirally twisted, ceolate leaves distantly arranged along the stem. patent when wet, constricted above ovate base Anoectangium hanedlii has much smaller leaves, and then lanceolate, usually slightly widened at 0.3-0.6 mm long vs. to 0.8-1.6 mm long and mid-leaf, narrowly acute or acuminate, strongly grows in very dense tufts and in xeric habitats, keeled, carinate, (1.0-)1.4-2.0(-2.2) mm long, while A. aestivum is rather a mesophytic species. 0.25-0.4 mm wide at base; margins plane, slightly Distribution and ecology. Anoectangium aes- uneven, sometimes slightly crenulate at shoul- tivum is a widespread species, sporadically oc- ders below constriction or entire throughout; costa curring almost throughout the world: in North, strong, 50-75 µm wide at base, excurrent into Central and South America, Africa, Eurasia and short apiculus to 5-6 cells long or percurrent, New Zealand (Zander & Eckel, 2007). However, densely or loosely papillose dorsally in distal 2/ in Russia it is a rare species known from several 3; upper and median laminal cells rounded-quad- localities in the Caucasus and northwestern Eu- rate, rounded-hexagonal or transversely widened, ropean Russia. Caucasian collections were made 6-8×5-12 µm, thin-walled, opaque, densely cov- in Teberda and Caucasian Nature Reserves. In ered by massive low bifid papillae; basal laminal the former the species grows mostly in calcare- cells hyaline, smooth, with moderately thickened ous areas in the alpine belt, at 2600-2750 m elev., walls, rectangular, 12-25×10-12 µm, becoming
Load more

Recommended publications
  • Antibacterial Activity of the Alcoholic Extracts of Entodon Nepalensis Mizush
    ISSN : 2349 – 1604 (Volume – 3, No. 1, January 2016) Research article Indexed in SIS (USA), ASI (Germany), I2OR & i-Scholar (India) and SJIF (Morocco) databases Impact Factor: 3.835 (SJIF) Antibacterial Activity of Anoectangium clarum Mitt. (Bryophyta: Pottiaceae) against some Pathogenic Bacteria Anshul Bishnoi, Vanshika Singh, Vinay Sharma and Afroz Alam* Department of Bioscience and Biotechnology, Banasthali University, Tonk -304 022 (Rajasthan), India *Corresponding author: Dr. Afroz Alam Phone: +919785453594 Manuscript received : 19.10.15 Manuscript accepted: 22.11.15 Abstract In recent years bryophytes have emerged as potential plants for many bioactive phytochemicals with antimicrobial efficacies. In present study antibacterial activity of Anoectangium clarum Mitt. has been assessed using various extracts. For this assessment agar diffusion method is used against selected pathogenic fungus. Tetracycline is used as positive control. The ethanolic extract of moss exhibited a superior effect than the extract prepared in methanol. The utmost effect was observed against Escherichia coli followed by Salmonella typhimurium and Bacillus subtilis. This study projects Anoectangium clarum as an eco-friendly antibacterial agent. Keywords: Anoectangium clarum, antibacterial activity, extract, moss, well diffusion method. 650 SMU Medical Journal, Volume – 3, No. – 1, January, 2016 Introduction Plants are treasured source of natural products for upholding normal health of human beings. Conventional therapeutic systems of Ayurveda, Unani and Sidha are the prevalent verification of their use in therapy. In the last few decades, with more rigorous researches for therapies based on natural systems, plants are used extensively to cure different diseases especially dermal ailments [1]. The search for unexplored plants or plant group with substantial antimicrobial action has attained massive importance these days, due to a growing concern about the attainment of antibiotic- resistance by the pathogenic microorganism.
    [Show full text]
  • Fossil Mosses: What Do They Tell Us About Moss Evolution?
    Bry. Div. Evo. 043 (1): 072–097 ISSN 2381-9677 (print edition) DIVERSITY & https://www.mapress.com/j/bde BRYOPHYTEEVOLUTION Copyright © 2021 Magnolia Press Article ISSN 2381-9685 (online edition) https://doi.org/10.11646/bde.43.1.7 Fossil mosses: What do they tell us about moss evolution? MicHAEL S. IGNATOV1,2 & ELENA V. MASLOVA3 1 Tsitsin Main Botanical Garden of the Russian Academy of Sciences, Moscow, Russia 2 Faculty of Biology, Lomonosov Moscow State University, Moscow, Russia 3 Belgorod State University, Pobedy Square, 85, Belgorod, 308015 Russia �[email protected], https://orcid.org/0000-0003-1520-042X * author for correspondence: �[email protected], https://orcid.org/0000-0001-6096-6315 Abstract The moss fossil records from the Paleozoic age to the Eocene epoch are reviewed and their putative relationships to extant moss groups discussed. The incomplete preservation and lack of key characters that could define the position of an ancient moss in modern classification remain the problem. Carboniferous records are still impossible to refer to any of the modern moss taxa. Numerous Permian protosphagnalean mosses possess traits that are absent in any extant group and they are therefore treated here as an extinct lineage, whose descendants, if any remain, cannot be recognized among contemporary taxa. Non-protosphagnalean Permian mosses were also fairly diverse, representing morphotypes comparable with Dicranidae and acrocarpous Bryidae, although unequivocal representatives of these subclasses are known only since Cretaceous and Jurassic. Even though Sphagnales is one of two oldest lineages separated from the main trunk of moss phylogenetic tree, it appears in fossil state regularly only since Late Cretaceous, ca.
    [Show full text]
  • An Overview of Family Pottiaceae (Bryopsida) in Central India with Special Reference to Pachmarhi Biosphere Reserve (PBR)
    Lindbergia 34: 30–39, 2011 ISSN 0105-0761 Accepted 11 January 2011 An overview of family Pottiaceae (Bryopsida) in central India with special reference to Pachmarhi Biosphere Reserve (PBR) Virendra Nath, A. K. Asthana and Reesa Gupta V. Nath, A. K. Asthana ([email protected]) and R. Gupta, Bryology Laboratory, National Botanical Research Inst., 226 001 Lucknow, India. The family Pottiaceae is one of the dominant acrocarpous moss family of India, with approximately 26 genera and 130 species being encountered. The present work intends to take up distribution of the members of the family in the central Indian region with nearly 22 taxa being reported earlier. Achanakmar – Amarkantak Biosphere Reserve, Bhimbetka world heritage site, Gujarat, Mt. Abu and Pachmarhi Biosphere Reserve (PBR) are some important hotspots of bryophyte vegetation in central India. Among these, PBR listed under UNESCO’s Man and Biosphere (MAB) Programme is an important site of bryophyte vegetation. Mosses of PBR however have not been studied until now. Here we assess the status of family Pottiaceae in central Indian zone along with providing an illustrated account of eight species encountered from PBR. Among these, Hyophila spathulata (Harv.) Jaeg. and Oxystegus teneuirostre (Hook. et Tayl.) A.J.E. Smith are new additions to central India, while Semibarbula ranuii Gangulee and Weissia edentula Mitt. are being reported from PBR for the first time. India is well known for its floristic diversity of various plant betka world heritage site (Nath and Bansal 2009), 11 taxa groups. Indian researchers have broadly divided India into from Mt. Abu (Chaudhary and Deora 2001) and eight eight bryo-geographical regions (Singh 1997).
    [Show full text]
  • Liverworts, Mosses and Hornworts of Afghanistan - Our Present Knowledge
    ISSN 2336-3193 Acta Mus. Siles. Sci. Natur., 68: 11-24, 2019 DOI: 10.2478/cszma-2019-0002 Published: online 1 July 2019, print July 2019 Liverworts, mosses and hornworts of Afghanistan - our present knowledge Harald Kürschner & Wolfgang Frey Liverworts, mosses and hornworts of Afghanistan ‒ our present knowledge. – Acta Mus. Siles. Sci. Natur., 68: 11-24, 2019. Abstract: A new bryophyte checklist for Afghanistan is presented, including all published records since the beginning of collection activities in 1839 ‒1840 by W. Griffith till present. Considering several unidentified collections in various herbaria, 23 new records for Afghanistan together with the collection data can be added to the flora. Beside a new genus, Asterella , the new records include Amblystegium serpens var. serpens, Brachythecium erythrorrhizon, Bryum dichotomum, B. elwendicum, B. pallens, B. weigelii, Dichodontium palustre, Didymodon luridus, D. tectorum, Distichium inclinatum, Entosthodon muhlenbergii, Hygroamblystegium fluviatile subsp. fluviatile, Oncophorus virens, Orthotrichum rupestre var. sturmii, Pogonatum urnigerum, Pseudocrossidium revolutum, Pterygoneurum ovatum, Schistidium rivulare, Syntrichia handelii, Tortella inflexa, T. tortuosa, and Tortula muralis subsp. obtusifolia . Therewith the number of species increase to 24 liverworts, 246 mosses and one hornwort. In addition, a historical overview of the country's exploration and a full biogeography of Afghan bryophytes is given. Key words: Bryophytes, checklist, flora, phytodiversity. Introduction Recording, documentation, identification and classification of organisms is a primary tool and essential step in plant sciences and ecology to obtain detailed knowledge on the flora of a country. In many countries, such as Afghanistan, however, our knowledge on plant diversity, function, interactions of species and number of species in ecosystems is very limited and far from being complete.
    [Show full text]
  • Evolutionary Leverage of Dissilient Genera of Pleuroweisieae (Pottiaceae) Evaluated with Shannon-Turing Analysis
    Hattoria 12: 9–25. 2021 Evolutionary leverage of dissilient genera of Pleuroweisieae (Pottiaceae) evaluated with Shannon-Turing analysis Richard H. ZANDER Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63116, U.S.A. Author for correspondence: Richard H. ZANDER, [email protected] Abstract Evolutionary leverage is the average of total transformational trait changes among species in a dissilient (radiative) genus or larger, integral taxonomic group. It is proposed as a measure of evolutionary success in modern environments. Comparing evolutionary leverage aids in measuring potential evolutionary coherency and ecological success among taxonomic groups, here exemplified with Pleuroweisieae and Barbuloideae of Pottiaceae and genera of Streptotrichaceae (all Bryophyta). Shannon-Turing analysis—calculation using summed informational bits and sequential Bayes— translates to relative support for order and coherence of species in dissilient (radiative) genera. Hymenostylium gracillimum (Pottiaceae) is transferred to Ardeuma, while H. xerophilum and H. hildebrandtii (Merceyopsis hymenostylioides as synonym) are transferred to Eobryum. These three possess the primitive trait of a stem central strand in Pleuroweisieae. The genus Ozobryum is resurrected from synonymy with Molendoa. Introduction “Evolutionary leverage” is a term used for suites of adaptations that presumably allow better competition against other species, e.g. angiosperm success over competing groups (Brodribb & Feild 2010). Evolutionary leverage is more exactly defined here as a comparison of the average number of trait transformations among newly generated species in a dissilient genus as a closed causal group (Zander 2018: 36). A dissilient genus is one characterized by a progenitor species with apparent descendant species radiating from it without reversals in key traits (reversed from those in an outgroup or the inferred progenitor of a progenitor) or at least very few reversals.
    [Show full text]
  • 2447 Introductions V3.Indd
    BRYOATT Attributes of British and Irish Mosses, Liverworts and Hornworts With Information on Native Status, Size, Life Form, Life History, Geography and Habitat M O Hill, C D Preston, S D S Bosanquet & D B Roy NERC Centre for Ecology and Hydrology and Countryside Council for Wales 2007 © NERC Copyright 2007 Designed by Paul Westley, Norwich Printed by The Saxon Print Group, Norwich ISBN 978-1-85531-236-4 The Centre of Ecology and Hydrology (CEH) is one of the Centres and Surveys of the Natural Environment Research Council (NERC). Established in 1994, CEH is a multi-disciplinary environmental research organisation. The Biological Records Centre (BRC) is operated by CEH, and currently based at CEH Monks Wood. BRC is jointly funded by CEH and the Joint Nature Conservation Committee (www.jncc/gov.uk), the latter acting on behalf of the statutory conservation agencies in England, Scotland, Wales and Northern Ireland. CEH and JNCC support BRC as an important component of the National Biodiversity Network. BRC seeks to help naturalists and research biologists to co-ordinate their efforts in studying the occurrence of plants and animals in Britain and Ireland, and to make the results of these studies available to others. For further information, visit www.ceh.ac.uk Cover photograph: Bryophyte-dominated vegetation by a late-lying snow patch at Garbh Uisge Beag, Ben Macdui, July 2007 (courtesy of Gordon Rothero). Published by Centre for Ecology and Hydrology, Monks Wood, Abbots Ripton, Huntingdon, Cambridgeshire, PE28 2LS. Copies can be ordered by writing to the above address until Spring 2008; thereafter consult www.ceh.ac.uk Contents Introduction .
    [Show full text]
  • A Miniature World in Decline: European Red List of Mosses, Liverworts and Hornworts
    A miniature world in decline European Red List of Mosses, Liverworts and Hornworts Nick Hodgetts, Marta Cálix, Eve Englefield, Nicholas Fettes, Mariana García Criado, Lea Patin, Ana Nieto, Ariel Bergamini, Irene Bisang, Elvira Baisheva, Patrizia Campisi, Annalena Cogoni, Tomas Hallingbäck, Nadya Konstantinova, Neil Lockhart, Marko Sabovljevic, Norbert Schnyder, Christian Schröck, Cecilia Sérgio, Manuela Sim Sim, Jan Vrba, Catarina C. Ferreira, Olga Afonina, Tom Blockeel, Hans Blom, Steffen Caspari, Rosalina Gabriel, César Garcia, Ricardo Garilleti, Juana González Mancebo, Irina Goldberg, Lars Hedenäs, David Holyoak, Vincent Hugonnot, Sanna Huttunen, Mikhail Ignatov, Elena Ignatova, Marta Infante, Riikka Juutinen, Thomas Kiebacher, Heribert Köckinger, Jan Kučera, Niklas Lönnell, Michael Lüth, Anabela Martins, Oleg Maslovsky, Beáta Papp, Ron Porley, Gordon Rothero, Lars Söderström, Sorin Ştefǎnuţ, Kimmo Syrjänen, Alain Untereiner, Jiri Váňa Ɨ, Alain Vanderpoorten, Kai Vellak, Michele Aleffi, Jeff Bates, Neil Bell, Monserrat Brugués, Nils Cronberg, Jo Denyer, Jeff Duckett, H.J. During, Johannes Enroth, Vladimir Fedosov, Kjell-Ivar Flatberg, Anna Ganeva, Piotr Gorski, Urban Gunnarsson, Kristian Hassel, Helena Hespanhol, Mark Hill, Rory Hodd, Kristofer Hylander, Nele Ingerpuu, Sanna Laaka-Lindberg, Francisco Lara, Vicente Mazimpaka, Anna Mežaka, Frank Müller, Jose David Orgaz, Jairo Patiño, Sharon Pilkington, Felisa Puche, Rosa M. Ros, Fred Rumsey, J.G. Segarra-Moragues, Ana Seneca, Adam Stebel, Risto Virtanen, Henrik Weibull, Jo Wilbraham and Jan Żarnowiec About IUCN Created in 1948, IUCN has evolved into the world’s largest and most diverse environmental network. It harnesses the experience, resources and reach of its more than 1,300 Member organisations and the input of over 10,000 experts. IUCN is the global authority on the status of the natural world and the measures needed to safeguard it.
    [Show full text]
  • A Revised Checklist of Hawaiian Mosses
    Tropical Bryology 25: 35-69, 2004 A revised checklist of Hawaiian mosses G. W. Staples, C. T. Imada, W. J. Hoe, and C. W. Smith Correction for Page 38: Hawaiian Moss Flora in Summary Endemic Indigenous Alien Total Families 0 41 1 42 Genera 2 126 7 135 Species 75 166 14 255 Total named taxa 90 169 14 273 Correction for Page 39: Frahm et al. 2000, cited under Amphidium tortuosum, should correctly refer to the following: Frahm, J.-P., T. Klöcker, R. Schmidt, and C. Schöter. 2000. Revision der Gattung Amphidium (Musci, Dicranaceae). Tropical Bryology 18: 173-184. Revised checklist of Hawaiian mosses 35 Tropical Bryology 25: 35-69, 2004 A revised checklist of Hawaiian mosses1 G. W. StaplesA, C. T. ImadaA, W. J. Hoe,B and C. W. SmithC A – Hawaii Biological Survey, Bishop Museum, 1525 Bernice St., Honolulu, Hawaii 96817 U.S.A. B – deceased; C – Department of Botany, The Natural History Museum, Cromwell Road, London, SW7 5BD, U.K. Abstract. A revised and updated literature-based checklist of Hawaiian mosses is presented. Geographic coverage includes the eight main Hawaiian Islands; the Northwestern Hawaiian Islands are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography of supporting references is included. Introduction cryptogamic plants—the mosses. Also in The Hawaii Biological Survey (HBS) was preparation by HBS staff members are checklists established as a program of Bishop Museum by and bibliographies for the Hawaiian anthocerotes the Hawaii State Legislature in 1992, specifically (hornworts) and hepatics (liverworts).
    [Show full text]
  • Botanice Est Scientia Naturalis Quae Vegetabilium Cognitiorem Tradit. — Linnaeus Page 2
    Number 55 August 15, 2011 A Newsletter for the flora MOSSES OF NEW MEXICO of New Mexico, from the Range Science Herbarium and Kelly W. Allred Cooperative Extension Service, College of Range Science Herbarium, Department of Animal & Range Sciences New Mexico State University, Las Cruces, NM 88001 [email protected] Agricultural, Consumer, and Environmental Sciences, New Total familes = 38 Mexico State University. Total genera = 130 Total species = 302 Total taxa = 312 Family AMBLYSTEGIACEAE Amblystegium fluviatile (Hedwig) Bruch, Schimper, & Gumbel = Hygroamblystegium varium (Hedwig) Mönkemeyer Amblystegium humile (Beauvois) Bruch, Schimper, & Gumbel = Hygroamblystegium varium (Hedwig) Mönkemeyer Amblystegium juratzkanum = Amblystegium serpens (Hedwig) Bruch & Schimper var. juratzkanum (Schimper) Rau & In This Issue — Hervey Amblystegium noterophilum (Sullivant & Lesquereux) Holzinger = Hygroamblystegium varium (Hedwig) Mönkemeyer New Mexico Mosses .. 1 Amblystegium riparium (Hedwig) Schimper = Leptodictyum riparium (Hedwig) Warnstorf Amblystegium serpens (Hedwig) Bruch & Schimper var. juratzkanum (Schimper) Rau & Hervey = Amblystegium Plant Distribution serpens (Hedwig) Bruch & Schimper Reports ..................... 11 Amblystegium serpens (Hedwig) Bruch & Schimper [Bartram 1931; Mahler 1978] Amblystegium juratzkanum Schimper Amblystegium serpens (Hedwig) Bruch & Schimper var. juratzkanum (Schimper) Rau & Hervey Amblystegium tenax (Hedwig) C. Jensen = Hygroamblystegium varium (Hedwig) Mönkemeyer Amblystegium varium (Hedwig) Lindberg =
    [Show full text]
  • ANOECTANGIUM THOMSONII MITT. (POTTIACEAE, BRYOPHYTA) from UTTARAKHAND, INDIA 111 Geophytology 43(2): 111-116 November 2013 Anoectangium Thomsonii Mitt
    ISSN 0376-5561 ANOECTANGIUM THOMSONII MITT. (POTTIACEAE, BRYOPHYTA) FROM UTTARAKHAND, INDIA 111 Geophytology 43(2): 111-116 November 2013 Anoectangium thomsonii Mitt. (Pottiaceae, Bryophyta) from Uttarakhand, India Dinesh K. Saxena1, Dheeraj Gahtori1 and Richard H. Zander2 1Department of Botany, Bareilly College, Bareilly-243001, India 2Missouri Botanical Garden, P.O. Box 299, St. Louis, MO 63166, U.S.A. E-mail: [email protected]; [email protected]; [email protected] Manuscript received: 16 May 2013 Accepted for publication: 03 August 2013 ABSTRACT Saxena D. K., Gahtori D. & Zander R. H. 2013. Anoectangium thomsonii Mitt. (Pottiaceae, Bryophyta) from Uttarakhand, India. Geophytology 43(2): 111-116. The moss Anoectangium thomsonii Mitt. is widely distributed in almost all the bryo-geographical regions of the Kumaon and Garhwal regions of Uttarakhand state of India, where it grows luxuriantly especially on calcareous rocks and dry and arid places. Although this species has earlier been listed from this region, the present study is rst attempt for a GPS-based distribution and regional taxonomic description of the species. Key-words: Anoectangium thomsonii Mitt., Pottiaceae, moss, taxonomy, GPS based distribution, Uttarakhand, India. INTRODUCTION it amenable to rich plant diversity. The moss The large area and variety of phyto-climatic was collected from calcareous, rocky, dry and conditions within different biogeographical arid places. Plants were carefully separated out, zones contribute to the great and rich diversity and gametophytes appeared to be Anoectangium of the Indian ora. Bryophytes, like other plant thomsonii Mitt. of the family Pottiaceae. Only groups, are an important component of any forest scattered information is available for Anoectangium ecosystem (Carleton & Maycock 1981, Glime thomsonii in oristic lists (Barukial 2011).
    [Show full text]
  • The Pottiaceae S. Str. As an Evolutionary Lazarus Taxon
    J Hattori Bot. Lab. No. 100: 581-602 (Aug. 2006) THE POTTIACEAE S. STR. AS AN EVOLUTIONARY LAZARUS TAXON RICHARD H. ZANDER I ABSTRACT. Genomic studies have shown that evolution can be based on clusters of genes that may be silenced and reactivated much later. Thus, apparent phylogenetic homoplasy may sometimes rep­ resent a one-time developmental response to selection, involving many genes or gene clusters and one or more regulators, and as such may be called homoiology. Evidence for such exaptions (pre­ adaptions) in many organisms is cited. Distinctive features of the Pottiaceae (Bryopsida), the complex gametophytic morphology characterized by the pleisiomorphic Timmiella and the twisted peristome of 32 filamentous, spiculose teeth distal to a basal membrane are suggested to be likewise homoiolo­ gous. The characteristic peristome may have reappeared in the family four to seven times. Inasmuch as exaptions may skip across cladistic events tracked by non-coding or neutral gene traits, this paper re-evaluates past evolutionary schema in light of an apparent, theoretically acceptable exception to Dollo's Law that complex traits are never re-evolved. An evolutionary diagram based in part on in­ ferred process is offered in place of the usual c1adogram based on inferred events of genetic isolation. KEY WORDS: Exaption, Dolle's Law, evolution, homoiology, Pottiaceae, phylogenetics, Lazarus taxon. INTRODUCTION With a change during the 1700's and 1800's towards probabilistic thinking in science (Pap, 1962), philosophical or logical support for certainty or relative certainty about "truth" in science began to be replaced by a pragmatic attitude that science does not establish truth, but identifies theories that are so well supported by facts (well-corroborated observations) that they may be acted upon (used as a basis for additional research), while alternative the­ ories may be ignored even though some facts support them.
    [Show full text]
  • (Pottiaceae, Bryophyta) Based on Nrits Sequence Data
    54 (2) • May 2005: 361–368 Werner & al. • Phylogeny of Trichostomoideae Molecular phylogeny of Trichostomoideae (Pottiaceae, Bryophyta) based on nrITS sequence data Olaf Werner1, Rosa María Ros1 & Michael Grundmann2 1 Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus de Espinardo, 30100 Murcia, Spain. [email protected] (author for correspondence), [email protected] 2 Department of Botany, The Natural History Museum, Cromwell Road, London, SW7 5BD, U.K. m.grund- [email protected] Bayesian analysis of an alignment of 83 nrITS sequences belonging to 66 taxa of Pottiaceae revealed repre- sentatives of subfamily Trichostomoideae in a well supported monophyletic lineage. Two robust clades with- in Trichostomoideae include species of Tortella and Weissia, respectively. Eucladium verticillatum is part of Trichostomoideae, and Anoectangium, Gymnostomum, Hymenostylium, and Tuerckheimia form a robust basal clade. Affinities of Hyophila and Leptobarbula to Trichostomoideae are unclear. Within Trichostomoideae, Pleurochaete squarrosa is resolved in Tortella while Trichostomum tenuirostris, Pseudosymblepharis, and Chionoloma are placed as a separate genus or part of a broadly defined Tortella. The type species of Trichostomum, T. brachydontium, is nested in Weissia together with T. brittonianum, T. crispulum, and T. jamaicensis. Trichostomum unguiculatum and W. ayresii are resolved as closely related and separate from the rest of the Weissia clade. The genus Astomum, erected for cleistocarpous species of Weissia, is also nested with- in Weissia. Genetic divergence between selected accessions of Weissia controversa and Astomum is sometimes smaller than between accessions of Weissia controversa. KEYWORDS: bryophyta, molecular phylogeny, nrITS sequence data, Pottiaceae, Trichostomoideae. Beauv., Dialytrichia (Schimp.) Limpr., Ephemerum INTRODUCTION Hampe, Kingiobryum H.
    [Show full text]