DOCSLIB.ORG

Chalcidoidea: Chalcididae) from India with a Key to Oriental Species

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Chalcidoidea: Chalcididae) from India with a Key to Oriental Species ISSN 0973-1555(Print) ISSN 2348-7372(Online) HALTERES, Volume 10, 80-85, 2019 C. BINOY, P.M. SURESHAN, S. SANTHOSH AND M. NASSER doi: 10.5281/zenodo.3596046 Description of a new species of Lasiochalcidia Masi (Chalcidoidea: Chalcididae) from India with a key to Oriental species *C. Binoy1,3, P.M. Sureshan2, S. Santhosh3 and M. Nasser1 1Insect Ecology and Ethology Laboratory, Department of Zoology, University of Calicut, Kerala-673635, India. 2 Western Ghats Regional Centre, Zoological Survey of India, Eranhipalam, Kozhikode, Kerala-673006, India. 3 Systematic Entomology Laboratory, Malabar Christian College, (Affiliated to University of Calicut), Kozhikode, Kerala-673001, India. (Email: binoy_doz@uoc.ac.in) Abstract Lasiochalcidia Masi, 1929 (Hymenoptera: Chalcididae) is one of the rarest chalcid genera to have been recorded from the world. Association with antlions and the peculiar mode of oviposition makes the genus more interesting. Here we describe and illustrate a new species of Lasiochalcidia Masi with a key to Oriental species. Keywords: Chalcididae; Lasiochalcidia Masi; New Species; India; Oriental Region. Received: 11 July 2019; Revised: 26 December 2019; Online: 31 December 2019 Introduction Lasiochalcidia Masi, 1929 is one of innate ability to discover hidden hosts by the least common genera of hybothoracine perceiving the movements on loose soil made (Haltichellinae: Hybothoracini) tribe to occur by the antlion larva, using the specialised in any collection from the tropics. Presently mechanoreceptors on the antennae. The female constituting of 23 species worldwide, the parasitoid provokes the antlion larva to attack species is mostly associated as parasitoids of its hindlegs with the powerful and deadly antlion larvae (Neuroptera: Myrmeleontidae) mandibles of antlion larva. When the antlion (Nikol’skaya, 1978; Bouček 1988; Noyes, grabs on, the toothed mandibles get held on by 2019). The genus is represented by seven the rugate basal teeth of the hindlegs of the species from the Oriental region of which only parasitoid. Now, parasitoid stretches the two species are recorded from the Indian antlion’s jaws apart using its muscular legs subcontinent viz., L. dargelasii (Latreille, making the predator helpless and expose its 1805) and L. pilosella (Cameron, 1904). most vulnerable and least chitinous part of its Host association and the mode of body, the ventral side of its neck. The parasitisation of species in Lasiochalcidia parasitoid takes its time and carefully oviposit have been of much interest. Bouček (1956) an egg through the antlion's exposed throat not reported Lepidopteran pupae as hosts for harming any of its vital organs. The task when Lasiochalcidia species, but this is very completed, the parasitoid releases the helpless doubtful as Lasiochalcidia is commonly antlion larva from its hold and flies off. The regarded as a potential parasitoid of much antlion larva is thus left to rot while the voracious predators, the ant-lion larvae parasitoid lives within its body and the adult (Lotfalizadeh et al., 2012). The peculiarity in wasp emerges off the pits build by antlion oviposition is described by many as a larva when they fully mature. spectacle to watch. Stefan (1958, 1959, 1961, 1966) Materials and Methods studied the ovipositional behaviour of several The specimens for the present study species of Lasiochalcidia and found their was retrieved employing both sweeping from 80 Description of a new species of Lasiochalcidia Masi from India an abandoned vegetable field in Edakkara and Lasiochalcidia (Anoplochalcidia) Steffan, by passive yellow pan trap collections from 1953: 34. New status for Jafferkhan colony, Kozhikode district and near Anoplochalcidia Steffan (page 34) D. B. College, Shasthamkotta in Kollam Oxycoryhpus Cameron, 1904: 109. Type district of Kerala, India. The specimens were species: Oxycoryhpus pilosellus preserved in 70% ethyl alcohol and later card Cameron, by monotypy mounted for microscopic observation. The Oxycoryphiscus Ghesquiere, 1946: 368. parasitoids were examined under a Replacement name for Oxycoryphus stereoscopic binocular microscope of model Cameron, 1904 nec Fischer, 1853. LEICA M205 and the images were captured Diagnosis: Temples almost lacking in profile, with the camera model LEICA DFC 500. vertex above in anterio-posterior view very Measurements were obtained using Leica LAS thin; frons and gena covered with thick silvery (Leica Application Suite V3.80) microsystems bristles; posterior margin of pronotum with a by Leica (Heerburg, Switzerland). Images at border of minute bristles; scutellum apically varying depth were stacked using Leica Auto often bidentate, propodeum sloping steeply montage Software V3.80 and the final onto gaster; scape in males often with illustrations were post-processed for contrast prominent horn like dent below. and brightness using Adobe® Photoshop® CS5 (Version 12.0 x64) software. The type Hosts: Most species of the genus are parasites specimens are deposited in the National on antlion larvae (Neuroptera: Myrmeleon- Zoological collections of Zoological Survey of tidae) (Bouček, 1988; Noyes, 2019). India, Western Ghat Regional Centre, Kozhikode (ZSIK). Distribution: This genus is distributed in Africa, Europe and Asia (Noyes, 2019). Terms and measurements: The terminology used is mainly that of Narendran (1989) and Key to Oriental species of Lasiochalcidia Narendran & van Achterberg (2016). The Masi nomenclature for cuticular sculpturing follows (Modified from Narendran, 1989) Harris (1979).The general abbreviations of the 1. Females....................................................2 terms are as follows: – Males........................................................6 F1–F7: First to seventh funicular segments 2. Hind femora less than 2× as long as wide; MV: Marginal vein distal half not narrower than proximal OD: Diameter of median ocellus (equal to subequal) half; scrobal striations OOL: Minimum distance between posterior narrow, slightly convex...........................3 ocelli and compound eye – Hind femora distinctly more than 2× as PMV: Postmarginal vein long as wide; distal half a trifle narrower POL: Distance between two posterior ocelli than proximal half; other characters partly STV: Stigmal vein or completely different............................4 T1-T6: Abdominal tergites one to six 3. Hind femora distinctly much less than 2× as long as wide; scrobal striae strong and Results and Discussion narrowed; apex of scutellum well Genus Lasiochalcidia Masi, 1929 emarginate; T1 smooth and shiny.............. Anoplochalcidia Steffan, 1951: 2. Type ……………………….…L. pilosella (Cam.) species: Anoplochalcidia guineensis – Hind femora less than or equal to 2× as Steffan, original designation and long as wide; scrobal striae weak and monotypy wider than in alternate; apex of scutellum Dromochalcidia Masi, 1929: 185. Type weakly emarginate to rounded; T1 species: Dromochalcidia moluccensis shagreened ……….............L. narendrani Masi, by monotypy Binoy and Sureshan sp.n. Lasiochalcidia Masi, 1929: 209-220. Type 4. Apex of scutellum prominently bilobate; species: Euchalcis rubripes Kieffer, by hind femora black; propodeum with subsequent designation of Nikol'skaya, median area with several longitudinal M. (1952). carinae...................L. moluccensis (Masi) 81 C. Binoy, P.M. Sureshan, S. Santhosh and M. Nasser – Apex of scutellum weakly emarginate; brownish to black, pedicel brown, F1 and F2 hind femora usually red, propodeum brownish black, remaining flagellar segments without longitudinal carinae ...................5 brownish black with apices brown, clava 5. Striations of scrobe almost straight; apex basally liver brownish, remaining portion of scutellum produced in the form of a brownish black; fore and mid femur and tibia flange; propodeum with lateral teeth reddish brown, fore and mid coxa black, all weak; antennae and legs completely tarsi testaceous; hind coxa dorsally shiny black rufous…………....L. thresiae Narendran with fine setose punctures ventrally, hind – Striations of scrobe convex; apex of femora reddish brown with black tooth at scutellum bidentate; propodeum with ventral margin and scattered pits on the disc prominent lateral teeth; antennae, fore and arising from which are thin silvery white long mid femora and hind tibiae usually hairs, hind tibia reddish brown dorsally, blackish...................... L. dargelasii (Lat.) ventrally black, carinate, surface with scattered 6. Antenna with scape without a prominent thin white hairs; gaster black with dense dent below...........................L. narendrani golden yellow pubescence laterally on T2 to Binoy and Sureshan sp.n. T5; T6 completely covered with rows of – Antenna with scape having a prominent golden yellow pubescence; pubescence on the dent below................................................7 body golden. 7. Apex of scutellum deeply or well incised Head (Fig. 1–3): Head a trifle wider than and bilobate..............................................8 width of thorax (excluding tegula), 1.86× as – Apex of scutellum hardly emarginate, high as wide in lateral view, moderately almost entire…….......L. dargelasii (Lat.) punctate with thin golden yellow pubescence 8. Hind leg black........L. birmanus (M.&D.) on the upper face extending to the occiput; – Hind leg red...............L. pilosella (Cam.) lower face moderately punctate with dense silvery white pubescence extending to the Lasiochalcidia narendrani Binoy and gena; POL 3.25× OOL, median ocellus Sureshan sp.n. slightly
Load more

Recommended publications
  • New Records of the Family Chalcididae (Hymenoptera: Chalcidoidea) from Egypt
    Zootaxa 4410 (1): 136–146 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2018 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4410.1.7 http://zoobank.org/urn:lsid:zoobank.org:pub:6431DC44-3F90-413E-976F-4B00CFA6CD2B New records of the family Chalcididae (Hymenoptera: Chalcidoidea) from Egypt MEDHAT I. ABUL-SOOD1 & NEVEEN S. GADALLAH2,3 1Zoology Department, Faculty of Science (Boys), Al-Azhar University, P.O. Box 11884, Nasr City, Cairo, Egypt. E-mail: medhatibrahiem_23@yahoo.com 2Entomology Department, Faculty of Science, Cairo University, Giza, Egypt 3Corresponding author. E-mail: n_gadallah@hotmail.com Abstract In the present study, a checklist of new records of the family Chalcididae of Egypt is presented based on a total of 180 specimens collected from 24 different Egyptian localities between June 2011 and October 2016, mostly by sweeping and Malaise traps. Nineteen species as well as the subfamily Epitraninae and the genera Bucekia Steffan, Epitranus Walker, Proconura Dodd, and Tanycoryphus Cameron, are newly recorded from Egypt. A single species previously placed in the genus Hockeria is transferred to Euchalcis Dufour as E. rufula (Nikol’skaya, 1960) comb. nov. Key words: Parasitic wasps, Chalcidinae, Dirhininae, Epitraninae, Haltichellinae, new records, new combination Introduction The Chalcididae (Hymenoptera: Chalcidoidea) is a medium-sized family represented by more than 1500 described species in 93 genera (Aguiar et al. 2013; Noyes 2017; Abul-Sood et al. 2018). A large number of described species are classified in the genus Brachymeria Westwood (about 21%), followed by Conura Spinola (20.3%) (Noyes 2017).
    [Show full text]
  • Hymenoptera: Chalcididae) in the European Continent
    Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 64 (1): 61–65 (2021) doi: 10.3897/travaux.64.e66165 FAUNISTIC NOTE First record of the subfamily Epitraninae (Hymenoptera: Chalcididae) in the European Continent Evangelos Koutsoukos1, 2, Gerard Delvare3 1 Section of Ecology and Systematics, Department of Biology, National and Kapodistrian University of Athens, 15784 Athens, Greece 2 Museum of Zoology, National and Kapodistrian University of Athens, 15784 Athens, Greece 3 Centre de Coopération Internationale en Recherche Agronomique pour le Développement (CIRAD), Montpellier SupAgro, INRA, IRD, Univ. Montpellier, Montpellier, France Corresponding author: Evangelos Koutsoukos (vag18000@gmail.com) Received 19 March 2021 | Accepted 25 May 2021 | Published 30 June 2021 Citation: Koutsoukos E, Delvare G (2021) First record of the subfamily Epitraninae (Hymenoptera: Chalcididae) in the European Continent. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 64(1): 61–65. https:// doi.org/10.3897/travaux.64.e66165 Abstract Epitraninae Burks (Hymenoptera: Chalcididae) is a subfamily with a single recognised genus, Epitranus Walker, known to be distributed throughout the tropical areas of the Old World. Whilst recent studies have reported the presence of Epitraninae in countries of the Middle East, there are no published records from the European continent. A female specimen belonging to the Epitranus hamoni species complex was collected in Salamis island, Attica, Greece, and deposited at the Museum of Zoology (Athens). This record constitutes an important addition to the Greek and European Chalcidoidea fauna. Keywords Chalcidoidea, Chalcididae, new record, Epitranus, Greece. Introduction Chalcidid wasps (Chalcidoidea: Chalcididae) are a moderate sized family regarding species number (Aguiar et al.
    [Show full text]
  • Phylogenetic Analysis of Eurytominae (Chalcidoidea: Eurytomidae) Based on Morphological Characters
    Blackwell Publishing LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082© 2007 The Linnean Society of London? 2007 1513 441510 Original Article PHYLOGENETIC ANALYSIS OF EURYTOMINAEH. LOTFALIZADEH ET AL. Zoological Journal of the Linnean Society, 2007, 151, 441–510. With 212 figures Phylogenetic analysis of Eurytominae (Chalcidoidea: Eurytomidae) based on morphological characters HOSSEINALI LOTFALIZADEH1, GÉRARD DELVARE2* and JEAN-YVES RASPLUS2 1Plant Pests and Diseases Research Institute, Evin, Tehran 19395–1454, Iran 2CIRAD – INRA, Centre de Biologie et de Gestion des Populations (CBGP), Campus International de Baillarguet, CS 30 016, F-34988 Montferrier-sur-Lez, France Received February 2006; accepted for publication December 2006 A phylogenetic study of the Eurytominae (Hymenoptera: Chalcidoidea) treating 178 taxa and based on 150 mor- phological characters is given. Several cladograms using the complete species sample, but obtained with different weightings, are presented. Local studies were also carried out to provide possible alternate topologies. The deep nodes of the trees were unstable and were never supported, but most of the superficial nodes were stable and robust. The results therefore provide support for a generic classification of the subfamily. The large genus Eurytoma – which includes about half of the described species of the subfamily – proved to be polyphyletic, and is redefined in a nar- rowed sense using putative synapomorphies. Bruchophagus and Prodecatoma were similarly redefined. The genera Philolema and Aximopsis are reconsidered and defined in a broader concept. A number of the species presently included in Eurytoma were transferred to these genera. Finally, 22 new generic synonymies are proposed and 33 spe- cies are transferred. The study also demonstrates that the Eurytomidae are polyphyletic.
    [Show full text]
  • A Phylogenetic Analysis of the Megadiverse Chalcidoidea (Hymenoptera)
    UC Riverside UC Riverside Previously Published Works Title A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera) Permalink https://escholarship.org/uc/item/3h73n0f9 Journal Cladistics, 29(5) ISSN 07483007 Authors Heraty, John M Burks, Roger A Cruaud, Astrid et al. Publication Date 2013-10-01 DOI 10.1111/cla.12006 Peer reviewed eScholarship.org Powered by the California Digital Library University of California Cladistics Cladistics 29 (2013) 466–542 10.1111/cla.12006 A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera) John M. Heratya,*, Roger A. Burksa,b, Astrid Cruauda,c, Gary A. P. Gibsond, Johan Liljeblada,e, James Munroa,f, Jean-Yves Rasplusc, Gerard Delvareg, Peter Jansˇtah, Alex Gumovskyi, John Huberj, James B. Woolleyk, Lars Krogmannl, Steve Heydonm, Andrew Polaszekn, Stefan Schmidto, D. Chris Darlingp,q, Michael W. Gatesr, Jason Motterna, Elizabeth Murraya, Ana Dal Molink, Serguei Triapitsyna, Hannes Baurs, John D. Pintoa,t, Simon van Noortu,v, Jeremiah Georgea and Matthew Yoderw aDepartment of Entomology, University of California, Riverside, CA, 92521, USA; bDepartment of Evolution, Ecology and Organismal Biology, Ohio State University, Columbus, OH, 43210, USA; cINRA, UMR 1062 CBGP CS30016, F-34988, Montferrier-sur-Lez, France; dAgriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, ON, K1A 0C6, Canada; eSwedish Species Information Centre, Swedish University of Agricultural Sciences, PO Box 7007, SE-750 07, Uppsala, Sweden; fInstitute for Genome Sciences, School of Medicine, University
    [Show full text]
  • Fauna of Chalcid Wasps (Hymenoptera: Chalcidoidea, Chalcididae) in Hormozgan Province, Southern Iran
    J Insect Biodivers Syst 02(1): 155–166 First Online JOURNAL OF INSECT BIODIVERSITY AND SYSTEMATICS Research Article http://jibs.modares.ac.ir http://zoobank.org/References/AABD72DE-6C3B-41A9-9E46-56B6015E6325 Fauna of chalcid wasps (Hymenoptera: Chalcidoidea, Chalcididae) in Hormozgan province, southern Iran Tahereh Tavakoli Roodi1, Majid Fallahzadeh1* and Hossien Lotfalizadeh2 1 Department of Entomology, Jahrom branch, Islamic Azad University, Jahrom, Iran. 2 Department of Plant Protection, East-Azarbaijan Agricultural and Natural Resources Research Center, Agricultural Research, Education and Extension Organization (AREEO), Tabriz, Iran ABSTRACT. This paper provides data on distribution of 13 chalcid wasp species (Hymenoptera: Chalcidoidea: Chalcididae) belonging to 9 genera and Received: 30 June, 2016 three subfamilies Chalcidinae, Dirhininae and Haltichellinae from Hormozgan province, southern Iran. All collected species are new records for the province. Accepted: Two species Dirhinus excavatus Dalman, 1818 and Hockeria bifasciata Walker, 13 July, 2016 1834 are recorded from Iran for the first time. In the present study, D. excavatus Published: is a new species record for the Palaearctic region. An updated list of all known 13 July, 2016 species of Chalcididae from Iran is also included. Subject Editor: George Japoshvili Key words: Chalcididae, Hymenoptera, Iran, Fauna, Distribution, Malaise trap Citation: Tavakoli Roodi, T., Fallahzadeh, M. and Lotfalizadeh, H. 2016. Fauna of chalcid wasps (Hymenoptera: Chalcidoidea: Chalcididae) in Hormozgan province, southern Iran. Journal of Insect Biodiversity and Systematics, 2(1): 155–166. Introduction The Chalcididae are a moderately specious Coleoptera, Neuroptera and Strepsiptera family of parasitic wasps, with over 1469 (Bouček 1952; Narendran 1986; Delvare nominal species in about 90 genera, occur and Bouček 1992; Noyes 2016).
    [Show full text]
  • Ichneumonid Wasps (Hymenoptera, Ichneumonidae) in the to Scale Caterpillar (Lepidoptera) [1]
    Central JSM Anatomy & Physiology Bringing Excellence in Open Access Research Article *Corresponding author Bui Tuan Viet, Institute of Ecology an Biological Resources, Vietnam Acedemy of Science and Ichneumonid Wasps Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam, Email: (Hymenoptera, Ichneumonidae) Submitted: 11 November 2016 Accepted: 21 February 2017 Published: 23 February 2017 Parasitizee a Pupae of the Rice Copyright © 2017 Viet Insect Pests (Lepidoptera) in OPEN ACCESS Keywords the Hanoi Area • Hymenoptera • Ichneumonidae Bui Tuan Viet* • Lepidoptera Vietnam Academy of Science and Technology, Vietnam Abstract During the years 1980-1989,The surveys of pupa of the rice insect pests (Lepidoptera) in the rice field crops from the Hanoi area identified showed that 12 species of the rice insect pests, which were separated into three different groups: I- Group (Stem bore) including Scirpophaga incertulas, Chilo suppressalis, Sesamia inferens; II-Group (Leaf-folder) including Parnara guttata, Parnara mathias, Cnaphalocrocis medinalis, Brachmia sp, Naranga aenescens; III-Group (Bite ears) including Mythimna separata, Mythimna loryei, Mythimna venalba, Spodoptera litura . From these organisms, which 15 of parasitoid species were found, those species belonging to 5 families in of the order Hymenoptera (Ichneumonidae, Chalcididae, Eulophidae, Elasmidae, Pteromalidae). Nine of these, in which there were 9 of were ichneumonid wasp species: Xanthopimpla flavolineata, Goryphus basilaris, Xanthopimpla punctata, Itoplectis naranyae, Coccygomimus nipponicus, Coccygomimus aethiops, Phaeogenes sp., Atanyjoppa akonis, Triptognatus sp. We discuss the general biology, habitat preferences, and host association of the knowledge of three of these parasitoids, (Xanthopimpla flavolineata, Phaeogenes sp., and Goryphus basilaris). Including general biology, habitat preferences and host association were indicated and discussed.
    [Show full text]
  • The Use of the Biodiverse Parasitoid Hymenoptera (Insecta) to Assess Arthropod Diversity Associated with Topsoil Stockpiled
    RECORDS OF THE WESTERN AUSTRALIAN MUSEUM 83 355–374 (2013) SUPPLEMENT The use of the biodiverse parasitoid Hymenoptera (Insecta) to assess arthropod diversity associated with topsoil stockpiled for future rehabilitation purposes on Barrow Island, Western Australia Nicholas B. Stevens, Syngeon M. Rodman, Tamara C. O’Keeffe and David A. Jasper. Outback Ecology (subsidiary of MWH Global), 41 Bishop St, Jolimont, Western Australia 6014, Australia. Email: Nick.Stevens@mwhglobal.com ABSTRACT – This paper examines the species richness and abundance of the Hymenoptera parasitoid assemblage and assesses their potential to provide an indication of the arthropod diversity present in topsoil stockpiles as part of the Topsoil Management Program for Chevron Australia Pty Ltd Barrow Island Gorgon Project. Fifty six emergence trap samples were collected over a two year period (2011 and 2012) from six topsoil stockpiles and neighbouring undisturbed reference sites. An additional reference site that was close to the original source of the topsoil on Barrow Island was also sampled. A total of 14,538 arthropod specimens, representing 22 orders, were collected. A rich and diverse hymenopteran parasitoid assemblage was collected with 579 individuals, representing 155 species from 22 families. The abundance and species richness of parasitoid wasps had a strong positive linear relationship with the abundance of potential host arthropod orders which were found to be higher in stockpile sites compared to their respective neighbouring reference site. The species richness and abundance of new parasitoid wasp species yielded from the relatively small sample area indicates that there are many species on Barrow Island that still remain to be discovered. This study has provided an initial assessment of whether the hymenoptera parasitoid assemblage can give an indication of arthropod diversity.
    [Show full text]
  • Appendix S4. the Tentorium and Its External Landmarks in the Chalcididae
    Appendix S4 . The tentorium and its external landmarks in the Chalcididae This study is the first in the whole superfamily Chalcidoidea to investigate the tentorium as a phylogenetic character and to establish the connection between the inner skeleton of the cephalic capsule and its external landmarks on the back of the head. In this section, details are provided on the methodology used by GD to examine and code the different bridges. S4.1. Context Phylogenetic informativeness of the characters of the head capsule in Hymenoptera was recently highlighted (Vilhelmsen 2011; Burks & Heraty 2015; Zimmermann & Vilhelmsen 2016). However, interpretation is difficult and requires landmarks (Burks & Heraty, 2015). More precisely, the identity of the sclerotized structures between the occipital foramen and the oral fossa are still debated. Homology and nomenclature of these structures were established by Snodgrass (1928, 1942 and 1960) and reassessed by Vilhelmsen (1999) and Burks & Heraty (2015). These authors describe various types of ‘bridges’, such as postgenal, hypostomal and subforaminal bridges, according to the cephalic part – postgena or hypostoma – from which they putatively originate. In his phylogenetic analyses of the Chalcididae, Wijesekara (1997a & 1997b) used the back of the head – reduced to a single character – and distinguished an ‘hypostomal bridge’ and a ‘genal bridge’. The detailed examination of the back of the head in the Eurytomidae (Lotfalizadeh et al. 2007), probable sister group of the Chalcididae, provided useful characters for their phylogeny and prompted GD to also investigate these characters in the Chalcididae. Chalcididae exhibit variable and puzzling structures that may be phylogenetically informative but request a thorough identification of homologies among the subfamilies and more largely with other families of Chalcidoidea.
    [Show full text]
  • Fauna Europaea: Hymenoptera – Apocrita (Excl
    Biodiversity Data Journal 3: e4186 doi: 10.3897/BDJ.3.e4186 Data Paper Fauna Europaea: Hymenoptera – Apocrita (excl. Ichneumonoidea) Mircea-Dan Mitroiu‡§, John Noyes , Aleksandar Cetkovic|, Guido Nonveiller†,¶, Alexander Radchenko#, Andrew Polaszek§, Fredrick Ronquist¤, Mattias Forshage«, Guido Pagliano», Josef Gusenleitner˄, Mario Boni Bartalucci˅, Massimo Olmi ¦, Lucian Fusuˀ, Michael Madl ˁ, Norman F Johnson₵, Petr Janstaℓ, Raymond Wahis₰, Villu Soon ₱, Paolo Rosa₳, Till Osten †,₴, Yvan Barbier₣, Yde de Jong ₮,₦ ‡ Alexandru Ioan Cuza University, Faculty of Biology, Iasi, Romania § Natural History Museum, London, United Kingdom | University of Belgrade, Faculty of Biology, Belgrade, Serbia ¶ Nusiceva 2a, Belgrade (Zemun), Serbia # Schmalhausen Institute of Zoology, Kiev, Ukraine ¤ Uppsala University, Evolutionary Biology Centre, Uppsala, Sweden « Swedish Museum of Natural History, Stockholm, Sweden » Museo Regionale di Scienze Naturi, Torino, Italy ˄ Private, Linz, Austria ˅ Museo de “La Specola”, Firenze, Italy ¦ Università degli Studi della Tuscia, Viterbo, Italy ˀ Alexandru Ioan Cuza University of Iasi, Faculty of Biology, Iasi, Romania ˁ Naturhistorisches Museum Wien, Wien, Austria ₵ Museum of Biological Diversity, Columbus, OH, United States of America ℓ Charles University, Faculty of Sciences, Prague, Czech Republic ₰ Gembloux Agro bio tech, Université de Liège, Gembloux, Belgium ₱ University of Tartu, Institute of Ecology and Earth Sciences, Tartu, Estonia ₳ Via Belvedere 8d, Bernareggio, Italy ₴ Private, Murr, Germany ₣ Université
    [Show full text]
  • The Taxonomy of the Side Species Group of Spilochalcis (Hymenoptera: Chalcididae) in America North of Mexico with Biological Notes on a Representative Species
    University of Massachusetts Amherst ScholarWorks@UMass Amherst Masters Theses 1911 - February 2014 1984 The taxonomy of the side species group of Spilochalcis (Hymenoptera: Chalcididae) in America north of Mexico with biological notes on a representative species. Gary James Couch University of Massachusetts Amherst Follow this and additional works at: https://scholarworks.umass.edu/theses Couch, Gary James, "The taxonomy of the side species group of Spilochalcis (Hymenoptera: Chalcididae) in America north of Mexico with biological notes on a representative species." (1984). Masters Theses 1911 - February 2014. 3045. Retrieved from https://scholarworks.umass.edu/theses/3045 This thesis is brought to you for free and open access by ScholarWorks@UMass Amherst. It has been accepted for inclusion in Masters Theses 1911 - February 2014 by an authorized administrator of ScholarWorks@UMass Amherst. For more information, please contact scholarworks@library.umass.edu. THE TAXONOMY OF THE SIDE SPECIES GROUP OF SPILOCHALCIS (HYMENOPTERA:CHALCIDIDAE) IN AMERICA NORTH OF MEXICO WITH BIOLOGICAL NOTES ON A REPRESENTATIVE SPECIES. A Thesis Presented By GARY JAMES COUCH Submitted to the Graduate School of the University of Massachusetts in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE May 1984 Department of Entomology THE TAXONOMY OF THE SIDE SPECIES GROUP OF SPILOCHALCIS (HYMENOPTERA:CHALCIDIDAE) IN AMERICA NORTH OF MEXICO WITH BIOLOGICAL NOTES ON A REPRESENTATIVE SPECIES. A Thesis Presented By GARY JAMES COUCH Approved as to style and content by: Dr. T/M. Peter's, Chairperson of Committee CJZl- Dr. C-M. Yin, Membe D#. J.S. El kin ton, Member ii Dedication To: My mother who taught me that dreams are only worth the time and effort you devote to attaining them and my father for the values to base them on.
    [Show full text]
  • Zootaxa, Fossil Eucharitidae and Perilampidae
    Zootaxa 2306: 1–16 (2009) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2009 · Magnolia Press ISSN 1175-5334 (online edition) Fossil Eucharitidae and Perilampidae (Hymenoptera: Chalcidoidea) from Baltic Amber JOHN M. HERATY1 & D. CHRISTOPHER DARLING2 1Department of Entomology, University of California, Riverside, CA, USA, 92521. E-mail: john.heraty@ucr.edu 2Department of Entomology, Royal Ontario Museum, Toronto, Ontario, Canada, M5S 2C6 and Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada, M5S 1A. E-mail: cdarling@rom.on.ca Abstract Palaeocharis rex n. gen. and sp. (Eucharitidae: Eucharitinae) and Perilampus pisticus n. sp. (Perilampidae: Perilampinae) are described from Baltic amber. Perilampus renzii (Peñalver & Engel) is transferred to Torymidae: Palaeotorymus renzii n. comb. Palaeocharis is related to Psilocharis Heraty based on presence of one anellus, linear mandibular depression, dorsal axillular groove, free prepectus and a transverse row of hairs on the hypopygium. This fossil is unique in comparison with extant Chalcidoidea because there are two foretibial spurs instead of a single well- developed calcar. Perilampus pisticus is placed into the extant Perilampus micans group because the frenum and marginal rim of the scutellum are visible in dorsal view and the prepectus forms a large equilateral triangle. The phylogenetic placement of both genera is discussed based on an analysis of both a combined morphological and molecular (28S and 18S) and morphology-only matrix. Morphological characters were used from an earlier study of Eucharitidae (Heraty 2002), with some characters revised to reflect variation in Perilampinae. Baltic amber is of Eocene age, which puts the age of divergence of these families at more than 40 mya.
    [Show full text]
  • LIST of PRIVATE LABS APPROVED by STATE for COVID TESTING AS on 21-08-2020 Cost of Tests Fixed by Government of Kerala in Private Sector
    LIST OF PRIVATE LABS APPROVED BY STATE FOR COVID TESTING AS ON 21-08-2020 Cost of tests fixed by Government of Kerala in Private Sector. TYPE OF RESULT RATE( Inclusive of Tax) TEST RT-PCR CONFIRMATORY Rs 2750/- OPEN CBNAAT CONFIRMATORY Rs 3000/- TRUENAT If STEP1 is positive, require step 2 for confirmation STEP 1- Rs 1500/- STEP 1 negative is confirmatory STEP2- Rs1500/-( required only if STEP1 turns positive) ANTIGEN Positive results are confirmatory. Rs 625/- Negative results in a symptomatic person require + cost of further RT-PCR/CBNAAT/TRUENAT test RT- PCR/CBNAAT/TRUENAT if required Private Labs approved for RT-PCR open system 1. DDRC SRL Diagnostics Pvt Ltd, Panampilly Nagar, Ernakulam 2. MIMS Lab Services, Govindapuram, Kozhikode 3. Lab Services of Amrita Institute of Medical Sciences & Research Centre, AIMSPonekkara, Kochi 4. Dane Diagnostics Pvt Ltd, 18/757 (1), RC Road, Palakkad 5. Medivision Scan & Diagnostic Research Centre Pvt Ltd, Sreekandath Road, Kochi 6. MVR Cancer Centre & Research Institute, CP 13/516 B, C, Vellalaserri NIT (via), Poolacode, Kozhikode 7. Aza Diagnostic Centre, Stadium Puthiyara Road, Kozhikode 8. Neuberg Diagnostics Private Limited, Thombra Arcade, Ernakulam 9. Jeeva Specialty Laboratory, Thrissur 10. MES Medical College, Perinthalmanna, Malappuram Private Labs approved for XPERT/CBNAAT Testing 1. Amrita Institute of Medical Science, Kochi 2. Aster Medcity, Aster DM Healthcare Ltd, Kutty Sahib Road, Kothad, Cochin 3. NIMS Medicity, Aralumoodu, Neyyattinkara, Thiruvananthapuram 4. Rajagiri Hospital Laboratory Services, Rajagiri Hospital, Chunangamvely, Aluva 5. Micro Health LAbs, MPS Tower, Kozhikode 6. Believers Church Medical College Laboratory, St Thomas Nagar, Kuttapuzha P.O., Thiruvalla 7.
    [Show full text]