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Cicadina10:3369(2009) 33

AnannotatedcatalogueoftheAuchenorrhynchaof NorthernEurope (Insecta,:FulgoromorphaetCicadomorpha)

GuySöderman1,GöstaGillerfors2,AndersEndrestöl3

Abstract:AnannotatedcatalogueoftheandofNor thern Europe, with marked occurrences for each country (Iceland, Norway, DenmarkwithoutGreenland,Sweden,Estonia,Latvia,Lithuania)andadminis trativeregionsofadjoiningRussia(Kaliningrad,Murmansk,Karelia,St.Peters burgandPskov)ispresented.Thecatalogueincludesaltogether513specieswith comments on several new species hitherto unrecorded. The recent northward expansionofsomeCentralEuropeanspeciesisbrieflydiscussed. Zusammenfassung:Kommentierte Artenliste der Zikaden Nordeuropas (Insecta, Hemi- ptera, Fulgoromorpha et Cicadomorpha).–EswirdeinekommentierteArtenlisteder ZikadenNordeuropasvorgelegt,mitAngabenderNachweisefürdienordischen und baltischen Staaten (Island, Norwegen, Dänemark ohne Grönland, Schwe den, Estland, Lettland, Litauen) und die angrenzenden Verwaltungsregionen vonRussland(Kaliningrad,Murmansk,Karelien,St.PetersburgundPskov).Die Listeenthält513ArtenmiteinigenunveröffentlichtenNeufunden.DieNord ausbreitungmittleuropäischerArteninjüngererZeitwirdkurzdiskutiert. Keywords:faunalchecklist,leafhoppers,planthoppers,NordicandBalticcoun tries,NorthwestRussia

1.Introduction No catalogue of Northern European Auchenorrhyncha (hereafter shortly named “hop pers”)haspreviouslybeenpublished.Existingcataloguescoveringpartsofthearea(Vil baste 1974, Ossiannilsson 1983) have become old and new species have been found in more thorough taxonomic research and faunistic inventories. New trapcollecting tech niqueshaveincreasedthematerialinmanyofthecountriesinthe21stcenturyandrevealed anorthwardexpandingtrendofmanyspecieslivingincentralEurope.Wehereintendto updatetheknowledgeofhopperrecordsofthecountriesandotheradministrativeareasin NorthernEuropeandtodiscusssomeoftheexpandingtrendsinthefauna.

2.Geographicconcept TheconceptofNorthernEuropeincludesinthisworkalltheNordiccountries,viz.Ice land,Norway,Sweden,FinlandandDenmark(excepttheautonomicareaofGreenland),all theBalticcountries,viz.Estonia,LatviaandLithuania,andtheadjacentRussianareasto

1FinnishEnvironmentInstitute,Helsinki,Finland;email:[email protected] 2Maskrosvägen23,Varberg,Sweden;email:[email protected] 3NorwegianInstituteforNatureResearch,Oslo,Norway;email:[email protected] ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

34 G.Söderman,G.Gillerfors,A.Enderstöl

Fig.1:Administrativeareasofthecatalogue,Icelandomitted.Forabbreviationsofgeographicre gionsseechapter4.1. thesepreviouslymentionedcountries,viz.theMurmanskregion(oblast),RussianKarelia (autonomousrepublic),theLeningradregion(incl.St.PetersburgandtheIsthmusofKare lia),thePskovregion(oblast)andtheenclaveofKaliningrad(seeFig.1).Byelorussia,bor deringinthewesttoLithuania,hasbeenleftout,becauseachecklistincluding331hopper speciesrecordedinthiscountryhasquiterecentlybeenpublishedbyBorodin2004).

3.Presentknowledgebase Iceland Nonationalcollecting activityofhoppershasdeveloped(ErlingOlafsson,pers.comm.). Most data stem from two sources, the collecting of the Swedish entomologist C.H. Lin droth(reportedbyOssiannilsson1947b)andtheGermanentomologistM.v.Tschirnhaus in2001(reportedbyRemane,2005).ThetotalknownspeciesnumberfromIcelandthus sumsonlyto8. Norway The bulk of previous hopper information has been compiled by H. Holgersen (1944a, 1944b,1945,1946,1954)andF.Ossiannilsson(1943,1962,1974,1977,1978,1981,1983). RecentcollectingactivitiesinNorwayarerelativelyhighincludingcollectingwithdifferent ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 35 traptechniques.Publicationsofnewspecieshavealsobeenfrequent(Huldén1982,Hol gersen 1985, 1992, Raatikainen &Ylönen 1988, Hansen & Borgersen 1991, Olsen 1999, Hansen2000,Endrestöl2008).Asoftoday291hopperspecieshavebeenrecorded. Denmark Most hopper recordings in Denmarkwere made by N.P. Kristensen (1965a, 1965b) and L.Trolle(1966,1968,1973,1974,1982)andhavebeencompiledintheappendixofOs siannilssons book series (1983). The recentmost list of Danish hoppers was given by L.Skipper(2001)andincludesnonewspeciessinceOssiannilssonswork.Someadditional recordsofexpandingspecieshavebeenpublishedontheweb.Thereisaverymodesthop percollectingactivityinDenmarkatpresent.TheknownhoppernumberforDenmarkis now295. Sweden ThebasisforhopperknowledgeinSwedenhasbeencompiledbyF.Ossiannilsson(1947a, 1978,1981,1983).Afterwardsrecordsofnewspeciesandregionalfindingshavebeenpub lishedbyG.Gillerfors(2002,2003,2005,2008).Therecentcollectingactivityislow,but Swedish trappingprojects haveproducedasubstantialamountofnewmaterialofwhich onlyaparthasbeenanalysedyet.Swedenholdsthehighestnumberofrecordedhoppersin NorthernEuropewith417species. Finland RegularlistsofhoppershavebeenproducedformorethanacenturyinFinland(Sahlberg 1871,Lindberg1935,1947,Linnavuori1969a,1969b,Huldén&Heikinheimo1984).The latestoverviewwasgivenbyG.Söderman(2007).Collectingactivityhasincreasedsincethe establishmentoftheExpertGrouponHemiptera.Extensivetrappingschemesincluding lighttraps(seeSöderman2005),Malaisetraps,colorpansandpitfalltraps(seeSöderman 2007)havebeenusedinthefaunisticinventoriesofthe21stcentury.Thenumberofspe ciesrecordednowamountsto401. Estonia TheEstonianhopperfaunawassurveyedformanyyearsbyJuhanVilbaste,whostartedto publishanationalbookseriesofthegroup(1971).Itwas,unfortunately,nevercompleted, butrecordsofEstonianspecieswerelatergivenbyhiminhisarticleontheLatvianand Lithuanianfauna(1974).Inthelate20thcenturyandinthebeginningofthe21stcentury someFinnishentomologisthavecollectedasmallmaterialofhoppersalsofromnorthern Estonia,butthismaterial,locatedintheZoologicalMuseumofHelsinki,doesnotinclude manynewspecies.CollectingactivityinEstoniaisnowadaysmodest.Thenumberofspe ciesrecordedfromEstoniaisnow305. Latvia ThebasicknowledgeofthehopperfaunawasgivenbyVelce&Danka(1970).Later,Vil baste(1974)revisedsomeolddataandaddedasubstantialamountofnewspeciesbasedon hisfieldcollectinginthe1960´s.SincethenonlyVarzinska(1983)haspublishedonLatvian leafhoppers.Thelatestlistpublishedontheweb(Spungis&Kalnins2002)doesnotac knowledgeVilbaste´sworkandisthereforemisleading.Thereisnoactivityonhopperre searchin thecountry. However,hopperrecordsfrompitfall trappingofthe coastalgrey ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

36 G.Söderman,G.Gillerfors,A.Enderstöl dunes in 2006 have recently been published (Söderman & Spungis 2008). Presently, 333 speciesareknownfromLatvia. Lithuania OldknowledgeofhoppersisfoundinFlor(1861)whoidentifiedmaterialfromthecollec tionsofGimmerthal(1846).SomeofFlor´srecordshavebeenrevisedbyVilbaste(1973) whoalsoundertookextensivefieldcollectionsinLithuaniainthe1960´s.Theresultswere publishedmuchlater(Vilbaste1974).Morerecentdataonlystemfromlighttrappinginthe CepkeliaiNatureReservein2007(Söderman&Dapkus,unpublished)andshortfieldvisits in the southeastern part of the country (Söderman & Rintala, unpublished). The species numberforLithuaniaisnow331. NorthwesternRussia OldinformationonhopperrecordsfromNorthwesternRussiaisdifficulttofindinlitera turebecausetheformerinformationbaseonlyincludedmajorsubdivisionsoftheSoviet Union:NorthwesternRussiaextendingeasttotheKomiRepublic,WesternRussiainclud ing the Baltic countries, and Southwestern Russia including parts of Byelorussia and Ukraine(seee.g.Emeljanov1964).Recordsfromparts,whichinprewartimesbelongedto FinlandsuchasPetsamointheMurmanskregion(Lindberg1932),Kareliaproperandthe IsthmusofKarelia(Sahlberg1871,Lindberg1947)areavailableforthesubdivisionused here. Furthermore, the new inventories made by Anufriev (2000a, 2000b) and Dmitriev (2002)intheMurmanskregionareveryuseful.Thenumberofhopperrecordsforother different administrative regions in this catalogue is low, as present collecting interest in theseregionshasbeenlow.Manyoftherecordsinthecataloguehavebeenproducedwith trappingusedincooperativemonitoringbetweenFinlandandRussiain19932008. TheonlyreportofthisactivityhasbeenpresentedbySöderman&Goltsova(2005).Asof now104speciesareknownfromKaliningrad,allrecordedbyA.P.Shapoval&Söderman in20072008,108fromtheMurmanskarea,192fromRussianKarelia,171fromtheLen ingradregionand109fromthePskovregion.

4.Thecatalogue 4.1Introductoryremarks NomenclatureandsystematicsmainlyfollowNast(1987)withsomelateramendmentsof Nickel(2003)andothers.Speciesrecordsintheareashavebeenmarkedwith“X”.Coun triesandregionsincolumnheadsaregivenasinternationallyacceptedabbreviations,viz. IC=Iceland,NO=Norway,DK=Denmark(withoutGreenland),SE=Sweden,FI= Finland,EE=Estonia,LV=Latvia,LT=Lithuania, KG = Kaliningrad, MM = Mur mansk,KR=Karelia,LG=St.PetersburgandtheLeningradregionandPV=Pskov.Re cordsofuncertaintyhavebeenmarkedwithanadditionalquestionmarkintherespective columns. ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 37

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV FULGOROMORPHAEvans,1946 CIXIIDAESpinola,1839 CixiiniSpinola,1839 1 Cixius nervosus (Linnaeus,1758) . X X X X X X X . X? X X . 2 Cixius cunicularius (Linnaeus,1767) . X X X X X X X X X X X X 3 Cixius distinguendus Kirschbaum,1868 . X X X X X X X . . X X X 4 Cixius cambricus China,1935 . . . . X . . . . . . . . 5 Cixius similis Kirschbaum,1868 . X X X X X X X . X X X . 6 Cixius stigmaticus (Germar,1818) . X X X . . X? . . . . . . 7 Tachycixius pilosus (Olivier,1791) . . X X . . . X . . . . . PentastiriniEmeljanov,1971 8 Pentastiridius leporinus (Linnaeus,1761) . . X X X X . X . . X . . Leach,1865 AsiracinaeMotschulsky,1863 9 Asiraca clavicornis (Fabricius,1794) . . . . . . X? . . . . . . KelisiinaeWagner,1963 10 Kelisia guttula (Germar,1818) . X X X X X X X . . X X . 11 Kelisia guttulifera (Kirschbaum,1868) . . . . . . . X . . . . . 12 Kelisia vittipennis (J.Sahlberg,1868) . X X X X X X X . . X . . 13 Kelisia pallidula (Boheman,1847) . X X X X X X X . . . . . 14 Kelisia praecox Haupt,1935 . . . . X X X X . . . . . 15 Kelisia confusa Linnavuori,1957 . . . X X . . X . . . . . 16 Kelisia monoceros Ribaut,1934 . X . X X X X X . . . . . 17 Kelisia ribauti Wagner,1938 . X X X X X X X . . X X . 18 Kelisia sabulicola Wagner,1952 . . . X X X X X X . . . . 19 Kelisia sima Ribaut,1932 . . . X . . . . . . . . . 20 Anakelisia fasciata (Kirschbaum,1868) . . X X . . . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

38 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 21 Anakelisia perspicillata (Boheman,1845) . . X X X X X . . . . . . StenocraninaeWagner,1963 22 Stenocranus fuscovittatus (Stål,1858) . . . . X X X . . . . . . 23 Stenocranus major (Kirschbaum,1868) . X X X X . X . . . . X X 24 Stenocranus minutus (Fabricius,1787) . X X X X X X X X . . X X DelphacinaeWagner,1963 DelphaciniLeach,1815 25 Acanthodelphax denticauda (Boheman,1847) . X X X X X X X . X X X . 26 Acanthodelphax spinosa (Fieber,1866) . . . X X X X X . . . . X 27 Achorotile albosignata (Dahlbom,1850) . X X? X X . . . . X X X . 28 Achorotile longicornis (J.Sahlberg,1871) . . . . X . . . . . X . . 29 Calligypona reyi (Fieber,1866) . . X X X X . . . . . . . 30 Chloriona chinai Ossiannilsson,1946 . . . X X X X . . . X . . 31 Chloriona dorsata Edwards,1898 . . X X . . X X . . . . . 32 Chloriona glaucescens Fieber,1866 . X X X X X X . . . . . . 33 Chloriona smaragdula (Stål,1853) . X X X X X X X . X X . . 34 Chloriona stenoptera (Flor,1861) . . . . . X X . . . . . . 35 Chloriona vasconica Ribaut,1934 . . X X . . X . . . . . . 36 Conomelus anceps (Germar,1821) . X X X X X X X . . . X . 37 Criomorphus albomarginatus Curtis,1833 . X X X X X X X . . X X 38 Criomorphus borealis (J.Sahlberg,1871) . X . X X X X . . X X X . 39 Criomorphus moestus (Boheman,1847) . . . X X X X . . . X X . 40 Delphacinus mesomelas (Boheman,1850) . . X X X X X X . . . . . 41 Delphacodes capnodes (Scott,1870) . . . X X X . . . . . . . 42 Delphacodes venosus (Germar,1830) . X X X X X X . . . X X . 43 Delphax crassicornis (Panzer,1796) . . X X X X X X . . X . X 44 Delphax pulchellus (Curtis,1833) . X X X X X . X . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 39

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 45 Dicranotropis hamata (Boheman,1847) . X X X X X X X . . X X . 46 Ditropis pteridis (Spinola,1839) . X X X . . . . . . . . . 47 Euconomelus lepidus (Boheman,1847) . X X X X X X X . . . . . 48 Euides basilinea (Germar,1821) . X X X X X X . . . X . . 49 Eurybregma nigrolineata Scott,1870 . . . X . . . . . . . . . 50 Eurybregma porcus (Emeljanov,1964) . . . . X . . . . . X . . 51 Eurysa lineata (Perris,1857) . X . X . . . . . . . . . 52 Eurysula lurida (Fieber,1866) . X X X X X X X . . X . . 53 Ditropsis flavipes (Signoret,1865) . . . . X . . . . . . . . 54 Florodelphax leptosoma (Flor,1861) . . X X X X X . . . . . . 55 Florodelphax paryphasma (Flor,1861) . . . X X X X X . . X . . 56 Gravesteiniella boldi (Scott,1870) . . X X X X X X X . . . . 57 Hyledelphax elegantula (Boheman,1847) . X X X X X X X . X X X . 58 Toya propinqua (Fieber,1866) X . . . . . . . . . . . . 59 Javesella alpina (J.Sahlberg,1871) . X . X X . . . . X . . . 60 Javesella bottnica Huldén,1974 . . . X X . . . . . X . . 61 Javesella discolor (Boheman,1847) . X X X X X X X . X X X . 62 Javesella dubia (Kirschbaum,1868) . X X X X X X X . . X X . 63 Javesella forcipata (Boheman,1847) . X X X X X X X . X X . . 64 Javesella obscurella (Boheman,1847) . X X X X X X X . X X X X 65 Javesella pellucida (Fabricius,1794) X X X X X X X X X X X X X 66 Javesella salina (Haupt,1924) . . . X . X . X . . . . . 67 Javesella simillima (Linnavuori,1948) . . . . X X . . . . X . . 68 Javesella stali (Metcalf,1943) . . . X X X X X . X X X . 69 Kosswigianella exigua (Boheman,1847) . X X X X X X X . X X . 70 Laodelphax striatella (Fallén,1826) . X . X X X X X X X X X X 71 Megadelphax haglundi (J.Sahlberg,1871) . . . X . . . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

40 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 72 Megadelphax sordidula (Stål,1853) . . . X X X X . . . X . . 73 Megamelodes quadrimaculatus (Signoret,1865) . . X . . . . . . . . . . 74 Megamelus notula (Germar,1830) . X X X X X X X . X X X . 75 Muellerianella brevipennis (Boheman,1847) . X X X X X X X . . X X . 76 Muellerianella extrusa (Scott,1871) . X X X X . . . . . . . . 77 Muellerianella fairmairei (Perris,1857) . X X X X X X X . . . . . 78 Muirodelphax aubei (Perris,1857) . . X X X X X X . . . . . 79 Nothodelphax albocarinata (Stål,1858) . . . X X X X . . X . . . 80 Nothodelphax distincta (Flor,1861) . X X X X X X X . . . . . 81 Oncodelphax pullula (Boheman,1852) . X X X X X X X . . X X . 82 Paradelphacodes litoralis (Reuter,1880) . . . . X . . . . . . . . 83 Paradelphacodes paludosa (Flor,1861) . . X X X X X X . . X . . 84 Paraliburnia adela (Flor,1861) . . X X X X X X . . . . X 85 Paraliburnia clypealis (J.Sahlberg,1871) . . . X X . . . . . . . . 86 Ribautodelphax albostriata (Fieber,1861) . X . X X X X X . X X X . 87 Ribautodelphax angulosa (Ribaut,1953) . . X X X . X . . . . . . 88 Ribautodelphax collina (Boheman,1847) . X X X X X X X . . X . . 89 Ribautodelphax pallens (Stål,1854) . X . X X X X X X X X . . 90 Ribautodelphax pungens (Ribaut,1953) . . . X . . . . . . . . . 91 Ribautodelphax vinealis Bieman,1987 . X . . X . . . . . . . . 92 Stiroma affinis Fieber,1866 . X X X X X X X . X X . . 93 Stiroma bicarinata (HerrichSchäffer,1835) . X X X X X X X . X X X X 94 Stiromella obliqua (Wagner,1948) . . . . . X X . . . . . . 95 Stiromoides maculiceps (Horváth,1903) . . . . X X . . . . . . . 96 Struebingianella lugubrina (Boheman,1847) . . X X X X X X . . X X . 97 Unkanodes excisa (Melichar,1898) . X X X X X X X X . . X . 98 Xanthodelphax flaveola (Flor,1861) . X X X X X X X . . X . X ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 41

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 99 Xanthodelphax straminea (Stål,1858) . . X X X X X X . . X X . 100 Xanthodelphax xantha Vilbaste,1965 . . . . X . X . . . . . . ACHILIDAEStål,1866 101 Cixidia confinis (Zetterstedt,1838) . X . X X X . X . . . . . 102 Cixidia lapponica (Zetterstedt,1838) . X . X X . . . . . X . . TETTIGOMETRIDAEGermar,1821 103 Tettigometra atra Hagenback,1825 . . . . . X X X . . . . . CALISCELIDAEAmyot&Serville,1834 OmmatidiotiniFieber,1875 104 Ommatidiotus dissimilis (Fallén,1806) . X X X X X X X X . X X . 105 Ommatidiotus inconspicuus Stål,1863 . . . . X . . . . . . . . ISSIDAESpinola,1839 IssiniSpinola,1839 106 Issus muscaeformis (Schrank,1781) . X X X . . . . . . . . . CICADOMORPHAEvans,1946 CICADIDAELatreille,1802 TibicininaeDistant,1905 107 Cicadetta montana (Scopoli,1772) . X . X X . . . . . X X . APHROPHORIDAEAmyot&Serville,1843 108 Aphrophora alni (Fallén,1805) . X X X X X X X X . X X X 109 Aphrophora corticea Germar,1821 . X . X . . . . . . . . . 110 Aphrophora major Uhler,1896 . . . . . . . X . . . X . 111 Aphrophora pectoralis Matsumura,1903 . X . X X X X X X . . X X 112 Aphrophora salicina (Goeze,1778) . X X X X X X X . . . . X 113 Lepyronia coleoptrata (Linnaeus,1758) . X X X X X X X . . X . . 114 Neophilaenus albipennis (Fabricius,1798) . . . . . X . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

42 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 115 Neophilaenus campestris (Fallén,1805) . . X X . X X X . . . X . 116 Neophilaenus exclamationis (Thunberg,1784) . X X X X X X X . X X X . 117 Neophilaenus lineatus (Linnaeus,1758) . X X X X X X X X X X X . 118 Neophilaenus minor (Kirschbaum,1868) . . . . X . X X . . . . . 119 Peuceptyelus coriaceus (Fallén,1826) . . . . X X X X . . X X . 120 Philaenus spumarius (Linnaeus,1758) X X X X X X X X X X X X . MEMBRACIDAERafinesque,1815 121 Centrotus cornutus (Linnaeus,1758) . X X X X X X X . . X X . 122 Gargara genistae (Fabricius,1775) . . X . . . X . . . . . . CICADELLIDAELatreille,1825 AgalliinaeKirkaldy,1901 123 Agallia brachyptera (Boheman,1847) . X X X X X X X . . X X . 124 Agallia consobrina Curtis,1833 . . X X . . . . . . . . . 125 Anaceratagallia estonica Vilbaste,1959 . . . . X X X . . . . . . 126 Anaceratagallia lithuanica Vilbaste,1974 . . . . X . X X . . . . X 127 Anaceratagallia ribauti (Ossiannilsson,1938) . X X X X X X X . . . . . 128 Anaceratagallia venosa (Fourcroy,1785) . X X X X X X X X X X X . AphrodinaeHaupt,1927 129 Anoscopus albifrons (Linnaeus,1758) . X X X X . X . . . X X . 130 Anoscopus albiger (Germar,1821) . . X X . X X . X . . . . 131 Anoscopus flavostriatus (Donovan,1799) . X X X X X X X X . X X . 132 Anoscopus histrionicus (Fabricius,1794) . X X X X . X X . . . X . 133 Anoscopus limicola Edwards,1908 . X . X . . . . . . . . . 134 Anoscopus serratulae (Fabricius,1775) . . X X . X X X . . . . . 135 Aphrodes bicincta (Schrank,1776) . X X X X X X X X . . . . 136 Aphrodes diminuta Ribaut,1952 . X . X X X X? X . . . . . 137 Aphrodes makarovi Zachvatkin,1948 . X X X X X X X X . X X . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 43

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 138 Planaphrodes bifasciata (Linnaeus,1758) . X X X X X X X . X X X . 139 Planaphrodes laeva (Rey,1891)(=trifasciataFour . X X X X X X X . X X . . croy) 140 Planaphrodes nigrita (Kirschbaum,1868) . X . X X . X X . . . . . 141 Stroggylocephalus agrestis (Fallén,1806) . X X X X X X X . . X X X 142 Stroggylocephalus livens (Zetterstedt,1840) . X X X X X X X . . X . . CicadellinaeLatreille,1825 143 Batysmatophorus reuteri J.Sahlberg,1871 . X . X X . . . . X X . . 144 Cicadella lasiocarpae Ossiannilsson,1981 . . X X X . . . . . . . X 145 Cicadella viridis (Linnaeus,1758) . X X X X X X X X . X X X 146 Graphocephala fennahi Young,1977 . . X X . . . . . . . . . 147 Evacanthus acuminatus (Fabricius,1794) . X X X X X X X . . X X . 148 Evacanthus interruptus (Linnaeus,1758) . X X X X X X X . . X X . IassinaeAmyot&Serville,1843 149 Batracomorphus allionii (Turton,1802) . . X . X X . X . . X . X 150 Batracomorphus irroratus Lewis,1834 . . X? . . . X X . . . . . 151 Iassus lanio (Linnaeus,1761) . X X X X X X X X . . . . IdiocerinaeBaker,1915 152 Acericerus heydenii Kirschbaum,1868 . X X X X . . . . . . . . 153 Acericerus ribauti Nickel&Remane,2002 . . . X . . . X . . . . . 154 Balcanicerus larvatus (HerrichSchäffer,1835) . . . X . . . . . . . . . 155 Idiocerus herrichii (Kirschbaum,1868) . X . X X . . . . . X . . 156 Idiocerus lituratus (Fallén,1806) . X X X X X X X . . X X . 157 Idiocerus similis Kirschbaum,1868 . . . X . . . X . . . . . 158 Idiocerus stigmaticalis Lewis,1834 . X X X X X X X X . . . . 159 Metidiocerus crassipes (J.Sahlberg,1871) . . . X X . . . . . X . . 160 Metidiocerus elegans Flor,1861 . X X X X X X X X X X X . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

44 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 161 Populicerus albicans Kirschbaum,1868 . . X X X . X X . . . . . 162 Populicerus confusus Flor,1861 . X X X X X X X . . X X X 163 Populicerus laminatus Flor,1861 . X X X X X X . X . X X X 164 Populicerus nitidissimus (HerrichSchäffer,1835) . X X X X . . X . X . . . 165 Populicerus populi (Linnaeus,1761) . X X X X X X X X X X X X 166 Rhytidodus decimusquartus (Schrank,1776) . X . X . . . X X . . . . 167 Sahlbergotettix salicicola (Flor,1861) . . . . X X . . . . . . . 168 Stenidiocerus poecilus (HerrichSchäffer,1835) . X . X X . X X . . . . . 169 Tremulicerus distinguendus (Kirschbaum,1868) . . X X X . . X X . . . . 170 Tremulicerus fulgidus Fabricius,1775 . X . X . . . X . . . . . 171 Tremulicerus tremulae (Estlund,1796) . X X X X . X X . . X X . 172 Tremulicerus vitreus (Fabricius,1803)sensuRibaut . . X X . . . . . . . . . 173 Viridicerus ustulatus (Mulsant&Rey,1855) . . . X . . . X . . . . . LedrinaeKirschbaum,1868 174 Ledra aurita (Linnaeus,1758) . X X X . . . . . . . . . MacropsinaeEvans,1935 175 Hephathus achilleae Mitjaev,1967 . . . . X X X X . . X X . 176 Macropsidius sahlbergi (Flor,1861) . . . . X X X X . . . . . 177 Macropsis albae Wagner,1950 . . . . X X X X . . . . . 178 Macropsis cerea (Germar,1837) . X X X X X X X . . X X . 179 Macropsis flavida Vilbaste,1980 . . . . X . . . . . . . . 180 Macropsis fuscinervis (Boheman,1845) . X X X X X X X . . X . X 181 Macropsis fuscula (Zetterstedt,1828) . X X X X X X X . . X X . 182 Macropsis glandacea (Fieber,1868) . . . . . . . X . . . . . 183 Macropsis graminea (Fabricius,1798) . X . X . . . . . . . . . 184 Macropsis gravesteini Wagner,1953 . . . . X . . . . . . . . 185 Macropsis impura (Boheman,1847) . X X X X X X X . X . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 45

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 186 Macropsis infuscata (J.Sahlberg,1871) . X X X X X X X . . X X . 187 Macropsis marginata (HerrichSchäffer,1836) . . . . . X X X X . . . . 188 Macropsis megerlei (Fieber,1868) . X . . . . . . . . . . 189 Macropsis prasina (Boheman,1852) . X X X . X X X . . X X X 190 Macropsis scutellata (Boheman,1852) . . X X X . . . . . . . . 191 Macropsis vicina (Horvath,1897) . . . . . . . X . . . . . 192 Macropsis viridinervis Wagner,1950 . . . . . . . X . . . . . 193 Oncopsis alni (Schrank,1801) . X X X X X X X . . . X . 194 Oncopsis appendiculata Wagner,1944 . X . X X . X X X . . . . 195 Oncopsis avellanae Edwards,1920 . . . . X . . . . . . . . 196 Oncopsis carpini (J.Sahlberg,1871) . . X X . . . X . . . . . 197 Oncopsis flavicollis (Linnaeus,1761) . X X X X X X X X X X X . 198 Oncopsis planiscuta (Thomson,1870) . X . X X . . . . X . . . 199 Oncopsis subangulata (J.Sahlberg,1871) . X X X X X X X . . X . . 200 Oncopsis tristis (Zetterstedt,1940) . X X X X X X X X X X X . 201 Pediopsis tiliae (Germar,1831) . X X X X . X . . . . . . MegophthalminaeKirkaldy,1906 202 Megophthalmus scanicus (Fallén,1806) . X X X X X X X X . X . X TyphlocybinaeKirschbaum,1868 AlebriniMcAtee,1926 203 Alebra albostriella (Fallén,1826) . X X X X X X X X . X . . 204 Alebra neglecta Wagner,1940 . . . X X . . X . . . X . 205 Alebra wahlbergi (Boheman,1845) . X X X X X X X . . X . X DikraneuriniMcAtee,1926 206 Dikraneura aridella (J.Sahlberg,1871) . X . X X . . . . X X . . 207 Dikraneura variata Hardy,1950 . . X X X X X X X . . . . 208 Liguropia juniperi (Lethierry,1876) . . . X . . . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

46 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 209 Emelyanoviana mollicula (Boheman,1845) . X X X X X X X . . . . . 210 Erythria aureola (Fallén,1806) . X X X X X X X . X X . . 211 Forcipata citrinella (Zetterstedt,1828) . X X X X X X X X X X . X 212 Forcipata forcipata (Flor,1861) . X X X X X X X . X . . . 213 Forcipata palustris Holgersen,1992 . X . . . . . . . . . . . 214 Igutettix oculatus (Lindberg,1929) . . . . X X . . . . . X . 215 Micantulina micantula (Zetterstedt,1840) . X . X X . . . . . . . . 216 Micantulina pseudomicantula (Knight,1966) . . . . X . . X . . . . . 217 Notus flavipennis (Zetterstedt,1828) . X X X X X X X X X X X . 218 Wagneriala incisa (Then,1897) . . . X . . . . . . . . . 219 Wagneriala minima (J.Sahlberg,1871) . X X X X X X X . . . . . EmpoasciniDistant,1908 220 Austroasca vittata (Lethierry,1884) . . . . . . X? X? . . . . . 221 Chlorita dumosa Ribaut,1933 . . . X X X X X . . . . . 222 Chlorita paolii Ossiannilsson,1939 . . . . X X X X X . X X X 223 Chlorita viridula (Fallén,1806) . X X X X X X . X . X X . 224 Empoasca affinis Nast,1937 . . . . . . . X . . . . . 225 Empoasca apicalis (Flor,1861) . . . . X X X . . X . . . 226 Empoasca decipiens Paoli,1930 . X X X . . X X . . . . . 227 Empoasca kontkaneni Ossiannilsson,1949 . X . X X X X . X . . X X 228 Empoasca ossiannilssoni Nuorteva,1948 . X . X X . X . . . . X . 229 Empoasca pteridis (Dahlbom,1850) . X X X X X X X . . . X X 230 Empoasca vitis (Göthe,1875) . X X X X X X X X . X X X 231 Kyboasca bipunctata (Oshanin,1871) . . . X X . . . . . . . . 232 Kybos abstrusus (Linnavuori,1949) . X . X X . X X . . . . . 233 Kybos butleri (Edwards,1908) . X X X X X X X . X X X X 234 Kybos limpidus (Wagner,1955) . . . X . . . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 47

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 235 Kybos lindbergi (Linnavuori,1951) . X . X X X X . . X X X X 236 Kybos mucronatus (Ribaut,1933) . . . . . . X . X . . . . 237 Kybos populi (Edwards,1908) . X X X X X X X X . . X X 238 Kybos rufescens Melichar,1896 . . X X X . . . . . . . X 239 Kybos smaragdula (Fallén,1806) . X X X X X X X X X X X X 240 Kybos sordidulus (Ossiannilsson,1941) . X . X X . . . . X X . X 241 Kybos strigilifer (Ossiannilsson,1941) . X X X X X X . . . X . . 242 Kybos virgator (Ribaut,1933) . X X X X . X X . X . . . 243 Kybos volgensis Vilbaste,1961 . . . X X . . . . . . . . ErythroneuriniYoung,1952 244 Alnetoidia alneti (Dahlbom,1850) . X X X X X X X X . X X X 245 Arboridia erecta (Ribaut,1931) . . . . . . X . . . . . . 246 Arboridia parvula (Boheman,1845) . X X X X . . . . . X X X 247 Arboridia ribauti (Ossiannilsson,1937) . . . . . . X X . . . . . 248 Arboridia velata (Ribaut,1952) . . . . . . . X . . . . . 249 Hauptidia distinguenda (Kirschbaum,1868) . . . X . . . . . . . . . 250 Zygina angusta Lethierry,1874 . X . X X . . X X . . . . 251 Zygina flammigera (Geoffroy,1785) . X X X X X X X X . X X X 252 Zygina hyperici (HerrichSchäffer,1836) . X X X X X X X . . X . X 253 Zygina nigritarsis Remane,1994 . . . . X . . . . . . X X 254 Zygina ordinaria (Ribaut,1936) . . X X X X X X . . . X X 255 Zygina rosea (Flor,1861) . X . X X X X . . . X X . 256 Zygina rosincola (Cerutti,1939) . X X X X . . . . . . . . 257 Zygina rubrovittata (Lethierry,1869) . X X X X X X X X . . . . 258 Zygina salicina Mitjaev,1975 . . X . . . . . . . . . . 259 Zygina schneideri (Günthart,1974) . X . X X . . . . . . . . 260 Zygina suavis Rey,1891sensuOssiannilsson . . . X X X X X . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

48 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 261 Zygina tiliae (Fallén,1806) . X X X X X X X . . X X X 262 Zyginidia mocsaryi (Horváth,1910) . . . X . X X . . . . . . 263 Zyginidia pullula (Boheman,1845) . . X X X . . . . . . . . 264 Zyginidia viaduensis (Wagner,1941) . . . . . X X X . . . . . Kirschbaum,1868 265 Aguriahana pictilis (Stål,1853) . X X X X X X X . . X . . 266 Aguriahana stellulata (Burmeister,1841) . X X X X . X X X . X . . 267 Edwardsiana alnicola (Edwards,1924) . X X X X X X . . . . . X 268 Edwardsiana ampliata (Wagner,1947) . . . . X . X X . . . . . 269 Edwardsiana avellanae (Edwards,1888) . . X X X X X X . . . . . 270 Edwardsiana bergmani (Tullgren,1916) . X X X X X X X . X X X X 271 Edwardsiana candidula (Kirschbaum,1868) . . . X X . X X . . . . . 272 Edwardsiana crataegi (Douglas,1876) . X X X X . . . . . . X X 273 Edwardsiana diversa (Edwards,1914) . . . X . . . . . . . . . 274 Edwardsiana flavescens (Fabricius,1794) . . X X . . . X . . . . . 275 Edwardsiana frustator (Edwards,1908) . X X X X . . X . . . . . 276 Edwardsiana geometrica (Schrank,1801) . X X X X X X X X . X X X 277 Edwardsiana gratiosa (Boheman,1852) . . X X X X X X . . . . X 278 Edwardsiana ishidai (Matsumura,1932) . X X X X . . . . . . . . 279 Edwardsiana kemneri (Ossiannilsson,1942) . . . X . . . . . . . . . 280 Edwardsiana lanternae (Wagner,1937) . . . X X . . . . . . . X 281 Edwardsiana lethierryi (Edwards,1881) . X . X X . . X . . . . . 282 Edwardsiana menzbieri Zachvatkin,1948 . X . X X . X X . X X . X 283 Edwardsiana nigriloba (Edwards,1924) . . . X . . . . . . . . . 284 Edwardsiana plebeja (Edwards,1914) . X X X X . X X . . . . . 285 Edwardsiana plurispinosa (Wagner,1935) . X . X X . . X . . . . . 286 Edwardsiana prunicola (Edwards,1914) . X X X X X X X . . . . X ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 49

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 287 Edwardsiana rosae (Linnaeus,1958) . X X X X X X X X . . X X 288 Edwardsiana salicicola (Edwards,1885) . X X X X X X X . . . . . 289 Edwardsiana sociabilis (Ossiannilsson,1936) . X X X X . . . . . . X X 290 Edwardsiana soror (Linnavuori,1950) . X . X X X X X X . . . X 291 Edwardsiana spinigera (Edwards,1924) . . . . . X . . . . . . . 292 Edwardsiana staminata (Ribaut,1931) . X . X . . X X . . . X . 293 Edwardsiana stehliki Lauterer,1958 . . . X X . . . . . . . . 294 Edwardsiana tersa (Edwards,1914) . X X X X . . X . . . . . 295 Edwardsiana ulmiphagus Wilson&Claridge,1999 . . X X X . . X . . . . . 296 Eupterycyba jucunda (HerrichSchäffer,1837) . X X X X X X X . . . . . 297 Eupteryx adspersa (HerrichSchäffer,1838) . . X? . . . . X . . . . . 298 Eupteryx artemisiae (Kirschbaum,1868) . . X X . . . . . . . . . 299 Eupteryx atropunctata (Goeze,1778) . X X X X X X X X . X X X 300 Eupteryx aurata (Linnaeus,1758) . X X X X X X X . . X . X 301 Eupteryx calcarata Ossiannilsson,1936 . X X X X X X X X . . X X 302 Eupteryx collina (Flor,1861) . . . . X . X X . . . . . 303 Eupteryx cyclops Matsumura,1906 . X X X X . X X X X X X X 304 Eupteryx decemnotata Rey,1891 . . X X X . . . . . . . . 305 Eupteryx florida Ribaut,1936 . . X X . . . . X . . . . 306 Eupteryx heydenii (Kirschbaum,1868) . . X . . . X . . . . . . 307 Eupteryx notata Curtis,1837 . X X X X X X X X X X X X 308 Eupteryx origani Zachvatkin,1948 . . . X X X X X . . . . X 309 Eupteryx signatipennis (Boheman,1847) . X X X X X X X . . X . X 310 Eupteryx stachydearum (Hardy,1850) . X X X X X X X . . . . . 311 Eupteryx tenella (Fallén,1806) . X . X X X X X X . X X X 312 Eupteryx thoulessi Edwards,1926 . . X X . . . . . . . . . 313 Eupteryx urticae (Fabricius,1803) . X X X X X X X . . . . X ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

50 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 314 Eupteryx vittata (Linnaeus,1758) . X X X X X X X X X X . X 315 Eurhadina concinna (Germar,1831) . X X X X X X X . . . . . 316 Eurhadina kirschbaumi Wagner,1837 . X . X X . X X . . . . . 317 Eurhadina loewii (Then,1886) . . X X . X . . . . . . . 318 Eurhadina pulchella (Fallén,1806) . X X X X X X X X . X X X 319 Eurhadina ribauti Wagner,1935 . X X X X . . . . . . . . 320 Eurhadina saageri Wagner,1937 . . . . . . X X . . . . . 321 Fagocyba carri (Edwards,1914) . X X X X . X X . . . . . 322 Fagocyba cruenta (HerrichSchäffer,1838) . X X X X X X X X . . . X 323 Linnavuoriana decempunctata (Fallén,1806) . X X X X X X X . X . X X 324 Linnavuoriana intercedens (Linnavuori,1949) . X . X X . . . . . X X X 325 Linnavuoriana sexmaculata (Hardy,1850) . X X X X X X X . X X X X 326 Ossianilssonola callosa (Then,1886) . . X X . . . . . . . . . 327 Ribautiana scalaris (Ribaut,1931) . X X X . . . . . . . . . 328 Ribautiana tenerrima (HerrichSchäffer,1834) . X X X . . . X X . . . . 329 Ribautiana ulmi (Linnaeus,1758) X X X X X X X X X . . X . 330 Typhlocyba quercus (Fabricius,1777) . X X X X X X X . . . X X 331 Wagneripteryx germari (Zetterstedt,1840) . X X X X X X X X X X X . 332 Zonocyba bifasciata (Boheman,1851) . . X X X X X X . . . . . 333 Zyginella pulchra P.Löw,1885 . . X X . . . . . . . . . UlopinaeLePeletier&Serville,1825 334 Ulopa reticulata (Fabricius,1784) . X X X X X X X X X X . . 335 Utecha trivia (Germar,1821) . . . . . . . X . . . . . Fieber,1869 VanDuzee,1892 336 Allygidius commutatus (Fieber,1872) . X X X X X X X . . X X . 337 Allygus communis (Ferrari,1882) . X X X . . . X . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 51

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 338 Allygus maculatus Ribaut,1948 . . X X . . . . . . . . . 339 Allygus mixtus (Fabricius,1794) . X X X X X X X X . X X X 340 Allygus modestus Scott,1876 . . X X . . . X . . . . . 341 Athysanus argentarius Metcalf,1955 . X X X X X X X X . . . . 342 Athysanus quadrum Boheman,1845 . X X X X X X X . . . X . 343 Cicadula albingensis Wagner,1940 . X . . X X . X . . . . . 344 Cicadula ciliata (Osborne,1898) . . . . X . . . . . . . . 345 Cicadula flori (J.Sahlberg,1871) . . X X X X X X . . X X X 346 Cicadula frontalis (HerrichSchäffer,1835) . . X X X X X X X . . . X 347 Cicadula intermedia (Boheman,1845) . X . X X X . . . X X . . 348 Cicadula rubroflava Linnavuori,1952 . . . . X X X X . . . . X 349 Cicadula nigricornis (J.Sahlberg,1871) . . . X X X X X . . . X . 350 Cicadula ornata (Melichar,1900) . X . X X X X . . . . . X 351 Cicadula persimilis (Edwards,1920) . X X X X X X X . . X X X 352 Cicadula quadrinotata (Fabricius,1794) X X X X X X X X X X X X X 353 Cicadula quinquenotata (Boheman,1845) . X X X X X X . . . X X . 354 Cicadula saturata (Edwards,1915) . X X X X X X X . . X . . 355 Colladonus torneellus (Zetterstedt,1828) . X X X X X X X . X X . . 356 Colobotettix morbillosus (Melichar,1896) . . . . X . . . . . . . . 357 Conosanus obsoletus (Kirschbaum,1858) . X X X . X X X . . . . . 358 Doliotettix lunulatus (Zetterstedt,1840) . X . X X X X X . X X . X 359 Ederranus discolor (J.Sahlberg,1871) . . . . X . . . . . X X X 360 Ederranus sachalinensis (Matsumura,1911) . . . . X . . . . . . . . 361 Elymana kozhevnikovi (Zachvatkin,1938) . . . X X X X X . . . X X 362 Elymana sulphurella (Zetterstedt,1828) . X X X X X X X X X X X X 363 Euscelidius schenckii (Kirschbaum,1868) . X X X X X X X . . X X . 364 Euscelis distinguendus (Kirschbaum,1858) . X . X X X X X . . X . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

52 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 365 Euscelis incisus (Kirschbaum,1858) . . X X . . . X X . . . . 366 Euscelis ohausi Wagner,1939 . . X . . . . . . . . . . 367 Euscelis venosus (Kirschbaum,1868) . . . . . . X . . . . . . 368 Graphocraerus ventralis (Fallén,1806) . X X X X X X X X . X X . 369 Handianus flavovarius (HerrichSchäffer,1835) . . . . . X X X . . . . . 370 Hardya signifier (Then,1897) . . . . . . . X . . . . . 371 Hardya tenuis (Germar,1821) . . . X X . X? X . . X X . 372 Hesium domino (Reuter,1880) . X . X X X X X X . X X X 373 Idiodonus cruentatus (Panzer,1799) . X X X X X X X . X X X . 374 Laburrus impictifrons (Boheman,1852) . . X X X X X X X . . X . 375 Lamprotettix nitidulus (Fabricius,1787) . . X X . X X X X . . . . 376 Limotettix aviger Emeljanov,1966 . . . . . . . . . . . . X 377 Limotettix atricapillus (Boheman,1845) . X . X X X . X . . X . . 378 Limotettix ochrifrons Vilbaste,1973 . . . . X X X X . . . . . 379 Limotettix sphagneticus Emeljanov,1964 . . . . X X X X . . X . . 380 Limotettix striola (Fallén,1806) . X X X X X X X X X X X X 381 Macustus grisescens (Zetterstedt,1828) . X X X X X X X . X X X X 382 Mocydiopsis attenuata (Germar,1821) . . X X . . . . . . . . . 383 Mocydiopsis parvicauda Ribaut,1939 . . . X . . . . . . . . . 384 Ophiola cornicula Marshall,1866 . . . X X X X X . X X X . 385 Ophiola decumana Kontkanen,1949 . X X X X X X X . . X X . 386 Ophiola identica (Tischechkin,2003) . . . . X . X? X? . . . . . 387 Ophiola orichalcea (Thomson,1869) . . . X X . . . . . . . . 388 Ophiola russeola (Fallén,1806) . X X X X X X X . X . . . 389 Ophiola transversa (Fallén,1826) . X X X X X X X . . X X . 390 Ophiolix paludosa (Boheman,1845) . X . X X X X . . . X X . 391 Paluda flaveola (Boheman,1845) . X X X X X X X . X X X . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

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No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 392 Perotettix pictus (Lethierry,1880) . . . . X . . . . . . . . 393 Perotettix orientalis (Anufriev,1971) . X . X X . . . . . . . . 394 Pithyotettix abietinus (Fallén,1806) . X X X X X X X . X X . . 395 Platymetopius guttatus Fieber,1869 . X . X . . . X . . . . . 396 Platymetopius major (Kirschbaum,1868) . . X X . . . . . . . . . 397 Platymetopius undatus (DeGeer,1773) . X X X X X X X . . X X . 398 Rhopalopyx adumbrata (C.Sahlberg,1842) . X X X X X X X . . . . . 399 Rhopalopyx preyssleri (HerrichSchäffer,1838) . X X X X X X X . . X X . 400 Rhopalopyx vitripennis (Flor,1861) . X? X X X X X X . . X . . 401 Rhytistylus proceps (Kirschbaum,1868) . . X X . . . . . . . . . 402 Speudotettix subfusculus (Fallén,1806) . X X X X X X X X X X X X 403 Stictocoris picturatus (C.Sahlberg,1842) . . . X X X X X . . X X . 404 Streptanus aemulans (Kirschbaum,1868) . X X X X X X X . . . X . 405 Streptanus confinis (Reuter,1880) . X X X X X X X . . . . . 406 Streptanus marginatus (Kirschbaum,1858) . X X X X X X X . X X X . 407 Streptanus okaensis Zachvatkin,1948 . . X X X . . . . X . . . 408 Streptanus sordidus (Zetterstedt,1828) . X X X X X X X . X X X X 409 Thamnotettix confinis Zetterstedt,1840 . X X X X X X X . X X X . 410 Thamnotettix dilutior (Kirschbaum,1868) . . X X . . . . . . . . . CoryphaeliniEmeljanov,1962 411 Coryphaelus gyllenhalii (Fallén,1826) . . . X X X X X . . X . X DeltocephaliniFieber,1869 412 Deltocephalus maculiceps Boheman,1847 . . X X . . . . . . . . . 413 Deltocephalus pulicaris (Fallén,1806) . X X X X X X X . X X X . 414 Endria nebulosa (Ball,1900) . . X X X X . . . . . . . DoraturiniRibaut,1952 415 Doratura exilis Horváth,1903 . . . X X X X X X . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

54 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 416 Doratura homophyla (Flor,1861) . . X X X X X X . . X X . 417 Doratura impudica Horváth,1907 . . X X . X X X . . . . . 418 Doratura littoralis Kuntze,1937 . . . . . . X . . . . . . 419 Doratura stylata (Boheman,1847) . X X X X X X X X X X X . DorycephaliniOman,1943 420 Eupelix cuspidata (Fabricius,1775) . X X X X X X X . . X X . FieberielliniWagner,1951 421 Fieberiella septentrionalis Wagner,1963 . . . X . . X X . . . . . GrypotiniEvans,1947 422 Grypotes puncticollis (HerrichSchäffer,1834) . X X X X X X X X . X X . MacrosteliniKirkaldy,1906 423 Balclutha arhenana Dlabola,1967 . . . . X . . X . . . X X 424 Balclutha boica Wagner,1950 . . . X X X X X . . X . . 425 Balclutha calamagrostis Ossiannilsson,1961 . . . X X X X X X . . . X 426 Balclutha punctata (Fabricius,1775)sensuWagner . X X X X X X X X X X X X 427 Balclutha rhenana Wagner,1939 . . X X X . . . X . . . X 428 Erotettix cyane (Boheman,1845) . X X X X X X . . . . . . 429 Macrosteles alpinus (Zetterstedt,1828) . X . X X . X . . X . X . 430 Macrosteles cristatus (Ribaut,1927) . X X X X X X X . . . X X 431 Macrosteles empetri (Ossiannilsson,1935) . X . X X . . . . X . . . 432 Macrosteles fieberi (Edwards,1889) . X . X X X . . . X . . . 433 Macrosteles frontalis (Scott,1875) . X . X X X X X . X X X . 434 Macrosteles horvathi (Wagner,1935) . X X X X X X X . . X X . 435 Macrosteles laevis (Ribaut,1927) X X X X X X X X X X X . X 436 Macrosteles lividus (Edwards,1894) . X X X X X X X . X . . . 437 Macrosteles maculosus (Then,1897) . . . X . . . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

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No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 438 Macrosteles nubilus (Ossiannilsson,1936) . . . X X . . X . X X . . 439 Macrosteles oshanini Razvyazkina,1957 . . . X X . . X . . . . . 440 Macrosteles ossiannilssoni Lindberg,1954 . X X X X X X X . . . . . 441 Macrosteles quadripunctulatus (Kirschbaum,1868) . . X X X X X X . . . . . 442 Macrosteles septemnotatus (Fallén,1806) . X X X X X X X . . X X X 443 Macrosteles sexnotatus (Fallén,1806) X X X X X X X X X X X X X 444 Macrosteles sordidipennis (Stål,1858) . X X X X . . . . . . . . 445 Macrosteles variatus (Fallén,1806) . X X X X X X X . X X . X 446 Macrosteles viridigriseus (Edwards,1922) . X X X X X X X . X . X . 447 Sagatus punctifrons (Fallén,1826) . X X X X X X X X . X X X 448 Sonronius anderi (Ossiannilsson,1948) . . . X X . . . . X? . . . 449 Sonronius binotatus (J.Sahlberg,1871) . X . X X . X X . X X X . 450 Sonronius dahlbomi (Zetterstedt,1840) . X . X X . X X . X X X . OpsiiniEmeljanov,1962 451 Circulifer haematoceps Mulsant&Rey,1855 . . . . X . . . . . . . . 452 Neoaliturus fenestratus (HerrichSchäffer,1834) . . . . . . X X . . X . . 453 Opsius stactogalus Fieber,1866 . X . X . . . . . . . . . Distant,1908 454 Adarrus bellevoyei (Puton,1877) . . . . . X . . . . . . . 455 Adarrus multinotatus (Boheman,1847) . . . X X X X . . . . . . 456 Arocephalus languidus (Flor,1861) . . . X X X X X X X? X . . 457 Arocephalus longiceps (Kirschbaum,1868) . . X X . . . . . . . . . 458 Arocephalus punctum (Flor,1861) . X X X X X X X . . X . . 459 Arthaldeus pascuellus (Fallén,1826) . X X X X X X X . X X X . 460 Arthaldeus striifrons (Kirschbaum,1868) . . X X X X X X X . . . . 461 Boreotettix bidentatus (DeLong&Davidson, . X . X X . . . . . . . . 1935) ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

56 G.Söderman,G.Gillerfors,A.Enderstöl

No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 462 Calamotettix taeniatus (Horváth,1911) . . . . X . . . . . . . X 463 Cosmotettix aurantiacus (Forel,1859) . . . . . . . X . . . . . 464 Cosmotettix caudatus (Flor,1861) . X X X X X X . . . X X X 465 Cosmotettix costalis (Fallén,1826) . X X X X X X X . X . X X 466 Cosmotettix edwardsi (Lindberg,1924) . . X X X X . . . . . . . 467 Cosmotettix evanescens Ossiannilsson,1976 . . . X X . . . . . . . . 468 Cosmotettix panzeri (Flor,1861) . X X X X X X X . X X X . 469 Diplocolenus bohemani (Zetterstedt,1840) . X X X X X X X . . X X . 470 Ebarrius cognatus (Fieber,1869) . X . X X . . . . X . . . 471 Errastunus ocellaris (Fallén,1806) . X X X X X X X X X X X X 472 Erzaleus metrius (Flor,1861) . X X X X X X X X X . X X 473 Henschia collina (Boheman,1850) . . X X X X X X . . X . . 474 Jassargus allobrogicus (Ribaut,1936) . . X X X . X . . . . . X 475 Jassargus alpinus (Then,1896) . X . X X X X X . X . . . 476 Jassargus flori (Fieber,1869) . X X X X X X X X X X . X 477 Jassargus pseudocellaris (Flor,1861) X X X X . X X X . . . . . 478 Jassargus sursumflexus (Then,1902) . X X X X X X . . X . . . 479 Lebradea flavovirens (Gillette&Baker,1895) . . . X X . . . . X . X . 480 Mendrausus pauxillus (Fieber,1869) . . . X . . . . . . . . . 481 Metalimnus formosus (Boheman,1845) . . . X X X X X . . X X . 482 Metalimnus marmoratus (Flor,1861) . . . . X X X X . . X X . 483 Metalimnus obtusus Emeljanov,1966 . . . X X . . X . . . . . 484 Metalimnus steini (Fieber,1869)sensuAnufriev& . . . . X . . X . . . . X Emeljanov,1988 485 Mongolojassus bicuspidatus (J.Sahlberg,1871) . . . . . X . . . . X X . 486 Mongolojassus sibiricus (Horvath,1901) . . . . X X . . . . . . . 487 Paralimnus phragmitis (Boheman,1847) . X X X X X X . . . X . . 488 Paralimnus rotundiceps (Lethierry,1885) . X . X X . . . . . . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

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No. Taxon IC NO DK SE FI EE LV LT KG MM KR LG PV 489 Paralimnus zachvatkini Emeljanov,1964 . . . . X . . X . . . . . 490 Paramesus obtusifrons (Stål,1853) . X X X X X X . . . . . . 491 Parapotes reticulatus (Horváth,1897) . . . X X . . . . . . . . 492 Pinumius areatus (Stål,1858) . . . . X X X X . . . X . 493 Psammotettix albomarginatus Wagner,1941 . X? X X X . . . . . . . . 494 Psammotettix alienus (Dahlbom,1850) . X X X X X X X X . X . . 495 Psammotettix cephalotes (HerrichSchäffer,1834) . X X X X X X X . . X . . 496 Psammotettix confinis (Dahlbom,1850) . X X X X X X X X X . X X 497 Psammotettix dubius Ossiannilsson,1974 . X . X X X . X . . . . . 498 Psammotettix excisus Matsumura,1906 . . X X X X . X? . . . . . 499 Psammotettix frigidus (Boheman,1845) . X . X X . . . . X . . . 500 Psammotettix koeleriae Zachvatkin,1948 . . . . . . X X . X . . . 501 Psammotettix lapponicus (Ossiannilsson,1938) . X . X X . . . . . . . . 502 Psammotettix nodosus (Ribaut,1925) . X X X X . X X . X . . . 503 Psammotettix pallidinervis (Dahlbom,1851) . X . X X X X X X X X . . 504 Psammotettix poecilus (Flor,1861) . . . X X X X X . . X X . 505 Psammotettix putoni (Then,1898) . X X X . . . . . . . . . 506 Psammotettix sabulicola (Curtis,1837) . X X X X . X X X X . X . 507 Psammotettix slovacus Dlabola,1948 . . . . X X X . . . . . . 508 Rosenus laciniatus (Then,1896) . X . X X . . . . . . . . 509 Sorhoanus assimilis (Fallén,1806) . . X X X X X X . . . X . 510 Sorhoanus xanthoneurus (Fieber,1869) . X X X X X X X X X X . . 511 Turrutus socialis (Flor,1861) . . X X X X X X . . . . . 512 Verdanus abdominalis (Fabricius,1803) . X X X X X X X X X X X . 513 Verdanus limbatellus (Zetterstedt,1828) . X . X X . . . . X . . . ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

58 G.Söderman,G.Gillerfors,A.Enderstöl

4.2Shortcommentsonsomespecies 6. Cixius stigmaticusisreportedfromLatviabySpungis&Kalnins(2002).TheLatvian recordisbyVilbaste(1974)believedtobeamisidentificationofGimmerthal(1846). However,thisspeciesisnotuncommoneastofLatviainwesternRussia(Söderman, inprep.)andwethereforeregarditaspossiblethatGimmerthal’srecordisaccept able. 9. Asiraca clavicornisisreportedfromLatvia.Nast(1987)considers therecordunreli able,becauseVilbaste(1974)regardsitasbeingamisidentificationmadebyGim merthal(1846).Asthishardlycanbemisinterpretedforanyotherspe cies,weconsideritpossiblethatGimmerthalsidentificationwascorrect.Thespe ciesisquitewidespreadinareasclosetoLatvia,i.a.Byelorussia(seeBorodin2004) andtheBelgorodoblast(Söderman,inprep.). 15. Vilbaste (1974) described Kelisia nervosa from Lithuania. This taxon has later (Holzingeret al.2003)beenregardedasajuniorsynonymofKelisia confusa.Thespe cies has recently been found in Finland (Söderman 2007) and Sweden (Gillerfors 2008)andmaybeexpandingitsrange. 23. Stenocranus major is recorded new for Norway by Endrestöl in 2007 (unpublished data). 24. Stenocranus minutusisrecordednewtoLithuaniabySöderman&Rintalain2008(un publisheddata). 26. Acanthodelphax spinosaisrecordednewtoEstoniabyKoponenin1990(unpublished data). 27.Achorotile albosignata is reported from Denmark by Nast (1987), but is not acknowl edgedinOssiannilsson(1978). 33. Chloriona smaragdulaisrecordednewforNorwaybyÖdegaardin2005(unpublished data). 51. Eurysa lineataisrecordednewtoNorwaybyÖdegaard(unpublisheddata). 53. Ditropsis flavipesisrecordednewtoFinlandbySödermanin2007(unpublisheddata). 54. Florodelphax leptosomaisrecordednewtoFinlandbyAlbrechtin2007(unpublished data). 58. Toya propinquaisawidespreadspeciesintheEthiopianandOrientalregionsandthe southern parts of the Holarctic. The specimen from Iceland (Remane 2005) is probablyduetolongrangemigrationwithjetstreams. 91. Ribautodelphax vinealishasrecentlybeendescribedandfoundtobequitecommonin Finland(Söderman2007).ItwasrecentlyalsorecordedfromNorwaybyEndrestöl (unpublisheddata). 97. Unkanodes excisa is recorded new to Norway by Ödegaard in 2007 (unpublished data). 101. Cixidia confinisisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 103. Vilbaste (1974) and Nast (1987) report the taxon Tettigometra atrata Fieber, 1872 fromallBalticcountries.Thetaxonappearstobeunclearandisregardedasapos siblevarietyofeitherT. atraorT. impressifronsbyHolzingeret al.(2003).AsVilbaste alsoreportsT. atrafromEstonia,itislikelythatthespecimensidentifiedasatrata belongtothisspecies. ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 59

105. ThespecimensofOmmatidiotus inconspicuusfromsouthernFinlandareaccordingto Söderman(2007)regardedaslongrangemigrants. 107. Old European records of Cicadetta montana may according to recent investigations belongtothreedifferentspecies.OnthebasisofsonghearingstheNordicpopula tionsshouldbelongtoC. montanas.str.(seeSöderman2007). 110. Aphrophora majorisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 112. Aphrophora salicina is recorded new to Finland by Albrecht in 2006 (unpublished data). 125. Anaceratagallia estonicaisrecordednewtoLatviabySöderman&Spungis(2008). 126. Anaceratagallia lithuanicaisrecordednewtoLatviabySöderman&Spungis(2008). 133. Anoscopus limicola is recorded new to Norway by Endrestöl in 2008 (unpublished data). 136. ThedistinctionbetweenAphrodes diminutaandA. bicinctahasnotbeenmadeinall oldcollections.TherecordforLatviaisthereforeuncertain. 140. Planaphrodes nigrita is recorded new to Norway by Endrestöl in 2008 (unpublished data). 146. Graphocephala fennahiisrecordednewtoSwedenbyGillerfors(2008). 149. Batracomorphus allionii is recorded new to Estonia by Tarlap in 2007 (unpublished data). 150. Batracomorphus irroratus has been reported from Denmark by Nast (1972), but itis notincludedforthiscountrybyOssiannilsson(1981)orNast(1987)intheirlater works.RecordednewtoLatviain2006bySöderman&Spungis(2008). 152. Acericerus heydenii has been reported from Denmark by Nast (1972), but later he states the record to be dubious. It is not included by Ossiannilsson (1981) in his books on the Auchenorrhyncha of Denmark and Fennoscandia. Gillerfors (2002) reportsA. vittifrons(Kirschbaum,1868)fromSweden,butthisrecordhasprovedto beamisidentificationofA. heydenii(seeGillerfors2008).RecordednewtoNorway by Endrestöl in 2007 (unpublished data) and found to be common in Denmark (U.H.Nielsen,pers.comm.) 153. Acericerus ribautiisrecordednewtoLithuaniabySöderman&Rintalain2008(un publisheddata). 154. Balcanocerus larvatus (HerrichSchäffer, 1835) has been reported as Idiocerus notatus (Fabricius,1803)fromNorwaybyStrand(1905).Thisisanobviousmisidentifica tionasalreadystatedbyHolgersen(1944b).ThespeciesisstilllistedfromNorway byNast(1972),butitisnotincludedbyOssiannilsson(1981)inhisbooksonthe Auchenorrhyncha of Denmark and Fennoscandia. Later, Nast (1987) reported it fromDenmark,but thishasnotbeenconfirmed.However,Gillerforsrecordedit newtoSwedenin2008fromaMalaisetrap(unpublisheddata). 157. Idiocerus similisisrecordednewtoSwedenbyGillerfors(2008). 159. Metidiocerus crassipesisrecordednewtoFinlandbySödermanin2006(unpublished data). 161. Populicerus albicansisrecordednewtoLithuaniabySöderman&Rintalain2008(un publisheddata). 163. Populicerus laminatusisrecordednewtoEstoniabyTarlapin2007(unpublisheddata). 164. Populicerus nitidissimusisrecordednewtoLithuaniabySöderman&Rintalain2008 (unpublisheddata). ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

60 G.Söderman,G.Gillerfors,A.Enderstöl

166. Rhytidodus decimusquartusisrecordednewtoKaliningradbySödermanin1998(un publisheddata)andNorwayin2007(Endrestöl2008). 168. Stenidiocerus poecilus is recorded new to Lithuania by Söderman & Dapkus in 2007 (unpublisheddata). 169. Tremulicerus distinguendusisrecordednewtoLithuaniabySöderman&Rintalain2008 (unpublisheddata). 170. Nast(1987)considerstherecordofTremulicerus fulgidusfromLatviaasdubious,but isstilloccursinSpungis&Kalninslist(2002)basedonDanka&Velce(1970).Vil baste (1974) believes it to be a misidentificationof Gimmerthal (1846). Recorded newtoNorwaybyEndrestöl(2008). 172. Tremulicerus vitreus(Fabricius,1803)sensuRibaut1952hasbeenreportedfromLatvia byNast(1972),butitisnotincludedbyVilbaste(1974)inhiscatalogueofLatvian Auchenorrhyncha.VilbasteregardsittobeamisidentificationmadebyGimmerthal (1846).RecordednewtoSwedenbyGillerfors(2008). 173. Viridicerus ustulatus is recorded new to Sweden by Gillerfors (2008) and new to LithuaniabySöderman&Rintalain2008(unpublisheddata). 175. Hephathus nanus is in all older works (Linnavuori 1969, Nast 1972, 1987, Vilbaste 1974,Ossiannilsson1981)reportedfromNorthernEurope.Söderman(2007)states thatatleasttheFinnishspecimensbelongtoH. achilleae,whichisaneasternspecies andnottoH. nanusoccurringinCentralEurope. 177. Macropsis albaeisreportedbyNast(1987)underthenameM. ocellata.Recordednew toFinlandbyAlbrechtin2007(unpublisheddata). 179. Macropsis flavidaisrecordednewtoFinlandandEuropebySödermanin2007(un publisheddata). 182. Macropsis glandaceaisreportedfromLithuaniabyVilbaste(1974)underthenameM. mendax, which is regarded as a junior synonym of M. glandacea. The record from Norway(Nast1987)appearstobeverydubious(Schulz1976). 184. Macropsis gravesteiniisrecordednewtoFinlandbySödermanin2008(unpublished data). 186. Macropsis infuscata is recorded new to Norway by Ödegaard in 2003 (unpublished data). 191. Macropsis vicinaisrecordednewtoLithuaniabySöderman&Rintalain2008(unpub lisheddata). 194. Oncopsis appendiculataisrecordednewtoNorwaybyEndrestölin2007(unpublished data). 195. Oncopsis avellanaemighthavealargerdistribution,sinceithasonlyrecentlybeenac knowledgedtooccurinNorthernEurope(Söderman2007). 204. Alebra neglectaisrecordednewtoSwedenbyGillerforsin2008(unpublisheddata). 207. Dikraneura variata is recorded new to Sweden by Gillerfors in 2008 (unpublished data). 208. Liguropia juniperiisrecordednewtoSwedenbyGillerforsin2008(unpublisheddata). 213. Forcipata palustrishasbycheckingHolgersensseriesprovedtobeagoodspecies.Itis sofarendemictoNorway. 214. Igutettix oculatus is recorded new to Estonia by Mannerkoski in 2006 (unpublished data). 220. Austroasca vittataisreportedbyVilbaste(1974)fromLithuania,butfromLatviaby Nast(1987). ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

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224. Empoasca affinisisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 228. Empoasca ossiannilssoniisrecordednewtoNorwaybyEndrestölin2007(Endrestöl 2008). 232. Vilbaste(1974)reportsKybos oshaniniZachvatkin,1953fromLatviaandLithuania. The species is difficult to discern fromKybos abstrusus and the specimens seen by Vilbastearepossiblymisidentifications.K. abstrususisrecordedasanewspeciesto NorwaybyEndrestöl(2008)andtoLithuaniabySöderman&Rintala(unpublished data). 234. Kybos limpidusisrecordednewtoSwedenbyGillerforsin2008(unpublisheddata). 236. Vilbaste (1974) reports Kybos verbaeZachvatkin,1953fromLatvia.Thetaxonisa junior synonym of K. mucronatus. Recorded new to Kaliningrad by Söderman & Shapovalin2007(unpublisheddata). 237. Kybos populiisrecordednewtoLithuaniabySöderman&Rintalain2008(unpub lisheddata). 238. Kybos rufescensisrecordednewtoFinlandbyAlbrechtin2008(unpublisheddata). 250. Zygina angustaisrecordednewtoLithuaniabySöderman&Rintalain2008(unpub lisheddata). 253. Zygina nigritarsishasbeenconfusedwithZ. flammigeraintheNordiccountries.Itis probablymuchmorewidespread. 259. Zygina schneideriisrecordednewtoFinlandbySödermanin2007(unpublisheddata). 260. Zygina suavisishereregardedasthetaxonlivingonRhamnus frangula(ie.sensuOs siannilsson). 268. Edwardsiana ampliataisrecordednewtoLithuaniabySöderman&Rintalain2008 (unpublisheddata) 270. Edwardsiana bergmaniisrecordednewtoLithuaniabySöderman&Dapkusin2007 (unpublisheddata). 275. Edwardsiana frustratorisrecordednewtoLithuaniabySöderman&Rintalain2008 (unpublisheddata). 279. ThespecificidentityofEdwardsiana kemnericanbequestioned.Specimensreported belongingtothistaxonmaybeparasitizedspecimensofE.tersa(seeNickel2003). 281. Edwardsiana lethierryiisrecordednew toLithuaniabySöderman&Rintalain2008 (unpublisheddata). 285. Edwardsiana plurispinosa is recorded new to Norway (Endrestöl, unpublished data) andLithuaniabySöderman&Rintala(unpublisheddata) 286. Vilbaste(1974)reportsEdwardsiana barbatafromLatvia,whichisregardedaformof E. prunicola. 292. Edwardsiana staminatahasbysomeauthorsbeensynonymisedwithE. avellanaeonthe samebasisaskemneri/avellanae. 295. SincethesynonymisationofEdwardsiana hippocastaniwithE. lethierryi(seeWilson& Claridge1999)previousrecordsofE. hippocastanihavebecomeunclearasthereare twospeciesE. ulmiphagusandE. plurispinosathathasbeenmixedbefore.Revisions ofmaterialfromNorway,DenmarkandSwedenrevealthatbothspeciesoccurin thesecountries.TheLithuanianrecordremainsunclearasVilbaste(1974)reportsit tohavebeencapturedfromSalix,notbeingthehostplantofeitherofthespeciesin question. ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

62 G.Söderman,G.Gillerfors,A.Enderstöl

297. Eupteryx adspersaisreportedfromDenmarkbyNast(1987)butnotbyOssiannilsson (1983). 300. Eupteryx aurataisrecordednewtoFinlandbySödermanin2008(unpublisheddata). 304. Eupteryx decemnotata is recorded new to Denmark by U.H. Nielsen on the web in 2007andnewtoFinlandin2008bySöderman(unpublisheddata)andSwedenin 2008byGillerfors(unpublisheddata). 305. Eupteryx floridaisrecordednewtoDenmarkbyU.H.Nielsenonthewebin2007. The species was recorded new to Kaliningrad in 2007 by Söderman & Shapoval (unpublisheddata). 306. Nast(1972)reportedthetaxonEupteryx ornata(Lethierry,1874)fromDenmarkand Latvia.ThisisajuniorsynonymofEupteryx heydenii.Inhiscatalogueof1987hehas omitted the species from NEurope because Vilbaste (1974) regards the Latvian findtobemuchtoofarnorth. 309. Eupteryx signatipennisisrecordednewtoLithuaniabySöderman&Rintalain2008 (unpublisheddata). 317. Eurhadina loewiihasbeenreportedfromEstonia(Vilbaste1974)andDenmarkand Sweden(Ossiannilsson1982)underthenameE. unticaDlabola,1969. 324. Linnavuoriana intercedens is regarded a separate species (Lauterer & Novotný 1991, Söderman2007)fromL. decempunctata. 328. Ribautiana tenerrima is recorded new to Lithuania by Söderman & Rintala in 2008 (unpublisheddata). 333. Zyginella pulchraisrecordednewtoDenmarkbyU.H.Nielsenonthewebin2007. RecordednewtoSwedenbyGillerforsin2008(unpublisheddata) 335. Utecha trivia(Germar,1821)isreported(asamemberofthegenusUlopa)byNast (1972, 1987) from Denmark, but the species is not mentioned by Ossiannilsson fromthiscountryagain(1981).ThespecieswasrecordednewtoLithuaniain2008 bySöderman&Rintala(unpublisheddata). 337. Allygus communisisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 338. Allygus maculatusisreportedbyNast(1987)fromFinland.Thisisanobviousmis print. 340. Allygus modestusisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 346. Cicadula frontalisisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 350. Cicadula ornataisrecordednewtoNorwaybyEndrestölin2006(unpublisheddata). 365. Euscelis plebejus(Fallén,1806)isreportedfromLatviabyNast(1972).Accordingto Vilbaste(1973)FlormisidentifiedtheLatvianspecimenanditshouldaccordingto himbeE. distinguendus. 370. TherecordofH. signiferfromNorthernEuropeissomewhatquestionablesincethe revelation of its biology (see Nickel 2003). We have not been able to check the specimeninquestion. 371. Nast(1987)regardstherecordofHardya tenuisfromLatviaasdubious,asVilbaste (1974)regardsitamisidentificationmadebyGimmerthal(1846). 386. Ophiola identica is probably identical with Limotettix plutonius recorded from Latvia andLithuaniabyVilbaste(1974)(seeSöderman2007) ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

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387. Ophiola orichalceaisbySöderman(2007)regardeddistinctfromO. cornicula.Thetaxa havenotbeenseparatedintheBalticcountriesyet. 393. Perotettix orientalisisreportedfromNorwaybyNast(1987)basedontherecordof Holgersen(1985). 395. Platymetopius guttatus is recorded new to Lithuania by Söderman & Rintala in 2008 (unpublisheddata). 400. Rhopalopyx vitripennisisreportedasadubiousrecordfromNorwaybyNast(1987).It isnotreportedbyOssiannilsson(1983)fromthiscountry. 414. Vilbaste (1974) reported the first Endria nebulosa records from Estonia under the nameAmphicephalus nebulosa. 417. Doratura exilisisrecordednewtoFinlandbySödermanin2007(unpublisheddata). 418. Doratura littoralisisrecordednewtoLatviain2006bySöderman&Spungis(2008). The specific statusof thistaxonmaybequestionedasthereareveryfewreliable charactersforidentification. 421. Fieberiella macchiaeisreportedfromLatviabyNast(1972).Thisspecieswasoriginally described from Israel by Linnavuori (1962). Later Nast (1987) omits it from the EuropeanlistbecauseVilbaste(1974)doubtsfindingsofFieberiellathisfarnorth.G. SödermanhascollectedspecimensofFieberiellafromCarnikavainLatviawhichcor respondtoF. septentrionalisanditisobviousthatearlierrecordsfromthesamere gionbelongtothisspecies.RecordednewtoSwedenbyGillerforsin2007(2008) andtoLithuaniain2008bySöderman&Rintala(unpublisheddata). 423. Balclutha arhenanaisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 438. Macrosteles nubilusisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 443. Macrosteles pygmaeuswasdescribedfromLithuaniabyVilbaste(1974).Basedonthe description its taxonomic status is critical and the specimen is question may be a parasitizedformofM. sexnotatus.It seemstotheauthorsthatanendemicspecies fromLithuaniaisnotveryprobable.M. sexnotatusisreportedfromIcelandbyNast (1987),butisnotknowntooccurthereaccordingtoErlingOlafssonfromtheIce landicMuseumofNaturalHistory. 448. Anufriev(2000a,2000b)reportsSonronius anderifromtheKolaPeninsula.Thefind ingsappeartobefarapartfromotherknownsites(locustypicusinsouthernSwe den and two sites in southern Finland) of the species.We have not been able to confirmAnufriev’sidentification. 451. Söderman(2007)regardsthespecimensofCirculifer haematocepsfromFinlandtobe longrangemigrants. 452. Neoaliturus guttulatusisregardedtobeaformofN. fenestratus(seeNickel2003). 456. Arocephalus languidus is reported from the Petsamo area in Murmansk (Lindberg 1932).Thisrecord,farfromanyotherinEurope,mustbeconsidereddubious. 461. Boreotettix bidentatusisrecordednewtoNorwaybyEndrestölin2007(unpublished data). 472. Erzaleus metriusisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 475. Jassargus alpinushasbeenamuchconfusingtaxonintheBalticcountries.Thenomi nalformlivesintheAlpsanditssubspeciesneglectusintheNorthernBorealRegion ofFennoscandia.J. neglectushasbeenreportedfromallBalticcountriesbyVilbaste ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

64 G.Söderman,G.Gillerfors,A.Enderstöl

(1974),alltheserecordsareacceptedbyOssiannilsson(1983)butneglectedbyNast (1987).Asthespeciesisknowntohaveseveralsubspecies,theisolatedareainthe Balticcountriesmaybelongtoaseparatesusbspecies“baltica”. 477. Jassargus distinguendusisreportedfrommanyareasbyVilbaste(1974),Ossiannilsson (1983)andNast(1987).InallthesecasesJ. distinguenduscorrespondstoJ. pseudocel- laris. 483. Metalimnus obtusus has recently been identified from Finland and western Russia (Söderman2007).RecordedfromSwedenbyGillerforsalreadyin2004butonlyre centlyidentified(unpublished).AlsorecordedinlighttrapmaterialfromLithuania in2007(Söderman&Dapkus,unpublished).ItisclosetoM. formosusandthemate rial of the latter has not been revised elsewhere in the eastern parts of Northern Europe. 484. Metalimnus steiniisrecordednewtoLithuaniabySöderman&Dapkusin2007(un publisheddata). 486. Mongolojassus sibiricus isreportedfromEstoniabyNast(1972)andVilbaste(1974). Later Nast (1987) reports both M. sibiricus and M. bicuspidatus from Estonia. The FinnishrecordsconcerningM. bicuspidatus(seeAlbrechtet al.2003)haveinarecent revisionturnedouttobelongtothisspecies. 489. Paralimnus zachvatkinihasrecentlybeenrecordedfromFinland(Albrechtet al.2006). ItisclosetoP. phragmitisandthematerialofthelatterhasnotbeenstudiedinten sivelyelsewherein NorthernEurope.RecordednewtoLithuaniain 2007(Söder man&Dapkus,unpublisheddata). 490. Paramesus in NorthEuropemay consistof twospecies.P. majormayoccurin the easternpartoftheregion.SofaronlyspecimensfromtheBalticcoasthavebeen checkedandtheyallbelongtoP. obtusifrons,butextensiveinlandmaterialfromRus siahasnotbeencheckedyet. 493. Psammotettix albomarginatus is reported from Norway as an “unreliable record” by Nast(1987)basedonSchulz(1976). 497. Psammotettix dubius is in earlier works mentioned as P. exilis. However, Söderman (2007)showedthatthenameexilishascausedmuchconfusionandthereforeallre cordsreferringtoP. exilisareinneedofrevision. 498. Psammotettix excisus is reported from Latvia by Nast (1987). Due to the confusion withP. dubius,theLatvianrecordisdubious. 499. Psammotettix frigidusisreportedfromLatviabySpungis&Kalnins(2002)basedon thelistofVelce&Danka(1970).Vilbaste(1974)regardsitasamisidentificationof P. pallidinervis. 507. Psammotettix slovacusisidentifiedfromEstoniaandLatvia(Vilbaste1974,Nast1987, Spungis&Kalnins2002)asP. makaroviMoravskaja,1952.AccordingtoSöderman (2007)thisisajuniorsynonymofP. slovacus. 4.3Dubiousspeciesrecordsomittedfromthecatalogue 1. Cixius simplex (HerrichSchäffer, 1835) has been reported from Denmark by Nast (1972),butitisnotincludedinOssiannilsson(1981)orNast(1987). 2. Tettigometra virescens(Panzer,1799)isreportedtooccurinLatviabyNast(1972),butis notincludedinhislaterlist(1987).Vilbaste(1974)believesittobeamisidentification byGimmerthal(1846).Theidentificationofthisspeciesisnoteasyandthespecimen inquestionmightbeamisidentifiedT. atra. ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

AnnotatedcatalogueoftheAuchenorrhynchaofNorthernEurope 65

3. Cercopis vulnerataRossi,1807hasbeenreportedfromNorwaybyNast(1972),butitis notincludedinlaterworks(Ossiannilsson1981,Nast1987). 4. Idiocerus aaliensisStrand,1913hasbeenreportedfromNorwaybyNast(1972),butitis notincludedbyOssiannilsson(1981)inhisbooksontheAuchenorrhynchaofDen mark and Fennoscandia. It is regarded a taxonomically unclear species. The type specimenmaybelostbecauseithasnotbeenfoundinOslo,RiganorBerlin,where Strand’scollectionissupposedtobe.Basedonthegivendescriptionthetypeisafe male, which Ossiannilsson regarded as a possible form of Metidiocerus elegans. Hol gersen(1944b)concurswiththisandconsidersI. aaliensis anomen confusum. 5. Idiocerus varius(Fabricius,1794)isreportedfromLatviabyVelce&Danka(1970).Itis regardedasdistinctfromI. stigmaticalisbyEmeljanov(1964),butVilbaste(1974)re gardsittobeajuniorsynonym.Ribaut(1952)mentionsI. variusasasynonymtoI. similis. 6. Metidiocerus impressifrons(Kirschbaum,1868)hasbeenreportedfromDenmarkbyNast (1972),butitisnotincludedinlaterworksbyOssiannilsson(1981)orNast(1987). 7. Penthimia nigra(Goeze,1778)hasbeenreportedfromLatviabyNast(1972),butitis notincludedbyVilbaste(1974)inhiscatalogueofLatvianandLithuanianAuchenor rhyncha.VilbasteregardsittobeamisidentificationmadebyGimmerthal(1846). 8. Kybos austriacusW.Wagner,1949isreportedfromLatviabySpungis&Kalnins(2002), butisevidentlyamisidentification. 9. Allygidius atomarius(Fabricius,1794)isreportedfromDenmarkbyNast(1972),butis notreportedinhislaterwork(1987).TherecordfromLatviabyFlor(1861)isanin correctnamingofA. commutatusaccordingtoVilbaste(1973). 10. Platymetopius henribautiisacriticaltaxon(seeNickel2003,Söderman2007).Theaeda gusdepictedbyOssiannilsson(1983)correspondstothisspecies,butitisuncertainif thespecimendrawnbyhimhasbeencollectedinSweden. 11. Cicadula aurantipes(Edwards,1894)isreportedfromDenmarkbyNast(1972).After Ossiannilsson’swork(1983)heregardstheDanishrecordunreliable.Thetaxonitself isquitecritical(seeSöderman2007). 12. Jassargus obtusivalvis (Kirschbaum,1868)isreportedfromDenmarkbyNast(1972),but helater(1987),afterOssiannilssonswork(1983)omitsthespeciesfromthiscountry. 13. Jassargus repletus(Fieber,1869)isreportedfromLatviaandtheKaliningradregionby Nast(1972,1987)andVilbaste(1974).Thespeciesismostlyconfinedtoalpinere gionsinEurope.TherecordsfromtheeasternpartsoftheBalticsareproablydueto misidentifications. 14. Psammotettix helvolus (Kirschbaum, 1868) is reported from Denmark by Nast (1972), but he later (1987), after Ossiannilssons work (1983) omits the species from this country. 15. Psammotettix ornaticeps(Horvath,1897)isreportedfromLatviabySpungis&Kalnins (2002)basedonthelistofVelce&Danka(1970).Vilbaste(1974)statesthisisamis identificationofP. poecilus. 16. Deltocephalus ignoscusFieber,1869isdescribedfromLatvia.Thestatusofthistaxonis veryunclear. ©Arbeitskreis Zikaden Mitteleuropas e.V. - download unter www.biologiezentrum.at

66 G.Söderman,G.Gillerfors,A.Enderstöl

5.Discussion ThetotalnumberofrecordedspeciesinNorthernEuropeisnow513,whichislowif compared to e.g. Germany, holding more than 600 species alone (Nickel & Remane 2002).ThenumberofspeciesdecreaseswithlatitudeandisolationfromcoresofPleisto ceneglacialrefugia(e.g.Iceland).ThenumberofspeciesinNorthernEuropeis,however, increasingatafastpace.Thirtyonenewspeciesfortheregionhasbeenrecordedinthe 21stcenturyandforsomecountriestheproportionofnewspeciesincreases1020percent ofthefauna. Thereareatleast3majorexpansionroutesforleafhopperstoNorthernEurope.One goesfromNorthwesternGermanyovertheDanishislandstotheeasterncoastofKat tegatt(e.g.Graphocephala fennahi, Acericerus heydenii, A. ribauti, Tremulicerus fulgidus, Eupteryx decemnotata, E. florida, Zyginella pulchra, Acanthodelphax spinosa, Eurybregma nigrolineata), an otherfromwesternRussiaoverPskovandtheLeningradregiontosoutheasternFinland (e.g.Xanthodelphax xantha, Calamotettix taeniatus, Psammotettix slovacus, Mongolojassus sibiricus, Balclutha arhenana, Igutettix oculatus).AthirdonegoesovertheBalticcountriesreachingthe Baltic isles (Gotland, Alandia) and the southwestern Finnish coast (e.g. Kelisia confusa, Acanthodelphax spinosa, Acericerus heydenii, Fieberiella septentrionalis, Idiocerus similis, Tremulicerus distinguendus, Viridicerus ustulatus). This route might consist of two different routes, one overLithuaniaandLatviareachingGotland,andoneoverEstoniareachingFinland,but thepresentknowledgeofthefaunaoftheBalticcountriesdoesnotallowustodrawany conciseconclusionofthis,yet. Aboutonethirdoftheexpandingspecieslivesonornamentalbushesandtreesnot indigenoustoNorthernEurope.Nevertheless,noneofthesespeciescanbeconsidered importedtotheregion.Theyhaveadaptedtoliveinanthropogenoushabitatsandsome areshowingtrendsofhostplantchangetowildspecies,e.g.Igutettix oculatusfromSyringa toFraxinusandTremulicerus distinguendusfromPopulus albatoP. tremula. Acknowledgements Wewishtothankthefollowingpersonsforgivingustheopportunitytoanalysefreshly collectedmaterial:NilsRyrholm(GävleHighschool),PeeterTarlap(Tallinn), Voldemars Spungis (University of Riga), Dalius Dapkus (Vilnius Pedagogic University) AnatolyiP.Shapoval(St.PetersburgUniversity),NataliaGoltsova(St.PetersburgUniver sity), Natalia Kutenkova (Kiwach Nature Reserve in Petroskoi) andTeemu Rintala (Jy väskylä). The following colleagues kindly provided us with new records and help to study musealspecimens:U.H.Nielsen(Copenhagen),FrodeÖdegaard(NorwegianInstitutefor Nature in Trondheim), Lars Ove Hansen (Natural History Museum in Oslo), Anders Albrecht(CentralMuseumofNaturalHistoryinHelsinki),VladimirGnedzilov(St.Pe tersburgUniversity)andDimitryA.Dimitriev(ZoologicalInstituteinSt.Petersburg).

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