INSECTA MUNDI, Vol. 15, No.3, September, 2001 151

A checklist of the Limnichidae and the Lutrochidae (Coleoptera) of the world

P. J. Spangler, C. L. Staines, P.M. Spangler, and S.L. Staines Department of Entomology National Museum of Natural History Smithsonian Institution Washington, DC 20560

Abstract. A checklist of the world species of Limnichidae (35 genera, 345 species) and Lutrochidae (1 genus, 11 species) is presented. The author, year of publication and page number, synonyms, distribution by country, and a terminal bibliography are given for each genus and species. Biological information is also reviewed.

Introduction scribed the species and adult behavior of Lutrochus arizonicus Brown and Murvosh. Hilsenhoff and The Limnichidae Erichson (1847) and Lutro­ Schmude (1992) reported that adults of L. laticeps chidae Kasap and Crowson (1975) are small semi­ Casey (1893) were found in small to large, warm, aquatic beetles; adults of both families most fre­ calcareous streams; that adults oviposit on sub­ quently live in riparian habitats. Both families merged wood or on travertine rocks; and larvae were previously assigned to the superfamily Dry­ were aquatic and were collected on the travertine opoidea (Crowson, 1978); however, they are now rocks. placed in the Byrrhoidea (Lawrence and Newton, Previous world catalogs that included the lim­ 1995). From 1975 through 1993, D. P. Wooldridge nichids and lutrochids are by Zaitzev (1910) and published numerous limnichid descriptions and von Dalla Torre (1911) who recorded 11 genera and revisions but much systematic work remains to be 75 species. Blackwelder (1944) published a check­ done for the Limnichidae, especially in the tropical list of the Coleoptera of Mexico, Central America, regions of the Eastern Hemisphere. the West Indies, and South America in which he Not much has been published dealing with the reported 11 genera and 44 species of Limnichidae biology of the families. Paulus (1970) described the and 8 species of Lutrochidae in the nominate genus. larva of the European species Limnichus sericeus Wooldridge (1986b) published a catalog of the Lim­ Duftschmid (1825) and the larva and pupa of Pelo­ nichidae (including Lutrochus) of North America in chares versicolor Waltl (1838). Paulus also reported which he included 6 genera and 29 species of that the larvae of both genera are terrestrial and Limnichidae and the single genus, Lutrochus with live in the soil under the "Chlorophyceae-Bryophy­ 3 species. Some recent revisonary studies on the ta layers," and reportedly feed on those plants. Limnichidae are those by Wooldridge (1976, 1977a, Adults may be found near the waterline on 1977b, 1978, 1979, 1980, 1981b, 1982, 1983, 1987b). branches and grasses hanging over stream banks Additional studies that have added to our knowl­ or in riffles, on driftwood, in logjams, and on rocks edge of the Limnichidae are by Darlington (1936), in streams; some are found in splash zones beside fauna of the West Indies; Wu (1937), China; Sato cascades and waterfalls and on dam breasts. Spec­ (1965), Formosa; Sato (1966, 1976, 1997), Japan; imens of at least five limnichid genera, Pseudeu­ Deleve (1968, 1974), Herando and Ribera (2000a), cinetus Heller (1921), Martinius Spilman (1959), Africa; Deleve (1971,1973), Asia; Wooldridge (1975), Hyphalus Britton (1971, 1973, 1977), Throscinus Western Hemisphere; Piitz (1991b), Germany; LeConte (1874) (fide Wooldridge, 1981), and Mexico Mascagni (1995), Italy; Piitz (1998a, 1998b), China Spilman (1972) are known to live on coral reefs and neighboring countries; Ribera and Hernando (Sate, 1997; Hernando and Ribera, 2000b), coastal (1999), and Hernando and Ribera (2000b), Oriental lagoons, mangrove swamps, or edges of mud or Region. rocky flats along coastal areas. This checklist is an effort to bring the limnichid Brown (1965) discussed characteristic micro­ .and lutrochid names and pertinent taxonomic liter­ habitats of various dryopoids including Lutrochus ature into a single publication and was compiled by Erichson (1847). Brown and Murvosh (1970) de- examining the literature for descriptions, synonyms, 152 Volume 15, No.3, September, 2001, INSECTA MUNDI and distributional data plus examination of the biroi Pic, 1956:75 collection of the Smithsonian Institution. New Guinea bomansi (Deleve), 1968:245 A checklist of the Limnichidae Tanganyika of the world borneensis Pic, 1923b:7 Borneo ACONTOSCELES Champion, 1924a:27 boucardi Pic, 1923b:7 hydroporoides Champion, 1924a:29 Sumatra Taiwan [Formosa], India brevior (Pic), 1927a:65 tagalog Spilman, 1959:116 Sumatra Philippines convexus (Pic), 1923b:5 yorioi Sato, 1966:60 Philippines Japan (Ryukyus) cribrosus Champion, 1923:253 India, Sri Lanka AFROLIMNICHUS Deleve, 1968:259 curtus Pic, 1935b:471 brevicornis Deleve, 1968:260 Sierra Leone Congo densepunctatus (Pic), 1923b:6 oblongus Deleve, 1974:40 Ivory Coast South Africa (Cape Province) diversus (Pic), 1923b:5 Ivory Coast BABALIMNICHUS Sato, 1994:174 ellypticus Motschulsky, 1858:51 masamii Sato, 1994:175 India Japan (Ryukyus) euryaspis Champion, 1923:250 taiwanus Sato, 1994:174 India [Nilgiri Hills] Taiwan ferax Wooldridge, 1993b:359 Philippines, Borneo BOTHRIOPHORUS Mulsant and Rey, 1852:19 flavicornis Champion, 1923:251 atomus Mulsant and Rey, 1852:20 India (syn., ventetus Ferrari, 1864:108) formosanus Pic, 1935a:2 Europe Taiwan [Formosa] fuscus Motschulsky, 1858:52 BYRRHINUS Motschulsky, 1858:50 India syn., BYRRHININUS Pic, 1922:6 goldbachi Wooldridge, 1987b:310 syn., CYPHONICHUS Sharp, 1902a:679 Ecuador, Paraguay syn., NOTIOCYPHON Blackburn, 1896:272 gracilicornis Pic, 1922:5 syn., PELOCHEROPS Pic, 1923a:3 Brazil, Paraguay, Venezuela amoenus Wooldridge, 1987a:310 grandis (Pic), 1956:74 Belize, British Guiana, Colombia, Ecuador New Guinea andamanus (Pic), 1935a:2 grossepunctatus (Deleve), 1968:249 Andaman Islands Congo angustatus Motschulsky, 1858:51 hargreavesi (Pic), 1935b:470 India, Thailand Sierra Leone antipodium Fauvel, 1903:348 hirsutulus Champion, 1923:250 New Caledonia India [Kumaon] apicalis Pic, 1922:5 hirsutus Pic, 1922:5 Katanga French Guiana, Trinidad atricornis (Pic), 1935a:3 hulstaerti (Pic), 1952:9 Congo, Sierra Leone Kinshasa [Congo] atrolucidus (Pic), 1923a:3 improcerus Wooldridge, 1987b:310 New Guinea Ecuador binhanus (Pic), 1923b:5 inlineatus Pic. 1923c:15 Vietnam India INSECTA MUNDI, Vol. 15, No.3, September, 2001 153 irregularis Pic, 1922:5 plenus Wooldridge, 1987b:305 Philippines Boliva, Ecuador jacobsoni Pic, 1927a:65 pubiventris Lea:1920:277 Sumatra Australia latus Motschulsky, 1858:50 punctaticeps Pic, 1927b:49 Bangladesh, Burma, India, Sri Lanka Sumatra leleupi (Deleve), 1968:257 punctatus (Pic), 1922:5 Kivu [Zaire] Philippines leonensis (Pic), 1923b:6 punctulatus (Deleve), 1968:243 Congo, Sierra Leone Upemba [Zaire] maculatus Wooldridge, 1987b:307 reitteri (Pic), 1923b:6 Ecuador Ghana magnus Wooldridge, 1987b:307 rhomboideus Champion, 1923:253 Ecuador, Peru, India marginatus Champion, 1923:249 rufoapicalis (Pic), 1955:132 India [Kumaon], Pakistan Ruanda, Kiva [Zaire] maroniensis Pic, 1923b:7 rufus Pic, 1935a:2 Brazil, Colombia, French Guiana, Trinidad­ Thailand Tobago, Venezuela schoutedeni (Pic), 1952:9 mimicus Wooldridge, 1987b:308 Congo Colombia, Panama semirufus Pic, 1923b:8 misellus (Sharp), 1902a:681 Trinidad -Tobago, Venezuela Colombia, Costa Rica, Mexico, Nicaragua, seydeli (Deleve), 1968:252 Panama Elisabethville [Zaire] monilicornis Champion, 1923:251 sikkimensis Champion, 1923:249 India [Kumaon] India [Sikkim] neoguineeneis (Pic), 1956:75 subglobosus (Deleve), 1968:254 New Guinea Elisabethville [Zaire] niger Pic, 1922:5 subglobosus upembanus (Deleve), 1968:256 Vietnam Upemba [Zaire] nitidicollis Pic, 1956:75 subregularis Pic, 1923c:15 New Guinea Vietnam nitidulus Motschulsky, 1858:51 subtestaceous Pic, 1923b:8 India, Thailand Philippines noctivagus Lea 1920:277 tarawakanus Deleve, 1973:23 Australia Borneo [Sabah], Philippines (Tawi Tawi) oculatus Motschulsky, 1858:52 testaceicornis (Pic), 1923b:5 India Philippines olibroides (Sharp), 1902a:679 testaceipes Pic, 1922:5 (syn., celatus Sharp, 1902a:680) subregularis Pic, 1923b:7 Lapsus Calami Belize, El Salvador, Guatemala, Mexico Sumatra orientalis (Sharp), 1902b:63 tschoffeni (Deleve), 1968:235 Taiwan [Formosa], Malaysia (Borneo) Boma [Congo] padangus Pic, 1927b:49 vatovai (Pic), 1953:113 Sumatra Ethiopia panamensis (Sharp), 1902a:679 vestitus (Sharp), 1902a:681 (syn., mollis Sharp, 1902a:681) Guatemala, Mexico (syn., rufipennis Pic, 1923b:7) vicinus (Deleve), 1968:248 Belize, Colombia, Costa Rica, Guatemala, Congo Panama wittei (Deleve), 1968:253 peguensis Pic, 1923c:15 Upemba [Zaire] India 154 Volume 15, No.3, September, 2001, INSECTA MUNDI

CACCOTHRYPTUS Sharp, 1902b:63 nepalensis Piitz, 1998b:360 syn., MACROBYRRHINUS Pic, 1922:4 Nepal [Myagdi] compactus Sharp, 1902b:63 schillhammeri Piitz, 1998b:362 Borneo, Vietnam China [Sichuan] laosensis Pic, 1928:2 schuelkei Piitz, 1998b:363 Laos China [Sichuan] maculosus Pic, 1923a:3 sichuanensis Piitz, 1998b:365 Tonkin China multiseriatus Champion, 1923:224 subelongatus Pic, 1923b:4 India [Bengal] India [Sikkim] ripicola Champion, 1923:223 India [W. Almora] CHIBIDORONUS SaW, 1966:59 rouyeri (Pic), 1922:4 aureus Sat6, 1966:60 Sumatra Japan (Ryukyus) testudo Champion, 1923:222 India [W. Almora], Nepal CORRINEA Wooldridge, 1980:69 admirandus (Sharp) 1902a:677 CEPHALOBYRRHINUS Pic, 1923b:6 Mexico brevis Wooldridge, 1986a:512 audax Wooldridge, 1980:75 Ecuador Mexico curticornis Pic, 1923b:6 aurata Wooldridge, 1980:71 Costa Rica, Mexico, Nicaragua, Panama, Brazil, Ecuador Venezuela delicata Wooldridge, 1980:75 impressopunctatus Wooldridge, 1977b:32 Colombia, Costa Rica, Mexico, Panama Bolivia edita Wooldridge, 1993a:36 lineatus Wooldridge, 1986a:513 Venezuela Costa Rica, Panama incondita Wooldridge, 1980:73 robustus Wooldridge, 1986a:512 Bolivia, Colombia, Ecuador Ecuador, Peru lecta Wooldridge, 1980:74 sedatus Wooldridge, 1986a:513 Bolivia Ecuador parva Wooldridge, 1980:74 Bolivia, Ecuador, Peru CEPHALOBYRRHUS Pic, 1923b:4 placida Wooldridge, 1980:76 bertiae Piitz, 1998b:344 Trinidad China [Yunnan] salebrosa Wooldridge, 1980:75 brevipalpus Piitz, 1998b:346 Guatemala China [Yunnan] spicula Wooldridge, 1980:73 emeishanensis Piitz, 1998b:347 Ecuador China [Sichuan] splendida Wooldridge, 1980:71 gibbicollis Champion, 1925a:175 Ecuador, Peru India [Kumaon] vianai (Pic), 1939:249 jaechi Piitz, 1998b:351 Argentina China [Sichuan] japonicus Champion, 1925a:176 CYCLOLIMNICHUS Deleve, 1968:270 Japan dentoni Hernando and Ribera 2000a:216 jiangxiensis Piitz, 1998b:354 Cameroon China [Jiangxi] jaechi Hernando and Ribera 2000a:215 jinggangshanensis Piitz, 1998b:355 Kenya China [Jiangxi] ovalis Hernando and Ribera 2000a:215 latus Pic, 1923b:4 Cameroon Taiwan [Formosa] presignis Deleve, 1968:272 longipalpis Piitz 1998:359 Congo China [Jiangxi] INSECTA MUNDI, Vol. 15, No.3, September, 2001 155

ERSACHUS Erichson 1847:497 Mexico, USA (AZ, CA, CO, NM, OR) erichsonianus Sharp, 1902a:683, pl. 20 nitidulus (LeConte), 1854:117 Guatemala, Mexico, Panama Argentina, Brazil, Costa Rica, Ecuador, lugubris Sharp, 1902a:684 Mexico, Nicaragua, Panama, USA (AL, FL, Guatemala GA, IN, ME, MO, NC, NH, PA, SC, VA) mexicanus Sharp, 1902a:684 obscurus (LeConte), 1854:116 Mexico USA (DC, IL, IN, KS, MD, MS, NC, NY, TN, variegatus Erichson, 1847:497 TX, VA) Colombia optatus Sharp, 1902a:675 Guatemala, Mexico, USA (AL, GA, LA, TX) EULIMNICHUS Casey, 1889:146 pellucidus Wooldridge, 1979:5 acutus Wooldridge, 1979:7 Venezuela Costa Rica, Colombia, EI Salvador, Guatemala, perpolitus Casey, 1889:151 Honduras, Mexico, Panama USA (AZ, CA, NM, TX) analis (LeConte), 1879:515 plebius Sharp, 1902a:675 (syn., analis rugiceps Casey, 1912:47) (syn., Limnichoderus convexus Pic, 1922:4) (syn., analis coloradensis Casey, 1912:47) (syn., Limnichoderus breyeri Pic, 1930:43) Mexico, Panama, USA (AR, AZ, CA, CO, MO, (syn., Limnichoderusperuvianus Pic, 1954:178) OR, TX, UT, WA) Argentina, Brazil, Colombia, Dominican ater (LeConte), 1854:117 Republic, Ecuador, EI Salvador, Guatemala, (syn., guatemalicus Sharp, 1902a:676) Haiti, Honduras, Mexico, Puerto Rico, Peru, (syn., sculpticeps Casey, 1912:50) Trinidad, Venezuela (syn., Limnichoderus oblongus Pic, 1922:4) rugulosus Wooldridge, 1979:6 Argentina, Belize, Bolivia, Brazil, British USA (IL, MI, WI) Guiana, Colombia, Costa Rica, Cuba, rusticus Wooldridge, 1979:7 Ecuador, Guatemala, Honduras, Jamaica, Mexico Mexico, Panama, Peru, USA (FL, GA, IN, sharpi Wooldridge, 1979:5 LA, MA, MS, NC, SC, TX), Venezuela Mexico californicus (LeConte), 1879:515 sordidus Sharp, 1902a:676 USA (CA, OR) Guatemala coheni Wooldridge, 1979:4 spangleri Wooldridge, 1979:8 Bolivia, Ecuador EI Salvador, Guatemala, Honduras, Mexico confertus Sharp, 1902a:676 sub titus Wooldridge, 1979:8 Mexico, Guatemala USA (AL, FL) corrineae Wooldridge, 1979:3 sublaevis Sharp, 1902a:674 Argentina, Brazil, Venezuela Guatemala, Panama epistemus Sharp, 1902a:673 uisendus Wooldridge, 1979:1 Costa Rica, Guatemala, Nicaragua, Mexico Mexico evanescens Casey, 1912:48 USA (CA) EUTHRYPTUS Sharp, 1902a:682 expeditus Wooldridge, 1984b:57 orbicularis Sharp, 1902a:682 Guatemala, Honduras Guatemala, Mexico, Panama impostus Wooldridge, 1978:172 trigonalis Sharp, 1902a:682 (syn. elongatus Pic, 1923b:7; preoccupied) Guatemala Brazil improcerus Wooldridge, 1979:4 HYPHALUS Britton, 1971:88 USA (MA) crowsoni Hernando and Ribera 2000b:240 incultus Wooldridge, 1979:6 Seychelles [Aldabra] Bolivia, Ecuador insularis Britton, 1971:90 langleyae Wooldridge, 1979:3 Australia Ecuador, Peru kuscheli Britton, 1977:82 montanus (LeConte), 1879:514 New Zealand (syn., laeticulus Sharp, 1902a:674) 156 Volume 15, No.3, September, 2001, INSECTA MUNDI prolixus Britton, 1977:85 angustus Wooldridge, 1981b:180 New Zealand Colombia, Ecuador, Panama, Venezuela taekoae Sat6, 1997:110 barberi Wooldridge, 1981b:174 Japan (Ryukyus) Haiti, Puerto Rico, Trinidad-Tobago ultimus Britton, 1977:85 conspectus Wooldridge, 1981b:181 New Zealand Ecuador, Peru wisei Britton, 1973:121 cupidus Wooldridge, 1981b:178 New Zealand Brazil curtulus Sharp, 1902a:678 JAECHOBYRRHINUS Piitz, 1991a:132 (syn., minutus Pic, 1922:4) amanosius Piitz, 1991a:132 Costa Rica, Panama Turkey durus Wooldridge, 1981b:182 Brazil LICHMINUS Casey, 1889:147 excelsus Wooldridge, 1993a:35 tenuicornis Casey, 1889:155 Venezuela USA (CA) fragosus Wooldridge, 1981b:178 Ecuador LIMNICHITES Casey, 1889:147 imprioris Blackwelder, 1944:274 browni Wooldridge, 1977a:187 (syn., oblongus Brethes, 1922:297; preoccupied) EI Salvador, Mexico Argentina confertus (Sharp), 1902a:676 insularis Wooldridge, 1981b:188 Costa Rica, Guatemala, Honduras, Jamaica, Dominican Republic, Haiti, Puerto Rico Mexico, Nicaragua lutrochinus (LeConte), 1879:515 foraminosus Casey, 1912:43 (syn., indiscretus Sharp, 1902a:678) Mexico, USA (AZ, CA, NM, OR, TX, UT) (syn., lanosus Casey, 1912:55) huronicus Casey, 1912:43 Belize, Costa Rica, EI Salvador, Guatemala, Canada, USA (IL, MA, MI) Honduras, Mexico, Panama, USA (AZ, CA, imparatus Wooldridge, 1977a:181 IL, OK, TX) Mexico magnus Wooldridge, 1981b:179 nebulosus (LeConte), 1879:515 Costa Rica, Guatemala, Mexico (syn., densissimus Casey, 1912:41) modicus Wooldridge, 1981b:176 Mexico, USA (AR, AZ, CA, OK, OR, TX) Costa Rica, Ecuador, EI Salvador, Guatemala, olivaceus (LeConte), 1854:116 Honduras, Panama (syn., virginicus Casey, 1912:42) moratus Wooldridge, 1981b:185 USA (AL, AR, CT, DC, lA, IL, IN, MA, MD, MI, Colombia, Costa Rica, Guatemala, Mexico, MO, NC, OH, OK, PA, TX, VA, WV) Panama perforatus (Casey), 1889:155 naviculatus (Casey), 1889:156 Mexico, USA (CA) Costa Rica, EI Salvador, Guatemala, Honduras, porrectus Wooldridge, 1977a:182 Mexico, Nicaragua, Panama, Puerto Rico, Mexico USA (AZ, CA, NM, NV, OK, TX) punctatus (LeConte), 1854:116 ovatus (LeConte), 1854:117 (syn., austinianus Casey, 1912:42) (syn., pulvereus Casey, 1912:54) Canada, USA (AL. DC, GA, lA, IL, MA, MD, Mexico, USA (AL, FL, GA, LA, NC, SC, TX) ME, MI, MN, MS, NH, NJ, NY, OH, OK, PA, placidus Wooldridge, 1987a:330 RI, TX, UT, VA, WI) Costa Rica rudis Wooldridge, 1977a:183 plenus Wooldridge, 1981b:185 Cuba, Jamaica Mexico simplex Wooldridge, 1977a:188 punctiventris (Casey), 1889: 159 Costa Rica, Mexico USA (FL) seclusus Wooldridge, 1987a:330 LIMNICHODERUS Casey, 1889:147 Ecuador amoenus Wooldridge, 1981b:179 seriatus (Casey), 1889:158 Ecuador (syn., floridanus Casey, 1912:55) INSECTA MUNDI, Vol. 15, No.3, September, 2001 157

USA (AL, FL, GA, LA, MS, NC, SC, VA) inornatus Weise, 1877:301 similis Wooldridge, 1981b:182 (syn., punctipennis Baudi, 1870:64) Peru Cyperus vicinus Pic, 1922:4 javanicus Pic, 1922:3 Brazil, Colombia, Ecuador, Panama, Venezuela Java latiusculus Reitter, 1901:72 LIMNICHOMORPHUS Pic, 1922:4 Turkestan curtulus Pic, 1922:4 lederi Weise, 1877:302 Vietnam (Tonkin) Caucasus ohbayashii Sato, 1966:58 lewisi Nakane, 1963:43 Japan (Ryukyus) (syn., paululus Lewis, 1879:12) China, Japan, Russia LIMNICHUS Latreille, 1829:510 longitarsis Champion, 1923:275 aeneipennis Pic, 1935a:l India [Sarda] Vietnam marmoratus Fairmaire, 1898:224 africanus Pic, 1923a:3 Madagascar Ghana micraspis Champion, 1923:276 angustulus Weise, 1877:300 India [Sarda] Sardinia monticola Deleve, 1968:220 astruci Pic, 1922:3 Kenya India murinus Baudi, 1870:63 ater Lea 1920:279 Cyperus Australia nigripes Broun, 1893:1449 aurosericus Jacquelin du Val, 1857 (1859):268 New Zealand (syn., leprieuri Perris, 1864:282) obliteratus Champion, 1923:275 Algeria, France, Italy, Spain India [Kumaon] australis Erichson, 1842: 153 orientalis Motschulsky, 1861:112 Australia, Tasmania Sri Lanka brevior Fairmaire, 1898:224 palawanus Deleve, 1973:18 Madagascar Philippines (Palawan) coomani Pic, 1935a:l picinus Broun, 1881:671 Vietnam New Zealand cuneatus Champion, 1923:275 pruinosus Fleischer, 1909:244 India [Kumaon] Samarkand curtus Pic, 1922:3 pumilio Obenberger, 1916:29 Java Corfu decorus Broun, 1880:247 punctatus Broun, 1880:247 New Zealand New Zealand demoulini DeIENe, 1968:217 punctipennis Kraatz, 1858:148 Congo Greece, Italy ellipticus Lea 1920:280 pygmaeus Sturm, 1807:114 Australia (syn., sericeus Stephens, 1832:411) [alsus Deleve, 1971:371 (syn., auricomus Reitter, 1877 (1878):180) India (syn., minutus Rey, 1889:66) fragilicornis Wollaston, 1868:80 (syn., variegatus Guillebeau, 1886:2) Cape Verde Islands (syn., pygmaeus var. Reitter, 1887:104) fulvopubens Pic, 1922:3 Turkestan, middle Europe, Caucasus China ruficolor Pic, 1935a:2 globosus Champion, 1923:276 N. Borneo India [Kumaon] rufipes Pic, 1923b:6 incanus Kiesenwetter, 1851:584 Ivory Coast Mediterranean area, southern Europe, rufulopubens Fairmaire, 1897:368 Caucasus, Turkestan Madagascar 158 Volume 15, No.3, September, 2001, INSECTA MUNDI rufus Pic, 1923b:7 PARATHROSCINUS Wooldridge, 1984c:121 Philippines (Luzon) corrinae Wooldridge, 1990:254 saegeri Deleve, 1968:218 Singapore Congo magnus Wooldridge, 1990:254 sericeus Duftschmid, 1825:24 Singapore Europe, Asia murphyi Wooldridge, 1990:251 subchalybeus Reitter, 1881:85 Singapore Caucasus oculatus (Motschulsky), 1858:52 subelongatus Pic, 1922:3 Burma, India Sumbawa orientalis Wooldridge, 1984c:122 subnotatus Champion, 1923:273 Philippines, Vietnam India [Kumaon] sulcatulus Pic, 1922:3 PELOCHARES Mulsant and Rey, 1869:161 Guadeloupe acuminatus Champion 1923:271 tonkineus Fairmaire, 1888:337 India [Sikkim] Tonkin, Vietnam acutipennis Pic, 1935a:2 trachyformis Pic, 1922:3 Malaysia India [Kumaon] alluaudi Deleve, 1968:226 turkestanicus Pic, 1905:97 Africa Orientale Turkestan basilewskyi (Pic), 1955:131 unistriatus Champion, 1923:274 Ruanda India [Kumaon] biroi (Pic), 1956:75 vagegluttatus Fairmaire, 1898:223 New Guinea Madagascar congoensis Deleve, 1968:221 waelbroecki Deleve, 1968:215 Congo Kinshasha densatus Pic, 1922:3 Java MANDERSIA Sharp, 1902b:62 diversus (Pic), 1922:3 scymnoides Sharp, 1902b:62 Philippines (Los Banos) Burma, China [Shan states], India emarginatus Mulsant and Rey, 1869:161 Europe MARTINIUS Spilman, 1959:118 gibbipennis Champion, 1923:269 ripisaltator Spilman, 1966:124 Europe, Burma, India [Bengal], Sri Lanka Cuba minutus Deleve, 1968:224 tellepontis Spilman, 1959:119 Kivu [Zaire] Panama murinus Baudi, 1870:63 temporalis Wooldridge, 1988:314 Cyperus, Caucasus Ecuador oblongulus Champion, 1925b:263 India (Kumaon) MEXICO Spilman, 1972:113 orientalis (Motschulsky), 1861:112 litoralis Spilman, 1972:114 (syn., horni Deleve, 1971:368) Mexico Sri Lanka philippinensis Deleve, 1973:21 PARALIMNICHUS Deleve, 1973:24 Philippines (Tawi Tawi) castaneus (Lea), 1920:279 rugiventrus Champion, 1923:270 Australia (syn., ceylanicus Deleve 1971:370) panamensis Wooldridge, 1983:232 India [Kumaon], Sri Lanka Panama ryukyuensis Sato, 1966:57 punctatus (Pic), 1923b:5 China, Japan (Ryukyus) (syn., punctatus var. obscurior Pic, 1923b:5) sulciger Champion, 1923:270 (syn., rufeecens Pic, 1923b:5) Sri Lanka Borneo, Guadalcanal, Philippines [Banguey] trifidus Champion, 1923:272 Burma INSECTA MUNDI, Vol. 15, No.3, September, 2001 159

versicolor Waltl, 1838:273 PLATYPELOCHARES Champion,1923:271 (syn., emarginatus Mulsant and Rey, 1869:161) syn. WOOLDRIDGEUS Spangler, 1999:181 (var., opacus Reitter, 1881:84) latimargo Champion, 1923:272 Europe, New Caledonia India [Kumaon] vitalisi Pic, 1935a:1 perforatus (Spangler), 1999:182 Cambodia Malaysia periculosissimus Ribera and Hernando, 1999:107 PHALACRICHUS Sharp, 1902a:678 Loas, Thailand, Vietnam syn., LIMNICHALIA Casey, 1912:52 petrus Ribera and Hernando, 1999:108 atomarius Sharp, 1902a:679 Malaysia (syn., colonica Casey, 1912:52) trifidus Champion, 1923:272 Belize, Bolivia, Colombia, Costa Rica, Ecuador, Burma El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Surinam, PSEUDEUCINETUS Heller, 1921:155 Trinidad, Venezuela syn., THAUMASTODUS Champion, 1924a:25 diligens Wooldridge, 1982:389 PSEUDOEUCHINETUS Sate, 1994:173 Bolivia, Brazil, Colombia, Venezuela (misspelling) durus Wooldridge, 1982:387 novabritannica Deleve, 1973:18 Bolivia, Brazil, Ecuador, Paraguay, Peru, Bismarck (New Britain) Venezuela spilmani Spangler, 1995:399 golbachi Wooldridge, 1982:388 Indonesia Paraguay uenoi Spangler, 1995:396 latus Wooldridge, 1982:386 Malaysia Brazil, Colombia, Ecuador, Peru, Venezuela zygops Heller, 1921:156 punctatus Wooldridge, 1982:389 (syn., Thaumastodus fueiformis Champion, Bolivia, Brazil, Colombia, Panama, Venezuela 1924a:27) rudis Wooldridge, 1982:385 India, Malaysia, New Ireland, Philippines, Surinam, Venezuela Solomon Islands simplex Wooldridge, 1982:387 Bolivia, Brazil, Venezuela RESACHUS Deleve, 1968:263 spangleri Wooldridge, 1982:386 lineatifrons Deleve, 1968:266 Ecuador, Venezuela Congo sublimus Wooldridge, 1993a:36 striatellus Deleve, 1968:263 Venezuela Congo vianai (Pic), 1938:77 Argentina, Brazil, British Guiana, Trinidad SIMPLOCARINA Pic, 1922:4 syn., CYRTOLIMNICHUS Deleve, 1968:267 PHYSEMUS LeConte, 1854:117 curticollis Pic, 1922:4 syn., DITAPHRUS Casey, 1886:250 (syn., punctulatus Deleve, 1968:267) excavatus Wooldridge, 1976:182 Congo El Salvador, Guatemala, Honduras latifrons Wooldridge, 1976:180 THROSCINUS LeConte, 1874:51 Panama aethiops Darlington, 1936:75 levis Wooldridge, 1984a:538 Cayman Islands, Cuba, Dominican Republic, Brazil, Trinidad, Venezuela Haiti, Jamaica, Puerto Rico minutus LeConte, 1854:117 crotchi LeConte, 1874:52 (syn., Ditaphrus scymnoides Casey, 1886:250) Mexico, USA (CA) Mexico, USA (AZ, CA, TX) politus Casey, 1889:162 mirus Wooldridge, 1984a:538 USA (TX) Ecuador punctatus Wooldridge, 1981a:220 punctatus Wooldridge, 1976:180 Colombia Colombia schwarzi Schaeffer, 1904:204 Jamaica, USA (TX) 160 Volume 15, No.3, September, 2001, INSECTA MUNDI simplex Wooldridge, 1981a:220 bus dam additis. Berliner Entomologische Colombia Zeitschrift 14: 49-90. spangleri Wooldridge, 1981a:218 Blackburn, T. 1896. Coleoptera (exclusive of the Ecuador Carabidae). pp. 254-308 in B. Spencer (ed.). Report of the work of the Horn Scientific Expe­ TRICHOLIMNICHUS Hernando and Ribera, dition to Central Australia. Part II. Zoology. 2001:153 Dalau and Company. London. maior Hernando and Ribera, 2001:159 Blackwelder, R.E. 1944. Checklist of the Co­ Malaysia leopterous Insects of Mexico, Central America, minor Hernando and Ribera, 2001:160 the West Indies, and South America. United Malaysia States National Museum Bulletin 185, Part 2: sabahensis Hernando and Ribera, 2001:160 189-341. Malaysia Brethes, J. 1922. Descripci6n de Varios Coleopteros de Buenos Aires Annales de la Sociedad de Checklist of the Lutrochidae of the World Ciencias de Argentina 94: 263-305. Britton, E.B. 1971. A new intertidal beetle (Co­ LUTROCHUS Erichson, 1847:509 leoptera: Limnichidae) from the Great Barrier BYRROMORPHUS Sharp, 1882:125 Reef. Journal of Entomology (B)40(2): 89-91. acuminatus Grouvelle, 1889:163 Britton, E.B. 1973. Hyphalus wisei sp. n., a new Venezuela intertidal beetle from New Zealand (Coleoptera: arizonicus Brown and Murvosh, 1970:1031 Limnichidae). Records of the Auckland Insti­ USA (AZ) tute and Museum 10: 119-122, 6 figures. geniculatus Chevrolat, 1864:406 Britton, E.B. 1977. Three new intertidal species of Cuba, Haiti Limnichidae (Coleoptera) from New Zealand. germari Grouvelle, 1889:163 Records of the Auckland Institute and Museum Brazil 14: 81-85, 9 figures. gigas Hinton, 1939:23 Broun, T. 1880. Manual of the New Zealand Co­ Peru leoptera. Part 1. pp. 1-651. Government Print­ laticeps Casey, 1893:580 er. Wellington. USA (AL, AR, DC, IL, IN, MD, MI, MO, OH, Broun, T. 1881. Manual of the New Zealand Co­ MS, OK, PA, TN, WI, WV) leoptera. Part 2. pp. 653-744. James Hughes. luteus LeConte, 1852:42 Wellington. USA (NM, OK, TX) Broun, T. 1893. Manual of the New Zealand Co­ misellus Grouvelle, 1896: 17 leoptera. Part 7. pp. 1395-1504. Government Brazil Printing Office. Wellington. montanus Grouvelle, 1896:16 Brown, H. P. 1965. Microhabitats of some Neotro­ Bolivia pical riffle beetles (Coleoptera: Dryopoidea). pilula Erichson, 1847:510 American Zoologist 5(2): 151. Brazil Brown, H.P., and C. M. Murvosh. 1970. Lutro­ vestitus Sharp, 1882:126 chus arizonicus New Species, with notes on Guatemala ecology and behavior (Coleoptera, Dryopoidea, Limnichidae). Annals of the Entomological So­ Acknow ledgments ciety of America 63(4): 1030-1035. Casey, T.L. 1886. Descriptive notices of North D. P. Wooldridge, Pennsylvania State Univer­ American Coleoptera I. Bulletin of the Califor­ sity, Ogontz, and W. E. Steiner, Smithsonian Insti­ nia Academy of Sciences 2(6): 157-264. tution, reviewed earlier drafts of this manuscript. Casey, T.L. 1889. Coleopterological notices I. With an appendix on the Termitophilous Staphylin­ Literature Cited idae of Panama. Annals of the New York Acad­ emy of Sciences 5: 39-198. Baudi de Selve, F. 1870. Coleopterorum messis in Casey, T.L. 1893. Coleopterological notices V. insula Cypro et Asia minore ab Eugenio Trugui Annals of the New York Academy of Sciences 7: congregatae recensitio: de Europaeis notis qui- 281-606. INSECTA MUNDI, Vol. 15, No.3, September, 2001 161

Casey, T.L. 1912. Descriptive catalogue of the Duftschmid, C.E. 1825. Fauna Austriae. Oder American Byrrhidae. Memoirs on the Coleoptera Beschreibung Osterreichischen Insecten, fur III: 1-69. Angehende Freunde der Entomologie 3: 1-289. Champion, G.C. 1923. Some Indian Coleoptera Linz. (12). Entomologist's Monthly Magazine 59: 219­ Erichson, W.F. 1842. Bericht tiber die Leistungen 224,247-253,269-277. in der Naturgeschichte die Insecten, Arach­ Champion, G.C. 1924a. On a new subfamily of niden, Crustaceen und Entomostraceen Clavicorn Coleoptera. Entomologist's Monthly wahrend des Jahres 1841. Archiv fur Naturge­ Magazine 60(717): 25-29. schichte 8: 83-287. Champion, G.C. 1924b. Thaumastodus fusiform­ Erichson, W.F. 1847. Naturgeschicte der Insecten is Champ.: Synonymical note. Entomologist's Deutschlands. Abteilung 1. Coleoptera 3(4): Monthly Magazine 60(720): 116-117. 481-640. Berlin. Champion, G.C. 1925a. Some Indian (and Tibet­ Fairmaire, L. 1888. Descriptions de Coleopteres an) Coleoptera (17). Entomologist's Monthly de l'Indo-Chine. Annales de la Societe Ento­ Magazine 61: 169-181. mologique de France (6)8: 333-378. Champion, G.C. 1925b. Some Indian (and Tibet­ Fairmaire, L. 1897. Materiaux pour la faune an) Coleoptera (18). Entomologist's Monthly Coleopterique de la region Malgache. Annales Magazine 61: 260-264. de la Societe Entomologique de Belgique 41: Chevrolat, L.A.A. 1864. Coleopteres de I'lle de 367-406. Cuba (suite). Notes, synonymies et descriptions Fairmaire, L. 1898. Materiaux pour la faune d'especes nouvelles. Ci nqu ieme memoir e. Coleopterique de la region Malgache. Annales Famille des parnides, heterocerides, passalides de la Societe Entomologique de Belgique 42: et lamellicornes (Tribus des coprides, aphodi­ 222-260. ides, hybosorides, geotrupides et trogides). Fauvel, C.A.A. 1903. Faune analytique des Annales de la Societe Entomologique de France Coleopteres de la Nouvell-Caledonie. Revue (4)4: 405-418. d'Entomologie 22: 203-378. Crowson, R.A. 1978. Problems of phylogenetic Ferrari, J.A. 1864. Kleiner Beitrag zur Kaferfau­ relationships in Dryopoidea (Coleoptera). En­ na Venedigs und des Lido, mitgetheilt von tomologica Germanica 4(3/4): 250-257. einem suddeutschen Entomologen. Wiener von Dalla Torre, K. W. 1911. Coleopterorum Entomologische Monatschrift 8: 108. Catalogus, pars 33: Nosodendridae, Byrrhidae, Fleischer, A. 1909. Ein neuer Limnichus. Wiener Dermestidae. W. Junk. Berlin. 96 pp. Entomologische Zeitung 28: 244. Darlington, P.J. 1936. A list of the West Indian Grouvelle, A.H. 1889. Voyage de M. E. Simon au Dryopidae (Coleoptera), with a new genus and Venezuela (Decernbre 1887 - Avril 1888). eight new species, including one from Colom­ Coleopteres, 2e memoire. Cucujidae, Rhysodid­ bia. Psyche 43: 65-82. iae, Dryopidae, Cyathoceridae, Heteroceridae. Deleve, J. 1968. Contribution a l'etude des «Dry­ Annales de la Societe Entomologique de France opoidea»>, Les Limnichinae d'Afrique (Co­ (6)9: 157-166. leoptera Limnichidae). Bulletin and Annales de Grouvelle, A.H. 1896. Description de deux dry­ la Societe Royale d'Entomologie de Belgique opides du genre Lutrochus Er. avec tableau 104: 212-274. general des especes (Col.). Bulletin de la Societe Deleve, J. 1971. Contribution a l'etude des Dry­ Entomologique de France 1896: 16-17. opoidea 23: De quelques Ptilodactyidae et Lim­ Guillebeau, F. 1886. Notes entomologiques sur la nichidae asiatiques de la collection Pic. Bulletin faune coleopterologique de Sinaia (Valachie) et de la Societe Entomologique de Belgique 107: de la Dobroudja (Roumanie). L'Echange 2(19): 352-376. 1-2. Deleve, J. 1973. Limnichidae, Dryopidae et Elmin­ Heller, K.M. 1921. Philippinische Melandryidae. thidae des Iles Philippines et de l'Archipel Bis­ Entomologische Blatter 17(10-12): 155-157. marck (Insecta, Coleoptera, Dryopoidea). Hernando, C. and I. Ribera. 2000a. Taxonomic [Noon a Dan Papers No. 122]. Steenstrupia 3(3): revision of the Afrotropical genus Cyclolimni­ 17-30. chus Deleve (Coleoptera: Limnichidae). Afri­ Deleve, J. 1974. Coleoptera: Limnichidae. South can Entomology 8: 211-216. African Animal Life 25: 40-41. 162 Volume 15, No.3, September, 2001, INSECTA MUNDI

Hernando, C. and I. Ribera. 2000b. The first 80th birthday of Roy A. Crowson. Museum i species of the intertidal genus Hyphalus Brit­ Instytut Zoologii PAN. Warszawa. ton from the Indian Ocean (Coleoptera: Lim­ Lea, A. M.1920. On Australian Coleoptera. Family nichidae: Hyphalinae). Annales de la Societe Byrrhidae. Part II. Records of the South Aus­ Entomologique de France (n.s.) 36(3): 239-243. tralian Museum 1: 273-290. Hernando, C. and I. Ribera. 2000c. Notes on LeConte, J.L. 1852. Synopsis of the Parnidae of Limnichidae (Coleoptera): Crytolimnichus the United States. Proceedings of the Academy punctulatus Deleve, new junior synonym of of Natural Sciences of Philadelphia 6: 41-45. Simplocarina curticollis Pic. Coleopterists Bul­ LeConte, J.L. 1854. Synopsis of the Byrrhidae of letin 54: 291. the United States. Proceeding of the Academy Hernando, C. and I. Ribera. 2001. Tricholimni­ of Natural Sciences of Philadelphia 7: 113-117. chus gen. n. and three new species from Borneo LeConte, J.L. 1874. Descriptions ofnew Coleoptera (Coleoptera: Limnichidae). Koleopterologische chiefly from the Pacific Slope of North America. Rundschau 71: 153-161. Transactions of the American Entomological Hilsenhoff, W.L. and KL. Schmude. 1992. Riffle Society 5: 43-72. beetles of Wisconsin (Coleoptera: Dryopidae, LeConte, J.L. 1879. The Coleoptera of the alpine Elmidae, Lutrochidae, Psephenidae) with notes Rocky Mountain Region. Part II. Bulletin ofthe on distribution, habitat, and identification. U.S. Geological and Geographical Survey of the Great Lakes Entomologist 25: 191-213. Territories 5(3): 499-520. Hinton, H.E. 1939. On some new genera and Lewis, G. 1879. A catalogue of Coleoptera from the species of Neotropical Dryopoidea (Coleoptera). Japanese Archipelago. Taylor and Francis. Transactions of the Entomological Society of London. 31 pp. London 89: 23-46. Mascagni, A. 1995. Nuovi dati alcuni coleotteri Jacquelin, P.N.C. du Val. 1857. Manuel ento­ Italiani (Limnichidae, Dryopidae, Elmidae et mologique. Genera des Coleopteres d'Europe, Georissidae). Bollettino della Societa Entomo­ par Jacquelin du Val (Camille) [et L. Fair­ logica Italiana 127: 122-126. maire]. Et plus de treizecents types par Jules Motschulsky, V. 1858. Insectes des Indes Orien­ Migneaux [et Theophile Deyrolle]. Deyrolle. tales. Etudes Entomologiques 7: 20-122. Paris. Volume II. 285 pp. Motschulsky, V. 1861. Essai d'un Catalogue des Kasap, H. and R. A. Crowson. 1975. A compara­ Insectes de d'Ile Ceylon, Lief. Bulletin de la tive anatomical study of Elateriformia and Societe Imperiale des Naturalistes de Moscou Dascilloidea (Coleoptera). Transactions of the 34(2): 95-155. Royal Entomological Society, London 126(4): Mulsant, E. and C. Rey. 1852. Description d'un 441-495, 91 figures. genre noveau de la famille des Byrrhiens. An­ von Kiesenwetter, M.H. 1851. Enumeration des nales de la Societe Linneenne de Lyon (n.s.) 1: Coleopteres trouves dans le midi de la France et 19-21. en Cataloge. Annales de la Societe Ento­ Mulsant, E. and C. Rey. 1869. Histoire naturelle mologique de France (2)9: 577-656. des Coleopteres de France. Piluliformes. An­ Kraatz, G. 1858. Beitrag zue Kaferfauna Griech­ nales de la Societe Linneenne de Lyon 17: 201­ enlands. Berliner Entomologische Zeitschrift 378. 2: 123-148. Nakane, T. 1963. [untitled]. Fragmenta Coleopter­ Latreille, P.A. 1829. Les Crustacea, les Arach­ ologica 11: 43-46. nides et les Insectes, distribues en familles Obenberger, J. 1916. II. Beitrag zur Kenntnis der naturelles, ouvrage formant les tomes 4 et 5 de palaearktischen Kaferfauna. Archiv fur Celui de M. le Baron Cuvier sur le Regne Naturgeschite 82A: 9-45. Animal (Deuxieme Edition). Vol. 1. Paris. 584 Paulus, H.F. 1970. Zur Morphologie und Biologie pp. der Larven von Pelochares und Limnichus. Lawrence, J.L. and A.F. Newton. 1995. Fami­ Senckenbergiana Biologica 51: 77-87. lies and subfamilies of Coleoptera (with select­ Perris, E. 1864. Description de quelques especes ed genera, notes, references and data on family nouvelles de Coleopteres et notes diverses. group names. pp. 779-1006 in J. Pakaluk and A. Annales de la Societe Entomologique de France Slipinski (eds.). Biology, phylogeny and classi­ (4)4: 275-310. fication of Coleoptera. Papers celebrating the INSECTA MUNDI, Vol. 15, No.3, September, 2001 163

Pic, M. 1905. Especes et varietes nouvelles de Pic, M. 1956. Nouveaux Coleopteres de diverses Coleopteres palearctiques. L'Echange 21: 97­ familIes. Annales Historico-Naturales Musei 100. Nationalis Hungarici 7: 71-92. Pic, M. 1922. Nouveautes Diverses. Melanges Ex­ Piitz, A. 1991a. Jaechobyrrhinus - eine neue Gat­ otico-Entomologiques 36: 1-32. tung der Familie Limnichidae aus der Turkei Pic, M. 1923a. Nouveautes Diverses. Melanges (Col., Limnichidae). Entomologische Nachrich­ Exotico-Entomologiques 39: 1-32. ten und Berichte 35(2): 132-134. Pic, M. 1923b. Nouveautes Diverses. Melanges Piitz, A. 1991b. Beitriige zur Insektenfauna der Exotico-Entomologiques 40: 1-32. DDR: Coleoptera- Limnichidae. Beitrage Ent. Pic, M. 1923c. Coleopteres exotiques en partie Berlin 41: 375-381. nouveaux. L'Echange 39: 15-16. Piitz, A. 1998a. I. Check list and bibliography of Pic, M. 1927a. Fauna Sumatrensis. Malacoderma­ the Limnichide of China and neighbouring coun­ ta (Col.). Supplementa Entomologica, Berlin tries (Coleoptera). pp. 337-339 in M. A. Jiich 15: 62-65. and L. Ji (eds.). Water beetles of China. Vol. 2. Pic, M. 1927b. Fauna Sumatrensis. Malacoderma­ Wien: Zoologisch-Botanische Gesellschaft in ta (Col.). Supplementa Entomologica, Berlin Osterreich and Wiener Coleopterologenverein. 16: 45-49. Piitz, A. 1998b. II. Taxonomic revision ofthe genus Pic, M. 1928. Nouveautes Diverses. Melanges Ex­ Cephalobyrrhus Pic (Coleoptera). pp. 341-371 otico-Entomologiques 52: 1-32. in M. A. Jiich and L. Ji (eds.). Water beetles of Pic, M. 1930. Nouveaux Coleopteres de diverses China. Vol. 2. Wien: Zoologisch-Botanische familles. Revista de la Sociedad Entomol6gica Gesellschaft in Osterreich and Wiener Co­ de Argentina 3: 43-46. leopterologenverein. Pic, M. 1935a. Nouveautes Diverses. Melanges Rey, C. 1889. Remarques en passant. L'Echange 5: Exotico-Entomologiques 66: 1-36. 66-67. Pic, M. 1935b. Nouveaux Coleopteres exotiques. Reitter, E. 1877[1878]. in O. Schneider and H. Annals and Magazine of Natural History (10)6: Leder. Beitriige zur Kenntniss der kaukasis­ 470-473. chen Kaferfauna, Verhandlungen des Natur­ Pic, M. 1938. Nouveaux Coleopteres de la Repub­ forschenden Vereines in Brunn 16: 1-258. lique Argentine. Revista de la Sociedad En­ Reitter, E. 1881. Bestimmungs-Tabellen der eu­ tom6logica Argentina 10: 77-79. ropaischen Coleopteren. IV. Enthaltend die Pic, M. 1939. Nouveaux Coleopteres de la Repub­ Familien: Cistelidae, Georyssidae und 'I'horic­ lique Argentine. Revista de la Sociedad En­ tidae. Verhandlungen der kaiserlich-konigli­ tom6logica Argentina 10: 249-251. chen zoologisch-botanischen Gesellschaft in Pic, M. 1952. Coleopteres nouveaux du Congo Wien 31: 67-96. Belge. Annales du Musee Congo belge, 'I'er­ Reitter, E. 1901.Weitere Beitriige zue Coleopter­ vuren, in-S", Sciences Zoologiques 12: 9 en-Fauna des russichen Reiches. Deutsche Pic, M. 1953. Coleopteres nouveaux de l'Mrique Entomologische Zeitschrift 1901: 65-84. orientale. Atti Museo Civico di Storia Naturale Ribera, I. and C. Hernando. 1999. Taxonomic di Trieste 19(2): 110-114. revision of Platypelochares Champion (Co­ Pic, M. 1954. Ptinidae, Anobiidae, Dermestidae, leoptera: Limnichidae). Koleopterologisch Byrrhidae (Col.). Beitrag zur Fauna Perus, Rundschau 69: 103-110. Jena 4: 176-178. Sato, M. 1965. The limnichid-beetles of Formosa. Pic, M. 1955. Contributions a l'etude de la fauna Special Bulletin of the Lepidopterists Society of entomologique du Ruanda-Urundi (Mission P. Japan 1965: 121-125. Basilewsky 1953). XII Coleoptera Rhip i­ Sato, M. 1966. The Limnichid-Beetles of Japan. phoridae, Oedemeridae, Scaptiidae, Pedilidae, Transactions of the Shikoku Entomological Aderidae, Anthicidae, Dermestidae, Byrrhidae, Society 9(2): 55-62. Anobiidae, Ptinidae, Cleridae, Lymexylonidae Sato, M. 1976. Family Limnichidae. Check-list of et Helodidae. Annales du Musee Congo Belge, the Coleoptera of Japan. 7: 1-2. Tervuren, in-S", Sciences Zoologiques 63: 121­ Sato, M. 1994. Note on the genus Pseudeucinetus 135. [sic] Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa En­ tomological Society, Niigata 2: 173-177. 164 Volume 15, No.3, September, 2001, INSECTA MUNDI

Sato, M. 1997. Occurrence of a new Hyphalus Wollaston, T.V. 1868. Coleoptera Hesperidum: (Coleoptera: Limnichidae) from the Ryukyu being an enumeration of the coleopterous in­ Archipelago, Japan. Elytra, Tokyo 25(1): 109­ sects ofthe Cape Verde. J. Van Voorst. London. 112. 285 pp. Schaeffer, C.F.A. 1904. New genera and species of Wooldridge, D. P. 1975. A key to the New World Coleoptera. Journal of the New York Entomo­ genera of the beetle family Limnichidae. Ento­ logical Society 12: 197-236. mological News 86(1-2): 1-4. Sharp, D. 1882. Biologia Centrali-Americana, In­ Wooldridge, D.P. 1976. New World Limnichinae secta, Coleoptera, Haliplidae, Dytiscidae, Gy­ I: A revision of the genus Physemus LeConte rinidae, Hydrophilidae, Heteroceridae, (Coleoptera: Limnichidae). Coleopterists Bul­ Parnidae, Georissidae, Cyanthoceridae. Vol­ letin 1976; 30(2): 177-182. ume 1, Part 2. 144 pp. Wooldridge, D.P. 1977a. New World Limnichinae Sharp, D. 1902a. Insecta, Coleoptera, Cryptoph­ III: A revisionofLimnichites Casey (Coleoptera: agidae, Lathriidae, Mycetophagidae, Dermes­ Limnichidae). Great Lakes Entomologist 10(4): tidae, Byrrhidae. Biologia Centrali-Americana 179-189. 2(1): 625-688. Wooldridge, D.P. 1977b. New World Limnichinae Sharp, D. 1902b. Descriptions of Oriental Limni­ II: Cephalobyrrhinus Pic (Coleoptera: Lim­ chini (Coleoptera, Fam. Byrrhidae). Entomolo­ nichidae). Entomological News 88(1 and 2): 29­ gist's Monthly Magazine (2)13: 61-64. 32. Spangler, P.J. 1995. A review and two new species Wooldridge, D.P. 1978. New World Limnichinae ofthe genus Pseudeucinetus Heller from south­ IV. Eulimnichus Casey. A. Synonymies, lecto­ east Asia and a world checklist of the Thaumas­ type designations, and redescriptions (Co­ todinae (Coleoptera: Limnichidae). Special Bul­ leoptera: Limnichidae). Great Lakes Entomol­ letin of the Japanese Society of Coleopterology ogist 11(3): 163-173. 4: 395-405. Wooldridge, D.P. 1979. New World Limnichinae Spangler, P. J. 1999 [1998]. A new tribe, genus, IV. Eulimnichus Casey. B. Descriptions of new and species of limnichid beetles, Wooldridgeus species (Coleoptera: Limnichidae). Great Lakes perforatus, from Malaysia (Coleoptera: Lim­ Entomologist 12(1): I-II. nichidae: Wooldridgeini). Insecta Mundi Wooldridge, D.P. 1980. New World Limnichinae 12(3and4): 181-187, 32 figures. V. Corrinea, a new Neotropical genus (Lim­ Spilman, T. 1959. A study ofthe Thaumastodinae, nichidae). Coleopterists Bulletin 34(1): 69-78. with one new genus and two new species (Lim­ Wooldridge, D.P. 1981a. Three new species of nichidae). Coleopterists Bulletin 13: 111-122. Throscinus LeConte, with notes on other spe­ Spilman, T. 1966. A new species of Martinius from cies (Coleoptera: Limnichidae: Cephalobyrrhi­ Cuba (Coleoptera: Limnichidae). Coleopterists nae). Coleopterists Bulletin 35(2): 217-22I. Bulletin 20: 123-125. Wooldridge, D.P. 1981b. New World Limnichinae Spilman, T. 1972. A new genus and species of VI. A revision of Limnichoderus Casey (Co­ jumping shore beetle from Mexico (Coleoptera: leoptera: Dryopoidea: Limnichidae). Journal of Limnichidae). Pan-Pacific Entomologist 48(2): the Kansas Entomological Society 54(1): 171­ 108-115. 19I. Stephens, J.F. 1832. Illustrations of British ento­ Wooldridge, D.P. 1982. New World Limnichinae mology. Baldwin and Cradock, London. Vol­ VII: Phalacrichus Sharp-Arecharacterization ume 5. 240 pp. and eight new species (Coleoptera: Dryopoidea: Sturm, J. 1807. Deutschlands Insecten von Jacob Limnichidae). Coleopterists Bulletin 36(2): 381­ Sturm. Kafer. Deutschlands fauna in abildun­ 389. gen nach der natur mit beschreibungen von Wooldridge, D.P. 1983. New World Limnichinae Jacob Sturm. Nurnberg. Volume 2. 279 pp. VIII. Paralimnichus Deleve (Coleoptera: Dry­ Waltl, J. 1838. Verzeichniss der um Passau vork­ opoidea: Limnichidae). Journal of the Kansas ommenden seltenen Kafer nebst Beschreibung Entomological Society 56(2): 232-233. der neuen Arten. Isis Oken 1838(4): 263-273. Wooldridge, D.P. 1984a. Two new species of Phy­ Weise, J. 1877. Kurze Revision der Limnichus­ semusLeContefrom South America (Coleoptera: und Pelochares Arten. Deutsche Entomolo­ Dryopoidea: Limnichidae). Journal of the Kan­ gische Zeitschrift 21: 299-302. sas Entomological Society 57(3): 538-539. INSECTA MUNDI, Vol. 15, No.3, September, 2001 165

Wooldridge, D.P. 1984b. Eulimnichus expeditus, Wooldridge, D.P. 1988. Martinius temporalis, a a new speciesfrom Central America (Dropoidea: new species from Ecuador (Coleoptera: Lim­ Limnichidae: Limnichinae). Coleopterists Bul­ nichidae: Thaumastodinae). Journal ofthe New letin 38(1): 57-58. York Entomological Society 96(3): 314-315. Wooldridge, D.P. 1984c. Parathroscinus, a new Wooldridge, D.P. 1990. Three new Paraihrosci­ genus of beetles from Southeast Asia (Lim­ nus from Singapore (Coleoptera: Limnichidae). nichidae: Cephalobyrrhinae). Journal of the Raffles Bulletin of Zoology 38(2): 251-255. New York Entomological Society 92(2): 121­ Wooldridge, D.P. 1993a. Three new Limnichidae 124. from Cerro de la Neblina,Venezuela (Coleoptera: Wooldridge, D.P. 1986a. Four new Neotropical Dryopoidea). Coleopterists Bulletin 47(1): 35­ Cephalobyrrhinus Pic (Coleoptera: Dryopoidea: 37. Limnichidae). Journal of the New York Ento­ Wooldridge, D.P. 1993b. A new species of Byrrhi­ mological Society 94(4): 512-515. nus Motschulsky from the Philippine Islands Wooldridge, D.P. 1986b. A catalog of the Co­ and Borneo (Coleoptera: Limnichidae). Coleop­ leoptera of America North of Mexico. Family: terists Bulletin 47(4): 359-361. Limnichidae. Agricultural Research Service, Wooldridge, D. P. 1995. Nomenclatural changes United States Department of Agriculture, Ag­ in Pelochares Mulsan t of Sri Lanka (Coleoptera: riculture Handbook 529-48. 8 pp. Limnichidae: Limnichinae). Coleopterists Bul­ Wooldridge, D.P. 1987a. Two new Neotropical letin 49(4): 326. species of Limnichoderus Casey (Coleoptera: Wu, C.F. 1937. Catalogus insectorum sinenium. Dryopoidea: Limnichidae). Journal of the Kan­ Fan Memorial Institute of Biology. Volume III. sas Entomological Society 60(2): 330-331. 1312 pp. Wooldridge, D.P. 1987b. New World Limnichinae Zaitzev, P. 1910. Coleopterorum Catalogus, pars IX: A revision of Neotropical Byrrhinus Mots­ 17: Dryopidae, Cyathoceridae, Georyssidae, chulsky (Coleoptera: Dryopoidea: Limnichidae). Heteroceridae. W. Junk. Berlin. 68 pp. Coleopterists Bulletin 41(4): 303-314.