Déterminisme Génétique De La Couleur Chez Les Espèces De Perruches Et De Perroquets Élevés En

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Déterminisme Génétique De La Couleur Chez Les Espèces De Perruches Et De Perroquets Élevés En ÉCOLE NATIONALE VÉTÉRINAIRE D’ALFORT Année 2011 DÉTERMINISME GÉNÉTIQUE DE LA COULEUR CHEZ LES ESPÈCES DE PERRUCHES ET DE PERROQUETS ÉLEVÉS EN CAPTIVITÉ THÈSE Pour le DOCTORAT VÉTÉRINAIRE Présentée et soutenue publiquement devant LA FACULTÉ DE MÉDECINE DE CRÉTEIL le…………… par Anna FÜRST Née le 4 Septembre 1984 à Chatenay-Malabry (Hauts-de-Seine) JURY Président : Pr. Professeur à la Faculté de Médecine de CRÉTEIL Membres Directeur : Monsieur COURREAU Jean-François, Professeur à l’École Nationale Vétérinaire d’Alfort Assesseur : Mademoiselle ABITBOL Marie, Maître de conférences à l’École Nationale Vétérinaire d’Alfort LISTE DES MEMBRES DU CORPS ENSEIGNANT Directeur : M. le Professeur MIALOT Jean-Paul Directeurs honoraires : MM. les Professeurs MORAILLON Robert, PARODI André-Laurent, PILET Charles, TOMA Bernard Professeurs honoraires: MM. et Mme : BRUGERE Henri, BRUGERE-PICOUX Jeanne, BUSSIERAS Jean, CERF Olivier, CLERC Bernard, CRESPEAU François, DEPUTTE Bertrand, LE BARS Henri, MOUTHON Gilbert, MILHAUD Guy, POUCHELON Jean- Louis, ROZIER Jacques DEPARTEMENT D’ELEVAGE ET DE PATHOLOGIE DES EQUIDES ET DES CARNIVORES (DEPEC) Chef du département : M. POLACK Bruno, Maître de conférences - Adjoint : M. BLOT Stéphane, Professeur - UNITE DE CARDIOLOGIE - UNITE DE PARASITOLOGIE ET MALADIES PARASITAIRES Mme CHETBOUL Valérie, Professeur M. CHERMETTE René, Professeur * Melle GKOUNI Vassiliki, Praticien hospitalier M. POLACK Bruno, Maître de conférences - UNITE DE CLINIQUE EQUINE M. GUILLOT Jacques, Professeur M. AUDIGIE Fabrice, Professeur* Mme MARIGNAC Geneviève, Maître de conférences M. DENOIX Jean-Marie, Professeur M. HUBERT Blaise, Praticien hospitalier Mme GIRAUDET Aude, Praticien hospitalier M. BLAGA Radu Gheorghe, Maître de conférences (rattaché au DPASP) Mme CHRISTMANN Undine, Maître de conférences - UNITE DE PATHOLOGIE CHIRURGICALE Mme MESPOULHES-RIVIERE Céline, Maître de conférences contractuel M. FAYOLLE Pascal, Professeur * Mme PRADIER Sophie, Maître de conférences M. MOISSONNIER Pierre, Professeur Melle DUPAYS Anne-Gaëlle, Assistant d’enseignement et de recherche M. MAILHAC Jean-Marie, Maître de conférences contractuel M. NIEBAUER Gert, Professeur contractuel - UNITE D’IMAGERIE MEDICALE Mme VIATEAU-DUVAL Véronique, Maître de conférences Mme BEDU-LEPERLIER Anne-Sophie, Maître de conférences contractuel Mme RAVARY-PLUMIOEN Bérangère, Maître de conférences (rattachée au Mme STAMBOULI Fouzia, Praticien hospitalier DPASP) - UNITE DE MEDECINE M. ZILBERSTEIN Luca, Maître de conférences M. BLOT Stéphane, Professeur* - UNITE DE REPRODUCTION ANIMALE M. ROSENBERG Charles, Maître de conférences M. FONTBONNE Alain, Maître de conférences* Mme MAUREY-GUENEC Christelle, Maître de conférences M. NUDELMANN Nicolas, Maître de conférences Mme BENCHEKROUN Ghita, Maître de conférences contractuel M. REMY Dominique, Maître de conférences (rattaché au DPASP) - UNITE DE MEDECINE DE L’ELEVAGE ET DU SPORT M. DESBOIS Christophe, Maître de conférences M. GRANDJEAN Dominique, Professeur * Mme CONSTANT Fabienne, Maître de conférences (rattachée au DPASP) Mme YAGUIYAN-COLLIARD Laurence, Maître de conférences contractuel Mme MASSE-MOREL Gaëlle, Maître de conférences contractuel (rattachée au - DISCIPLINE : NUTRITION-ALIMENTATION DPASP) M. PARAGON Bernard, Professeur M. MAUFFRE Vincent, Assistant d’enseignement et de recherche contractuel, (rattaché au DPASP) - DISCIPLINE : OPHTALMOLOGIE - DISCIPLINE : URGENCE SOINS INTENSIFS Mme CHAHORY Sabine, Maître de conférences Mme ROUX Françoise, Maître de conférences DEPARTEMENT DES PRODUCTIONS ANIMALES ET DE LA SANTE PUBLIQUE (DPASP) Chef du département : M. MILLEMANN Yves, Maître de conférences - Adjoint : Mme DUFOUR Barbara, Professeur - DISCIPLINE : BIOSTATISTIQUES - UNITE DE PATHOLOGIE MEDICALE DU BETAIL ET DES ANIMAUX DE M. DESQUILBET Loïc, Maître de conférences BASSE-COUR - UNITE D’HYGIENE ET INDUSTRIE DES ALIMENTS D’ORIGINE M. ADJOU Karim, Maître de conférences * ANIMALE M. MILLEMANN Yves, Maître de conférences M. BOLNOT François, Maître de conférences * M. BELBIS Guillaume, Assistant d’enseignement et de recherche contractuel, M. CARLIER Vincent, Professeur M. HESKIA Bernard, Professeur contractuel Mme COLMIN Catherine, Maître de conférences - UNITE DE ZOOTECHNIE, ECONOMIE RURALE M. AUGUSTIN Jean-Christophe, Maître de conférences Mme GRIMARD-BALLIF Bénédicte, Professeur* - UNITE DES MALADIES CONTAGIEUSES M. COURREAU Jean-François, Professeur M. BENET Jean-Jacques, Professeur* M. BOSSE Philippe, Professeur Mme HADDAD/HOANG-XUAN Nadia, Professeur Mme LEROY-BARASSIN Isabelle, Maître de conférences Mme DUFOUR Barbara, Professeur M. ARNE Pascal, Maître de conférences Melle PRAUD Anne, Assistant d’enseignement et de recherche contractuel, M. PONTER Andrew, Professeur DEPARTEMENT DES SCIENCES BIOLOGIQUES ET PHARMACEUTIQUES (DSBP) Chef du département : Mme COMBRISSON Hélène, Professeur - Adjoint : Mme LE PODER Sophie, Maître de conférences - UNITE D’ANATOMIE DES ANIMAUX DOMESTIQUES - UNITE DE PATHOLOGIE GENERALE MICROBIOLOGIE, M. CHATEAU Henry, Maître de conférences* IMMUNOLOGIE Mme CREVIER-DENOIX Nathalie, Professeur Mme QUINTIN-COLONNA Françoise, Professeur* M. DEGUEURCE Christophe, Professeur M. BOULOUIS Henri-Jean, Professeur Mme ROBERT Céline, Maître de conférences M. MAGNE Laurent, Maître de conférences contractuel - DISCIPLINE : ANGLAIS - UNITE DE PHARMACIE ET TOXICOLOGIE Mme CONAN Muriel, Professeur certifié M. TISSIER Renaud, Maître de conférences* - UNITE DE BIOCHIMIE Mme ENRIQUEZ Brigitte, Professeur M. MICHAUX Jean-Michel, Maître de conférences* M. PERROT Sébastien, Maître de conférences M. BELLIER Sylvain, Maître de conférences - UNITE DE PHYSIOLOGIE ET THERAPEUTIQUE - DISCIPLINE : EDUCATION PHYSIQUE ET SPORTIVE Mme COMBRISSON Hélène, Professeur* M. PHILIPS, Professeur certifié M. TIRET Laurent, Maître de conférences - UNITE DE GENETIQUE MEDICALE ET MOLECULAIRE Mme PILOT-STORCK Fanny, Maître de conférences Mme ABITBOL Marie, Maître de conférences* - UNITE DE VIROLOGIE M. PANTHIER Jean-Jacques, Professeur M. ELOIT Marc, Professeur * -UNITE D’HISTOLOGIE, ANATOMIE PATHOLOGIQUE Mme LE PODER Sophie, Maître de conférences M. FONTAINE Jean-Jacques, Professeur * - DISCIPLINE : ETHOLOGIE Mme LALOY Eve, Maître de conférences contractuel Mme GILBERT Caroline, Maître de conférences Mme CORDONNIER-LEFORT Nathalie, Maître de conférences M. REYES GOMEZ Edouard, Assistant d’enseignement et de recherche contractuel, * responsable d’unité REMERCIEMENTS Merci au Professeur de l’Université de Créteil d’avoir accepté de présider le Jury de cette thèse. Merci au Professeur Jean-François Courreau de m’avoir proposé ce sujet et d’avoir accepté d’être mon directeur de thèse. Merci également pour l’aide apportée dans la réalisation de ce travail. Merci au Docteur Marie Abitbol d’avoir accepté d’être mon assesseur et merci pour tous les précieux conseils de rédaction. Merci à la Commission Nationale des Juges de France section Psittacidés de m’avoir proposé un sujet aussi intéressant. J’espère sincèrement que ce travail sera utile et permettra, à terme, d’obtenir une unification des appellations de mutations. Merci à l’UOF, pour m’avoir permis d’utiliser certaines photographies issues de leur banque de données. Je remercie tout particulièrement M. Channoy pour toute l’aide dont j’ai pu bénéficier au cours de ces années de travail. Merci également pour les précieux documents dont j’ai pu disposer et qui m’ont été d’une aide très précieuse. Un grand merci à M. Fleurier pour ses remarques de spécialiste et de passionné, elles m’ont été bien utiles. Je le remercie aussi pour avoir pris du temps à rechercher dans la banque de photographies de l’UOF celles qui m’étaient utiles. Je remercie également M. de Lavergne pour ses suggestions et commentaires. Enfin, merci à M. Campagne et M. Dulière pour avoir répondu rapidement à ma demande et m’avoir autorisée à utiliser certaines de leurs photographies afin d’illustrer cette thèse. Merci à toute ma famille, pour m’avoir supportée et soutenue pendant toutes ces années d’étude. Merci également pour leur soutien durant la réalisation de cette thèse. Merci à Pierre pour son soutien et pour son expertise informatique. Merci à tous mes amis, alforiens ou non. TABLE DES MATIÈRES LISTE DES ABREVIATIONS ................................................................................................. 7! LISTE DES FIGURES ............................................................................................................... 9! LISTE DES TABLEAUX ........................................................................................................ 11! INTRODUCTION PRESENTATION DES PSITTACIFORMES ...................................... 13! PREMIERE PARTIE : DONNEES BIOLOGIQUES NECESSAIRES A L’ETUDE DES COULEURS CHEZ LES PSITTACIFORMES .......................................................... 17! I.! RAPPELS GENETIQUES DE BASE ......................................................................................... 17! 1.! Présentation du génome .............................................................................................. 17! a.! Constitution du chromosome ................................................................................... 17! b.! Caractéristiques des chromosomes .......................................................................... 18! 2.! Notion de gène et d’allèle ............................................................................................ 21! a.! Du chromosome au gène ........................................................................................
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