Lista Faunistica De Los Hydradephaga De La Península Ibérica Y Baleares

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Lista Faunistica De Los Hydradephaga De La Península Ibérica Y Baleares ASOCIACION ESPAÑOLA DE LIMNOLOGIA Lista faunística y bibliográfica de los Hydra­ dephaga (Coleoptera: Haliplidae, Hygrobiidae, Gyrinidae, Noteridae, Dytiscidae) de la Penín• sula Ibérica e Islas Baleares E. RICO L. C. PEREZ C. MONTES USTAS DE LA FLORA Y FAUNA DE LAS AGUAS CONTINENTALES DE LA PENINSULA mERICA PUBLICACION N! 7 • 1990 LISTA FAUNISTICA Y BffiLIOGRAFICA DE LOS HYDRADEPHAGA (COLEOPTERA: HALIPLI­ DAE, HYGROBIIDAE, GYRINIDAE, NOTERI­ DAE, DYTISCIDAE) DE LA PENINSULA IBERICA E ISLAS BALEARES por EUGENIO RICO Dpto. Biología Animal y Genética Universidad del País Vasco. Bilbao LUIS CARLOS PEREZ y CARLOS MONTES Dpto. lnteruniversitario de Ecología. Madrid ASOCIACION ESPAÑOLA DE LIMNOLOGIA 1990 Depósito Legal M-40307-1990 I.S.B.N.: 84-404-7987-5 Impreso en FASTER S.Francisco de Sales, 1 28003 MADRID INDICE INTRODUCCION •••••••• , , ...... , .. , ... , • , , , • , , , •• , , • , , .. , • , , •• , • , , • , , •• , •• Familia HALIPLIDAE •••••••••••••••••••••••••••••••••••••••••• , •• , , • , 3 Género Peltodytes •••••••••••••••••••••••••••••••••••••••••••• 3 Género Brychius •••••••••••••••••••••••••••••••••••••••••••••• 5 Género Haliplus •••••••••••••••••••••••••••••••••••••••••••••• 6 Familia HYGROBIIDAE ............................................... 19 Género Hygrobia ••••••••••••• , • , ••••••••••••••••••••• , ••••••• 19 Familia GYRINIDAE ............. , ................................... 21 Subfamilia GYRININAE •••••••••••••••••••••••• , •••••••••••••••••• 21 Género Aulonogyrus ••••••••• , •••••• , ••••••••••••••••••••••••• 21 Género Gyrinus • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 23 Subfamilia ORECTOCHILINAE , ... , •••••••• , ........................ 31 Género Orectochilus ••••••• , ••••• , • , ••••••••• , ••••••••••••••• 31 Familia NOTERIDAE ••••••••••••••••••• , ••••••••••••••••••••• , •••••• , 33 Género Canthydrus ••••••••••••••••••••••••••••••••••• , , ••• , •• 33 Género Note rus ••••••••••••••••••••••••••••••••••••••• , , ••••• 33 Familia DYTISCIDAE • , .......... , ••••• , •• , ................... , .. , ... 37 Subfamilia METHLINAE .......................................... , 37 Género Hethles .............................................. 37 Subfamilia HYDROPORINAE •••••••••••••• , ......................... 37 Género Hyphydrus •••••••••••••••••••••••••••••••••••••••••••• 37 Género Hydrovatus ••••••••••••••••••••••••••••••••••••••••••• 39 Género Yola • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 42 Género Bidessus ••••••••••••••••••••••••••••••• , •••••••••• , • • 43 Género Hydroglyphus ••••••••••••••••••••••••••••••••••••••••• 48 Género Coelambus •••••••••••••••••••••••••••••••••••••••••••• 51 Género Hygrotus •••••••••••••••••••••••••••••••••••••• , •••••• 56 Género Herophydrus •••••••••••••••••••••••••••••••••••••••• , • 59 Género Suphrodytes •••••••••••••••••••••••••••••••••••••••••• 59 Género Hydroporus ••••••••••••••••••••••••••••••••••••••••••• 60 Género Porhydrus •••••••••••••.••••••••••••••••••••••••••••••• 84 Género Graptodytes •••••••••••••••••••••••••••••••••••••••••• 86 Género Rhithrodytes ••••••••••••••••••••••••••••••••••••••••• 92 Género Hetaporus •••••••••••••••• , •••••••••••••••••••• , •• , , • • 93 Género Scarodytes • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 94 Género Stictonectes ••••••••••••••••••••••••••••••••••••••••• 95 Género Deronectes •••••••••••••••••••••••••••••••••••••••••• 100 Género Stictotarsus ••••••••••••••• , •••••••••••••••••••••••• 109 Género Potamonectes ••••••••••• , •••••••••••••••••••••••••••• 110 Género Oreodytes • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 117 Subfamilia LACCOPHILINAE •••••••••••••••••••••••••••••••••••••• 119 Género Laccophil us • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 119 Subfamilia COLYMBETINAE ••••••••••••••••••••••••••••••••••••••• 123 Género Copela tus ........................................... 123 Género Platambus ........................................... 125 Género Agabus •••••••••••••••••••••••••••••••••••••••••••••• 126 Género Ilybius ••••••••••••••••••••••••••••••••••••••••••••• 146 Género Rhantus • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 151 Género Colymbetes •••••••••••••••••••••••••••••••••••••••••• 154 Género Heladema ............................................ 158 Subfamilia DYTISCINAE ••••••••••••••••••••••••••••••••••••••••• 159 Género Eretes •••••••••••••••••••••••••••••••••••••••••••••• 160 Género Hydaticus ........................................... 161 Género Graphoderus ••••••••••••••••••••••••••••••••••••••••• 163 Género Acilius ••••••••••••••••••••••••••••••••••••••••••••• 164 Género Dytiscus ............................................ 166 Género Cybister •••••••••••••••••••••••••••••••••••••••••••• 172 RELACION GENERAL DE ESPECIES • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 176 BIBLIOGRAFIA • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 186 INDICE ALFABETICO DE NOMBRES CIENTIFICOS ••••••••••••••••••••••••••••• 209 INTRODUCCION El grupo de coleópteros acuáticos agrupados bajo la denominación, sin categoría taxonómica, de Hydradephaga, y que está representado en nuestro área de estudio por las familias Haliplidae, Hygrobiidae, Gyrinidae, Noteridae y Dytiscidae, ha recabado tradicionalmente la atención de numerosos estudiosos; sin duda, tanto su gran capacidad de adaptación a diferentes ambientes acuáticos como sus llamativas pautas de comportamiento, son factores que han contribuido a ello. Así, nos encontramos con un grupo, en general, bien conocido taxonómicamente y que ha dado lugar en numerosos países europeos a obras generales de recopilación y determinación sistemática (Guignot 1931- 33, 194 7; Balfour-Browne, 1940, 1950; Zaitsev, 1953; Freude, 1971; Franciscolo, 1979; Holmen, 1987). La Península Ibérica no ha escapado a este interés. Ya desde mediados del siglo XIX existen referencias de esta taxocenosis, muchas de ellas resultado de viajes de naturalistas extranjeros (Rosenhauer, 1856; Heyden, 1870, 1880). El interés de los investigadores extranjeros a seguido a partir de entonces de forma continua, destacando trabajos como los de Champion, Breit, Falkenstrom, Lindberg, Bertrand, Franciscolo, y recientemente los de Brancucci, Fery, etc. El movimiento naturalista espaHol, que se formó en torno a la Sociedad EspaHola de Historia Natural y la Institución Catalana de Historia Natural, también realizó numerosas contribuciones al conocimiento de este grupo, sobre todo en las zonas de CataluHa, Levante y Baleares. En la actualidad son numerosas las personas o equipos distribuidos por gran parte de la geografía peninsular que continúan estudiando este grupo taxonómico. El primer catálogo donde se recopilaron estos insectos en todo el área de estudio que nos ocupará es el de Fuente (1921), si bien existian otros como el de Oliveira (1894), para el área portuguesa, y los de Cardona (1872), Estelrich et al. (1885), Moragues (1889) y Tenenbaum (1915), para las Baleares; posteriormente aparecen, para la zona espaHola, el de Pardo (1932, 1933), y los de Seabra (1943) y Serrao & Azevedo (1970), para la zona portuguesa. A pesar de todo, no existe todavía ninguna obra que realice una revisión sistemática conjunta. La necesidad de este trabajo se hace más importante, si cabe, en tanto en cuanto la Península Ibérica resulta ser una encrucijada de faunas de distinta procedencia en la que, además, existen numerosos endemismos, algunos de los cuales son considerados por los científicos de dudosa validez sistemática. La elaboración de esta lista faunistica tiene como 2 fin el servir de base al desarrollo de ese trabajo que se presenta como una asignatura pendiente desde hace décadas, necesaria de abordar. La estructura de esta lista faunistica y bibliográfica es, básicamente, la misma que la seguida en los volúmenes anteriores de esta serie. El área de estudio abarca la peninsula, considerando como limite septentrional la frontera franco-española, Andorra, y las Islas Baleares. Para cada especie se señala la descripción original, asi como sinonimias y variedades descritas en nuestro área de estudio. A continuación se presentan aquellos trabajos básicos que permitan su identificación sistemática. La información faunistica se ha organizado, dentro de lo posible, por provincias, incluyendo una referencia a su distribución geográfica general. En ocasiones se añade un apartado de observaciones en donde se comentan aspectos de diversa indole. 3 Familia HALIPLIDAE Thomson, 1860. Genero PELTODYTES Régimbart, 1878. Peltodytes (Peltodytes) caesus (Duftschmid, 1805). Dytiscus caesus DUFTSCHMID, 1805: 284. Dytiscus impressus PANZER, 1794: pl. 7. Cnemidotus caesus: AUCTT •• Cnemidotus impressus: AUCTT •• Peltodytes impressus: AUCTT •• Peltodytes caesus; GUIGNOT 1931-33: 201-203, figs. 84, 86, 88; pl. 1, fig. l. Peltodytes caesus; GUIGNOT 1947: 36-37, figs. 15 bis A. Peltodytes caesus; GUIGNOT 1959-61: 21-23, figs. 3, 5, 8. Peltodytes caesus; FRANCISCOLO 1979: 69, figs. 26a, 39-40, 44, 52, 54-55, 58, 85, 95-96, 120, 140, 171, 189. Peltodytes caesus; HOLHEN 1987: 88-89, figs. 128-130, 143-148, pl. 1:1. ES PARA: Catalufta: Fuente 1921. Regi6n valenciana: Boscá 1916. Alicante: Torres Sala 1962. Barcelona: Lagar 1951, 1968; Sabater et al. 1986. Castell6n: Pardo 1924. Ciudad
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