Geosuffolk Notes 52 Trilobites – a Selection Geosuffolk at Ipswich
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A Simple Generative Model of Trilobite Segmentation and Growth
paleo.peercommunityin.orG Open Access A SIMPLE GENERATIVE MODEL OF TRILOBITE Peer-ReVIEWED SEGMENTATION AND GROWTH RESEARCH ARTICLE Melanie J Hopkins1 1 Division OF Paleontology (INVERTEBRates), American Museum OF NaturAL History – NeW York, USA Cite as: Hopkins MJ (2020). A SIMPLE GENERATIVE MODEL OF TRILOBITE SEGMENTATION THIS ARTICLE HAS BEEN PEER-REVIEWED BY PCI PALEO AND GROwth. PaleorXiv VERSION 3, Read THE Editor’S RECOMMENDATION AND ReferEES’ REPORTS AT DOI:10.24072/pci.paleo.100004 peer-rEVIEWED BY PCI Paleo. DOI: 10.31233/osf.io/zt642 Published: 27 January 2020 ABSTRACT Recommender: GenerATIVE GROWTH MODELS HAVE BEEN THE BASIS FOR NUMEROUS STUDIES OF MORPHOLOGICAL DIVERSITY AND Evolution. Christian Klug Most WORK HAS FOCUSED ON MODELING ACCRETIONARY GROWTH systems, WITH MUCH LESS ATTENTION TO DISCRETE GROWTH systems. GenerATIVE GROWTH MODELS FOR MOLTING ORganisms, SUCH AS ARTHRopods, HAVE REMAINED PARTICULARLY ReVIEwers: Kenneth De Baets AND Lukáš Laibl elusive. HoWEver, OUR UNDERSTANDING OF post-embryonic GROWTH IN TRILOBITE SPECIES IS SUFFICIENTLY MATURE THAT IT IS NOW POSSIBLE TO MODEL GROWTH IN A WAY THAT INCORPORATES THE ADDITION OF NEW PARTS AS WELL AS DIFFERENTIAL Correspondence: GROWTH RATES FOR EXISTING PARTS ACROSS THE TRILOBITE BODY plan. Furthermore, BODY SIZE DATA FOR A LARGE SAMPLE OF [email protected] SPECIMENS OF THE TRILOBITE SPECIES AulacopleurA KONINCKII (Barrande, 1846) MAKE IT POSSIBLE TO GENERATE ROBUST ESTIMATES FOR MODEL PARameters. Although THE GENERATIVE MODEL DESCRIBED HERE WAS BASED ON A RELATIVELY SIMPLE SEGMENTATION schedule, A DIVERSE ARRAY OF OBSERVED BODY SIZES AND RELATIVE PROPORTIONS OF BODY REGIONS CAN BE Open Data ATTAINED BY ALTERING ONLY A FEW PARAMETERS AT A time. -
Morphology and Developmental Traits of the Trilobite Changaspis Elongata from the Cambrian Series 2 of Guizhou, South China
Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China GUANG-YING DU, JIN PENG, DE-ZHI WANG, QIU-JUN WANG, YI-FAN WANG, and HUI ZHANG Du, G.-Y., Peng, J., Wang, D.-Z., Wang, Q.-J., Wang, Y.-F., and Zhang, H. 2019. Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China. Acta Palaeontologica Polonica 64 (4): 797–813. The morphology and ontogeny of the trilobite Changaspis elongata based on 216 specimens collected from the Lazizhai section of the Balang Formation (Stage 4, Series 2 of the Cambrian) in Guizhou Province, South China are described. The relatively continuous ontogenetic series reveals morphological changes, and shows that the species has seventeen thoracic segments in the holaspid period, instead of the sixteen as previously suggested. The development of the pygid- ial segments shows that their number gradually decreases during ontogeny. A new dataset of well-preserved specimens offers a unique opportunity to investigate developmental traits after segment addition is completed. The ontogenetic size progressions for the lengths of cephalon and trunk show overall compliance with Dyar’s rule. As a result of different average growth rates for the lengths of cephalon, trunk and pygidium, the length of the thorax relative to the body shows a gradually increasing trend; however, the cephalon and pygidium follow the opposite trend. Morphometric analysis across fourteen post-embryonic stages reveals growth gradients with increasing values for each thoracic segment from anterior to posterior. The reconstruction of the development traits shows visualization of the changes in relative growth and segmentation for the different body parts. -
1414 Hughes.Vp
The depositional environment and taphonomy of the Homerian Aulacopleura shales fossil assemblage near Lodìnice, Czech Republic (Prague Basin, Perunican microcontinent) NIGEL C. HUGHES, JIØÍ KØÍ, JOSEPH H.S. MACQUAKER & WARREN D. HUFF Excavation of Joachim Barrande’s classic fossil locality of the “Aulacopleura shales” exposed on Na Černidlech Hill, near Loděnice reveals that most specimens were recovered from a 1.4 m interval exposed in “Barrande’s pits”. These are located at the eastern end of a 0.4 km trench dug in the mid 1800’s to expose the interval along strike. Over an hundred bedding planes occur within the 1.4 m interval, and thousands of articulated trilobites have been collected at the site. In- dividual bed surfaces vary in the density, size, and taxonomic composition of the fossils contained. Some preserve a di- verse benthic shelly fauna, others are almost exclusively dominated by the trilobite Aulacopleura koninckii, and a third variety is apparently barren of all shelly fossils. Isolated sclerites of A. koninckii are rare, and on almost all bedding sur- faces exoskeletons are predominantly partially articulated and lack both alignment and sclerite fragmentation. The oc- currence of A. koninckii conforms in many ways to the characteristics of a Type I trilobite lagerstätte of Brett et al. (2012). The presence of enrolled A. koninckii suggests that final burial may have resulted from relatively rapid obrution, although the condition of partial articulation indicates that many carcasses or exuviae partially disaggregated before burial. The mean size and density of A. koninckii specimens varies markedly among bedding planes, with some assem- blages entirely comprised of juveniles, suggesting that notably dense trilobite clustering was not restricted only to repro- ductively mature individuals. -
Instituto De Geociências Revisão Sistemática E
UNIVERSIDADE DE SÃO PAULO INSTITUTO DE GEOCIÊNCIAS REVISÃO SISTEMÁTICA E PALEOBIOGEOGRÁFICA DE TRILOBITAS PHACOPIDA (HOMALONOTIDAE E CALMONIIDAE) DO DEVONIANO DAS BACIAS DO PARNAÍBA E AMAZONAS, BRASIL Felipe van Enck Meira Tese apresentada ao Programa de Pós- Graduação em Geoquímica e Geotectônica do Instituto de Geociências da Universidade de São Paulo, como parte dos requisitos para a obtenção do título de doutor. Orientadora: Prof. Dra. Juliana de Moraes Leme TESE DE DOUTORAMENTO Programa de Pós-graduação em Geoquímica e Geotectônica São Paulo 2016 “Kites rise highest against the wind - not with it.” - Winston Churchill Agradecimentos Agradeço a Deus, por sempre iluminar o caminho durante esses anos de altos e baixos do Doutorado. Agradeço a Ele também por enviar dois verdadeiros anjos da guarda à minha vida – minha esposa Angela Faleiros van Enck Meira e meu filho, Thomas Faleiros van Enck Meira. Agradeço a meus pais, José Carlos e Sylvia, e à minha irmã, Patrícia, pelo apoio durante a jornada, ainda que à distância, por vezes. Sou grato aos meus sogros, Jair e Lucia, que sempre foram como verdadeiros pais e conselheiros. À FAPESP (Processo n° 2012/07075-3), pelo suporte financeiro, sem o qual o Doutorado não seria viável. À minha orientadora, Drª. Juliana de Moraes Leme, pela orientação, discussões e esclarecimentos pertinentes ao projeto. Ao Doutorando Fabio Carbonaro e ao Dr. Renato Ghilardi (UNESP-Bauru), pela parceria e por discussões importantes na realização do trabalho. À Drª. Niède Guidon (FUMDHAM), pelo empréstimo de fósseis da região de São João Vermelho (Piauí), estudados aqui. Sou grato às seguintes pessoas, por permitirem meu acesso às instituições para visita de acervos, e por sua disponibilidade, atenção e ajuda durante minha permanência: Bushra Hussaini (AMNH), Flávia Alessandra Figueiredo, Mônica de Medina Coeli, Dr. -
Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
The Annals and Magazine of Natural History
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A Case Study of Eyeless, Dimorphic Devonian Trilobites from Poland
[Palaeontology, Vol. 59, Part 5, 2016, pp. 743–751] ANCIENT ANIMAL MIGRATION: A CASE STUDY OF EYELESS, DIMORPHIC DEVONIAN TRILOBITES FROM POLAND † by BŁAZEJ_ BŁAZEJOWSKI_ 1, CARLTON E. BRETT2,ADRIANKIN3, , † † ANDRZEJ RADWANSKI 4, and MICHAŁ GRUSZCZYNSKI 3, 1Institute of Palaeobiology, Polish Academy of Sciences, Twarda 51/55, 00-818, Warszawa, Poland; [email protected] 2Department of Geology, University of Cincinnati, Cincinnati, OH 45221-0013, USA; [email protected] 3‘Phacops’ – Association of Friends of Geosciences, Grajewska 13/40, Warszawa, 02-766, Poland 4Institute of Geology, University of Warsaw, Zwirki_ i Wigury 93, 02-089, Warszawa, Poland Typescript received 15 February 2016; accepted in revised form 8 July 2016 Abstract: We report evidence of one of the oldest known have migrated periodically to shallow marine areas for mass animal migratory episodes in the form of queues of the eye- mating and spawning. The sudden death of the trilobites in less trilobite Trimerocephalus chopini Kin & Błazejowski,_ the queues may have been caused by excess carbon dioxide from the Late Devonian (Famennian) of central Poland. In and hydrogen sulphide introduced into the bottom water by addition, there is evidence for two morphs in this popula- distal storm disturbance of anoxic sediments. This study tion, one with nine segments and the other with ten. We demonstrates the potential for further research on the evolu- infer that these queues represent mass migratory chains tion and ecology of aggregative behaviour in marine arthro- coordinated by chemotaxis, comparable to those observed in pods. modern crustaceans such as spiny lobsters, and further sug- gest that the two forms, which occur in an approximately Key words: animal migration, eyeless trilobite, Phacopinae, 1:1 ratio, may be dimorphs. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
The Viruan (Middle Ordovician) of Öland
The Viruan (Middle Ordovician) of Öland By Valdar Jaanusson ABSTRACT.-The stratigraphy and lithology of the Viruan (Middle Ordovician) Iimestones of the bed-rock of Öland are described based on three bares and on field work in the outcrop area. A combined litho- and bio-stratigraphic classification (termed topo-stratigraphic) is introduced for the described sequence. The names of the Estonian stages (Aserian, Lasnamägian, Uhakuan, and Kukrusean) are used as chrono-stratigraphic references instead of the previous Swedish names of the units of stage category (Platyurus, Schroeteri, Crassicauda, and Ludibundus, re spectivcly). New topo-stratigraphic divisions are Segerstad Limestone (of Aserian age), Skärlöv, Seby, and Folkeslunda Limestones (of Lasnamägian age), Furudal, Källa, and Persnäs Lime stones (of Uhakuan age), and Dalby Limestone (of Kukrusean age in the bed-rock of Öland). The Aserian Lasnamägian topo-stratigraphic divisions have the same lithological characteris and tics throughout Öland. The Uhakuan beds are developed as calcilutites (Furudal Limestone) on southern Öland continuing as a tongue (Källa Limestone) on northern Öland. The middle and upper part of the Uhakuan beds of northern Öland consist of calcarenites (Persnäs Limestone) lithologically indistiguishable from the Kukrusean Dalby Limestone which forms the bed-rock only on northern Öland. Within the Segerstad Limestone two zones are distinguished (z. of Angelinoceras latum and z. of Illaenus planifrons).H ouvr ' s zones of Lituites discors, L. lituus, and L. perfectus are of Lasna mägian age, and their stratigraphic position and fauna! characteristics are described. Contents Introduction . 207 Methods .............. 209 Classification of the Viruan rocks of Öland 2I2 Historical survey . 2I9 Taxonornie and nomenclatural notes 22I Viruan rocks of northern Öland . -
Type and Figured Fossils in the Worthen Collection at the Illinois
s Cq&JI ^XXKUJtJLI 14oGS: CIR 524 c, 2 TYPE AND FIGURED FOSSILS IN THE WORTHEN COLLECTION AT THE ILLINOIS STATE GEOLOGICAL SURVEY Lois S. Kent GEOLOGICAL ILLINOIS Illinois Department of Energy and Natural Resources, STATE GEOLOGICAL SURVEY DIVISION CIRCULAR 524 1982 COVER: This portrait of Amos Henry Worthen is from a print presented to me by Worthen's great-grandson, Arthur C. Brookley, Jr., at the time he visited the Illinois State Geological Survey in the late 1950s or early 1960s. The picture is the same as that published in connection with the memorial to Worthen in the appendix to Vol. 8 of the Geological Survey of Illinois, 1890. -LSK Kent, Lois S., Type and figured fossils in the Worthen Collection at the Illinois State Geological Survey. — Champaign, III. : Illinois State Geological Survey, 1982. - 65 p. ; 28 cm. (Circular / Illinois State Geological Survey ; 524) 1. Paleontology. 2. Catalogs and collections. 3. Worthen Collection. I. Title. II. Series. Editor: Mary Clockner Cover: Sandra Stecyk Printed by the authority of the State of Illinois/1982/2500 II I IHOI'.MAII '.I 'II Of.ir.AI MIHVI y '> 300 1 00003 5216 TYPE AND FIGURED FOSSILS IN THE WORTHEN COLLECTION AT THE ILLINOIS STATE GEOLOGICAL SURVEY Lois S. Kent | CIRCULAR 524 1982 ILLINOIS STATE GEOLOGICAL SURVEY Robert E. Bergstrom, Acting Chief Natural Resources Building, 615 East Peabody Drive, Champaign, IL 61820 TYPE AND FIGURED FOSSILS IN THE WORTHEN COLLECTION AT THE ILLINOIS STATE GEOLOGICAL SURVEY CONTENTS Acknowledgments 2 Introduction 2 Organization of the catalog 7 Notes 8 References 8 Fossil catalog 13 ABSTRACT This catalog lists all type and figured specimens of fossils in the part of the "Worthen Collection" now housed at the Illinois State Geological Survey in Champaign, Illinois. -
2Nd International Trilobite Conference (Brock University, St. Catharines, Ontario, August 22-24, 1997) ABSTRACTS
2nd International Trilobite Conference (Brock University, St. Catharines, Ontario, August 22-24, 1997) ABSTRACTS. Characters and Parsimony. Jonathan M. Adrain, Department of Palaeontology, The Natural History Museum, London SW7 5BD, United King- dom; Gregory D. Edgecombe, Centre for Evolutionary Research, Australian Museum, 6 College Street, Sydney South, New South Wales 2000, Australia Character analysis is the single most important element of any phylogenetic study. Characters are simply criteria for comparing homologous organismic parts between taxa. Homology of organismic parts in any phylogenetic study is an a priori assumption, founded upon topological similarity through some or all stages of ontogeny. Once homolo- gies have been suggested, characters are invented by specifying bases of comparison of organismic parts from taxon to taxon within the study group. Ideally, all variation in a single homology occurring within the study group should be accounted for. Comparisons are between attributes of homologous parts, (e.g., simple presence, size of some- thing, number of something), and these attributes are referred to as character states. Study taxa are assigned member- ship in one (or more, in the case of polymorphisms) character-state for each character in the analysis. A single char- acter now implies discrete groupings of taxa, but this in itself does not constitute a phylogeny. In order to suggest or convey phylogenetic information, the historical status of each character-state, and of the the group of taxa it sug- gests, must be evaluated. That is, in the case of any two states belonging to the same character, we need to discover whether one state is primitive (broadly speaking, ancestral) or derived (representative of an evolutionary innovation) relative to the other. -
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More By Michelle M. Casey1, Perry Kennard2, and Bruce S. Lieberman1, 3 1Biodiversity Institute, University of Kansas, Lawrence, KS, 66045, 2Earth Science Teacher, Southwest Middle School, USD497, and 3Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Middle level laboratory exercise for Earth or General Science; supported provided by National Science Foundation (NSF) grants DEB-1256993 and EF-1206757. Learning Goals and Pedagogy This lab is designed for middle level General Science or Earth Science classes. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They only lived in the oceans but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter across. They were once one of the most successful of all animal groups and in certain fossil deposits, especially in the Cambrian, Ordovician, and Devonian periods, they are extremely abundant.