A Field Guide to Common Puget Sound Native Bees: Southern Region Introduction 3

Total Page:16

File Type:pdf, Size:1020Kb

A Field Guide to Common Puget Sound Native Bees: Southern Region Introduction 3 Contents A Field Guide to Common Puget Sound Native Bees: Southern Region Introduction 3 How to Use Guide 3 Making Observations 4 Classification 5 Recording Data 6 Parts of the Bee 8 Section I: Color Plates 10 Section II: Group Descriptions 22 Group Index 34 Data Sheet 36 Contributors 38 Acknowledgements 38 The Author 39 Author Elias H. Bloom, Washington State University [email protected] Level: Beginner 1 2 Introduction Making Observations This field guide is intended to be a tool When: for the identification and monitoring of The best time to make observations of floral visitors, in particular, native bees in bees is at temperatures greater than 70° the urban gardens and farms of the Puget Fahrenheit, and wind speeds less than 5 Sound Region. The relatively compact mph. Bees will be harder to observe in size of the guide was purposeful in that windy conditions, and may not visit flow- we hope you take this guide with you into ers when temperatures are too cool. the garden when you are working, or on Where: trips around the region. Before getting For the purpose of data collection, we ask started with the pictures and identifica- that you only make observations of bees tion, lets go over some basics about this or other flower visitors on flowers in a field guide, and the proper way to assess garden. You may also see bees on leaves bees. or on the ground. These bees may be rest- How to Use the Guide ing, or looking for a place to nest. Bees and other floral visitors are notori- How: ously difficult to identify in the field. De- Bees and other floral visitors are easily tails on how best to observe floral visitors disturbed. Avoid sudden movements, can be found at the front of the guide, loud noises, or casting your shadow over while pictures and habitat information are them. Observe bees first, then less mobile sandwiched in the middle. The final sec- flower visitors like beetles and bugs. Flies tion includes a data sheet and details for will often come back if disturbed, but oth- tracking your observations over time. er insects may not. Please, read each section carefully. 3 4 Classification Recording Data To simplify classification, we’ve created Data sheets are at the back of the guide. 5 groups of bees and 6 groups of floral We recommend photocopying the data visitors. Use these groups for your obser- sheets and replenishing these as you need vations and notes. Example images of more. Before starting your observations, each group can be found in Section I. On- take note of the date, time, and location. ly classify floral visitors to the level at Step 1: which you are most confident. Pick a spot in a garden where you can Groups of Bees: clearly see a plant and it’s flowers. Sit or Honey bees (HB) stand stationary near this plant and record any floral visitors you may see. If you ob- Bumble bees (BB) serve the same plant or plants multiple Other large bees (OLB) times over a year, or over many years, Green bees (GB) this will give us the best information Small bees (SB) about what types of floral visitors are im- Groups of Other Floral Visitors: portant for those plant groups. Butterflies (BTF) Step 2: Wasps (W) If the floral visitor is a bee, and you can Bugs (BG) confidently classify it, then note the bee Flies (F) group. Make sure to use the pictures, Beetles (BTL) habitat information, sizing guide, and Spiders (S) other details in Section II, to substantiate your observations. 5 6 Step 3: Parts of the Bee Insects are generally broken down into If possible, photograph the floral visitor three segments, the head, thorax, and ab- and plant. If your camera has a macro domen. The head is home to sensory ap- mode, use that setting. Otherwise, try to pendages including the eyes and anten- take pictures that are in focus. Bees will nae. The mouth of the insect is located on fly away if you get too close, so experi- the head too. Almost all insects are bilat- ment with the distance that is appropriate. erally symmetrical, meaning they have Every bee is a little different! When pho- the same appendages on both sides of the tographing the plant, take a picture of the body. Bees have two sets of wings, a flower, leaf, and where the leaf attaches to the stem. Pictures are not necessary, Abdomen but they help us to verify your observa- pygidial area tions. Step 4: Add additional notes about the site, plant, terga hind wing and floral visitor. Your notes are very ocelli helpful in the monitoring process and will compound eye allow you to build a more complete data antenna middle leg set. clypeus Step 5: fore wing Submit your observations, images, and hind leg notes about your garden to our website, and track your pollinators over time! 7 8 hind wing and fore wing, and three pairs Section I: Color Plates In this guide, each bee group represents of legs. The legs and wings are attached many different species. The only excep- to the thorax which is the center of loco- tion to this is the honey bee. We’ve in- motion of insects. Bees may or may not cluded some of the most common bees have legs that are modified for carrying for each group that you may see in this pollen. In honey bees, this is called the region. There are at least 20 different gen- corbicula, or pollen basket. Additional era of bees in the South Puget Sound, and markings or groups of hairs are often likely dozens of species. These plates are found on the abdomen of the insect. not intended to be a taxonomic identifica- Many bees carry pollen on the underside tion guide, but rather a means to familiar- of the abdomen instead of on the leg. Abdomen ize yourself with the native bees of our region. Common names are given, and Thorax Latin names are included to Genus or Species. Head Key Characteristics: Some images have arrows which point to key characteristics. Those characteristics are described below the picture. Additional Information: fore leg For additional information on each group, see the group description pages. 9 10 Honey bees (HB) Bumble bees cont. (BB) Common Name: honey bee Common Name: yellow bumble bee Family: Apidae Family: Apidae Genus: Apis Genus: Bombus Species: Apis mellifera Species: Bombus fervidus Key Characteristic: flattened hind leg Key Characteristic: yellow body, single black band between wings Group Description: p. 24 Group Description: p. 26 Bumble bees (BB) Common Name: yellow-faced bumble Common Name: tricolored bumble bee bee Family: Apidae Family: Apidae Genus: Bombus Genus: Bombus Species: Bombus mixtus Species: Bombus vosnesenskii Key Characteristic: three distinct colors, Key Characteristic: yellow face, most- orange rear end ly black body Group Description: p. 26 Group Description: p. 26 11 12 Other large bees (OLB) Other large bees cont. (OLB) Common Name: long-horned bee Common Name: sand/mining bee Family: Apidae Family: Andrenidae Genus: Melissodes Genus: Andrena Key Characteristic: very hairy hind leg Key Characteristic: non-descript black abdomen, sometime with lighter thorax Group Description: p. 28 Group Description: p. 28 Common Name: digger bee Common Name: mason/leaf-cutter bee Family: Apidae Family: Megachilidae Genus: Habropoda Genus: Osmia Key Characteristic: long antenna are Key Characteristic: blue shiny body, common hair on bottom of abdomen Group Description: p. 28 Group Description: p. 28 13 14 Other large bees cont. (OLB) Green bees cont. (GB) Common Name: mason/leaf-cutter bee Common Name: sweat bee Family: Megachilidae Family: Halictidae Genus: Anthidium Genus: Lasioglossum Key Characteristic: thick hair on bottom Key Characteristic: bright blue/green of abdomen body Group Description: p. 28 Group Description: p. 30 Small bees (SB) Green bees (GB) Common Name: masked bee Common Name: sweat bee Family: Colletidae Family: Halictidae Genus: Hylaeus Genus: Agapostemon Key Characteristic: yellow markings on Key Characteristic: green to black head face and thorax, green to black stripped abdo- men Group Description: p. 32 Group Description: p. 30 15 16 Small bees cont. (SB) Small bees cont. (SB) Common Name: sweat bee Common Name: sand/mining bee Family: Halictidae Family: Andrenidae Genus: Halictus Genus: Panurginus Key Characteristic: hairy hind legs and Key Characteristic: non-descript black stripped abdomen body Group Description: p. 32 Group Description: p. 32 Common Name: cuckoo sweat bee Family: Halictidae Common Name: small carpenter bee Genus: Sphecodes Family: Apidae Key Characteristic: red abdomen, black Genus: Ceratina head and thorax Key Characteristic: protrusion at end of Group Description: p. 32 abdomen Group Description: p. 32 17 18 Butterflies (BTF) Flies (F) Common Name: hover flies Common Name: skipper butterfly Family: Syrphidae Family: Hesperiidae Key Characteristics: one set of wings, Key Characteristics: clubbed antennae, very large eyes scale covered wings Bugs (BG) Wasps (W) Common Name: common wasp Common Name: stink bugs Family: Vespidae Family: Pentatomidae Genus: Vespula Key Characteristics: triangular area be- Species: Vespula vulgaris tween leathery wings Key Characteristic: obvious pinched abdomen, aggressive looking 19 20 Beetles (BTL) Section II: Group Descriptions Sizing and Shape Guide: Size and shape are often useful tools to identify bees. Each group description includes an sizing guide for your reference. Sizes: Large Medium Common name: chafer beetle Family: Scarabidae Small Key Characteristics: hardened front Shapes: wings, distinct mouthparts Narrow Spiders (S) Robust Round Seasonality and Range Maps: In addition to size, all bees have a unique geographic distribution that changes over the course of the season. Us- Common Name: crab spider ing previously Family: Thomisidae collected data, Key Characteristics: four pairs of legs we were able to prepare a bar chart for each group, and generate a series 21 22 of maps that show the abundance of each Honey bees (HB) bee group over the course of any given Group Description: Honey bees are me- season.
Recommended publications
  • Journal of Hymenoptera Research
    c 3 Journal of Hymenoptera Research . .IV 6«** Volume 15, Number 2 October 2006 ISSN #1070-9428 CONTENTS BELOKOBYLSKIJ, S. A. and K. MAETO. A new species of the genus Parachremylus Granger (Hymenoptera: Braconidae), a parasitoid of Conopomorpha lychee pests (Lepidoptera: Gracillariidae) in Thailand 181 GIBSON, G. A. P., M. W. GATES, and G. D. BUNTIN. Parasitoids (Hymenoptera: Chalcidoidea) of the cabbage seedpod weevil (Coleoptera: Curculionidae) in Georgia, USA 187 V. Forest GILES, and J. S. ASCHER. A survey of the bees of the Black Rock Preserve, New York (Hymenoptera: Apoidea) 208 GUMOVSKY, A. V. The biology and morphology of Entedon sylvestris (Hymenoptera: Eulophidae), a larval endoparasitoid of Ceutorhynchus sisymbrii (Coleoptera: Curculionidae) 232 of KULA, R. R., G. ZOLNEROWICH, and C. J. FERGUSON. Phylogenetic analysis Chaenusa sensu lato (Hymenoptera: Braconidae) using mitochondrial NADH 1 dehydrogenase gene sequences 251 QUINTERO A., D. and R. A. CAMBRA T The genus Allotilla Schuster (Hymenoptera: Mutilli- dae): phylogenetic analysis of its relationships, first description of the female and new distribution records 270 RIZZO, M. C. and B. MASSA. Parasitism and sex ratio of the bedeguar gall wasp Diplolqjis 277 rosae (L.) (Hymenoptera: Cynipidae) in Sicily (Italy) VILHELMSEN, L. and L. KROGMANN. Skeletal anatomy of the mesosoma of Palaeomymar anomalum (Blood & Kryger, 1922) (Hymenoptera: Mymarommatidae) 290 WHARTON, R. A. The species of Stenmulopius Fischer (Hymenoptera: Braconidae, Opiinae) and the braconid sternaulus 316 (Continued on back cover) INTERNATIONAL SOCIETY OF HYMENOPTERISTS Organized 1982; Incorporated 1991 OFFICERS FOR 2006 Michael E. Schauff, President James Woolley, President-Elect Michael W. Gates, Secretary Justin O. Schmidt, Treasurer Gavin R.
    [Show full text]
  • Insects and Related Arthropods Associated with of Agriculture
    USDA United States Department Insects and Related Arthropods Associated with of Agriculture Forest Service Greenleaf Manzanita in Montane Chaparral Pacific Southwest Communities of Northeastern California Research Station General Technical Report Michael A. Valenti George T. Ferrell Alan A. Berryman PSW-GTR- 167 Publisher: Pacific Southwest Research Station Albany, California Forest Service Mailing address: U.S. Department of Agriculture PO Box 245, Berkeley CA 9470 1 -0245 Abstract Valenti, Michael A.; Ferrell, George T.; Berryman, Alan A. 1997. Insects and related arthropods associated with greenleaf manzanita in montane chaparral communities of northeastern California. Gen. Tech. Rep. PSW-GTR-167. Albany, CA: Pacific Southwest Research Station, Forest Service, U.S. Dept. Agriculture; 26 p. September 1997 Specimens representing 19 orders and 169 arthropod families (mostly insects) were collected from greenleaf manzanita brushfields in northeastern California and identified to species whenever possible. More than500 taxa below the family level wereinventoried, and each listing includes relative frequency of encounter, life stages collected, and dominant role in the greenleaf manzanita community. Specific host relationships are included for some predators and parasitoids. Herbivores, predators, and parasitoids comprised the majority (80 percent) of identified insects and related taxa. Retrieval Terms: Arctostaphylos patula, arthropods, California, insects, manzanita The Authors Michael A. Valenti is Forest Health Specialist, Delaware Department of Agriculture, 2320 S. DuPont Hwy, Dover, DE 19901-5515. George T. Ferrell is a retired Research Entomologist, Pacific Southwest Research Station, 2400 Washington Ave., Redding, CA 96001. Alan A. Berryman is Professor of Entomology, Washington State University, Pullman, WA 99164-6382. All photographs were taken by Michael A. Valenti, except for Figure 2, which was taken by Amy H.
    [Show full text]
  • A Catalogue of the Family Andrenidae (Hymenoptera: Apoidea) of Eritrea
    Linzer biol. Beitr. 50/1 655-659 27.7.2018 A catalogue of the family Andrenidae (Hymenoptera: Apoidea) of Eritrea Michael MADL A b s t r a c t : In Eritrea the family Andrenidae is represented by two species of the genus Andrena FABRICIUS, 1775 (subfamily Andreninae) and one species each oft he genera Borgatomelissa PATINY, 2000 and Meliturgula FRIESE, 1903 (both subfamily Panurginae). The record of the genus Andrena is doubtful. K e y w o r d s : Andrenidae, Andreninae, Andrena, Panurginae, Borgatomelissa, Meliturgula, Eritrea. Introduction In the Afrotropical region the family Andrenidae is represented by more than 30 species (EARDLEY & URBAN 2010). The Eritrean fauna of of the family Andrenidae is poorly known. The knowledge is based on the material collected by J.K. Lord in 1869 and by P. Magretti in 1900. No further material is known. Up to date two species each of the subfamilies Andreninae (genus Andrena FABRICIUS, 1775) and Panurginae (genera Borgatomelissa PATINY, 2000 and Meliturgula FRIESE, 1903) have been recorded from Eritrea. Abbreviations Afr. reg. .................................. Afrotropical region biogeogr. ......................................... biogeography biol. .......................................................... biology cat. ......................................................... catalogue descr. ................................................... description fig. (figs) ....................................... figure (figures) syn. ......................................................... synonym tab.
    [Show full text]
  • A Survey of Alfalfa Pollinators and Polination in the San Joaquin
    AN ABSTRACT OF THE THESIS OF Philip Frank Torchio for the M. S. in Entomology (Name) (Degree) (Major Date thesis is presented January 10, 1966 Title A Survey of Alfalfa Pollinators and Pollination in the San Joaquin Valley of California with Emphasis on Establishment of the Alkali Bee Abstract-- approved --Redacted for privacy (Major professor) The study involves the establishment, management, and behavioral observations of Nomia melanderi Cockerell and Megachile rotundata (Fabricius) in California, a survey of native bees within western Fresno County, a determination of selfing and /or parthenocarpy in two fields of alfalfa, and observations on the honeybee as an alfalfa pollinator. The alkali bee is characterized. It is an endemic species limited to western North America. Eight artificial alkali bee nesting sites were constructed during the establishment program. Details of materials used during construction and their importance are discussed. Alkali bee cores used for introduction into California were procured in eastern Oregon and western Idaho, trucked to California, and planted in artificial sites. The heavy usage of insecticides in the area of study proved to be the greatest problem in the survival of the alkali bee. Observed toxicities of Dimethoate, Sevin, Malathion, Systex -Toxaphene, Toxaphene, DDT, Phosdrin, Dibrom, TEPP, Dylox, and Kelthane on the alkali bee were compared. The occurrence of multiple generations of the alkali bee in California is reported. Explanation of this behavior is based upon high soil temperatures (82° F. and above) during the nesting season. The importance of multiple generations is discussed. Biologies and importance of alkali bee parasites in California are described.
    [Show full text]
  • A New Species of Andrena (Trachandrena) from the Southwestern United States (Hymenoptera, Andrenidae)
    JHR 77: 87–103 (2020) A new species of Trachandrena 87 doi: 10.3897/jhr.77.53704 RESEARCH ARTICLE http://jhr.pensoft.net A new species of Andrena (Trachandrena) from the Southwestern United States (Hymenoptera, Andrenidae) Cory S. Sheffield1 1 Royal Saskatchewan Museum, 2340 Albert Street, Regina, Saskatchewan, S4P 2V7, Canada Corresponding author: Cory S. Sheffield ([email protected]) Academic editor: Jack Neff | Received 27 April 2020 | Accepted 31 May 2020 | Published 29 June 2020 http://zoobank.org/21056983-AAB7-42A5-A08B-E61139F474C2 Citation: Sheffield CS (2020) A new species ofAndrena (Trachandrena) from the Southwestern United States (Hymenoptera, Andrenidae). Journal of Hymenoptera Research 77: 87–103. https://doi.org/10.3897/jhr.77.53704 Abstract A new species of Andrena Fabricius, 1775, subgenus Trachandrena Robertson, 1902 is described and illus- trated, A. hadfieldi sp. nov., from Arizona, United States. The new species, presently known only from the female holotype, was collected in a Malaise trap in 1994, and remained unstudied until recently. In addition, Trachandrena is compared to similar subgenera in North America to assist in recognizing new members. Keywords Bee, new species, Trachandrena, North America, Arizona Introduction Andrena Fabricius, 1775 is one of the largest genera of bees, with 1,556 species (Ascher and Pickering 2020). Dubitzky et al. (2010) estimated that there are likely ca 2,000 species, suggesting there are many undescribed species, especially in Mesoamerica and in the dry regions of Central Asia. Though the genus is mainly Holarctic, it extends into Mesoamerica, parts of Africa and tropical Asia (Michener 2007). The subgenus Trachandrena Robertson, 1902 is represented by 30 species glob- ally (Gusenleitner and Schwarz 2002; Michener 2007), 24 of which occur in the Copyright Cory S.
    [Show full text]
  • Appendix 1 the 615 Bee Species That Were Used in the Anlayses
    Oikos OIK-05260 Gérard, M., Vanderplanck, M., Franzen, M., Kuhlmann, M., Potts, S. G., Rasmont, P., Schweiger, O. and Michez, D. 2018. Patterns of size variation in bees at a continental scale: does Bergmann’s rule apply? – Oikos doi: 10.1111/oik.05260 Appendix 1 The 615 bee species that were used in the anlayses. The nomenclature of the bees follows Kuhlmann (2014). Species are sorted alphabetically. Species ITD (mm) Family Sociality Nesting Andrena abbreviata 1.90 Andrenidae solitary ground Andrena aberrans 2.93 Andrenidae solitary ground Andrena abrupta 1.84 Andrenidae solitary ground Andrena aciculata 1.40 Andrenidae solitary ground Andrena aegyptiaca 1.61 Andrenidae solitary ground Andrena aeneiventris 1.39 Andrenidae solitary ground Andrena aerinifrons 2.00 Andrenidae solitary ground Andrena afrensis 2.92 Andrenidae solitary ground Andrena agilissima 2.94 Andrenidae solitary ground Andrena agnata 2.23 Andrenidae solitary ground Andrena albopunctata 3.00 Andrenidae solitary ground Andrena alfkenella 1.32 Andrenidae solitary ground Andrena alfkenelloides 1.48 Andrenidae solitary ground Andrena allosa 1.94 Andrenidae solitary ground Andrena alluaudi 1.97 Andrenidae solitary ground Andrena anatolica 1.94 Andrenidae solitary ground Andrena angustior 1.88 Andrenidae solitary ground Andrena anthrisci 1.30 Andrenidae solitary ground Andrena antigana 2.36 Andrenidae solitary ground Andrena apicata 2.64 Andrenidae solitary ground Andrena apiformis 3.17 Andrenidae solitary ground Andrena argentata 1.86 Andrenidae solitary ground Andrena asperrima
    [Show full text]
  • Bee Diversity in Iowa City, Iowa
    Urban Pollinators: Bee diversity in Iowa City, Iowa By: Cameron, Ashley, Sam, Will, Chang, and Alex http://blog.nwf.org/2013/05/in-the-buzz-about-bees-dont-forget-the-natives/ Urban bees Managed bees - Honey bees (Apis)/bumble bees/solitary bees - Agriculture Wild native bees - Bumble bees, mining bees, squash bees and leaf cutting bees (The Xerces Society for Invertebrate Conservation, 2014) - Urban gardens/yards E.g. Bumble bees- tomato and pepper; bottle gentian E.g. Solitary blue orchard bee (Osmia lignaria)-cold weather, willow, apples, cherries E.g. Squash bee (Peponapis pruinosa) –pumpkins, squash and other cucurbits Specialist vs generalists - Bumble bees depend on succession of plants flowering from early spring when the queen emerges to late summer – early fall when the colony dies - squash bees (cucurbit plants) Ground-nesting Bees in Iowa - Live underground in burrows and tunnels - Bumble bees: pre-existing cavities made by rodents or small mammals - Digger bees: Andrenidae, Halictidae and Colletidae bees (e.g. yellow-faced bees), genera Lassioglossum and Agapostemon - thin and sparse vegetation + loose soil - Native solitary bees: females dig cylindrical tunnels in shady areas for offspring - docile and less likely to sting than social bees Hole-nesting bees in Iowa - Take advantage of already existing holes - Most members of the family Megachilidae, e.g. mason and leafcutter bees, are hole nesters who use certain materials to modify their nest chambers - Mason bees (Osmia): use mud to construct partition walls between adjacent cells and a thicker plug to seal the nest entrance from parasites - leafcutter bees: cut rounded leaf pieces for the same purpose, to line the inner walls of the nest burrow Carpenter bees in Iowa - Create burrows, or holes, of their own making - Have powerful jaws called mandibles with which they can excavate tunnels in wood Prefer soft wood and dislike paint or other finishing materials.
    [Show full text]
  • OTHER BEES and WASPS Advanced Level Training Texas Master Beekeeper Program
    OTHER BEES AND WASPS Advanced Level Training Texas Master Beekeeper Program Introduction • As a beekeeper, you are often treated as the expert on all things with wings or stings. • The knowledge gained from this presentation should help you to confidently field questions from the general public, identify a few of the common bees and wasps of Texas and discuss their biology and importance as beneficial insects or as pests. Bees and Wasps Bees Wasps • More body hair • Very little hair • Flattened hindlegs, usually • Rounded legs containing a pollen basket • Are predators of other insects, or will • Feed on pollen and nectar scavenge food scraps, carrion, etc. • Generally can only sting once • Can (and will) sting repeatedly • Includes hornets and yellowjackets 1 Yellowjackets and Hornets • General biology • Colonies founded in spring by a single‐mated, overwintered queen • Constructs the paper brood cells • Forage for food • Lay eggs • Feed her progeny • Defend the nest Yellowjackets and Hornets • When the first offspring emerge they assume all tasks except egg laying. • Workers progressively feed larvae • Masticated adult and immature insects • Other arthropods • Fresh carrion • Working habits apparently are not associated with age as they are with honey bees. Yellowjackets and Hornets in Texas • Eastern yellowjacket • Vespula maculifrons Buysson • Southern yellowjacket • Vespula squamosa Drury • Baldfaced hornet • Dolichovespula maculata Linnaeus 2 Yellowjackets and Hornets in Texas • Eastern yellowjacket (Vespula maculifrons) • Family: Vespidae • Mostly subterranean nests, but aerial nests do occur. • Largest recorded nest: • 8 levels of comb with over 2800 wasps present (Haviland, 1962) Yellowjackets and Hornets in Texas • Southern yellowjacket (Vespula squamosa) • Family: Vespidae • Construct both terrestrial and aerial nests.
    [Show full text]
  • Bees of Ohio: a Field Guide
    Bees of Ohio: A Field Guide North American Native Bee Collaborative The Bees of Ohio: A Field Guide (Version 1.1.1 , 5/2020) was developed based on Bees of ​ Maryland: A Field Guide, authored by the North American Native Bee Collaborative ​ Editing and layout for The Bees of Ohio : Amy Schnebelin, with input from MaLisa Spring and Denise Ellsworth. Cover photo by Amy Schnebelin Copyright Public Domain. 2017 by North American Native Bee Collaborative Public Domain. This book is designed to be modified, extracted from, or reproduced in its entirety by any group for any reason. Multiple copies of the same book with slight variations are completely expected and acceptable. Feel free to distribute or sell as you wish. We especially encourage people to create field guides for their region. There is no need to get in touch with the Collaborative, however, we would appreciate hearing of any corrections and suggestions that will help make the identification of bees more accessible and accurate to all people. We also suggest you add our names to the acknowledgments and add yourself and your collaborators. The only thing that will make us mad is if you block the free transfer of this information. The corresponding member of the Collaborative is Sam Droege ([email protected]). ​ ​ First Maryland Edition: 2017 First Ohio Edition: 2020 ISBN None North American Native Bee Collaborative Washington D.C. Where to Download or Order the Maryland version: PDF and original MS Word files can be downloaded from: http://bio2.elmira.edu/fieldbio/handybeemanual.html.
    [Show full text]
  • Missouri Bee Identification Guide Edward M
    Missouri Bee Identification Guide Edward M. Spevak 1, Michael Arduser 2, 1 Saint Louis Zoo 2 Missouri Department of Conservation Bees are Beneficial Honey bees (Apis mellifera) Leafcutter and Mason bees (Megachile spp. & Osmia spp.) Bees play an essential role in natural and agricultural systems Family: Apidae. Heart-shaped head; black Family: Megachilidae. Head as broad as as pollinators of flowering plants that provide food, fiber, to amber-brown body with pale and dark thorax; large mandibles; black body most spices, medicines and animal forage. Plants rely on pollinators stripes on abdomen; pollen baskets on hind with pale bands on abdomen (metallic green to reproduce and set seed and fruit. In fact, approximately legs; 10-15 mm. or blue for Osmia); pollen carrying hairs three-quarters of all flowering plants rely on pollinators to ● Large social colonies, 30,000 or more; live under abdomen; 5-20 mm. reproduce. Honey bees pollinate crops, but native bees also in man-made hives and natural cavities like ● Solitary, but nest in aggregations in have a role in agriculture and are essential for pollination in tree hollows. Swarm to locate new nests. natural or man-made holes such as beetle natural landscapes. There are over 425 native species of ground- ● Honey bees are not native to the U.S., but holes, nesting blocks, stems, or soil. nesting, wood-nesting and parasitic bees found within Missouri. were brought over by Europeans in the ● Females cut circular pieces from leaves This guide identifies 10 groups of bees commonly observed in 17th century. to line their nests.
    [Show full text]
  • Insects of Grand Canyon: an Overview for Hiking Guides
    INSECTS OF GRAND CANYON: AN OVERVIEW FOR HIKING GUIDES Larry Stevens Museum of Northern Arizona [email protected] Giants on Whose Shoulders We Stand Nothing in biology makes sense except in the lightof evolution Theodore Dobzhansky Basin and Range Rocky Mtn. Geologic Prov. Geol. Prov. Virgin R. Rkm 0 Grand Canyon Rkm Rkm 400 Rkm 100 Rkm 200 300 Basin and Range Geologic Prov. Grand Canyon Rims Map (Billingsley and Hampton 1999) Eastern Basin Western Basin Muav Gorge THE BIOGEOGRAPHY OF LARGE, DEEP CANYONS (STEVENS 2012) Barrier/Filter Refugia North Rim Barrier/ Corridor Barrier/ Filter Flow Direction Filter South Rim Barrier/Filter ARTHROPODA: Half a Billion Years of World Dominance Segmented worm ancestor Chelicerata Mites Bark Scorpion (Buthidae: Centruroides) Scorpiones Silurian (430 m.y. ago) AZ Giant Hairy Scorpion (Vaejovidae: Hadrurus) Scolopendron viridis Centipede Hyallela Amphipod Orthoporus ornatus Millepede AQUATIC AND RIPARIAN INSECTS Belostoma ~fluminium Mayflies: ♂ Stoneflies: Plecoptera Ephemeroptera Naucoridae Ranatra Masked Clubskimmer Rhagovelia (Brechmorhoga pertinax) distincta Aquarius remigis Odonata – 89 spp, dominated by neotropical fauna, with ~equivalent boreal, range-centered, and Pacific coast elements; 78% of fauna is exogenous 13 5 18 20 3 25 endemic 1 29 ANT LIONS: MYRMELEONTIDAE Thermonectus Chrysina Dryopoid Beetles Prionus Tiger Beetles Ground Beetles DIPTERA Nematocera: “Two Winged Insects” Ancestral flies with aquatic larvae (e.g., Culicidae, Simuliidae, Chironomidae) Brachycera: Tabanomorpha
    [Show full text]
  • The Bee Genus Andrena (Andrenidae) and the Tribe Anthophorini (Apidae) (Insecta: Hymenoptera: Apoidea)
    Studies in phylogeny and biosystematics of bees: The bee genus Andrena (Andrenidae) and the tribe Anthophorini (Apidae) (Insecta: Hymenoptera: Apoidea) Dissertation zur Erlangung des Doktorgrades der Fakultät für Biologie der Ludwig-Maximilians-Universität München vorgelegt von Andreas Dubitzky Hebertshausen, 16. Dezember 2005 Erstgutachter: Prof. Dr. Klaus Schönitzer Zweitgutachter: PD Dr. Roland Melzer Tag der Abgabe: 16.12.05 Tag der mündlichen Prüfung: 23.5.06 Disclaimer All nomenclaturically relevant acts in this thesis have to be regarded as unpublished according to Article 8 of the International Code of Zoological Nomenclature, and will become available by separate publications. This dissertation is dedicated to my parents Heinz and Christine Dubitzky, who gave me the opportunity to carry out these studies and continuously supported me with their love and patience. Contents 1. Introduction............................................................................................................1 2. Material and methods............................................................................................4 2.1 Material examined ......................................................................................4 2.1.1 Morphological studies.......................................................................4 2.1.2 Molecular analysis ............................................................................5 2.2 Preparation of male genitalia and female head capsule including mouthparts...................................................................................5
    [Show full text]