DOCSLIB.ORG

Hemiptera: Cicadidae) and a Key to Their Species

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Hemiptera: Cicadidae) and a Key to Their Species Journal of Asia-Pacific Biodiversity xxx (2016) 1e7 HOSTED BY Contents lists available at ScienceDirect Journal of Asia-Pacific Biodiversity journal homepage: http://www.elsevier.com/locate/japb Original article Description of three new genera, Paratibicen, Megatibicen, and Ameritibicen, of Cryptotympanini (Hemiptera: Cicadidae) and a key to their species Young June Lee* Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, CT, USA article info abstract Article history: Three new genera, Paratibicen gen. nov., Megatibicen gen. nov., and Ameritibicen gen. nov., are erected for Received 18 August 2016 the former members of Neotibicen Hill & Moulds, 2015 from North America, designating Cicada similaris Received in revised form Smith & Grossbeck, 1907, Cicada auletes Germar, 1834, and Cicada dorsata Say, 1825 as their respective 4 September 2016 type species. Morphological comparisons are made among the three new genera, Lyristes Horváth, 1926 Accepted 7 September 2016 (¼ Tibicen Latreille, 1825), Auritibicen Lee, 2015, Neotibicen sensu stricto, and other related genera. A key to Available online xxx the species of the genera Neotibicen, Paratibicen, Megatibicen, and Ameritibicen is provided. Copyright Ó 2016, National Science Museum of Korea (NSMK) and Korea National Arboretum (KNA). Keywords: Auritibicen Production and hosting by Elsevier. This is an open access article under the CC BY-NC-ND license (http:// Hadoa creativecommons.org/licenses/by-nc-nd/4.0/). Lyristes Neotibicen new combinations North America Tibicen Introduction species: H. bifida (Davis, 1916), H. chihuahuaensis (Sanborn, 2007), H. chiricahua (Davis, 1923), H. chisosensis (Davis, 1934), H. distanti By the erections of Auritibicen Lee, 2015 by Lee (2015) and (Metcalf, 1963), H. duryi (Davis, 1917), H. fusca (Davis, 1934), Neotibicen Hill & Moulds, 2015 and Hadoa Moulds, 2015 in Hill et al H. hidalgoensis (Davis, 1941), H. inaudita (Davis, 1917), (2015), and by the generic reassignments by Sanborn (2015), there H. longiopercula (Davis, 1926), H. minor (Davis, 1934), H. montezuma have been relocations of many species of the former Lyristes Hor- (Distant, 1881), H. neomexicensis (Stucky, 2013), H. paralleloides váth, 1926 (type species: Cicada plebeja Scopoli, 1763 from (Davis, 1934), H. parallela (Davis, 1923), H. robusta (Distant, 1881), Europe; ¼ Tibicen Latreille, 1825 nomen nudum) into different H. simplex (Davis, 1941), H. sugdeni (Davis, 1941), H. texana (Metcalf, genera. After synonymization of Lyristes altaiensis Schmidt, 1932 1963), and H. townsendii (Uhler, 1905). with Neotibicen resh (Haldeman, 1852) by Wang et al (2015) (also However, it was found through an alpha taxonomic study that see Lee 2015 and Sanborn 2015), the genus Lyristes is now there still should exist multiple groups or genera within the current comprised of only five species from sub-Mediterranean and Med- concept of the genus Neotibicen. iterranean areas in the Western Palaearctic Region (in Europe). Among the species of the genus Neotibicen, the following 11 Most of the former “Tibicen” species from North America are species form a monophyletic group: N. auriferus (Say, 1825), now placed in the two genera Neotibicen and Hadoa. Neotibicen N. canicularis (Harris, 1841), N. davisi (Smith & Grossbeck, 1907), includes 21 species, excluding an extinct species [Neotibicen ber- N. latifasciatus (Davis, 1915), N. linnei (Smith & Grossbeck, 1907), mudianus (Verrill,1902)] and an invalid name [Neotibicen variegatus N. lyricen (Degeer, 1773), N. pruinosus (Say, 1825), N. robinsonianus (Fabricius, 1794) nomen nudum]. Hadoa includes the following 20 (Davis, 1922), N. superbus (Fitch, 1855), N. tibicen (Linnaeus, 1758), and N. winnemanna (Davis, 1912). These 11 species remain in Neotibicen. Neotibicen similaris (Smith & Grossbeck, 1907) has its own * Corresponding author. peculiar morphology as discussed later and thus should be placed E-mail address: [email protected]. Peer review under responsibility of National Science Museum of Korea (NSMK) and in a separate monotypic genus. A new genus, Paratibicen gen. nov., Korea National Arboretum (KNA). is described for this species. http://dx.doi.org/10.1016/j.japb.2016.09.002 pISSN2287-884X eISSN2287-9544/Copyright Ó 2016, National Science Museum of Korea (NSMK) and Korea National Arboretum (KNA). Production and hosting by Elsevier. This is an open access article under the CC BY-NC-ND license (http://creativecommons.org/licenses/by-nc-nd/4.0/). Please cite this article in press as: Lee YJ, Description of three new genera, Paratibicen, Megatibicen, and Ameritibicen, of Cryptotympanini (Hemiptera: Cicadidae) and a key to their species, Journal of Asia-Pacific Biodiversity (2016), http://dx.doi.org/10.1016/j.japb.2016.09.002 2 YJ Lee / Journal of Asia-Pacific Biodiversity xxx (2016) 1e7 The following three species form another monophyletic group: Hadoa townsendii (Uhler, 1905): 1_, Sierra County, NM, USA, 15 VI N. auletes (Germar, 1834), N. resh (Haldeman, 1852), and N. resonans 2012 (UCONN). (Walker, 1850). A new genus, Megatibicen gen. nov., is described for Cacama valvata (Uhler, 1888): 1_, Bent County, CO, USA, 4 VII these three species. 2004 (UCONN). The following six species form another monophyletic group: Photographs and illustrations provided by Cole (2008), Davis N. cultriformis (Davis, 1915), N. dealbatus (Davis, 1915), N. dorsatus (1915, 1916, 1917, 1923, 1926, 1927, 1934, 1938, 1941), Lawson (Say, 1825), N. figuratus (Walker, 1858), N. pronotalis (Davis, 1938), (1920), Stucky (2013), and others were also referenced. and N. tremulus (Cole, 2008). Another new genus, Ameritibicen gen. The digital images of the male genitalia were captured using a nov., is described for these six species. Leica stereo-zoom microscope attached to a computer-assisted im- A key to the males of all the 21 species of Neotibicen, Paratibicen, aging system, Auto-Montage by Syncroscopy (Cambridge, UK), at Megatibicen, and Ameritibicen is provided in subsequent sections. the collection facility of the University of Connecticut. Terminology for morphological features generally follows that of Moulds (2005). Materials and methods Genus Neotibicen Hill & Moulds, 2015 Many specimens of the species of the new genera and their Type species. Cicada canicularis Harris, 1841 (type locality: Mas- closely related genera including Lyristes, Auritibicen, Neotibicen sachusetts; Figure 1). sensu stricto, Hadoa Moulds, 2015, and Cacama Distant, 1904 were Included species. N. auriferus (Say, 1825), N. canicularis (Harris, examined for this study, which are specimens from Kathy B.R. Hill 1841), N. davisi (Smith & Grossbeck, 1907) [with the subspecies of and David C. Marshall’s cicada collection at the collection facility of N. davisi davisi (Smith & Grossbeck, 1907) and N. davisi hardeni the University of Connecticut, Storrs, CT, USA (UCONN), and from (Davis, 1918)], N. latifasciatus (Davis, 1915), N. linnei (Smith & the present author’s private collection (YJL), including the speci- Grossbeck, 1907), N. lyricen (Degeer, 1773) [with the subspecies of mens listed below. N. lyricen engelhardti (Davis, 1910), N. lyricen lyricen (Degeer, 1773), Neotibicen auriferus (Say, 1825): 1_, Meade County, KS, USA, 05 and N. lyricen virescens (Davis, 1935)], N. pruinosus (Say, 1825) [with IX 2007 (UCONN). Neotibicen canicularis (Harris, 1841): 1_, Man- the subspecies of N. pruinosus fulvus (Beamer, 1924) and chester, CT, USA, 10 VIII 2008 (YJL); 1_, Mansfield, CT, USA, 07 VIII N. pruinosus pruinosus (Say, 1825)], N. robinsonianus (Davis, 1922), 2008 (YJL); 1_, Storrs, CT, USA, 19 VIII 2008 (YJL). Neotibicen davisi N. superbus (Fitch, 1855), N. tibicen (Linnaeus, 1758) [with the (Smith & Grossbeck, 1907): 1_, Citrus County, FL, USA, 22 IX 2011 subspecies of N. tibicen australis (Davis, 1912) and N. tibicen tibicen (UCONN). Neotibicen latifasciatus (Davis, 1915): 1_, Cape May (Linnaeus, 1758)], and N. winnemanna (Davis, 1912). County, NJ, USA, 06 VIII 2010 (UCONN). Neotibicen linnei (Smith & Diagnosis. Head including eyes wider than base of mesonotum. Grossbeck, 1907): 1_, Kent County, DE, USA, 28 VIII 2004 (UCONN). Pronotum with lateral margin weakly ampliated, not dentate. Male Neotibicen lyricen (Degeer, 1773): 1_, Storrs, CT, USA, 30 VII 2008 opercula about as long as wide or longer than wide, overlapped at (YJL); 1_, Storrs, CT, USA, 6 VIII 2009 (YJL). Neotibicen lyricen inner margins. Wings hyaline. Forewing comparatively long and engelhardti (Davis, 1910): 1_, Albemarle County, VA, USA, 17 VIII slender (ratio of forewing length to width 2.65e3.20). Forewing 2003 (UCONN). Neotibicen pruinosus (Say, 1825): 1_, Barber County, radial cell slightly slenderer than ulnar cell 3. Male abdomen about KS, USA, 5 IX 2007 (UCONN). Neotibicen robinsonianus (Davis, as long as or slightly longer than head and thorax together. Male 1922): 1_, Culpeper County, VA, USA, 17 VIII 2003 (UCONN). Neo- abdominal sternite VIII widely rounded or V-shaped in posterior tibicen superbus (Fitch, 1855): 1_, Freestone County, TX, USA, 23 VI view. Timbal cover completely covering timbal. Pygofer barrel- 2002 (UCONN). Neotibicen tibicen (Linnaeus, 1758): 1_, Bergen shaped (with a little widened distal part) in ventral view. Dorso- County, NJ, USA, 15 VIII 2015 (YJL); 1_, Culpeper County, VA, USA, 17 distal part (between distal shoulders, including dorsal beak and VIII 2003 (UCONN). inward curvatures on both sides of dorsal beak) of pygofer lower Paratibicen similaris (Smith & Grossbeck, 1907) comb. nov.:
Load more

Recommended publications
  • A New Neotibicen Cicada Subspecies (Hemiptera: Cicadidae)
    Zootaxa 4272 (4): 529–550 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2017 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4272.4.3 http://zoobank.org/urn:lsid:zoobank.org:pub:C6234E29-8808-44DF-AD15-07E82B398D66 A new Neotibicen cicada subspecies (Hemiptera: Cicadidae) from the southeast- ern USA forms hybrid zones with a widespread relative despite a divergent male calling song DAVID C. MARSHALL1 & KATHY B. R. HILL Dept. of Ecology and Evolutionary Biology, University of Connecticut, 75 N. Eagleville Rd., Storrs, CT 06269 USA 1Corresponding author. E-mail: [email protected] Abstract A morphologically cryptic subspecies of Neotibicen similaris (Smith and Grossbeck) is described from forests of the Apalachicola region of the southeastern United States. Although the new form exhibits a highly distinctive male calling song, it hybridizes extensively where it meets populations of the nominate subspecies in parapatry, by which it is nearly surrounded. This is the first reported example of hybridization between North American nonperiodical cicadas. Acoustic and morphological characters are added to the original description of the nominate subspecies, and illustrations of com- plex hybrid song phenotypes are presented. The biogeography of N. similaris is discussed in light of historical changes in forest composition on the southeastern Coastal Plain. Key words: Acoustic behavior, sexual signals, hybridization, hybrid zone, parapatric distribution, speciation Introduction The cryptotympanine cicadas of North America have received much recent attention with the publication of comprehensive molecular and cladistic phylogenies and the reassignment of all former North American Tibicen Latreille species into new genera (Hill et al.
    [Show full text]
  • Arthropods of Elm Fork Preserve
    Arthropods of Elm Fork Preserve Arthropods are characterized by having jointed limbs and exoskeletons. They include a diverse assortment of creatures: Insects, spiders, crustaceans (crayfish, crabs, pill bugs), centipedes and millipedes among others. Column Headings Scientific Name: The phenomenal diversity of arthropods, creates numerous difficulties in the determination of species. Positive identification is often achieved only by specialists using obscure monographs to ‘key out’ a species by examining microscopic differences in anatomy. For our purposes in this survey of the fauna, classification at a lower level of resolution still yields valuable information. For instance, knowing that ant lions belong to the Family, Myrmeleontidae, allows us to quickly look them up on the Internet and be confident we are not being fooled by a common name that may also apply to some other, unrelated something. With the Family name firmly in hand, we may explore the natural history of ant lions without needing to know exactly which species we are viewing. In some instances identification is only readily available at an even higher ranking such as Class. Millipedes are in the Class Diplopoda. There are many Orders (O) of millipedes and they are not easily differentiated so this entry is best left at the rank of Class. A great deal of taxonomic reorganization has been occurring lately with advances in DNA analysis pointing out underlying connections and differences that were previously unrealized. For this reason, all other rankings aside from Family, Genus and Species have been omitted from the interior of the tables since many of these ranks are in a state of flux.
    [Show full text]
  • Southern Gulf, Queensland
    Biodiversity Summary for NRM Regions Species List What is the summary for and where does it come from? This list has been produced by the Department of Sustainability, Environment, Water, Population and Communities (SEWPC) for the Natural Resource Management Spatial Information System. The list was produced using the AustralianAustralian Natural Natural Heritage Heritage Assessment Assessment Tool Tool (ANHAT), which analyses data from a range of plant and animal surveys and collections from across Australia to automatically generate a report for each NRM region. Data sources (Appendix 2) include national and state herbaria, museums, state governments, CSIRO, Birds Australia and a range of surveys conducted by or for DEWHA. For each family of plant and animal covered by ANHAT (Appendix 1), this document gives the number of species in the country and how many of them are found in the region. It also identifies species listed as Vulnerable, Critically Endangered, Endangered or Conservation Dependent under the EPBC Act. A biodiversity summary for this region is also available. For more information please see: www.environment.gov.au/heritage/anhat/index.html Limitations • ANHAT currently contains information on the distribution of over 30,000 Australian taxa. This includes all mammals, birds, reptiles, frogs and fish, 137 families of vascular plants (over 15,000 species) and a range of invertebrate groups. Groups notnot yet yet covered covered in inANHAT ANHAT are notnot included included in in the the list. list. • The data used come from authoritative sources, but they are not perfect. All species names have been confirmed as valid species names, but it is not possible to confirm all species locations.
    [Show full text]
  • First Host Plant Record for Pacarina (Hemiptera, Cicadidae)
    Neotropical Biology and Conservation 15(1): 77–88 (2020) doi: 10.3897/neotropical.15.e49013 SHORT COMMUNICATION First host plant record for Pacarina (Hemiptera, Cicadidae) Annette Aiello1, Brian J. Stucky2 1 Smithsonian Tropical Research Institute, Panama 2 Florida Museum of Natural History, University of Florida, Gainesville, FL, USA Corresponding author: Brian J. Stucky ([email protected]) Academic editor: P. Nunes-Silva | Received 4 December 2019 | Accepted 20 February 2020 | Published 19 March 2020 Citation: Aiello A, Stucky BJ (2020) First host plant record for Pacarina (Hemiptera, Cicadidae). Neotropical Biology and Conservation 15(1): 77–88. https://doi.org/10.3897/neotropical.15.e49013 Abstract Twenty-nine Pacarina (Hemiptera: Cicadidae) adults, 12 males and 17 females, emerged from the soil of a potted Dracaena trifasciata (Asparagaceae) in Arraiján, Republic of Panama, providing the first rearing records and the first definitive host plant records for any species of Pacarina. These reared Pacarina appear to be morphologically distinct from all known species of Pacarina and likely repre- sent an undescribed species. In light of this finding, we also discuss the taxonomy, biogeography, and ecology of Pacarina. Keywords cicada, Dracaena, host plant, rearing, taxonomy Introduction As far as is known, all cicadas are herbivores that spend the vast majority of their long life cycles as nymphs, living deep underground and feeding on the xylem sap of plant roots (Beamer 1928; Cheung and Marshall 1973; White and Strehl 1978). Be- cause of their relative inaccessibility to researchers, very little information is availa- ble about the host plant associations of juvenile cicadas. Consequently, even though adult cicadas are among the most conspicuous and familiar of all insects, the host plants of most cicada species’ nymphs remain unknown.
    [Show full text]
  • New Combinations for Six Species Belonging to Cryptotympanini Handlirsch (Hemiptera: Cicadidae), Former Members of the Genus Tibicen Latreille, 1825
    Zootaxa 4027 (2): 447–450 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Correspondence ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4027.3.9 http://zoobank.org/urn:lsid:zoobank.org:pub:2C08AA8A-8FBB-4291-9A29-63CEB7BD77BA New combinations for six species belonging to Cryptotympanini Handlirsch (Hemiptera: Cicadidae), former members of the genus Tibicen Latreille, 1825 ALLEN F. SANBORN Department of Biology, Barry University, 11300 NE Second Avenue, Miami Shores, FL 33161-6695, USA. E-mail: [email protected] Abstract. The generic status of several species formerly placed in the genus Tibicen is reconsidered based on the recent erection of new genera as well as historical evidence. Subsolanus Moulds, 2015 syn. nov. is shown to be a junior synonym of Auritibicen Lee, 2015. Six species are reassigned from Tibicen to appropriate genera based on morphological characteristics. The genus Tibicen Latreille, 1825 has had a confusing past (see summary in Boulard & Puissant, 2014; Marshall & Hill, 2014; Sanborn, 2014a). A major factor in the confusion was the use of different species as the type species of the genus in Latreille (1825) (Cicada plebeja Scopoli, 1763) and Latreille (1829) (Cicada haematodes Scopoli, 1763), species that are currently classified in different subfamilies. This combined with the limited original descriptions led to confusion as to what taxa should be assigned to the genus. As a result, Tibicen became a convenient place to assign species when it was not known where else the species should be placed and species from all three of the current subfamilies were at one time or another classified in the genus Tibicen.
    [Show full text]
  • Cicada Ovipositors Enhanced with Metals and Other Inorganic Elements Matthew S
    www.nature.com/scientificreports OPEN An augmented wood-penetrating structure: Cicada ovipositors enhanced with metals and other inorganic elements Matthew S. Lehnert1*, Kristen E. Reiter1,2, Gregory A. Smith1 & Gene Kritsky3 Few insect species are as popular as periodical cicadas (Magicicada spp.). Despite representing an enormous biomass and numbers that exceed 370/m2 during mass emergences, the extended time period of the underground nymphal stages (up to 17 years) complicates investigations of their life history traits and ecology. Upon emergence, female cicadas mate and then use their ovipositors to cut through wood to lay their eggs. Given the ability to penetrate into wood, we hypothesized that the ovipositor cuticle is augmented with inorganic elements, which could increase hardness and reduce ovipositor fracturing. We used scanning electron microscopy and energy dispersive x-ray spectroscopy to evaluate the material properties of ovipositors of four cicada species, including three species of periodical cicadas. We found 14 inorganic elements of the cuticle, of which P, Ca, Si, Mg, Na, Fe, Zn, Mn, Cl, K, and S show the highest concentrations (%wt) near the apex of the ovipositor, where other structural modifcations for penetrating wood are present. To the best of our knowledge, this is the frst report of metal deposits in the cuticle of true bugs (Hemiptera, >80,000 described species). Te independent origin of traits that perform similar functions represents a cornerstone of natural selection. Examples of such convergent evolution can be found across animal taxa: intelligence among birds and apes1, echolocation among bats and dolphins2, and fuid-feeding mechanisms among fies and butterfies3.
    [Show full text]
  • An Appraisal of the Higher Classification of Cicadas (Hemiptera: Cicadoidea) with Special Reference to the Australian Fauna
    © Copyright Australian Museum, 2005 Records of the Australian Museum (2005) Vol. 57: 375–446. ISSN 0067-1975 An Appraisal of the Higher Classification of Cicadas (Hemiptera: Cicadoidea) with Special Reference to the Australian Fauna M.S. MOULDS Australian Museum, 6 College Street, Sydney NSW 2010, Australia [email protected] ABSTRACT. The history of cicada family classification is reviewed and the current status of all previously proposed families and subfamilies summarized. All tribal rankings associated with the Australian fauna are similarly documented. A cladistic analysis of generic relationships has been used to test the validity of currently held views on family and subfamily groupings. The analysis has been based upon an exhaustive study of nymphal and adult morphology, including both external and internal adult structures, and the first comparative study of male and female internal reproductive systems is included. Only two families are justified, the Tettigarctidae and Cicadidae. The latter are here considered to comprise three subfamilies, the Cicadinae, Cicadettinae n.stat. (= Tibicininae auct.) and the Tettigadinae (encompassing the Tibicinini, Platypediidae and Tettigadidae). Of particular note is the transfer of Tibicina Amyot, the type genus of the subfamily Tibicininae, to the subfamily Tettigadinae. The subfamily Plautillinae (containing only the genus Plautilla) is now placed at tribal rank within the Cicadinae. The subtribe Ydiellaria is raised to tribal rank. The American genus Magicicada Davis, previously of the tribe Tibicinini, now falls within the Taphurini. Three new tribes are recognized within the Australian fauna, the Tamasini n.tribe to accommodate Tamasa Distant and Parnkalla Distant, Jassopsaltriini n.tribe to accommodate Jassopsaltria Ashton and Burbungini n.tribe to accommodate Burbunga Distant.
    [Show full text]
  • A Guide to Arthropods Bandelier National Monument
    A Guide to Arthropods Bandelier National Monument Top left: Melanoplus akinus Top right: Vanessa cardui Bottom left: Elodes sp. Bottom right: Wolf Spider (Family Lycosidae) by David Lightfoot Compiled by Theresa Murphy Nov 2012 In collaboration with Collin Haffey, Craig Allen, David Lightfoot, Sandra Brantley and Kay Beeley WHAT ARE ARTHROPODS? And why are they important? What’s the difference between Arthropods and Insects? Most of this guide is comprised of insects. These are animals that have three body segments- head, thorax, and abdomen, three pairs of legs, and usually have wings, although there are several wingless forms of insects. Insects are of the Class Insecta and they make up the largest class of the phylum called Arthropoda (arthropods). However, the phylum Arthopoda includes other groups as well including Crustacea (crabs, lobsters, shrimps, barnacles, etc.), Myriapoda (millipedes, centipedes, etc.) and Arachnida (scorpions, king crabs, spiders, mites, ticks, etc.). Arthropods including insects and all other animals in this phylum are characterized as animals with a tough outer exoskeleton or body-shell and flexible jointed limbs that allow the animal to move. Although this guide is comprised mostly of insects, some members of the Myriapoda and Arachnida can also be found here. Remember they are all arthropods but only some of them are true ‘insects’. Entomologist - A scientist who focuses on the study of insects! What’s bugging entomologists? Although we tend to call all insects ‘bugs’ according to entomology a ‘true bug’ must be of the Order Hemiptera. So what exactly makes an insect a bug? Insects in the order Hemiptera have sucking, beak-like mouthparts, which are tucked under their “chin” when Metallic Green Bee (Agapostemon sp.) not in use.
    [Show full text]
  • Cicadas Fact Sheet No
    Cicadas Fact Sheet No. 5.590 Insect Series|Trees and Shrubs by W.S. Cranshaw and B. Kondratieff* Cicadas are among the largest Colorado Quick Facts insects in the order Hemiptera, which includes other sap-sucking groups with • Cicadas are large insects with prominent beaks such as leafhoppers, aphids, young stages that burrow and and spittlebugs. Twenty-nine species occur in develop underground and the state (Table 1). The largest (Megatibicen feed on fluids from roots of spp.) are stout-bodied insects that are 1 to grasses, shrubs, and trees. 1 ½ inches but Colorado’s cicadas range considerably in size. Beameria venosa, • The adult cicadas found Cicadettana calliope and C. kansa are small in Colorado emerge 3 to 5 species (ca. ½ inch) that is develop on grasses years, sometimes longer, after and shrubs growing in hillsides of canyons Figure 1: Megatibicen dealbatus, a large eggs have hatched. and arroyos of both eastern and western “dog-day” type of cicada that is expanding populations along the Front Range. Colorado. • Periodical cicadas, such Although abundant, cicadas are far more as the 17-year and 13-year often heard than seen. Males make a variety cicadas, do not occur in of sounds to attract females. Most commonly Colorado. heard are loud, often shrill, buzzing calls, • Male cicadas “sing” to attract sometimes with several individual insects females. Many produce loud, synchronizing their songs. Other cicadas make rustling or clicking noises. shrill buzzing noises. Despite their large size, cicadas cause • Cicadas do little if any injury little injury. The immature stages (nymphs) while feeding on plants.
    [Show full text]
  • Elk and Wolves in Yellowstone
    Western Winter 2014 Issue 01 NATURAL RESOURCE SCIENCE AND MANAGEMENT IN THE WEST The Bird that Brought the West Together Cattle as Ecosystem Engineers The Ecology of Fear A publication of the Ruckelshaus Institute, a division of the University of Wyoming’s Haub School of Environment and Natural Resources Western Confluence Editor, Emilene Ostlind DIRECTOR’S NOTE Editorial Fellow, Kelly Hatton Graphic Designer, Tana Stith By Indy Burke Website Designer, Jessica Perry “I’m weary and tired. I’ve done my day’s riding. Nighttime is rolling my way. Ruckelshaus Institute The sky’s on fire and the light’s slowly fading. Peaceful and still ends the day. Director, Indy Burke Associate Director, Nicole Korfanta And out on the trail the night birds are calling, singing their wild melody. Spicer Chair in Collaborative Practice, Steve Smutko Down in the canyon the cottonwood whispers a song of Wyoming for me.” Collaboration Program Director, – Chris LeDoux Jessica Clement Communications Coordinator, Grazing in the west has been an important way of life for well over a century, supporting families, inspiring Emilene Ostlind Marketing Coordinator, Fred Schmechel poetry and song, and maintaining open spaces. Over recent decades, grazing has also provoked intense controversy. Senior Office Assistant, Differing grazing practices, variable impacts to public lands, and livestock influences on wildlife habitat trigger Mandy Bohlender opposing views. Even the scientific literature is contradictory, with recent articles both demonstrating the positive Accountant
    [Show full text]
  • LISTADO DE ESPECIES DE CIGARRAS (HEMIPTERA: CICADIDAE) EN EL ESTADO DE OAXACA, MÉXICO Y NOTAS SOBRE LA BIOLOGÍA DE Quesada Gigas (OLIVIER)
    SISTEMÁTICA Y MORFOLOGÍA ISSN: 2448-475X LISTADO DE ESPECIES DE CIGARRAS (HEMIPTERA: CICADIDAE) EN EL ESTADO DE OAXACA, MÉXICO Y NOTAS SOBRE LA BIOLOGÍA DE Quesada gigas (OLIVIER) José Antonio Sánchez-García1 *, Roselia Jarquín-López1, Héctor Miguel Guzmán- Vásquez1, Fernando Ruiz-Ortiz1, Elia Jirón-Pablo1, Laura Martínez-Martínez1 y Hansen Duvan Meneses-Agudelo2 1 Instituto Politécnico Nacional. CIIDIR Unidad Oaxaca. Hornos 1003, Col. Nochebuena, Santa Cruz Xoxocotlán Oaxaca, México, C. P. 71230. 2Universidad Politécnica de Madrid, Escuela Técnica Superior de Ingeniería Agronómica, Av. Complutense 3 y Av. Puerta Hierro, Madrid, España, C. P. 28040. Autor de correspondencia: [email protected] RESUMEN. Se presenta un listado de ocho especies de cigarras presentes en el estado de Oaxaca y se proporciona información sobre la biología de Quesada gigas (Olivier) en Valles Centrales de Oaxaca. Palabras clave: Taxonomía, guamúchil, base de datos. Checklist of cicadas (Hemiptera: Cicadidae) from Oaxaca, Mexico and notes on biology of Quesada gigas (Olivier) ABSTRACT. A checklist of eigth species of cicadas present in the state of Oaxaca is presented and information is provided on biology of Quesada gigas in Central Valley of Oaxaca Keywords: Taxonomy, guamuchil, data base. INTRODUCCIÓN Las cigarras o chicharras son insectos de tamaño mediano a grande, de 2 a 7 cm, muy familiares por su canto. En México se conocen 117 taxa (113 especies y cuatro subespecies), de las cuales ocho están registradas en Oaxaca (Sanborn, 2007). Sin embargo, no se han realizado estudios faunísticos en el estado de Oaxaca; los registros que se tienen provienen de la revisión de colecciones principalmente extranjeras y de la UNAM, por lo que ahora se plantea realizar un estudio más completo de la fauna de cigarras de la región y proveer un listado actualizado, además de proporcionar información de la biología, que prácticamente en México se desconoce, así como registro de nuevos hospederos.
    [Show full text]
  • An Appraisal of the Cicadas of the Genus <I>Abricta</I> StÅL and Allied Genera
    © Copyright Australian Museum, 2003 Records of the Australian Museum (2003) Vol. 55: 245–304. ISSN 0067-1975 An Appraisal of the Cicadas of the Genus Abricta Stål and Allied Genera (Hemiptera: Auchenorrhyncha: Cicadidae) M.S. MOULDS Invertebrate Zoology Division, Australian Museum, 6 College Street, Sydney NSW 2010, Australia [email protected] ABSTRACT. The cicada genus Abricta Stål currently contains a heterogeneous group of species which is considered best divided into four genera. Abricta sensu str. includes only A. brunnea (Fabricius) and A. ferruginosa (Stål) which are confined to Mauritius and neighbouring islands. The monotypic genus Chrysolasia n.gen., is proposed for a single Guatemalan species, A. guatemalena (Distant). Another monotypic genus, Aleeta n.gen., is proposed for the species A. curvicosta (Germar) from eastern Australia. Fourteen Australian species are placed in Tryella n.gen.: castanea Distant, noctua Distant, rubra Goding & Froggatt, stalkeri Distant, willsi Distant, adela n.sp., burnsi n.sp., crassa n.sp., graminea n.sp., infuscata n.sp., kauma n.sp., lachlani n.sp., occidens n.sp. and ochra n.sp. The five remaining species currently placed in Abricta (borealis Goding & Froggatt, burgessi Distant, cincta Fabricius and occidentalis Goding & Froggatt from Australia plus pusilla Fabricius of unknown locality) do not belong to Abricta or closely allied genera. Cladistic analyses place C. guatemalena basally on all trees. The Mauritian genus Abricta sensu str., and the genera, Abroma Stål and Monomatapa Distant, form a sister group to all Australian species. There is strong evidence suggesting that Abricta and Abroma are synonymous. Keys to genera and species and maps of distribution are provided.
    [Show full text]