The malacologicalsocietymalacological society of Japan

Rn VENUS (Jap. Jour. Malac.) VoL 43, No, 4 (1984):301-304

pP as jij: } ti ,v jv- I Ii}o 1 MK*

us Z･wt tw 4

($nax[11flilliJ.fieVft71eqt}htr) (#ipaTiF*waenig-tNMD

A New Clausiliid Genus GTanainenia for Steatonenia

miTifoca Chen & Gao, 1982 (: )"

Hiroshi MINATO

(Kumano Senier High School, Wakayama-ken, Japan) and

De-niu CHEN

(Institute of Zoology, Academia Siniea, China)

Chen and Gao (1982) described a Chinese Clausiliid as Steatonenia mirifica,

which obtained from the Guangxi Zhung Autonomous Region, China. This

species is characterized by having an abruptly constricted and tortuous body

, forming a narrow neck with wrinkles. The structures of plica and

were not deseribed in the original description. We here reviewed

the structures of plica and clausilium and other external and internal chaTacters

"Stea･tonenia", to eompare with which is a Peruvian clausiliid genus, and con-

elude that the present species requires a new genus, Grandinenia.

Before going further, the authors wishes to express their hearty thanks

to Dr. Takashi Okutani, Professor of the Tokyo University of Fisheries, for

his kindness in reading the original manuscript.

Family

Grandinenia gen. noy.

Type species: Steatonenia mirifiea Chen & Gao, 1982

Shell thick, obesely fusiform, with about 8 whorls, of which first two lustrous with the conical upper part of the . Penultimate whorl widest,

mueh inflated, while abruptly eonstrieted, tortuous, forming a

narrow neek, with coarse and irregular wrinkles. Principal plica and lunella

well developed. Clausilium narrow, tapering toward a slenderly acute end and

not excised near the filament. tt t tt tt tttttt ttt ttttttttt - ..tmN * Received July 13; revised August 21, 1984.

NII-Electronic Library Service The malacologicalsocietymalacological society ofJapanof Japan

302 VENUS: Vol, 43, No. 4 (1984)

Grandinenia･ 7n･irifica･ (Chen & Gao, 1982)

(Figs. 1-7)

1982 Steatonenia mirifi.ea Chen & Gao. Acta. Zootaxonomica Sinica, 7(1): 32-34.

Shell mediurn in size, sinistral, thick, obesely fusiform and vDiametervellowish-brown

in eolor. Surface with faint traces of fine striation and dull. about

2 tirnes height and widest at the penultimate whorl. Body whorl abruptly

3 4[

[

-`-A

, StL /7 K 6

Figg. 1-7. Grandinenia mirijica (Chen & Gao, 1982) fi tfSz y;' L-ij'HiiL･ (ptfi) [Scale == 1mm] 1. Ventral view; 2. Right lateral view (H 21mmXB 10mm) ; 3-4. Clausi- lium ; 5. ; 6. Plica and lunella; 7. Side view of mouth, with the palatal area removed to show internal folds and the clausilium. A: Aperture; C : Clausilium; IL : Inferior lamella; L: Lunella ; PP : Principal plica ; SCL : Subcolurnellar lamella ; SL : Superior Iamella ; SPL: Spiral larnella.

NII-Electronic Library Service The malacologicalsocietymalacological society of Japan

Minato'Chen:A New Clausiliid Genus Crandinenia 303

somewhat axial constricted, tortuous, forming a narrow neck with irregular number, convex, ribs, wrinkled behind the outer of aperture. Whorls 81/{] in

with shallow . wit,h 2 whorls, smooth, lustrous, pale yellowish continuous, and mamillated. Aperture nearly eircular in outline and and subeolumellaT well expanded, whitish glossy at the edge. Superior, inferior thin, lamellae visible in front view of the aperture. Superior lamella vertical,

rather strong and continuous with the spiral lamella. Inferior lamella developed, emerging, conspieuous and gratly spiral within. Subcolumellar lamella weakl.v from the but not reaehing the peristome. Prineipal pliea lateral, running elosely con- middle of the right side nearly to the lip, where its inner end tiguous to the upper end of the lunella. Lunella dorso-lateral, well developed and and strongly arched. Clausilium occupies the whole space between lunella

subeolumellar lamella. Subeolumellar lamella narrow, spindle-shaped, parallel-

sided, aeute at the distal end and not excised on the columellar side of the filament. was reported by Chen & Gao (1982). Genitalia are unknown. Type Loeality: Guangxi Zhunang Automous Region, China. Type specimen: Height 22mm, breadth 11mm (Holetype in Institute of

Zoology, Aeademia Sinica) .

Paratypes exatnined: (1) Height 21 mm, breadth 10 mm; (2) Height 18.6 mm, breadth 9.3mm; (3) Height 18.6mm, breadth 8,8 mm; C4) Height 19.8 mm, breadth 8.5mm; (5) Height 20,lmm (deeollated), breadth 10.3mm; all

from the type loeality. Peruvian Re?na,rks: In the general profile, this speeies is easily separated from

speeies, Stea-tonenia･ eooki. (Pilsbry, 1919) in the following shell characters: (1) smaller size, (2) spire consisted of 8YEi whorls, (3) emerging subeolumellar lamella, (4) abruptly constricted and tortuous body whorl, forming a narrow and wrinkled neck, (5) fine striation on shell surface, and (6) ¢ lausilium acuminated at the distal end. But, no species eomparable to this speeies with

such a bizar're shell morphology has been known from the Asia.

s ts

C･rk, iL- Chen & Gao C1982)Vcz.ltovCtt'd'ewS2zt;ilil':eMzzrk SteatoneniamiriLrica t`Dua"ZeLAL' 'Jy'(fiCi}6 rk SteatoneniaPilsbry, 19264xfo'CrCL,k. ?llEc･tlEElldi"a) Chen t#}lc?liptii tw1tsteAt -E･ pm lc wt .ti･I- Lk b L- 6, )ts fi IC S{t L-C refM ts it i7.,. t. -･C g 2 L i 5 #. ma wa .ee l}f elk. Zii as ÷t JR' :E & Vtl[: l･t x-C 'd. ( L ikML, eefiP va fF OfiY ma･- DL ts k Lk.

Grandinenia gen. nov. .a tfJ?; i)' viS" tt,vM (Xi'Mi) rc'N1fiop2,NaiiVtifllh;th fiecosg<, ijE'SIS'e･=･ ,HEIJ71Uk{ihutffr;. mal)4tzfitYsJti, -a).E:h"･ILFJsuJf 6. tth:tasNemptttJA'<, k<"E2i.6. ff;ne}mJitieetceqvaL, t･nV.tht.6. scizzex.an<, lfU

ongB (,ttigh{Hti) trcMtht j: lkblfitL･.

NII-Electronic Library Service The malaoologioalmalacological societysooiety of Japan

304 VENUS ： VoL 43 ， No ．4 （1984 ）

ー 本 新 属 は ペ ル Steatonenia の 産 模 式 種 S ． cooki （Pilsbry，1919） と は ，〔ユ）貝 殻 が 小 形 ，（2）螺 層 が い 多 （約 8 層 ）， 〔3）唐 突 に し つ 緊 縮 ， ね じれ た 体 層 を も こ と ， 殻 表 に 粗 い 波 状 肋 が な い こ と ， 閉 弁 が 細 長 い こ と な ど で 区 別 で き る 。

模 式種 ： Steatonenia mirifica Chen ＆ Gao 1982 ． ， ： 22mm 模 式 慓 本 殻 高 ，殻 径 11mm （中 国 科 学 院 動 物 研 究 所 蔵 ） ． ． 模 式 産 地 ： 中 国 広 西 壮 族 自治区 貴 県樟 木 公 社 良 占大 隊 （石 灰 岩 地 ）

References

Chen D ・and ・ ・ G … J・1982 ・ On a n ・w ・ ・・i… fClau ・ ・ ・ ・α・t・ p iliid ， St n ・n iα 跚 ぜ短 加 ．

ノ40‘α Zoot − α xonomica Sinic α ，7 （1）： 32 34 ．

LQosjes ， F ． E ． and Loosjes −van ． ． Bemmel ， A C W ．1984 ． On a collection of Peruvian

Neniinae ： Gastropoda ： Clausiliidae check ． （ ）， with a 1ist and a provisional key to all the Peruvian species known ． Zoologische y θ沈 ｛鵬 （telingen 212 ： 1− 38 ． ， （ ） Pilsbry ， H ． A ．1919 ． Neni α coo んi n ． sp ． Nautilus ： 80 −81 ， 32 （3 ） ． Pilsbry ， H ．A ． 1926 ． South Ameriean land and freshwater mollusks ： Notes and

description， VI ． P ア oo ． Ac α d ． 祕 ． 痂 〜α ． 78 ： 1−15 N P ， ．

一 NII-ElectronicN 工 工 Eleotronio Library Service