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(Reptilia: Archosauromorpha) from the Upper Triassic of West Texas and Its Biochronological Significance Andrew B

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(Reptilia: Archosauromorpha) from the Upper Triassic of West Texas and Its Biochronological Significance Andrew B New Mexico Geological Society Downloaded from: http://nmgs.nmt.edu/publications/guidebooks/52 New occurrence of Trilophosaurus (Reptilia: Archosauromorpha) from the Upper Triassic of West Texas and its biochronological significance Andrew B. Heckert, Spencer G. Lucas, Robert Kahle, and Kate Zeigler, 2001, pp. 115-122 in: Geology of Llano Estacado, Lucas, Spencer G.;Ulmer-Scholle, Dana; [eds.], New Mexico Geological Society 52nd Annual Fall Field Conference Guidebook, 340 p. This is one of many related papers that were included in the 2001 NMGS Fall Field Conference Guidebook. Annual NMGS Fall Field Conference Guidebooks Every fall since 1950, the New Mexico Geological Society (NMGS) has held an annual Fall Field Conference that explores some region of New Mexico (or surrounding states). Always well attended, these conferences provide a guidebook to participants. Besides detailed road logs, the guidebooks contain many well written, edited, and peer-reviewed geoscience papers. These books have set the national standard for geologic guidebooks and are an essential geologic reference for anyone working in or around New Mexico. Free Downloads NMGS has decided to make peer-reviewed papers from our Fall Field Conference guidebooks available for free download. Non-members will have access to guidebook papers two years after publication. Members have access to all papers. This is in keeping with our mission of promoting interest, research, and cooperation regarding geology in New Mexico. However, guidebook sales represent a significant proportion of our operating budget. Therefore, only research papers are available for download. Road logs, mini-papers, maps, stratigraphic charts, and other selected content are available only in the printed guidebooks. Copyright Information Publications of the New Mexico Geological Society, printed and electronic, are protected by the copyright laws of the United States. No material from the NMGS website, or printed and electronic publications, may be reprinted or redistributed without NMGS permission. Contact us for permission to reprint portions of any of our publications. One printed copy of any materials from the NMGS website or our print and electronic publications may be made for individual use without our permission. Teachers and students may make unlimited copies for educational use. Any other use of these materials requires explicit permission. This page is intentionally left blank to maintain order of facing pages. New Mexico Geological Society Guidebook, 52nd Field Conference, Geology of the Llano Estacado, 200 I 115 NEW OCCURRENCE OF TRILOPHOSAURUS (REPTILIA: ARCHOSAUROMORPHA) FROM THE UPPER TRIASSIC OF WEST TEXAS AND ITS BIOCHRONOLOGICAL SIGNIFICANCE ANDREW B. HECKERT', SPENCER G. LUCAS2, ROBERT KAHLE3 AND KATE ZEIGLER' 1Department of Earth and Planetary Sciences, University of New Mexico, Albuquerque, NM 87131-11 16; l'New Mexico Museum of Natural History, 1801 Mountain Rd NW, Albuquerque, 87104; 34305 Roosevelt Road, Midland. Texas 79703 Abstract.- We document a rich Upper Triassic bonebed in Borden County, Texas in either the uppermost Tecovas Formation or, more likely, the Trujillo Formation ofthe Chinle Group, that represents one ofthe highest stratigraphic occurrences of the aberrant archosauromorph reptile Trilophosaurus. The associated fauna includes indeterminate coprolites, unionid bivalves, osteichthyans, indeterminate reptiles, a possible ornithischian dinosaur, and the putative theropod dinosaur Spinosuchus caseanus Huene. INTRODUCTION NMMNH3775 Numerous workers have collected and described Upper Trias­ :~~~~~neerate sic fossils from various localities in West Texas, including Cope siltstone (1892), Case (1922, 1928, 1932), Gregory (1945), Sawin (1947), mudstone/shale 6 Murry (1982, 1986) and Chattetjee (1986) (Fig. 1). Among the limestone D-:3~ fossil bone fossils collected were the original specimens of the aberrant archosauromorph reptile Trilophosaurus buettneri Case, 1928, 5 1 meter subsequently the subject of a monograph by Gregory (1945) and described further by Parks (1969) and Murry (1982). To date, all other verifiable Trilophosaurus records come from low in 4 the Chinle Group in Texas (Gregory, 1945; Parks, 1969; Murry, 1982) or in Arizona and New Mexico (Long and Murry, 1995), 3 with the exception of isolated teeth collected by Kirby {1989, \_ 2 1991 , 1993) in the Owl Rock Formation of north-central Arizona. .-i NMMNH locality 3775 Here, we outline the stratigraphy ofa new locality yielding fossils of Trilophosaurus, describe the fauna of this locality, and com­ ment on its biostratigraphic and biochronologic significance. In this paper MNA = Museum of Northern Arizona, Flagstaff; and NMMNH = New Mexico Museum of Natural History and Science, Albuquerque. FIGURE I. Index map and measured stratigraphic section ofNMMNH locality 3775. For a description of stratigraphic units in the section see STRATIGRAPHY the Appendix. Detailed locality information is on file at NMMNH. One of us (RK) discovered New Mexico Museum ofNatural History and Science (NMMNH) locality 3775 while working Formation of the Chinle Group (Lucas et al., 1994, 2001). How­ in West Texas in the early 1990s (Fig. 1). This locality yields ever, the possibility exists that these strata instead represent an numerous pinkish-gray disarticulated bones from a matrix of outcrop of the more areally limited coarse-grained channel bodies intraformational conglomerate. The principal fossiliferous hori­ within the stratigraphically lower Tecovas Formation (Lucas et zon consists of a moderate brown, clast-supported conglomerate al., 1994 and sources cited therein). Because these are some of of rounded, flattened, intraformational mud pebbles up to 2 em in the only outcrops in an area characterized by broad flatlands, we diameter. The matrix is principally a rounded, moderately poorly remain unsure as to the exact stratigraphic position of these strata. sorted sublitharenite. This horizon is separated from the underly­ However, we suspect that these deposits represent the base of ing moderate brown mudstone by an irregular contact that sug­ the Trujillo Formation locally, and summarize arguments both for gests slight scouring and erosion (Fig. 1). Most of the overlying and against that assignment in the following paragraphs. strata are interbedded sandstone, conglomerate and mudstone. Trujillo Formation strata overlie a regional erosional surface Unionid bivalves were collected from the uppermost conglomer­ termed the Tr-4 unconformity by Lucas (1993) that approximates ate, which is a moderate yellowish-brown intraformational con­ the Carnian-Norian boundary. Although exposure in the vicinity glomerate dominated by clasts of calcrete and siltstone up to 1.5 of NMMNH locality 3775 is poor, strata to the north and west em in diameter. clearly pertain to the Bull Canyon Formation, which overlies the The bedded intraformational conglomerates that dominate this Trujillo throughout West Texas and east-central New Mexico. outcrop strongly resemble the dominant lithologies of the Trujillo Similarly, the Trujillo Formation throughout this region is consid- 116 HECKERT, LUCAS, KAHLE AND ZEIGLER A s-c --- D-E G-H----- FIGURE 2. Photographs ofrepresentative fossils from NMMNH locality 3775. A-E. Trilophosaurus sp., A. NMMNH P-29959, maxillary? fragment in occlusal view; B-C. NMMNH P-34101 left humerus in (B) lateral and (C) medial views, D-E. NMMNH P-34102, right tibia in lateral (D) and medial (E) views; F-H. NMMNH P-29954, indeterminate uniorud bivalves, incomplete valves in interior (F) and exterior (G-H) views. All scales= 2 em. erably more resistant than either the overlying Bull Canyon For­ fore, we tentatively assign NMMNH locality L-3775 to the Tru­ mation or the underlying Tecovas Formation. Thus, the Trujillo jillo Formation. Formation crops out as either a ledge-former above Tecovas For­ mation badlands, such as at Muchakooyoo Peak slightly to the PALEONTOLOGY southeast or, alternately, as a bench flooring broad plains beneath localized exposures and badlands of the Bull Canyon Formation. We have collected numerous macro- and microvertebrate fos­ The strata at NMMNH locality L-3775 appear to represent the sils from NMMNH locality 3775. Although many of these fos­ latter, as they are part of a broad upland surface that extends from sils have yet to be prepared, it is evident that this locality yields a Gail to the northeast southward to just northwest of Westbrook. number of tetrapod taxa, dominated by the archosauromorph rep­ The only other possibility is that these strata represent a chan­ tile Trilophosaurus (Fig. 2). The associated fauna includes inde­ nel deposit in the Tecovas Formation. However, we note here that terminate coprolites, unionid bivalves, osteichthyans, indetermi­ the broad fiats of which this outcrop is a part suggest that it is nate reptiles, a possible omithischian dinosaur, and probable the­ not composed of isolated channel-fill lithologies, and instead rep­ ropods, including a possible occurrence of the putative theropod resents more regional deposition of coarse-grained strata. There- dinosaur Spinosuchus caseanus Huene. TRILOPHOSAURUS AND ITS BIOCHRONOLOGICAL SIGNIFICANCE 117 Trilophosaurus Trilophosaurus and appears to be unique to Tri/ophosaurus spec­ imens from NMMNH locality 3775. As with the tooth spacing, We have collected abundant material of Trilophosaurus from this may have taxonomic significance, although we refrain from NMMNH locality 3775; these fossils include skull or
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