Biodiversity Journal, 2019, 10 (4): 499–516 https://doi.org/10.31396/Biodiv.Jour.2019.10.4.499.516 MONOGRAPH

Notes on some interesting species of of the Monterosato collection from the Museum of Paleontology (Catania University, Italy)

Agatino Reitano1*, Fabio Liberto2, Maria Stella Colomba3, Ignazio Sparacio4 & Rossana Sanfilippo5

1Natural History Museum, via degli Studi 9, 97013 Comiso (Ragusa), Italy; e-mail: [email protected] 2Via del Giubileo Magno 93 - 90015 Cefalù, Italy; e-mail: [email protected] 3Università di Urbino, DiSB, via Maggetti 22 (loc. Sasso), 61029 Urbino, Italy; e-mail: [email protected] 4Via Principe di Paternò 3, 90144 Palermo, Italy; e-mail: [email protected] 5Department of Biological, Geological and Environmental Sciences, University of Catania, Corso Italia 57, 95129 Catania, Italy; e-mail: [email protected] *Corresponding author

ABSTRACT In the year 2008, the then named Department of Geological Sciences of the University of Catania came into possession of malacological material belonging to Tommaso Di Maria, baron of Allery and marquis of Monterosato (Palermo, 1841–1927), an important malacolo- gist specialized in the systematics of continental and mostly marine and fossil molluscs from the Mediterranean Sea and north east Atlantic. The small collection also includes interesting lots of shells belonging to other Sicilian naturalists such as the abbot Giuseppe Brugnone (Caltanissetta, Italy) and Pietro Calcara (Palermo, Italy). In this malacological collection, some interesting taxa little known and difficult taxonomic interpretations have been found. The rediscovery of the Aghatina mandralisci Calcara, 1840 and Lachesis retifera Brugnone, 1880 probable lectotypes, and the of Helix schwerzenbachi Calcara, 1841 and Helix cupani Calcara, 1842 (syntypes) are discussed. Particularly, A. mandralisci is proposed as a synonym of Allopeas gracilis (Hutton, 1834), H. schwerzenbachi is proposed as a synonym of Punctum (Punctum) pygmaeum (Draparnaud, 1801), and H. cupani is proposed as a syn- onym of Xerotricha conspurcata (Draparnaud, 1801).

KEY WORDS Taxonomy; Mollusca; Gastropoda; Monterosato; malacological collection; Museum.

Received 12.07.2018; accepted 10.09.2018; published online 28.12.2019 Proceedings of the 4th International Congress on Biodiversity “Man, Natural Habitats and Euro-Mediterranean Biodiver- sity”, November 17th-19th, 2017 - Malta

INTRODUCTION mostly marine and fossil molluscs from the Mediter- ranean and north east Atlantic. He described a very Tommaso Di Maria, baron of Allery and marquis large number of taxa, still valid today. of Monterosato (Palermo June 27th, 1841 - ibidem, Much of the Monterosato specimens, since March lst, 1927), was one of the most influential 1942, are hold in the malacological collections of malacologists in the second half of the 19th century, the Museo Civico di Zoologia di Roma [= Town- specialized in the systematics of continental and ship Zoological Museum of Rome (ZMR)] . These 500 AGATINO REITANO ET ALII

collections have always been considered a great then placed inside cardboard boxes with plastic lids. amount of information because they contain not Each box contains: only the specimens of Monterosato himself, but the original vial still closed, for lots containing also acquired lots belonging to other contemporary uninspected shells; malacologists, such as Giovanni Battista Adami, the original vial, emptied of the shells, with its Nicola Tiberi, Giuseppe Antonio Brugnone, which cap, both inside a zipped plastic bag, plus the shells in turn included shells belonging to Luigi Benoit, in a plastic box; Ignazio Libassi and Pietro Calcara (Oliverio & the cm-sized shells, inside zipped plastic bags Tringali, 2001). and/or relative cardboard boxes; In recent years, this important collection has un- the original labels, inside plastic boxes when of dergone new studies and a complete list of new ma- compatible size, or inside zipped plastic bags; rine mollusc taxa introduced by Tommaso Allery a label bearing the words “Laura Ryolo Dona- Di Maria Marquis of Monterosato has been com- tion, Monterosato Collection, Sample n° (from 1 to piled (Oliverio & Tringali, 2001; Appolloni et al., 140)” and also the Latin name of the mollusc 2018). species, in square brackets when assigned by the re- In February 2008, Ms. Laura Ryolo heir of the viser and finally the locality; Marquis of Monterosato, donated library and a label printed on a strip of paper, folded into malacological material belonging to him, to the three parts, with the header “University of Catania then Department of Geological Sciences of the - Museum of Paleontology” bearing the inventory University of Catania. In the same year, the mala- number of the collections on its lateral side. In the cologist Stefano Palazzi cataloged the collection central part the complete inventory number of the (Table 1, Fig. 1) allocated in the then Section of Museum is reported, together with the progressive Oceanology and Paleoecology, now belonging to number of the Monterosato collection (as per the those of the today's Museum of Palaeontology of label mentioned above), the specific name (in the Department of Biological, Geological and En- square brackets when assigned by the reviser) fol- vironmental Sciences, University of Catania (Re- lowed by the number of specimens (in round brack- itano, 2016). ets). The age, the location, and finally the initials of In the present work, we will examine some the reviser and the year of the revision (S[tefano] species of this collection, which are poorly known P[alazzi] 2008) are also reported. and of particular taxonomic interest. In the present work are examined four lots we considered more interesting: 1136/13–33 Bul[imus] mandraliscae, 1173/13–33 H[elix] schwerzenbachi, MATERIAL AND METHODS 1178/13–33 H[elix] cupaniana, 1198/13-33 Lach- esis retifera (Calcara, 1840a, b, 1841, 1842, 1845) The Monterosato collection kept at the Museum (Figs. 2, 3). of Palaeontology of the University of Catania Low magnification photos were acquired with a (MPCU) consists of 140 lots: 136 Gastropoda (ma- Canon EOS700D digital camera equipped with rine, terrestrial, and freshwater recent and fossils Tamron SP 60mm f/2 Di II LD Macro, in order to species), 3 Bivalvia, and a mixed collection includ- document the general features of the shells. Speci- ing molluscs and other taxonomic groups. mens were also examined uncoated under a Tescan They arrived to the MPCU in a good state of Vega 2 LMU Scanning Electron Microscope in preservation, inside a cardboard box. The shells Low Vacuum modality to investigate tube micro- with one or more labels were contained mostly in- morphology. All images were acquired at the side variously shaped and sized glass vials and Department of Biological, Geological and Environ- partly inside rectangular handmade cardboard mental Sciences (University of Catania, Italy). The boxes. The vials were closed with a cork stopper or photographed specimens, although with some dirt cotton balls. Some of the most interesting shells of organic substances, have not undergone any were extracted from the original vial, to be pho- cleaning, in order to avoid their damage. tographed, and then placed back in a plastic box, to ACRONYMS. MPCU: Museum of Paleontol- avoid harm from Byne’s disease. The 140 lots were ogy, Catania University (Italy). Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 501

Table 1/1. Catalogue of the Monterosato malacological collection of the MPCU (the content of the square brackets refers to the Stefano Palazzi’s revision). 502 AGATINO REITANO ET ALII

Table 1/2. Catalogue of the Monterosato malacological collection of the MPCU (the content of the square brackets refers to the Stefano Palazzi’s revision). Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 503

Table 1/3. Catalogue of the Monterosato malacological collection of the MPCU (the content of the square brackets refers to the Stefano Palazzi’s revision). 504 AGATINO REITANO ET ALII

Table 1/4. Catalogue of the Monterosato malacological collection of the MPCU (the content of the square brackets refers to the Stefano Palazzi’s revision).

Figure 1. The Monterosato malacological collection of the Museum of Palaeontology, Catania University (Italy). Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 505

Figure 2. Caption on page 46 of the table attached to the work of Calcara (1845) where Bulimus mandralisci, Helix schwerzenbachii, and Helix cupani, found in the Monterosato malacological collection (MPCU), are drawn. 506 AGATINO REITANO ET ALII

Figure 3. Table attached to the work of Calcara (1845) where Bulimus mandralisci, Helix schwerzenbachii and Helix cupani, found in the Monterosato malacological collection (MPCU), are drawn. Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 507

RESULTS purtroppo, è andata distrutta durante la Spedizione dei Mille perché la casa del Testa era vicina al Aghatina mandralisci Calcara, 1840 (Figs. 4–7) Palazzo Reale e fu distrutta da un incendio. Si sal- varono solo alcuni rari pezzi che il Testa aveva LOCUS TYPICUS. Calcara, 1840a: “Conchiglia scambiato con il Brugnone [the passion for mala- unica ritrovata dall'ornatissimo signor Testa nei con- cology was nourished by friendship with the af- torni di Palermo ed esistente nella sua collezione” firmed malacologist Domenico Testa who boasted a [= surroundings of Palermo, Sicily, Italy]. rich collection considered superior to all the others at the time and which he made available to the abbot. DESCRIPTION. Calcara, 1840a: “Testa cilindracea A collection that, unfortunately, was destroyed dur- subfusiformis, subdiaphana flava apice obtuso, lon- ing the Expedition of the Thousand because the gitudinaliter tenuissime striata; anfractibus septem, Testa’s home was close to the Royal Palace and was convexo planis; sutura profunda divisis, apertura destroyed by fire. Only a few rare pieces were saved oblongo-ovata, labro simplici . Lunghezza 6 linee - that Testa had exchanged for the Brugnone]”. Larghezza 2 1/6 - Altezza dell’apertura 2 linee”. Also Benoit (1882: 76) reports that Brugnone Conical elongated slender whitish-opaque shell had the opportunity to take shells from the collec- 9.50 mm high and 2.79 mm wide, consisting of tion of Domenico Testa. The specimen we analyzed seven whorls; last whorl and aperture high respec- corresponds in size and morphological characters tively the 49% and the 31% of the total shell. The to the original description of B. mandralisci and protoconch is rounded with weak longitudinal could constitute its lectotype. This shell could then striae, on the first whorl; a regular sutural crenula- be passed from Testa to Brugnone and by the latter tion starts with the second protoconch whorl. The labeled with the name of B. mandraliscae. After teleoconch whorls are evenly rounded with a deep the death of Brugnone (1884) the collection of the suture, which is crenulated by minute papillae. The Nissen abbot was sold to the Marquis of Monteso- surface of the whorls is covered by fine and dense rato (Ryolo & Palazzi, 2008; Cimino, 2011). axial striae with the widest curvature in the median We attribute the shell of B. mandralisci from the region of each whorl. The last whorl is more flat- MPCU to the species Allopeas gracilis (Hutton, tened than the previous ones. The aperture is ob- 1834) (Subulinidae). The species (graceful awlsnail) long-oval; the columella is straight and folded to is widely distributed in subtropical and tropical areas close the umbilicus. of Asia, Africa, Australia, Polynesia, central-south- REMARkS. Calcara (1840a, b) refers to a single ern America, Caribbean islands and southeastern shell, on which he based the institution of this United States (Dundee, 1971; Capinera, 2017), also taxon, belonging to the collection of “signor introduced in the Peninsula Arabica (Neubert, 1998) Domenico Testa” of Palermo, and found by him and Iraq (Naser, 2010). Although this species has “nei contorni di Palermo [in the surroundings of been described from India, Neck (1976) and Auffen- Palermo]”. Calcara himself (1840b, in note 2) says berg & Stange (1988) suggest that its origin is South to have never found it in so many researches and America. for this reason he considers it “specie dubbiosa per It is probable that the shell found in the sur- la Sicilia [a doubtful species for Sicily]”. roundings of Palermo by Testa and described by Lancia (1853) considers the shell of Bulimus Calcara (1840a) was introduced to Sicily with ex- mandralisci to a “var. monstruosa” of Hypnophila otic plant species. It is more difficult to hypothesize cylindracea (Calcara, 1840). Based on this, Benoit whether this shell belonged to a living and accli- (1862: 227) excludes B. mandralisci from the Sicil- mated population, albeit small or localized. Nowa- ian species. days, little subulinids like A. gracilis have been As reported by Cimino (2011), in Abbot found again in Palermo, inside the Botanical Gar- Brugnone “la passione per la malacologia fu ali- den (Roberto Viviano, in verbis) but, also in this mentata dall'amicizia con l’affermato malacologo case, without evidence of living populations. Domenico Testa che vantava una ricca collezione Taking into account the above, we propose B. ritenuta allora superiore a tutte le altre e che met- mandralisci as a younger synonym of A. gracilis teva a disposizione dell’abate. Una collezione che, (Hutton, 1834). 508 AGATINO REITANO ET ALII

Lachesis retifera Brugnone, 1880 (Figs. 8–12) REMARkS. For a correct geological dating of this species, it should be considered that the presence of LOCUS TYPICUS. Contrada “Giannettello” (today Chlamys (Aequipecten) scabrella (Lamarck, 1819) Giannittello), near Caltanissetta, Sicily, Italy. (Bivalvia Pectinidae) mentioned by Brugnone among the fauna found in Contrada Giannittello, DESCRIPTION. Brugnone, 1880: “Conchiglia would indicate it belong to the MPMU3 area minuta, ovato-turrita, con apice ottuso e levigato, (Monegatti & Raffi, 2001). In the geological map lunga mm. 6¼, larga mm. 2½: anfratti 7, legger- of Caltanissetta (http://www.isprambiente.gov.it/ mente convessi, separati da suture poco profonde, Media/carg/631-Caltanissetta-Enna/Foglio.html) sottilmente reticolati: reticolo formato da linee near Contrada Giannittello (near Cozzo di Naro, poco elevate o costicine longitudinali e spirali, le SW of Caltanissetta) are outcropping at the bottom une e le altre uguali tra loro ed ai loro intervalli, the marly clays of Geracello (Upper Pliocene) and perlate nel loro incrociamento e formanti tante pic- towards the other the sands of Lannari (Upper cole areole incavate e quadrate come le maglie Pliocene-Lower Pleistocene?). Since the exact col- d'una rete; linne longitudinali rette, esistenti dalla lection point of the fossil is unknown, we attribute seconda metà del primo giro, 24 nell'ultimo, quasi its age, as a precaution, to the Upper Pliocene- mancanti nella coda; linee spirali o transverse pro- Lower Pleistocene. tratte dal secondo giro sino a tutta la coda, 13 nel- The taxonomic aspects relating to Chauvetia l'ultimo giro, 5 nel penultimo, 3 negli altri: apertura retifera have already been extensively discussed rotondata, coda cortissima, columella contorta; (Micali, 1998; Tringali, 2001). In the present work, labbro destro acuto, internamente munito di 5 the taxon Chauvetia retifera is considered applica- pieghe grossette, che cominciano sopra il canaletto ble only to the fossil species treated here. Therefore ma non arrivano in alto; labbro sinistro tenue, dis- we believe that the name Chauvetia elongata F. tinto e continuo col destro”. Nordsieck & Garcia-Talavera, 1979 is available and Shell buccinoid, 6.30 mm high, 2.6 mm wide, applicable for the living species, both in the Atlantic solid, with seven moderately convex whorls and a and Mediterranean (Micali, 1998; Gofas & Oliver, marked suture. Protoconch 750 µm in maximum 2010), unlike what is reported in MolluscaBase diameter. The nucleous is smooth and 400 µm in (www.molluscabase.org/aphia.php?p=taxdetails&i diameter; the first whorl is markedly sculptured by d=138890; last consultation 21/12/2019). As al- axial ribs, narrower than the spaces separating ready believed by Tringali (2001) the taxa described them, where spiral cords are obvious at the inter- by Monterosato (1889) are not available, having section with the ribs generating small tubercles. been published as synonyms for Lachesis retifera The teleoconch ornamentations are spiral cords (ICZN, 2019: Art. 11.6), and as varieties are not and axial ribs, both narrower than the interspaces. available (ICZN, 2019: Art. 45). At present Chau- Interspaces between the ribs and cords form a vetia retifera is not known for any other Plio-Pleis- rather uniform mesh, which form beaded tuber- tocene deposit in the Mediterranean area. cles. The first whorl consists of three spiral cords. The species differs in general from Chauvetia In the last whorl there are 13 spiral cords, of which elongata in having a smaller shell and less slender 4 from the penultimate whorl. The size of the profile (see also Table 2). cords is constant for almost the entire surface of the last whorl. Axial ribs on the last whorl 24 in Helix schwerzenbachii Calcara, 1841 (Figs. 13– number and prosocline. Last whorl and aperture 15) high respectively the 60% and 42% of the total height of the shell. Outer lip enlarged, showing LOCUS TYPICUS. Calcara, 1841: “Si rinviene co- traces of reconstruction due to trauma. Inside the munemente attaccata sopra le pietre della contrada lip there are five weak denticles, the adapical one di Bellolampo e come ancora nelle vicinanze delle most evident; the abapical tooth forms the outer sponde dell'Oreto [near Palermo]”. edge of the siphonal canal. White-yellowish color with some presumed traces of original coloring DESCRIPTION. Calcara, 1841: “Testa minuta, or- near the tubercles. biculata, tenui, pellucida, glabra, corneo fulva, late Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 509

Table 2. Morphological differences between Chauvetia retifera and C. elongata. umbilicata; anfractibus 3 convexis, ultimo magno he too compares it with “Helix pygmoea”. However, inflato; apertura rotundata patula, labro simplici, the description of Benoit (1859) for some characters acuto. Diametro 1/3 di linea [equivalent to about is different from that of Calcara. In particular, Benoit 0.8 mm]”. (1859) describes the last angled and dilated whorl, Shell discoidal in shape and very small, light the rounded lunar aperture, the smallest umbilicus horny brown and shiny, teleoconch consisting of and the presence of a lamella that originates between three convex whorls; external surface with dense the two edges of the peristome and continues inside and regular ribs, on the ventral side around the um- the coil. These characteristics lead us to believe that bilicus about 20 rows of very prominent, blunt spi- H. schwerzenbachiana Benoit, 1859 is based on ju- ral ribs crossed by axial ribs, followed by an area venile specimens of a different species from that of with only axial sculpture towards the outer edge of Calcara (1841), probably Lauria cylindracea (Da the body whorl; aperture rounded, margin thin (but Costa, 1778). broken), umbilicus wide. The two specimens preserved in the Mon- REMARkS. According to Calcara (1841) this terosato collection of the MPCU are juveniles since taxon differs from the similar “Helix” pygmaea their heights are respectively 1.03 mm (1173/13- Draparnaud, 1801 in having a shell one third 33a) and 1.027 mm (1173/13-33b). Both show apex smaller, for a smaller number of whorls, for the or- and peristome heavily damaged. The shape of the bicular shape and for having a wider umbilicus. shell and the ornamentation of these specimens lead The species was accepted by Aradas & Mag- us to classify them as Punctum (Punctum) pyg- giore (1841: 60) and Calcara (1845: 17) but most maeum (Draparnaud, 1801), a species with a wide of the later authors considered the taxon H. schw- Holarctic distribution, present all throughout Italy erzenbachii Calcara, 1841 doubtful and based on (Manganelli et al., 1995) and mainly linked to un- juvenile specimens: Philippi (1844: 218), Pfeiffer dergrowth environments of wooded areas. L. (1848: 218; 1853: 272; 1868: 492), kobelt These two specimens are probably syntypes of (1875: 10), Tryon (1887: 30), Taylor (1909: 160). the species described by Calcara because, in the Benoit (1859: 144–145, Pl. 4, f. 8) describes the label inserted in this lot, the abbot Brugnone re- species of Calcara sub H. schwerzenbachiana and ported the wording “avuta dall’autore [had by the 510 AGATINO REITANO ET ALII

author]”, moreover Brugnone bought shells directly From a taxonomic point of view, H. cupaniana from Calcara (Cimino, 2011), which subsequently Calcara, 1842 was prevalently considered a doubt- merged in the Monterosato collection (Ryolo & ful species, with a description based on juvenile Palazzi, 2008). specimens. Philippi (1844: 218) hypothesizes that In consideration of the above, Helix schwerzen- H. cupani may be based on juvenile specimens of bachii is proposed here as a synonym of Punctum Xerotricha apicina (Lamarck, 1822), as also shared (Punctum) pygmaeum (Draparnaud, 1801). by Pilsbry (1894: 254). Benoit (1859: 146, Pl. 4, f. 9) accepts the species of Calcara (1842) and pub- Helix cupani Calcara, 1842 lishes a description and drawing compatible with young specimens of Xerotricha Monterosato, 1892. LOCUSTYPICUS. Calcara, 1842: “Trovasi nelle However, figures 9 and 10 of Benoit’s Plate 4 sponde di fiume Oreto vicino Ponterrotto” [near (1859) are reversed (see kobelt, 1877: 285). Rein- Palermo, Sicily, Italy]. hard (1877: 285) and kobelt (1877: 285), on the basis of specimens from the Berlin Museum, place DESCRIPTION. Calcara, 1842: “testa parva, orbico- H. cupaniana in synonymy of Discus (Gonyodis- lato-depressa, corneo-fulva, inferne convexa late um- cus) rotundatus rotundatus (O.F. Müller, 1774). bilicata; anfractibus 2 a suturis impresis, superficie This synonymy was followed by later authors as puctulato-scabra, subpilosa, labro tenui, simplici. Di- Westerlund (1890: 3, 49), Pilsbry (1894: 46–47) ametro ½ linea [equivalent to about 1.15 mm]”. and Taylor (1909: 180). Shell dextral, very small, pale light brown, glo- The lot of the MPCU is made up of 43 rather bose; 2–2.1 convex whorls. The protoconch, about uniform specimens in terms of size and morpholog- 1.5 whorls, is regular spirally striated, sometimes, ical characteristics. These characters allow us to at- with faint ribs more evident near the sutura. On the ventral side, around the umbilicus, spiral striae are tribute these shells to the species Xerotricha weaker, and shorts ribs have a concentric shape. conspurcata (Draparnaud, 1801). The teleconch whorl is globose with irregular ribs. The original description of H. cupaniana Cal- The surface of the shell is covered by irregular pe- cara, 1842 shows the term “subpilosa”, and there- riostracal hairs. Aperture rounded, margin thin fore corresponds better to the specimens of X. slightly reflected towards the umbilicus. conspurcata of the MPCU, rather than to the spec- imens of D. rotundatus of the Berlin Museum. Fur- REMARkS. From a nomenclatural point of view, thermore, the locality “Orfanello near Oreto” Benoit (1859: 146) re-describes and depicts the written on the label of the MPCU specimens is very Calcara’s species by amending the name H. cupani close to the type locality of H. cupaniana: “banks to H. cupaniana and citing the original spelling in of the Oreto river near Ponterotto”. the text. Subsequently, Pfeiffer (1868: 431–432) Considering that, Brugnone purchased shells di- places H. cupaniana Benoit, 1859 in synonymy rectly from Calcara (Cimino, 2011) the specimens with H. cupani Calcara, 1842. However, the name of the MPCU, ex Monterosato collection, ex H. cupaniana attributed to Calcara (1842) contin- Brugnone collection can be considered syntypes. ued to be used by most of subsequent authors: In consideration of the above, Helix cupaniana Benoit (1875: 138; 1882: 40), Westerlund (1876: Calcara, 1842 is proposed here as a synonym of Xe- 39; 1890: 3, 49), Reinhard (1877: 285), kobelt rotricha conspurcata. (1877: 285), Pilsbry (1894: 47), Taylor (1909: Bourguignat (1878?) established the new 180), Ruhoff (1980: 227). Allerya for three species of small molluscs from Although the taxon H. cupani Calcara, 1842 was Sicily: A. monterosati n. sp., A. brocchi (Calcara, used by some authors of the 1800s, including 1842) and A. cupanii (Calcara, 1842). It is likely Aradas & Maggiore (1841: 60), Calcara (1845: 18) that the description of A. monterosati and A. broc- and Pfeiffer (1848: 424), the taxon H. cupaniana chi, as for H. cupaniana, was based on juvenile has remained in predominant use, attributed to the specimens of already described species. Thus, fur- original author Calcara (1842). Therefore, it can be ther studies are necessary for a correct taxonomic considered a justified amendment of Helix cupani interpretation of both A. monterosati and H. brocchi Calcara, 1842 (I.C.Z.N. Art. 33.2.3.1.). and of the genus Allerya. Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of paleontology (Catania University) 511

Figures 4–7. Aghatina mandralisci Calcara, 1840 from Monterosato malacological collection (MPCU). Figs. 4, 5: shell and original label. Figs. 6, 7: protoconch. 512 AGATINO REITANO ET ALII

Figures 8–12. Lachesis retifera (Brugnone, 1880) from Monterosato malacological collection (MPCU). Figs. 8, 9: shell and original label. Figs. 10–12: protoconch. Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 513

Figures 13–15. Helix schwerzenbachii Calcara, 1841 from Monterosato malacological collection (MPCU). Fig. 13: shells 1173/13–33a and 1173/13–33b and original label. Fig. 14: shell 1173/13–33a. Fig. 15: shell 1173/13–33b. 514 AGATINO REITANO ET ALII

Figures 16–21. Helix cupani Calcara, 1842 from Monterosato malacological collection (MPCU). Fig. 16: shell 1198/13– 33a. Fig. 17: shell 1198/13–33b. Fig. 18: shell 1198/13–33c. Fig. 19: shell 1198/13–33d. Fig. 20: shell 1198/13–33e; pro- toconch nucleus. Fig. 21: shell 1198/13–33e, detail of the periostracal hairs and microsculpture. Some interesting Mollusca Gastropoda of the Monterosato collection from the Museum of Paleontology (Catania University) 515

Figure 22. Helix cupani Calcara, 1842 from Monterosato malacological collection (MPCU), shells and original label.

REFERENCES Benoit L., 1882. Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia o continuazione alla illus- Appolloni M., Smriglio C., Amati B., Lugliè L., Nofroni trazione sistematica critica iconografica de' testacei I., Tringali L.P., Mariottini P. & Oliverio M., 2018. estramarini della Sicilia Ulteriore e delle isole cir- Catalogue of the primary types of marine molluscan costanti. D’Amico, Messina, VI+176 pp. taxa described by Tommaso Allery Di Maria, Mar- Bourguignat J.R., 1878. Monografia del nuovo génère si- quis of Monterosato, deposited in the Museo Civico ciliano Allerya. Atti dell'Accademia di Scienze, let- di Zoologia, Roma. Zootaxa, 4477(1): 1–138. https:// tere ed arti di Palermo, 4: 1–7 (1876). doi.org/10.11646/zootaxa.4477.1.1. Brugnone G., 1880. Le conchiglie plioceniche delle vic- Aradas A. & Maggiore L., 1840–1843. Catalogo ragion- inanze di Caltanisetta - Bullettino della Società Mala- ato delle conchiglie viventi e fossili di Sicilia. Atti Ac- cologica Italiana 6: 85-158, tav. 1. cademia Gioenia di Scienze Naturali di Catania, Ser. Calcara P., 1840a - Nuove specie di conchiglie. Effemeridi I, Mem. I, 15 (2), pp. 187–217 (1840); Mem. II, 15 scientifiche e letterarie per la Sicilia, 82: 35–37. (2), pp. 349–366 (1840); Mem. III, 16 (1), pp. 49-87 Calcara P., 1840b. Monografie dei generi Clausilia e Bu- (?1841); Mem. IV, 17 (1), pp. 53–106 (?1841); Mem. limo coll'aggiunta di alcune nuove specie di V, 17 (2), pp. 163- 205 (?1842); Mem. VI, 20, pp. conchiglie siciliane esistenti nella collezione della 101–142 (1844); Mem. VII, 20, pp. 343–360 (1844). Sig. Teresa Gargotta in Salinas, pp. 1–54. Auffenberg k. & Stange L.A., 1988. The Subulinidae of Calcara P., 1841. Monografie dei generi Spirorbis e Suc- Florida. Florida Department of Agriculture and Con- cinea seguite da alcune nuove specie di conchiglie si- sumer Services, Division of Plant Industry, Entomol- ciliane esistenti nella collezione della sig. Teresa ogy Circular 305. Benoit L., 1857-1862. Illustrazione sistematica critica Gargotta in Salinas. Giornale Letterario, 226: 1–11. iconografica de’ testacei estramarini della Sicilia Ul- Calcara P., 1842. Delle nuove specie di conchiglie ter- teriore e delle isole circostanti. Gaetano Nobile, restri. An unnumbered page in the appendix to Cal- Napoli, 248 pp., 11 pls. [Quaderno 1.o: pp. i–xvi, 1– cara P., Cenno topografico dei dintorni di Termini. 52, pls.1–2 (1857); Quaderno 2.o: pp. 53–116, pls. 3- Stamperia Roberti, Palermo, 31 pp. [+1]. 4 (1857); Quaderno 3.o: pp. 117–180, pls. 5–6 Calcara P., 1845. Esposizione dei molluschi terrestri e (1859); Quaderno 4o: pp. 181–248, pls. 7–8 (1862); fluvitiali dei dintorni di Palermo. Atti Accademia pls. 9, 11, 12 unknown publication date. Scienze, Lettere e Arti di Palermo, 1: 1–46 + 1 tav. Benoit L., 1875. Catalogo delle conchiglie terrestri e flu- Capinera J.L., 2017. Biology and food habits of the in- viatili della Sicilia e delle Isole circostanti. Bullettino vasive snail Allopeas gracile (Gastropoda: Subulin- della Società Malacologica italiana, 1: 129–163. idae). Florida Entomologist, 100, 116–123. 516 AGATINO REITANO ET ALII

Cimino V., 2011. Giuseppe Antonio Brugnone: Malacol- huius familiae generum et specierum hodie cogni- ogo di Caltanissetta, Ed. WWF di Caltanissetta - SBN tarum. Volumen primum. Brockhaus, Lipsiae, 484 pp. Pal0232284, 38. Pfeiffer, L. 1853. Monographia heliceorum viventium. Dundee D.S., 1971. Subulinids and veronicellids in the Sistens descriptiones systematicas et criticas omnium United States. The Biologist, 53: 128–132. huius familiae generum et specierum hodie cogni- Gofas S. & Oliver J.D., 2010. Las especies del género tarum. Volumen tertium. Brockhaus, Lipsiae, 711 pp. Chauvetia (Gastropoda, , ) Pfeiffer, L. 1868. Monographia heliceorum viventium. del área ibero-marroquí, con descripción de cuatro Sistens descriptiones systematicas et criticas omnium especies nuevas Iberus 28: 23–60. huius familiae generum et specierum hodie cogni- Manganelli G., Bodon M., Favilli L. & Giusti F., (1995). tarum. Volumen quintum. Brockhaus, Lipsia, 565 pp. Gastropoda Pulmonata. In: Minelli A., Ruffo S., & Philippi R.A., 1836–1844. Enumeratio molluscorum si- La Posta S. (Eds.), Checklist delle specie della fauna ciliae cum viventium tum in tellure tertiaria fossilium, italiana. Calderini, Bologna. 16: 60 pp. quae in itinere suo observavit. Berolini, XIV+267 pp., I.C.Z.N. International code of zoological nomenclature, 12 tav. (1836); Halis Saxonum, IV+303 pp., 16 tav. fourth edition. Available at: https://www.iczn.org/the- (1844). code/the-international-code-of-zoological-nomencla- Pilsbry H.A., 1894. Manual of Conchology; structural and ture/the-code-online/ Last access: 16.11.2019 systematic. With illustrations of the species. Second Se- kobelt W., 1875. Zur Fauna Italiens. Jahrbücher der ries: Pulmonata. 9 (Helicidæ, Vol. 7.): 1–366, Pl. 1–71. Deutschen Malakoologischen Gesellschaft, 2: 192– Reinhardt O., 1877. Bemerkungen über einige sicilianis- 213. che. Jahrbücher der Deutschen Malakozoologischen kobelt W., 1877. Bemerkungen über einige sicilianische Gesellschaft, 4: 277–295, Pl. 9. Helix-Arten. Jahrbücher der Deutschen Malakoolo- Reitano A., 2016. Le collezioni paleontologiche del gischen Gesellschaft, 4: 277–287. Museo di Scienze della Terra di Catania. Incontri, 14: Lancia F., 1853. Elogio di Pietro Calcara. Atti dell'Ac- 19–22. cademia di Scienze e Lettere di Palermo. Nuova Ruhoff F.A., 1980. Index to the Species of Mollusca In- serie, 2: 1–18. troduced from 1850 to 1870. Smithsonian Institution Monterosato T. A., 1889. Coquilles marines marocaines Press, Washington, 640 pp. Journal de Conchyliologie 7 (1): 20–40, (2): 112–121 Ryolo L. & Palazzi S., 2008. Tommaso Di Maria, March- Micali P., 1998. Note sulle specie di Chauvetia dell’At- ese di Monterosato. Documenti inediti sulla sua vita lantico nord-orientale. Bollettino Malacologico, 34: e sulla sua opera. DVD available through the Società 53–68. Italiana di Malacologia web site. Available from: Monegatti P. & Raffi S., 2001. Taxonomic diversity and http://www.societaitalianadimalacologia.it/index.php? stratigraphic distribution of Mediterranean Pliocene option=com_content&view=article&id=342&Itemid bivalves. Palaeogeography, Palaeoclimatology, =56 (Accessed 10 June 2019) Palaeoecology, 165: 171–193. Taylor J.W., 1909. Monograph of the land and freshwater Naser M.D., 2010. New record of the land snail Allopeas Mollusca of the British Isles. Part 16: 145–224, Pl. gracilis (Hutton,1834) (Gastropoda: Subulinidae) 16–22. from Basrah area, Iraq. Jordan Journal of Biological Tryon G.W., 1887. Manual of Conchology; structural and Sciences, 3: 55–56. systematic. Second Series: Pulmonata, 3. Helicidæ, Neck R.W., 1976. Adventive land snails in the I: 1–313, Pl. 1–63. Brownsville, Texas area. The Southwestern Naturalist Tringali L.P., 2001. Marine malacological records from 21: 133–135. Torres de Alcalá, Mediterranean Morocco, with the Neubert E. 1998. Annotated checklist of the terrestrial description of a new philinid species. Bollettino and freshwater mollusks of the Arabian Peninsula Malacologico, 37: 207–222. with descriptions of new species. Fauna of Arabia 17: Westerlund C.A., 1876–1878. Fauna Europæa mollusco- 333–461. rum extramarinorum. Prodromus. Sistens descrip- Nordsieck F. & García-Talavera F., 1979. Moluscos tiones systematicas et criticas omnium generum et Marinos de Canarias y Madera (Gastropoda). Aula specierum horum animalium in Europa viventium et de Cultura, Tenerife, 208 pp., 46 pls. hodie cognitarum. Lundae, Berlingia, 320 pp. Oliverio M. & Tringali L., 2001. The types of marine Westerlund C.A., 1890. Fauna der in der paläarctischen molluscan species described by Monterosato, in the Region (Europa, kaukasien, Sibirien, Turan, Persien, Museo Civico di Zoologia, Roma. General scope of kurdistan, Armenien, Mesopotamien, kleinasien, the work, and part 1: the opisthobranch gastropods. Syrien, Arabien, Egypten, Tripolis, Tunesien, Alge- Bollettino malacologico, 37: 121–142. rien und Marocco) lebenden Binnenconchylien. VII. Pfeiffer L., 1848. Monographia heliceorum viventium. Malacozoa acephala, pp. 1–319, 1–16, 1–15. Berlin. Sistens descriptiones systematicas et criticas omnium (Friedländer).