Flora of Glacier National Park, Montana
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0K17S .37 SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM CONTRIBUTIONS FROM THE United States National Herbarium Volume 22, Part 5 FLORA OF GLACIER NATIONAL PARK MONTANA By PAUL C. STANDLEY WASHINGTON GOVERNMENT PRINTING OFFICE 1931 SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM CONTRIBUTIONS FROM THE United States National Herbarium Volume 22, Part ^ FLORA OF GLACIER NATIONAL PARK MONTANA By PAUL C. STANDLEY iiian)rv9 "IVDIWVJ.f>3 WASHINGTON \ GOVERNMENT PRINTING OFFICE 1921 Si BULLETIN OF THE UNITED STATES NATIONAL MUSEUM. PREFACE. Among the many thousands of persons who visit the national parks each year a large nmnber are deeply interested in the plant life, including not only the wealth of conspicuous flowering plants but the trees of the forest also. All the national parks offer excep- tional opportunity for the study and enjoyment of primeval vege- tation, and in this respect none of them perhaps is better provided with attractive and refreshing material than Glacier National Park, Montana. The present paper has been written for the purpose of ena- bling visitors to this park to become more familiar with its wild plants and to derive greater enjoyment from them. This publication will be useful not only in Glacier Park but elsewhere in the mountains of Idaho, Alberta, and British Columbia. Most of the common plants of Yellowstone National Park occur also in Glacier Park. The present flora will be found helpful therefore to travelers in the Yellowstone region. Frederick V. Coville, Curator of the United States Natiojial Herbarium. CONTENTS. Introduction 235 General features of the flora 236 Life 7.ones 238 Transition Zone 239 Canadian Zone 241 Hudsonian Zone 245 Arctic-Alpine Zone 247 Scope and plan of the flora 249 Earlier botanical exploration 251 Acknowledgments 253 Bibliography 253 Systematic treatment . 255 Key to the families 255 Annotated catalogue of species . 263 Index IX ILLUSTRATIONS. PLATES. Following page. Plate 33. View south from the Garden Wall, showing the mountains of the west slope 438 34. View southeast from Iceberg Tiake 438 35. Swiftciurent Valley 438 36. Pond near Swiftcurrent Creek below Lake McDermott 438 37. View from a point near Granite Park; Heavens Peak at the right . 438 38. A. Limber pine {Pinus flexilis) on Altyn Peak. B. Dense forest ^ of hemlock and giant cedar near Lake McDonald 438 39. Mountain side below Sperry Chalets 438 40. A. Shore of Lake McDonald. B. Sphagnum bog at Johns Lake . 438 41. View on the trail to Piegan Pass 438 42. A. Dwarfed trees at Swiftcurrent Pass. B. Meadow near Granite Park 438 43. Ptarmigan Lake 438 * 44. A. Lake Ellen Wilson and Gunsight Pass. B. An Arctic-Alpine . rock slide 438 45. A. Fir clubmoss {Lyco-podiuvi selago). B. Stiff clubmoss (Li/ropo- dium annotinum) 438 46. A. Running-pine (Lycopodium, clavatum). B. Queencup {Clintonia unifiora) 438 47. A. Western yew {Taxus brevifolia). B. Ladies'-tresses (Ibidium romanzoffianum) 438 48. A. FuTTple clevasitis {Clematis Columbiana) . B. White dryad (Dryas octopetala) 438 49. A. Alpine bistort {Polygonum, viviparum). B. Spiny currant {Ribes lacustre) 438 50. A. Red raspberry {Rubus strigos^is). B. Blue phacelia {Phacelia lyallii) 438 51. A. Canada buffaloberry {Lepargyrea canadensis). B. Cow parsnip {Heracleum lanalum) . 438 52. A. Rough aster {Aster conspicuus). B. Showy fleabane (Erigeron salsuginosii^s) 438 VII wew Vs.. FLORA OF GLACIEE NATIONAL PARK, MONTANA. By Paxtl C. Standley. INTRODUCTION. Glacier National Park lies in northwestern Montana along tlie main range of the Rockies. It embraces an area of 1,534 square miles, nearly all of which consists of masses of high mountains. On the north it adjoins British Columbia and Alberta; eastward stretch the prairies of the Blackfoot Indian Reservation, and to the west lie the mountains and heavy forests of the Flathead Valley. The Continental Divide runs along the crest of the chief mountain ridge. The drainage of the west (Pacific) slope of the park is conse- quently into the Columbia River, while on the east (Atlantic) slope it is partly to Hudson Bay and partly to the Missouri River. From one point, Triple Divide Peak, the drainage is partly into each of the three systems. The highest peaks of the region reach an altitude of but little more than 3,000 meters (10,000 feet), but the elevation of the surrounding country is comparatively low (950 meters on the west slope and 1,440 meters on the east slope), so that the mountains are quite as imposing in appearance as many of those in the southern Rockies which have a much greater elevation. The rocks of the park are stratified and of Algonkian age. They consist chiefly of shale, limestone, sandstone, and argillite, their pre- vailing colors being rich reds and dull greens. In many places the stratification is very regular and certain strata can be traced with the _^eye for many miles, but in other places the strata are folded and con- torted in an interesting fashion. Besides the stratified rocks, a con- ^ spicuous feature is an intrusion of diorite—an igneous rock—which can be followed for distance along the Garden Wall as a well- J a long - marked band of black. ^ The whole region of the park is extremely rugged (see plate 33), the 9 mountains usually having sharp summits and precipitous sides. The main ridge is broken only infrequently by passes, whose altitudes range from 1,800 to about 2,100 meters. The slopes of the mountain masses have been plowed by ancient glaciers, and numerous lateral valleys have been cut, along which streams now run. In many of the valleys lie large or small lakes whose waters are wonderfully trans- parent and beautifully colored in blue or green. Fed directly by 235 236 CONTRIBUTIONS FROM THE NATIONAL, HERBARIUM. streams from the snow fields and present glaciers, the water of these lakes is ice cold. High up on the peaks, in glacier basins, are often found miniature jewel-like lakes of intense colors, freqviently sur- rounded by the ice walls of the glaciers themselves. Most spectac- ular of them is Iceberg Lake, on the east slope, which all summer long is full of huge blocks of floating ice which have broken off from the overhanging glacier. In some of these small alpine lakes the water is milky white from the particles of rock, finely ground by glaciers, held in suspension. One of the most striking features of the park is found in the glaciers which have given it its name. These lie in depressions at the head of some valley or hang high up on steep rock slopes. None of them are very large—Blackfoot Glacier is about three miles wide—but there are over 60 of them in the park. Each one possesses all or most of the features of the largest glaciers, and in a small glacier one can see the processes of glacial action much more readily than in a large one. Streams of water flow away from each of these ice masses, often falling abruptly over high cUffs, carrying down chunks of ice and rock which are heard constantly crashing upon the rocks below. The streams themselves, which abound everywhere in the park, are one of its many attractive features. To most people another source of interest is the profusion of animal life.^ Deer, elk, moose, mountain sheep and goats, bears, and many smaller animals are found in varying abundance. Sheep and goats are particularly plentiful and may be seen by the visitor in any of the higher regions, sometimes even in the vicinity of the hotels and chalets. The streams and lakes are well stocked with trout and other fishes. The winters are long in northern Montana. Snow falls by middle September or even earlier; indeed, at high altitudes, it frequently falls even in midsummer. At low elevations it remains until late in the season. In 1919, although the snowfall had been extremely light the previous winter, numerous snow banks remained at middle or even at low altitudes the first of July. Because of the short summers, the tourist season is limited; it extends from June 15 to September 15. GENERAL FEATURES OF THE FLORA. All the national parks of the West possess many attractions for anyone interested in plants, whether from the esthetic or from the scientific standpoint, and for the study of plants none offers greater advantages than Glacier Park. The flora is rich in number of species, and the vegetation is luxuriantly developed. All through * See Wild animals of Glacier Park: The mammals, by Vernon Bailey; The birds, by Florence Merriam Bailey. Department of the Interior, National Park Ser^dce. 11)18. STANDLEY FLORA OF GLACIER PARK. 237 the summer there is a lavish display of color, which attains its climax about the first of July. The flowers follow close upon the retreating snow banks, and thus even when they are somewhat faded at lower altitudes they may be seen in all their vernal freshness on the high slopes. The growing season is so brief that plants must bloom and fruit quickly. At high altitudes, and to some extent at middle ones, there are no "spring," "sumLmer," or "fall" flowers, for nearly all plants are in bloom at once, asters and goldenrod mingling with violets, springbeauties, and anemones. At low altitudes, however, there are marked differences between the spring and autumn floras. The flora of Glacier Park is in general typical of the Rocky Mountain region, but like that of any restricted area it shows certain special characteristics. The Continental Divide, which traverses the park, is to a certain extent a barrier to plant migra- tion, and there are noticeable differences between the plants of the east and west slopes.