u 17 December 1993 PROC. BIOL. SOC. WASH. 106(4), 1993, pp. 705-713 ON A NEW GENUS AND SPECIES OF XANTHID (CRUSTACEA: : BRACHYURA) FROM CHESTERFIELD ISLAND, CORAL SEA

Peter K. L. Ng

Abstract.—A new genus and species of xanthid crab, Cranaothus deforgesi, is described from Chesterfield Island. Cranaothus appears to be closely related to Paramedaeus, Metaxanthops, Macromedaeus, Medaeops, Neoxanthops, Glyptoxanthus and Lipaesthesius, but differs in the form and sculptures on the carapace, as well as structures of the sternum, male abdomen and male first pleopod. The larger cheliped of Cranaothus also possesses a specialized basal cutting tooth on its dactylus.

A collection of Brachyura from Chester­ eroded vermiform granulated ridges; front field Island was deposited in the Museum distinctly produced; lobes truncatiform, national d'Histoire naturelle (MNHN), Par­ separated by deep fissure extending to epi­ is, by ORSTOM (Institut Francais de Re­ gastric region; external orbital angle low, in­ cherche Scientifique pour le Developement distinct, not clearly demarcating beginning en Cooperation). Among these specimens of anterolateral margin, joining series of was an interesting crab from Chesterfield smaller granules curving gently downwards Island with several peculiar features distin­ below supraorbital margin, across subor­ guishing it from all other known genera and bital region and towards buccal cavity; an­ species in the family MacLeay, terolateral margin not lobulated or toothed, 1838 (sensu Guinot 1978). anterior % arcuate, posterior xh subparallel The description of this new genus and to median longitudinal carapace axis; pos­ species forms the text of the present paper. terolateral margins gently concave. Ster- Abbreviations G1 and G2 are for the male nites 2-4 broad, sternal suture 1 and 2 com­ first and second pleopods respectively. plete, sternal sutures 2 and 3, and 3 and 4 Measurements are reported in millimeters, interrupted medially. Chelipeds distinctly in the sequence carapace width by carapace asymmetrical, fingers sharp, without pig­ length. mentation, larger cheliped with pronounced molariform basal cutting tooth on dactylus. Lateral margins of fused male abdominal Systematic Account segments 3-5 entire, continuous; segment 7 semicircular, lateral margins strongly con­ Family Xanthidae MacLeay, 1838 vex, tip rounded. (sensu Guinot, 1978) Type species. — Cranaothus deforgesi, new Subfamily Euxanthinae Alcock, 1898 species, by monotypy. (sensu Serene, 1984) Etymology.—The generic name is de­ Cranaothus, new genus rived from the Greek "kranaos" for "rugged Diagnosis.— Carapace quadrate, regions and rocky" (alluding to the eroded carapace not well defined; dorsal surfaces with very surface), in arbitrary combination with a fi­ small squamiform granulations; branchial, nal syllable of many xanthid genera. Gender gastric, cardiac and intestinal regions with masculine. 706 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

Remarks.— In its external features Cra­ The carapace shape of Cranaothus is per­ naothus, new genus, appears to be closest haps closest to Neoxanthops, especially in to the genus Paramedaeus Guinot, 1967; species like N. cavatus (Rathbun, 1907) however, it differs in that the anterolateral which also has ridges on the dorsal surface. margin is not cut into distinct teeth (weak In N. cavatus, however, the frontal lobes are or otherwise), the front is very produced and separated only by a shallow cleft, not a deep lamellar, with the median fissure very deep; fissure, and the anterolateral margin is cut sternites 2-4 distinctly broader, sternal su­ into distinct lobes. The Gl of Cranaothus ture 1 and 2 being distinct (absent in Para­ is also proportionately shorter, with a dif­ medaeus), sternal suture 2 and 3 interrupted ferent shape, and the distal part is tapered medially (entire in Paramedaeus); male ab­ and very slender, compared to that of typ­ dominal segment 7 semicircular in shape ical Neoxanthops (see Serene 1984: figs. 127, (distinctly triangular in Paramedaeus); dis­ 128). Neoxanthops cavatus is not a typical tal part of the Gl is long, slender and strong­ member of Neoxanthops and should prob­ ly tapering, the tip being relatively sharp ably be transferred to a new genus. It differs (distal part stout and tip rounded in Para­ markedly from the type species of the genus, medaeus) (Guinot 1967, Serene 1984). N. lineatus Milne Edwards, 1867, in many With regard to the shape of the carapace, aspects, viz. the anterolateral margin grad­ Cranaothus resembles Indo-West Pacific ually becomes more obscure and gently genera such as Metaxanthops Serene, 1984, curves to end below the orbits, and does not Macromedaeus Ward, 1942, and Neoxan- meet the external orbital angle or supraor­ thops Guinot, 1968, but differs in many key bital margin; the frontal margin is not dis­ aspects. The front margin of Cranaothus is tinctly produced beyond the internal angle somewhat similar to that of Metaxanthops, of the supraorbital margin; the anterolateral with two broad and truncate lobes separated margin is not distinctly cristiform; the sur­ by a deep median fissure. Metaxanthops face is more distinctly domed, distinctly however, differs from Cranaothus in having sculptured, appears eroded, and instead of well developed and distinct epibranchial gently convex and completely smooth; the teeth and a much smoother carapace sur­ fingers of the chelipeds are very short in­ face. The general shape of the Gl in Meta­ stead of pigmented; and the subdistal part xanthops, although similar, is distinctly of the Gl has only a few short, simple hairs, stouter and the distal part is not slender and moderately long, setose hairs. tapering (Serene 1984: fig. 129). The an­ Cranaothus also differs from Macrome­ terolateral margin of Cranaothus resembles daeus, Neoxanthops and Metaxanthops in Macromedaeus nudipes (Milne Edwards, that the dactyls of the larger cheliped have 1867), but the structure of the front is dif­ a molariform basal cutting tooth absent in ferent, with the lobes more sinuous and less the other genera. projected forward in Macromedaeus. In With regard to the sculpturing on the car­ Macromedaeus, the two frontal lobes are apace surface, Cranaothus somewhat re­ also separated only by a cleft, without the sembles the genus Glyptoxanthus Milne Ed­ deep fissure present in Cranaothus. While wards, 1873-1881, which is represented in the regions are well defined in Macrome­ the Indo-West Pacific region by G. mean- daeus, they are only vaguely so in Crana­ drinus (Klunzinger, 1913). The anterolat­ othus. .The Gl of Cranaothus differs sub­ eral margin of Glyptoxanthus, however, is stantially from those of Macromedaeus, cut into distinct teeth, the front is not pro­ being proportionately shorter, stouter and jecting but is about level with the orbits, the different in shape (Serene 1984: figs.101 — frontal lobes are not separated by a deep 104). fissure, the dactylus of the cheliped lacks the VOLUME 106, NUMBER 4 707 special basal cutting tooth, and the last male the Xanthinae (cf. Neoxanthops, Metaxan- abdominal segment is distinctly triangular, thops, Macro medaeus). The genus is placed with a sharp tip and the lateral margins al­ in the Euxanthinae based on the absence of most straight (Odhner 1925: pi. 4 fig. 1, a clearly defined anterolateral margin be­ Guinot 1979: pi. 6 fig. 7). In Cranaothus, hind the external orbital angle in Crana­ the last male abdominal segment is semi­ othus, with the row of granules curving along circular. There is also some resemblance to the suborbital region toward the buccal cav­ of the genus Medaeops Guinot, 1967, ity. although the shapes of the carapaces differ. Interestingly, Medaeus granulosus (Has- Cranaothus deforgesi, new species well, 1882) has a weak basal cutting tooth Figs. 1-3 on the dactyls of the chelipeds. Paramedaeus noelensis. — ? Serene & Umali, The shape and form of the carapace of 1972:68, pi. 7 figs. 7-9 [nee Paramedaeus Cranaothus also bears a marked similarity noelensis (Ward, 1934)]. to an eastern Pacific monotypic genus, Li- paesthesius Rathbun, 1898, represented by Material examined. — Holotype male L. leeanus Rathbun, 1898. Like Crana­ (carapace 8.0 by 5.9 mm), (MNHN), Ches­ othus, the surface of the carapace in Li- terfield Island, Coral Sea, station 144, paesthesius is covered by many small gran­ 19°27.73'S, 158°23.28'E, ca. 50 m depth, ules, forming uneven patterns. Lipaesthesius sand and Halimeda algae substrate, dredge, differs from Cranaothus in the following as­ leg. B. Richer de Forges, 30 Aug 1988. pects: point of attachment of antennal fla- Description of holotype male.— Carapace gellum concealed by upper margin of basal regions not well defined, grooves separating segment (point of attachment of flagellum gastric and branchial regions shallow, 2F, distinct and not concealed in Cranaothus); 1M, 3M regions low but discernible, 4M granule patterns on dorsal surface of cara­ indistinct, L and R not defined; H-shaped pace not vermiform; front not produced for­ groove separating cardiac and gastric regions ward and without deep median cleft; pos­ distinct; dorsal surfaces covered with very terolateral margin strongly concave (more small squamiform granulations; branchial, so than in Cranaothus), forming distinct gastric, cardiac and intestinal regions with "waist"; junction between antero- and pos­ uneven eroded ridges, forming vermiculat- terolateral margins not clearly demarcated, ed pattern; pterygostomial, suborbital and the anterolateral margin gradually curving sub-branchial regions granulose. Density of posteriorly; carpus of cheliped very elon­ granules and granulations somewhat ob­ gate; chelae symmetrical; fingers of cheliped scures sutures between pterygostomial, long and slender and pigmented dark brown sub-branchial and suborbital regions as well (Rathbun 1930:272, pi. 112). as base of chelipeds, ambulatory legs and Serene's (1984) separation the subfami­ sternum. Front distinctly produced beyond lies Euxanthinae Alcock, 1898, and Xan- imaginary line connecting internal supra­ thinae MacLeay, 1838, is not satisfactory as orbital angle; distinctly bilobed, lobes sep­ there appears to be a degree of overlap in arated by very deep fissure extending back some of the characters used by him. The to epigastric region; surfaces smooth, mar­ establishment of Cranaothus further com­ gin truncatiform, gently concave. Supraor­ plicates matters because, while the genus bital margin with clearly defined rounded seems to belong to what Serene (1984) de­ inner angle, separated from front by distinct fined as Euxanthinae (cf. Medaeops, Para- groove; external orbital angle low, indis­ medaeus, Glyptoxanthus), it also bears a tinct, not clearly demarcating beginning of striking resemblance to some members of anterolateral margin; rest of supraorbital 708 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

margin entire, gently sinuous. Internal in­ ered with very small scale-like granulations fraorbital angle with distinct tooth visible and vermiform eroded ridges; carpus short, from dorsal view. Anterolateral margin ar­ rounded, inner distal angle with distinct cuate along anterior %, becoming straight blunt tooth and uneven anterior subdistal along posterior xh (approximately parallel to serrations; fingers not pigmented black or median longitudinal axis of carapace); not brown, appearing white in preservative; divided into distinct lobes or teeth, without larger cheliped with pronounced molari- any distinct cristate borders or clefts; 1 tu­ form basal cutting tooth on dactylus di­ bercle visible shortly behind external orbital rected obliquely outward; minor cheliped angle, followed by 3 larger ones on arcuate with elongated fingers. part of margin; straight part of anterolateral Abdomen with segments 3-5 completely margin marked by blunt tubercle on each fused, sutures separating these segments not edge; anterolateral margin distinctly sepa­ discernible, lateral margins entire, without rated from gently concave, converging pos­ any clefts or discontinuity; segments 1-3 terolateral margins. Posterior margin of car­ trapezoidal, segment 6 squarish, lateral apace gently convex. Antennules folding margins straight, parallel; segment 7 semi­ transversely, antennular fossae partly cov­ circular, lateral margins strongly convex, tip ered by protruding front from frontal view. rounded; surfaces of all segments slightly Antennal flagellum short, attached to stout rugose to squamate. basal segment occupying entire space be­ Gl relatively short and stout, proximal tween antennular fossa and internal orbital part gradually tapering, distal part straight, angle. Endostome with weak median lon­ very slender, distinctly tapering to sharp tip; gitudinal ridge and strong, oblique ridge on lateral margins of slender distal part lined either side of posterior part, adjacent to with short spines; subdistal part with nu­ mouth. merous long, stout setiferous hairs. G2 short, Entire outer surface of third maxilliped slender, distal part with petaloid process. finely granulose, that on merus appearing Etymology.— The species is named after more eroded; ischium rectangular, median Bertrand Richer de Forges, who so kindly oblique sulcus very shallow, indistinct; me­ made the ORSTOM specimen available for rus quadrate, with median oblique patch of study. eroded granules; outer surface of carpus Remarks. — The single known specimen rounded, granuliform; exopod reaching an­ of Cranaothus deforgesi, new species, is ma­ terior edge of merus, with blunt triangular ture despite its small size because the gon- subdistal tooth on inner margin, flagellum opods are fully developed. The vermiform long. ridges on the carapace and chelipeds are Sternum broad, entire surface covered formed by patches of very small granules with eroded granules, appearing squamate; and are easily chipped and scraped off. This suture between sternites 1 and 2 distinct, accounts for the species' eroded appearance. complete, sutures between sternites 2 and Cranaothus deforgesi bears a striking re­ 3, and 3 and 4 shallow, interrupted medi­ semblance to a specimen from the Philip­ ally, sutures between sternites 4 and 5, 5 pines identified as Paramedaeus noelensis and 6, and 6 and 7 incomplete; abdomen (Ward, 1934) by Serene & Umali (1972). reaching to imaginary line joining posterior Serene & Umali's (1972:68, pi. 7 figs. 7-9) bases of chelipeds. Gonopore coxal, open­ specimen, was a male 8.5 by 5.5 mm from ing below abdominal segment 3. Maluso Bay, collected from a depth of 25 Chelipeds distinctly asymmetrical; outer m by the Pele Sulu Expedition in 1964. The surfaces of merus, carpus and chelae cov­ specimen has a very produced and lamellar VOLUME 106, NUMBER 4 709

Fig. 1. Cranaothus deforgesi, new genus and species. Holotype male, carapace 8.0 by 5.9 mm, MNHN, Chesterfield Island. A, dorsal view; B, chelae. front, and the median fissure appears to be P. noelensis, noting that in their specimen very deep. The shape of the carapace, sculp­ "... the postero-lateral border is concave ture of the surface (covered with granulated instead of being straight. . . [t]he breadth is vermiculations) and form of the postero­ 1.41 its length instead of 1.47, the propor­ lateral margin, also resemble C. deforgesi. tion in the specimen of Forest & Guinot Serene & Umali (1972:69) commented on (1961). . . . the front in our specimen is more the differences between their Philippine salient, more pointed medially with the si­ specimen and Ward's (1934) description of nus more open." It appears that Serene & 710 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

Fig. 2. Cranaothus deforgesi, new genus and species. Holotype male, carapace 8.0 by 5.9 mm, MNHN, Chesterfield Island. A, dorsal view of carapace; B, frontal view of carapace; C, buccal cavity showing endostomial ridges; D, left third maxilliped (denuded, most of granulation not shown); E, sternites 1-4; F, abdomen; G, coxa, basis-ischium and merus of left cheliped; H, right carpus of cheliped (dorsal view); I, right fourth ambulatory leg; J, right third ambulatory leg. Small granules and squamate structures on surfaces of carapace, third maxilliped, sternum, chelipeds and abdomen omitted. Scales: A, B, E-J, 1.0 mm; C, D, 0.5 mm. VOLUME 106, NUMBER 4 711

Fig. 3. Cranaothus deforgesi, new genus and species. Holotype male, carapace 8.0 by 5.9 mm, MNHN, Chesterfield Island. A-D, left Gl; E, F, left G2. A, C, F, ventral view; B, D, E, dorsal view. Scales: A, B, E, 0.5 mm; C, D, F, 0.25 mm.

Umali's (1972) specimen is actually con- whether all the specimens of this species specific with Cranaothus deforgesi. Only a reported are conspecific. Certainly, Sakai's re-examination of their specimen (suppos­ (1976:426, fig.224 ) descriptions and figures edly in the National Museum of the Phil­ of the species differ from those by Forest & ippines) will resolve this matter with cer­ Guinot (1961) in having a shorter front, tainty. It is clear, however, that Cranaothus presence of distinct anterolateral teeth, and deforgesi is not conspecific with Parame- the smoother carapace. daeus noelensis (Ward, 1934). Unlike Cranaothus deforgesi, the cara­ As regards Paramedaeus noelensis (Ward, pace surface of Paramedaeus noelensis is 1934), the species was described from smoother and has no granulated vermicu- Christmas Island, Indian Ocean, by Ward lations; the carapace regions are more dis­ (1934:17, pi. 1 fig. 1) as Medaeus, but his tinct; the front is not lamellar in appearance, descriptions are brief and his figures rather is less produced and lacks the deep median schematic. Forest & Guinot (1961:56, pi. 1 fissure; the posterolateral margin is almost fig. 1, text figs. 42, 43, 44a, b) redescribed straight to slightly convex (distinctly con­ and refigured the species after examining the cave in C deforgesi); the outer surfaces of type and additional specimens from Upolu the carpus of the cheliped are less rugose; and Tahiti. Guinot (1967) subsequently the sternal structure has a distinctly wider proposed transferring the species to a new space between sternal sutures 2 and 3, and genus, Paramedaeus, and this was followed 3 and 4; the second male abdominal seg­ by Sakai (1976) and Serene (1984). The spe­ ment has no transverse ridges; the lateral cies is known from Christmas Island, Mad­ edges of the fused male abdominal segments agascar, Upolu, Tahiti, Japan and the Phil­ three and four have a distinct deep cleft on ippines. However, there are doubts as to each side (entire and continuous in C. de- 712 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON forgesi); the last male abdominal segment have improved the paper substantially. The is distinctly triangular in shape (not round­ study has been partially supported by RP ed); and the distal part of the Gl is not 900360 from the National University of produced into a long, slender projection (see Singapore. Forest & Guinot 1961:56, figs. 42, 43, 44a, b, pi. 1 fig. 1; Serene 1984: fig. 51, pi. 12F). Literature Cited Paramedaeus noelensis and Cranaothus Alcock, A. 1898. Materials for a carcinological fauna deforgesi seem to be closely related. The car­ of India. No. 3. The Brachyura Cyclometopa. apace of P. noelensis is remarkably similar Part I. The Family Xanthidae.—Journal of the to that of C. deforgesi in shape and general Asiatic Society of Bengal 67(2)(l):67-233. armature, the posterior part of the anterior Forest, J., & D. Guinot. 1961. Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In Ex­ lateral margin in both species is almost pedition francaise sur les recifs coralliens de la straight and parallel with the median axis Nouvelle-Caledonie. Volume preliminaire. Par­ of the . This confers a rather quadrate is, editions de la Fondation Singer-Polignac, ix- appearance to the carapace of both species. xi + 1-194, figs. 1-178, pis. 1-18, tabls. 1-3, 7 The general form of the Gls in both species maps. Guinot, D. 1967. Recherches preliminaires sur les is also similar. Differences in the form of groupements naturels chez les Crustaces Deca­ the male abdomen and sternum, however, podes Brachyoures. II. Les anciens genres Mi- suggest that it might be premature to trans­ cropanope Stimpson et Medaeus Dana. —Bul­ fer P. noelensis to Cranaothus, at least for letin du Museum national d'Histoire naturelle, the time being. Paris (2)39(2):345-374. . 1968. Recherches preliminaires sur les The unusual molariform basal cutting groupements naturels chez les Crustaces Deca­ tooth on the dactylus of the cheliped is rem­ podes Brachyoures. IV. Observations sur quel- iniscent of that in crabs of the genus Ca- ques genres de Xanthidae. —Bulletin du Mu­ lappa (Calappidae) which is used for "peel­ seum national d'Histoire naturelle, Paris (2)39(4): ing" gastropod shells (Shoup 1956, Ng & 695-727. . 1978. Principes d'une classification evolutive Tan 1984). In these crabs, the right cheliped des Crustaces Decapodes Brachyoures.—Bulle­ is almost always the larger one and pos­ tin du biologique Francaise et Belgique, new se­ sesses the basal cutting tooth. Ng & Tan ries 112(3):211-292. (1985) suggested that this was because ma­ . 1979. Donnees nouvelles sur la morphologie, rine gastropods have dextral coiling. The la phylogenese et la Crustaces Decapodes Brachyoures. — Memoires du Museum national well developed condition of the cutting tooth d'Histoire naturelle, Paris (A) Zoology 112:1- in Cranaothus strongly suggests that the crab 354, pis. 1-27. is also a "peeler" like Calappa. Interesting­ Haswell, W. A. 1882. On some new Australian ly, in the specimen recorded by Serene & Brachyura.—Proceedings of the Linnean Soci­ Umali (1972) (as Paramedaeus noelensis) ety of New South Wales 6(3):540-551. from the Philippines, the right chela is also Klunzinger, C. B. 1913. Die Rundkrabben (Cyclo­ metopa) des Roten Meeres.—Nova Acta Leo­ the larger and has a basal cutting tooth. pold Carolia 99(2):97-402, pis. 5-11. MacLeay, W. S. 1838. Illustrations of the Annulosa of South Africa; being a portion of the objects Acknowledgments of natural history chiefly collected during an ex­ pedition into the interior of South Africa, under The author is very grateful to A. Crosnier the direction of Dr. Andrew Smith, in the years and B. Richer de Forges (ORSTOM) for for­ 1834, 1835, and 1836; fitted out by the "Cape warding the material to him for study, and of Good Hope Association for Exploring Cen­ their kind help. A. Crosnier, D. Guinot, P. tral Africa." In A. Smith, ed., Illustrations of the zoology of South Africa investigations, Lon­ Clark, A. B. Williams and R. B. Manning don, Smith, Elder and Co., pp. 1-75, pis. 1-4. kindly read through the manuscript, and Milne Edwards, A. 1867. Descriptions de quelques their many useful suggestions and criticisms especes nouvelles de Crustaces Brachyures.— VOLUME 106, NUMBER 4 713

Annales de la Societe entomologique Francaise Sakai, T. 1976. Crabs of Japan and the adjacent seas. (4)7:263-288. In three volumes; English Text, pp. xxix + 773 . 1873-1881. Etudes sur les Xiphosures et les pp., Japanese Text, pp. 1-461, Plates Volume, Crustaces de la region Mexicaine. In Mission pp. 1-16, pis. 1-251. Kodansha Ltd., Tokyo. scientifique du Mexique et dans l'Amerique cen- Serene, R. 1984. Crustaces Decapodes Brachyoures trale, Recherches Zoologiques pour servir a de l'Ocean Indien occidental et de la Mer Rouge. Thistoire de la faune de l'Amerique centrale et Xanthoidea: Xanthidae et Trapeziidae. Adden­ du Mexique, 5:1-368, pis. 1-61. dum Carpiliidae et Menippidae —A. Cros- Ng. P. K. L..&L.W. H.Tan. 1984. The'shell peeling' nier.-Faune Tropicale (ORSTOM) 24:1^00, structure of the box crab Calappa philargius (L.) pis. 1-48. and other crabs in relation to mollusc shell ar­ , & A. F. Umali. 1972. The family Raninidae chitecture.—Journal of the Singapore National and other new and rare species of Brachyuran Academy of Science 13:195-199. Decapods from the Philippines and adjacent , & . 1985. 'Right Handedness'in het- regions. —Philippine Journal of Science 99(1-2): erochelous calappoid and xanthoid crabs—sug­ 21-105, pis. 1-9. gestion for a functional advantage.—Crusta- , & C. Vadon. 1981. Crustaces Decapodes: ceana 49:98-100. Brachyoures. List preliminaire, description de Odhner, T. 1925. Monographic Gattungen der Krab- formes nouvelles et remarques taxonomiques. benfamilie Xanthidae. I. —Goteborgs Kungl Ve- Resultats des Campagnes MUSORSTOM. I. tenskaps-och Vitterhets-Samhalles Handlingar Philippines. —Collection Memoires ORSTOM, (4)29(1): 10-92, pis. 1-5. Paris 91:117-140, pis. I—III. Rathbun, M. J. 1898. The Brachyura collected by the Shoup, J. B. 1956. Shell opening by crabs of the genus United States Fish Commission steamer Alba­ Calappa.- Science 160:887-888. tross on the voyage from Norfolk, Virginia, to Ward, M. 1934. Notes on a collection of crabs from San Francisco, California, 1887-1888.-Pro­ Christmas Island, Indian Ocean. —Bulletin of ceedings of the United States National Museum the Raffles Museum 9:5-28, pis. 1-3. 21:567-616, pis. 41-44. . 1942. Notes on the Crustacea of the Desjar- . 1907. Report on the Brachyrhyncha, Oxysto- dins Museum, Mauritius Institute, with descrip­ mata and Dromiacea. In Report on the crabs tions of new genera and species. —Mauritius In­ obtained by F.I.S. "Endeavour" on the coasts stitute Bulletin 2(2):49-l 13, pis. 5, 6. of Queensland, New South Wales, Victoria, South Australia and Tasmania, Biological Re­ Department of Zoology, National Uni­ sults of the Fishing Experiments carried on by the F.I.S. "Endeavour" 1909-14, Sydney 5(3): versity of Singapore, Kent Ridge, Singapore 95-156, pis. 16-42. 0511, Republic of Singapore. . 1930. The cancroid crabs of America. — Unit­ ed States National Museum Bulletin 152:i-xvi + 1-609, figs. 1-85, pis. 1-230.