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The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Comissão Organizadora
ISSN 2175-7720 Paleontologia 2 Livro de Resumos ISSN 2175-7720 Paleontologia 3 Livro de Resumos Comitê Editorial: Valéria Gallo Hilda Maria Andrade da Silva Capa e Identidade Visual: Mapinguari Design Projeto gráfico interno e editoração: Rafael Fernandes Lopes da Silva XXI Congresso Brasileiro de Paleontologia: A paleontologia e os eventos globais. 2009, Belém, Pará, Brasil. ISSN: 2175-7720 1. Paleontologia 2. Geociências 3. Congresso Brasileiro de Paleontologia Paleontologia 4 Sobre a Logomarca A Logomarca do XXI Congresso Brasileiro de Paleontologia O gênero Orthaulax é um representante extinto da família Strombidae, endêmico da província paleobiogeográfica Caribeana, vivente entre o Oligoceno Superior e o Mioceno Inferior. Do ponto de vista paleoambiental teria vivido em ambiente marinho de águas rasas, quentes, límpidas, agitadas e com salinidade normal. O processo envolvente da sua ultima volta tornou a concha bastante sólida e maciça, permitindo que habitasse ambiente de grande agitação, como os biohermas, pequenas edificações recifais. Na Formação Pirabas, a espécie Orthaulax pugnax (Heilprin, 1887) foi reconhecida por Maury (1925), e corroborada em pesquisas subseqüentes. Estudos realizados por Cândido Simões Ferreira, delimitaram nos calcários aflorantes no litoral nordeste do Estado do Pará e noroeste do Estado do Maranhão, uma zona caracterizada por elementos estenobiônticos, típicos de recifes de corais. A espécie O. pugnax, associada com algas coralíneas, corais hermatípicos e equinóides regulares são as formas mais características que contribuíram para a edificação do bioherma. Assim, este gastrópode constitui-se em um elemento importante da Formação Pirabas, por ter sido o primeiro táxon utilizado para datar esta unidade litoestratigráfica como oligo-miocênica, bem como seu decisivo papel para correlação com outras unidades sincrônicas da Província Biogeográfica Caribeana, permitindo delimitar no norte do Brasil, a sua extremidade sul. -
Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships
438 Article 438 by Saswati Bandyopadhyay1* and Sanghamitra Ray2 Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships 1Geological Studies Unit, Indian Statistical Institute, 203 B. T. Road, Kolkata 700108, India; email: [email protected] 2Department of Geology and Geophysics, Indian Institute of Technology, Kharagpur 721302, India; email: [email protected] *Corresponding author (Received : 23/12/2018; Revised accepted : 11/09/2019) https://doi.org/10.18814/epiiugs/2020/020028 The twelve Gondwanan stratigraphic horizons of many extant lineages, producing highly diverse terrestrial vertebrates India have yielded varied vertebrate fossils. The oldest in the vacant niches created throughout the world due to the end- Permian extinction event. Diapsids diversified rapidly by the Middle fossil record is the Endothiodon-dominated multitaxic Triassic in to many communities of continental tetrapods, whereas Kundaram fauna, which correlates the Kundaram the non-mammalian synapsids became a minor components for the Formation with several other coeval Late Permian remainder of the Mesozoic Era. The Gondwana basins of peninsular horizons of South Africa, Zambia, Tanzania, India (Fig. 1A) aptly exemplify the diverse vertebrate faunas found Mozambique, Malawi, Madagascar and Brazil. The from the Late Palaeozoic and Mesozoic. During the last few decades much emphasis was given on explorations and excavations of Permian-Triassic transition in India is marked by vertebrate fossils in these basins which have yielded many new fossil distinct taxonomic shift and faunal characteristics and vertebrates, significant both in numbers and diversity of genera, and represented by small-sized holdover fauna of the providing information on their taphonomy, taxonomy, phylogeny, Early Triassic Panchet and Kamthi fauna. -
A Beaked Herbivorous Archosaur with Dinosaur Affinities from the Early Late Triassic of Poland
Journal of Vertebrate Paleontology 23(3):556±574, September 2003 q 2003 by the Society of Vertebrate Paleontology A BEAKED HERBIVOROUS ARCHOSAUR WITH DINOSAUR AFFINITIES FROM THE EARLY LATE TRIASSIC OF POLAND JERZY DZIK Instytut Paleobiologii PAN, Twarda 51/55, 00-818 Warszawa, Poland, [email protected] ABSTRACTÐAn accumulation of skeletons of the pre-dinosaur Silesaurus opolensis, gen. et sp. nov. is described from the Keuper (Late Triassic) claystone of KrasiejoÂw in southern Poland. The strata are correlated with the late Carnian Lehrberg Beds and contain a diverse assemblage of tetrapods, including the phytosaur Paleorhinus, which in other regions of the world co-occurs with the oldest dinosaurs. A narrow pelvis with long pubes and the extensive development of laminae in the cervical vertebrae place S. opolensis close to the origin of the clade Dinosauria above Pseudolagosuchus, which agrees with its geological age. Among the advanced characters is the beak on the dentaries, and the relatively low tooth count. The teeth have low crowns and wear facets, which are suggestive of herbivory. The elongate, but weak, front limbs are probably a derived feature. INTRODUCTION oped nutrient foramina in its maxilla. It is closely related to Azendohsaurus from the Argana Formation of Morocco (Gauf- As is usual in paleontology, with an increase in knowledge fre, 1993). The Argana Formation also has Paleorhinus, along of the fossil record of early archosaurian reptiles, more and with other phytosaurs more advanced than those from Krasie- more lineages emerge or extend their ranges back in time. It is joÂw (see Dutuit, 1977), and it is likely to be somewhat younger. -
Supplementary Information Niche Partitioning
Supplementary Information Niche partitioning shaped herbivore macroevolution through the early Mesozoic Suresh A. Singh1*, Armin Elsler1, Thomas L. Stubbs1, Russell Bond1, Emily Rayfield1 & Michael J. Benton1. 1. School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS8 1TQ, UK. *email: [email protected] Contents: Supplementary Note 1..………………………………………………………………………...………3 Alternative data transformation methods and results Supplementary Table 1…………………………………………………………………………………5 Functional character descriptions and functional utility to feeding behaviour. Supplementary Table 2…………………………………………………………………………………7 Character loadings for functional principal component (fPC) scores in main text. Supplementary Table 3…………………………………………………………………………………8 Character loadings for functional principal component (fPC) scores in supplement. Supplementary Table 4…………………………………………………………………………………8 Symmetric Procrustes analysis results. Supplementary Table 5…………………………………………………………………………………9 Internal validation statistics for different cluster configurations. Supplementary Table 6…………………………………………………………………………………9 External validation statistics for different cluster configurations. Supplementary Table 7………………………………………………………………………………..10 Feeding functional group conflicts within early Mesozoic assemblages. Supplementary Table 8………………………………………………………………………………..11 Total herbivore shape and functional disparity at stage level. Supplementary Table 9…………………………………………………………………………..……12 Archosauromorph shape and functional disparity -
Geological Explorations in the Triassic of Southern Tanzania
Supradapedon revisited: geological explorations in the Triassic of southern Tanzania Max C. Langer1, Átila A.S. da Rosa2 and Felipe C. Montefeltro3 1 Laboratório de Palentologia, FFCLRP, Universidade de São Paulo, Ribeirão Preto, SP, Brazil 2 Laboratório de Estratigrafia e Paleobiologia, Universidade Federal de Santa Maria, Santa Maria, RS, Brazil 3 Departamento de Biologia e Zootecnia, FEIS, Universidade Estadual Paulista, Ilha Solteira, SP, Brazil ABSTRACT The upper Triassic deposits of the Selous Basin in south Tanzania have not been prospected for fossil tetrapods since the middle of last century, when Gordon M. Stockley collected two rhynchosaur bone fragments from the so called ``Tunduru beds''. Here we present the results of a field trip conducted in July 2015 to the vicinities of Tunduru and Msamara, Ruvuma Region, Tanzania, in search for similar remains. Even if unsuccessful in terms of fossil discoveries, the geological mapping conducted during the trip improved our knowledge of the deposition systems of the southern margin of the Selous Basin during the Triassic, allowing tentative correlations to its central part and to neighbouring basins. Moreover, we reviewed the fossil material previously collected by Gordon M. Stockley, confirming that the remains correspond to a valid species, Supradapedon stockleyi, which was incorporated into a comprehensive phylogeny of rhynchosaurs and found to represent an Hyperodapedontinae with a set of mostly plesiomorphic traits for the group. Data gathered form the revision and phylogenetic placement of Su. stockleyi helps understanding the acquisition of the typical dental traits of Late Triassic rhynchosaurs, corroborating the potential of hyperodapedontines as index fossils of the Carnian-earliest Norian. -
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0 www.journalsofindia.com February 2021 1. 0 www.journalsofindia.com February 2021 0 www.journalsofindia.com February 2021 0 www.journalsofindia.com February 2021 0 www.journalsofindia.com February 2021 ENVIRONMENT ................................................................................................................................................................ 6 1. ENHANCED TRANSPARENCY FRAMEWORK OF PARIS PROTOCOL ................................................................................... 6 2. TAJ TRAPEZIUM ZONE (TTZ) ............................................................................................................................................. 7 3. MICROBIAL DECOMPOSER CAPSULE ............................................................................................................................... 7 4. ACTION PLAN IMPROVING AIR QUALITY ......................................................................................................................... 8 5. INTERNATIONAL SOLAR ALLIANCE (ISA) .......................................................................................................................... 8 6. UNESCO GLOBAL GEOPARKS ........................................................................................................................................... 9 7. ARCTIC AMPLIFICATION ................................................................................................................................................... 9 8. MADHUCA DIPLOSTEMON ............................................................................................................................................ -
Environment of the Upper Omingonde Formation
The postcranium of the carnivorous cynodont Chiniquodon from the Middle Triassic of Namibia and the palaeo- environment of the Upper Omingonde Formation Helke Brigitte Mocke A Dissertation submitted to the Faculty of Science, University of the Witwatersrand, in fulfilment of the requirements for the Degree of Master of Science Johannesburg, 2015 Declaration I declare that this thesis is my own, unaided work, except where specific acknowledgement is made to the work of others. It is being submitted for the Degree of Master of Science at the University of the Witwatersrand, Johannesburg, and has not been submitted before for any degree or examination at any other university. I would like to dedicate this thesis to my parents, who have inspired me throughout my growing up years to always follow my passion. †Hartmut Sören Wagner Bridget Ann Wagner Abstract The Chiniquodontidae is a family of Triassic carnivorous cynodonts well represented in the Middle-Upper Triassic of Argentina and Brazil. Chiniquodontids were more recently discovered in Madagascar and central Namibia, representing the only record of the family outside South America. The Namibian specimen was discovered in the Upper Omingonde Formation and is represented by the skull and a partial skeleton. The new chiniquodontid was identified as Chiniquodon and is diagnosed by the postcranial characteristics identified; a strong bend in the proximal portion of thoracic ribs, reduced curvature of the clavicle, although this may be due to deformation, robustness of the neck of the ilium, differences in the angulation between the edge of the posterior lamina of the ilium and the margin of the neck, and a large ischium, which is more than twice the size of the pubic plate. -
The Karoo Basins of South-Central Africa
Journal of African Earth Sciences 43 (2005) 211–253 www.elsevier.com/locate/jafrearsci The Karoo basins of south-central Africa O. Catuneanu a,*, H. Wopfner b, P.G. Eriksson c, B. Cairncross d, B.S. Rubidge e, R.M.H. Smith f, P.J. Hancox e a Department of Earth and Atmospheric Sciences, University of Alberta, 1-26 Earth Sciences Building, Edmonton, Alta., Canada T6G 2E3 b Geologisches Institut, Universitat zu Koln, Zulpicher Str. 49, 50674 Koln, Germany c Department of Geology, University of Pretoria, Pretoria 0002, South Africa d Department of Geology, University of Johannesburg, P.O. Box 524, Auckland Park, Johannesburg 2006, South Africa e Bernard Price Institute for Palaeontological Research, University of Witwatersrand, PO Wits, Johannesburg 2050, South Africa f Department of Karoo Palaeontology, Iziko: South African Museum, Cape Town, South Africa Received 1 June 2004; accepted 18 July 2005 Available online 25 October 2005 Abstract The Karoo basins of south-central Africa evolved during the first-order cycle of supercontinent assembly and breakup of Pangea, under the influence of two distinct tectonic regimes sourced from the southern and northern margins of Gondwana. The southern tec- tonic regime was related to processes of subduction and orogenesis along the Panthalassan (palaeo-Pacific) margin of Gondwana, which resulted in the formation of a retroarc foreland system known as the ‘‘main Karoo’’ Basin, with the primary subsidence mechanisms represented by flexural and dynamic loading. This basin preserves the reference stratigraphy of the Late Carboniferous–Middle Jurassic Karoo time, which includes the Dwyka, Ecca, Beaufort and Stormberg lithostratigraphic units. North of the main Karoo Basin, the tec- tonic regimes were dominated by extensional or transtensional stresses that propagated southwards into the supercontinent from the divergent Tethyan margin of Gondwana. -
Enigmatic Dinosaur Precursors Bridge the Gap to the Origin of Pterosauria
Published in "Nature 588(7838): 445–449, 2020" which should be cited to refer to this work. Enigmatic dinosaur precursors bridge the gap to the origin of Pterosauria Martín D. Ezcurra1,2ಞᅒ, Sterling J. Nesbitt3, Mario Bronzati4, Fabio Marco Dalla Vecchia5,6, Federico L. Agnolin7,8, Roger B. J. Benson9, Federico Brissón Egli7, Sergio F. Cabreira10, Serjoscha W. Evers9,11, Adriel R. Gentil7, Randall B. Irmis12,13, Agustín G. Martinelli1, Fernando E. Novas7, Lúcio Roberto da Silva10, Nathan D. Smith14, Michelle R. Stocker3, Alan H. Turner15 & Max C. Langer4 Pterosaurs were the first vertebrates to evolve powered flight1 and comprised one of the main evolutionary radiations in terrestrial ecosystems of the Mesozoic era (approximately 252–66 million years ago), but their origin has remained an unresolved enigma in palaeontology since the nineteenth century2–4. These flying reptiles have been hypothesized to be the close relatives of a wide variety of reptilian clades, including dinosaur relatives2–8, and there is still a major morphological gap between those forms and the oldest, unambiguous pterosaurs from the Upper Triassic series. Here, using recent discoveries of well-preserved cranial remains, microcomputed tomography scans of fragile skull bones (jaws, skull roofs and braincases) and reliably associated postcrania, we demonstrate that lagerpetids—a group of cursorial, non-volant dinosaur precursors—are the sister group of pterosaurs, sharing numerous synapomorphies across the entire skeleton. This finding substantially shortens the temporal and morphological gap between the oldest pterosaurs and their closest relatives and simultaneously strengthens the evidence that pterosaurs belong to the avian line of archosaurs. Neuroanatomical features related to the enhanced sensory abilities of pterosaurs9 are already present in lagerpetids, which indicates that these features evolved before flight. -
10-537-548-GVR PRASAD.Pmd
Proc Indian Natn Sci Acad 82 No. 3 July Spl Issue 2016 pp. 537-548 Printed in India. DOI: 10.16943/ptinsa/2016/48466 Review Article An Overview of Recent Advances in the Mesozoic–Palaeogene Vertebrate Paleontology in the Context of India’s Northward Drift and Collision with Asia G V R PRASAD1,* and S BAJPAI2 1Department of Geology, Centre for Advanced Studies, University of Delhi, Delhi 110 007, India 2Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow 226 007, India (Received on 10 April 2016; Accepted on 20 May 2016) This review article summarizes the recent work done in India on fossil vertebrates from the Mesozoic-early Paleogene interval, with focus on the past five years. Advances made during this period highlight the importance of India’s vertebrate fossil record in our understanding of the origin and evolutionary history of several vertebrate groups and the dynamics of intercontinental faunal dispersal and vicariance, especially in the context of India’s former position in the Gondwanaland and its subsequent separation from the different Gondwana landmasses as it moved northward to finally collide with Asia in the early Paleogene. These studies have led to a better understanding of the mode of evolution of tetrapods on the drifting Indian plate and provide independent constraints to test the traditional models favouring India’s physical isolation from all other landmasses, especially during the late Cretaceous phase of its northward drift. Recent data points to faunal interchanges between India, Africa and Europe at or near the K-Pg boundary and also suggests that the Indian landmass was the centre of origin/early evolution for several orders of modern mammals, including cetaceans (whales), perissodactyls (horses, tapirs) and primates. -
The Oldest North American Prosauropod, from the Upper Triassic Tecovas Formation of the Chinle Group (Adamanian: Latest Carnian), West Texas
Heckert, AB" and Lucas, S.G., cds., 2002, Upper Triassic Stratigraphy and Paleontology. New Mexico Museum of Natural History and Science Bulletin No. 21. 249 THE OLDEST NORTH AMERICAN PROSAUROPOD, FROM THE UPPER TRIASSIC TECOVAS FORMATION OF THE CHINLE GROUP (ADAMANIAN: LATEST CARNIAN), WEST TEXAS SUSAN K. HARRIS, ANDREW B. HECKERT, SPENCER G. LUCAS and ADRIAN P. HUNT New Mexico Museum of Natural History, 1801 Mountain Road NW, Albuquerque, 87104-1375 Abstract-An isolated tooth from the Tecovas Formation of the Chinle Group, Crosby County, West Texas, represents the oldest definitive record of a prosauropod dinosaur from North America. Its age is Adamanian (latest Carnian, approximately 225 Ma) based on vertebrate biostratigraphy, palynostratigraphy and other data. It is clearly distinguished from isolated ornithischian teeth by a combination of prosauropod dental synapomorphies. The addition of this tooth to other recent discov eries of late Carnian prosauropod material from Brazil and Madagascar, as well as the Moroccan Azendohsaurus, indicates a Pangea-wide distribution for prosauropods near their oldest occurrences. Keywords: Carnian, prosauropod, Tecovas Formation, tooth INTRODUCTION Kalgary conglomerate The Prosauropoda are poorly represented in the Upper Tri sandstone assic of North America. Among the oldest known prosauropod ~ siltstone body fossil specimens from the continent is a single dorsal cen ~.c- mudstone/shale trum from the Revueltian (early-middle Norian) Bull Canyon ; 6 =-=; limestone Formation of New Mexico (Hunt et al., 1998) and part of the holo [/=::3 fossil bone type of the putative ornithischian Technosaurus smalli Chatterjee, 1984 (Sereno, 1991). All other Triassic prosauropod records in North America are restricted to tracks.