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MICROCOPY RESOLUTION TES, CHART MICROCOPY RESOLUTION TEST CHART NATIONAL BUREAU Of SlANDARDS-1963·A NATIONAL BUREAU Of STANDARDS-1963-A NEARCTIC SAWFLIES II. Selandriinae: Adults _ (Hymenoptera: . Tenthredinidae)_
By DAVID R. SMITH
Technical Bulletin No. 1398
Agricultural Research Service U.S. DEPARTMENT OF AGRICULTURE
Washington, D.C. Issued September 1969
For sale by the SuperlnitenOOIlJ!: of Documents, U.S. Government Printing Office Washington, D.C'. 20402 - Price 40 cents Contents Page Systematic arrangement______3 Key to Selanc.iriinae genera ______.______5 Tribe Strungylogasterini ______6 Genus Eriocampidea Ashmead______6 Genus Hemitaxonus Ashmead______7 Genus Slrongylogasler Dal:.lboffi______------______12 Tribe AneugmeninL ______. ______25 Genus Aneugmenus Hartig______25 Genus Birka Malaise______28 Genus Eustromboceros Rohwer______30 Genu!:! Nesoselandria RohweL______31 Genus Slromboceridea Rohwer______33 Tribe SelandriinL ______34 Genus Brachylhops Haliday______34 Tribe AdelestinL ______37 Genus Adelesla Ross______37 Tribe HeptamelinL______38 Genus Heptamelus Haliday______38 Literature cited______40 Indcx______47 IV NEARCTIC SAWFLIES II. Selandriinae: Adults (Hymenoptera: Tenthredinidae)
By D.WID R. S~UTH, cntomologist, S'ystemaf,;o Entomology Laboratory, Agricultural Researoh Servioe Tho subfamily Selandriinae includes 2'i spedes in 11 genera and five tribes in North America. It is poorly represented in the Ne.'Ll'ctic ,'~egion, but it is one of the dominant sawfly groups in the Tr~pics, especially Central America, South America, and eastern Asia. Members of the Selandriinae m!!.y be of sporadic economic impor tance to agriculture. All species except He]Jtamel~/'8 ochr'.'le~f,(JU8 (Stephens) are external feeders in the 1arval stage on various ferns, sedges, and grasses. They are especially 11. nuisance in ornamental and landscape plantings, ,yhere their defoliation of the host can be very unsightly. They may also be destructive to forage gras.o:;es. Some may be beneficial by destroying ferns, 'which are considered weeds, or by killing bracken, a. plant toxic to livestock (Beer, 1955).1 The subfamily as treated here is as proposed by Ross (1951) with the addition of the tribe Heptamelini, which has been found only recently in North America. Ross' 1951 classification is similar to that he proposed in 1937 except for reducing the Adelestinae to tribal rank. Benson's (195fJ) concept of the Se1andriinae is slightly broader and includes the Dolerinae as a tribe. - The subfamily Selandriinae includes those sawflies of the Tenthre dinidae that either lack or possess an anal crossve5.n in the forewing and have vein Rs+i11 cUTTed. They may be separated from the Dol erinae by the presence of vein 2t'-m in the forewing. Other winO" vena tion characters that may ~)e used to separate some of the Selandriinae from other subfamilies are veins M and m-C1b of the forewing being parallel, except in Brachytho]Js and HeptameZ~/..8, and vein ill meeting Rs+nJ at or near the junction of Rr+ R. (In Adele8ta and seve.ral other genera vein 111 meets S c +R before R8 +111.) The larvae are not treated in this bulletin. Very few are lrnown or have been associated with adults. Dyar (1894, 1895, 1897) was the first to describe several selandriine larvae. Yuasa (19fJfJ) included descriptions of several genera and species under two subfamnies, the Emphytinae and Selandriinae. Beer (1955) gave detailed descrip tions of the biology of two western species of Strongylogaster and described their larvae. The most suitable generic key is that by Lorenz and Kraus (1957), and it is adequate for determining some Nearctic genera.
1 'rh~ year in italic aft~'r thp author'!' name is the key to the reference in Literature Cited, p. 40. 1 2 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE
The tribal arrangement for the Selandriinae is still slightly arti ficial, although wit.h the information currently ava.ilable it does seem to express relationships to some degree. Benson (1938,1952), Ta.keu chi (1.9.4-1), and Ross (1951) reported [1, tribal classification for this subfamily. The five tribes p:'esented here appear to be t:lhe best ttrrange ment and haye been based primarily on adult characters. The most unique t.ribes are the Adelestini -\lnd Heptamelini, eaeh represented by ,"ery few species and separated from each other and fl'om other tribes by deeided morphological differences. The other tribes, the Strongy logasterini, Anellgmenini, and Selandriini, appear to be closely re lated H..rod their degree of separation is not so distinct.; however, ~fax well (1955) supported the separation of these three tribes basec1 on the structure of the salivary glands of the larvae. This triba.l arrange ment will have to be evaluated again when more larVtle are a.vailable for study and the Neot,ropical and Ol'iental species aTe better known. The five tribes ttre defined as follows: Strongylogasterini.-Benson (1988) erected several tribes, but simJ2ly shtted that t.his tribe included those species not in the ot·hers unhl t.hey could. be furt.her divided. In general, this is still the situa tion. All members of this tribe are n5..sociated with ferns. The morpho logiettl characters are highly nwia.ble, but ft11 members are ra.ther long, slender insects with the costa not distinctly swollen at its apex, t.he anal ('Toss,"ein of the fore;wing present or absent, the prepectns usually IHtrrow and differentiated by a furrow (in some Hcmita;c01H1S and in E1'iorampidea the prepectusis wide and separated by a suture), the postgenal ("arina present (ex('ept in E?>iorampidea) , and the antennae long and slender (ex("ept in E1'io('([7npidca). Genera.: E1'io('wnpidea. H emHa;'f.'onu.~, 8f'rongy7oga,~tf'l'. Aneugmenini.-Members of this tribe are associated \yith ferns and graSS('s. Charaeters include the absence of the anal ("ro5..')\'ein in the forewing, t.he prepectus differentiated by a furrow, t.he cost.al vein swollen at its apex, the. presence or absence of a postgenal carina, and the short, stoeky antennae. Genera: A17cu,qmcnu8. Birl.'a, E11.~t?'01n OOCeI'08, N ('.w8eland1in. 8h'omooocridea. Selandriini.-This tribe is similar to t.he AneugmeninL but it in cludes those spc('ies in ",hi("h the mesepisternum is divided by an invertt'd V-shal:>€cl "11ture, th<.' ])ostgenal carintt is absent, veins !If and 1m.-C1/.. of the forewing are divergent, and vein 11'-111.. of the hind wing arises from vein 8c+ R instead of from vein Rs. The species are associated with sedges. Genus: B1'a('hytlwps. (The genusSe7rr.1L d'ria is Palaearctir.) Adelestini.-This tribe includes one monot.ypic genus. It is sepa rated by the subserrate antenna, the presence of an anal crossvein, the long distal anal cell and constriction of the basa1 anal ceU of the forewing, the absence of a prepectus, and the indistinctly defined post tergite. The host is unknown. Genus: Ad.e[csta. Heptamelini.-The larvae. of the g-enus He7)tamel1l8 bore in the stems of ferns. In the adult the antennn, is short with seven or eight segments, the prepootus is absent, the a.nal crossvein of the forewing is present, the basa.l section of 9A and 3A is faint., veins J[ and 11n-cu of the forewing n,re markedly cli,-ergent, and veins L1l and Olb-a of the forewing a,'e interstitin,l. Genus:Heptmnel,u.<;. NEARCTIC SAWFLIES II. SELANDRIINAE 3 According to Ross (1937) and as accepted by Maxwell (1955), members of the Selandriinae are the most generalized of the Tenthre dinidae, and representatives of the Strongylogasterini most closely resemble the hypdLhetical 2.ncestor of the tenthredinids. From avail able information it is difficult to support or disprove this hypothesis. If we consider those characters usually accepted as most generalized in the Tenthredinidae, members of the Strongylogasterini must be re garded as the most primitive, especially some Hemitawonu8 species that possess the anal crossvein in the forewing and have a well-dif ferentiated prepecLl1s. The species lacking the anal crossvein and with the prepectus present as a raised sholilder, as in most Strongylogaster, are probably more specialized and likely gave rise to the Aneugmenini and Selandriini. The latter two tribes also include members that feed on grasses and sedges rather than ferns !md thus represent a host divergence. The Adelestini. artd Heptamelini may be considered as highly advanced. owing to their more specialized morphological characters, although both tribes have retained the anal crossvein in the forewing.
Systematic Arrangement Family TENTHREDINIDAE Subfamily SELANDRIINAE Tribe STRONGYLOGASTERINI Genus E1'iocampidea Ashmead
(1) Erioca1npidea arizonen8is Ashmead; Adzona, New Mexico.
Genus Hcmitawonws Ashmead
(2) Hemtitaxoml8 aZbidopicfAl8 (Norton); Alabama, Connecticut, District of Columbia, Illinois, Indiana, Iowa, Kansas, Maine, Mary land, Massachusetts, Michigan, :Minnesota, ~fissouri, New Brunswick, Xew Hampshire, New ,Jersey, New York, Nova Scotia, Ohi.o, Ontario, Pennsylvania, Quebec, Virginia, 'Wisconsin; on Onoclect and OS1Tllunda. (3) Hmnitaxonu8 d'l.(bUat~ls (Norton); Connecticut, District of Columbia, 111inois, Indiana, Maine, Maryland, Massachusetts, Michi gan, Minnesota, New Brunswick, New Hampshire, New .Jersey, New York, North Carolina, Nova Scotia, Ohio, Ontado, Pennsylvania, Quebec, Texas, Virginia, 'Visconsin; on Onoclca. and OS1n1l11da. (4) Hemitax0n1l8 1lHllticinct1l8 Hall; Maine, Maryland, Ohio, On tario; on Athyerilum. (5) Hemitawon~l,~ pri?na7'iu8 Smith; British Columbia, California, Oregon. Genns Strongyloga.'1te1' Dahlbom
(6) St?'ongylo,qaBter di.~tan.'1 Norton; BrHish Columbia, California, Colorado, Idaho, Nevada, Oregon, Utah, 'Vashington; on Pteridium. 4 'l'ECENICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE
(7) Str'ongyl'()ga,'fter iwpressata, Provancher; Colorado, Connecti cut, Illinois, Iowa, 'Maine, Massachusetts, New Bnmswick, New Mexico, New York, North Carolina, Nova Scotia, Ontario, Pennsylvania, Quebec, Tennessee, Wisconsin. (8) Stl'on gyloga8ter 1nflC'ltla, (KIug) ; British Colllm1)in, Ontll,Tio; Palae!trctic. (9) StrongyZogaBte'r 1111llticinrt{( Norton; Florida, Maine, Mary land, ~{assachusntts, Michig'lllll Minnesota, New Brunsw;kk, New Hampshire. New .Tersey, New Y()rk, North Oarolina, Nova Scotia, Ontario, Pennsylvania, Quebec, V('rmollt, Vir~il1ia; on Ptr'1'icliwn. (10) StrongylogaJ3ter polita C'rescion; Connecticut, New York, Ontario; on Pteridi1tm.. (11) 8t1'01l.gylo.q{(.~te:r 'remota Rohwer: Pennsylvania, Quebec. (12) Strongyloga.'ft£'I' I'llfigastm (Kincaid); Alaska, British Columbia. (13) St1'01tgyloga8te1' 81,.. 'a:ulatipes Cresson; Illinois, Maine, Mary land, Massachusetts, Minnesota., New Brunswick, New York, Ontario, Pennsylvania, Quebec; on Pteridiwn. (14) Str01l.gyloga8ter' taf"ita (Norton); Alabama, Oonnecticut, Georgia, Illinois, Maine, Mitl'ylnnc1, Massachusetts, Michigan, New Brunswick, New Hampshire, New .T(,l'sey, New York, Nort'h Carolina, Ontario, Pennsylvania, Quebec, South Carolina, Vermont, Virginia, Wisconsin. (15) ,Rtrongylo,qasff')' tibiali8 Cresson; British Columbia, Califor ni11, Ida.ho, Oregon, 'Vashington; on Ptf'I'idhl1n. (16) Strongylogaste1' tllbe1'Guliceps Rohwer; Florida.
Tribe ANEUGMENINI
Genus Aneugme111i-~ Hartig
(17) Ane1tgmen11,~ jlavipe8 (Norton) ; Colorado, Connecticut, Flor i(ht, Georgia, Illinois, Maine, Manitoba, Maryland, Mas!'iachusetts, Miehign.n, Minn'l!sota, New' Brunswick, Newfoundland, New Hamp shire, New .Tersey, New York, North CfLl"Olina, Nova, Scot.ia., Ohio, Ontario, Pennsylvania, Quebec, Vermont, Virginia, "Test. Virginia, 'Wisconsin; on Ptf'ridimn. (18) Ane1t.qmen1l8 7)adi (Linnaeus); British Columbia, Oregon, 'Washington; Pnlaearctic; on Pteridium. (19) Aneu,gmen1t8 sC'lttellat11.Y, new species; Arizona.
Genus Birka Malaise
(20) BirkCL n01'dica, new species; Alaska, Alberta, British Columbia, Manitoba, Yukon Territory.
Genus E11stl'Omboce1'08 Rohwel'
(21) E'U8tr'omboce1'08 1'llfoca'U Genus Str01nboceridea Rohwer (23) Stromb.oce'ridea nigricans (Norton); Arizon&, Mexico. Tribe SELAN"DRIINI Genus Brachythops Haliday (24) Brachythops flavens (KIug) ; Alaska, Albertit, Colorado, Man itoba, Newfoundland (Labrador), New York, Northwest Territories, Quebec, Saskatchewan, 'Wisconsin; Palaearctic; on Oarex. (25) B'rachythops l..mtestneii (Konow) ; Alaska, Alberta, Manitoba, Northwest Territories, Quebec, Yukon Territory; Palaearctic; on Oarex. Tri'be ADELESTINI Genus AdeZesta Ross (26) Adelesta nova (Norton) ; New Brunswick, Nova Scotia, Que bec. Tribe HEPTAMELINI Genus Heptamelu.~ Haliday (27) Heptamehl8 ochroleuclls (Stephens) ; British Columbia; Palae arctic; larva bo res in stems offerns. Key to Selandriinae Genera 1. Forewing with anal crossvein present (pI. 1,2)______.______~ Forewing with anal crossvein absent (pI. 1,1)______6 2. Antenna seven- or eight-segmented (pI. III, 30); forewing with 2A and 3A basal to anal crossvein faint (pI. II, 4); prepectus absent; tarsal claw with long, slender inner tooth (pI. III, 12) ______Heptamelu8 Antenna nine-segmented; forewing with 2A and 3A distinct for its en tire length (pI. I, 2; pI. II, 5); prepectus present or absent; tarsal claw simple or with small inner tooth (pI. III, 6, 9)______3 3. Prepectus absent; forewing with proximal anal cell as long as distal anal cell (pI. II, 5); antenna slightly serrate (pI. III, 29); tarsal claw simple______Adelesta Prepectus present as either narrow raised shoulder or fiat. sclerite; fore wing with proximal anal cell twice as long as distal anal cell (pI. I, 2) ; antenna filiform (pI. III, 20-22) j tarsal claw simple or with small inner tooth______4 4. Prepectus present as narrow raised shoulder, separated from mesepi sternum by furrow; second antennal segment wider than long (pI. III, 22) ______StrongylogWiter (pt.) Prepectlls present as flat sclerite, separated from mesepisternum by suture, or second antennal segment as long as wide (pI. III, 20, 21) __ 5 315-625 0-60-2 6 TECHNICAL BULLETIN 1398) U.S. DEPT. OF AGRICULTURE 5. Postgenal carina present; antenna long and slender with sixth segment more than twice as long as wide (pI. III, 21} ______Hemitaxonus Postgenal carina absent; antenna short with sixth segment less than twice as long as wide (pI. III, 20) ______Eriocampidea 6. Mesepisternum with inverted V-shaped suture, dividing sclerite into two sections (pI. lII, 19); vein lr-m of hindwing arising from vein Sc+ R (pI. II, 3); malar space wide, about twice diameter of front ocellus______Brachythops Mesepisternum without inverted V-shaped suture; vein lr-m of hind wing arising from vein Rs (pI. I, 1); malar space usually less than twice diameter of front ocellus______7 7. Prepectus present as narrow raised ~ Tribe STRONGYLOGASTERINI Genus ERIOCAMPIDEA Ashmead Eriocampidca Ashmead, 1898, p. 256; Rohwer, 1911a, p. 224; Rohwer, 1911b, p. 109; Malaise, 1933, p. 56·; Conde, 1934, p. 177; Ross, 1W7, p. 64; Ross, 1951, p.22. 'rypl': Eriocumpidea ari.::onel!sis Ashmead. Original designation. ('ocT.:crcllonis ~racGilIlvray, 1908, p. 365; Rohwer, 1911b, p. 109 (=Eriocampiclca Ashmead) ; ~Ialaise, 1933, p. 56. Type: aockerelwni.~ occidentali.~ MacGillivray. Original designation. Description.-Antenna short, second segment globular, as long as wide; third segment subequal in length to fOLrth segment; sixth seg ment less than twice as long as wide (pl. III, 20). Postgenal carina absent; clypeus slightly emnrginate, flat; malar space slightly wider than diameter of front ocellus. Prepectus .present as flat sclerite, sepa rated from mesepisternum by suture. Tarsal claw simple (pI. III, 6). Forewing wiMl anal crossvein present; proximal anal cell twice length of distal anal cell. Hindwing with annl cell petiolate (pI. I, 2). The sheath and lancet of the single species in this genus are re markably close to those of some species of Hemitamonus, but Eriocam pidea. may be separated by the lack of a genal carina and the shorter antenna. NEARCTIC SAWFLIES II. SELANDRIINAE 7 Description of Eriocampidea Species Eriocampidea, a?'izonen. Eriocampitleu arizonensis Ashmead, 1898, p, 256, S! ; Rohwer, 1911b, p. 109; Ross, 1007, p. 66; Ross, 1951, p. 2"2. ('ocl"crellonis oooidcntalis MaeGHlivray, 1908, p. 365; Rohwer, 1911b, p. 109 (=arizonensis Ashmead) ; Frison, 1927, p. 230. Female.-AverfLocre length, 5.5 mm. Antenna black. Hen.d black wit.h line on inner orbits, outer orbits, supraclypeal area, clypeus, labrum, and labial and maxillary palpi whitish. Thorax black 'with tegula and upper angles of pronoturri whitish. Legs black with apical third of each femur, each tibia, and forebasitarsus whitish. Abdomen black. 1Vings hyaline. Sheath slender, pointed at apex (pI. V, 59). Lancet long and slender with seven to nine teeth irregularly placed along apical fourth (pI. VII, 84). Male.-Unknown. Types.-Erlocampidea arizonens-ts ..Ashmead ( S!) is type No. 4136 at the U.S. National Museum with the data "Ariz." {!ockerel7.oni8 o('cidentali8 is at the Illinois Natural History Survey with the data "Ruidosa Creek, New Mexico, 6,600 feet elevation, on fronds of Pterls aquilina, .July 1, E. O. 1Yooton." The abdomen of occidentalis is miss ing and the sex is not known, but all other characters agree with this species. Distribution.-A'l'izona: Douglas, July 4, 1933, 1Y. W. Jones; Co chise Co., Rustlers Park, Chiricahua Mts., 8500', June 10, 1964, on hi~ mi.w)lwiensis Nutt., J. H. Puckle, M. A. Mortenson, M . .A. Cazier; Chiricahua Mts., '7-8-32, R. H. Beamer.2 Ne'l.o Mexico: Ruidosa. Host.-Unknown except for preceding adult collection records. Discussion.-This species has been 'taken only from southern Ari zona and New :Mexico at high elevations. Genus HEMITAXONUS Ashmead JJem.ita. .conIl8 Ashmead, 1808, p. 311; Konow, 1905, p. 102; Rohwer, 1911e, p. 397; l~nslin, 1914, I>. 206; MacGillivray, 1916, p. 46; ~Ialaise, 1931, p. 140; Malaise, 1933, I>. 00; Conde, 1934, p. 177; Ross, 1037, p. (14; Takeuehi, 1941, p. 24(;; Ross, 1951, p. 22; Smith, 1006a, p. 113. Type: Tuwonlls dllbitutlls Norton. Original designation. Epitaxonlls ~IaeGilliYray, 1908, p. 365; Rohwer, 1911e, p. 397 (=H('mitaa;olllls Ashmead) ; MaeGillivray, 1916, p. 46; Malaise, 1933, p. 56. 'l'ype: Tua:onli8 o:lbidopictll.~ Norton. Original designation. l'tahlbcrgia ForsiLls, 1010, p. 49; Enslin, 1914, p. 206 (=Hcmitawonus Ashmead). Type: Sahlbergi/.: .~trllthwpteri(H8 Forsius. ~Ionotypie. Description.-.Antelma long and slender; second segment globular, as long as wide; third segment sllbeqnal in length to fourth segment; sixth segment at least two times longer tluln wide (pI. III, 21). Post genal carina present; clypeus slightly emarginatc; malar space equal to or slightly wider than diameter of front ocellus. Pl'epectus present as Aat sclerite, separated fl'Oll1 mesepisternnm by suture, or present as , 'rhroughout this bulletin all information pertaining to d Key to Hemitaxonu8 Species 1. Female____ , ______.. ______2 J\fale______. ______Ii 2. S healh short, truncate at apex (pI. V, 64); pre'pectuB present as raised shoulder, separated from mesepisternum by furrow; abdomen black with central rufous band ______albidopiclus (Norton) Sheath long, pointed Ilt apex (pI. V, 65-67); prepectus present as flat sclerite, separated from mesepisternum by suture; abdomen uniformly light or dark______.. ______~______3 3. Sheath narrow at base, dorsal and ventral margins aubpara!lel (pI. V, 67); western species______primari7ls Smith Sheath broad at base, dorsal and ventral margins not subparallel (pI. V, 65, (6); eastern species.. ______4 4. P(!ctIlS nnd mesepisternum uniformly colored, either entirely black or entirely rufOllS______.clubilal7ls (Norton) Pectus black; mesepisternlllll with large trianglllar rufous arelL_rnulticinclus Ha1l :i. Prepectlls present as raised shoulder, sepnrated frolll mesepisternum by furrow. ______.. _.. ______allJidopiclus (Norton) PrepectllS present as flat llclerite, separated from mesepisternum by Sll til ro _, ______• _.. _.. __ . _.... __ .. ______.... _.. ______6 U. Western species .... ___ .. _. _... ______prirnariu8 Smith Eastern. species. __ .. _...... __ .. ______...... ______.. ______7 7. Abdomen entirelYi'lIfolis or ye1l0wish ______1nulticinclus Hall Abdomen blnck wit.h rufous arells on tcrgitm; 2-5 Ilnd sternites 1-u. _.. . _._... _. _____ . _. _. _____ . ______. _____ d1lbilalu8 (Norton) Descriptions of Hemitaxonus Species fiemita,1'01l/18 albidopic1718 (Norron) '1'1/:1:011-'18 (/llJirlo[Ji<~tll.~ ~orton, 1.8GR, p. 213, 3, ~ ; l'rovllncht'r, 1878. II. lOG: 1'1'0 ,'nr1('h('r, lRB~, p. 211;; Dnlla ~'orrt'. IBIH, I'. 110. J[cmit(/J·onll.~ IIl1Jir1olli('tll.<, Konow, l!)();;, II. l0'2; Hohwl'r, 10Ue, II. aOR; HOi-IS, 10:37, p. Gr;; Hoss, l!Y.il. p. 22; ~mith, 10(l(;n. p. 114. l~fliI(/.1'OIIlIH (lllJiriopict'I,<, i\fn('GiIlinny, 1\)08, p. 8(;;;; lfnrGiIlinny, 10](1, Jf. 4H. UEARCTIC SAWFLIES II. SELANDRIINAE 9 Ta:conU8 amiclls Norton, 1868, p. 213, ~, ~ ; Provuncher, 1878, p. 166; ProV'ancher, 1883, p. 215; Dalla Torre, 1894, p. 110; Konow, 1905, p. 102; MacGillivray, 1016, p. 47; Ross, 1951,1>. 22 (=albidopictu8 Norton). FIemita:COI!U8 rutopectu8 Rohwer, 191Oc, p. 204, ~ ; Rohwer, 1911c, p. 398 (=albi dopictll8 Norton). Female.-Average length, '7.0 mm. _Antenna and head black; labrum and clypeus white. Thorax black withtegula and upper angles of pro notum white, pectus and mesepisternum (except upper angle and pre pectus) rufous. T... egs orange yellow with apex of each coxa, each t~·o ('hanter, basal one-fourth of front and middle femora, :basal one-thIrd of hindtibia, and ba.c;al one-half of hindbasitarsus white, and extreme apex of hindfemur, apical two-thirds of hindtibia, and hindtarsus (ex cept basal one-half of basitarsus) black. Abdomen black with sternites 1 and 2 and segments 3 and 4- rufous. "Wings hyaline. Prepectus present as raised shoulder, separated from mesepisternum by furrow. Tarsal claw with minute inner tooth. Sheath ShOlt and truncate (p1. V, (i4). Lancet as in plate VII, 85. Male.-A\rerage length, 5.7 mm. 'Color similar to that of female except for abdomen, which is entirely black with rufous areas on ster nites 1-3. Structure as for female. Genitalia 'as in plate X, 126 and 127. Types.-Both of Norton's types are at the Academy of Natural Sciences of Philadelphia. T. aZbidolJictu8 (9, type No. 266) has the data "Virginia," and T. amicus (~, type No. 265) has -the data "Can ada.:' R .. mfopectu8 Rohwer (~) is U.S. National Museum type No. 12928 with the data "Nerepis, N.B., 22 Aug. [?], A. G. Leavitt, Col lector." The date is not legible. Distribution.-Enstern North America: Alabama, Connecticut,. Dis trid of Columbia, Illinois, Indiana, Iowa, Kansas, Maine, Maryland, ~fassac:husetts, Michigan, Minnesota, Missouri, New Brunswick, New Hampshire, New .Tersey, New York, Nova Scotia, Ohio, Ontario, Penn sylvaniu., Quebec, Virginia, 1Yiscom,in, Hosts.-On()den 8en.~ibili8 Ij. and 081nunda. Discussion.-This is the most distinct species in the genus and may be separated by the pl'epectus, female sheath, genitalia of both sexes, and coloration. lJemita,r01W8 rZubitat118 (Norton) 'I'UJ.·OIUI8 rlllbitutll,Y Xorton, 186211, p. 110. ~, ~; PrO\'ancher, 1878, p. 16;:;; Pro vancher, 188.'3, p. 21v; Dalla Torre, 1894, p. 110; Dyar, 1897, p. 20. }[emita:conll.~ Ilubitatu8, Ashmead, 1898, p. 311; Konow, 1005, p. 102; MacGilli vray, 1916, p. 46; Ross, 1951, p. 22; 1\Iaxwell, 1955, p. 48; Krombein, 1000, p. 30; Smith, 1006:1, p. 117. TaxonU8 albidopictu8 Dyar, nec Norton, 1897, p. 20. Hemitaxonll8 dubitatu8 var. amicu8 Rohwer, nec Norton, 1911c, p. 397. Female.-Average length, 7.2 mm. Antenna and head black; cly pens, labrum, and base of eaeh mandible white. Dorsum of thorax mostly rufous or mostly black, with aU intermediates; pectus and me sepisternul1l either entirely black or entirely rufous, not with contrast ing ('olol's. Legs ora.nge yellow with extreme apices of front and middle tibiae, enelt trochanter, and extreme base of hindtibia white, and base of hincIcoxa and hindtibia (except e:\'i:reme apex) black. Abdomen rufous with basal plates and anterior half of second tergite black; in fuseate areas sometimes on apical segments and sheath. Wings hyaline. 10 TECHNICAL BULLETIN 1398) U.S. DEPT. OF AGRICULTURE Prepectus present as flat sclerite, separated from mesepisternum by suture. Titrl'ial clll,w with smull inner tooth. Sheath with base broad, poiut.ed at apex (pI. V, 65), Lancet as in plate VII, 86. Male.-Average Jerl,.~th, 6.3 mm. Color as for female except for legs, which arc ('ntiI'ely orange yellow, and abdomen, whi('.h is b1aC'k with rufous al'eas 011 tergites 3-5 lind sternitC's 1-5. Genitalia as ill plate X, 128 itlld 12\). Tupes.-A specimen In,be.led as the type is in t.he Pea.body Musnelll of Na,tuml History, Yale TTnivel'sity. H. bears t.he data "Ct.187ii." Also, there is n,l1othm' specimen with the labels "Alotype" illlc1 "Mass, 1876." Although the localities on these specimens agree with those in Norton's deseL'iption, the dates areU~ and H years Itrt.er the description. Ilmve not seen these specimens and Wit..'> not nNe to loca;ie the type elsewhere, Distribution.-Eltstem North 1.\.meriClt: Connectieut, District, of Columbia, Illinois, Indiana, Maine, ~farY]ltnd, Massachusetts, Michi gan, Minnesota., New Brunswiek, New Hampshire, New .Tersey, New York, North Cal'olina, NO\'lt S(~tia, Ohio, Ont.ario, Pl'nnsylva,nia" Que heC', Texas, 'Virginia., 'Vis('ollsin. Hosts.-OnocZea. 8engibilis L., OSlIHl.nda, Pttl1'idiwn (?) (Kl'ombein 1.160) , Discussion.-This spceies Illily l)i! dist.in~nishedby coloration. Tho pcetus Itncl meseplsternmll aL'e always It uniform (~lor, whereas ill m'll7tirin('I1(.~ and In'imm'ill'~ they arc eonrtmsting colors. The abdomen of the !lIlLIe is most.ly black, exC'ept the rnfOlls spots as noted, whereas in 11wZti('inct'll8 !tnd prim(l}'i'll.~ the Itbdomenis mostly light III color. 1'11(', shcat,h of I'he femn.1e is broader t,llltU that. of eit.her 1miltin/nci1l8 or 7wim(ll'iu8. Itnd the genitalia. 01' both sexes ids<> difrer. The dorsum of the thomx 1'ltn~es fl'om !t1most, entirely black to entirely rufolls with \'arions degrees 01' lntennec1inltes, Dyn,r (18{J7) I'eltrcd t.his spec.ies from the l)l'e\rionsly ment,ioned hosts and described t.he la.n'lt under t.wo nltm(~s, du})it Tfcmitaa:O/l.II,~ Inlilticillctll.q Hall, 1017, p. 28, egg, lnrva.; Hoss, J,!):}l, II, 22; Smith, lOOGn. II. 118. Female.-Avemge 1enbrt,h , 7,0 Il1m, AntennlL and head bluek, clypens itIld labrllm white. Thot'ltx black with tegultt Itnd upper angles of pronotuIll whit'e and l:trge tl'ianl-,,,t1:u' rufous area on mesepisterllllIll; ped,lIs ancl mesepist.el"Imm with ('.()nr.rasting colors. Legs entirely ornnl-,l'() yellow; en,ch t~bia and tlWSIIS infuscn~e, Abdomen light. yellow to L'Ufolls with blt,sa,l phtt~, shen.th, and eerei bla,ck; ItI'(~as 011 Jatcl'Il.l NEARCTIC SAWFLIES II. SELANDRIINAE 11 margins and basal and apical tergites sometimes infuscate. Wings hya.line. Prepectus present as flat sclerite, sepa.rated from mesepisternum by suture. Tarsal claw simple. Sheath broad rut base and pointed at apex, somewhat narrower than that of dttbitat1t8 (pI. V, 66). Lancet as in plate VII, 87. Male.-A'Verage length, 6.1 mm. Color and structure as for female. Gen italia as inplate X, 130 and 131. Type.-A leCtotype was designa.ted by Smith (1966a). It is a hti'va, U.S. Na.tional Museum type No. 18701, and bea.rsthe data "·Wa.kemlUl, Ohio, ""V. B. Hall." . Distribution.-Maine~ Maryland, Ohio, Ontario. Host.-L1thyr-ium, thelypte1'ioides (Michx.) Desv. Discussion.-This species may be separated from dubitatttS by the contrasting colors of 'the pectus and mesepisternum, the entirely light c.olored abdomen in the male, the slightly narrower female sheath, and the genitaliiL of'both sexes. H differs from p1'ima1'i-U8 by the shape of the female sheath, but there are few reliable characters to separate the males from those of pr'imari118. This is not a commonly collected species. Hall (H)17), who inad \'ertently received credit for this species by describing the la.rva, reared adults from larvae feeding on ..tithyc1'iu1n tlwlyptcr'oldes in Ohio. The larvae were in tL bed of cultivated ferns. He stated that .the "eggs are attached on end to the upper side of the lea.i, often as many as ten or twelve on a frond." I colleoted severa.l specimens of this species in :~fn.ine by sweeping ferns, most of which 'were Onoclea. H. dubitat118 specimens were colleoted in the same area. I1emitamon1lB 7J1'i1na1iu,g Smith IIcmita.l'0nllb' primarill8 Smith, 1966u, p. 110, ~, Ij!. Female.-Average length, 7.2 mm. A.lltenna, black with first two segments light rufous or black; head black with c1ypeus, base of man dibles, and rest of mouth pa,l'ts white. Dorsum of thorax black or dark rufous with various amounts of each color; mesepisternum with large triangular rufous area,; In,ternl Plut of pronotum light rufous; tegula rufous; upper margins of pronotnm white. Legs entirely white to omnge yellow; each tarsus infuscate. Abdomen orange; sheath and cerci black. 'Wings llyn,line. Prepectus present as flat sclerite, separated from mescpisternum by suture. Tarsal claw simple. Sheath long and narrow, dorsal and ventral margins subparallel (pI. V, 67). Lancet as in plate VII, 88. Male.-Avemge length, 7.0 nun. Color similar to that of female; thora.x usually with dorsum black; abdomen usually darker. St.ructure as for female. Genitalia as in plate X, 132 and 133. Type.-Female, U.S. National Museum type No. 67983, with t.he data "Lagunitas, California, April 7, 1946, elevation 1000', H. P. Chandler." Distribution.-British Columbia., California, Oregon. Host.-Unknown. Discussion.-The long, narrow female sheath ..,,,ill separate the {ema.Io of this species from all ot.her llemitamonu.q females. The m:1.le 12 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE is easily separated from that of albidopict'll.S by the prepectus and light colored a,bdomen and from that of d1tbitat'll.S by the light abdomen; however, only slight genitalic characters will separate the male from that of mlutir:irwt'll8. This species is known from British Columbia to the Bay area in Califormu., and all the specimens examined were collected within 100 miles of the coast. It is the only known western species of this genus. Genus STRONGYLOGASTER Dahlbom Tenthredo subgenus Strongylogaster Dahlbom, 1835, p. 13; Hartig, 1837, p. 299. Strollgylogaster, Costa, 1859, p. 107; Thomson, 1870, p. 292; Andre, 1879, p. 410 ; Cresson, 1880b, p. 62; Cameron, 1882, p. 187; Cameron, 1883, p. 6; Konow, 1887, p. 26; Dalla Torre, 1894, p. 133; MaeGillivray, 1894, p. 325; Kono,,", 1005, p. 00; MacGillivray, 1908, p. 369; Euslin, 1914, p. 204; l\lacGillivruy, 11)16, p. 67; Enderlein, 1920, p. 3.'55; Yuasa, 1922, p. 53; Ross, 1937, p. 67; Takeuchi, 1941, p. 240 ; Berland, 1947, p. 203 ; Ross, 1951, p. 22; Zhelokhovtsev, 1951, p. 149; Benson, 1952, p. 58; Lorenz and Kraus, 1957, p. 44 ; Burks, 1958, p.ll; 'Malaise, 1983, p. 170. Type: Tenthredo cingulata Fabricius. Designated by MacGillivray, 1008. Thrilla:c Konow, 188.5a, p. 19; Dalla Torrt', 1894, p. 139; Konow, 1905, p. 9;:;; Mac Gillivray, 1008, p. 369; Enslin, 1913, p. 201; MacGillivray, 1916, p. 67; Yuasa, 1922, p. 53; Ross, 1937, p. 67 (=Strongylogaster Dahlbom) ; Takeuchi, 1941, p. 234; Berland, 1947, p. 205; ZhelokhovtseY, 1951, p. 146; Malah;e, 1968, p.170. Type: Strr:mgylogaster co-ntigu4 Konow. Designated by MacGillivray, 1908. Pscudota:ronus Costa, 1894, p. 157; Enslin, 1914, p. 205; Ross, 1937, p. 67 (=Strongyloga8ter Dahlbom) ; Takeuchi, 1941, p. 234; Berland, 1947, p. 210; l\[alaise, 1963, p. 187. Type: Tenthredo filicis Klug. l\Ionotypic. Polyatichophagus Ashmead, 1898, p. 310; Konow, 1905, p. 96 (=Strongyloga8tcr Dahlbom) ; Enslin, 1914, p. 205 (=Pseudota:cnus A. Costa). Type: TenthreiW filicis Klug. Original designation. Protota:co'nIlS Rohwer, 1910a, p. 49; MalaiRE', 1933, p. 56; Ross, 1937, p. 67 (=Strrmgyloga8tcr Dahlbom). Type; Prot()ta:co-nll.'J t-ypiC1l8 Rohwer. l\Ionotypic. Description.-Antenna filiform; second segment disklike, wider than long; third and fourth segments subequal in length; segments beyond third gradually decreasing in length (pI. III, 22). Postgenal earina present; clypeus very shallowly to deeply emargin!lJte; malar space na,rrower or wider than diameter of front ocellus. Prepect.us present as narrow raised shoulder, separated from mesepisternum by furrow. Tarsal claw simple or with small or large inner tooth; basal lobe absent (pl. III, 1,9, 10). Forewing with anal crossvein present or n:bsent. Hindwing with anal cell seesile or with short petiole. The differences within this genus, such as presence or absence of the anal crossvein in the forewing, size of the inner tooth of the tarsal claw, and degree of emargination of the clypeus, have led to the de scription of the several genera previously listed. Owing to lack of other differences, however, Ross (1937) grouped them together, and I see no reason to separate them on the basis of such characters that appear to obscure relationships and for which there are several inter mediates. Pseuriotaxo7lfU8, Polyst·iehophagu. Key to Strongylogaster Species 1. Fema~______2 1-fale ___ • _____ .. ______• ______.. ___ 13 2. Anal crossvein present in forewing______3 Anal crossvein absent in forcwing______4 3. Fcmora orange; eastcrn ______.. ______soriculatipes Cresson (pt.) Femora mOptiy black; wcstern______tibialis Cresson ,j. Sheath without scopa or dorsoapical projections,______~______5 Sheath with scopa or dorsoapical projections______i 5. Clypeus white; sheath truncate at apex (pI. IV, 38) __soriculatipcs Cresson (pt.) Clypells black; sheath rounded or pointed at apex (pI. IV, 42, 50)---- 6 6. Sheath broadly rounded at apex (pI. IV, 50); mesopleuron and mesona tum with dark rufous markings; abdomen mostly orange__ remota Rohwer Sheath narrow and pointed at apex (pI. IV, 42); mesothorax black; abdomen mostly black ______polita Cresson 7. Shcath slender, with small dorsoapical projections (pI. IV, 46-49) i anal cell of hindwing petiolute______8 Sheath broad, with smnll or large leaflikc scopa (pI. IV, 31-37, 44, 45); nnal cell of hindwing sessilc______9 (l15-6250-60--a 14 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE 8. Sheath______with narro,,", cylindrical dorsoapical projections (pi. IV, macula46, 47) (Klug) Sheath with broad, rounded, shorter dorsoapical projections (pi. IV, 48, 49) ______rufigastra (Kincaid) 9. Sheath with broad, large leaflike scopa (pi. IV, 31-34); clypeus deeply emarginate (pi. III, 13)______10 Sheath with small scopa (pi. IV, 35-37, 44, 45); clypeus shallowly cmarginate (pi. III, 14)______11 10. In postcrior view, wings of scopa widcly separatcd (pi. IV, 32) ; castern ______mlliticincta N ortoll In posterior vicw, wings of scopa closer together (pi. IV, 33); wcstern ______distans Norton 11. Wings hyali'.1e; head without large, coarse punctures__ impressata Provancher Wings dark;!y and uniformly infuscatc; hcad with numerous, large, coarse punctures______• ______12 12. Thorax usudly black; clypeus black; north of Florida ______lacita (Norton) Thora, usually rufous, with prothorax and tegula black; clypeus U!'".,.:\!y partly light; Florida______tllbercrdiceps Rohwer 13. Anal crossvein present in forewing______14 Anal crossvcin absent in forewing______15 14. Abdomcn entirely black except for narrow white margin on posterior edge of cach scgmcnt; rastern______soriculatipes Cresson (pt.) Abdomrn usually with ventcr yellowish; western______.. _libialt's Cresson 15. Wings darkly and uniformly infuscate______16 Wings hyaline___ • _. ______17 16. Abdomcn black; hypandrium emarginate (pi. VIII, 106) __ tuberculicllps Rohwer Abdomcn rufous; hypandriulll rounded (pi. VIII, 107) ______tacita (Norton) 17. Clypeus deeply cmarginate (pi. III, 13); abdomcn usually orange____ 18 Clypcus shallowly emarginate (pi. III, 14, 15) ; abdomcn black, at most with some rufous areas on basal stcrnites or central segments______19 IS. Inner margin of harpe strongly angulate (pi. VII, 92); wcstcrn__ d1:stans Norton Inner margin of harpe straight (pL VII, 94); eastern__ ___ multicincta Norton 19. Clypcus white; head and mesopleuron shining_____ soriculatipes Cresson (pt.) Clypeus black; head and mesoplcuron shining or dull and shagreencd_ 20 20. Hcad and mesopleuron dull and shagreened; abdominal tergites 3-7 orange; penis valve long and slender (pi. VIII, 101) __ impressata Provancher Head and mesoplcuron shining; abdomen mostly black; penis valve shorter and broader (pi. IX, 109) _____ (?) remota Rohwer or polita Cresson Descriptions of Strongylogaster Species S t1'onf/yl of/aster diRtan.~ Norton 8tron{}ylofll/_~t('r IIi.~tan8 :s'orton, 1868. p. 2'20. ~ ; Dalla Torre, 1894, p. 134; KOllo\\', 100;;, p. 96; Ross, 1937, p. 67; Ross, Hl51,p. 22; Beer, 1955, p. 238; Burks, 1958, p.ll. Strcmgyloga.stcr paoif/CIM )IacGiIlivray, 1893, p. 241, ~, ~; Dyar, 1894, p. 42; Konow, 1905, p. 96; Frison, 1927, p. 260; Ross, 1951, p. 22 (=distans Norton). Female.-Average length, 8.2 mm ..Antenna aml hen.d black; maxil lary and labial palpi whitish. Thorax black with tegula and most of pronotum white. Legs, except coxa, mostly light orange with base of each femur, apex of hindtibia, and each tarsus infuscate. Abdomen mostly orange with first and second tergites usually black and black areas sometimes present on sternltes anel other tergites; sheath black. Wi ngs hyaline. Head roughened and shagreened; clypeus deeply and angularly emarginate (pI. TIl, 13); malar space equal to diameter of front ocellus. Mesopleuron roughened, sparsely punctate; mesonotum shin ing; scutellum with several large punctures. Tarsal claw with long NEARCTIC SAWFLIES II. SELANDRIINAE 15 inner tooth (pI. III, 7). Forewing with anal crossvein absent. Hind wing with annl cell sessile. Sheath with ]ar~e leaflike scopa, with lateral halves not widely separated (pI. IV, D3). Serrulae of lancet with 10-13 subbasal teeth i serrlllae each separated by large circular notch (pl. V, 68). illale.-Average length, 7.4. nun. Color simihtl· to that of female except for legs, which are entirely light orange to white, and abdomen, whi('h is usually entirely omnge except for first two tergites. Structure s.imilar to that of female. HarI?e with inner margin strongly angulate; parapenis rounded at n,pex (pl. 'TIl, 92); penis yn,lye as in plate VII, 93. Types.-S. di.stan,'1 Norton is at the Museum of Comparative Zool ogy, Harvn,rcl F niversity. There are two specimens ( 'jl 'jl ), each bearing the labels "Cal. ('. Mus.," "Type 14007," "Strongylogaster distans." One has been chosen as the lectotype. ~[acGillivray's type ('jl) is at tho Illinois Natural History SUtTey and bears the data "Olympia, ·Wash., May 21, 1892, T. Kincaid." Distribution.-YlTester·n Nort.h America: British Columbia, Cali fornia, Colorado, Idaho, Nevada, Oregon, Utah, 1Vashington. Host.-Ptei'idiwn aquilinnJ11. (L.) Kuhn. Discussion.-This speeies is closely related to 1I11tlticinct(~ of eastern )forth Ameri('a, but it may be septlrated by the orange abdomen of the :female and genitalic characters of both sexes. The long inner tooth of the tarsal (']aw, the deeply emurginate clypeus, and the large scopa of the sheath will distinguish (li.~t(ln.s from other species. Beer (7965) discussed the life history and behavior and described the.!tU·Vll of t.his spe('ies. Adults are commonly collected in early spring near thei r host plant. 8tronUillogast("1" illlpI"C.~8atll.~ Provnnch('r, ISiS, 11. 170, 'jl; Or('S801l, 188On, p. cl2; Promnrher, 18S3, [1. 219; Ollila Torre, 18.'H, p. 135; Gallan and Rohwer, 1()1R, p. 107; Ros~, lO~1, p. 22; BurkH.l!N8, p.ll. 'I'II/·ina.c imJirC.YfJ(/tll.~, KOIlO\\', 1005, p. oo;~IacGiIliYrny, 1016, p. 67; Yllnsa, 1022, p. (;,3• .'{tron{JylogcM/CI" II/("tIl08l1.~ 1'ro\'nll("her, 1S,'I;}, p.12, Rtl'ongJ/logaBtC1' macula (KIng) 7,'enthrc({o (.:lllantll,~) mac/lla, Klug, 1814, p. 217; Hartig, 1837, p. 301; Kriech baumer, 1884, p. 202 . .'!tronUllloua,~ter m.acula" Thomson, 1870, 'p. 293; Thomson, 1871, p. 243; Andre, 1881, p. 413; Oameron, 1882, p. 100; Cameron, 1887, p. 45; Benson, 1952, p. ~9; )Iuxw('ll, 1955, p. 49; Lorenz and Kraus, 1957, p. 44; Smith, 1967, p. 95. Thrillax man/l.[a, Konow, 1885a, p. 23; Dalla Torre, 1894, p. 140; Konow, 1905, p. 96; .Enslin, 1M3, p. 202; Taylor, 1931, 'p. 41; Takeuchi, IMl, p. 236; B('rland, IM7, p. 200; Zhelokho\'tse\', 1951, p. 148. Female.-Avern.ge length, 6.6 mm. AllteIU1aa,Jlcl head black; cly peus, labrum, and maxillary and labial palpi white. Thorax black with tegllla and upper angles of pronotnm white. Legs, except extreme base of each coxa, (·ntirely orange yellow; tarsi infuscate. Abdomen NEARCTIC SAWFLIES II. SELANDRIINAE 17 black with various amounts of orange on third to seventh tergites, on lateral margins of each tergite, and usually several central sternites. ·Wings hyaline. Head mostly shagreened, shining only in ocellar basin; clypeus shallowly emn,rginate; malar space wider than diamet~r of front ocel lus. Thorn,x shining except for dull areas on pronotum, lateral n,reas of prescutum, and mesopleuron and metapleuron. Tarsal claw simple. Forewing with anal crossvein absent. Hindwing withn,nal een petio late. Sheath Harrow and pointed, ·with slender, cylindrical dorso apical projections (pl. IV, 46, 47). Lancet as in plat~ VI, 73. Male.-I have not seen males of tllls species. Type.-Klng's type is probably ill the Zoological Museum of Berlin. Distribution.-Europe, .Japan. North America,: B'ritish Oolumbia.: )lission City, .Tune 5,1963, ·W. R. M. Mason. Ontario: Ottawa, May 15, l!Hl. O. Peck. Host.-In Europe, Athyriwn filix-fellvi?la (L.) Roth and Pteridi~mL nqldlinwn (1..) Kuhn (Benson, 1952). Discussion.-The identity ofthis species is based on descriptions by Enslin (1918) and Benson (19592). I have seen only two females from widely sepltmted localities in North America. They are very similar to those of rlLjigaRtm, but .they differ in the more slender sheath and the more slender, cylindrical, dorsoapical projections of the sheath. StrongyZogastC1' nvuZticincta Nort.on Strongylogastcl' IIWltiCi.1l0tll-8 .'forton, 1862b, p. 143, 5,? j Norton, 1868, p. 221; Dulla Torre, 1894, p. 135 j Konow, 1905, p. 96; MucGilliYTuy, 1916, p. 68; Cresson, 1m, p. 7; Ross, 1951, 1). 23 j Hogh, 1966, p. 3±7. Stl'ongylogast(?r anll-IIlo811S Norton, 1868, p. 211, 5,?; ProYllncher, 1878, p. 169 j Provancher, 18H3, p. 210; Dulla Torre, 1894, p. 133; Dyar, 189i5, p. 310 (?) ; Konow, 1005, p. 00; ~IucGillivray, 11)16, p. 68; Yuusu, 1922, p. 53; Cresson, 1028, p. 3; Ross, 1951, p. 23 (=multicillctlls Norton). Strongylogaster lOllglllll.~ Norton, 1868, p. 216, ~; ProYIUlcher, 1878, p. 169; Provancher, 1883, p. 218; Dllllll Torre, 1894, p. 135; Konow, 1905, p. 00 j MucGillivray, 1916, p. 68; Cresson, 1928, p. 7; Ross, 1937, p. 6'7 j Ross, 1!);31, p. 28 (=-multicinctll.~ Norton). StrOIl-UlIloguster ulboanlllllatu.~ Rohwer, ]1)12, p. 237, 5,? j Ross, 1951, 1). 23 (= IItl/lUcillctu8 Norton). ,'itr01tUJllogu8tcr unnll10811.~ Yllr. c:inglllc8ccn.~ Ross, 1932, p. 250, ~, 5,? ; Ross, 1951, p. 23 (=mlllticillctll.~ Norton). Female.-Average lengt.h, 8.3 mm. Antenna anel head black; labrum and maxillary and labial palpi whitish; in specimens from Florida, first two antenn!LI segments, clypeus, supraclypeal area, gena, and SlJots on upper inner orbits may be yellowish to reddish brown. Thorax black with tegula white and pronotum white or yellowish. Le.l-,rs, except coxae, mostly light orange to white with black to infuscate areas on each femur and each. tarsus. Abdomen extremely variable, nearly entirely black with narrow to wide white margins on posteriol' edge of each segment to ne:Lrly entirely orange with similar ·white margins on each segment, with all intermediates; sheath black. ·Wings -hyaline to lightly infuscate, infusea,tion most common in southern specimens; \'cnation black ex('ept for eosta, subcosta, and stigma, which are usually yellowish infuscate; in southern specimens st.igma entirely yellowish, or black with lLpex n,ndlower margin yellowish. 18 TECHNICAL RULLETIN 13'98, U.S. DEPT. OF AGRICULTURE Head roughene.fl and dull, with large scattered punctures; c1ypeus deeply and anguiarly emarginate (pl. III, 13) ; malar space equal to diameter of front ocellus..Mesopleuron, sides of prescutum, :md hind margin of scutellum with large shallow punctures; remainder of thorax shagreened. Tarsal claw with long inner tooth. Forewi,ng with anal cross\"ein absent. Hindwing with anal cell sessile. Shf.!ath with large leaflike scopa, lateral halves widely separated (pl. IV, 32, 34). Serrulae of lancet fiat, with distinct subbn.sal teeth; each serrula not separated by deep notch (pl. V, 69). Male.-Average length, 7.7 mm. Color similar to tha.t. of female except for legs, which are entirely orange, including basal half of each coxa, and abdomen, which is entirely orange in most cases, sometimes with black areas on first two or three tergites. In sonthcl11 specimens clypells may be light, but head is usually entirely black and abdomen is US\Htlly d!trker. Structure as for female. Harpe more gently rounded than in diManR,· ha,rpe and parl"tpenis as in plate VII, 94; penis valve as in plate VII, 9n. Types.-l{ 1nultici-nciu8 Norton lmdS. annul-08u~ Norton are both at the Academy of Natural Sciences of Philadelphia; the former bears the data" Ii', Va.," and is type No. 270 and the latter has the data" 2 • Mass." and is type No. 271. There are specimens labeled a..', types of 8. longuhl.~ }'':orton at both the Museum of Comparative Zoology, Harvard University, and at the Philadelphia museum. The one at Harvard has n0 locality data and is a male; the one at Philadelphia has the data" 2 , Mass." and is a female. Since the original description is from three females ltnd since most of Norton's later work was done at Philadelphia, T Il.m hereby designating the specimen at Philadelphia as the lectotype; it bears the type No. 268. S. alboannu7aMl8 Rohwer (2) is type No. 10410 at the F.S. National Museum with the data "Brown's Mills ,Jet., N.J., .June 9, 1907." The type of the variety Ross desf'rihed is at the Illinois Natural Hist.ory Survey, and the type locality is Douglas Lake, Mich. Distribution.-Eastern North America: Florida, ~faine, Maryland. :Mass!lchusetts, Michigan, :Minnesota, New Brunswick, New Hamp shire, New .Jersey, New York, North Carolina, Nova Scotia, Ontario, Pennsylvania, Quebec, VeI,nont, Virginia. Host.-Pteridiu1n aqu,ilinwrn (T.J.) Kulm. Discussion.-This species is most closely related to distan8 of west ern North America, hut it may be separ,ated by the coloration of the female and by the genitalia of both sexes. The considerable color variation, especially of the abdomen, led to the description of the several species listed. Several specimens from Florida show n, distinct color variant, having more yellow on the ihead and having infuscate wings with the stigma and costa a contrasting yellow; however, I could not find structural differences between these and the northern specimens and am considering them only as a color variation. Hogh (1966) presented an ecological study of this species in Michigan. St7'ongyloga.ste1' 7wZita, Cresson 8tronU1l1oua,~tcr poliltls Cresson, 1880n, p. 20, 2; Provnncher, 1882, p. 294; Provnncher, 1883, p. 743; Dnlln Torre, 1894, p. 137; Konow, 1905, p. 97; NEARCTIC SAWFLIE~, II. SELANDRIINAE 19 Cresson, 1916, p. 8; )IacGillivray, 1916, p. 68; Gahan and Rohwer, HUS, p. 197; Yuasa, 1922, p. 54 ; Ross, 195;[, p. 23. Female.-Avemge length, 6.0 nun. Head and an'tenna alack; la.brum and maxillary and labial palpi whitish. Thorax black with tegula and upper an~les of prollotum white. Each coxa black with apex whitish; each trocl1anter whitish; fn:mt and middle femora black to infuscate, if black then apex is white; hindfemur black ,dt.h apex white; each tibia with basal one-third white, front :tnd middle tibiae with apical two-thiL"Cls ot"ange infuscate, hindtibia with apical two-thirds black; each tarsus infuscate to black with hindtarsus darkest. Abdomen black with narrow white band on posterior margin of each segment; cen tral areaS of abdomen sometimes orange. ,\Vings hyaline. Head shining, with yeryfine punctures; clypeus yery shallowly emarginate (pI. III, 15); malar space eqnal to diameter of front ocellus. Thorax shining; prothorax dun. Tarsal cIaw with small inner tooth. Forewing with !lmd cross\Oein absent. Hindwing with anal cen sessile or with extremely short petiole. Sheath narrow and pointed at apex, usually with apical tuft of hairs (pI. IV, 42, 43) ; scopa absent. Lancet very lightly sclerotized; serrulae 'lot distinet, covered with fine hairs (pl. VI, 77). ~Jale.-Unknown. Type.-The type of 'poZit11S ( Cj1 ) is No. 274 at the Academy of Na tural Sciences of Philadelphia I.;ith the data "Canada." Distribution.-Eastern North America: Oonn~'otUnlt: Lyme, May Hi, 1018, apple foliage, 'Y. S. Fisher. New York: L.I., Riverhead, August 1, 1950, R. Latham. Ontm'io: Blackburn, May 22, 1934, O. Peck; Ottawa. Host.-Yunsa (192)2) reported the host as Pte1'idi1(,'lJ1, aquili1l'ltln (L.) Kuhn. Discussion.-Cres.'50n (1880a) ascribes t.hic:; species to Provancher, but Pt"Qmnrher had not described it previously. Provancher used the <;ame name for the same species when describing it as new in 1882. 1 was not able to loeate Pronwcher's specimen, and it may be the same one designated as Cresson's type at Philadelphia. Provancher did not c1eseribe the Illttle, but Gahan and Rohwer (1,918) noted It male in the Pronlllcher eolJeetioll with yellow label 498 and said it was not the type. This species Illay be separated from 1'emota by the narrow pointed sheath and eharaeters of the laneet. Fnfortunately yery few speeimens are n,mihble for study. ~[ales lutve not been associated with the females of this species, and, ;tlthough I ha\'e seen se\'eral mn,les that may be th is speeies, I hesitate to describe them because of the close resem l)lallCe of )'('mota to 1Jolita. The males I have seen, however, are un doubtedly either this species or ?'(,lnota, and they may be separated from othrr spe('ies by the key. /';"h'onr;!Ilor;a.~t(,1' l"emf)ta Rohwer gtrOIl[lYlog(lstcr rcmotll.Y Rohwet·, 1912, p. 237, ~ ; Ross, 1051, p. 23. Female.-Ayemge length, 6.6 mm. A~ntenna and lle:tdhlaek; labrum and maxillary ancllabi:tl palpi whitish. Thorax ,yith tegnla and upper nngles of pronotllm white, rest of prothorax orange yenow; mesoplen 20 TECHNICAL BULLETIN 13'98, U.s. DEPT. OF AGRICULTURlll ron and mesonotum, especially lateral lobes and lateral margins of prescutum, reddish brown to dark rufous; rest of thorax black. Legs, except for basal half of each coxa, entirely orange yellow; tarsi infuscate. Abdomen mostly orange with white to yellow bands on posterior edge of each segment and with basal plates and usually apical segments blending toward black. Wings hyaline. Head shining with very fine punctures; clypeus very shallowly emarginate; malar space less than diameter of front ocellus. Thorax shining; prothorax dull. Tarsal claw with small inner tooth. Fore wing with anal crossvein absent. Hindwing with anal cell sessile or with extremely short petiole. Sheath short, broadly rounded at apex; scopa absent (pI. IV, 50). Lancet with serrulae low and distinct, each with about five or six subbasal teeth; numerous long hairs lacking (pl. VI, 75). Male.-Unknown. Type.-The ~ype ( ct) is at the U.S. National Museum, No. 14411, "lith the data "Germ't'n, Pa., May 2, 1908." Distribution.-Eastern North America: Penn.sylvania: Data as on holotype.Q'llebeo: Hull, May 21, 1924, C. H. Curran. Host.-Probably ferns. Discussion.-Although close to polita, this species may be separated by the broader and blunter she:\Jth and by the lancet that has distinct serrulae and lacks numerous long hairs. Several males have been examined that belong to either tlus species or palitaj however, none have been associated. I have seen very few specimens of this species. Strongyloga.ster 1'U;jigCI-stra. (Kincaid) Pa.ra·selan.dria rujiflastra Kincaid, 1900, p. 346, ct. Selandria ruji!Jastra, Konow, 1905, p. 95. Strongylofluster rttjiflaster, Ross, 1951, p. 23. Female.-Average length, 5.3 mm. Antenna and head bla.ck; cly pellS, labnun, a,nd nu\.xillary and labial palpi white. Thorax black with tegula and upper angles of pronoturn white. Legs entirely orange with basal half of each coxa black; tarsi infuscate. Abdomen varies from nearly entirely orange to nearly entirely bla.ck with various amounts of omnge; narrow white band usually present on posterior margin of each segment. Wings hyaline. Head shagreened, faintly shining, ocellar basin usually shining; clypeus shallowly ema,rginatej malar space more than twice width of front ocellus. Thorax mostly shining with mesopleuron and pronotum dull. Tarsal cla.w simple. Forewing with anal crossvein absent. Hind wing wit.h anal cell petiolate. Sheath narrow and pointed with short, blunt dorsoapica.l projections (pI. IV, 48, 49). Laneet as in plate VI, i4:. Male.-Unknown. Type.-Type No. 5284 at the U.S. National Museum with the data "Kukak Bay, Alaska, .July 4, 1899, Harriman Expedition '99, T. Kin ('aid, collector." Distribution.-Known OIlly from Alaska and British Columbia. Aln,8ka: Data as on holotype. B'litiBh Oolmnoi{),: Mt. Seymour, .Jlme 7, 1959,.A.. T. Wj]kinson; Roule:t.u L.,May 27, 1956, R. E. Leech; N. Pe:l.k, .Jade Pass,Mt. Revel stoke Nat. Pk., 7400', truly 16,1952, G. P. Holland. NEARCTIC SAWFLIES II. SELANDRIINAE 21 Host.-UnkTtoWIl. Discussion.-This species is similar to macula, being separated only by the wider and bhmter projections of the sheath and slight differ ences in t.he lancet. I IHLve seen only four speeimens; when more mate rial is twai1able, this and llWmhl may pro\-e to ~ conspecific. l'Jtl'ongyZogruitel' 80riculcdipes Cl'esson StrcmgylogU8tcr 80riclIlatipcs Cresson, 1880a, p. 20, 2 ; Dalla Torrt:', 1894, p. 1S8; Konow, 1905, p. fl7 ; Cre&<;oll, 1!n6, p. (); Ross, 1051, p. 23. Ntrollgylogustcr 80riClliatuil ProYan('her, 1&<';2, p. 21)6, 2 ; Prfwancher, 18H3, p. 7045; Harrington, 1893, p. 61; Dalla Torre, 18W, p. 138 (=,wriclIlatipes Cresson) : Dyar, 1805, p. :no; Gahan and Rohwer, 1018, p. 11)7. Female.-Average Ienf,rth, 8.2 mm...Antenna a.nd head black; ely [>ellS, labrum, and maxilla.ry and labial palpi whit{'. Thol·a.x blaock with t{'gula and upper a,ngles of pronotum white. Each coxa, trochanter, and femur orange; front and middle tibiae orange, hindtibia with basal otie-third whit.e, tlpieal t\Yo~thirds black; front and middle tarsi infus rate, hindtarSliS blaek. Abdomen bla('k except for omnge a,reus on vent{'r anel sides of segments 1-3. lYing;; hyaline. Head shining; elYPellS shallowly ell1aq!'inate (pI. III, 14); malar spfLce equa.l to cliamete.r of front ocellus. Thorax smooth a.nd shining. Tar~ml da.\\' with small inn81' tooth. Forewing- with anal crossvein pres {'nt. or absent. Hinc1wing with Rna I ('ell sessile. Sheath short, trunca.te at H.pex; SCOp,L absent (pI. IV, ~18, 39). La.ncet with serrulae flat and low at base, indistinct at apex; sllbhasa1teeth smRU and continuous (pI. VI, 71). Male,-Average, length, 6.4 mm. Color similar to that of female except for following: Legs entirely orange except for hindtibia and tl11 tarsi, which are mfuscate;abdomen entirely black except for very narrow white band on posterior margin of each segment. Structure as for female. GenHalia as in plate 'TIn, 96 and 97. Types.-.''i. 8oriC'ulatipps ( 2) is type No. 275 at the Academy of Xat1ll',tl Sci{'Jl('es of Philadelphia with the da.ta "I11inois." B. s07'icu llltll.~ ( 2 ) is at the Jfnseum of Quebec, Laval UniYersity,and bears the following data: Yellow label "692," name label "Strongylogaster sori('ulntus Pro\T." This specimen is desiJpmted here as the lectotype. Distribution.-Enstern North America: Illinois, Maine, Maryland, ~rassa('husetts, ~Iinnesota, New Brunswick, New York, Ontario, Pennsylvania, Quebec. Host,-I have collected adults of this spocies from Pte1idiwn aqui linutn (L.) Kuhn. Dyar (1895) reared this species from the same host.. Discussion.-This species mlly be distinguished by the white ely peltS, laek of orange on the dorsum of the abdomen, shining thora.'{, and lack of a scopa on the sheath of the female. It is most close.ly relatNl to tibia1i.~. The anal crossyein of the forewing may be either prN;{'nt or absent in this species, and in a series collected in l\faryland the ("'oRRyein waR pres{'ntin about one-third of the specimens. C"{,SROII (J880a) ga\'{' Pl'oY~lI1('her credit for thi.s species in his orig inRI dNwription; howe.vpr, Pronmch{'l' had not published 'a description of it. ProYanrh{'r (188i~) later descl'ibecl a spec-ies, 8o~'iculatu.s. wllieh may be based on the same specimens to which Cresson referred. X{'\'ertheless both speei{'s are the same. 315-625 O-{JO--t 22 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE St-rongyZ1Jgaster taeita (Norton) Tenthredo (Allantus) tacita Harris, 1835, p. 583. Nomen nudum. ~lllantus tacitus Norton, 1860, p. 246, ~, 'i1. Strollgylogaster tacitu Str-ongywga8te1' tibialis Cresson Strongylogaster tibialis Cresson, 1880a, p. 19, !? ; Dalla Torre, 1894, p. 138; Cres son, 1916, p. 9; Ross, 1937, p. 67; Ross, 1951, p. 23; Beer, 1955, p. 2Zi; Burks, 1958, p.ll. Taxoltl/,~ tibialis, Konow, 1905, p. 110. Selandria tl/argina.ta Provancher, 1R83, p. 8, !? ; Dalla Torre, 1894, p. 143; Konow, 1905, p. 95; Gahan Ilnd Rohwer, 1918, p. In; Ross, 1951, p. 23 (=tibialis Cresson) . Strrmgylogastpr fJrimativl/s l\IacGillivray, 1893, p, 241, !?; MacGillivray, 1894, p. 327; Konow, 1905, p. 97 j Frison, 1927, p. 260; Hoss, 1951, p. 2:3 (=Ubialis Cr('s80n) . Prototaxoll1l8 typiCIl8 Rohwer, 19lOa, p.-50, t, !?; Ross, 1951, p. 23 (=tibialis Cresson) . I'ltrongylogaster typiCU8, Ross,1937, p. 67. Female.-Averagelength, 7.3 mm. Antenna and head black; labrmn and maxil1ary and labial palpi whitish; C'1ypeus yaries from black to whitish. Thorax black with tegula and most of pronotum white. Legs with each coxa black; each femur black with extreme apex white; front and middle tibiae white, hindtibia black with extreme base white; front tarsus white, middl€ tarsus and hindtarsus black. Abdomen vari able, usually black with central three or four segments reddish yellow, hut may be nearly entirely yellowish or reddish brown or entirely black; narrow white band present on posterior margin of each seg mE'nt. 'Wings hyaline. Head moderately shining on postoC'ular and postocellar areas, other wise shagI'0l'necl; clypeus shallmY1y emaqdnate (pl. III, 14) ; malar space equal to diameter of front ocellus. Thorax moderately shining; pronotllm roughened. Tarsal claw with small inner tooth. Forewing with anal Cross\'ein present. Hindwing with anal cell sessile. Sheath short, truncate at apex; scopa absent (pI. IV, 40, -11). Lancet with ser ntlae low and flattened, without anteriol' subbasal teeth and with many fine posterior subbasal teeth (pI. YI,i2). Male.-Average length, 6.5 mm. Color similar to that of female ('xC'ept for following: Clypeus usually white; legs, except coxae, most of hindfE'mur, and all tarsi, orange; abdomen usually bhlck above and ,,-hitish hE'nE'ath. Structure as for female. Genitalitt as in plate VIII, DS and !l!l, Types.-8. tibialis Cresson (!?) is type No. 272 at the Academy of Xatul'al SciencE'S of Philadelphia with the clatl1 "Ne,-." S. mal'ginata PW'>!Ulcher ( 9) isin the Canadian "National Collection with the fol lowing labE'ls: "Type, Selandria marginata Pl'. No. U8," "Selandr!a marginata. Pro\'. Type?, J.F.Vic. B.C. '85," "Lectotype, Selandrla marginata. Prov., Comeau, Apr. U) ..W." This specimen was evidently C'olleC'tNl in l.'lHfj and the species described in 1883: consequently, it is probably not the specin1E'n Provaneher de!"C'ribed. However, t11is speci men fit;:; Pro\itnC'hE'l"s deseription, and if therE' is another specimen, it could not 1)(' lo(·ated. I am consillering this specimen as the type. s. JH'inwti1'1l8 ~Ia('Gi11ivr:ty ( !?) is at the Illinois Natural History SUITey with thE' data "Olympia, ",Yash., ).fay 18, 18!l2, T. Kincaid." P. t!fpi('""~ RohwC']' is represented in the r.s. National Museum by two cotypC's, a male and It female. The female is hE'l'e designated as the lee'totype, and it beal's the following data: "'Mts. lIear Claremont, Cal. 24 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE Baker," "U.S.N.M. Type No. 12817," "Prototaxollus typicus Roh., Ootype Stronuylogusfcr tllbcrc/lUccpS Rohw{'r. 1011e, p. 306, ~; Ross, 1951, p. 23 (= (?) tueit/l,y Rohwer). Stro-nUylollu,,'Jter lIIelulw(]w!ter Rohwer, 1911e, p. 397, ~ ; Ross, 1951, p. 2~. New synonymy. Female.-Average length, 7.2 mm. Antenna. black; head black with clypeus and malar areas yellowisl\ or black. Thorax entirely rufous or with celTical sclerites, pronotum and teg-ula black. Legs entirely black, at times with front tibia and femur light. Abdomen rufous; sheath black. 'Yings darkly and uniformly infuscat€. Head moderately shillIng, covered with large coarse punctures; cl.vpeus shallowly and angularly emarginate (pI. III, 14) ; malar space slight1y wider than diameter of front ocellus. Thorax shining, coarse pnnctures on angles of prothorax, mesopleuron, and scutellum, at times with several present on lateral margins of pl'escutum. Tarsal claw with small inner tooth. Forewing with anal crossvein absent. Hindwing with anal cen sessile. Sheath short anel rounded, with small scopa (as in pI. IV, 44-, 45). Lancet very lightly sclel'otized, without serrnlae, and with long lateral spines separating segments (as in pl. 'VI, 76). Male.-Average length, 6.4 mm..Antenna and head black; clypens may be partly light; thorax with mesonotul11, mesopleuror;.., and most of pectus rufous, otherwise black; legs and abdomen entirely black; wings darkly infuscate. Structure as for f(,l11ale. Hypandrium slightly emarginate at apex (pI. VIII, 106); penis vah'e with apex much broader than long (pI. VIII, 105) ; harpe and parapenis as in plate VIII,10;t Types.-S. tuberC1.iliceps ( Tribe ANEUGMENINI Genus ANEUGMENUS Hartig Emphytus subgenus Aneugmenlls Hartig, 1837, p. 253. ,lneugmen1ts, Andre, 1879, p. 241; Konow, 1886, p. 110; Dalla Torre, 1894, p. 140 (=Seiandria Leach) ; Rohwer, 1911c, p. 390; Ross, 1937, p. 65; Takeup.hi, 1041, p. 250; ilIaiaise, 1944, p. 4; Ross, 1951, p. 23; Zhelokhovtsev, 1951, p. 125 ; Benson, 1952, p. 60; Lorenz and Kraus, 1957, p. 49; Burlrs, 1958, p. 11; Malaise, 1963, p. 173. Selandria. subgenus Ancugmenus, Enslin, 1913, p.l95; Ross, 1930, p. 185. Type: :J.'entltredo (Emphytus) coronata Klug. Monotypic. C'otposelwn17l'ia. Enslin, 1912, p. 110; ilIalaise. 1944. p. 4 (=Anellgmenlls Hartig). Type: Oolposelandria jacobsoni Enslin. Original designation. Selal/rlria subgenus A.toposelandria EnSlin, 1913, p. 197; Malaise, 1944, p. 4 i Zhelokhovtsev, 1&51, p.140 (subgenus of A.neugmenus). Type: Selundl'ia tii,r8tenbergenHi.'J Konow. Original designation. Polyselandria MacGillivray, 1914, p. 104; Ross, 1930, p. 185 (=.Aneugmenu8 Hurtig). 'l'ype: Selanclria florida'lla MacGillivray. Original designation. Description.-Antenna short and stout; second segment as long as wide; third segment longer than fourth se.ooment; apical four seg ments reduced in length, together slightly less than third plus fourth segments (pI. III, 27). Postgenal carina present; clypeus truncate 01' slightly emarginate; malar space linear or less than one-half di.ameter of front ocellus. Prcpectus present as flat sclerite, separated from mescpisternum by suture. Tarsal claw with inner tooth and rounded basal lobe (pI. III, 8). Forewing with anal crossvein absent. Hind wing with anal cell sessile. 26 TECHNICAL BULLETIN 13'98, U.S. DInPT. OF AGRICULTURE This genus is most likely to be confused with Birka, but it may be separated by the presence of 11 postgenal carina and a basal lobe of the tarsal claw and by the anal cell of the hindwing lacking a petiole. The postgenal carina, basal lobe of the tarsal claw, presence of a pentag onal area on the head, and presence of a distinct prepectus will separate A neugmenus from N esoselandria. Probably a dozen species are in this genus, three of which occur in North America. Ross (1930) revised the North American forms as a subgenus of Selandria, but there have been considerable changes since then. The species of this genus are associated with ferns. Key to Aneugmenus Species 1. Pronotum entirely black; western______padi (Linnaeus) Pronotum with upper angles white or yellow______2 2. Scutellum black; clypeus truncate; eastern______flavipes (Norton) Scutellum yellow; clypeus slightly emarginate; southern Arizona ______~ _scutellatus, n. sp. Descriptions of Aneugmenus Species Aneugme1lJU8 flavipes (Norton) Beland-ria /lavipes Norton, 1861., p. 222, ~, ~ ; Norton, 1867, p. 258; Norton, 1872, ,po 85; Provancher, 1878, p. 101; Provancher, 1883, p. 202; Konow, 1905, p. 95 ; Ross, 1980, p. 188 ; Yuasa, 1922, p. 54. Aneugmenus /lttvipes, Ross, 1951, p. 23. Belandria /loridana MacGillivray, 1895, p. 281, ~ ; Konow, 1905, p. 95; Frison, 1927, p. 260; Ross, 1980, p. 189 (preoccupied by Belandria jloridana Cresson, 188Oa). Aneugmenus /lavitar8is Rohwer, 1911c, p. 390, S!; Ross, 1951, p. 23 (=/lavipe8 Norton). Belandria fiavipes var. /lcwitar8is, Ross, 1930, p. 188. Aneugmenus /lavipes occidentalis Rohwer, 1911c, p. 390, ~; Ross. 1930, p. 188 (= fiavipes Norton). Beland-ria fiavipes var. floridella Ross, 1930, p. 189 (new name for Belandria ftoridana MacGillivray) ; Ross, 1951, p. 23 (=jlavipes Norton). Female.-Average loogth, 5.2 mm. Antenna and head black; labrum and maxillary and labial palpi white. Thorax black with tegula and upper angles of pronotum white. Legs, except coxae, entirely yellow; tarsi infuscate at their apices. Abdomen black. Wings hyaline to very lightly infuscate. Clypeus truncate. Sheath short and rounded (pI. V, 51, 52). Lancet as in plate VI, 78. Male.-Average length, 4.0 mm. Color and st.ructure as for female. Genitalia as in plate IX, 110 and 111. Types.-S. flavipes Norton is at the l~luS€um of Comparative Zool ogy, Harvard University, and bears the la:bels "Type 14011" and "Selandria flavipes N., Ct., ~." S. floridana MacGillivray (~) is at the Illinois Natural History Survey with the d!l.ta "Ormond, Flor ida." A. fla~'itarsis Rohwer (~) is U.S. National Museum type No. 13941 with the data "Fla." and "collection C. F. Baker." A. flavipes occidentaUs Rohwer (~) is U.S. National Museum type No. 13939 with the data "Colo, 1212" and collection C. F. Baker." NEARCTIC SAWFLIES II. SELANDRIINAE 27 Distribution.-Eastern North America: Colorado, Connecticut, Florida, Georgia, Illinois, Maine, Manitoba, Maryland, Massachusetts, Michigan, Minnesota, New Brunswick, Newfoundland (insular) , New Hampshire, New Jersey, New York, North Carolina, Nova Scotia, Ohio, Ontario, Pennsylvania, Quebec, Vermont, Virginia, West Virginia, Wisconsin. Host.-Pte1id·iwm aq1tilinwm (L.) Kuhn. Discussion.-The variation in the infuscation of the wings, the presence or absence of yellow Rreas on the mesopleuron, and the black or yellow clypeus resulted in the description of the species by Mac Gillivray and Rohwer and the recognition of several varieties by Ross. .. Although the more lightly marked individuals and those with darker wings usually occur farther south, there is no character other than color to recognize varieties. This is a common species widely distributed east of the Rocky ~rotmtains. Specimens may be collected in Jarge numbers in early spring about the host. .tlneugmenu.'5 padi (Linnaeus) Tenthredo padi Linnaeus, 1761, p. 390 ; l\Ialaise and Benson, 1934, p. 5. Selan(/ria padi, Malaise and Benson, 1934, p. 5; Berland, 1947, p. 205. Aneugmf?nus padi, Zhelokovteser, 1951, p. 139; Benson, 1952, p. 60; Lorenz and Kraus, 1957, p. 49; Benson, 1962, p. 387 ; Burkl>, 1967, p. 15. Tenthredo strwmineipes Klug, 1814, p. 75, Cj1 ; Hartig, 1837, p. 282; Eversmann, lS47, p. 11; Malaise and Benson, 1934, p. fi (=padi Linnaeus). Sei(Inliri{L &tramineipes, Stephens, 1835, p. 41l; McLachlan, 1867, p. XC; Thomson, 1870, p. 291; Thomson, 1871, p. 238; Cameron, 1877, p. 197; Andre, 1880, p. 296; Cameron, 1882, p. 197; Brischke and Zaddarh, 1883, p. 291; Konow, 1885a, p. 24; Dalla Torre, 1894, p. 146; Konow, 1905, p. 95; Enslin, 1913, p. 201 ; Ross, 1932, p. 248; Hellen, 1935, p. 20; Ce,i>allos, 1941, p. 48; Zirngiebl, 1956, p. 322. ."tneuymentts stramineipes, Konow, 1886, p. 110; Ross, 1951, p. 23. Sclandria u,.bi.~ Ross, 1930, p. 186, Cj1 ; Ross, 1932, p. 248 (=stramincipes Klug) ; Benson, 1962, p. 387 (=padi Linnaeus). In 1934: Malaise a.nd Benson studied the Linnean sawfly types. Tenthl'edo pculi was found to belong in the genus .t1neugmeml8, and all references to padi Linnaeus of authors previous to 1934: do not; pertain to padiLinnaeus. Female.-Average len~h, 6.1 mm. Antenna and head black; labrum white; maxillary and labIal palpi infuscate. Thorax black with tegula white. Legs, except coxae, entIrely yellow; tarsi infuscate at their apices. Abdomen black. 'Wings very lightly infuscate. Clypeus shallowly emarginate. Sheath short and rounded (pl. V, 53). Lancet as in plate VI, 79. Male.-Unknown. Types.-The type is in the Linnean collection of the I ...innean SO('iety of London. KIng's type is probably at the Zoological "Museum of Berlin. The type of 8. 'I./,rbiB Ross (Cj1) is at the Illinois Natural History Survey with the data. "Vancouver, British Columbia, .Tune 20,1029, coIl. H. H. Ross." Distribution.-Throllg-hout Enrope, Xol'th Africa, Asia :Minor, and Transcaucasia. N ortIr America.: British Columbia, Oregon, 1Yashington. Host.-Pteridiwn aquilinwn (L.) Kuhn. Discussion.-This species ma.y be separated from flavipe.~ by the '28 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE emarginate clypeus, entirely black pronotum, and lancet characters. It differs from scu,tellatu-s by the black scutellum and lancet characters. This is a parthenogenetic species and was undoubtedly introduced into western North America, where it has become rather common in the coastal regions. A'1leugmenus s(.·ulellalu-s: new spe{!ies Female.-Length, 6.8 mm. Antenna, and head black; ]a,brum and maxillary and labial pll]pi white.. Thorax black with teguht, upper angles of pronotum, lwd scutellllln white to yell!)\\". Legs, except coxae, entirely yellow; tarsi infuscate at their apices. Abdomen black. 'Wings hyaline to very lightly infuscn.te. .. Antenna ShOlt and stout; second segment as long as wide; third segment longer tha.n fourth se~1'Jnent: apical four segments reduced in length, together only slightly shorter than third plus fourth segments. Postgenal carina, pl.·esent; clypens shallowly emaqrina.te; malar space less than one-half diametel' of front ocellus. Prepe<.'lflls present as flat sclerite, separated from rnesepisternurn by suture. Tarsal claw with moderate inner tooth lwd rOllluled basal lobe (as in p1. III, 8). Fore wing with ana.l crossvein absent. Hindwing with a.nal cell sessile. Sheath short and rounded (pI. V, 54, 55). Lllnce.t as in plate VI, 80. Male.-Length, 6.0 mm. Color similar to that. of female except for abdomen, whieh has tergites 2-6 rufous and sternites 1-3 yellow to rufous. Structure as for female. Penis yah-e with dorsoapicallobe (pI. IX, 118) ; harpe a.nd parapenis as in plate IX, 112. Holotype.-Female, Rusflers Park, Chir Birka )[lllnise, 1944, p. 4; Zhl'lokhoYtseY, 1951, p. 12;}; Mnlnisl', 100:3, p. Ii;:; (=Mclisandra Bl'Il!;On). Type: Tenthredo (AllantIlJl) cillcreipe,y Klu!,\'. Originnl dl'signatioll. NEARCTIC SAWFLIES n. SELANDRUNAE 29 Descript'ion.-Antenna short, second segment as long as wide; third and fourth segments subequal in length; apical four segments not no ticeably reduced (pl. ITI, 25). Postgenal carina absent; c1ypeus trun cate; malar space equal to or less than diameter of front ocellus; penta gonal arel\ of head present. Prepectus present as flat sclerite, separated from mesepisternwn by suture. TarsaJ c1a.w with small inner tooth (pI. III, 9). Forewing with anal crossvein absent; first abscissa of Rs pres ent. Hindwing With a.nal cell ~tiola,te. The species included in tIns genus were previously placed in Meli S(t1/(Z1'Cl. The characters for differentiation of this gellus and Neso uland'l'i(t a,re discussed undel' the latter. This genus differs from Aneug menU8 in iacking a postgena,l carina a.nd not having the l1J?ical four antenna1 segments reduced. Besides these morphological dIfferences, the species of Bi1'!.:a are associated with !,,1Tasses and sedges, whereas the species of Aneugmenu8 ILre associated with ferns . •\.bout five speeies are in this genus, one of which hn<; been ta.ken in Alaska. and CILlutda. Sinee I haNe been unable to find any described spooics thRt fit this fo1'111, I am describing it as new. Description of Birka Species Bil'lal -nol'dira. new species Female.-Lengt:h, 5.8 nun..A.ntetma. and head black. Thorax black with upper nUl.l'gins of pronotwu white ILnd outer ma.rgins of tegulae brown. Legs With (·.oxae, trocha.llters, and femora. black; ext.reme apex of eadl femlll' and front and middle tibiae white; hindt.ibia with basal two-thirds white, a,pica.1 one-third black; eaeh basitarsus whit.ish, rema.ining ta,rsal se~"llents infuscMe. Ahdomen black. 'Vings hya,line . •\ntenna short: sec.oud seg"ment. as long as wide; third segment slight.ly longer and nearly subequal in length to fourth segment; seg ment:{ beyond third gmdually decreasing i1l1en~rth (pI. III, 25). Post~ gena.l earina absent; clypeus truncate: malar space equal to diameter of front ocellus. Preped,us present as flat sclerite, separated from me..~pisternum by suture. Tarsa.l claw with small inner tooth. Forewing wit.h anal cros..<;vein absent; first absc.is5a. of RR present. Hindwing with anal cell petiolate. Sheath long, narrow, rounded at apex (pl. Y, 56, 57). Lancet as in plate VI, 81. Male.-Length, 5.1 mm. Color anel structure similar to t.hose of female. GenitalilL as in plate IX, 114 and 115. Holotype.-Female, North Fork Crossing, Mi. 43, Peel PIt. Rd., 3500', .July 3, 1062, Yukon Territory, P . .T. Skitsko. Deposited in the Canadian National Collection. Allotype.-Male. Same data, as holotype. Deposited with the ho10 type. Paratypes.-Alaska: Mile 315, Richard Hwy., June 7, 1951 W. R. ~L ).[1150n (899); Mile 115, Steese Hwy., June 22, 1951, Mason I1nd~[('Gil1is (l d"); Big Delta, June 6, 1951, W. R. M. Mason (1 9) ; Anehornge, June 27, 1951, R. S. Bigelow (1 9); Anchorage, June 24, 1951, R. S. Bigelow (1 9); Isabel Pass, Mi. 206, Richardson Hwy., 2900', July 18,1962, P. J. Skitsko (299,1 d"); Isabel Pass, Mi. 206, Richardson Hwy., 2900', July 17, 1962, P. J. Skitsko (1 9); Isabel 30 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE! Pass, Mi. 206, Richardson Hwy., 2900', July 16, 1962, P. J. Skitsko (1 9); Curry, June 29, 1952, W. R. )'Iason (1 9) ; Ourry, June 28, 1962, W. R. Mason (1 9); Tanana, June 8, 1951, R. 1. Sailer (1 d'I); Little Susitana, June 6, 1959, R. H. Washburn (1 9) i Palmer, June 1, 1953, R H. Washburn (1 d'I); McKinley Hotel, June 18, 1954, W. O. F. (1 9). Alberta: Edmonton, June 12, 1937, E. H. Strickland (1 d'I). British Columbia: Summit Lake, )'fi. 392, Alaska Hwy., July 2-4, 1959, 4500', R. E. Leech (1 9); Summit Lake, )'fi. 392, Alaska Hwy., July 17-19,1959,4500', E. E. :\[acDougall (1 d'I); Summit Lake, Mi. 392, Alaska Hwy., July 11-14, 1959, 4500', E. E. ~[acDougall (1 9); Atlin, 2200', June 13,1955, H. Huckel (1 9).l.VIanitoba: Gillam, July 5, 1960, J. F. ).·[cAlpine (1 ci"'). Y1lkon Territory: Same data as holotype (299) J 1:'orth Fork CJ~ossing, ).[i. 43, P~el PH.. Rd., 3500', July 4, 1..96~, P. J. Skltsko (299);:::\ orth Fork OrossIng, ),11. 42, Peel PIt. Rd., 3:>00 , June 25, 1962, R. E. Leech (1 d'I); Korth Fork Pass, Ogilvie )'Hs., 4100', June 17,1962, R. E. Leech (1 d'I); Otter Lake, 130°25',62°30', 4000', July 15, 1960, E. W. Rockburne (1 9); Ross Ri\Ter, 132°30', 61 °56', June 19, 1960, E. W. Rockburne (1 9); Ross River, 132°30', 61 °56', June 22,1960, J. E. H. Martin (19). Deposited in the Oanadian National Collection and the U.S. N ational ~!1useum. Host.-N0 host data are il\'ailable. This species may feed on grasses or sedges as do the European species of this genus. Discussion.-I ha\'e been unable to find any Palaearctic forms that conform to this species. It is close to cinereipes (Klug) of Enrop e; however, nordica may be separated by thelighter tegula and pronotum. Genus EUSTROMBOCEROS Rohwer Stromb()ccro.~ subgenus Ellstrombocero8 Rohwer, 1011e, p. 394. Eustromboccr08, Ross, 1037, p. 66; Malaise, 1942, p. 94; Malaise, 1963, p. 167. Anellylltenu,y subgenus EII,~trombocc"'n, Ross, 1!)51, p. 24. '.rype: Str(}lI~bocero8 (Elistromboceros) melallopleruiJ Rohwer. Origiual desig' nation. Description.-A.ntenna moderately long, usually thickened in male; second segment disklike, wider than long; third and fourth segments snbequal in length; segments beyond third gradually decreasing in length (pI. III, 24). Postgenal carina absent; clype.us truncate; malar space linear. Prepectus present as flat sclerite, separated from mesepis ternum by suture. Tarsal claw simple or with small or moderate inner tooth; basal lobe absent (pl. III, 10). Forewing with anal crossvein absent. Hindwing with anal cen sessile. Several species are in this genus from )Iexico and Central America. Only one species reaches .Arizona. Description of Eustromboceros Species EU8tl'Olllbort')'08 ?"lljorall(/atIl8. new species FemaZe.-rnknown. MaZe.-Length, 6.5 l11m. Antenna and head black; labrum and maxil lary and labial palpi whitish. Thomx black with most of prothorax, postspiracular sclerite, and tegulae white to light oran!:,re. Legs with ('oxae and most of eaeb femur bl1tck, apex of ear:h femur whitish; NEARCTIC SAWFLIES II. SELANDRIINAE 31 front and middle tibiae and tarsi whitish; hindtibia and tarsus black except for whitish line on inner sm'face of tibia. Abdomen entirely orange with basal plates and hypandrium blackish. 'Wings uniformly and moderately infuscate. A.ntenna about twice width of head with segments thickened and slightly expanded at their apices, appearing subserrate; second seg ment disklike, wider than long; third and fourth segments subequalm len~rth ; remaining segments decreasing gradually in length and slightly tapering in thickness toward apex (pI. III, 24.). Postgenal carina absent; clypeus truncate; malar space linear. Prepectus present as flat sclerite, separated from mesepisternum by suture. Tarsal claw simple. Forewing with anal crossvein absent. Hindwing with anal cell sessIle. Genitalia as in plate IX, 118 and 119. Holotype.-Male, Post Creek Cn., Pinaleno Mts., Fort Grant, Ari zona, July 15-18, 1917. Deposited in the U.S. National Museum, type No. 69158. Paratype.-One male, same data as holotype. Deposited with the holotype. Distribution.-Known only from the typelocality. Host.-Unknown. Discu88ion.-I was llilable to find any described species resembling this form in Cameron (1883) or other sources. Further collecting in this area will be needed to locate the female and associate the sexes. Genus NESOSELANDRIA Rohwer Ncsosclandria Rohwer, 1910b, p. 657; Forsius, 1933, p. 182; Takeuchi, 1941, p. 262; Malaise, 1944, p. 11; Zhelokhovtsev, 1951, -p. 131; Malaise, 1963, p. 173. Type: Parasclandria immatatri:ll Ashmead. Original dp.·signation. Neobusarbia Takeuchi, 1928, p. 40; Takeuchi, 1941, p. 262 (=Nesoselandria Rohwer). Type: Neobu.,arbw ftavipcs Takeuchi. Ori~~nal designation. Melisandra Benson, 1939, p. 110; Benson, 1940, p. 211; Takeuchi, 1941, p. 262 (=Ncsosclandria Rohwer) ; ZhelokhQvtsev, 1951, p. 131 (=Ncso8cla.ndria Rohwer) ; Benson. 1952, p. 60; Malaise, 1963, p. 175; Smith, 1967, p. 95. Synonymy reestablished. Type: Selandria morio (Fabricius) Enslin. Original designation. Description.-Antenna short; first and second segments longer than wide; third segment llh times length of fourth segment; segments beyond third gradually decreasing in length (pl. III, 26). Postgenal carina absent; clypeus truncate to shallowly emarginate; malar space linear -to one-half diameter of front ocellus; pentagonal area of head obsolete. Prepectus indistinctly defined by suture. Tarsal claw with long inner tooth j basal lobe r..bsent. Forewing with anal crossvein absent; first abscissa of Os atrophied or very faint. Anal cell of hind wing with short petiole. This genus is close to Birka, but it may be separated by the longer second antennal segment, the length of the third antennal segmeut, the absence of a pentagonal area on the head, the long inner tooth of the tarsal c1li;w, the near absence of the prepectus, and the faint first abscissa of 08 of the forewing. Benson (1939) erected }.[eli.'Jandra to include those specie.'l that pre\Tiously had been placed in A.neugmenus that lack a postgenal earina and are associated with grasses and sedges, not ferns. However, Benson designated m.o'rio as the type spe 32 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE cies, and moria agrees very closely with the type species of NeSo. Stromoocero~ subgenus Stromooceridca Rohwer, 1911e, p. 392. A.lIcugmelluB subgenus Stromboceridea, Ross, 1937, p. 68; Ross, 1951, p. 24. Stromboocridea, 'Malaise, 1942, p. 91; Malaise, 1963, p.167. Type: StrcnnbOCf)7"OB (Stro-mboceridea) pilOBUluB Rohwer. Original designation. Description.-Antenna moderately long; second segment longer than wide; third segment longer than fourth se~~_ent; apical four se~ents distinctly reduced in length, together suoequal in length to thml segment (pI. III, 23). Postgenal cal'ina present; clypeus trun cate to shallowly emarginate, strongly convex in profile (pI. III, 18) ; malar space linear. Prepectus present as flat sclerite, separated from mesepisternum by suture. Tarsal claw with long inner tooth situated clo88 to large, acute basal lobe (pI. III, 11). Forewing with anal ('rossvein absent. Hindwing with anal cell sessile. Sewn:.] Neotropical species are in this genus, only one of which reaches Arizona. Description of Stromboceridea Species Strmnboce'ridea nig1'ican~ (Norton) Strol/gy/ogaster lIigricans Norton, 1868, p. 223, ~, ~; Cameron, 1883, p. 14; Dalla Torre, 1894, p. 137. Strolnoocer08 nigrical/,'j, Kono\\', 1905, p. 98. A,IlCIIUlllcnll.~ (Swomboceri4ea,) nigricall.~, Ross, 1951, p.24. fic/andria, cllrialis Cresson, 188011, p. 15, Cj? : Dalla Torre, 1894, p. 142; Konow, 1905, p. 95; Cresson, 1916, p. 4: Ross, 1!)51, p. 24 (=nigricans Norton). Stromboceros (Eustromboccros) /Cllco.~tomIl8 Rohwer, 10lle, p. 395, Cj?; Ross, 1951 p.24 (=lI'igricane Norton). NeoanapcjJtam('1I Distribution.-Mexico. Southern Arizona (Pa~'lgonia, VIII-I-'24, E. P. VanDuzee). Host.-UnknOWll. Discussion.-The relationslLip of this species to other members of this genus will be difficult to determine until the Mexican fauna is better known. The use of Norton's name nigl·ican.'i is based on the original description and the traditional concept of this name. Tribe SELANDRIINI Genus BRACHYTHOPS Haliday ('or/lna Lepeietier aml Sen'ille, 1828, p. 569; Dalla Torre, 1894, p. 14-0 (=Sciandria Leach) ; Ross, 1930, p. 18.'): Benson, 1936, p. 204; Ross, 193'.... p. U6 (=SeirlIIdrirL L{'aeh). Pr{'Occupied hy ('or/llla Bose, 180'2. Type: Tcnthredo scapularis Lepeletier and Serville. Designated hy Rohwer 19l1h. Brachythops Haliday, 1839, p. 764; Benson, 1936, p. 204; Benson, 1940, p. 210; ~falaise, 1044, ll. 3 (=8el~lIIdria. Leach); Zhelokhoytsev, 1051, p. 128 (a subgenus of Sclandria Leaeh) ; Benson. 1052. p. 61. Type: Bruchythops ,Yeminfgra Haliday. l\[onotypic. Pu·ra.~ela1Ulria Ashmead, 1898, p. 255; Konow, 1905, p. 04 (=Selandria. r_each) : Ross, 1930, p. 185 (=('oryna Lepeletier and Serville); Ro!'s, 1937, p. 66 (=Selandria Leach) ; Zhe)okhovtse,', 19ii1. p. 128 (=Brachythops Haliday). Type: Selanrlria flavcn.~ King. Original designation. Rclan(iririea Rohwer, 1911e, p. 388; Ross, 1930, p. 185 (=Ooryna Lepeletier and Serville) ; Ross, 1937, p. 66 (=Selandria Leach) ; Zhelokhovtsev, 1951, p. 128 (=BrachythoP8 RaHday). Selandria suhgenus Selandridea, Enslin, 1914, p. 194. Type: Selanrlriclca vanrlllzeci Rohwer. Original designation. P8eurioselU714ria ~racGillivrar, 1914, p. 103; Ross, 1930, p. 185 (=Oor/ll1U Lepe letier and Serville) ; Ross, 1937, p. 66 (=Sclanclria Leach) ; Zhelokhovtsev, 1951, p. 128 (=Brachyfhop8 Halidar). Type: Plleuriosela71dria oxalata ~racGillivray. ~Ionotypic. Description.-Antenna short and stocky; second segment globular, as long as wiele; third segment slightly longer than fourth segment; segments beyond third gradually decreasing in length (pI. III, 28)_ Eyes short and oval; postgenal carina absent; clypeus truncate to shallowly emarginate; malar space more than twice diameter of front ocellus; mouth parts reduced, labial palpi three-segmented, maxillary palpifoul'-segmented. Prepectus present as flat sclerite, separated from mesepistel'num by deep groove j inverted V-shaped suture present on mesoplenron, separating sclerite into two parts (pI. III, 19). Tarsal r}aw with smalI inner tooth; basal lobe absent. Hindbasitarsus shorter than three :following segments together. Forewing with anal crossyein absent; costa distinctly swollen at apex (pI. II, 3). Anal cel] of hind wing with short petiole; "ein 11'-711 arises :from vein S('+R instead of from vein Rs (pI. II, 3). The genera Belamh'ia and BmchytllOp,c; are generiral1y distinct as rlefi!H'd by Benson (1.936. 195f) . .All the genera listed here are isogeno tYPIC'. So far as is known, the genus )'\flandria does not o(,(,ur in North ~\meri('a. This genus may be separated from Selandr'ia by the short, on1l eyes, wide malar space, shallowly emlu'ginate ('lypeus, SllOrt hindbasitarsus, and the reduced mouth parts with the labial palpi three-segmented NEARCTIC SAWFLIES II. SELA.J.'1"DRIINAE 35 and the maxillary palpi four-segmented. The labial and maxillary palpi are four- and six-segmented, respectively, in 8elandria. Bmchy t!lOps may be separated from other North American genera by the wide malar space, lark of a postgenal carina, and the presence of an in \'erted V-shaped suture on the mesopleuron. Two species occur in Xorth America, both of which are Holarctic. Key to Brachythops Species Front lobe of mesonotum densely pubeseent and with fine surface sculp ture; clypeus and antennu entirely blnck; pronotum mostly black; wings illfus('ute with venution bluck except at extreme buse, stigma with pos terior half yellowish brown ______'ICIIClltlleii (Konow) .Front looe of lII('Sollotum with very sparse pubescence at sides only and sllrfll('(' shining, without fine punctures; rlypeus usually with front lJlaq;'in yellow; nntenliU uSllully with ut leust basal segment yellow; pronotuUl mostly ye'llow; wings yellowish hyaline with yellow venation at Ie-.lst on oaslil half of forewing, stigma usually all bla(·k____ fiUVCIIII (h."1ug) Descriptions of Brachythops Species Brachytlwp8 flaven.g (Klug) TC'nthn!(lo (}UlUlltllll) fiUl·C/!•• Klug, 18H, p.48. 0, S1. 8('lalldr;a /tur(I/.'!, Cameron, 1882, p. 100: Dalla Torr!', 1R94, p. 142; Konow, 1905, p. 05; EIl!;lill, 1013, p. 1\}5; ~ruluise. 1921, p. 99; Berland, 1947, p. 200; Zh('lokhovtse\', 1!r.i1, p. 128: Burks. 1007, p. 16. Bru('hythops fian>ns, B('nson, 1936, p. :.!O!; Hl'lIen, 1948, p. 41; Pasteels, 1951, p. 134; Bt'nson, 1952, p. 61; Lorenz and Kraus, 195i, p. 4i; Benson, 1962, p. 3Si. •r .:;('{(IIldria dceolo"ulu CresMn. 1R&>a. p. 15, i5 : Dalln Torre. 1894, p. H2; Konow, 1905, p. 95 ; Cresson, 1916, p. 4 ; Ross, W51, p. 24. New synonymy• •t .{elalliirid('a. decolaratu. Rohwer, 1911c, p. &~8. ('orYlla decolorutu, Ross, 1930, p. 185. Sclundridea ·L'undllzeei Rohw!'r, 1911c, p. 388, 0, S1.; Ben;:on, 1962, p. 387 (= fiuVC7!,'J Klug), Corynu runduzeei, Ross, 1930. p. 18.'5. Rl'iundr;a 'L"UmiIIZCci, Hos':!, 1951, p.24. P,YC'I/do,~('lulldrin. o;ra/utu MacGillivray. 1914. p. 103, S1 ; Frison, 1927, p. 258; Ross, 19aO, p.l85 (='l:ulldllzeci Rohwer). New synonymy. The European synonymy for tills species is given by Kono\\' (1.905) and Enslin (1913). Female.-A.verage .length, 6.5 mm. ~;\ntel111a black with first segment or first and see-ond segments yellow. Head black with labrum, maxil lary and. labial pa.lpi, and at least apical half of c1ypeus whitish. Thorax black with tegula, most of pronotum, and sometimes meso pleuron yellowish. Legs beyond trochanters entirely yellowish; ex treme base or each femur and tarsus usually infuscate. Abdomen en tirely orall~e; basal plates may he black and sometimes basal sternites and tergites infuscate. 1Yings light yellowish hyaline with yellow Yellation at least on basal half of forewing. Front lobe of mesonotum with "ery sparse pubescence on side only; :mr.fnee shining, without fine punctures. Sheath with large scopa (pI. V, 60-62), Lancet as in plate VII, 90. Male.-Average length, 6.1 nun. C{)lor as for female except for following: Mesopleul'on usually pa.le; sides of mesonotmn and metano tum pale; legs entirely yellowish except for black at base of ench coxa; 36 TECH1>.'ICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE abdomen usual1y entirely omn~. Structure as for female. Genitalia as in plate X, 120 and 121. Types.-S. decolo1'at(t Cresson ( 0) is type No. 189 at the Academy of Natural Seiene'es of Philadelphia with the data "Col." P. orealata ~IaeGilliYmy (<;» is at the Illinois Xatural History Survev and bears the data "\Visconsin." .'.,t. 1'rtnrluz(,f'i Rohwer (<;» is U.S. NationaJ Museum type No. 13998 with the clata "Buffalo, N.Y., .June 4,1910, M. Yan Duzee." Klug's type is probably at the Zoologieal ~IuFellln of Berlin. Distribution.-Central and northern Enrope; Siberia. North America: Alaska, Alberta, Colorado, Manitoba, Newfoundland (Goose Bay, Labrador), New York, NorthweRt Territories (Ft. Smith), Quebec, Saskatche.wan. \.\"i5<'on5111. Host.-{}a;·ex spp. Discussio~.-rr;his species may be separated from wllestlleii by tho5(' chamc·teI·s .f!1\·en 1Il the key. The ('olor of the mesoplenron ranges from entirely black to entirely rufous and WlIS 11 charadeI' IH·e\·ioli;;;)y llsed to separn,te dc('o7ol'tda nnd I'lInrluz('('i.S. jlr(I'{'l1S is apparently most common in th(\ northem lat.itude;;;, beeause. most of Ule. speeimens have been collected in northern Canada. BJ'{(('hytlWP8 11'1U'stnfii (Konow) Seiandria 'lC'u('sflleii Konow, 1885b. p. 122, ¢. <;>; Dalla Torre, lRfi4. p. 147; Konow, U)();'5, p. 95; Enslin. 1\)13, p. ]95: Benson, 1934. p. 202; Berland, HH7, p. 201; Zhelokhoytsev, 1951, p. 129; Burks, 1967, p. 16. Bruchythopll 1{'ucsfll('ii, Benson, 1936. p. 2().t; Ben!'on, 10;;2. p. 61; HelUill, 1!l:;:;. p. 113; Lorenz and Kraus, 1957, p. 48; Benson, 1962, p. 387. Female.-Ave-rage len¢h, 6.7 mm. Antennfi and head black: labrum and maxillary and labial palpi white. Thorax black; tegula sometimes light brown. Legs with each coxa, trochanter, and femur black; ex treme apex of each femnr and each tibia orange yellow; tarsi infuscate. Abdomen mostly orange with black at base and extending apically on dorsum and venter, amount of black find orange \rariable. Win~ with venation black e.xcept at extreme base; stigma with npper half black, lower hnH light; wings lightly infuscate. Prescutnm densely pubescent and with fine snrface punctures. Sheath with broad scopa (as in pI. V, 60-62). I,ancet as in plate VII, 91. Mate.-Average length, 5.6 mm. Color similar to that of female except eacll femur is orange yellow except at extreme base and abdo men is usually darker. Structure as for female. Genitalia a;;; In plate X, 122 and 123. Type.-Konow's type is at the Del1tsches Entomologisches Institut, Ebers,valde. Germany. Distribution.-('(>ntral and 110l-tllel'1l Eumpe. NOlth America: Alaska, Alberta, ~ranitoha, Northwest Territories (Aklavik), Quebec, Yukon Territory. Host.-('flrf.).~ spp. Discussion.-This speeies may be. separated from jl{/l'fll.~ by those characters given in the key. It is usually smaller and darker than NEARCTIC SAWFLIES n. SELANDRIINAE 37 flavens. Benson (1962) first reported this species from North America, a.nd it has been taken only .in the northern parts of Canada and in Alaska. Tribe ADELESTINI Genus ADELESTA Ross Adelesta Ross, 1937, p. 68; Benson, 1938, p. 362; Ross, 1951, p. 24; Malaise, 100.3, p. 169. Type: Se/andria. n01Ja Norton. Original designation. Description.-Antenna subserrate with each segment slightly ex panded at its apex; second segment as wide as lona ; third segment slightly longer than fourth segment; segments beyon~ third gradually decreasing J1l length (pI. III, 29). Postgenal carina absent; c]ypeus truncate; malar space equal to diameter of front ocellus. Prepectus absent; posttergite not clearly dill"erentiated. Tarsal claw simple, long, and slender. Basal plates narrow, widely separated, leaving large membranous area. Forewing with anal crossvein present; distal anal cell subequal in length to proximal anal cell; proximal anal cell mark edly constricted near its base (pI. II, 5). Hindwing with anal cell petiolate. The subserrate antenna, absence of a prepectus, and presence of the anal rross\"ein in the forewing will separate this genus. Ross (1937) erected a new subfamily, the Adelestinae, for this monotypic genus although he stated that it is in the selandriine group. Benslll (1938) placed Adele8ta in the Selandriinae, and subsequently Ross (1951) retained this genus in a separate tribe in this subfamily. The curved R8+ill vein of the forewing places this genus in the Selandriinae, a position that seems most adequate at present. Description of Adelesta Species Adelesm 'Twva (Norton) 8e'lcmdria nora Norton, 1867, I). 254, 9 ; Dnlln Torre, 18M, p. 144. Hoplocllmpll ? nO'va, Kirby, 1882, p. 168. Lyeaota n01Ja·, Konow,19(};), p. 102. At1ele,~ta noea, Ross, 1937,p. 67; Ross, 1951, p. 24. Female.-Averngelength, 4.6 mm. Antenna and heacl black. Thorax blaek with teg-ula light. Legs with earh roxa, trorhanter, and basal ludf of each femur black; apiral half of each femur, each tibia, and tarsus white j tarsi infusrate. Abdomen blnck. "rings hyaline. Front tarsus only slightly longer than front tibia; hindbasihu'Sus subequal in length to three following tarsal segments. Sheath short and rounded. Male.-D"nknown. Type.-Norton's type has not been located. Ross (1937) based the USe of this name on a ft'male identified as 1W!'a. by Harrington. In the Ol"iginal cleRcription Nortollstated "Maine, 1. 9." 38 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE Distribution.-Northeastern North .America: New Brunswick, Nova Scotia, Quebec. Host.-Unlmown. Discussion.-Ihave seen only four specimens of this unusual species. It is easily separated from other Selanc1riinae by the antenna, wing venation, and lack of a pre pectus. - Tribe HEPTAMELINI Genus HEPTAMELUS Haliday Mplicprt(L Stephens, lR:.{ij. p. M. Preoccupied by .Melirerta, Schrank, 1808. Type: .I[elicer/II o('/trolell('a Stf'(lhemi. :\fonotypic. [[('))tU11lelll.~ HlllidllY. IS-X;. p. no; CamETon, 1.<;82, p. 301; Dalla Torre, 1894, p. 196; Kono\\,. 100:;, p. n: Enslin, 1014, p. 256; Benson, 1088, p. 368; Berland, 1!).17, p. 277; Drnson, 19;)2, p. 57; Takeuchi, 10;;2, p. 5'1: Lorenz and Kraul', l!}.i7. p. 47; Togashi, 1061, p. 264; :\falail;'e, 10H3, p. 1<'>2. TSlX': J!('litC'rta ochrolCI/('(I. :';tPphPllf>. :\Ionotypic. C'acnollcllrlL ~'homs()n, 1870, p. 270; Cumeron, 18~2, p. 801 (=Hcpt(/ml'lll.~ HulidllY). ~'ype: ('U(,1I0)lCIII'(£ dahl/Jomi Thortt,on. :\!onotypic-. Description.-Antenna seven- or eight-segmented; second segment longer than wide; third segment longer than fourth segment; seg ments beyond third subequal in length (pI. IIT, 30). Postgenal carina absent; clypeus shallowly emarginate; malar space equal to diameter of front ocellus. Prepectus absent. Tarsal claw with long slender inner tooth i basal lobe absent (pI. III, 12). Forewing with anal crossvein present; vein 2.tl and 3A basal to crossvein faint; stigma nearly as broad as length of 2r; ,-eins J.1I and In-(''/b divergent; veins l1l and C'/,t-a interstitial. Hindwing with cell R8 one-half or less size of l1l " anal cell wit h long petiole (pl. II, 4). This genus may be. recognized by the short antenna and wing vena tion. It was previously placed in the Heterarthrinae until Benson (1/)62) coneC'tly pl!lC'ec1 it ill the Selanclriinae. A separnte tribe, the Heptamelini, has been set up to include this unique group. ~\.bout £h'e or six species are in this genus, only one of which has been recorded from North America. Togashi (1961) revised the four species known to occur in Japan. Description of Heptameius Species BeptameZU$ och1'oleuc1l8 (StephPTIs) .Ifclicrrta. oeftrolruca. Stc-phenl', ]83.5, p. 05; Westwood, 1840, p. 54. /{1'[Jt(//IIl'lll.~ o('hrol(,lIclI.~, Hali£1ay. lS:j'1, p. GO; Cameron, 1882, p. 308; Dalla 'l'orr!~, 11"04, p. W7; Kono\\', 1!)();), p. 7!3; de )Ieijere, lOll, p. 95; Enslin, 1914, II. 2:;0; :\Illlais(', 1081, p. 1-12; ('ond(', 1084, p. 180; Benson, W85, p. 7H7; Berland, 10-l7. p. 277: B('llson. ]!):;2, p. ;'7; IJ\lrellz and Kraus, 10:;7, p. 47; Tognshi, l!)(ll, p. 2n!); Dplll'On, 1002, p. 387; Burks, 1007, p. IH. ('a(,1I0Il('lIra. dah II/O/lli Tholm;on. 11"70, [I. 271; Thomson, ]871, p. 182; Cameron, 1k7:3. 11. So,; Cameron, 1874, p. 108; Andre, 1880, p. 280; Cameron, 1882, p. 303 (=oehrolcllca St('phens). NEARCTIC SAWFUES ll. SELANDRIINAE 39 The synonymy of dahlbomi is generally accepted in literature. Female.-Length, 5.0 mm. Antenna with flagellum black, first seg ment whitish, second segment infuscate. Head black; clypeus whit ish. Thorax black with tegula whitish. Legs, including coxae, entirely yellowish white. Abdomen and sheath black. Wings hyaline. Antenna, head, and body covered 'with fine hairs. Sheath slender, truncate at apex, and dorsal and ventral margins subpa.rallel (pI. V, 63). Lancet as in plate VII, 89. Male.-Length, 4.5 mm. Paler than female, with pronotum, meso notum, mesosternum, and metapleuron reddish yellow; stigma yellow; flagellum of antenna brownish. Structure as for female. Genitalia as in plate X, 124 and 125. Type.-Stephen's type is in the British Museum (Natural History), London. Distribution.-Europe, Japan. North America: British Oolumbia: Squamish, Diamond Head Trail, 3,300 It., August 5, 1953, E. Mason (1 ~, in the Canadian National Collection). Host.-1n Europe the larva bores downward in the stems of fern'>: Ath!jri-ztm, Blechrmm" and Polypodhl1J'b (Benson, 19513). Dlscussion.-This species may be recognized by the seven- or eight segmentNl antenna, long sheath, and unusual wing venation. Only one specimen has been taken in North America.. The female is described from this one specimen, and the male is described from European specimens. On one occasion it was reported as a pest among cultivated ferns in England (Benson, 19513). 40 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE Literature Cited ANDRE, E. 1879-82. SPECIES DES HY~rENOPTEREB D'EUROPE ET D'ALGERIE. V. 1, 642 pp. Beaune (Cote-d'Or), France. ASHMEAD, W. H. 1898. CLASSIFICATION OF THE HORNTAILS AND SAWFLIES, OR THE SUBORDER PHYTOPHAGA. Canad. Ent. 30: 141-145, 177-183, 205-213, 225-232. 249-257, 281-287, 305-316. BEOHSTEIN, J. M., and SOHARFENBeRG, G. L. 1805. VOLLSTANDIGE lIlATURGESOHICHTE DER SCHADLICHEN FORSTINSEKTEN. V. 3, 1046 pp. Leipzi6. BEER, R. E. 1955. BIOLOGICAL STUDIES AND TAXOSOMlC NOTES ON THE GENUS STRONGYLO GASTER DAHLBOM (HYMENOPTERA: TENTHREDINIDAE). Kans. Univ. Sci. Bul. 37, pp. 223-249. BENSON, R. B. 1934. SO~IE MORE NEW OR LI'rTLE KNOWN BRITISH SAWFLIES (HYMENOPTERA, SYMPHYTA). Ent. Monthly Mag. 70: 201-204. 1&35. THE _\LIEN ELEMENT IN THE BRITISH SAWFLY FAUNA. Ann. App!. BioI. 22: 754-768. 1936. SOME MORE NEW OR LITTLE KNOWN BRITISH SAWFLIES (HYMENOPTERA, SY}[PIIYTA). Ent. Monthly Mag. 72: 203-207. 1938. ON THE CLASSIFICATION OF SAWFLIES (HYMENOPTERA: SYMPHYTA). Roy. Ent. Soc., London, Trans. 87: 353-384. 1939. FOUR NEW GENERA OF BRITISH SAWFLIES (HYMENOPTERA, SYMPHYTA). Ent. Monthly Mag. 75: 110-113. 1940. SAWFLIES OF THE BERKHA].[STED DISTRICT WITH A LIST OF THE SAW FLIES OF HERTFORDSHIRE AND BUCKINGHAMSHIRE, AND A St..-IWEY OF THE BIUTISH SPECIES (HYMENOPTERA: SYMPHYTA). Hertfordshire Nat. Hist. Soc. Trans. 21: 177:"'231. 1952. HYMENOPTERA (SYMI'HYTA). FAMILY TENTHREDINIDAE. In Royal Entomological Society of London, Handbooks for the Identification of Rritish Insects, v. 6, pt. 2 (b), pp. 51-137. London. 1962. HOLARCTIC SAWFLIES (HYMENOPTERA: SYMPHYTA). Brit. Mus. (Nat. Hist.) Ent. Bul. 12, pp. 381-409. BERLAND, L. 1947. FAUNE DE FRANOE. V. 47. HY]'[ENOPTERES TENTHREDOIDES. 496 pp. Paris. BRISCHKE, C. G. A., and ZADDACH, G. 1883. BEOBAOHTUNGEN UBER DIE ARTEN DER BLATT- UND HOLZWESPEN. Schr. Naturf. Gesell. Danzig 5, pp. 201-328. BmtKs, B. D. 1958. SYMPHYTA. In Krombein, K. V., and Burks, B. D., eds., Hymenop tera of America North of l'tiexico, Synoptic Gatalog, 1st Sup., U.S. Dept. Agr. Agr. Monog. 2, pp. 8-17. 1967. SYMPHYTA. In Krombein, K. V., ed., Hymenoptera of America North of Mexico. Synoptic Catalog, 2<1 SU))., U.S. Dept. Agr. Agr. )[onog. 2, pp. 15-26. CA:MERON, P. 1873. OCCURRENCE 0]' COF.NONEUltA DAHLBo~rr, THO:MS., A GENUS AND SPECIES OF TENTHREDINIDAE NEW TO BRITAIN. Ent. Monthly Mag. 10: 85. 1874. NOTES ON BRITISH TENTHREDINIDAE, WITH DESCRIPTIONS OF A NEW NEMATUS. Ent. Monthly Mag. 11: 107-109. NEARCTIC SAWFLIES II. SELANDRIINAE 41 1877. 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I. 1961. ON THE JAP,\NESE SPECIES OF THE GENUS HEPTAMELUS HALIDAY (IIYM.: SYMPHYTA). KontyO 29: 2G-l-270. VILLERS, C••T DE. 1789. O,\ROLILINNAEI ENTOlfOLOGlA. V. 3, 656 pp. Lugduni. WESTWOOD, J. O. 1840. SYNOPSIS OF THE GENERA OF BRITISH INSECTS. 158 pp. London. (His Introduction to the Modern Classification of Insects, T". II, 587 pp.) YUASA, H. 1922. A CLASSIFICATION OF THE LARVAE OF THE TENTHREDINIDAE. Ill. BioI. }Ionog. 7, pp. 1-172. ZETTERSTEDT,J. W. 1838. INSECTA LAPP(J;'iICA DESCRIPTA. HY~fENOPTERA. 868 PI>. LiJJsiae. ZHELOKIIO\'TSEV; A. N. 1951. [SURVEY OF THE PALAEARCTIC SAWFLIES OF THE SUJlFA~!ILY SELAN DRUNAE (RYlr., TENTHR.).] Moscow Gosud. Univ. Zool. Muz. Sborn. Trudy 7: 123-152. [In Russian.) ZmNGIEBL, L. 1956. BLATTWESPEN AUS mAN. )Iiinchen. Ent. Gesell. Mitt. 46: 322-326. Index This index contains both superspecific and trivial names. Valid names are in roman and synonyms in Italic. Puge Page Adelesta Ross______37 immatatrixdria______Ashmead, Paraselan _ AdelestinL ______2, 37 31 albidopictusonus______(Norton), Hemitax- _ impressatalogaster______Provancher, Str(lngy _ 8 15 alboannulatu8logaster______Rohwer, Strongy _ 17 jacobsoni Enslin, Colposelandria_ 25 amiclts N O:t~lfl, Taxonl1S ______9 Aneugmefllfll______2,25 leucoslomusceros______Rohwer, Strombo _ Aneugmenus Hartig______25 33 annulosus Nortoll, Strongy longulus Norton, Strongylogl.l.8 logaster______17 17 lu;t~~o"s~~ -P~;~~~~h~;'- Sti;~gy~: arizonensispidea______Ashmead, Eriocam- _ 6 gaster______15 Aloposelandria Enslin______25 macula (Klug), Strongylogaster_ 16 Birka Malaise______marginata Provancher, Selandria_ 23 23 melanogaster Rohwer, Strongylo- Brachythops Haliday______34 gaster______24 CaenoncltTa Thomson______melanopterus Rohwer, Strombo- 30 cinereipes Klug, Tenthredo____ _ 38 cingulata Fabricius, Tenthredo__ 31 cinguiescenster [v.l______Ross, Strongylogas _ ~i ~}:f~~~!t~~~~~~~~======morio (Fabricius), N esose 17 landria______32 Cockerellonis MacGillivray______Coiposelandria Enslin______7 multicincta Norton, Strongylo- 25 gaster______~------17 contiguater______Konow, Strongylogas _ multicinctus Hall, Hemi 12 taxonus______10 coronata Klug, Tenthredo______25 Coryna Lepeletier and Serville__ 34 Neobusarbia TakeuchL______31 curialis Cresson, Selandria______33 Nesoselandria Rohwer______31 mgncans (Norton), Strombo dahlbomi Thomson, Caenoneura_ 38 ceridea______33 decolorala Cresson, Selandria___ _ 35 nordica, n. sp., Birka______29 distans Norton, Strongylogaster_ 14 nova (Norton), Adeiesta______37 dubitatusnuB______(Norton), Hemitaxo _ occidenlalis MacGillivray, Cock 9 erellonis ______6 occidentalis Rohwer, Aneug- Epilaxonus MacGillivray______Eriocampidea Ashmead______~ oc~~I~~cJ~sp·lSt~Ph~~~): --ii~P: 26 Eustromboceros Rohwer______30 tameius______38 oxalala MacGillivray, Pseudo filicis Klug, Tenthredo______12 selandria______.______35 flavens (Klug), Brachythops___ _ 35 flavipes (Norton), Aneugmenus _ 26 pacificus MacGillivray, Strongy flavipes Takeuchi, Neobusarbia_ 31 iogaster______14 jlauilarsis Rohwl.'r, Aneugmenus_ 26 padi (Linnaeus), Aneugmenus_ 27 Paraselandria Ashmead______jloridanadria. ______MacGillivray, Selan- _ 34 26 pilosuius Rohwer, Strombo jloridella Ross, Selandria [v.l ___ _ 26 ceros ______33 filrstenbergensisdria______Konow, Selan- _ polita Cresson, Strongyiogaster_ 18 25 Polyselandria MacGillivray___ _ 25 Polyslichophagus Ashmead______12 Hemitaxonus Ashmead______7 primarius Smith, IIemitaxonus_ 11 HeptamelinL______2,38 primatiuusylogaster MacGillivray,______Strong_ Heptamelus HaHday______38 23 47 48 TECHNICAL BULLETIN 1398, U.S. DEPT. OF AGRICULTURE Page Page Prototaxonus Rohwer______12 soriculatus Provancher, Strongy Pseudoselandria MacGillivray__ 34 logaster_------ 21 Pseudotaxonus A. Costa______12 stramineipes Klug, Tenthredo___ 27 pullatus MacGillivray, Thrinax_ 15 Stromboceridea Rohwer______33 Strongylogaster Dahlbom______12 remota Rohwer, Strongylogaster 19 StrongylogasterinL______rufigastra (Kincaid), Strongylo 2,6 gaster.______struthiopteridis Forsius, Sahl 20 bergia______7 rujipe8 Harrington, Taxonus____ 15 rufocaudatus,boceros______n. sp., Eustrom tacita (Norton), Strongylogaster_ 22 30 Thrinax Konow______rufopectu8 Rohwer, Hemitaxonus 12 9 tibialis Cresson, Strong~logaster_ 23 Sahlbergia Forsius ______7 tuberculiceps Rohwer, trongylo gaster______'. ______24 scapularisTenthredo I..epeletier______and Serville, 34 tYP'iCU8 Rohwer, Prototaxonus_ 23 scutellatus, n. IIp., Ancugmenus_ 28 Selandridea RohweL______unicu8 Norton, Strongylogaster_ 22 34 urbi8 Ross, Sellmdria ______SelandriinL______2,34 27 seminigra Haliday, Brachythops_ 34 vanduzeei Rohwer, Selandridea_ soriculatipes Cresson, Strongy 35 logaster______21 wuestneii (Konow), Brachythopa 36 Tecbnlcal Bulletin 1308. V.S. Dept. of Agriculture PLAT. I 2 Forewing (top) and hlndwing of Strongyloga8ter mlllticincta (1) and Eriocam pi(lea arizolleIl8i.~ (2). [a=anal crossvein; A=anai cell; m-clI=medlocubitai eross\'ein; J[=me<1ia; lr-m, 2r-m=first an<1 second rlldlomedlai crossveins; ll.y==radia! seetor; RIl+J1=rndial 'sec.:tor and mediI!; Sc+R=;,'Ubcosta amI radius.] Technleal Bulletin 1398. U.S. Dept. of Agriculture PLATE n 5 Forewing (top) and hindwing ot Bracllythops flavens (3) and Heptamcluij oohrolcucus (4) ; forewing of. Adclcstan.(},lla (5). [cll-a=cubital anal crossvein; M=media; lr-m=first rlldiomcdilll crossveln; Rs=rndlal sector; Sc+R=sub costa and radius.] Technical BuUetli.1398, U.S. Dept. of Agriculture PUT& III 11 fb,. ~'2 \~15 ~16 ~17 28 29 30 20 21 22 23 2.t Tarsal claws of Erlocampidea arlzonemri8 (6), StronglllogaBter di8tans (7), Aneugmenus padi (8), Birka nordica (9), Eustromboceros ru!ocaudatus (10), Stromboceridca nigricans (11), and Heptamelus ochroleucus (12) ; clypeus of StronUlIlogaster distans (13), S. impressata (14), S. polita (15), Aneugmenus padi (16), and A. /lavipes (17); clypeus, lateral, of Stromboeeridea nigricans (18) ; mesopleuron of Brachllthops /lavcns (19); antennae of Eriocampidea urizonensil1 (20), Hcmita:z:onus dubitatus (21), Strong1l1ogaster multicincta (22), Siromboccridea niurlcurnI (23), Eustromboceros rutocaudatus (24), Birku nordicu (25), Ncsol1elandria morlo (26). Aneugmenus /lavipcB (27). Brachllthops /lavenB (28). Adelesta nova (29), and Heptamelus ochroleucus (30). Technical BulletIn 1308, U.S. Dept. 01 Agncultu!'e PLA.TlII IV 37 m'" ~ .. 39 41 ~-~7!l! 42 43 Female sheaths: Lateral (31) an~ posterior (32) of Strongylog(lster 1nlllticincta; posterior (38) of S. diatana; dorsal (34) of S. mUlticincta,' lateral (35), pos terior (36), and dorsal (37) of S. improBata,' laternl (38) and dorsal (39) of S. soriclllatipc8; lateral (40) nnd dorsal (41) of S. tibialis; lnteral (42) and dorsal (43) of S. poUtQ;,'lllteral (44) and dorsal (45) ofS. tacita; lateral (46) and dorsal (47) of S. macula; lateral (48) and dorsal (49) of S. rllfigastra; lateral (50) of S. rCTrWta. Technical Bulletin 1398. U.S. Dept. of Agriculture PLATE V -. ~~ ) ...--.::: " ", .. , 64 7.66 67 ~\ \\ 69 Female sheaths: Lateral (51) and dorsal (52) of Aneugmenu8 fiavipe8; dorsal (53) of A. padi; lateral (54) and dorsal (55) of A. 8cntellatu8; lateral (56) and dorsa! (57) of Birka nordica; lateral (58) of Ne808elandria morio; lateral (59) ofEriocampitleaarizonensis;lateral (60), posterior (61), and dorsal (62) of Brach/It/IOI18 /lavcns; laternl (63) of Heptamelu8 ochroleltClI8; lateral (64) of HcmitaJ;onll.y aZbidopictlls; lateral (65) of H. dlloitatIl8,:lateral (66) of H. mlllticinctrt.~; lateral (67) of H. primariu8. Female lancets of Strongyloua.~ter di8tans (68) and S. multicincta (69). Technical Bulletin 1398, U.S. Dept. of Agriculture PLATS VI l!'emale lanc-ets of Strongylogaster impressa.ta (70), S. 80riculatipes (71), S. tibialis (72), S. macula (7~), S. rufigastra (74), S. 'remota (75), S. tacita (76), S. polita (77), Aneugmenus {lavipes (78), A. padi (79), A. scuteUatus (80), Birka nordica (81), and Nesoselandria mo-rio (82). ;:-. Technical Bulletin 130S, U.S. Dept. or Agriculture PLATE VII 89 90 -----~ Female lancets of Strc;m.bO'ceridea nigrica.n., (83), Eriocampidca arizO'lIemJia (84), JTcmitaxO''/lII.~ (/~bilT(;pictll,Y (85), 1I. (illbitatll.$ (86), 1I. tnulticillct1l8 (87), If. prima rill.~ (HS), J[(~{Jtamelll.Y O'ehrO'1cIICll8 (89), Brad/.llthops f!aver..{J (90), and H. '1/.·lIc.~tn('fi, (91); harpe lind parapenis (1)'2) and penis valve (93) of StronU/llo!la.~tcr (li.'JfIl1/8,· harpe and parapenis (94) and penis valve (1)5) of S.mlllticil/eta. Technical Bulletin 1398. U.S. Dept. ot A.griculture PLATE: VIU ) I ~~ 100 llOI o 106 107 Harpe and parapenis (00) and penis vnlve (9;) ot 8trongylogastcr soriolllatipcs; harpe and parapenis (98) and penis valve (99) of 8. tibialis; harpe and lIara penis (100) and penis valve (101) of S. imprClJllata;harpe and l)llrapenis (1(2) and penis valve (100) ot S. tacita.; harpe and parapenis (1M) and penis vaive (105) of S. tllberl.IUliceps; hypandrium ot S. tllbcroulWepIl (106) and S. tacita (107). Technical BuUetin 1398, U.S. Dept. of Agriculture PLATS Ill: " ···.". 0 118 119 Harpe and l'arapenis (108) and penis valve (109) of Strcl'ngyloga-llter sp.; harpe and parapenis (110) lind penis vah'c (111) of Anellgmelf.l18 {la'VipC8; hat'pc anll llarapenis (112) and {INtis vlllve (113) of "L .~eu.tcll(ltIl8; harpe lind pllra l)('nis (114) Ilnd penis vlllve (115) of Birkcl nonlica; hurpe und parupcnis (116) and penis vain' (117) of i.V(,.~o8clandria morio; hllrpe and parapenis (llS) Ilnd penh; valve (111) of Ew,tromllOcero8 rlllofX/./Idatlls, Technical Bulletin 1398. U.S. Dept. of Agriculture 127 Harpe and parapenis (120) and penis valve (121) of Brachythopsl';,a'Vens; harpe and parapenis (122) and penis valve (123) of B. wuestneii; harpe and para peni:! (124) and penis valve (125) of Heptamel1l-8 ochroleucus; harpe Ilnd [larnpenis (126) 'and penis valve (127) of Hemitaa:onus a.Z1ti4op-ictus; harpe nnd pnrapenis (128) and peni:! valve (129) of H. dtlbitatll8; harpe and parapenis (130) and penis valve (131) of H. multicinctus; harpe and parapenis (132) and penis valve (133) of 11. primarills. ), ., ," 'I ,"! I , . ; ~ I \ 1 J ..