Revision of the Australian Clivinini 3. the Procera and Elegans Groups of the Genus Clivina Latreille

SPIXIANA 40 2 197-304 München, Dezember 2017 ISSN 0341-8391

Revision of the Australian Clivinini 3. The procera and elegans groups of the genus Clivina Latreille

(Coleoptera, Carabidae, Scaritinae)

Martin Baehr

Baehr, M. 2017. Revision of the Australian Clivinini 3. The procera and elegans groups of the genus Clivina Latreille (Coleoptera, Carabidae, Scaritinae). Spixiana 40 (2): 197-304. As the third part of a revision of the Australian species of the scaritine tribe Clivinini the procera and elegans groups of the genus Clivina Latreille are revised. To maintain stability of nomenclature, lectotypes and for certain species also paralectotypes are designated for the following nominal species: Clivina abbreviata (Putzeys), C. brevisterna major Sloane, C. foveiceps (Macleay), C. froggatti Sloane, C. macleayi Sloane, C. marginata (Putzeys), C. monilicornis Sloane, C. nyctosyloides Putzeys, C. obliquicollis Sloane, C. oblonga (Putzeys), C. prominens Putzeys, C. regularis Sloane, C. robusta Sloane. Clivina prominens Putzeys, 1866 is synonymized with C. procera Putzeys, 1866. Clivina brevisterna major Sloane, 1917 is resurrected as a separate taxon from synonymy with C. brevisterna Sloane, 1916 and is raised to specific status. Clivina interstitialis Sloane, 1896 is renamed C. propinqua, nom. nov., because the name interstitialis is a junior homonym of C. interstitialis Kolbe, 1883. Clivina foveiceps (Macleay, 1863) is renamed C. macleayana, nom. nov., because the name foveiceps is a junior homonym of C. foveiceps Putzeys, 1846. Some new species belonging to groups mentioned in the 2nd part of the revision, which were sampled too late to be included in that part, are added to this 3rd part. Following new taxa are described in this paper: bankae, biseriata, carnabyi, cobourgiana, conicollis, cooindae, crassipennis, dubia, foveifrons, gemina, gemina nigripes, gerstmeieri planior, glabripennis, goldingi, gracilipes longior, heros, horaki, horneri, incurvicollis, infans, inopinata, interposita, laevigata, mahoni, micans, montisbelli, moretona, newcastleana, nitescens, normanbyensis, ovalior, pachysoma, platynota, profundestriolata, rectipennis, rugosifrons, ryaceki, semirubra, sinuicola, subrufipes, thoracica, uncinata, variseta, victoriae, vixsulcata, windjanae. A key to the species of the procera and elegans species groups of Clivina s. l. is provided. For the additional species relevant parts of the key in part 2 of the revision are repeated and extended to cover the new species. Male genitalia are figured. Female genitalia are only figured of certain groups and subgroups. A checklist is provided for the Australian Clivinini mentioned in this third paper, including notes on the recorded ranges. Clivina bulirschi Baehr, 2015 is renamed C. bulirschiana, because Dostal also described a Clivina (Pjysoclivina) bulirschi, 2015 in a paper (Dostal 2015) which was printed a month earlier than C. bulirschi Baehr. Martin Baehr, SNSB – Zoologische Staatssammlung München, Münchhausenstr. 21, 81247 München, Germany; e-mail: [email protected]

197 Introduction of prothorax and hind body. Also measurements of the protibia can rather vary according to the age of the This third part of the revision of the Australian specimen and the degree of abrasion of the external Clivinini covers two related species groups of com- teeth. monly large or very large, or at least bulky, species. Presence, or structure, of punctation and/or microreticulation of head, pronotum, elytra, and abdomen, or Style and format of descriptions etc. are similar to of one of these, are important characters for species those in the first and second parts of the revision. definition and distinction. For unambigeous recognition Any general information about way of life, char- of the commonly very faint or superficial microreticula- acters, collections, and relevant literature may be tion a binocular microscope with a bright lamp is taken from these parts which also include the key needed which should offer at least 64 ×, better 100 × to genera, subgenera, and species groups. magnification. Data of examined material in full length with exact labelling, including sex, all ciphers, notes of determiners Material and methods and curators, and printed labels, are given in most species, except some common and well known ones. Altogether 2497 specimens of the species mentioned in Original spelling of date of collecting, especially of the the present part have been examined. Apart from mate- month (arabic, roman, abbreviations), was generally rial borrowed from several museum and private collec- used. An / with a blank before and after it denotes tions, these include the very rich collections made in another label. recent times by S. Bilý and S. Jakl (Prague), A. Dostal Records not specified as to a state are included (Vienna), and by myself in the northern parts of North- under the heading ‘AUS’, as well as specimens bearing ern Territory, Western Australia, and Queensland. only ‘Australia’, or those without any labelling. States In spite of the amount of material, several species are recorded as under ‘abbreviations’. State records or are still represented by single or very few specimens or data like ‘Australia’, ‘Nov. Holl.’ etc. have not been were available from just a single locality. Additional repeated for non-type material. systematic sampling using light traps and pitfall traps in many localities throughout northern and inland Aus- Abbreviations of collections mentioned tralia would be needed to get a more authentic impres- in the present part of the revision sion of the actual number and distribution of the species. Methods of dissections and descriptions are similar AMS Australian Museum, Sydney to those used in parts 1 and 2 of the revision (Baehr 2008, ANIC Australian National Insect Collection, Can- 2015). However, length of pronotum has been measured berra (including the collection of B. P. Moore) from the most advanced part of the apex to the most ANIC-MMS W. Macleay Collection in Australian Na- advanced part of the base. To avoid overburding de- tional Insect Collection, Canberra scriptions with repetitions of similar character states, CBM Working Collection M. Baehr in Zoologische those states that are uniform within a group or subgroup Staatssammlung, München have been mentioned in the diagnosis of the group but CBP Collection P. Bulirsch, Praha were omitted from the descriptions of the species. These CDW Collection A. Dostal, Wien (including the Karel mainly cover certain characters of mouth parts, struc- Kult Collection) ture of prothorax and elytra, legs, and female gonocox- CGT Collection P. M. Giachino, Torino ites. CHA Collection P. Hudson, Adelaide Distinction of sexes in the procera and elegans groups CHP Collection L. Hovorka, Praha is easier than in most other groups, because in the males CJP Collection S. Jakl, Praha of all species the internal spur of the protibia is widened CKZ Collection V. Kabourek, Zlin at its apex, or it is at least not as acute as in females. CMP Carnegie Museum of Natural History, Pitts- This is particularly useful, because due to the deep black burgh colour of most species and to the generally very strong CNCI Agriculture Canada, Ottawa sclerotization, genitalia are not translucent when seen CSM Collection R. Sciaky, Milano from below, as in many of the smaller, more delicate, CTV Collection L. Toledano, Verona and less sclerotized species worked in previous parts of CWB Collection D. Wrase, Berlin the revision. Moreover, males of several species possess DEI Deutsches Entomologisches Institut, Münche- narrower, dorsally more convex protibiae with smaller berg external teeth than the females, which have more de- FMNH Field Museum of Natural History, Chicago pressed, more strongly toothed protibiae. In several IRSNB Institut Royal des Sciences Naturelles, Bruxelles species also the male metatarsus is narrower and MCNV Museo Civico di Storia Naturale, Venezia longer than that of the females. MCSN Museo Civico di Storia Naturale, Genova In many species of the procera and elegans groups, MCZ Museum of Comparative Zoology, Cambridge/ differences in body size are considerable, and this is Mass. commonly combined with some allometric proportions MMS Macleay Museum, Sydney

198 MNHP Museum National d’Histoire Naturelle, Paris Taxonomy NHM The Natural History Museum, London NHMB Naturhistorisches Museum, Basel Identification of groups NMNL National Museum of Natural History Natura- lis, Leiden A key to the species groups of the Australian Clivina NMPC National Museum of Natural History, Praha is in both previous parts of this revision. However, NMV Museum of Victoria, Melbourne for easier use, an extract from this key is repeated NTD Museum and Art Gallery of the Northern Ter- ritory, Darwin below. Both species groups mentioned in the present OUM Oxford University Museum, Oxford part have the setae at the terminal abdominal ster- QDPIB Queensland Department of Primary Industries, num widely distant and have at most 3 striae ended Brisbane free at the base of the elytra. They are distinguished QDPIM Queensland Department of Primary Industries, as following: Mareeba QMB Queensland Museum, Brisbane 25. Excision of anterior margin of clypeus more or SAMA South Australian Museum, Adelaide less deep, quadrangular (Figs 78, 79); body size SMNS Staatliches Museum für Naturkunde, Stuttgart commonly large, > 10 mm, or if small, body short SMTD Staatliches Museum für Naturkunde, Dresden and compact...... 26. UASM Strickland Museum, Edmonton – Excision of clypeus regularly concave; body size USNM United States National Museum, Washington, usually smaller, < 10 mm, body usually elongate D.C. and rather parallel-sided...... 28. WADAK Western Australia Department of Agriculture, Kununurra 26. Excision of anterior margin of clypeus shallow; WADAP Western Australia Department of Agriculture, eye semicircular, laterad markedly protruded; Perth head dorsally with a transverse ridge on middle WAM Western Australian Museum, Perth of frons and a large depression behind; hind ZMHB Museum für Naturkunde der Humboldt- Universität, Berlin body elongate, parallel; protibia short and wide, ZMUA Zoölogisch Museum of the University of Am- with 4 distinct, stout teeth; aedeagus with dense, sterdam elongate spines in the internal sac...... ZSM Zoologische Staatssammlung, München ...... impressiceps group – Excision of anterior margin of clypeus deep or Abbreviations not; but if hind body elongate and parallel, exci- ACT Australian Capital Territory sion deep; in parallel-sided species eye usually AUS Australia laterad less protruded; head without a similar CYP Cape York Peninsula transverse ridge and depression; protibia narrow KID Kimberley Division and elongate, with 3 rather delicate teeth; body NP National Park shape various; aedeagus usually without dense, NSW New South Wales elongate spines in the internal sac...... 27. NT Northern Territory QLD Queensland 27. Hind body elongate, almost parallel-sided; met SA South Australia episternum elongate, metathoracic wings long; VIC Victoria excision of anterior margin of clypeus always WA Western Australia deep, about quadrangular (Fig. 78); aedeagus c. central never with arrow-head shaped, or club-shaped, e. eastern or claw shaped apex (Figs 1-23)...... ec. eastern central ...... procera group n. northern nc. northern central – Hind body various but commonly shorter and ne. north-eastern not parallel-sided; metepisternum commonly nw. north-western short and metathoracic wings commonly short- s. southern ened; excision of anterior margin of clypeus sc. southern central variously shaped but usually rather shallow se. south-eastern (Fig. 79); aedeagus with arrow-head shaped, or sw. south-western club-shaped, or claw-shaped apex (Figs 24- w. western 53)...... elegans group > larger or longer than < smaller or shorter than

199 Characters inserted slightly inside of the basal curvature. Basal angle evenly rounded. Prosternum usually not visible Particularly the species of the procera group are at the basal angle. Prosternal process coarsely margined, generally very similar in their external characters, short, asetose at apex. Proepisternum usually striolate even when minor differences may be present in body and microreticulate, in some species also punctate. Elytra: Base smooth without distinct knobs except size, proportions of prothorax and elytra, size of eye, in middle. 3rd stria almost always 4-setose, usually with emargination of clypeus, and structure of surface large and distinct punctures. Scutellary stria indistinct, (impressions on the head surface, striation of elytra, extremely short, situated close to suture. Lateral margin punctation, microreticulation). In certain species, not crenulate. however, even in some sympatric or syntopic ones, Legs: Profemur large, upper surface convex, lower the differences in body shape and external structure surface somewhat depressed, but not margined. Proti- are so minimal that their distinction on the basis of bia elongate, 3-dentate, with comparatively short teeth external characters is very difficult. Hence, in those which usually are shorter in males than females. Lower species examination of the male aedeagus is the best surface depressed and smooth, with a group of elongate if not the sole tool for species distinction. Although setae near base. Mesotibia with more or less elongate, the aedeagus in most species is likewise quite similar preapical spur on the dorsal surface. Meso- and metatibiae sparsely setose. Lower surface of 1st tarsomeres shaped, it may exhibit obvious differences, e. g. in usually with a dense fringe of setae on both sides. size, width, curvature, and shape of the apex, while Male genitalia: Genital ring very uniform through- the structure of the internal sac, e. g. denticulation out, more or less oval-shaped. Aedeagus in basal half of folds and number and/or position of these folds, or two thirds with two sclerotized rods; orificium usu- is very similar. ally very elongate, slightly asymmetrically situated on Also the female gonocoxites are extremely similar the upper left surface, and in apical half open on the and differ usually only slightly in relative length of upper side. Internal sac with three denticulate and more gonocoxite 2 and in the number of setae on the gono- or less strongly sclerotized folds. coxites and the lateral plate, if they differ at all. Moreo- Female genitalia: Gonocoxites 1 and 2 usually com- ver, number and arrangement of the setae may vary to bined without a distinct border; laterally depressed and some extent within species, so that the gonocoxites are with a sharp edge along the medio-ventral margin. very difficult to use and usually do not allow unam- Arrangement of the bursa and other not sclerotized biguous identification of species. This implies that in parts of the genitalia usually as in fig. 236 of part 2. some species of very similar shape and structure, in All species of the procera group are fully winged, in particular when they are sympatric, identification of several species of the elegans group the metathoracic females and their affiliation to the appropriate species wings are abbreviated. Therefore the metepisternum is is difficult or even impossible by use of morphological elongate in the procera group, but usually shortened at characters. In those species, taxonomic methods using different degrees in the elegans group. molecular analysis may reveal a better distinction of the species, in particular of females. Some character states are very uniform in all species Key to the species of the procera and elegans worked in the present part. For not overburding the groups of the genus Clivina descriptions with repetitions, a number of characters are not mentioned in the descriptions, except when they deviate from the common condition. These are: Note. The new species not belonging to the procera Head: Paraorbital ridge usually elongate, convex, and elegans groups are inserted into the most recent paraorbital sulcus inside of paraorbital ridge elongate, existing keys at the end of the descriptive part. Posi- shallow, linear. Supraantennal plates laterally obliquely tion of folds or sclerites in the aedeagus are given in convex, coarsely margined. Labrum usually straight and the inverted state of the internal sac. 5- or 7-setose, but in some species the number of setae deviates. Mandible usually short, evenly curved. Only 1. Metepisternum elongate, metathoracic wings in few species elongate and in apical half narrow. Men- long and clypeal excision always deep, quad- tum with distinct, obtusely triangular tooth, bisetose rangular (Fig. 78) and hind body elongate, almost near tooth and also at base, submentum quadrisetose. parallel-sided and aedeagus never with arrow- Glossa bisetose, paraglossae fused to glossa, barely head shaped, club-shaped, claw-shaped, or surpassing the glossa. Lacinia with strong teeth, a row spatulate apex (Figs 1-23)...... 2. of long setae on the lower ventral surface, and a dense – Not all these characters combined; aedeagus tuft of setae on the dorsal surface. Maxillary and labial palpi elongate, barely widened apicad, impilose. An- almost always (except C. robusta Sloane) with tenna short, median and apical antennomeres about arrow-head shaped, or club-shaped, or claw- quadrate, densely pilose from 3rd antennomere. shaped, or spatulate apex (Figs 24-53), or aedea- Prothorax: Lateral margin and channel of prono- gus unknown...... 25. tum usually very narrow; the posterior marginal seta

200 2. All legs black or almost so...... 3. 9. Surface of pronotum and elytra markedly glossy; – Middle and hind legs rufous, distinctly paler prothorax rather quadrate, ratio base/apex than the anterior leg, or all legs unicolourous < 1.24; prothorax wider in relation to head, ratio rufous to rufo-piceous...... 12. prothorax/head > 1.42; apical part of aedeagus markedly curved left, not sinuate, at tip not in- 3. Body size very large, > 18 mm; lateral margin of curved (Fig. 18). w.QLD, ce.NT...... pronotum considerably convex (Fig. 124); both ...... micans, spec. nov. parameres in front of the narrow apical part – Surface of pronotum and elytra far less glossy; abruptly quadrangular (Fig. 14). n.WA: KID, prothorax less quadrate, ratio base/apex > 1.27, adjacent nw.NT...... heros, spec. nov. usually more; prothorax narrower in relation to – Body size at the average smaller, usually < 18 mm head, ratio prothorax/head < 1.39; apical part of (rarely, in C. ryaceki, spec. nov. up to 19 mm, but aedeagus less curved left, or sinuate, or at tip then lateral margin of pronotum almost straight); incurved and crotched (Figs 2, 10, 15). n.NT, lateral margins of pronotum more or less oblique nw.WA: sw.KID...... 10. but rather straight (Fig. 123); both parameres in front of the narrow apical part more gently slop- 10. Body size larger, length 16.5 mm; aedeagus re- ing (Fig. 13). n.NT, n.WA: KID...... 4. markably sinuate, tip acute but not incurved (Fig. 15). n.NT...... goldingi, spec. nov. 4. Proepisternum more or less densely and coarse- – Body size smaller, length < 13.5 mm; aedeagus ly punctate...... 5. not sinuate, tip either acute and incurved, or – Proepisternum not or but feebly punctate...... 6. obtuse and straight (Figs 2, 10)...... 11. 5. Body size < 14 mm; proepisternum densely 11. Legs deep black; apex of aedeagus elongate, at punctate; apical part of aedeagus markedly tip acute and incurved (Fig. 10). n.WA: asymmetric, apex narrow, at tip curved down sw.KID...... windjanae, spec. nov. or even crotched (Fig. 8). n.WA: s. + w.KID ...... – Legs usually less deep black; apex of aedeagus ...... gemina nigripes, subspec. nov. shorter, at tip more or less obtuse and not or – Body size > 15.5 mm; proepisternum less dis- barely incurved (Fig. 2). n. + nw.NT...... tinctly punctate; apicial part of aedeagus sym- ...... obscuripes Blackburn, 1890 metric, apex rather wide, straight and at tip obtuse (Fig. 11). nw.NT...... mahoni, spec. nov. 12. Head with several distinct transverse sulci (Fig. 126); legs uniformly rufo-piceous; apex of 6. Prothorax elongate, ratio l/w > 1.06, and strong- aedeagus short, rather straight, at tip fairly acute ly conical (Fig. 127)...... 7. but not incurved (Fig. 19). n.QLD, ne.NT...... – Prothorax shorter, ratio l/w < 1.06, and far less ...... rugosifrons, spec. nov. conical (Fig. 81); if ratio > 1.04, surfaces of pro- – Head without perceptible transverse sulci; legs notum and elytra markedly glossy...... 8. variously coloured; aedeagus variously 7. Prothorax shorter, ratio l/w < 1.09; elytra short- shaped...... 13. er, ratio l/w < 2.02; apex of aedeagus shorter and 13. Elytra extremely elongate, ratio l/w 2.22; labrum wider (Fig. 20); labrum 5-setose. n.QLD, n. NT. 7-setose; eye laterad markedly protruded ...... monilicornis Sloane, 1896 (Fig. 84); head and pronotum finely but densely – Prothorax longer, ratio l/w > 1.10; elytra longer, punctulate; aedeagus unknown. n.WA: w.KID. ratio l/w > 2.06; apex of aedeagus longer and ...... montisbelli, spec. nov. narrower (Fig. 21); labrum usually 7-setose. – Elytra shorter, ratio l/w < 2.12; labrum 5-setose; n.QLD: CYP, n. NT...... conicollis, spec. nov. eye laterad less protruded (Figs 83, 85, 122); head 8. Anterior part of frons with distinct transverse and pronotum usually less densely punctulate; sulci; body size 15.0-16.5 mm; aedeagus wide, aedeagus variously shaped...... 14. apex deeply bisinuate at the left side and strong- 14. Pronotum and elytra remarkably glossy, without ly turned left, at tip not incurved (Fig. 9). n.WA: microreticulation; apical part of aedeagus mark- s. + w.KID...... carnabyi, spec. nov. edly curved down, on the left side with deep, – Anterior part of frons without transverse sulci; semicircular excision, apex straight, at tip obtuse body size varied; aedeagus varied (Figs 2, 10, 15, (Fig. 17). sw.QLD, ne.SA...... 18)...... 9...... glabripennis, spec. nov.

201 – Pronotum and/or elytra with distinct microre- – Head sparsely punctate; apical part of aedeagus ticulation, less glossy; aedeagus differently markedly asymmetric, apex narrow, at tip curved shaped (Figs 1, 3-7, 12, 16, 22, 23). Distribution down or even crotched (Fig. 7). nw.NT, ne.WA: different...... 15. ne.KID...... gemina gemina, spec. nov. 15. Pronotum elongate and considerably narrowed 20. Comparatively small species, length 12.8 mm; apicad, ratio l/w 1.11 (Fig. 122); all legs rufous, prothorax fairly elongate and slightly conical, anterior leg not darker than middle and hind ratio l/w 1.05; eye lateral little produced, orbit 1 legs; apex of aedeagus elongate, strongly sinuate, c. /3 of length of eye (Fig. 128); apical part of tip acute and markedly curved down (Fig. 6). aedeagus very asymmetric, but apex short, nw.QLD...... thoracica, spec. nov. straight, at tip obtuse and not incurved (Fig. 22). – Pronotum shorter and usually less narrowed AUS: locality not noted...... dubia, spec. nov. apicad, ratio l/w usually < 1.05 (Figs 80, 83, 125, – Size various, prothorax various, but less conical; 128), if up to 1.11, body size < 12 mm and body eye laterad well protruded, orbit short (Figs 80, depressed; either all legs uniformly rufous or 83, 125); aedeagus different (Figs 1, 3, 12, 13, anterior leg perceptibly darker than middle and 16)...... 21. hind legs; apex of aedeagus variously shaped, 21. Body size very large, length 15.2-18.9 mm; elytra but when all legs uniformly coloured, apex of rather elongate, ratio l/w 2.06-2.12; lower surface aedeagus shorter and obtuse at tip (Figs 1, 3-5, of aedeagus very concave, apical part almost 7, 12, 13, 16, 22, 23)...... 16. symmetrically triangular, acute, at tip not in- 16. All legs uniformly rufous or rufo-piceous; ratio curved (Fig. 13). n. + nw.NT, ne.WA: ne.KID. .... l/w of prothorax < 1.01; elytra parallel-sided; ...... ryaceki, spec. nov. aedeagus with rather short, not incurved apex – Body size various but usually smaller, if (Fig. 5). nw.QLD, ne.NT. .... sinuicola, spec. nov. > 15.0 mm, aedeagus on the left side deeply – Middle and hind legs distinctly paler than the bisinuate and species from eastern Australia; front leg; apex of aedeagus usually longer, either elytra various; lower surface of aedeagus less more sinuate and less obtuse, or narrow, paral- concave or bisinuate, apical part either deeply lel, with the tip suddenly bent down (Figs 1, 3, bisinuate or narrow and at tip markedly incurved 4, 7, 12, 13, 16, 22, 23); if apex short and not or crotched (Figs 1, 3, 12, 16)...... 22. sinuate, ratio l/w of prothorax > 1.05 and elytra 22. Prothorax generally slightly longer, ratio l/w slightly oval-shaped (Fig. 87)...... 17. > 1.02; lower surface of aedeagus regularly con- 17. Prothorax rather elongate, ratio l/w > 1.05 and cave; left side deeply bisinuate, apex barely in- elytra slightly oviform (Fig. 87) and protibia curved, not crotched (Fig. 1). e.SA, VIC, NSW, rather short, ratio in male < 2.85, in female < 2.5 e.QLD...... procera Putzeys, 1866 and apical part of aedeagus not curved down, – Prothorax generally slightly shorter, ratio l/w barely curved left, rather short and obtuse, at tip < 1.02; lower surface of aedeagus either regu- not incurved and parameres asetose (Fig. 23). larly concave but apex very narrow, or lower ne.NSW, se.QLD...... regularis Sloane, 1896 surface bisinuate; apex always crotched; left side – Not all character states combined; if from south- bisinuate or not (Figs 3, 12, 16). n. + nc. + nw. ern or south-eastern Australia, elytra parallel- NT...... 23. sided, protibia longer, ratio l/w in male > 4.0, in 23. Pronotum slightly shorter and more quadrate, female > 3.3; parameres unisetose...... 18. ratio l/w < 1.0 (Fig. 125); apex of aedeagus on 18. Proepisternum distinctly punctate; aedeagus as left side deeply bisinuate; parameres bisetose in Figs 4, 7...... 19. (Fig. 16). nc.NT...... newcastleana, spec. nov. – Proepisternum not punctate or only in lower – Pronotum usually slightly longer and less quad- part indistinctly punctate; aedeagus as in Figs 1, rate, ratio l/w usually > 1.0 (Figs 80, 83); apex of 3, 12, 13, 16, 22...... 20. aedeagus on left side not deeply bisinuate, either lower surface regularly concave (Fig. 3), or api- 19. Head rather densely punctate; aedeagus with cal part conspicuously incurved (Fig. 12); para- rather wide, almost regularly triangular apex, at meres asetose. n. + nw.NT...... 24. tip not incurved or crotched (Fig. 4). n.NT, n.QLD...... cooindae, spec. nov. 24. Elytra slightly longer, ratio l/w 2.04-2.08; lower surface of aedeagus regularly concave, apex narrow (Fig. 3). n.NT...... subrufipes, spec. nov.

202 – Elytra slightly shorter, ratio l/w 2.0-2.05; lower 33. 3rd, 5th, and 7th elytral intervals each with > 10 surface of aedeagus slightly bisinuate, apical part punctures and setae (Figs 92, 135); elytra soldered remarkably incurved (Fig. 12). nw. NT...... together; aedeagus unknown. n.NT: Kakadu ...... victoriae, spec. nov. NP...... triseriata, spec. nov. rd 25. Lateral margin of elytra widely yellow (Fig. 95); – Only 3 elytral interval with discal setae. .... 34. aedeagus with distinctly arrow-shaped apex, 34. 3rd interval with 5-7 discal punctures and setae; without serrate fold in the internal sac (Fig. 30). elytra free, but metathoracic wings almost com- n.QLD, n.NT, n.WA: KID...... pletely reduced; aedeagus narrow, with elongate ...... marginata (Putzeys, 1868) apical part and small, convexly arrow-shaped – Elytra unicolourous; aedeagus variously shaped. apex (Fig. 27). n.QLD...... variseta, spec. nov...... 28. – 3rd interval with 4 discal punctures and setae; 28. Metepisternum elongate, c. 2 × as long as wide elytra and aedeagus variously shaped...... 35. at apex, metathoracic wings long; hind body 35. Mandibles slender and elongate, pincer-like elongate, parallel-sided or almost so...... 29. (Fig. 94); aedeagus with narrow, gently knobbed – Metepisternum shortened, < 1.5 × as long as wide apex (Fig. 29)...... 36. at apex, metathoracic wings commonly abbrevi- – Mandibles short and wide; aedeagus variously ated; hind body not parallel-sided, more or less shaped...... 37. oval shaped and commonly shorter...... 33. 36. Elytra shorter, ratio l/w < 1.80; aedeagus see 29. Head with several irregularly transverse sulci, Fig. 29. nw.QLD, n.NT, ne.WA: ne.KID...... very densely punctate (Fig. 98, 199, 131); aedea- ...... gracilipes gracilipes Sloane, 1896 gus with short and wide, arrow-shaped apex – Elytra longer, ratio l/w >1.88; aedeagus un- (Figs 34, 35)...... 30. known. n.QLD: c. + n.CYP...... – Head without distinct transverse sulci, less ...... gracilipes longior, subspec. nov. densely punctate (Fig. 97, 132); aedeagus variously shaped (Figs 33, 36)...... 32. 37. Eye protruded but rather small, orbit large, oblique (Figs 114-116) and length < 8.8 mm and 30. Elytra longer, ratio l/w > 2.06; pronotum weak- labrum 5-setose; aedeagus with more or less ly microreticulate; aedeagus see Fig. 34; or un- elongate, claw-shaped apex (Figs 46-48)...... 38. known. nc. + sc.NT, ec.WA...... 31. – Eye large, orbit small (Figs 93, 100, 101, 104-113, – Elytra shorter, ratio l/w < 2.02; pronotum more 129) or size much larger or labrum 7-setose; strongly microreticulate; aedeagus see Fig. 35. aedeagus different, never claw-shaped (Figs 24- nw.WA...... incurvicollis, spec. nov. 26, 28, 31, 32, 37-45)...... 40. 31. Colour black; head with much denser, but finer 38. Smaller species, length < 6.5 mm; aedeagus with punctation; prothorax laterally less convex very narrow, elongate apical part and with small, (Fig. 131); aedeagus with obtusely triangular, claw-shaped apex; parameres conspicuously almost club-shaped apex (Fig. 34). nc.NT...... longitudinally striped (Fig. 46). n.NT...... bankae, spec. nov...... darwini Sloane, 1916 – Colour pale red; head with far less dense, but – Larger species, length > 7.5 mm; aedeagus with coarser punctation; prothorax laterally mark- shorter claw-shaped apical part; parameres not edly convex (Fig. 99); aedeagus unknown. sc.NT, longitudinally striped (Figs 47, 48). n.NT. .... 39. ec.WA...... rectipennis, spec. nov. 39. Temporal ridge and basal margin of clypeus both 32. Pronotum dorsally convex; both, pronotum and markedly raised, frons in middle with a longi- elytra, with fine but distinct microreticulation; tudinal groove (Fig. 115); prothorax slightly elytra shorter, ratio l/w < 2.0 (Fig. 97); aedeagus shorter, ratio l/w > 0.98; apex of aedeagus with triangular, arrow-shaped apex (Fig. 33). straight, parameres less stout (Fig. 47)...... nw.WA: KID...... obliquicollis Sloane, 1905 ...... macleayi Sloane, 1896 – Pronotum dorsally depressed; both, pronotum – Temporal ridge and basal margin of clypeus both and elytra, with very superficial microreticula- far less raised, frons in middle without longitu- tion, glossy (Fig. 132); elytra longer, ratio l/w dinal groove (Fig. 116); prothorax slightly long- > 2.10; aedeagus with slightly asymmetric, spat- er, ratio l/w > 1.02; apex of aedeagus strongly ulate apex (Fig. 36). nw.WA: Pilbara...... turned left, parameres stout (Fig. 48)...... platynota, spec. nov...... horneri, spec. nov.

203 40. Eye more or less large, but not hemispherical 45. Elytra generally slightly shorter and more and suddenly protruded, orbit short but always evenly oval-shaped, with narrower base, ratio visible (Figs 88-90, 96, 102, 103, 130)...... 41. l/w 1.94-2.02 (Fig. 89); ratio base/apex of pro- – Eye large, hemispherical and suddenly pronotum < 1.45; aedeagus slightly smaller, length truded, orbit extremely short, barely visible < 2.5 mm (Fig. 25). Area around Heathlands, far (Figs 93, 100, 101, 104-113, 129)...... 47. northern CYP...... interposita, spec. nov.

1 – Elytra generally longer and less evenly oval- 41. Eye rather small, orbit large and oblique, > /3 as shaped, with wider base, ratio l/w 2.03-2.11 long as eye, head markedly triangular (Figs 96, (Fig. 90); ratio base/apex of pronotum > 1.50; 131); aedeagus either with sinuate, at tip ob- aedeagus slightly larger, length > 2.7 mm tusely angulate apex (Fig. 32), or with short and (Fig. 26). Iron Range, ce. CYP...... wide, spatulate apex (Fig. 31). e.VIC, e.NSW, ...... kershawi Sloane, 1916 se.QLD...... 42. – Eye larger, orbit small and less oblique, at most 46. Elytra generally longer, more narrowed towards th th 1 /4 as long as eye, head not markedly triangular base, ratio l/w 1.96-2.07; 5 -7 striae weak or (Figs 88-90, 102, 103); aedeagus always with absent (Fig. 102); aedeagus with short, club- arrow-shaped or club-shaped apex (Figs 24-26, shaped apex (Fig. 39). n.NT around Darwin...... 39, 40)...... 43...... mastersi Sloane, 1896 – Elytra generally shorter, less narrowed towards 42. Generally larger species, length 12.5-17.2 mm, base, ratio l/w 1.93-1.98; 5th-7th striae distinct usually > 14 mm; eye smaller, orbit larger, c. half (Fig. 103); aedeagus with asymmetrically spatu- as long as eye (Fig. 130); aedeagus with sinuate, late apex (Fig. 40). n.NT: Cobourg Pen...... at tip obtusely angulate apex (Fig. 32). e.VIC, ...... cobourgiana, spec. nov. e.NSW...... robusta Sloane, 1905 – Generally smaller species, length 10.7-14.1 mm, 47. Labrum 5-setose and body size > 12.5 mm and 1 and usually < 13 mm; eye larger, orbit smaller, c. /3 elytra longer, ratio l/w > 1.75 elytra not as long as eye (Fig. 96); aedeagus with short and markedly oviform and striae but moderately wide, spatulate apex (Fig. 31). ne.NSW, deep and finely punctate (Fig. 93, 129)...... 48. se.QLD...... oblonga (Putzeys, 1873) – Labrum usually 7-setose; if 5-setose, either body size small, < 8 mm, or elytra shorter, ratio l/w 43. Elytra oblong-oval, considerably narrowed to- < 1.73 and markedly oviform and striae very wards base (Figs 88-90), free but metathoracal deep and very coarsely punctate (Figs 100, 101, wings reduced; aedeagus with narrow apical 104-113)...... 49. part and small, club-shaped apex; internal sac with a sclerotized and serrate fold (Figs 24-26). 48. Prothorax longer, less conical, ratio l/w 1.0 n.QLD...... 44. (Fig. 129); microreticulation of pronotum fine – Elytra slightly oval, but little narrowed towards but distinct, transverse striolation very weak; base (Figs 102, 103), soldered together; aedeagus elytra longer, ratio l/w 1.88; aedeagus with wide, either with large and short, club-shaped apex or spatulate but at tip convex apex (Fig. 28). with asymmetrically spatulate apex; internal sac ne.QLD...... inopinata, spec. nov. without a sclerotized and serrate fold (Figs 39, – Prothorax shorter, more conical, ratio l/w < 0.94 40). n.NT...... 46. (Fig. 93); microreticulation of pronotum barely perceptible, transverse striolation coarse; elytra 44. Aedeagus larger, length > 3.0 mm, with wide, shorter, ratio l/w < 1.80; aedeagus unknown. spade-shaped apex that is deeply separated from c. + sw.QLD...... profundestriolata, spec. nov. the preapical stalk, and with a strongly sclerotized and serrate fold in the internal sac 49. Labrum 5-setose; body length 7.7 mm; prothorax (Fig. 24); elytra elongate, ratio l/w > 2.02. n.QLD: rather elongate and but moderately conical, ratio tip of CYP north of Jardine River, and Torres l/w 1.06; elytra moderately oviform, ratio l/w Strait islands...... elegans Putzeys, 1862 1.83; striae finely punctate (Fig. 109); aedeagus – Aedeagus smaller, length < 2.8 mm, with nar- unknown. n.WA: sw.KID...... rower apex that is far less separated from the ...... froggatti Sloane, 1896 preapical stalk, and with a far less sclerotized – Labrum usually 7-setose; prothorax shorter, and serrate fold in the internal sac (Figs 25, 26); ratio l/w < 1.01; either much larger species, length and shape of elytra various. n.QLD: n. + ce. length > 9.5 mm, or prothorax very conical and CYP south of Jardine River...... 45. elytra short, ratio l/w < 1.73, remarkably oval-

204 shaped, striae very coarsely punctate (Figs 110- 55. Body size large, length > 14.2 mm; aedeagus with 113)...... 50. almost circular, club-shaped apex and elongate parameres (Fig. 38). n.NT...... 50. Generally smaller species, length 7.7-10.4 mm; ...... major Sloane, 1917 prothorax remarkably conical; elytra short, ratio l/w < 1.73; markedly oval-shaped and dorsally – Body size smaller, length < 13.0 mm; aedeagus convex (Figs 110-113)...... 51. either with club-shaped apex but shorter parameres (Fig. 37), or with arrow-shaped apex which – Generally larger species, length 9.6-16.8 mm; is asymmetrically attached (Figs 41, 42), or un- prothorax less conical; elytra usually longer, known...... 56. ratio l/w > 1.80, if less, body length > 12 mm; elytra less oval-shaped, dorsally rather depressed 56. Elytra elongately oval-shaped, markedly nar- (Figs 100, 101, 104-108)...... 54. rowed at base; striae in apical third very weak, 6th and 7th striae inconspicuous (Fig. 107); aedea- 51. Discal punctures of elytra foveiform (Fig. 111); gus unknown. n.QLD...... aedeagus with about circular, club-shaped apex ...... ovalipennis Sloane,1905 (Fig. 44). n.NT...... demarzi Baehr, 1987 – Elytra less oval-shaped, less narrowed at base; – Discal punctures of elytra not foveiform (Figs 110, striae complete, deeply impressed even at apex 112, 113); aedeagus with convexly triangular (Figs 100, 104-106)...... 58. apex (Fig. 45), or unknown...... 52. 58. Prothorax and elytra without any traces of mi- 52. Odd elytral intervals conspicuously raised; croreticulation, glossy (Figs 104, 106); aedeagus pronotum markedly depressed, with fine but with asymmetric, arrow-shaped apex (Figs 41, distinct microreticulation, dull, also with distinct, 42)...... 59. irregular strioles (Fig. 112); aedeagus with convexly triangular apex (Fig. 45). n.NT, n.WA: – Prothorax and elytra with fine microreticulation, ne.KID...... crassipennis, spec. nov. duller (Figs 100, 105); aedeagus with club-shaped apex (Fig. 37), or unknown...... 60. – Odd elytral intervals not raised; pronotum more convex, with less distinct microreticulation and 59. Eltyra little oviform, elongate, ratio l/w 1.84-1.92 very superficial strioles (Figs 110, 113); aedeagus (Fig. 106); whole surface very glossy; aedeagus unknown...... 53. with elongate, arrow-shaped apex (Fig. 42). n.NT, ? n.WA...... nitescens, spec. nov. 53. Elytra shorter, ratio l/w < 1.57; striation regular (Fig. 110); pronotum dorsally less convex; an- – Elytra distinctly oviform, usually shorter, ratio tenna and legs black. n.QLD...... l/w 1.80-1.85 (Fig. 105); dorsal surface less con- ...... hackeri Sloane, 1907 spicuously glossy; aedeagus with shorter, arrow- shaped apex (Fig. 41). ne.QLD...... – Elytra longer, ratio l/w 1.67; striation irregular, ...... nyctosyloides Putzeys, 1868 4th and 5th striae in basal half, 5th and 6th striae in apical half united (Fig. 113); pronotum dorsally 60. Microreticulation on pronotum and elytra fine very convex; antenna red, legs reddish-piceous. but distinct, both surfaces rather dull; elytra n.WA: sw.KID...... horaki, spec. nov. slightly longer, ratio l/w > 1.82 (Fig. 100); aedeagus with club-shaped apex (Fig. 37). n.NT...... 54. Pronotum short, ratio l/w < 0.90; elytra short, ...... brevisterna Sloane, 1916 ratio l/w < 1.78, conspicuously narrowed at base (Fig. 108); aedeagus with arrow-shaped apex – Microreticulation on pronotum and elytra very which is symmetrically attached (Fig. 43). fine and indistinct, both surfaces rather glossy; nw.WA: Pilbara...... pachysoma, spec. nov. elytra slightly shorter, ratio l/w 1.81 (Fig. 105); aedeagus unknown. ne.QLD...... – Pronotum usually longer, ratio l/w > 0.90, com- ...... ovalior, spec. nov. monly more; elytra longer, ratio l/w > 1.80, usually less narrowed at base (Figs 100, 101, 104-106); or elytra longer, elongately oval- procera group shaped, ratio l/w 1.85 (Fig. 107); aedeagus either with club-shaped apex (Figs 37, 38), or with ar- Diagnosis. Medium sized to very large species with row-shaped apex which is asymmetrically at- deep, quadrangular clypeal excision; hind body tached (Figs 41, 42), or unknown. n.QLD, n.NT. parallel-sided or almost so; clypeus 5- or 7-setose, but ...... 55. in some species the number of setae varies between 3-8. Aedeagus with more or less acute apex, but apex

205 never arrow-shaped, or club-shaped, or claw-shaped, differences mentioned in Blackburn’s and Sloane’s or spatulate. All species are fully winged. This is a descriptions and comments vary randomly in the very homogenous group with respect to external whole complex. and also genital morphology. As usual in such complex of very similarly Medium sized species are best distinguished shaped and structured populations, decisions about from species of the impressiceps group by the elongate the taxonomic value, or level, of the various different protibia that bears only three distinct though delicate populations are difficult. The reasons are a certain teeth. amount of variation within populations, the disput- able value of the character states mentioned above, Distribution. 23 species and one subspecies, most and, in the case of leg colour, the difficult decision of which are distributed in the northern part of about the colour in not fully pigmented or sclerotized Australia. specimens. Probably additional taxonomic methods, in particular molecular ones, would be useful and perhaps could help to elucidate the difficult taxo- The Clivina procera complex nomic situation in this complex. In the material available many specimens were According to the mentioned character states, I identified either as Clivina procera Putzeys, 1866, or believe that 12 populations can be distinguished in C. obscuripes (Blackburn, 1890). Usually southern and this complex, which differ in one or another of the eastern specimens were determinated as C. procera, mentioned character states: northern ones as C. obscuripes. C. procera was de- 1. Those specimens from southern and eastern scribed from a single specimen from Melbourne, and Australia which are attributed to C. procera, usually from Melbourne Putzeys (1866) in the same paper are comparatively large, the middle and hind legs are had also described C. prominens which was said to be definitely paler than the anterior leg, and the apex slightly smaller. However, comparison of the types of the aedeagus is more or less regularly triangular, reveals that both certainly belong to the same species. moderately acute, and at the very tip not or little C. obscuripes was described from a single female bent down. This population ranges from eastern SA from “Burrundie” in far Northern Territory and was through VIC, NSW, and southern and eastern QLD claimed by the author, and subsequently by Sloane up to CYP. For this population the name C. procera (1905), to be slightly more robust than C. procera, is still used, because the type specimens of C. procera having slightly shorter elytra with slightly deeper and C. prominens clearly belong to this population. striae, and a slightly more quadrate prothorax. As 2. In the far northern part of NT (environments the name suggests, C. obscuripes has dark legs, while of Darwin down to about Katherine) a population all southern and eastern specimens attributed to of moderately large specimens exists which pos- C. procera have at least the middle and hind legs sess fairly dark legs and barely paler middle and yellow or pale red, at any rate considerably paler hind legs. The apical part of the aedeagus usually than the front leg. is slightly sinuate, but the tip is rather short and All specimens belonging to both nominal spe- quite obtuse, and is more or less, but not suddenly cies have in common the deep, rather quadrangular or markedly, curved down. By comparison with excision of the clypeus, almost impunctate and not the type of C. obscuripes that name is attributed to transversely sulcate head, more or less quadrate pro- this population. thorax, and elongate, almost parallel-sided, dorsally 3. A population of fairly large specimens occurs somewhat depressed elytra. in north-eastern NT and adjacent north-western QLD Having seen several hundreds of specimens from around the coast of the Gulf of Carpentaria. The southern, eastern, and northern Australia attributed apex of the aedeagus, in the few male specimens to either C. procera or C. obscuripes, I recognize that recorded, usually is little or not sinuate and more or the taxonomy is not as easy as it was previously less obtuse. The legs are rather uniformly reddish- believed, because apparently several populations piceous to piceous. It is the connecting link between exist which differ in certain character states of ex- C. procera and C. obscuripes. ternal and genital morphology, and usually possess 4. A population occurring mainly in Kakadu NP definite and more or less well separated areals. The and at some localities west of the park, is compara- main characters that can be used for the distinction tively small, has definitively paler middle and hind of those populations are: body size, colour of legs, legs, and the aedeagus is moderately sinuate, acute, surface structure of head and pronotum, shape of and very shortly bent down at the tip. pronotum, degree of punctation of proepisternum, 5 and 6. Specimens from north-western NT and and shape of the apical part of the aedeagus. All other the Kimberleys commonly are comparatively large

206 and the apical part of the aedeagus is sinuate, very depressed, legs are pale, and the aedeagus is elon- acute, and distinctly bent down at the tip. Because gate, markedly asymmetric, and at tip not incurved. specimens from north-western NT and adjacent For the present paper, mainly for heuristic reasons north-eastern WA (the north-eastern part of KID) and to provide a frame for future taxonomic surveys, have decidedly paler middle and hind legs, whereas I have decided to give preliminary names to the the population from the southern and western part mentioned populations. However, which taxonomic of KID (from about Fitzroy Crossing to Derby, and level should be attributed to these populations? inland to Windjana Gorge) have very dark, even Taking shape and structure of the aedeagus as the completely black legs, they are regarded as two most important character with respect to repro- subspecies of a common species. ductive separation, two groups of populations are 7. A population distributed from far Northern distinguished: namely those with a short, more or Territory to extreme north-eastern Western Australia less obtuse and not much bent down apex of the (the north-eastern part of KID) is large, decidedly aedeagus, and those with an elongate, acute, and larger than any other group except one population distinctly bent down apex. Because in some areas that is even larger (see below under 10.), has pale populations of both groups are sympatric, those middle and hind legs, elongate, acute apex of the populations at any rate must be considered species, aedeagus, and the pronotum has slightly oblique whereas populations with rather similar aedeagi and lateral margins which are almost straight or only that geographically are not widely separated, might little convex. In its whole range it occurs together be considered subspecies. However, as long as no with other populations. other character sets are available, preferably those 8. A population that occurs in northern central of molecular genetics, for heuristic reasons I think NT includes comparatively large specimens with it the best to attribute to all populations the rank of shorter and more quadrate pronotum than usual. species, except two populations in the north-eastern, The aedeagus is of rather similar shape as that of respectively south-western Kimberleys which only the specimens from northern WA, but the middle differ in the leg colour, and which, therefore, are and hind legs are definitely paler than the anterior provisionally regarded subspecies. leg. Most specimens have been recently collected in The result is the following preliminary clas- the area from about Katherine to Elliott. sification: 9. A single male specimen from north-western QLD likewise has a very sinuate and acute, at tip markedly bent down aedeagus, but the legs are Clivina procera Putzeys rather uniformly reddish-piceous and the protho- Figs 1, 54, 78, 80 rax is unusually elongate and depressed. Therefore the specimen is quite different from those of the Clivina procera Putzeys, 1866a: 34. – Putzeys 1866b: 180; population around the Gulf of Carpentaria, as well 1868: 7; Blackburn 1890: 1248; Sloane 1896a: 181, 227, in external, as in genital morphology. 228, 231, 238, 244; 1896b: 275; 1905a: 730, 732; Csiki 10. A very large species, actually the largest 1927: 493; Moore et al. 1987: 75; Lorenz 2005: 144. found in Australia and perhaps the largest Clivina Clivina prominens Putzeys, 1866: 35 (syn. nov.). – Putzeys in general, occurs in KID in north Western Australia. 1866b: 182; 1868: 8; Blackburn 1890: 1249; Sloane It is characterized by completely black legs, short, 1896a: 229; 1905a: 730; Csiki 1927: 493; Moore et al. laterally markedly convex prothorax, and elongate, 1987: 75; Lorenz 2005: 144. remarkably downcurved apex of the aedeagus. Examined types. Holotype of procera: M, Scol. procerus. Moreover, the subapical margin of both parameres is M / Ex Musaeo Chaudoir (MNHP). – Lectotype of conspicuously quadrangular. It is sympatric with one prominens (by present designation): W, Ex Musaeo or even two other populations, according to the area. Chaudoir, no label but sub “prominens Putz. Australie 11. A population of rather large specimens bears Melbourne Bakewell” (MNHP). several more or less distinct transverse sulci on the Type localities. Of procera: “Melbourne”, Victoria. – Of frons. The head is sparsely but distinctly punctate, prominens: “Melbourne”, Victoria. the pronotum is depressed, middle and hind legs are pale, and the aedeagus is short and at tip not Note. As the type specimen of C. prominens does incurved. It occurs in north QLD up to the northern not exhibit any differences in body shape, apart part of CYP. from being slightly smaller than the type specimen 12. In a population of fairly large specimens from of C. procera, the name prominens is synonymized south-western QLD and adjacent north-eastern SA with procera. This procedure is corroborated by the the dorsal surface is not or barely microreticulate similar type localities and the considerable variation and remarkably glossy, the pronotum is wide and in body size of this species.

207 Other material (316 ex.). SA: Waikerie 24.viii.58 Clivina procera Putzeys (NMV COL-18909); Nr. Melbne B.P.Moore / Clivina procera Putz. Det. B.P.Moore’62 (NMV COL-13512/17); Murray R. / C. French’s Collec- (ANIC); Innamincka: Coopers Ck 24 Jan. 1976 M.S. & tion (NMV COL-5545); Kulkyne Dec. 1917 R.Thompson B.J.Moulds / Walford-Huggins Collection / Clivina / NM 7 Clivina procera Putz. (AMS K 149367); Vict. / procera Putz. det. B.P.Moore’77 (CMP); Type / Putzeys Clivina procera Putz. Id. by T. G. Sloane / Collectio Typ! / Gawler Odewahn / Compared with T Y P E K. Karel Kult (CDW); Victoria (NMV COL-10510/13); Kult 1946 / prominens Putz. typ! / Collectio Karel Clivina procera P. / Victoria (MMS). – NSW: Yandilla F. Kult (CDW); 27.02S, 140.02E unnamed Lake, 15.x.72 A. Gore. Don. 11262 / Clivina planifrons Putz. N. S. Wales Roffey & Mitchell (ANIC); 34.03S 140.43E GPS Calpe- (QMB); Dubbo 4 Dec. 1973 / Coll. A&M Walford- rum HS. 15km NNW Renmark, 7-8 Nov.1995, M.Do Huggins / Clivina procera Putz. det.M.Baehr’95 (CMP); minguez & K.R.Pullen / Calperum Station/Bookmann Australia, H.J.Carter 1912-483. / Windsor Hood 7.04 / Biosphere Reserve Invertebrate survey (ANIC); 34.02S C. procera Putz. (NHM); Windsor Hood 7.04 / H. J. 140.36E GPS 21km NW Renmark, 8 Nov.1995 Car Carter Coll. (NMV COL-10516); Bogan R. J. Armstrong dale,Lee,Pullen & Dominguez / Calperum Station/ / C. procera Putz. det. K. Kult 1948 / Collectio Karel Bookmann Biosphere Reserve Invertebrate survey Kult (CDW); Bogan R. J. Armstrong / Clivina procera (ANIC); 33.47S 140.42E GPS 24km NbyW Renmark 13 Putz. Id. by J.W.T.Armstrong (ANIC); Bogan R., S.of Dec 1995 Mallee grassland K.R.Pullen / Calperum Sta- Nyngan Oct’57 Darlingtons (MCZ); 10 km E of Nyngn, tion/Bookmann Biosphere Reserve Invertebrate survey 26.x.1978 K. J. Lambkin & D. R. Smith (QMB); 30.52S (ANIC); Adelaide / Adelaide Coll.Castelnau / Scolyptus 147.46E Sandy Camp Station, Macquarie Marshes, W. of procerus Putz. teste Putzeys 1868 (MCSG); Myponga A. Coonamble, 27 May 1952. J.Balderson (ANIC); Deni- H. Elston / C. procera Putz. Id. by A. M. Lea / A.H. liquin, 26.12.66 V. R. Squires (ANIC), Lake Cowal Elston Collection (AMS K 149365); Naracoorte 20.i.1967 10.iii.1970, 15-2-1971 W. Vestjens (ANIC); c.25km W. of G. Monteith (UQIC #90678-9 (QMB); Mt. Gambier Lea Wentworth, Moorna Station 13-18.ix.74, P.Meyer (SAMA 25-033615); Blanchetn (SAMA 25-033609); Man- (ANIC); Collector, N.S.W. 30.55S 149.26E 11.ix.1968, num (SAMA 25-033741); W.of Renmark Sept.’57 Dar- A.Clarke (ANIC); Mul / C. procera Putz Id. by Sloane lingtons (MCZ); R.Murray S. Australia A. H. Elston / 7.04 (NMV COL-5542); Murray R. / F. E. Wilson Col- 862 Clivina procera Putz. Id. by T.G. Sloane / A.H. Elston lection (NMV COL-5535); Canowindra Jan 56 F.E.Wilson Collection (AMS K 149366); R.Murray A. H. Elston / / F.E. Wilson Collection (NMV COL-10518); Richmond Clivina procera Putz. / A. H. Elston Collection (AMS); R. Austr. Mus. 5-02 / H. J. Carter Coll. (NMV COL- S.Australia Blackburn / 3105 (unreadable) / procerus 5426); Richmond R. SYNTYPE Clivina foveiceps, Macl. Putz. / Clivina procera Putz. S. Australia (SAMA); S.Aust (ANIC-MMS); Condobolin 29.10., 30.10., 31.10., 1.11.1958 (MMS); South Australia (MMS); S. Australia / Clivina / Clivina procera Putz. Det. B.P.Moore 1999 (AMS K procera Putz. S.Aust. (MMS); Scolyptus / S. Australia 147964-67); 301 JM 8.1.20 Coonamble / Clivina sp. (SAMA 25-033618). – VIC: Benalla 29.viii.60 B.P.Moore (AMS); Gwabegar 16.11.1979 / Clivina procera Putz. Det. (ANIC); Benalla viii.60 B.P.Morre (ANIC); Hattah Lakes B.P.Moore 1999 (AMS); S. procerus Putz Wentworth 25.xi.68 B.P.Moore (ANIC); 35°31'03S 141°18'56E Big (SAMA); Allroy N.S.W. / 7744 Clivina procera Putz. N. Desert Big Billy 7-18 Nov 1997 M.Naeve & M.Powell S. Wales (SAMA); C. procerus Putz. Wentworth / Sco- (ANIC); 34.35S 132.46E (wrong longitude, must be 142°!) lyptus procerus Putz. / 7742 Clivina procera Putz. N. S. Robinvale 25 Oct.-3 Nov. 1988 T.Weir, J.Lawrence & Wales (SAMA); Heatbon State Forest, 22 km SE Cess- M.Hansen (ANIC); 2km W Ulupna Island NNW Strath- nock St.64 2 XI 1980 / Australie New South Wales C. & merton 30 Oct 1990 P. A. Meyer (ANIC); Bonnetry (?) A. Jeekel (ZMUA, now NMNL); Round Hill district Jan Bend 26.viii.69 B.P.Moore (ANIC); Watchem 10.1.1989 1990 A.Sundholm J.Bugela (AMS K 225388); N.S.Wales Spurrell. (NMV COL 5540); Bermah For. 36°S, 145°E, / Scolyptus procerus Putz. Darling River (MMS); Lake 3.11.02, A.Ballinger / Clivina procera Putz. det.M.Baehr’03 Cowal 11 : 11 : 1972 D. A. Doolan / D. A. Doolan Col- (CBM); Lake Hattah 28.xi.67 G.W.Anderson (ANIC); lection / Clivina procera Putz. Det. B.P.Moore 1999 (AMS Kulkyne, Dec.34.7 R.Thompson / Clivina procera Putz K 147962); Mulwala T.G.S. (unreadable) / Clivian procera Id. by F. E. Wilson (ANIC); (unreadable) / Clivina procera Putz. Id. by T. G. Sloane (QMB); Australien N.S.Wales Putz. (Brisbane) (ANIC-MMS); Natia J.E.Dixon (NMV E.Plason / Sammlung H.Hesse (SMNS); (unreadable), COL-13550); Ouyen mallee 20.1912 / Pres by J.E.Dixon N.S.W. T.G.S. 8.10.25 (ANIC); N.S.Wales / Clivina pro- 19.11.16 (NMV COL-14050-1); Lake Hattah J. E. Dixon cera / Coll. Franklin Müller / Clivina procera N.S.Wales (NMV COL-13545); Lake Hattah Nov. 1924 C. Oke / / C. procera Putz. det. K. Kult 1948 (DEI); Clivina procera Clivina procera Putz. Id. by T. G. Sloane (NMV COL- Putz N.S.W. (SAMA); H. J. Carter Coll. / Clivina procera 5534/8); Lake Hattah Nov. 1924 C. Oke / Clivina procera Putz. N. S. Wales (NMV COL-14018); N.S.Wales (MMS); Putz. Id. by C. Oke / UQIC Reg. #90742 (QMB); Echuca, New South Wales UQIC Reg. # 907546 (QMB); N.S. 28.12.1989 E.M. (NMV COL-13619); Caulfield (NMV Wales / Scolyptus planiceps Putz. Australien O. Langen- COL-5514); Caulfield / Clivina procera Putz. det. by han / Sammlung O. Langenhan. Kauf 1931. 18 (SMTD); Sloane 7/04 (NMV COL-10515); Inglewood Dec. 1925 N.S.Wales / SYNTYPE (of C. foveiceps Macl.) (ANIC- C. Oke. / Clivina procera Putz. Id. by T. G. Sloane (NMV MMS). – QLD: “11 Mile Scrub”, 19 km N, of Moreton, COL-5536); Linga Oct. 1932 F.E.Wilson / F. E. Wilson C. York Pen. 15-16.vii.1973 G. B. Monteith (CBM, QMB); Collection (NMV COL-5537); Nagambie 12.34 J.C. 9 (?) Homestead Silver Plains East Coast Cape York Pen. (NMV COL-20836); Clivina procera Putz. Sea Lake / 20.IV.1960 J. L. Wassell (ANIC, CBM); Silver Plains Cape

208 York, Q. May-June ’58 Darlingtons (MCZ); Cooktown, “Allinga” Chinchilla Grace Lithgow (QMB); Emerald 30.12.48 E. Adams E.Sutton. / E. Sutton Collection. E.Allen 1916 (QMB); In flood debris / Rockhampton (QMB); Cooktown 19.4.53 C. Oke (NMV COL-13604); (SAMA 25-033612/19); Rockhampton / Rockhampton W. of Ravenshoe Atherton Tab., Q. c.3000’, Feb. 58 Coll.Castelnau / Scolyptus procerus Putz. teste Putzeys Darlingtons (MCZ); N. of Mareeba Feb.’58 N.Q.Dar- 1868 (MSCG); Coll. C. FelscheGeschenk, 1907 / Rock- lingtons (MCZ); 18.09S 144.20E Mount Surprise, Qld. 2.4 hampton (SMTD); P. Denison / Port Denison Coll. April 1995 J.Balderson P.K.Christensen (ANIC); Tolga Castelnau / Scolyptus procerus Putz. teste Putzeys 1868 N. Qld. 3-iv-1983 J.D. Brown, light trap (QDPIM); Tolga (MCSG); Caves nr. R/ton RLHiggins (ANIC); Clermont N. Qld. 5.iv.1981 N. Gough, J.D. Brown at light (QD- VII 29 Dr. K. K. Spence / C. Procera Putz. M Id by Dr. K. PIM); Cairns 3/53 GB. / M. 157 /J. G. Brooks Bequest K. Spence (AMS K 149368); Brisbane HJC 14/10 / H. J. 1976 (ANIC); Cairns (ANIC); v. Cairns, Q. Darlingtons Carter Coll. / Clivina procera Putz. (NMV COL-5544); (MCZ); Woodstock 3.9.53 E.Sutton A.Johnson / E. Sut- Gayndah K37093 / Clivina procera Putz Id. by T. G. ton Collection. (QMB); Woodstock 11.11.56 A.Johnson Sloane (AMS); Maryborough E. W. Fischer (SAMA 25- E.Sutton / E. Sutton Collection. (QMB); Woodstock N.Q. 033620); Milmerran 11/53 J.M. / M. 159 / J. G. Brooks 3/54 AJ / procera Putz. 339a / J.G. Brooks Bequest 1976 Bequest 1976 (ANIC); 5-4-55 / J.Sutherland Brisbane / (ANIC); Kowanyama, N. Qld. 9.i.1977 D.L.Hancock UQIC Reg. 90346 (QMB); Ex light trap St. George, 8. (QMB); Townsville G.F.Cook / Clivina obscuripes (un- ii.1974 L.Lloyd (QDPIB); Strathmore Stn. via George- readable) (SAMA 25-033772); Qld. Greenvale, 70km SW town 22.iv.1984 R.I. Storey at light (QDPIM); Kowany- at light. 28 Mar- 7 Apr. 1995 A. J. Watts (SAM); N. ama 9.i.1977 D. L. Hancock (QMB); Townsville, light Queensland 29.1.-3.2.2000 The Lynd Junction Sv.Bilý trap 23.iv.68 P. Ferrar (ANIC); Townsville 15.12.02 F. P. leg. (CBM); Wallaroo legit G. Hangay, 17.I.68 / Clivina Dodd / 2550 / Griffith Collection Id by A. M. Lea procera Putz. det.M.Baehr’96 (CBM); Wallaroo 17.I.68 (SAMA 25-033536); Queensland Br. Müller F. / C. pro- leg. G.Hangay / Clivina procera Putz. det.M.Baehr’89 cera Putz. det. K. Kult 1948 / Collectio Karel Kult (QMB); Mackenzie River 29.I.1968 leg.G.Hangay / (CDW); N.Qld / obscuripes Blackb. (???); Cape York VI. Clivina procera Putz. det.M.Baehr’89 (CBM); Foleyvale 1993M. & P. Uhla legit (CBM). – WA: W.Austr / Clivina Abor.Res. 20.-23.I.1968 lg.G.Hangay / Clivina procera procera P. (MMS); Swan Riv. + / 493 / Swan River Coll. Putz. det.M.Baehr’89 (CBM); SEQ: 25°26'SX150°01'E Castelnau / Scolyptus procerus Putz. / Scolyptus procerus Taroom District Boggomoss No.3 9 Sep-11 Nov1996, OF Putz. teste Putzeys 1868 (MCSG). – AUS: Sc. prominens P.Lawless o94 / QM Reg.No. T35621, 35624-8 / Bog- (unreadable) Putz. / Collectio Karel Kult (CDW); gomoss databased as Clivina sp.1 Temporary label only Aust. 73.7 / procera Ptz. det. K. Kult (NHM); 29910 / (QMB); SEQ: 25°27'SX150°03'E Taroom District Boggo- Nov.Holl. Austrs / Fry Coll. 1905-100 (NHM); Aus- moss No.21 18 Jun1996, H. Janetzki / QM Reg.No. tralien / Coll. Felsche, det. procera (SMTD); Montevideo T35629 / Boggomoss databased as Clivina sp.1 Tempo- 1889, Müller / 196, Scolyptus procerus Putz. (unreadable) rary label only (QMB); 6 km N Taroom25°36'E,149°46'S 80. (SMNS); Soc. Ent. Belg. Coll. Putzeys / Clivina procera 1 Mar 1991 mv lamp G. Daniels / UQIC Reg. 90253 Putzeys, 1866 (syn: Scolyptus procerus Putzexs, 1866) (QMB); 25°26'SX150°01'E Boggomoss Ck. BM03. (IRSNB); Soc. Ent. Belg. Coll. Putzeys / Sc. procerus Putz. Nov1996, 200m P.Lawless 10590 (QMB); 25°27'SX150° / Clivina procera Putzeys, 1866 (syn: Scolyptus procerus 03'E Boggomoss No.21 via Taroom. 11-12Nov1996, Putzeys, 1866) (IRSNB); Australie / Ex Coll. Chaudoir F.Cook o72 / QM Reg.No. T35621, 35622 / Boggomoss (MNHP); Coll. French Australia (ANIC); Trangie Stn databased as Clivina sp.1 Temporary label only (QMB); 8-9/9/77 / Clivina sp. det. T.A.Weir 1978 (ANIC); 296 26°23'Sx146°12'E Charleville, 8km SW 4-5Mar2003; / Carabidae Clivina elegans Putz. (ANIC); (defect), Mul. 300m G.Monteith, C.J.Burwell Mulga 51123 (QMB); 90 (ANIC); Clivina procera Putz. Id by Sloane 7/04 (NMV 26°26'Sx146°10'E Charleville, 5km NE 3-5Mar2003; COL-10512); Murch (unreadable) C. French’s Coll. / 310m G.Monteith, C.Burwell Box flat 51124 (QMB); 5km Clivina procera Putz. Vict & Queensland (NMV COL- NE Edungalba 18 Jan 1991 AStofarski (SAMA); Towns- 5543); Clivina procera Putz. Id. by C. Oke / E.T.Smith ville G. E. Bryant F.P.Dodd. 1911 / J. G. Bryant Coll. Collection (NMV COL-11913-4); 2872 (NMV COL- (NHM); Townsville Brown / H. J. Carter Coll. P.20.4.22. 10508); Murray River / Clivina procera P. (MMS); Darling (NMV COL-5411); Townsville to Mackay. Q Mar’58 R. (MMS); Riv. Paroo. / Riv. Paroo Coll. Castelnau / Darlingtons (MCZ); Injune CQ 2.3.38 E. Sutton / E. Scolyptus procerus teste Putzeys 1868 (MCSG); Ex J. J. Sutton Collection (QMB); Rockhampton A. M. Lea Walker bequest 1939 (OUM); (label faded) / M.416 (Sama 25-033518); Rockh / J. French’s Coll. / Clivina (QDPIM); Riv. Darling Coll.Castelnau / Scolyptus pro- obscuripes Blckb. Queensland (NMV COL-5140); Rock- cerus M / Scolyptus procerus Putz. teste Putzeys 1868 hampton Mar.’58, Q. Darlingtons (MCZ); Rockhampton (MCSG); Australia / Coll.Castelnau / Scolyptus procerus 9 2.1-44 E.Sutton C.Valles / E.Sutton Collection Don- Putz. t. Putz. / Scolyptus procerus Putz. teste Putzeys Dec.1964. (QMB); Dawson R. (ANIC); Townsville, 1873 (MCSG); Riv. Darling Coll. Castelnau / Clivina 15.12.02 F. P. Dodd (ANIC); Nelly Bay Magnetic Is procera Putz. teste Putzeys 1868 (MCSG); (unreadable) Mar97 S.Feam (QMB); Mackay / C. French’s Coll. / 916 14.11.91 (SAMA 25-044617); Wyandotte 16-4-46 / S.R.E. / Clivina procera Putz. Id. by T. G. Sloane (NMV COL- Brock Collection (ANIC); Relton / Relton Bequest / 5539); 7 / Dalby Q. Mrs. F. H. Hobler 7 Clivina procera Clivina quadraticeps Sloane (QMB); Clivina obscuripes Putz. Id. by T. G. Sloane (SAMA); Dalby Q. Mrs. F. H. (NMV COL-5399). Hobler (SAMA 25-033616/714-16); 26°41'Sx150°38'E

209 Diagnosis. Moderately large to large, black spe- Legs. Protibia narrow and elongate, in males cies with deep, quadrangular clypeal excision, not more than in females. longitudinal sulcus on upper sulcate, barely punctate head, rufous middle and surface barely perceptible; lower surface with a row hind legs, and in apical third asymmetric aedea- of 6-8 stout setae along basal part of inner margin. gus with the apex directed right and more or less Mesotibia moderately elongate, dorsal rim crenulate, curved down. with several rows of setae on the dorsal surface; subapical spur elongate. Metatibia elongate, rather Description sparsely setose. Metatarsomere 1 almost as long as the following three tarsomeres. Measurements. Length: 12.2-16.7 mm; width: 3.3- Male genitalia (Fig. 1). Aedeagus wide, fairly 4.7 mm. Ratios. Length/width of pronotum: 1.02- elongate, narrowed apicad; lower surface gently con- 1.05; base/apex of pronotum: 1.26-1.33; width cave, towards apex more concave. Apical part very pronotum/head: 1.33-1.40; length/width of elytra: asymmetric, deeply bisinuate at the left side; apex 2.01-2.06; length/width of protibia: M: 4.0-4.15, moderately short, slightly turned left, asymmetric, W: 3.3-3.35; length/width of metatibia: M; 5.6-5.8, tip not incurved, fairly obtuse. Both parameres very W: 5.5-5.8. elongate, the left one considerably stouter than the Colour (Fig. 80). Unicolourous black. Anterior narrower right one; both with elongate, very nar- leg piceous, median and posterior legs rufous. row, bisetose, apical part; setae very short, situated Head (Figs 78, 80). Eye large, laterally much right at tip. projected, orbit short, slightly oblique-convex. Cly Female gonocoxites (Fig. 54). Gonocoxites nar- peus with deep, almost quadrangular excision. row and elongate, little curved, with rather acute Labrum usually 5-setose. Clypeus not divided from apex; with one elongate but comparably slender frons; upper surface remarkably smooth, without seta basally at lateral surface and a smaller seta transverse sulci or strioles; frons in anterior part below, two elongate setae on the ventral surface with sparse, rather coarse punctures, otherwise very of gonocoxite 1, two or three setae at middle of the finely punctate. Surface without distinct microre- medio-dorsal surface, but without a nematiform seta ticulation, glossy. near apex. Lateral plate basally with one elongate Pronotum (Fig. 80). Comparatively short, con- seta, apically without or with one shorter seta. siderably narrowed apicad, dorsally rather de- Variation. Body size very varied, but the major- pressed. Lateral margin oblique, faintly concave ity of the specimens, particularly the southern ones, in apical third. Apex distinctly concave. Marginal are large. Also shape of pronotum varies to some channel narrow and median line shallow but well degree, being more or less narrowed apicad. impressed, impunctate. Basal sulcus narrow. An- terior marginal seta situated at apical fourth. Basal Distribution. Eastern Australia from eastern SA groove elongate, linear, straight, impunctate. Disk to north-eastern QLD. The few old records from with some very fine, transverse strioles in middle, ex- southern Western Australia certainly are wrong. tremely finely punctate, with only traces of extremely superficial, isodiametric microreticulation, glossy. Collecting circumstances. Specimens were collected Elytra (Fig. 80). Moderately elongate, almost by: “Mercury vapour light”, “at light”, “light trap”, parallel-sided, barely widened apicad, convex but in “Pitfall”, “in flood debris”, and in “dung trap”, but middle depressed. Basal angle little produced. Stria- almost all old specimens do not bear any information tion complete, but lateral striae less distinct. Striae about sampling methods. in basal half well impressed, becoming shallower towards apex, impunctate or almost so. Intervals slightly convex, impunctate, with some transverse Clivina obscuripes (Blackburn) strioles, with fine, isodiametric microreticulation, Figs 2, 81 moderately glossy. Epipleura narrow, basally with a sulcus formed by a row of punctures. Scolyptus obscuripes Blackburn, 1890: 1247. Metathoracic wings. Fully developed. Clivina obscuripes, Sloane 1905a: 730, 733; Csiki 1927: 508; Lower surface. Proepisternum impunctate, with Moore et al. 1987: 74; Lorenz 2005: 143. few irregular, elongate dorso-ventral strioles and Examined types. Holotype: W (probable), T. 2894 N.T. / many short, transverse strioles, also with distinct Type / Blackburn coll. 1910-236. / Scolyptus obscuripes, isodiametric microreticulation. The whole lower Blackb. / Clivina obscuripes Blackb. 47 det. K. Kult surface with fine, isodiametric microreticulation. (BMNH). Metepisternum elongate, c. 2.5 × as long as wide, Type locality. “Burrindie”, Northern Territory. in anterior half with sparse, rugose punctures. Ab- dominal sterna impunctate.

210 Other material (94 ex.). NT: 10 MM, 3 WW, 12.57S 132.33E Description Jim Jim Creek, 19km WSW of Mt. Cahill, N.T. 19.v.73 N.T. T. Weir & T. Angeles / 11236, -46, -54, -60, -64, Measurements. Length: 10.0-13.5 mm; width: 2.8- -66, -72-73, -75-78, -92 (NTD, CBM); 3 MM, 12 WW, 3/06, 3.85 mm. Ratios. Length/width of pronotum: 1.02- KAKADU NP, MUIRELLA 10.05.2006 40M S12°51'15" 1.03; base/apex of pronotum: 1.27-1.33; width E132°45'16" LIGHT LG. BERGER-DOSTAL (CBM, pronotum/head: 1.30-1.33; length/width of elytra: CDW); 1 M, 1 W, Kakadu Nat. Park, CSIRO Biodiversity 2.0-2.06; length/width of protibia: M: 3.5-3.65, W: 3.0- Survey, Site 40, 13.6062121 S, 132.959611 E, S. Oberprie ler, Feb-Mar 2014 (CBM); 5 MM, 4 WW, 13.03S 132.19E 3.25; length/width of metatibia: M; 6.0-6.1, W: 5.5-5.7. South Alligator River, 46km WSW of Mt. Cahill 20.v.73 Colour (Fig. 81). Unicolourous black. Anterior Matthews & Upton (ANIC, CBM, SAMA); 4 MM, 5 WW, leg dark piceous to black, median and posterior 13.45S 138.41E Daly River Mission 8-24.vi.1974 J. F. legs piceous to almost black, little or not paler than Hutchinson (ANIC); 3 WW, Daly River Mission 4.vii.1974 the anterior leg. J. F. Hutchinson (ANIC); 4 MM, 3 WW, S. Alligator River Head (Fig. 81). Eye large, laterally well project- area, 35 km W Jabiru, blacklight, 9.iv.1980, GFHevel ed, orbit short, oblique-convex. Clypeus with deep, & JAFortin (CBM, USNM); 1 W, 12.26S, 132.58E Cahills almost quadrangular excision. Labrum usually 5-se- Crossing, East Alligator River, 29.v.73, at light, E. G. tose. Clypeus not or barely divided from frons; upper Matthews (ANIC); 1 M, 12.57S 132.33E Jim Jim Creek, surface smooth, without transverse sulci or strioles; 19km WSW of Mt. Cahill, N.T. 19.v.73 N.T. T. Weir & frons in anterior part with or without sparse, rather T. Angeles / 11260 (NTD); 1 M, 1 W, Kakadu Nat. Park Cooinda30.III.1996 P.M.Giachino leg. (CGT); 1 M, 17m coarse punctures, otherwise very finely punctate. alt, Kakadu NP, Ubirr env, 12°25'S 132°57'E 25-26. Surface without distinct microreticulation, glossy. iv.2009, St. Jakl lgt. (CJP); 1 M, 2 WW, LITCHFIELD NP Pronotum (Fig. 81). Comparatively short, con- Adelaide River, 29.V.2000 leg. M. Langer (CBP); 1 W, siderably narrowed apicad, dorsally rather de- Windows of the Wetland Point Stuart, 25.V.2000 leg. pressed. Lateral margin oblique, faintly concave M. Langer / Wilderness Lodge Lichtfang am Rand von in apical third. Apex distinctly concave. Marginal Monsunregenwald (CBP); 1 M, Arnhem Land Urupunga channel narrow and median line shallow but well June 1975 Wendy Walsh / Clivina obscuripes Blbn det. impressed, impunctate. Basal sulcus narrow. An- P.P.Moore’82 / Clivina obscuripes Blackburn comp. w. terior marginal seta situated at apical fourth. Basal ’ type B.P.Moore 86 (ANIC); 1 M, 12.06S 133.04E Cooper groove elongate, linear, straight, impunctate. Disk Creek, 19km E by S of Mt. Borradaile, 5.vi.1973, T.Weir with some very fine, transverse strioles in middle, & A.Allwood (ANIC); 1 M, 1 W, Mary R. Arnhem Hwy. 29 Nov. 1978 R.I. Storey At Light Clivina obscuripes extremely finely punctate, with only traces of ex- B’bn det. B.P.Moore’79 (ANIC, QDPIM); 1 M, Katherine tremely superficial, isodiametric microreticulation, 2/3/93 Leg. Leech (CSM); 1 W, Katherine 28/3/93 Leg. fairly glossy. L. Toledano (CGT); 1 M, Mainoru Stn. 26.XI.81 Walford- Elytra (Fig. 81). Moderately elongate, almost Huggins / Walford-Huggins Collection Carnegie Mu- parallel-sided, barely widened apicad, convex but seum / Clivina obscuripes Blbn det.B.P.Moore’82 (CMP); in middle depressed. Basal angle little produced. 2 WW, 12.48S 132.44E 8 km NE by N of Mt. Cahill Nov. 26, Striation complete, but lateral striae less distinct. 1974 Coll – R. I. Storey at light / UQIC #90340-1 (QMB); Striae in basal half well impressed, becoming shal- 1 W, Kakadu N. Park S. Alligator River 19.xii.1990 lower towards apex, usually at least in basal half (MCNV); 1 W, Muirella Park, Kakadu 8.v.1987 Fay and finely punctate. Intervals slightly convex, impunc- Halfpapp at light (QDPIM); 1 W, Darwin 20.vi.1994 tate, usually without transverse strioles, with fine, (CHP); 1 W, Port Darwin (SAMA 25-033589); 1 M, Caught in flooded area King R. by W.Melliman 24-12-15 (?) / isodiametric microreticulation, moderately glossy. Not likely different from Clivina obscuripes Black. Id. Epipleura narrow, basally with a sulcus formed by by T. G. Sloane / COL-5407 (NMV); 4 MM, 5 WW, King a row of punctures. R. 24.12.15 Coll. W. Melliman / Clivina obscuripes Blkb Metathoracic wings. Fully developed. Det by T.G.Sloane Rec - 3 - 17 / COL-5400-6 (NMV); Lower surface. Proepisternum usually little or 1 W, Darwin Coll. by (?) R. Spencer 1912 / 826 Clivina not punctate, with few irregular, elongate dorso- obscuripes BlSl thinks this is not distinct from C. procera ventral strioles and many short, transverse strioles, Puz. / COL-5408 (NMV); 1 W, (largely unreadable) 14- also with distinct isodiametric microreticulation. 10-16 / Clivina obscuripes Blkb det. by T G Sloane Rec The whole lower surface with fine, isodiametric - 3 - 17 / COL-5409 (NMV); 1 M, Tortilla Flat 11.iii.1987 microreticulation. Metepisternum elongate, c. 2.5 × J. Waldock / 11301 (NTD). – AUS: 1 W, No locality label as long as wide, in anterior half with sparse, rugose S.R.E.Brock Collection (ANIC). punctures. Abdominal sterna impunctate. Diagnosis. Comparatively small to moderately Legs. Protibia narrow and elongate, in males large, black species with deep, quadrangular clypeal more than in females. longitudinal sulcus on upper excision, not sulcate, barely punctate head, dark legs, surface barely perceptible; lower surface with a row and short, in apical third slightly asymmetric aedea- of 6-8 stout setae along basal part of inner margin. gus with straight, barely curved, at tip obtuse apex. Mesotibia moderately elongate, dorsal rim crenulate,

211 with several rows of setae on the dorsal surface; of Mt. Borradaile, 5.vi.73, M.S. Upton (ANIC); 1 M, subapical spur elongate. Metatibia elongate, rather 12.26S, 132.58E Cahills Crossing, East Alligator River, sparsely setose. Metatarsomere 1 almost as long as 29.v.73, at light, E. G. Matthews (ANIC); 1 W, 12.17S the following three tarsomeres. 133.20E Cooper Creek, 11km S. by W. of Nimbuwah Male genitalia (Fig. 2). Aedeagus wide, fairly Rock, N.T. 3.vi.73, T.Weir & T.Angeles / 11234 (NTD); 3 MM, 12.51S 132.47E 8km E by N of Mt. Cahill, N.T. elongate, narrowed apicad; lower surface regularly 22.v.1973 T.Weir & T.Angeles / 11296-98 (NTD); 1 M, concave. Apical part very asymmetric, more or less 12.22S 133.01E 6km SW. by S. of Oenpelli, N.T. 30.v.73, deeply bisinuate at the left side; apex short, slightly E. G. Matthews (ANIC); 1 M, 12.17S 133.13E 18km E by turned left, asymmetric, tip not incurved, rather ob- N of Oenpelli, NT 1.vi.73, Matthews & Upton (ANIC); tuse. Lower surface in middle with a low ridge. Both 2 MM, 12.31S 133.19E Nabarlek Dam, N.T. 15km S. by parameres very elongate, the left one considerably W. of Nimbuwah Rock, 2.vi.73, at light, E. G. Matthews stouter than the right one; both with elongate, very (ANIC); 6 MM, 2 WW, 12.54S 132.32E Jim Jim Airstrip, N.T. narrow, uni- or, more commonly, bisetose apical 1.v.1973 T.Weir & T.Angeles / 11279-82, -85-88 (CBM, part; setae short, situated right at tip. NTD); 1 M, 1 W, 12.20S 132.54E 9km N by E of Mudg- Female gonocoxites. Very similar to those of inbarry HS, 26.v.73, Matthews & Upton (ANIC); 1 M, C. procera Putzeys. 12.31S 132.54E 9km N by E of Mudginbarry HS, N.T. 31.v.73, E.G.Matthews (ANIC); 2 MM, East Alligator R. Variation. Body size rather varied. Also shape 10 ml.S.W.Oenpelli Mission N.T. 22.vi.1971 A.Allwood of pronotum varies to some degree, being more or &T.Angeles / 11230-31, -62-63 (CBM, NTD); 1 W, 12.25S less narrowed apicad. 132.58E 1km N. of Cahills Crossing, N.T. East Alligator River, 7.vi.73, Upton & Feehan (ANIC); 2 WW, 12.25S Distribution. Northern part of NT, particularly in 132.58E 1km N. of Cahills Crossing (East Alligator R.) Kakadu NP. 29.v.1973 T.Weir & N.Forrester / 11245, -51 (NTD); 1 M, 12.36S 132.52E Magela Creek, N.T. 1km NNW. Collecting circumstances. Little recorded. Recently of Mudginbarry H.S. 25.v.1973, Matthews & Upton collected specimens were mainly obtained at light. (ANIC); 7 MM, 7 WW, AUSTRALIA N.T. Kakadu N.P. 22-25/3/93 Cooinda at light/al lume Leg. L. Toledano (CBM, CLT); 2 MM, 2 WW, AUSTRALIA N.T. Kakadu N.P. Clivina subrufipes, spec. nov. Cooinda 25-26/12/96 at light/al lume Leg. L. Toleda- Fig. 3 no, R. Olivieri (CBM, CLT); 5 MM, 2 WW, AUSTRALIA, N.T. Kakadu N.P.22-25/3/93 Leg. Leech (CBM, CSM); Examined types. Holotype: M, 12.23S 132.56E 7km NNW 3 MM, Australia N. T. Kakadu Nat. Park 30.III.1993 of Cahills Crossing, East Alligator River, N.T. 27.v.73, (CGT); 2 MM, AUSTRALIA N.T. KAKADU N.P.18/20. E. G. Matthews (ANIC). – Paratypes: 2 MM, same data XII (MCNV); 2 WW, AUSTRALIA N.T. Kakadu N. Park (ANIC); 4 MM, 1 W, 12.48S 132.42E Nourlangie Ck. N.T. Cooinda 20.xii.1990 (MCNV); 1 M, 1 W, Australia NT 8km N. of Mt. Cahill, 21.v.73, at light, E. G. Matthews Kakadu, 10/14.I.2004 Lgt. R. Novák (CKZ); 1 M, 1 W, (SAMA); 1 M, 12.48S 132.42E Nourlangie Ck. 8km N. of AUSTRALIA near Kakadu 1994 18/I E.&C.Balietto leg. Mt. Cahill, 26.xi.1974 N.T. T. Weir & T. Angeles / 11307 (CCS); 1 M, 1 W, AUSTRALIA NT 17m alt, Kakadu NP, (NTD); 1 M, 1 W, 12.48S 132.44E Nourlangie Ck. 8km NE Ubirr env, 12°25'S 132°57'E 25-26.iv.2009, St. Jakl lgt. by N of Mt. Cahill, N.T. 21.v.73,T.Weir & T.Angeles / (CBM, CJP); 1 W, AUSTRALIA Northern T. Windows of 11295, 11304 (NTD); 1 M, 12.49S 132.44E Nourlangie Ck. the Wetlands Point Stuart, 25.V.2000 leg. M. Langer / 6km NE. of Mt. Cahill,N.T. 2.v.73, T.Weir & T.Angeles Wilderness Lodge Lichtfang am Rand von Monsunre- / 12283 (NTD); 1 M, 6 WW, 12.46S 132.39E 12km NNW of genwald (CBP); 1 W, ARNHEM LAND NT URUPUN- Mt. Cahill, N.T. 20.v.73, E. G. Matthews (ANIC, CBM, GA JUNE 1975 W. WALSH / WALFORD-HUGGINS SAMA 25-024862-3); 1 M, 1 W, 12.46S 132.39E 12km NNW COLLEXCTION / Clivina obscuripes (Blackb.) Series det of Mt. Cahill, N.T. 20.v.73, Matthews & Upton (ANIC, by A. WALFORD-HUGGINS (CMP); 1 W, Wildman R. SAMA); 3 MM, 12.46S 132.39E 12km NNW of Mt. Ca- Cashew Porject, N.T. 12.VII.1989 Malipatil&Houston hill, N.T. 20.v.73 T.Weir & T.Angeles / 11257-59, -94 (QDPIM); 1 W, AUSTRALIEN (N.T.) LITSHFIELD (N.P.) (NTD); 2 MM, 2 WW, 12.23S 132.57E 5km NNW of Cahills 04.V.1992 leg. O.HILLERT (CWB); 2 MM, 2 WW, AUS- Crossing, N.T. East Alligator River, N.T. 28.v.73, E. G. TRALIA, Northern T. LICHTFIELD NP Adelaide River Matthews (ANIC); 1 M, 12.23S 132.56E 7km NW by N 29.V.2000 leg. M. Langer (CBM, CBP); 1 M, AUSTRALIA of Cahills Crossing (East Alligator R.) N.T. 27.v.73, N.T. Batchelor 26-27/3/93 at light/al lume Leg. L. T.Weir & N.Forrester / 12232 (NTD); 1 W, 12.23S 132.57E Toledano (CTV); 1 M, AUSTRALIA N.T. Batchelor 26- 5km NNW of Cahills Crossing (East Alligator R.) N.T. 27/3/93 Leg. Leech (SM); 1 W, Australia N.T. Batch- 87.v.73, T.Weir & N.Forrester / 12244 (NTD); 1 M, 12.06S elor Lake Bennet 30.III.1993 (CGT); 1 W, AUSTRALIA 133.04E Cooper Creek, 19km E. by S. of Mt. Borradaile, Douglas Hot Springs, 12.12.08 13°45'S, 131°26'E, 35m N.T. 31.v.1973, T.Weir & N.Forrester / 12471 (NTD); 1 M, L.Hovorka leg. (CBP); 1 M, 1 W, Burnside, N.T. May 1931 1 W, 12.06S 133.04E Cooper Creek, N.T. 19km E by SW. of HANDSCHIN (NHMB); 4 MM, 3 WW, S. Alligator River Mt. Borradaile, 31.vi.73, at light, E.G.Matthews (ANIC); area, 35 km W Jabiru, blacklight, 9.iv.1980, GFHevel & 1 W, 12.06S 133.04E Cooper Creek, N.T. 19km E by SW. JAFortin (USNM); 1 M, 12.17S 133.13E Birraduk Creek,

212 N.T. 18km E by N of Oenpelli. 4.vi.73, Upton & Feehan ric microreticulation, moderately glossy. Epipleura (ANIC); 1 M, 1 W, Tortilla Flat 11.iii.1982 J.Waldock / narrow, basally with a more or less distinct sulcus 11299-300 (NTD); 1 M, 12.48S 132.44E 8 km NE by N formed by a row of punctures. of Mt. Cahill, N.T. Nov. 26, 1974 Coll – R.I Storey at Metathoracic wings. Fully developed. light / UQIC #90342 (QMB); 1 M, Pt. Darwin (ANIC); Lower surface. Proepisternum little or not punc- 2 WW, AUSTRALIA N.T. Point Stuart Wilderness Lodge tate, with few irregular, elongate dorso-ventral stri- 26.V.2002 leg. M. Langer / S12°35,146 E131°45.661 h 25m 60km Arnhem Hwy. (CBP). oles and many short, transverse strioles, also with distinct isodiametric microreticulation. The whole Etymology. The name refers to the rufous middle and lower surface with fine, isodiametric microreticu- hind legs. lation. Metepisternum elongate, c. 2.5 × as long as Diagnosis. Comparatively small to moderately wide, in anterior half with sparse, rugose punctures. large, black species with deep, quadrangular clypeal Abdominal sterna impunctate. excision, not sulcate, barely punctate head, more or Legs. Protibia narrow and elongate, in males less pale middle and hind legs, and rather elongate, more than in females. longitudinal sulcus on upper in apical third asymmetric aedeagus with straight surface barely perceptible; lower surface with a row and very narrow, curved, at tip incurved or even of 6-8 stout setae along basal part of inner margin. hook-shaped apex. Mesotibia moderately elongate, dorsal rim crenulate, with several rows of setae on the dorsal surface; Description subapical spur elongate. Metatibia elongate, rather sparsely setose. Metatarsomere 1 almost as long as Measurements. Length: 10.6-13.5 mm; width: 2.95- the following three tarsomeres. 3.7 mm. Ratios. Length/width of pronotum: 0.99- Male genitalia (Fig. 3). Aedeagus rather wide, 1.02; base/apex of pronotum: 1.22-1.26; width fairly elongate, strongly and asymmetrically nar- pronotum/head: 1.25-1.33; length/width of elytra: rowed apicad; lower surface evenly concave. Lower 2.04-2.08; length/width of protibia: M: 3.7-3.8, W: 3.0- surface in apical half in middle slightly concave. 3.15; length/width of metatibia: M; 5.6-6.1, W: 5.6-6.0. Apical part very narrow, asymmetric, sinuate at Colour. Unicolourous black. Anterior leg dark the left side; apex very narrow, elongate, slightly piceous to black, median and posterior legs rufous, turned left, tip distinctly crotched. Both parameres considerably paler than the anterior leg. stout, the left one considerably stouter than the Head. Eye large, laterally well projected, orbit right one; both with moderately elongate, narrow, short, oblique-convex. Clypeus with deep, almost asetose, apical part. quadrangular excision. Labrum usually 5-setose, Female gonocoxites. Very similar to those of rarely 4-, 6-, or 7-setose. Clypeus not or barely C. procera Putzeys. divided from frons; upper surface smooth, without Variation. Body size rather varied. Also shape transverse sulci or strioles; frons with scattered, of pronotum varies to some degree, being more or rather fine punctures. Surface without distinct mi- less narrowed apicad. croreticulation, glossy. Pronotum. Comparatively short, considerably Distribution. Northern part of NT, particularly in narrowed apicad, dorsally rather depressed. Lateral Kakadu NP. margin oblique, faintly concave in apical third. Apex distinctly concave. Marginal channel narrow and Collecting circumstances. Little recorded. Recently, median line shallow but well impressed, impunctate. specimens were mainly collected at light. Basal sulcus narrow. Anterior marginal seta situated at apical fifth. Basal groove elongate, linear, straight, impunctate. Disk with some very fine, transverse Clivina cooindae, spec. nov. strioles in middle, not or sparsely and extremely Figs 4, 82 finely punctate, with very superficial, isodiametric microreticulation, fairly glossy. Examined types. Holotype: M, Douglas R. Crossing on Elytra. Moderately elongate, almost parallel- Oolloo Stn. Rd. NT. 26.iii.1972 A. Allwood & T. Angeles (NTD). – Paratypes: 1 W, Tindal, N.T. 14.31S 132.22E 1-20 sided, barely widened apicad, convex but on disk Dec. 1967 light trap W.J.M.Vestjens (ANIC); 7 MM, 5 WW, depressed. Basal angle little produced. Striation AUSTRALIA N.T. Kakadu N.P. 22-25/3/93 Cooinda at complete, but lateral striae less deep. Striae well im- light/al lume Leg. L. Toledano (CBM, CLT); 4 MM, 6 WW, pressed, becoming slightly shallower towards apex, AUSTRALIA N.T. Kakadu N.P. Cooinda 25-26/12/96 usually at least in basal half more or less coarsely at light/al lume Leg. L. Toledano, R. Olivieri (CBM, punctate. Intervals slightly convex, impunctate, usu- CLT); 3 MM, AUSTRALIA, N.T. Kakadu N.P.22-25/3/93 ally without transverse strioles, with fine, isodiamet- Leg. Leech (CBM, CSM); 1 W, AUSTRALIA N.T. Kakadu

213 N. Park Cooinda 20.xii.1990 (MCNV); 2 WW, AUSTRAL- Elytra (Fig. 82). Moderately elongate, almost IA, NT, Kakadu N.P., Gunlom, 13°26.02'S 132°24.85'E, parallel-sided, barely widened apicad, convex but on 60m alt, 6-8.IV.2009, light traps, leg. Vit Ryácek (CBM, disk depressed. Basal angle little produced. Striation CBP); 3 MM, AUSTRALIA, Northern T. LICHTFIELD NP complete, but lateral striae less distinct. Striae well Adelaide River 29.V.2000 leg. M. Langer (CBM, CBP); impressed, becoming slightly shallower towards 1 M, AUSTR.NT, Douglas Hot Springs, 1212.08 13°45'S apex, usually at least in basal half finely punctate. 131°26'E 35m L.Hovorka leg. (CBP); 1 W, Australia N.T. Katherine 21/12/98 leg. Daccordi (CGT); 1 W, Katherine Intervals convex, impunctate, without or with few / procera Putz. / Clivina procera Putz. (MCZ); 1 M, Pt. fine, transverse strioles, with distinct, isodiametric Darwin (ANIC); 3 MM, 13 WW, N. of Mareeba Feb.’58 microreticulation, moderately glossy. Epipleura N.Q. Darlingtons / procera Ptz. det.’59 Darlington narrow, basally with a shallow sulcus formed by a (CBM, MCZ); 3 MM, 1 W, Townsville Australia 5Feb1945 row of punctures. BMalkin / procera det.’48 Darlington (MCZ). Metathoracic wings. Fully developed. Lower surface. Proepisternum in basal half Etymology. The name refers to the locality Cooinda in Kakadu NP, where most specimens were collected. more or less distinctly, coarsely punctate, with few irregular, elongate dorso-ventral strioles and many Diagnosis. Comparatively small to moderately and short, transverse strioles, also with distinct large, black species with deep, quadrangular clypeal isodiametric microreticulation. The whole lower excision, not sulcate, densely punctate head, rufous surface with fine, isodiametric microreticulation. middle and hind legs, and stout, in apical third tri- Metepisternum elongate, c. 2.5 × as long as wide, angular aedeagus with almost straight, triangular, in anterior half with sparse, rugose punctures. Ab- at tip acute apex. dominal sterna impunctate. Legs. Protibia narrow and elongate, in males Description more than in females. longitudinal sulcus on upper surface barely perceptible; lower surface with a row Measurements. Length: 9.3-12.7 mm; width: 2.65- of 6-8 stout setae along basal part of inner margin. 3.7 mm. Ratios. Length/width of pronotum: 1.01- Mesotibia moderately elongate, dorsal rim crenulate, 1.04; base/apex of pronotum: 1.26-1.30; width with several rows of setae on the dorsal surface; pronotum/head: 1.27-1.32; length/width of elytra: subapical spur elongate. Metatibia elongate, rather 1.99-2.07; length/width of protibia: M: 3.65-3.8, W: sparsely setose. Metatarsomere 1 almost as long as 3.25-3.35; length/width of metatibia: M; 6.2-6.25, the following three tarsomeres. W: 6.0-6.1. Male genitalia (Fig. 4). Aedeagus very wide, Colour (Fig. 82). Unicolourous black. Anterior rather short, slightly asymmetrically narrowed api- leg dark piceous to black, median and posterior legs cad; lower surface in basal half deeply concave, then more or less bright rufous, considerably paler than convex, towards apex again concave. Apex triangu- the anterior leg. lar, slightly asymmetric, rather short, slightly curved Head (Fig. 82). Eye large, laterally well pro- left, tip not incurved, acute. Both parameres rather jected, orbit short, oblique-convex. Clypeus with elongate, the left one considerably stouter than the deep, almost quadrangular excision. Labrum usually right one; both with elongate, very narrow, unisetose, 5-setose. Clypeus not or barely divided from frons; apical part; setae elongate, situated right at apex; upper surface smooth, without transverse sulci or but the left paramere commonly with an additional strioles; frons very densely punctate, particularly in shorter seta on the lower surface of the apical part. anterior part. Surface without distinct microreticula- Female gonocoxites. Very similar to those of tion, moderately glossy. C. procera Putzeys. Pronotum (Fig. 82). Moderately elongate, con- Variation. Body size rather variable. Also shape siderably narrowed apicad, dorsally slightly de- of pronotum varies to some degree, being more or pressed. Lateral margin oblique, faintly concave less narrowed apicad. in apical third. Apex distinctly concave. Marginal channel narrow and median line shallow but well Distribution. Northern part of NT, particularly in impressed, impunctate. Basal sulcus narrow. An- Kakadu NP, north-eastern QLD. terior marginal seta situated at apical fifth. Basal groove elongate, linear, straight, impunctate. Disk Collecting circumstances. Little recorded. All re- with some very fine, transverse strioles in middle, cently collected specimens were sampled at light. not or extremely finely and sparsely punctate, with fine but distinct, isodiametric microreticulation, moderately glossy.

214 Clivina sinuicola, spec. nov. Pronotum. Comparatively short, considerably Fig. 5 narrowed apicad, dorsally slightly depressed. Lateral margin more or less oblique, faintly concave in api- Examined types. Holotype: M, Mornington Island Mis- cal third. Apex distinctly concave. Marginal channel sion 4 May 1960 P. Aitken N. Tindale (SAMA 25- narrow and median line shallow but well impressed, 033654). – Paratypes: 1 M, Qld, at light Mornington Is. impunctate. Basal sulcus narrow. Anterior marginal Mission 12 May,1963 P.Aitken N.B.Tindale (SAMA); seta situated at apical fifth. Basal groove elongate, 2 WW, Qld. Mornington Is. Mission At light. 28 April linear, straight, impunctate. Disk with some very 1960. P.Aitken, N.B. Tindale (SAMA); 1 W, Horn Islet, Sir Edward Pellew Group, N.T. 25-31.i.1968 B.Cantrell fine, transverse strioles in middle, not or barely / UQIC #90312 (QMB); 2 MM, 4 WW, 7/06 N-Austr, punctate, with more or less distinct, very superficial, NT, 80km w Roper Bar, 14.05.2006, 78m S14°54'18.8" isodiametric microreticulation, fairly glossy. E133°57'18.1" Light lg. Berger-Dostal (CBM, CDW); 1 Elytra. Moderately elongate, almost parallel- M, 8/06 N-Austr, NT, 40km v Port Roper, 15.05.2006, sided, barely widened apicad, convex but on disk S 14°54'04.8" E135°03'24.7" 134m, LT. Berger-Dostal depressed. Basal angle little produced. Striation (CD; W) 2 WW, Roper River, N.T. 6.iv.1976 T. Weir / complete, but lateral striae less distinct. Striae well , 11241, -65 (NTD); 1 M, 2 WW, 12/06 N-Austr. Qld. ca impressed, becoming shallower towards apex, at 50m. ca 25km E of Normanton, 19.05.2006, S17°24'54.2" least in basal half finely punctate. Intervals slightly E141°16'36.7"At Light Berger-Dostal (CBM, CDW); 1 M convex, impunctate, without or with few transverse AUS15, QLD13, Flinders R., 37 km wsw.Normanton, strioles, with fine but distinct, isodiametric micro- 12m, 17°52'35.4"S, 140°47'06.2"E, 18.4.2015, M.Baehr reticulation, moderately glossy. Epipleura narrow, (CBM); 3 MM, 2 WW, AUSTRALIA Queens- land. Flinders R., s. Normanton., u. v. - light 24-25.iv.1979 P. A. Meyer basally with a sulcus formed by a row of punctures. coll. (CBM, UASM); 2 WW, AUST: n.QLD 18km sth.Nor- Metathoracic wings. Fully developed. manton 19.Mar.2004 J.Hasenpusch T. light (QDPIM); Lower surface. Proepisternum in basal half usu- 1 M, 16.08S 136.06E 22 km WSW of Borroloola, N. T. ally slightly punctate, with few irregular, elongate 16.iv.1976, at light J.E. Feehan (ANIC); 2 WW, 15 ml. dorso-ventral strioles and many short, transverse S.W. of Normanton,Qld. 25.v.1972 G.B. & S.R.Monteith strioles, also with distinct isodiametric microreticula- /UQIC ‘90525/532 (QMB); 1 W, Walker CR., 25 ml. S.E. tion. The whole lower surface with fine, isodiametric of Karumba, Qld. 28.v.1972 G.B. & S.R.Monteith /UQIC microreticulation. Metepisternum elongate, c. 2.5 × ‘90387 (QMB); 2 WW, Gulf C.F. 23/4/93, 16.5.94 (ANIC). as long as wide, in anterior half with sparse, rugose Diagnosis. Comparatively small to large, black punctures. Abdominal sterna impunctate. species with deep, quadrangular clypeal excision, Legs. Protibia narrow and elongate, in males not sulcate, sparsely punctate head, more or less more than in females. longitudinal sulcus on upper dark rufous middle and hind legs, and in apical surface barely perceptible; lower surface with a row third very asymmetric aedeagus with rather short, of 6-8 stout setae along basal part of inner margin. slightly curved, at tip obtuse apex. Mesotibia moderately elongate, dorsal rim crenulate, with several rows of setae on the dorsal surface; Etymology. The name refers to the occurrence around subapical spur elongate. Metatibia elongate, rather the Gulf of Carpentaria. sparsely setose. Metatarsomere 1 almost as long as the following three tarsomeres. Description Male genitalia (Fig. 5). Aedeagus wide, fairly Measurements. Length: 10.1-15.9 mm; width: 2.8- elongate, narrowed apicad; lower surface deeply 4.5 mm. Ratios. Length/width of pronotum: 0.99- concave, near apex slightly bisinuate. Apical part 1.01; base/apex of pronotum: 1.22-1.33; width very asymmetric, bisinuate at the left side; apex pronotum/head: 1.28-1.35; length/width of elytra: moderately elongate, rather wide, curved left, tip 1.98-2.07; length/width of protibia: M: 3.4-3.8, W: 3.4- not incurved, obtuse. Both parameres stout but 3.5; length/width of metatibia: M; 6.25-6.6, W: 5.4-5.8. elongate, the left one considerably stouter than the Colour. Unicolourous black. Anterior leg rufo- right one; both with elongate, very narrow, unise- piceous to piceous, median and posterior legs rufous, tose, or rarely bisetose, apical part; setae very short, little paler than the anterior leg. situated right at tip. Head. Eye large, laterally well projected, orbit Female gonocoxites. Very similar to those of short, oblique-convex. Clypeus with deep, almost C. procera Putzeys. quadrangular excision. Labrum usually 5-setose. Variation. Body size very varied. Also shape of Clypeus not or barely divided from frons; upper pronotum varies to some degree, being more or less surface smooth, without transverse sulci or strioles; narrowed apicad; and punctation of proepisterna is frons with sparse punctures. Surface without distinct also somewhat varied. microreticulation, rather glossy.

215 Distribution. Area around the Gulf of Carpentaria, metric microreticulation, rather glossy. Epipleura from the south-eastern margin of Arnhem Land to narrow, without perceptible sulcus. about Normanton in north-western QLD; also on Metathoracic wings. Fully developed. Mornington Island. Lower surface. Proepisternum densely and coarsely punctate, with few irregular, elongate Collecting circumstances. Little recorded. Most dorso-ventral strioles, also with distinct isodiametric specimens apparently sampled at light. microreticulation. The whole lower surface with fine, isodiametric microreticulation. Metepisternum elongate, c. 2.25 × as long as wide, in anterior half Clivina thoracica, spec. nov. with sparse, rugose punctures, in posterior half Fig. 6, 122 with a deep, longitudinal sulcus. Abdominal sterna impunctate. Examined types. Holotype: M, Australia, QLD Nor- Legs. Protibia narrow and very elongate, teeth manton 1.1998 leg. Lamond (CBM). small, obtuse. Longitudinal sulcus on upper surface Etymology. The name refers to the unusually elongate shallow; lower surface with a row of 6-8 stout setae prothorax of the single specimen. along basal part of inner margin. Mesotibia moder- Diagnosis. Rather large, black species with deep, ately elongate, dorsal rim crenulate, with several quadrangular clypeal excision, not sulcate, sparsely rows of setae on the dorsal surface; subapical spur punctate head, dark rufous middle and hind legs, elongate. Metatibia elongate, rather sparsely setose. and in apical third very asymmetric aedeagus with Metatarsomere 1 slightly shorter than the following elongate, markedly sinuate, at tip crotched apex. three tarsomeres. Male genitalia (Fig. 6). Aedeagus wide, fairly Description elongate, narrowed apicad; lower surface in basal half concave, in apical third markedly convex, to- Measurements. Length: 14.5 mm; width: 4.0 mm. wards apex again deeply concave. Apical part very Ratios. Length/width of pronotum: 1.11; base/ asymmetric, deeply sinuate at the left side; apex apex of pronotum: 1.27; width pronotum/head: elongate, very narrow, bisinuate and at tip turned 1.32; length/width of elytra: 2.07; length/width of left, asymmetric, tip markedly curved down and protibia: M: 4.35; length/width of metatibia: M; 6.7. crotched, acute. Both parameres stout but elongate, Colour (Fig. 122). Unicolourous black. Anterior the left one considerably stouter than the right one; leg piceous, median and posterior legs dark rufous, both with rather elongate, very narrow, unisetose little paler than the anterior leg. apical part; setae very short, situated right at tip; but Head (Fig. 122). Eye large, laterally well pro- right paramere in some specimens with an additional jected, orbit short, oblique-convex. Clypeus with seta at the lower surface near apex. deep, almost quadrangular excision. Labrum 5-se- Female gonocoxites. Unknown. tose. Clypeus not divided from frons; upper surface Variation. Unknown. without transverse sulci or strioles; frons with sparse punctures. Surface without distinct microreticula- Distribution. North-western QLD. Known only tion, rather glossy. from type locality. Pronotum (Fig. 122). Elongate, considerably narrowed apicad, dorsally depressed. Lateral margin Collecting circumstances. Not recorded. oblique, faintly concave in apical third. Apex distinctly concave. Marginal channel narrow, median Clivina gemina, spec. nov. line shallow but well impressed, impunctate. Basal sulcus narrow. Anterior marginal seta situated at This species apparently occurs in two populations apical fifth. Basal groove elongate, linear, straight, which are only distinguished by the colour of their impunctate. Disk with some very fine, transverse middle and hind legs, but are geographically well strioles in middle, impunctate, very fine, superficial, separated. isodiametric microreticulation, fairly glossy. Elytra. Elongate, almost parallel-sided, barely widened apicad, convex but on disk depressed. Basal Clivina gemina gemina, spec. nov. angle little produced. Striation complete, but lateral Figs 7, 83 striae less distinct. Striae well impressed, becoming shallower towards apex, at least in basal half finely Examined types. Holotype: M, Wyndham WA 20.iv. punctate. Intervals slightly convex, impunctate, with 1976 K.&.E. Carnaby (ANIC). – Paratypes: 12 MM, 13 WW, few transverse strioles, with fine, superficial, isodia- Kakadu N.P., Gunlom, 13°26.02'S 132°24.85'E, 60m alt,

216 6-8.IV.2009, light traps, leg. Vit Ryácek (CBM, CBP, CRP); 2.5-2.10; length/width of protibia: M: 3.5-3.8, W: 2.75- 1 M, 3 WW, same data (ANIC); 1 W, Australia,NT,Gregory 2.85; length/width of metatibia: M; 6.3-6.8, W: 5.8-6.1. NP Victoria.R.Inn,24.IV.2004 16.36.95S, 131.07.73E 62m, Colour (Fig. 83). Unicolourous black. Anterior leg.M.Langer (CBM); 3 MM, 3 WW, AUSTRALIA N.T. m50 leg more or less dark piceous, median and posterior Victoria River Road House 20-21/24-25.III.1996 P. M. legs rufous, considerably paler than the anterior leg. Giachino leg. (CGT); 1 W, AUSTRALIA N.T. Victoria Head (Fig. 83). Eye large, laterally well pro- Crossing 28-29/3/93 at light/al lume Leg. L. Toledano (CTV); 1 W, AUSTRALIA N.T. Top Springs 29.XII.1998 jected, orbit short, oblique-convex. Clypeus with 180 km W of Dunmurra M. Daccordi leg. (CGT); 1 W, deep, almost quadrangular excision. Labrum usually Australia/NT Keep River N.P.: Gurandalg Camp, ca 5-setose. Clypeus not or barely divided from frons; 50 km E Kununurra 17. + 23.IV.2004 lgt. M.Langer / S upper surface smooth, without transverse sulci or 15.52.494 E 129.03.076 Höhe 99 m Lichtfang (CBM); 1 W, strioles; frons finely and sparsely punctate. Surface 15.44S 129.09E GPS 5.5km NE by E Jarnarm Keep River without distinct microreticulation, glossy. Nat. Pk. 27 May 2001 T.Weir, P.Bouchard / at light Pronotum (Fig. 83). Moderately elongate, con- open forest (ANIC); 1 M, 15.45S 129.05E Jarnarm Camp siderably narrowed apicad, dorsally moderately Keep River N.P. 5 jun. 2001 A. Calder at black light convex. Lateral margin oblique, faintly concave (ANIC); 3 MM, 11 WW, 100 mi. E. of Kununurra, W.A., in apical third. Apex distinctly concave. Marginal light trap, 27.3.66 J. A. Mahon (ANIC, CBM); 10 MM, channel narrow and median line shallow but well 12 WW, AUSTRALIA: WA: Kimberley 12m alt. Home Valley Station 14°42'S 127°51'E; 15-18.iv.2009 nr. Pen- impressed, impunctate. Basal sulcus narrow. Ante- tecoast river, S. Jakl lgt. (CBM, CBP, CJP); 11 MM, 4 WW, rior marginal seta situated at apical fifth. Basal groove AUSTRALIA: WA; Kimberley, Wyndham env., 20-21. elongate, linear, straight, impunctate. Disk without iv.2009, 15°31'S 128°10'E, salty bay of the sea, 3m alt. St. or with some very fine, transverse strioles in mid- Jakl lgt. (CBM, CBP, CJP); 1 W, AUSTRALIA: WA: 15-16. dle, barely punctate, with very fine and superficial, iv.2009; Hidden Valley NP Kununurra 15°46S 128°44E isodiametric microreticulation, fairly glossy. 64m alt. St. Jakl lgt. (CJP); 2 MM, AUSTRALIA: WA; Kim- Elytra (Fig. 83). Moderately elongate, almost berley, Home Valley St., 15°42'29"S 127°51'11"E, 12m parallel-sided, barely widened apicad, convex but on alt, 16-18IV.2009, light traps, leg. Vít Ryjácek (CRP); disk depressed. Basal angle little produced. Striation 2 MM, Wyndham-KRS 18.5.58.R.Lukins (WADAP); 1 M, complete, but lateral striae less distinct. Striae well Kimberley Res. Stn. 7.1.53 R.Lukins (WADAP); 1 W, WA impressed, becoming slightly shallower towards Wyndham 20 Apr 1975 Ex. Carnaby Coll. (ANIC); 3 WW, S15.64975 E128.71519 light trap in plowed field FWI, apex, usually at least in basal half finely punctate. Kununurra, W. Aust. 25.iii.2003 C.Y.Norwood (Dept Intervals slightly convex, impunctate, usually with- Agr WA 122165-6, 122181); 1 M, 3 WW, Australia/WA out transverse strioles, with fine, isodiametric micro- East Kimberley: Parry Creek Road, Parry Creek Farm reticulation, moderately glossy. Epipleura narrow, 22.IV.2004 lgt. M. Langer / S 15.35.882 E 128.16.7171 basally with a sulcus formed by a row of punctures. Höhe 33 m Lichtfang (CBM, CBP); 1 W, Australia, WA., Metathoracic wings. Fully developed. The Kimberley Parry Creek Farm , Parry Lagoons Nat. Lower surface. Proepisternum densely punctate, Res. 20 km S Wyndham / leg. M. Langer 15°35'47.3" with few irregular, elongate dorso-ventral strioles E 128°16'45.2" H = 11 m ( NF) (CWB); 1 W, Australia, and many and short, transverse strioles, also with WA, Parry Creek Farm, 20 km s.Wyndham 15°35'47.2"S. distinct isodiametric microreticulation. The whole 128°16'45.7"E 15.V.2009, 11m, M.Langer (CBM); 1 W, WA lower surface with fine, isodiametric microreticu- Wyndham 20 Apr 1975 Ex. Carnaby Coll. (ANIC). lation. Metepisternum elongate, c. 2.5 × as long as Etymology. The name refers to the similarity to C. pro- wide, in anterior half with sparse, rugose punctures. cera. Abdominal sterna impunctate. Legs. Protibia narrow and elongate, in males Diagnosis. Medium sized to rather large, black more than in females. longitudinal sulcus on upper species with deep, quadrangular clypeal excision, surface barely perceptible; lower surface with a row not sulcate, sparsely punctate head, rufous mid- of 6-8 stout setae along basal part of inner margin. dle and hind legs, densely and coarsely punctate Mesotibia moderately elongate, dorsal rim crenulate, proepisternum, and in apical third very asymmetric with several rows of setae on the dorsal surface; aedeagus with elongate, sinuate, at tip incurved or subapical spur elongate. Metatibia elongate, rather crotched apex. sparsely setose. Metatarsomere 1 slightly shorter Description than the following three tarsomeres. Male genitalia (Fig. 7). Aedeagus wide, fairly Measurements. Length: 11.3-13.5 mm; width: 2.8- elongate, narrowed apicad; lower surface in basal 3.9 mm. Ratios. Length/width of pronotum: 1.02- half deeply concave then convex, towards apex 1.05; base/apex of pronotum: 1.23-1.27; width again concave. Apical part very asymmetric, deeply pronotum/head: 1.26-1.34; length/width of elytra: bisinuate at the left side; apex elongate, narrow,

217 slightly turned left, slightly asymmetric, tip incurved W: 2.7-2.8; length/width of metatibia: M; 6.5-6.7, or slightly crotched, acute. Both parameres very W: 6.1-6.2. elongate, the left one considerably stouter than the Colour. Unicolourous black. All legs deep black. narrower right one; both with elongate, very narrow, Head. As in the nominate subspecies. asetose or unisetose, apical part; setae, of present, Pronotum. Much as in the nominate subspecies, very short, situated right at tip. but usually slightly more narrowed apicad. Female gonocoxites. Very similar to those of Elytra. As in the nominate subspecies. C. procera Putzeys. Metathoracic wings. Fully developed. Variation. Body size somewhat varied. Other- Lower surface. As in the nominate subspecies. wise little variation noted. Legs. Much as in the nominate subspecies. Male genitalia (Fig. 8). Much as in the nominate Distribution. Northern part of NT to north-eastern subspecies. Kimberleys in WA. Female gonocoxites. As in the nominate subspecies. Collecting circumstances. Almost all specimens Variation. Body size somewhat varied. Also were collected at light near rivers and billabongs. shape of pronotum varies to some degree, being more or less narrowed apicad. Clivina gemina nigripes, subspec. nov. Distribution. Southern part of the Kimberley Divi- Fig. 8 sion. The single specimen from “Armidale” certainly Examined types. Holotype: M, Fitzroy River W.A. is wrongly labelled. at light 16 March 1980 K. & E. Carnaby (ANIC). – Collecting circumstances. Little recorded, almost all Paratypes: 4 MM, 7 WW, same data (ANIC, CBM); 2 MM, specimens collected near Fitzroy River. Few more 3 WW, WESTERN AUSTRALIA Fitzroy River 16-4-1976 recently collected specimens sampled at light. (AMS, ANIC); 2 MM, WESTERN AUSTRALIA Fitzroy River 16-4-1980 (ANIC); 3 WW, W.Australia Fitzroy River 16.IV.1976 K.&E.Carnaby (ANIC); 2 MM, 2 WW, WA Fitzroy River 16 Apr 1976 Ex Carnaby Coll (ANIC); Clivina carnabyi, spec. nov. 2 MM, WA Fitzroy River 16 Apr 1980 K&E Carnaby Ex. Fig. 9 Carnaby Coll. (ANIC); 2 WW, WA Fitzroy River 16 Apr 1973 K&E Carnaby Ex. Carnaby Coll. (ANIC); 3 WW, Examined types. Holotype: M, Fitzroy Crossing, W.A WA Fitzroy River at light 16 Apr 1967 Ex. Carnaby 125:35E, 18:10S 17 Mar. 1984, at light K. & E. Carnaby Coll. (ANIC); 1 W, WA Fitzroy River 16 Apr 1980 K&E (ANIC). – Paratypes: 1 W, same data (CBM); 2 WW, Wil- Carnaby Ex. Carnaby Coll. (ANIC); 1 W, WA Fitzroy lare Bridge on Fitzroy River, W.A. 28 Feb.1985 at light River at light 16 Apr 1967 Ex. Carnaby Coll. (ANIC); K. & E. Carnaby (ANIC); 1 W, W.AUST., KIMBERLEY 1 W, WA Fitzroy River 16 Apr 1980 K&E Carnaby Ex. LENNARD RIVER XING GIBB RIVER ROAD 17.23S, Carnaby Coll. (ANIC); 2 WW, Fitzroy R. NWA 16 April 124.44E 1 APRIL 1988 T. F. HOUSTON 679-4 / AT 1976 K&E Carnaby (ANIC); 2 MM, 2 WW, Fitzroy Cross- LIGHT(MV) AT NIGHT (WAM 39158); 1 W, 16.28S ing WA 18.iv.76, at light K.&E.Carnaby (ANIC); 4 WW, 124.51E WA Walcott Inlet. CALM-SITE 19/2. 17-19 WA Fitzroy Crossing 16 Apr 1974 Ex. Carnaby Coll. Jun. 1988 J. Majer, closed forest (ANIC); 1 M, 1 W, Fitz- (ANIC); 1 M, 1 W, WA Fitzroy Crossing 23 Mar 1984 Ex. roy River W.A. at light 16 March 1980 K. & E. Carnaby Carnaby Coll. (ANIC); 1 M, 1 M, Australien,WA Fitzroy (ANIC, CBM); 1 M, WA Fitzroy Crossing 23 Mar 1984 Crossing 19.11.1984 M. Baehr (CBM); 1 M, 2 WW, Derby, Ex. Carnaby Coll. (ANIC); N.W.A. W. D. Dodd M (SAMA 25-033408/411/520); 1 W, Etymology. The name is a patronym in honour of the NSW Armidale 9 Dec1971 K&E Carnaby Ex. Carnaby collectors of most specimens, K. and E. Carnaby. Coll. (ANIC). Diagnosis. Large, black species with deep, quad- Etymology. The name refers to the black legs. rangular clypeal excision, transversely sulcate, Diagnosis. Similar to the nominate subspecies, but sparsely punctate head, deep black legs, not or all legs deep black. feebly punctate proepisternum, and in apical third asymmetric aedeagus with elongate, sinuate, at tip Description not incurved apex. Measurements. Length: 11.7-13.9 mm; width: 3.25- Description 3.9 mm. Ratios. Length/width of pronotum: 1.01- 1.03; base/apex of pronotum: 1.29-1.35; width Measurements. Length: 15.2-16.6 mm; width: 4.1- pronotum/head: 1.34-1.38; length/width of elytra: 4.55 mm. Ratios. Length/width of pronotum: 1.02- 2.03-2.09; length/width of protibia: M: 3.35-3.6, 1.04; base/apex of pronotum: 1.32-1.36; width

218 pronotum/head: 1.39-1.44; length/width of elytra: incurved, acute. Both parameres rather stout, the 2.09-2.15; length/width of protibia: M: 3.3-3.5, W: 3.0- left one considerably stouter than the right one; 3.1; length/width of metatibia: M; 6.35-6.5, W: 5.9-6.0. both with moderately elongate, very narrow, asetose Colour. Unicolourous black. All legs deep black. apical part. Head. Eye large, laterally well projected, orbit Female gonocoxites. Very similar to those of short, oblique-convex. Clypeus with deep, almost C. windjanae, spec. nov. quadrangular excision. Labrum usually 5-setose. Variation. Rather little variation noted, except Clypeus shallowly divided from frons; upper surface for length of elytra. with some more or less deep and complete transverse sulci; sparsely punctate. Surface without distinct Distribution. Western and southern part of the microreticulation, glossy. Kimberley Division. Pronotum. Moderately elongate, considerably Collecting circumstances. Most specimens were narrowed apicad, dorsally gently convex. Lateral collected at light near rivers. margin oblique, faintly concave in apical third. Apex distinctly concave. Marginal channel narrow and median line shallow but well impressed, impunctate. Clivina windjanae, spec. nov. Basal sulcus narrow. Anterior marginal seta situated Figs 10, 55 at apical fifth. Basal groove elongate, linear, straight, impunctate. Disk with some very fine, transverse Examined types. Holotype: M, Windjana Gorge W.A. 17 strioles in middle, extremely finely punctate or Mar. 1980 at light K. & E. Carnaby (ANIC). – Paratypes: impunctate, with fine, superficial, isodiametric 2 MM, 4 WW, same data (ANIC, CBM); 3 MM, 3 WW, Lennard microreticulation, fairly glossy. River W.A. at light 18 March 1980 K. & E. Carnaby Elytra. Elongate, almost parallel-sided, barely (ANIC, CBM); 1 M, Fitzroy River W.A. at light 16 March 1980 K. & E. Carnaby (ANIC). widened apicad, convex but on disk depressed. Basal angle little produced. Striation complete, but Etymology. The name refers to the type locality, Wind- lateral striae less distinct. Striae well impressed, jana Gorge in the Kimberley Division. becoming slightly shallower towards apex, at least in Diagnosis. Medium sized, black species with deep, basal half finely punctate. Intervals slightly convex, quadrangular clypeal excision, not sulcate, sparsely impunctate, usually without transverse strioles, with punctate head, deep black legs, not or feebly punc- fine, very superficial, isodiametric microreticulation, tate proepisternum, and in apical third asymmetric glossy. Epipleura narrow, basally with a shallow aedeagus with elongate, sinuate, at tip incurved or sulcus formed by a row of punctures. crotched apex. Metathoracic wings. Fully developed. Lower surface. Proepisternum usually barely Description punctate, with few irregular, elongate dorso-ventral strioles and many and short, transverse strioles, Measurements. Length: 12.0-13.4 mm; width: 3.4- also with distinct isodiametric microreticulation. 3.8 mm. Ratios. Length/width of pronotum: 0.97- The whole lower surface with fine, isodiametric 1.03; base/apex of pronotum: 1.32-1.36; width microreticulation. Metepisternum elongate, slightly pronotum/head: 1.30-1.38; length/width of elytra: > 2.5 × as long as wide, in anterior half with sparse, 2.03-2.07; length/width of protibia: M: 3.4-3.5, W: 2.8- rugose punctures. Abdominal sterna impunctate. 3.0; length/width of metatibia: M; 6.2-6.25, W: 6.1-6.2. Legs. Protibia narrow and elongate, in males Colour. Unicolourous black. All legs deep black. more than in females. longitudinal sulcus on upper Head. Eye large, laterally well projected, orbit surface barely perceptible; lower surface with a row short, oblique-convex. Clypeus with deep, almost of 6-8 stout setae along basal part of inner margin. quadrangular excision. Labrum 5-setose. Clypeus Mesotibia moderately elongate, dorsal rim crenulate, not divided from frons; upper surface without with several rows of setae on the dorsal surface; transverse sulci; sparsely punctate. Surface without subapical spur elongate. Metatibia elongate, rather distinct microreticulation, glossy. sparsely setose. Metatarsomere 1 slightly shorter Pronotum. Moderately elongate, considerably than the following three tarsomeres. narrowed apicad, dorsally gently convex. Lateral Male genitalia (Fig. 9). Aedeagus very wide, margin oblique, faintly concave in apical third. Apex fairly elongate, narrowed apicad; lower surface in distinctly concave. Marginal channel narrow and basal part concave, in apical third gently convex, median line shallow but well impressed, impunctate. towards apex again concave. Apical part very asym- Basal sulcus narrow. Anterior marginal seta situated metric, slightly bisinuate at the left side; apex fairly at apical fifth. Basal groove elongate, linear, straight, elongate, markedly turned left, asymmetric, tip not impunctate. Disk with some very fine, transverse

219 strioles in middle, extremely finely punctate or Collecting circumstances. All specimens were col- impunctate, without microreticulation or with fin- lected at light near rivers. est traces of extremely, superficial, isodiametric microreticulation, glossy. Elytra. Fairly elongate, almost parallel-sid- Clivina mahoni, spec. nov. ed, barely widened apicad, convex but on disk Fig. 11 depressed. Basal angle little produced. Striation complete, but lateral striae less distinct. Striae well Examined types. Holotype: M, 100 mi. E. of Kununurra, impressed, becoming slightly shallower towards W.A., light trap, 27.3.66 J. A. Mahon (ANIC). – Paratype: apex, almost impunctate. Intervals slightly convex, 1 W, same data (CBM). impunctate, without or with very weak transverse Etymology. The name is a patronym in honour of the strioles, with fine, extremely superficial traces of collector, J. A. Mahon. isodiametric microreticulation, glossy. Epipleura narrow, basally with a shallow sulcus formed by a Diagnosis. Large, black species with deep, quad- row of punctures. rangular clypeal excision, not sulcate, sparsely Metathoracic wings. Fully developed. punctate head, deep black legs, not or feebly punc- Lower surface. Proepisternum usually barely tate proepisternum, and in apical almost symmetric punctate, with few irregular, elongate dorso-ventral aedeagus with straight, parallel-sided, at tip slightly strioles and many and short, transverse strioles, incurved apex. also with distinct isodiametric microreticulation. The whole lower surface with fine, isodiametric Description microreticulation. Metepisternum elongate, c. 2.5 × Measurements. Length: 15.5-16.4 mm; width: 4.35- as long as wide, in anterior half with sparse, rugose 4.5 mm. Ratios. Length/width of pronotum: 1.03- punctures. Abdominal sterna impunctate. 1.08; base/apex of pronotum: 1.33-1.36; width Legs. Protibia narrow and elongate, in males pronotum/head: 1.36-1.39; length/width of elytra: more than in females. Longitudinal sulcus on upper 2.09-2.11; length/width of protibia: M: 4.5, W: 3.4; surface barely perceptible; lower surface with a row length/width of metatibia: M; 5.7, W: 5.6. of 6-8 stout setae along basal part of inner margin. Colour. Unicolourous black. All legs deep black. Mesotibia moderately elongate, dorsal rim crenulate, Head. Eye large, laterally well projected, orbit with several rows of setae on the dorsal surface; short, oblique-convex. Clypeus with deep, almost subapical spur elongate. Metatibia elongate, rather quadrangular excision. Labrum 5-setose. Clypeus not sparsely setose. Metatarsomere 1 slightly shorter divided from frons; upper surface smooth, without than the following three tarsomeres. transverse sulci or strioles; frons sparsely punctate. Male genitalia (Fig. 10). Aedeagus wide, fairly Surface without distinct microreticulation, glossy. elongate, narrowed apicad; lower surface regularly Pronotum. Rather elongate, considerably nar- concave. Apical part asymmetric, slightly bisinuate rowed apicad, dorsally gently convex. Lateral mar- at the left side; apex rather elongate, slightly turned gin oblique, faintly concave in apical third. Apex left, asymmetric, tip incurved or slightly crotched, distinctly concave. Marginal channel narrow and rather acute. Both parameres short and stout, the left median line shallow but well impressed, impunctate. one considerably stouter than the right one; both with Basal sulcus narrow. Anterior marginal seta situated very short, asetose or unisetose apical part; setae, if at apical fifth. Basal groove elongate, linear, straight, present, extremely short, situated right at tip. impunctate. Disk with some very fine, transverse Female gonocoxites (Fig. 55). Gonocoxites nar- strioles in middle, impunctate, with very fine, row and elongate, little curved, with rather acute though rather distinct, isodiametric microreticula- apex; with one elongate, rather stout seta basally at tion, fairly glossy. lateral surface and a smaller seta below, two elongate Elytra. Elongate, almost parallel-sided, barely setae on the ventral surface of gonocoxite 1, three widened apicad, convex but on disk depressed. Basal setae at middle of the medio-dorsal surface, but angle little produced. Striation complete, but lateral without a nematiform seta near apex. Lateral plate striae less distinct. Striae well impressed, becoming basally with one elongate seta, apically with three slightly shallower towards apex, at least in basal half or four shorter seta. finely punctate. Intervals slightly convex, impunc- Variation. Little varied. Only shape of pronotum tate, without or with very weak transverse strioles, and length of elytra vary slightly. with fine, isodiametric microreticulation, moderately glossy. Epipleura narrow, basal sulcus indistinct. Distribution. Western and southern part of the Metathoracic wings. Fully developed. Kimberley Division.

220 Lower surface. Proepisternum slightly punctate apex of pronotum: 1.20; width pronotum/head: in ventral half, with few irregular, elongate dorso- 1.22; length/width of elytra: 2.22; length/width of ventral strioles and many and short, transverse protibia: W: 2.8; length/width of metatibia: W: 7.3. strioles, also with distinct isodiametric microreticula- Colour (Fig. 84). Unicolourous rufous, probably tion. The whole lower surface with fine, isodiametric because of immaturity. Anterior leg piceous, median microreticulation. Metepisternum elongate, slightly and posterior legs rufous, considerably paler than > 2.5 × as long as wide, in anterior half with sparse, the anterior leg. rugose punctures. Abdominal sterna impunctate. Head (Fig. 84). Large in comparison with the Legs. Protibia narrow and elongate, in the male prothorax. Eye very large, almost semicircular, lat- more than in the female. Longitudinal sulcus on erad far projected, orbit very short, oblique-convex. upper surface barely perceptible; lower surface with Clypeus with deep, almost quadrangular excision. a row of 6-8 stout setae along basal part of inner Labrum 7-setose. Clypeus shallowly divided from margin. Mesotibia moderately elongate, dorsal rim frons; upper surface smooth, without transverse sulci crenulate, with several rows of setae on the dorsal or strioles; frons in middle with a distinct, oval im- surface; subapical spur elongate. Metatibia elongate, pression. Surface with dense, rather fine punctures, rather sparsely setose. Metatarsomere 1 slightly without microreticulation, glossy. shorter than the following three tarsomeres. Pronotum (Fig. 84). Comparatively elongate, Male genitalia (Fig. 11). Aedeagus very wide, almost parallel-sided, dorsally rather depressed. fairly elongate, narrowed apicad; lower surface con- Lateral margin almost straight, barely concave in cave, in apical third slightly convex, towards apex apical third. Apex feebly concave. Marginal channel again concave. Apical part almost symmetric, on narrow. Median line shallow but well impressed, both sides bisinuate; apex narrow, elongate, straight, impunctate. Basal sulcus narrow. Anterior marginal parallel-sided, tip slightly incurved, obtuse. Both seta situated slightly behind apical fifth. Basal groove parameres remarkably stout, the left one consider- elongate, linear, straight, impunctate. Disk with ably stouter than the right one; both with elongate, several fine, transverse strioles in middle, extremely very narrow, asetose apical part. finely but rather densely punctate, without microre- Female gonocoxites. Very similar to those of ticulation, very glossy. C. windjanae, spec. nov. Elytra (Fig. 84). Very elongate, parallel-sid- Variation. Little variation noted. ed, barely widened apicad, convex but on disk depressed. Basal angle little produced. Striation Distribution. North-western NT. Know only from complete, but lateral striae less distinct. Striae well the type locality. impressed, becoming slightly shallower towards apex, almost impunctate. Intervals slightly convex, Collecting circumstances. Both specimens were sparsely and extremely finely punctate, without collected in light trap. transverse strioles, with finest, almost invisible traces of isodiametric microreticulation, very glossy. Epipleura narrow, basally with a sulcus formed by Clivina montisbelli, spec. nov. a row of punctures. Fig. 84 Metathoracic wings. Fully developed. Lower surface. Proepisternum moderately Examined types. Holotype: W, W.AUST. KIMBERLEY densely punctate, with few irregular, elongate FERN CK NR MT BELL KING LEOPOLD RANGE 10 dorso-ventral strioles and many and short, transverse APRIL 1988 T. F. HOUSTON 688-4 / AT MV LIGHT AT strioles, also with distinct isodiametric microreticula- NIGHT (6.-7.40 PM) (WAM39176). tion. The whole lower surface with fine, isodiametric Etymology. The name refers to the type locality, Mt. microreticulation. Abdominal sterna in middle with Bell in the Kimberley Division. extremely fine, laterally coarser punctures. Metepis- Diagnosis. Medium sized, rufous species with deep, ternum very elongate, almost 3 × as long as wide, in quadrangular clypeal excision, not sulcate, densely anterior half with some dorso-ventral sulci. punctate head, rufous middle and hind legs, slightly Legs. Protibia rather wide. Longitudinal sulcus punctate proepisternum, and very elongate and on upper surface very distinct; lower surface with a remarkably parallel-sided elytra. row of 6 stout setae along basal part of inner margin. Mesotibia elongate, dorsal rim crenulate, with several Description rows of setae on the dorsal surface; subapical spur elongate. Metatibia very elongate, rather sparsely Measurements. Length: 12.1 mm; width: 3.15 mm. setose. Metatarsus remarkably elongate, tarsomere Ratios. Length/width of pronotum: 1.06; base/ 1 almost as long as the following three tarsomeres.

221 Male genitalia. Unknown. traces of about isodiametric microreticulation that is Female gonocoxites. Very similar to those of only perceptible at very high magnification; surface C. windjanae, spec. nov. rather glossy. Variation. Unknown. Elytra. Elongate, almost parallel-sided, barely widened apicad, convex but on disk depressed. Distribution. Central-western Kimberley Division. Basal angle little produced. Striation complete, but Known only from type locality. lateral striae less distinctly impressed. Striae well impressed, becoming slightly shallower towards Collecting circumstances. The holotype was col- apex, at least in basal half finely punctate. Intervals lected at light. slightly convex, impunctate, without or with very weak transverse strioles, with extremely superficial, isodiametric microreticulation that is only percep- Clivina victoriae, spec. nov. tible at very high magnification; surface glossy. Fig. 12 Epipleura narrow, basal sulcus indistinct. Examined types. Holotype: M, Victoria River Downs Metathoracic wings. Fully developed. 3 km SW of Stn NT 6 Aug. 1979 L.P.Kelsey at light Lower surface. Proepisternum impunctate, with (ANIC). – Paratypes: 2 MM, same data (ANIC, CBM); few irregular, elongate dorso-ventral strioles and 1 M, 6.4km W-SW of Victoria Riv. Downs, N.T. 13 June, many short, transverse strioles, also with distinct 1973 L.F. Kelsey (ANIC); 1 W, 8km ENE of Victoria Riv. isodiametric microreticulation. The whole lower Downs, N.T. 12 July, 1973 L.F. Kelsey (ANIC). surface with fine, isodiametric microreticulation. Metepisternum elongate, slightly > 2 × as long as Etymology. The name refers to the type locality, Vic- toria River Downs. wide, in anterior half with sparse, rugose punctures. Abdominal sterna impunctate. Diagnosis. Medium sized, black species with deep, Legs. Protibia narrow and elongate, in the male quadrangular clypeal excision, not sulcate, lit- more than in the female. Longitudinal sulcus on tle punctate head, rufous middle and hind legs, upper surface barely perceptible; lower surface with impunctate proepisternum, elongate and parallel- a row of 6-8 stout setae along basal part of inner sided elytra, and remarkably incurved apex of the margin. Mesotibia moderately elongate, dorsal rim aedeagus. crenulate, with several rows of setae on the dorsal surface; subapical spur elongate. Metatibia elongate, Description rather sparsely setose. Metatarsomere 1 as long as the following three tarsomeres. Measurements. Length: 12.3-13.3 mm; width: 3.4- Male genitalia (Fig. 12). Aedeagus wide, fairly 3.7 mm. Ratios. Length/width of pronotum: 0.99- elongate, narrowed apicad; lower surface in basal 1.01; base/apex of pronotum: 1.21-1.25; width half concave, in apical third slightly convex, to- pronotum/head: 1.28-1.32; length/width of elytra: wards apex again remarkably concave. Apical part 2.0-2.05; length/width of protibia: M: 2.85-3.25, asymmetric, slightly bisinuate at the left side; apex W: 2.75; length/width of metatibia: M; 5.7-5.9, W: 5.55. elongate, almost straight, remarkably curved down, Colour. Unicolourous black. Middle and hind tip slightly incurved, rather acute. Both parameres legs rufous, paler than the anterior legs. stout, the left one considerably stouter than the right Head. Eye large, laterally well projected, orbit one; both with fairly elongate, very narrow, asetose short, oblique-convex. Clypeus with deep, almost apical part. quadrangular excision. Labrum 5-setose. Clypeus not Female gonocoxites. Very similar to those of divided from frons; upper surface smooth, without C. windjanae, spec. nov. transverse sulci or strioles; frons sparsely punctate. Variation. Very little variation noted. Surface without distinct microreticulation, glossy. Pronotum. Moderately elongate, little narrowed Distribution. North-western NT. Only recorded apicad, dorsally gently convex. Lateral margin from a very limited area. oblique, faintly concave in apical third. Apex distinctly concave. Marginal channel narrow and me- Collecting circumstances. All specimens collected dian line shallow but well impressed, impunctate. at light near a large river. Basal sulcus narrow. Anterior marginal seta situated in front of apical fifth. Basal groove elongate, linear, straight, impunctate. Disk without transverse strioles, impunctate, with extremely fine and superficial

222 Clivina ryaceki, spec. nov. and remarkably curved aedeagus with elongate, Fig. 13, 123 straight, triagonal apex.

Examined types. Holotype: M, 15.46S 129.0E NT GPS Description Jarnarm Keep River N.P. 5 Jun. 2001 A. Calder at black Measurements. Length: 15.2-18.9 mm; width: 4.2- light (ANIC). – Paratypes: 4 MM, 7 WW, same data (ANIC, CBM); 2 MM, 3 WW, 15.46S 129.06E NT GPS Jarnarm 5.3 mm. Ratios. Length/width of pronotum: 1.0-1.03; Keep River Nat.Pk. 8 Jun. 2001 A. Calder Malaise trap base/apex of pronotum: 1.25-1.30; width pronotum/ (ANIC); 5 MM, 7 WW, 15.46S 129.0E NT GPS Jarnarm Keep head: 1.34-1.38; length/width of elytra: 2.06-2.12; River N.P. 5 Jun. 2001 A. Calder at black light (ANIC, length/width of protibia: M: 4.15-4.2, W: 3.3-3.5; CBM); 4 WW, 15.46S 129.10E NT GPS spring at 8 km E length/width of metatibia: M; 5.8-6.1, W: 5.7-5.9. by N Jarnarm Keep River Nat.Pk 26 - 28 May 2001 Colour (Fig. 123). Unicolourous black. Middle T.Weir, P.Bouchard (ANIC); 2 MM, 1 W, 15.44S 129.09E and hind legs rufo-piceous, perceptibly paler than NT GPS 5.5 km NE by E Jarnarm, Keep River Nat. the anterior leg. Pk, 27 May 2001 T.Weir, P.Bouchard (ANIC); 3 WW, Head (Fig. 123). Eye large, laterally well pro- Keep Creek, N.T. Victoria Hwy 29 Mar. 1984, at light. jected, orbit short, oblique-convex. Clypeus with K. & E. Carnaby (ANIC); 1 M, Keep River, N.T. 20.3.84 deep, almost quadrangular excision. Labrum 5-se- Coll Carnaby / A. Hiller Coll (QMB); 2 WW, NT Keep River NP Gorge carpark 15.50S, 129.06E 26 April 1996 tose. Clypeus not divided from frons; upper surface G.R.Brown mv lamp (NTD); 13 MM, 12 WW, AUSTRALIA, smooth, without transverse sulci or strioles; frons NT, Kakadu N.P., Gunlom, 13°26.02'S 132°24.85'E, 60m sparsely punctate. Surface without distinct micro- alt, 6-8.IV.2009, light traps, leg. Vit Ryácek (CBM, CRP); reticulation, glossy. 2 WW, Port Darwin N. Territory (SAMA 25-033607-8); Pronotum (Fig. 123). Moderately elongate, 1 M, 12.36S 132.52E Magela Creek, N.T. 1lm NNW of slightly narrowed apicad, dorsally gently convex. Mudginbarry HS. 25.v.73, Matthews & Upton (ANIC); Lateral margin oblique, faintly concave in apical 1 M, Australia/N.T. Keep River N.P.: Gurandalg Camp, third. Apex distinctly concave. Marginal channel ca 50 km E Kununurra 17. + 23.IV.2004 lgt. M.Langer / narrow and median line shallow but well impressed, S 15.52.494 E 129.03.076 Höhe 99 m Lichtfang (CBP); impunctate. Basal sulcus narrow. Anterior marginal 1 M, 2 WW, WYNDHAM. NWA 2ß.IV.76 K.CARNABY / seta situated at apical fifth. Basal groove elongate, WALFORD-HUGGINS COLLECTION / Clivina procera Putzeys [Series det. by A. Walford-Huggins] (CMP); linear, straight, impunctate. Disk with some very 1 M, 1 W, WA Wyndham 20 Apr. 1975 Ex Carnaby Colln fine, transverse strioles in middle, impunctate, with (ANIC); 1 W, Wyndham WA 20.IV.1976 K. & E. Carnaby or without finest and extremely superficial traces of (ANIC); 14 MM, 11 WW, AUSTRALIA: WA; Kimberley, isodiametric microreticulation, glossy. 12m, Home Valley Station, 14°42'S 127°51'E, 15-18. Elytra. Elongate, almost parallel-sided, barely iv.2009, nr Pentecost Riv., St. Jakl lgt. (CBM, CBP, CJP); widened apicad, convex but on disk depressed. Basal 1 W, AUSTRALIA, WA, Kimberley, Home Valley St., angle little produced. Striation complete, but lateral 15°42'29"S 127°51'11"E, 12m alt,16-18.IV.2009, light striae less distinct. Striae well impressed, becom- traps, lgt. Vít Ryjácek (CRP); 4 MM, 4 WW, AUSTRALIA: ing slightly shallower towards apex, impunctate. WA; Kimberley, Wyndham env., 20-21.iv.2009, 15°31'S Intervals slightly convex, impunctate, without or 128°10'E, salty bay of the sea, 3m alt. St. Jakl lgt. (CBM, with very weak transverse strioles, with fine, very CBP, CJP); 1 M, 1 W, AUSTRALIA WA; 15-16.iv.2009, superficial, isodiametric microreticulation, rather Hidden Valley NP, Kununurra 15°46S 128°44E 64 m alt. St. Jakl lgt. (CJP); 2 WW, AU Western Australia NE-Kim- glossy. Epipleura narrow, basal sulcus indistinct. berley: Wyndham 15°31.2'S 128°10.7'E 8m IV – 2009 lgt. Metathoracic wings. Fully developed. Vít Ryjácek (CBP); 1 M, 2 WW, 14.52S 125.50E 4km SbyW Lower surface. Proepisternum impunctate in Mining Camp, Mitchell Plat. WA. 13 May 1983 DCF. ventral half, with few irregular, elongate dorso- Rentz J.Balderson Stop 17 / Crusher at light (ANIC); ventral strioles, also with distinct isodiametric 1 M, 14.52S 125.50E 4km SbyW Mining Camp, Mitchell microreticulation. The whole lower surface with Plateau WA 13May 1983 I:D.Naumann J.C.Cardale at fine, isodiametric microreticulation. Metepisternum d light (ANIC); 1 M, Nov. Holl. Occi / 69650 (NHM). elongate, slightly < 2.5 × as long as wide, almost Etymology. The name is a patronym in honour of the impunctate. Abdominal sterna impunctate. collector of many specimens, Vit Ryácek. Legs. Protibia narrow and elongate, in the male more than in the female. Longitudinal sulcus on Diagnosis. Large, black species with deep, quad- upper surface barely perceptible; lower surface with rangular clypeal excision, not sulcate, little punctate a row of 6-8 stout setae along basal part of inner head, rufopiceous middle and hind legs, oblique margin. Mesotibia moderately elongate, dorsal rim but not convex lateral pronotal margin, impunctate crenulate, with several rows of setae on the dorsal proepisternum, elongate and parallel-sided elytra, surface; subapical spur elongate. Metatibia elongate,

223 rather sparsely setose. Metatarsomere 1 almost as Fitzroy River, W.A. 11 May 1989 ex coll. A.Hay (ANIC); long as the following three tarsomeres. 2 MM, 2 WW, Fitzroy River W.A. at light 16 March 1980 K. Male genitalia (Fig. 13). Aedeagus wide, fairly & E. Carnaby (ANIC); 2 WW, WA Fitzroy Crossing 16 Apr elongate, regularly narrowed apicad; lower surface 1974 Ex Carnaby Colln (ANIC); 1 M, FITZROY RIVER very concave throughout. Apical part barely asym- NWA: 26-4-1976 K+E.Carnaby (ANIC); 1 M, 1 W, Fitzroy Crossing, W.A 125:35E, 18:10S 17 Mar. 1984, at light K. metric, almost regularly triagonal; apex moderately & E. Carnaby (ANIC); 1 W, W.AUSTRALIA Fitzroy River elongate, straight, tip not incurved, triagonal and 16.IV.1976 K. & E. Carnaby (ANIC); 2 MM, WESTERN acute. Both parameres very elongate, the left one AUSTRALIA Fitzroy River 16-4-1976 K. & E. Carnaby considerably stouter than the narrower right one; (ANIC); 1 W, WESTERN AUSTRALIA Fitzroy River 16- the end of the stout part in both parameres oblique; 4-1980 K. & E. Carnaby (ANIC); 1 M, W.Australia Fitzroy both with elongate, very narrow, unisetose apical River 16.IV.1976 K.&E.Carnaby (CNCI); 1 W, Fitzroy R. part; setae very short, situated right at tip. NWA 16 April 1976 K.&E.Carnaby (ANIC); 1 M, Willare Female gonocoxites. Very similar to those of Bridge on Fitzroy River, W:A. 28 Feb. 1985, at light K. & C. procera Putzeys. E. Carnaby (ANIC); 1 M, Fitzroy River NWA: 26.IV.1976 Variation. Some variation noted in body size. K.&E.Carnaby / WALFORD-HUGGINS COLLECTION / Clivina obscuripes Blbn. det. B. P. Moore’77 (CMP); 1 W, Willare W.A. / 3-5’96 Knowles / Golding/Powell Distribution. Northern and north-western NT, Collection donated 12.Feb.2002 (WAM 39192); 1 M, AUS- north-eastern part of the Kimberley Division. TRALIA FITSROY R WA 07-02-1989 Ex Coll. F.Wachtel Collecting circumstances. Most specimens collected (CDW); 2 MM, 9 WW, Winjana Gorge W.A. 17 Mar.1980 at at light, some near rivers or billabongs. light K. & E. Carnaby / more specimens in spirit tube (ANIC); 1 M, 3 WW, W. Aust. At light. Winjana Gorge. Napier Range. 90 mi E. Derby .7th May 1973. T. and C. Clivina heros, spec. nov. Houston. (SAMA); 4 WW, Lennard River W.A. at light 18 Mar. 1980 K. & E. Carnaby (ANIC); 1 M, 2 WW, W.AUST., Figs 14, 56, 124 KIMBERLEY LENNARD RIVER XING GIBB RIVER ROAD 17.23S, 124.44E 1 APRIL 1988 T. F. HOUSTON Examined types. Holotype: M, W.AUST., KIMBERLEY 679-4 / AT LIGHT(MV) AT NIGHT (WAM 39156,-59); CA 9 KM E OF NAPIER DOWNS HS (17.20 S, 124.49 1 W, WA: 14°29'Sx126°46'E Kimberley,Carson R.std. E 5 & 9 APRIL 1988 T. F. HOUSTON 684-3 / AT 21-22 Apr 1998 D.J.Cook, woodland (QMB); 3 MM, 3 WW, LIGHT(MV) AT NIGHT (WAM 39167). – Paratypes: Derby, N.W.A. W. D. Dodd (SAMA 25-033593, -602, 1 M, 3 WW, same data (WAM 39164-66, 68); 1 M, Daly R. -603, -605, -606); Derby, N.W.A. W. D. Dodd / Clivina Mission 10.v.1974 J.Hutchinson (ANIC); 1 W (defect), obscuripes Blkb. ? 75 Large form S. Id. by T. G. Sloane Scolyptus procerus? Putz. Port Darwin (MMS); 1 W, Aus- (SAMA); 1 M, 16-5-1989 DERBY W.A. / Clivina nr. procera tralia/NT Keep River NP.: Gurandalg Camp, ca 50 km E Putz.det. B. P. Moore 1992 (DPIM); 1 W, 6-5-83 / DERBY Kununurra 17. + 23.IV.2004 lgt. M.Langer / S 15.52.494 E G.LAROND / Golding/Powell Collection donated 12 129.03.076 Höhe 99 m Lichtfang (CBM); 1 M, 3 WW, Keep Feb 2002 (WAM 39188); 1 M, N.W.Aust / Clivina procera Creek, N.T. Victoria Hwy 29 Mar. 1984, at light. K. & E. Putz. ? var.? (ANIC); 1 W, N.W.Austra / Clivina procera Carnaby (ANIC); 2 WW, NT. Keep River on Victoria Hwy ? Putz. N.W.Aust. (MMS); 1 (?sex, defect) N.W.Aust 29 Mar 1984 at light. K.&E.Carnaby (ANIC); 3 WW, Keep (MMS); 1 W (CBP); 1 W, (Wadak 117921). Creek, N.T. Victoria Hwy 29 Mar. 1984, at light. K. & E. Carnaby (ANIC); 1 W, AUSTRALIEN Northern Terri- Additional material (1 ex). WA: 1 (?sex, defect) N.W. tory Keep River Nat.Park 7. 3. 1991 leg. H.Schulenburg Aust (MMS). (NTD); 1 M, Keep River, M.T. 29-3-84 Coll: Carnaby (QMB); 1 M, Kununurra W.A. 12.2.62 K. S. Cole (WA- Etymology. The name refers to the very large body size. DAP 5300); 3 MM, 2 WW, Wyndham-KRS 10.2., 14.2., 22.2. Diagnosis. Very large, black species with deep, 53 .R.Lukins (WADAP 5292-96); 1 M, Wyndham-Goose quadrangular clypeal excision, not sulcate, little (?unreadable) 15.3. 53 .R.Lukins (WADAP 52997); 1 M, punctate head, deep black legs, oblique-convex WA, Wyndham20 Apr 1975 Ex: Carnaby Coll. (ANIC); 1 M, 1 W, Kimberley Res. Station W.A. Ex Light Trap. lateral pronotal margin, impunctate proepisternum, 8.3.60, 11.2.61 K. T. Richards (WADAP 5298-9); 2 MM, elongate and parallel-sided elytra, and remarkably Western Australie Ord River Valley Kimberley Res. Stn. curved aedeagus with elongate, straight, triagonal 18-ii-1983 . ESC Smith (WADAK 117919-20); 2 WW, 50-703 apex. Ord River (WAM 38939-40); 1 W, Kimberley Research Stn. Via Wyndham N.W. Aust 22.3.1965 E.C.B.Langfield Description (ANIC); 1 M, AUSTRALIA, WA, Kimberley Home Valley St. 15°42.29'S, 127°51.11'E, 12m alt,16.-18.IV.2009, light Measurements. Length: 18.2-23.9 mm; width: 4.9- traps. leg. Vit Ryácek (CRP); 1 M, Fitzroy River, WA 6.3 mm. Ratios. Length/width of pronotum: 0.98- 14 May 1989 ex. coll. A. Hay (ANIC); 1 W, Australien, 1.05; base/apex of pronotum: 1.30-1.34; width WA Fitzroy Crossing 19.11.1984 M. Baehr (CBM); 2 WW, pronotum/head: 1.36-1.44; length/width of elytra:

224 2.13-2.18; length/width of protibia: M: 3.9-4.05, the narrower right one; the end of the stout part in W: 2.95-3.15; length/width of metatibia: M; 6.2-6.5, both parameres quadrangular; both with moderately W: 6.4-6.6. elongate, very narrow, unisetose or asetose apical Colour (Fig. 124). Unicolourous black. All legs part; setae very short, situated right at tip. deep black. Female gonocoxites (Fig. 56). Gonocoxites com- Head (Fig. 124). Eye large, laterally well pro- paratively short and wide, little curved, with acute jected, orbit short, oblique-convex. Clypeus with apex; with one elongate, rather stout seta basally deep, almost quadrangular excision. Labrum 5-se- at lateral surface and a smaller seta below, three tose. Clypeus not divided from frons; upper surface elongate setae on the ventral surface of gonocoxite smooth, without transverse sulci or strioles; frons 1, three setae at middle of the medio-dorsal surface, sparsely punctate. Surface without distinct micro- and with a tiny nematiform seta near apex. Lateral reticulation, glossy. plate basally with one elongate seta, apically with Pronotum (Fig. 124). Moderately elongate, con- two or three shorter seta. siderably narrowed apicad, dorsally gently convex. Variation. Considerable variation noted in body Lateral margin oblique-convex. Apex distinctly size. concave. Marginal channel narrow and median line shallow but well impressed, impunctate. Basal Distribution. North-western NT, Kimberley Divi- sulcus narrow. Anterior marginal seta situated at sion in northern WA. apical fifth. Basal groove elongate, linear, straight, Collecting circumstances. Many specimens col- impunctate. Disk with some fine, transverse strioles lected at light, some near rivers or billabongs. in middle, impunctate, without microreticulation, glossy. Elytra. Elongate, almost parallel-sided, barely widened apicad, convex but on disk depressed. Basal Clivina goldingi, spec. nov. angle little produced. Striation complete, but lateral Fig. 15 striae less distinct. Striae well impressed, becom- Examined types. Holotype: M, 4 May 1997 / Knotts ing slightly shallower towards apex, impunctate. Crossing Katherine, N.T. / Golding/Powell Collection Intervals slightly convex, impunctate, without or donated 12 Feb 2002 (WAM 39193). with very weak transverse strioles, with very fine, superficial, isodiametric microreticulation, glossy. Etymology. The name is patronym and refers to the Epipleura narrow, basal sulcus shallow, punctate. collector M. Golding. Metathoracic wings. Fully developed. Diagnosis. Large, black species with deep, quad- Lower surface. Proepisternum impunctate, with rangular clypeal excision, not sulcate, little punctate few irregular, elongate dorso-ventral strioles, with head, deep black legs, straight and little oblique distinct isodiametric microreticulation. The whole lateral pronotal margin, impunctate proepisternum, lower surface with fine, isodiametric microreticula- elongate and parallel-sided elytra, and elongate, tion. Metepisternum elongate, c. 2.25 × as long as slightly sinuate, acute apex of the aedeagus. wide, in anterior half with sparse, rugose punctures, in posterior half with a deep longitudinal sulcus. Description Abdominal sterna impunctate. Legs. Protibia narrow and elongate, in the male Measurements. Length: 16.5 mm; width: c. 4.7 mm. more than in the female. Longitudinal sulcus on Ratios. Length/width of pronotum: 1.04; base/ upper surface deep; lower surface with a row of apex of pronotum: 1.35; width pronotum/head: 6-8 stout setae along basal part of inner margin. 1.39; length/width of elytra: c. 2.08; length/width Mesotibia moderately elongate, dorsal rim crenulate, of protibia: M: 3.3; length/width of metatibia: M; 5.8. with several rows of setae on the dorsal surface; Colour. Unicolourous black. All legs deep black. subapical spur elongate. Metatibia elongate, rather Head. Eye large, laterally well projected, orbit sparsely setose. Metatarsomere almost as long as the short, oblique-convex. Clypeus with deep, almost following three tarsomeres. quadrangular excision. Labrum 5-setose. Clypeus not Male genitalia (Fig. 14). Aedeagus wide, fairly divided from frons; upper surface smooth, without elongate, slightly asymmetrically narrowed apicad; transverse sulci or strioles; frons sparsely punctate. lower surface concave, extremely so in apical half. Surface without distinct microreticulation, glossy. Apical part moderately asymmetric, triagonal; apex Pronotum. Moderately elongate, very little nar- narrow and rather elongate, straight but slightly rowed apicad, dorsally gently convex. Lateral margin curved left, tip not incurved, acute. Both parameres straight and little oblique, faintly concave in apical very elongate, the left one considerably stouter than third. Apex barely concave. Marginal channel nar-

225 row; median line shallow, impunctate. Basal sulcus Clivina newcastleana, spec. nov. narrow. Anterior marginal seta situated at apical Figs 16, 57, 125 fifth. Basal groove very shallow, barely indicated, elongate, linear, straight, impunctate. Disk with some Examined types. Holotype: M, Elsey Ck. 10ml. E of very fine, transverse strioles in middle, impunctate, Mataranka on Roper Bar Rd. NT 22.iii.72, A. Allwood with very fine, isodiametric microreticulation, fairly & . W. Forrester / 11271 (NTD). – Paratypes: 2 MM, 7 WW, glossy. same data (NTD); 1 W, Elsey Ck. 10ml. E of Mataranka Elytra. Elongate, almost parallel-sided, barely on Roper Bar Rd. NT 19.iv.72, T. Weir A. Allwood / 11240 (NTD); 1 W, Elsey Ck. NT. Roper Bar Rd. 22.iii.1972 widened apicad, convex but on disk depressed. Allwood&Angeles / Clivina sp. / 11227 (NTD); 7 MM, Basal angle little produced. Striation complete, but 15 WW, AUSTRALIA NT: 190m alt. 70km SW of Mata- lateral striae less distinct. Striae well impressed, ranka, 15°19'S 132°50'E 22-23.xii.2008, St. Jakl lgt. (CBM, becoming slightly shallower towards apex, at least in CBP, CJP); 2 WW, AUSTRALIA NT: 190m alt. 70km SW basal half finely punctate. Intervals slightly convex, of Mataranka, 15°19'S 132°50'E 14-16.i.2009; St. Jakl impunctate, without transverse strioles, with fine, lgt. (CJP); 3 M, 9 WW, AUSTRALIA NT: 190m alt. 70km isodiametric microreticulation, moderately glossy. SW of Mataranka, 15°19'S 132°50'E 22-23.xii.2008; L. Epipleura narrow, basal sulcus indistinct. Hovorka lgt. (CBM, CBP, CHP); 1 W, AUSTR. NT, 70km Metathoracic wings. Fully developed. SW of Mataranka, 14-15.1.09, 15°19'S 132°50'E 190m, Lower surface. Proepisternum impunctate, with Sv. Bilý leg. (CBM); 2 MM, 2 WW, AUSTRALIA NT: 3km few irregular, elongate dorso-ventral strioles, with N of Mataranka, 14°43'S 132°01'E 18-19.xii.2008, St. Jakl lgt. (CBM, CBP, CJP); 7 MM, 10 WW, AUSTRALIA: distinct isodiametric microreticulation. The whole NT; 168m alt. 25 km S of Katherine, nr. Cutta Cutta lower surface with fine, isodiametric microreticula- Caves, 14°31'S 132°25'E, 23-31.xii.2008, St. Jakl lgt. tion. Metepisternum elongate, c. 2.25 × as long as (CBM, CBP, CJP); 2 WW, Cutta Cutta N.T. June 1987 S. wide, in anterior half with sparse, rugose punctures, Churchill (AMS K255334); 1 M, AUSTRALIA: NT; 100m in posterior half with a deep longituidinal sulcus. alt., 25 km S of Katherine, rd. to Kununurra, 14°44'S Abdominal sterna impunctate. 132°01'E, 17-31.xii.2008, St. Jakl lgt. (CBM); 26 MM, 36 WW, Legs. Protibia comparatively short and wide. AUS13, NT25, c.12 km wnw.Ellot, 204m 17°37'06"S, Longitudinal sulcus on upper surface faintly im- 133°28'41"E 17.4.2013, M.Baehr (CBM); 11 MM, 19 WW, pressed; lower surface with a row of 6-8 stout setae Australia: N.T. (am Stuart Hwy) ca. 25 km N Elliott along basal part of inner margin. Mesotibia moder- (Bauplatz) 20.IV.2011 LF; H = 274 m S 17°23'59.9" E ately elongate, dorsal rim crenulate, with several 133°27'12.2" leg. Michael Langer (CBM, CWB); 3 WW, AUS13, NT45, c.32 km e.Mataranka, Roper Bar Rd., rows of setae on the dorsal surface; subapical spur 14°59'16"S, 133°21'06"E 115m, 3.5.2013, M.Baehr (CBM); elongate. Metatibia elongate, rather sparsely setose. 1 M, 2 WW, AUS13, NT47, c.78 km e. Mataranka, Roper Metatarsomere 1 slightly shorter than the following Bar Rd., 14°54'59"S, 133°42'48"E, 74m, 5.5.2013, M.Baehr three tarsomeres. (CBM); 3 MM, 9 WW, AUS13, NT12, NT23, 5 km s. Lar- Male genitalia (Fig. 15). Aedeagus very wide, rimah, Birdum Crk., 184m, 15°32'02"S, 133°22'33"E, 8.4., fairly elongate, narrowed apicad and slightly sinu- 16.4.2013, M.Baehr (CBM); 10 MM, 9 WW, AUS13, NT44, ate; lower surface in basal two thirds gently concave, 5 km s. Larrimah, Birdum Crk., 500m e. NT12, 185m, then straight and slightly directed down. Apical part 15°32'02"S, 133°23'35"E, 2.5.2013, M.Baehr (CBM); 2 MM, elongate, conspicuously bisinuate at both sides, re- 1 W, AUSTRALIA NT 316m alt, 14°53'S (wrong latitude, markably sinuate; apex elongate, narrowed towards must be 18°53'S!), 132°01'E, Banka Banka env., road to tip, which is rather acute. Both parameres very stout, Tennant Creek, 12-14.i.2009, St. Jakl lgt. (CBM, CBP, CJP); 1 M, AUSTRALIA N. Territ. Katherine env. 10- the left one considerably stouter than the right one; 14.I.2004 Lgt. P. Machácek (CBM); 1 W, Col G. Lecourt both with rather short, narrow, uni- or bisetose apical 01/00 Katherine NT Australie / Clivina 1VK P. Bulirsch part; setae short, situated right at the apex; the left det. 2006 (CBP); 2 MM, 4 WW, AUSTRALIA N.T. Katherine paramere with a tiny second seta. 28/3/93 Leg. L. Toledano (CBM, CTV); 1 W, AUSTRAL- Female gonocoxites. Unknown. IA N.T. Katherine 28/3/93 (CGT); 2 WW, Austral. North Variation. Unknown. T. Katherine, XII.57 leg. H. Demarz (ZSM); 1 W, Australia N.T. Katherine Gorge30.III.93 (CGT); 1 M, Katherine, Distribution. Northern NT. Known only from type N.T. at light,7.ii.68 J.A.L.Watson (ANIC); 1 W, N.T. locality. Katherine Gorge NP M.V.Light 1 Apr 1981M. Malitpatil & J. Hawkins (NTD); 1 M, 1 W, Katherine (ANIC); 1 W, Collecting circumstances. Not recorded. AUS13, NT21, T/O Stuart Hwy./Edith River Rd., 145m, 14°11'10"S, 132°02'04"E, 15.4.2013, M.Baehr (CBM); 3 WW, NT: 80km S of Larrimah 24 Jan. 1977 M.S. & B.J. Moulds (AMS K225265, K225291, K225305); 1 M, 1 W, NT Elliot 10 March 1997 G. R. Brown at light (NTD); 1 M, Katherine,

226 N.T. 17 Apr.1962 I.F.H.Common (ANIC); 1 M, 1 W, Kath- or almost so, without or with very weak transverse erine Gorge M. V. Light 1 Apr 1981 M. Malipatil & J. strioles, with fine, isodiametric microreticulation, Hawkins (NTD); 1 M, N.T. Kulamindini Bore nr. Elliot moderately glossy. Epipleura narrow, basal sulcus 17.34S 133.34E M.V.Light 13 Apr. 1981 M. Malipatil & indistinct. J. Hawkins (NTD); 1 W, Newcastle Waters N. Australia 3 Metathoracic wings. Fully developed. June 1929 T. G. Campbell (ANIC), 1 W, N.T. Lake Woods, Lower surface. Proepisternum impunctate or 15 km SW Elliot, at light 3 Oct. 1977. G.F. Gross, J.A. Forrest (SAMA); 1 W, Newcastle Waters, N.Australia sparsely punctate in ventral half, with few irregular, June 1929 T. G. Campbell / Clivina quadratifrons Sln. Id. elongate dorso-ventral strioles and short, transverse by W.K.Hughes (ANIC). strioles, also with distinct isodiametric microreticulation. The whole lower surface with fine, isodiametric Etymology. The name refers to the Newcastle Waters microreticulation. Metepisternum elongate, c. 2.25 × area where many specimens were collected. as long as wide, in anterior half with sparse, rugose Diagnosis. Rather large, black species with deep, punctures, in posterior half with a deep longitudinal quadrangular clypeal excision, not sulcate, sparsely sulcus . Abdominal sterna impunctate. punctate head, rufous middle and hind legs, straight Legs. Protibia narrow and elongate, in the male and little oblique lateral pronotal margin, impunctate more than in the female. Longitudinal sulcus on or sparsely punctate proepisternum, elongate and upper surface faintly impressed; lower surface with parallel-sided elytra, and elongate, sinuate, acute and a row of 6-8 stout setae along basal part of inner incurved or slightly crotched apex of the aedeagus. margin. Mesotibia moderately elongate, dorsal rim crenulate, with several rows of setae on the dorsal Description surface; subapical spur elongate. Metatibia elongate, rather sparsely setose. Metatarsomere 1 almost as Measurements. Length: 11.2-14.4 mm; width: 3.15- long as the following three tarsomeres. 4.1 mm. Ratios. Length/width of pronotum:.0.96-1.0; Male genitalia (Fig. 16). Aedeagus wide, fairly base/apex of pronotum: 1.25-1.28; width pronotum/ elongate, narrowed apicad; lower surface in ba- head: 1.33-1.39; length/width of elytra: 2.0-2.05; sal half concave, in apical third slightly convex, length/width of protibia: M: 3.8-4.0, W: 2.9-3.2; towards apex again deeply concave. Apical part length/width of metatibia: M; 6.2-6.4, W: 5.8-6.2. very asymmetric, deeply bisinuate at the left side; Colour (Fig. 125). Unicolourous black. Middle apex elongate, narrow, slightly curved left, slightly and hind legs rufous, considerably paler than the asymmetric, tip incurved and crotched, acute. Both anterior leg. parameres stout but elongate, the left one consid- Head (Fig. 125). Eye large, laterally well pro- erably stouter than the right one; both with very jected, orbit short, oblique-convex. Clypeus with elongate, narrow, bisetose, apical part; setae very deep, almost quadrangular excision. Labrum 5-se- short, situated right at tip. tose. Clypeus not divided from frons; upper surface Female gonocoxites (Fig. 57). Gonocoxites very smooth, without transverse sulci or strioles, rather narrow and elongate, little curved, with acute apex; sparsely punctate. Surface without distinct micro- with one elongate, rather stout seta basally at lateral reticulation, glossy. surface and a smaller seta below, two elongate setae Pronotum (Fig. 125). Comparably short, little on the ventral surface of gonocoxite 1, three setae narrowed apicad, dorsally gently convex. Lateral at middle of the medio-dorsal surface, and a tiny margin straight and slightly oblique, faintly concave nematiform seta near apex. Lateral plate basally with in apical third. Apex distinctly concave. Marginal one elongate seta, apically apparently without setae. channel narrow; median line shallow but well im- Variation. Considerable variation noted only pressed, impunctate. Basal sulcus narrow. Anterior in body size. marginal seta situated at apical fifth. Basal groove elongate, linear, straight, impunctate. Disk with some Distribution. Central NT from about Katherine to fine, transverse strioles in middle, impunctate, with north of Tennant Creek. very fine, superficial, isodiametric microreticulation, glossy. Collecting circumstances. Many specimens col- Elytra. Elongate, almost parallel-sided, barely lected at light, some near rivers or billabongs. widened apicad, convex but on disk depressed. Basal angle little produced. Striation complete, but lateral striae less distinct. Striae well impressed, becoming slightly shallower towards apex, at least in basal half finely punctate. Intervals slightly convex, impunctate

227 Clivina glabripennis, spec. nov. impunctate, without or with very weak transverse Figs 17, 85 strioles, without microreticulation, remarkably glossy. Epipleura narrow, basal sulcus indistinct. Examined types. Holotype: M, AUSTRALIA: QLD: Metathoracic wings. Fully developed. 271 km West Windorah, 31.viii.1997, J. & A.Skevington, Lower surface. Proepisternum impunctate, with A.Zwick, S.Winterton, C.Lambkin / UQIC Reg. #90502 few irregular, elongate dorso-ventral strioles and (QMT234150). – Paratypes: 2 WW, same data / UQIC Reg. short, transverse strioles, also with distinct isodia- #90500-1 (CBM, QMB); 4 WW, AUSTRALIA SW Queens- metric microreticulation. The whole lower surface land 12km W of Windorah; 110m 28-31.i.2011; sanddunes 25°21'S, 142°32'E; St. Jakl lgt. (CBM, CBP, CJP); with fine, isodiametric microreticulation. Metepister- 1 W, INNAMINCKA: COOPERS CK S.A: 24 Jan. 1976 num elongate, c. 2.25 × as long as wide, in posterior M.S. & B.J.Moulds / WALFORD-HUGGINS COLLEC- half with a deep longitudinal sulcus . Abdominal TION / Clivina procera Putz. det. B.P. Moore’77 (CMP). sterna impunctate. Legs. Protibia narrow and elongate, in the male Etymology. The name refers to the glossy, not micro- more than in the female. Longitudinal sulcus on reticulate surface. upper surface deeply impressed; lower surface with Diagnosis. Large, black species with deep, quadran- a row of 6-8 stout setae along basal part of inner gular clypeal excision, not sulcate, sparsely punctate margin. Mesotibia moderately elongate, dorsal rim head, rufous middle and hind legs, straight and crenulate, with several rows of setae on the dorsal fairly oblique lateral pronotal margin, impunctate surface; subapical spur elongate. Metatibia elongate, proepisternum, elongate and parallel-sided elytra, rather sparsely setose. Metatarsomere 1 almost as and elongate, sinuate, acute, not incurved apex of long as the following three tarsomeres. the aedeagus. Male genitalia (Fig. 17). Aedeagus very wide, rather elongate, asymmetrically narrowed apicad; Description lower surface concave, extremely so in apical half. Apical half of lower surface with two longitudinal Measurements. Length: 13.7-17.0 mm; width: 3.8- ridges, surface between deeply concave. Apical part 4.8 mm. Ratios. Length/width of pronotum: 0.98- very asymmetric, deeply concave at the left side; 1.01; base/apex of pronotum: 1.29-1.31; width apex moderately elongate, rather narrow, straight pronotum/head: 1.37-1.41; length/width of elytra: but slightly curved left, tip not incurved ventrad, 2.02-2.06; length/width of protibia: M: 3.35, W: 2.85- obtuse. Both parameres elongate, the left one con- 3.1; length/width of metatibia: M; 6.7, W: 6.1-6.3. siderably stouter than the narrower right one; both Colour (Fig. 85). Unicolourous deep, glossy with moderately elongate, very narrow, bisetose black. Middle and hind legs rufous, anterior leg apical part; setae short, situated right at tip. rufo-piceous. Female gonocoxites. Very similar to those of Head (Fig. 85). Eye large, laterally well project- C. newcastleana, spec. nov. ed, orbit short, oblique-convex. Clypeus with deep, Variation. Considerable variation noted in body almost quadrangular excision. Labrum 5-setose. Cly size. peus not divided from frons; upper surface smooth, without transverse sulci or strioles, rather sparsely Distribution. South-western QLD, adjacent north- punctate. Surface without microreticulation, glossy. eastern SA. Pronotum (Fig. 85). Comparably short, fairly narrowed apicad, dorsally gently convex. Lateral Collecting circumstances. Little recorded. The margin straight and oblique, faintly concave in api- specimens from near Windorah were sampled in cal third. Apex distinctly concave. Marginal channel “sanddunes”, most probably at light. narrow; median line well impressed, impunctate. Basal sulcus narrow. Anterior marginal seta situated at apical fifth. Basal groove shallow, elongate, Clivina micans, spec. nov. linear, straight, impunctate. Disk with some fine, Figs 18, 86 transverse strioles in middle, impunctate, without microreticulation, very glossy. Examined types. Holotype: M, Q./N.T. border. Elytra (Fig. 85). Elongate, parallel-sided, not nr. Camooweal. W.Qld. 25 Mar. 84 Coll: A. Hiller widened apicad, convex but on disk depressed. (QMT234902). – Paratypes: 1 M, same data (CBM); 1 M, 1 W, Alexandria, N. Australia. W. Saalker. 18,20-3-06 Basal angle little produced. Striation complete, but 1907-261. / ovalipennis Sl. Det. K. Kult / Collectio lateral striae less distinct. Striae well impressed, Karel Kult Coll.A.Dostal,1989 (CDW). becoming slightly shallower towards apex, at least in basal half finely punctate. Intervals slightly convex, Etymology. The name refers to the very glossy surface.

228 Diagnosis. Rather large to large, black species with Legs. Protibia narrow and elongate, in the male deep, quadrangular clypeal excision, not sulcate, more than in the female. Longitudinal sulcus on sparsely punctate head, black legs, straight and upper surface distinct; lower surface with a row fairly oblique lateral pronotal margin, impunctate of 6-8 stout setae along basal part of inner margin. and very glabrous proepisternum, elongate but not Mesotibia moderately elongate, dorsal rim crenulate, absolutely parallel-sided elytra, coarsely punctate with several rows of setae on the dorsal surface; elytral striae, and elongate, straight, slightly turned subapical spur elongate. Metatibia elongate, rather left apex of the aedeagus with obtuse tip. sparsely setose. Metatarsomere 1 almost as long as the following three tarsomeres. Description Male genitalia (Fig. 18). Aedeagus wide, moderately elongate, very asymmetrically narrowed Measurements. Length: 12.9-15.6 mm; width: 3.7- apicad; lower surface concave, in apical third almost 4.35 mm. Ratios. Length/width of pronotum: 1.04- straight. Apical half of lower surface gently concave. 1.06; base/apex of pronotum: 1.16-1.24; width Apical part very asymmetric, bisinuate at the left pronotum/head: 1.42-1.50; length/width of elytra: side; apex elongate, rather narrow, slightly curved 2.0-2.03; length/width of protibia: M: 3.25-3.4, W: 2.5; left, tip obtusely angulate. Both parameres rather length/width of metatibia: M; 6.0-6.1, W: 5.1. short, the left one stouter than the narrower right Colour (Fig. 86). Unicolourous black. Legs deep one; both with very short, narrow, asetose apical part. black. Female gonocoxites. Very similar to those of Head (Fig. 86). Eye fairly large, laterally moder- C. newcastleana, spec. nov. ately projected, orbit short, oblique-convex. Clypeus Variation. Considerable variation noted in body with deep, almost quadrangular excision. Labrum size. Also apparently some sexual variation in pres- 5-setose. Clypeus not divided from frons; upper sur- ence/absence of microreticulation on pronotum face smooth, without transverse sulci but frons later- and elytra. ally with some short, faint strioles, rather sparsely punctate. Surface without microreticulation, glossy. Distribution. North-western QLD near QLD/NT Pronotum (Fig. 86). Moderately elongate, nar- border, adjacent central eastern NT. rowed apicad, dorsally gently convex. Lateral margin straight and oblique, faintly concave in apical third. Collecting circumstances. Not recorded. Apex distinctly concave. Marginal channel narrow; median line well impressed, impunctate. Basal sulcus narrow. Anterior marginal seta situated at apical Clivina rugosifrons, spec. nov. fifth. Basal groove shallow, elongate, linear, straight, Figs 19, 58, 126 impunctate. Disk with some fine, transverse strioles in middle, impunctate, in the males without, in the Examined types. Holotype: M, Moreton Tel. Stn., single female with very faint, superficial, isodiamet- Cape York Pen. N. Qld. 30.vi.1975 G. B. Monteith ric microreticulation, very glossy. (QMT234903). – Paratypes: 3 MM, 5 WW, same data (QMB, Elytra (Fig. 86). Elongate, but not absolutely CBM); 1 W, Batten Point 15.54S 136.32E 30 km NE by E of Borroloola, N.T. 18.iv.1976 J.E. Feehan (ANIC); 1 W, parallel-sided, or slightly widened apicad, convex 2754 NQU / C. procera Putz. var? Gulf of Carpentaria but on disk depressed. Basal angle little produced. (SAMA); 1 W, Cairns dist. E. Allen (SAMA 25-033611); Striation complete, but lateral striae less distinct. 1 W, AUSTRALIA: n. Qld. Pinnarondi Stn. 60 km W. Striae deep, becoming slightly shallower towards of Mt. Garnet 7.ii.1989 D. Heiner at light (QDPIM); apex, coarsely punctate. Intervals slightly convex, 1 W, NORTH QLD SPRING VALLEY / 20.Feb.1972A impunctate, without or with very weak transverse & M. Walford-Huggins 5902 / WALFORD-HUGGINS strioles, in males without, in the single female with COLLECTION / Clivina procera Putz. [Series det. by very faint, superficial, isodiametric microreticula- A.Walford-Huggins] (CMP); 1 W, Gulf. C.F. 16/6/94 tion, glossy. Epipleura narrow, basal sulcus distinct, N.W. (unreadable) W (ANIC); 1 W, Gulf C.F. 16/3/94 punctate. (ANIC); 1 W, Conjuboy2/42 / M. 158 / J. G. Brooks Be- Metathoracic wings. Fully developed. quest , 1976 (ANIC); 1 M, QLD: 21.138°Sx148.155°E Red- cliffe Tbld.Site 3. 13-14Apr2012, S.Monteith mushroom Lower surface. Proepisternum impunctate, and trap, open forest, 400m 35235 (QMB); 1 W, NQU / ob- almost without strioles, remarkably glabrous, with scuripes Blackb. (SAMA); 3 WW, Stewart R. Q W. D. Dodd faint isodiametric microreticulation. The whole / 19264 Clivina obscuripes Blkb. 75 S. Id by T. G. Sloane lower surface with fine, isodiametric microreticula- (SAMA); 1 M, 5 WW, W.of Ravenshoe AthertonTab.,Q. tion. Metepisternum elongate, slightly > 2 × as long c.3000’,Feb.’58 Darlingtons (CBM, MCZ). as wide, in posterior half with a deep longitudinal sulcus. Abdominal sterna impunctate. Etymology. The name refers to the rugose, transversely sulcate frons.

229 Diagnosis. Large, black species with fairly deep, Legs. Protibia narrow and elongate, in the male wide, quadrangular clypeal excision, transversely more than in the female. Longitudinal sulcus on sulcate, sparsely punctate head, rufous to rufo- upper surface distinct; lower surface with a row piceous middle and hind legs, straight and fairly of 6-8 stout setae along basal part of inner margin. oblique lateral pronotal margin, impunctate or Mesotibia moderately elongate, dorsal rim crenulate, sparsely punctate proepisternum, elongate, parallel- with several rows of setae on the dorsal surface; sided elytra, and elongate, sinuate, barely incurved subapical spur elongate. Metatibia elongate, rather apex of the aedeagus. sparsely setose. Metatarsomere 1 almost as long as the following three tarsomeres. Description Male genitalia (Fig. 19). Aedeagus wide, fairly elongate, narrowed apicad; lower surface in basal Measurements. Length: 14.1-15.9 mm; width: 3.9- half concave, in apical half bisinuate. Apical part 4.8 mm. Ratios. Length/width of pronotum: 1.01- asymmetric, bisinuate at the left side; apex short, 1.06; base/apex of pronotum: 1.26-1.33; width slightly turned left, slightly asymmetric, tip not pronotum/head: 1.32-1.38; length/width of elytra: incurved, more or less obtuse. Both parameres very 2.02-2.08; length/width of protibia: M: 3.15-3.25, elongate, the left one considerably stouter than the W: 2.9-3.05; length/width of metatibia: M; 6.1-6.2, narrower right one; both with elongate, very narrow, W: 5.7-6.1. asetose apical part. Colour (Fig. 126). Unicolourous black. Mid- Female gonocoxites (Fig. 58). Gonocoxites rather dle and hind legs rufopiceous, little paler than the narrow and elongate, little curved, with obtusely anterior leg. rounded apex; with one elongate, rather stout seta Head (Fig. 126). Eye large, laterally well pro- basally at lateral surface and a smaller seta below, jected, orbit short, oblique-convex. Clypeus with two elongate setae on the ventral surface of gono- deep, almost quadrangular excision. Labrum usually coxite 1, two setae at middle of the medio-dorsal 5-setose, rarely 4- or 7-setose. Clypeus divided from surface, and with a tiny nematiform seta near apex. frons by a sulcus, usually clypeus and anterior part of Lateral plate basally with one elongate seta, apically frons with additional transverse sulci. Surface rather with two or three shorter seta. sparsely punctate, without microreticulation, glossy. Variation. Some variation noted in body size, Pronotum (Fig. 126). Moderately elongate, relative length of prothorax and elytra, and distinct- slightly narrowed apicad, dorsally gently convex. ness of the frontal sulci. Lateral margin straight and oblique, faintly concave in apical third. Apex slightly concave. Marginal chan- Distribution. North QLD, adjacent north-eastern nel narrow; median line well impressed, impunctate. NT. Basal sulcus narrow. Anterior marginal seta situated at apical fifth. Basal groove shallow, elongate, linear, Collecting circumstances. Little recorded. Some straight, impunctate. Disk with some fine, transverse specimens were collected at light, one in “mushroom strioles in middle, impunctate, with very superficial trap in open forest”, several near river. traces of isodiametric microreticulation, glossy. Elytra (Fig. 127). Elongate, parallel-sided, barely widened apicad, convex but on disk depressed. Clivina monilicornis Sloane Basal angle little produced. Striation complete, but Fig. 20 lateral striae less distinct. Striae well impressed, becoming slightly shallower towards apex, at least in Clivina monilicornis Sloane, 1896a: 229. – Sloane 1896a: basal half finely punctate. Intervals slightly convex, 227, 231, 239; Csiki 1927: 508; Moore et al. 1987: 73; impunctate, without or with very weak transverse Lorenz 2005: 143. strioles, with very faint, superficial, isodiametric Examined types. Lectotype (by present designation): microreticulation, glossy. Epipleura narrow, basal W (probable, genitalia destroyed), Pt. Denison / SYN- sulcus indistinct. TYPE (red) / Clivina monilicornis Sl. Port Denison (AN- Metathoracic wings. Fully developed. IC-MMS). – Paralectotypes: 2 WW (probable, abdomen Lower surface. Proepisternum sparsely punctate of one specimen destroyed), Pt. Denison / SYNTYPE or impunctate, with few irregular, elongate dorso- (red) (ANIC-MMS); 1 (sex?, only abdomen left, but ventral strioles, with faint isodiametric microreticula- genitalia destroyed), Pt. Denison / Clivina monilicornis, tion. The whole lower surface with fine, isodiametric Sl. / PARATYPE (blue) (ANIC-MMS). microreticulation. Metepisternum elongate, c. 2.25 × Type locality. “Port Denison” (= Townsville), North as long as wide, in posterior half with a deep longi- Queensland. tudinal sulcus. Abdominal sterna impunctate.

230 Other material (13 ex.). QLD: 1 W, Cape York H. tion complete, but lateral striae less distinct. Striae Eigner / Clivina Cape York (QMB); 1 W, Silver Plains deep, becoming slightly shallower towards apex, Cape York, Q. May-June ’58 Darlingtons (MCZ); impunctate or almost so. Intervals slightly convex, 1 M, Qld:19°07.8'Sx145°20.2'E Gregory & Herveys Ra impunctate, without or with very weak transverse Dev. Rds crossing. 350m 14736 17Dec2006-15Feb2007. strioles, with very faint, superficial, isodiametric Monteith&Cook.pitfall.OF (QMB); 1 W, Endevour River microreticulation, glossy. Epipleura narrow, basal Queensland / Clivina monilicornis Sl. Id. by T. G. Sloane / Pres by T. G. Sloane Esq. 12-6-16 (NMV COL-5380); sulcus indistinct. 1 M, 1 W (defect), Pt. Denison (MMS); 1 W, Port Nales (?) Metathoracic wings. Fully developed. Queensland Elgner (SAMA 25-033590); 1 M, Queensland Lower surface. Proepisternum impunctate, with (SAMA); 1 W, Nat. Mus. Victoria Queensland (NMV); few irregular, elongate dorso-ventral strioles and 1 W, Cape York H. Eigner / Clivina Cape York (QMB); some short transverse strioles, with faint isodiametric 1 W, AUSTRALIEN Cape York. (SMNS); 1 M (defect), microreticulation. The whole lower surface with 5589 N. Qu. / sp. nov. near macleayi prostern sulcate at fine, isodiametric microreticulation. Metepisternum base / I. 6863 Clivina Queensland (SAMA 25-033694). elongate, slightly > 2 × as long as wide, in posterior – NT: 1 M, Brock’s Crk N.T. 13-1-32 T.G.Campbell. half with a deep longitudinal sulcus. Abdominal (ANIC). sterna impunctate. Diagnosis. Comparatively small to medium sized, Legs. Protibia narrow and elongate, in the male black species with wide and shallow, quadrangular more than in the female. Longitudinal sulcus on clypeal excision, More or less sulcate, sparsely punc- upper surface faintly impressed; lower surface with tate head, dark legs, conical prothorax, impunctate a row of 6-8 stout setae along basal part of inner proepisternum, elongate but not absolutely parallel- margin. Mesotibia moderately elongate, dorsal rim sided elytra, deeply impressed, barely punctate crenulate, with several rows of setae on the dorsal elytral striae, and short, straight, parallel-sided, at surface; subapical spur elongate. Metatibia moder- tip obtuse apex of the aedeagus. ately elongate, rather sparsely setose. Metatarsomere shorter than the following three tarsomeres. Description Male genitalia (Fig. 20). Aedeagus wide, fairly elongate, asymmetrically narrowed apicad; lower Measurements. Length: 9.1-13.0 mm; width: 2.6- surface almost regularly concave. Apical half of 3.6 mm. Ratios. Length/width of pronotum: 1.06- lower surface with two longitudinal, parallel ridges, 1.09; base/apex of pronotum: 1.35-1.37; width the surface between concave. Apical part bisinuate pronotum/head: 1.49-1.52; length/width of elytra: on the left side; apex fairly short, straight, narrow, 1.93-2.02; length/width of protibia: M: 3.1-3.2, W: 2.8- parallel-sided, tip obtusely angulate. Both parameres 2.85; length/width of metatibia: M; 5.1-5.2, W: 4.9-5.0. short and stout, the left one considerably stouter Colour. Unicolourous black. Legs dark piceous than the right one; both with very short but acute, to black. asetose apical part, Head. Eye fairly large, laterally moderately Female gonocoxites. Very similar to those of projected, orbit short, oblique-convex. Clypeus C. conicollis, spec. nov. with rather shallow, wide, almost quadrangular Variation. Considerable variation noted in body excision, though the lateral margins of the excision size. being rather oblique. Labrum usually 5-setose, rarely 6-setose. Clypeus usually divided from frons; upper Distribution. North QLD, doubtfully also in north- surface with more or less distinct transverse sulci, ern NT. rather sparsely punctate. Surface without microreticulation, glossy. Collecting circumstances. Not recorded. Pronotum. Rather elongate, rather conical, dorsally gently convex. Lateral margin straight and oblique, faintly concave in apical third. Apex distinct- Clivina conicollis, spec. nov. ly concave. Marginal channel narrow; median line Figs 21, 59, 127 well impressed, impunctate. Basal sulcus narrow. Examined types. Holotype: M, NT.A.L.Hertog Kakadu Anterior marginal seta situated at apical fifth. Basal NP Jabiru NT May 1998 (ANIC). – Paratypes: 1 M, groove shallow, rather punctiform, impunctate. Disk 1 (?sex, defect), King R’ N.T. 24.12.15 Nat. Mus. Victoria rarely with some fine, transverse strioles in middle, Coll.leg. W.McLennan / Clivina monilicornis Sl. (large impunctate, without microreticulation, very glossy. form) Id. by T. G. Sloane (ANIC, NMV COL-5386); 1 M, Elytra. Rather elongate, but not absolutely King R’ N.T. Nat. Mus. Victoria Coll.leg. W.McLennan parallel-sided, slightly widened apicad, convex but 24.12.15 / Clivina monilicornis Sl. Det by T. G. Sloane on disk depressed. Basal angle little produced. Stria- 3 - 17 / (NMV COL-14020); 1 M, King R’ N.T. 24.12.15 /

231 Coll. By W. Mc Lennan sp 64 H.L.White prp. 14.10-16 / excision, though the lateral margins of the excision Clivina monilicornis Sl. Det by T. G. Sloane 3 - 17 (NMV being rather oblique. Labrum usually 7-setose, rarely COL-5381); 1 M, King R’ N.T. 24.12.15 / Coll. By W. 5-setose. Clypeus usually shallowly divided from Mc Lennan sp 64 H.L.White prp. 14.10-16 (NMV COL- frons; upper surface without or with faint transverse 14024); 2 MM, 1 W, AUSTRALIA: Howard Springs N.T. sulci, rather sparsely punctate. Surface without 22.iii.68 B.P.Moore (ANIC); 1 M, Australia, NT Kakadu, microreticulation, glossy. 10/14.I.2004 lgt. R. Novák (CKZ); 1 M, 2 WW, AUS- TRALIA Elizabeth R. N.T. 21.iii.68 B.P.Moore / Clivina Pronotum (Fig. 127). Elongate, markedly coni- monilicornis Sl. det. B.P.Moore#79 (ANIC); 1 W, N.T. M. cal, dorsally gently convex. Lateral margin straight Armstrong Mary River Area N.T. PWCNT Study MS 01 and oblique, faintly concave in apical third. Apex Jun 1999 (ANIC); 1 W, 12.48S 132.42E Nourlangie Creek, distinctly concave. Marginal channel narrow; median N.T. 8km N. of Mt. Cahill, 16.vi.73, Upton & Feehan line well impressed, impunctate. Basal sulcus nar- (ANIC); 1 M, 1 W, AUS13, NT3, Leaning Tree Lagoon, row. Anterior marginal seta situated at apical fifth. 35 km e.Stuart Hwy.T/O 12°42'34"S,131°25'20"E 25m, Basal groove shallow, oval shaped, impunctate. Disk 4.4.2013, M.Baehr (CBM); 1 W, Groote Eylandt N. B. rarely with some fine, transverse strioles in middle, Tindale (SAMA No. 25-033513); 2 WW, AUSTRALIA impunctate, with faintest and extremely superficial NT; 190m alt. 70km SW of Mataranka 15°19'S 132°50'E traces of isodiametric microreticulation, very glossy. 14-16.I.2009 St Jakl (CBM, CJP); 1 M, 1 W, Kakadu Nat. Elytra. Elongate, but not absolutely parallel- Park, CSIRO Biodiversity Survey, Site 134B, 13.41257 S, 132.48407 E, S. Oberprieler, Feb-Mar 2014 (CBM); sided, slightly widened apicad, convex but on disk 13 MM, 31 WW, 3/06 N-Austr, NT, Kakadu NP Muire- depressed. Basal angle little produced. Striation lla, 10.06.2006 40m S12°51'15" E132°45'16" Light lg complete, but lateral striae less distinct. Striae Berger-Dostal (CBM, CDW); 2 MM, 3 WW 8/06 N- deep, becoming slightly shallower towards apex, Austr, NT, 40km v Port Roper, 15.05.2006, S14°54'04.8" impunctate or almost so. Intervals slightly convex, E135°03'24.7" 134m, LT. Berger-Dostal (CBM, CDW); impunctate, without or with very weak transverse 2 MM, Weipa NQ 12.35S 141.53E pitfall 17.i-30.vi.1993 strioles, with very faint, superficial, isodiametric A. Thomas (AMS K 225355); 1 M, QLD Weipa Cape microreticulation, glossy. Epipleura narrow, basal York 12°36'±7'S 141°53'±7'E 17 Jan. 1992-30 June, 1983 sulcus indistinct. A.J.Thomas, Pitfall Trap (AMS K 249708); 2 MM, Cape Metathoracic wings. Fully developed. York H. Eigner (QMB); 1 W, C. 2199 (QMB); 1 W, Horn Lower surface. Proepisternum impunctate, with Is.N. Queensl. 4/2/39 from wind / cf. monilicornis Sl. det.’59 Darlington / Clivina monilicornis Sl. (MCZ); 1 M, few irregular, elongate dorso-ventral strioles and Prince of Wales Is. Australia 2/19/39 (MCZ). some short transverse strioles, with faint isodiametric microreticulation. The whole lower surface with Etymology. The name refers to the elongate, markedly fine, isodiametric microreticulation. Metepisternum conical prothorax. elongate, slightly > 2 × as long as wide, in posterior Diagnosis. Medium sized to rather large, black spe- half with a deep longitudinal sulcus. Abdominal cies with wide and shallow, quadrangular clypeal sterna impunctate. excision, not or barely sulcate, sparsely punctate Legs. Protibia narrow and elongate, in the male head, dark legs, very conical prothorax, impunctate more than in the female. Longitudinal sulcus on proepisternum, elongate but not absolutely parallel- upper surface faintly impressed; lower surface with sided elytra, deeply impressed, barely punctate a row of 6-8 stout setae along basal part of inner elytral striae, and short, straight, parallel-sided, at margin. Mesotibia moderately elongate, dorsal rim tip slightly incurved apex of the aedeagus. crenulate, with several rows of setae on the dorsal surface; subapical spur elongate. Metatibia moder- Description ately elongate, rather sparsely setose. Metatarsomere shorter than the following three tarsomeres. Measurements. Length: 10.6-14.5 mm; width: 2.9- Male genitalia (Fig. 21). Aedeagus wide, fairly 3.75 mm. Ratios. Length/width of pronotum: 1.10- elongate, narrowed apicad; lower surface in basal 1.13; base/apex of pronotum: 1.40-1.51; width third concave, then almost straight, but the apex pronotum/head: 1.47-1.52; length/width of elytra: slightly bent down, surface longitudinally striolate. 2.06-2.12; length/width of protibia: M: 3.35-3.5, Apical half of lower surface with two longitudinal, W: 2.9-3.1; length/width of metatibia: M; 5.25-5.5, parallel ridges, the surface between concave. Api- W: 5.2-5.3. cal part asymmetrically narrowed; apex elongate, Colour (Fig. 127). Uniformly black. Legs dark straight, very narrow, parallel, near tip bent down; piceous to black. tip obtuse. Both parameres short and stout, the left Head (Fig. 127). Eye fairly large, laterally moder- one considerably stouter than the narrower right ately projected, orbit short, oblique-convex. Clypeus one; both with short but acute, asetose apical part. with rather shallow, wide, almost quadrangular

232 Female gonocoxites (Fig. 59). Gonocoxites nar- pressed, impunctate. Basal sulcus narrow. Anterior row and elongate, little curved, with rather acute marginal seta situated at apical fifth. Basal groove apex; with one elongate, rather stout seta basally at shallow, elongate, posteriorly deepened, impunctate. lateral surface and a smaller seta below, two elongate Disk with some fine, transverse strioles in middle, setae on the ventral surface of gonocoxite 1, three impunctate, without microreticulation, very glossy. or four setae at middle of the medio-dorsal surface, Elytra. Rather elongate, almost parallel-sid- and with a tiny nematiform seta near apex. Lateral ed, slightly widened apicad, convex but on disk plate basally with one elongate seta, apically with depressed. Basal angle little produced. Striation two or three shorter seta. complete, but lateral striae less distinct. Striae Variation. Considerable variation noted in body deep, becoming slightly shallower towards apex, size. impunctate. Intervals slightly convex, impunctate, without transverse strioles, without perceptible Distribution. Northern NT, north QLD: CYP; also microreticulation, very glossy. Epipleura narrow, on islands in the Gulf of Carpentaria and in Torres basal sulcus indistinct. Strait. Metathoracic wings. Fully developed. Lower surface. Proepisternum impunctate, with Collecting circumstances. Little recorded. More few irregular, very faint, elongate dorso-ventral recently sampled specimens collected at light near strioles and some short transverse strioles, markedly lagoons, and in pitfall traps. glabrous, with faint isodiametric microreticulation. The whole lower surface with fine, isodiametric Clivina dubia, spec. nov. microreticulation. Metepisternum elongate, slightly Figs 22, 128 > 2 × as long as wide, in posterior half with a deep longitudinal sulcus. Abdominal sterna impunctate. Examined types. Holotype: M, Austr / Soc.Ent.Belg. Legs. Protibia narrow and elongate. Longitudi- Coll. Putzeys (IRSNB). nal sulcus on upper surface well impressed; lower surface with a row of 6-8 stout setae along basal Etymology. The name refers to unknown type locality. part of inner margin. Mesotibia moderately elongate, Diagnosis. Medium sized, black species with deep, dorsal rim crenulate, with several rows of setae on quadrangular clypeal excision, impressed clypeal the dorsal surface; subapical spur elongate. Metatibia suture, sparsely punctate head, rufo-piceous legs, moderately elongate, rather sparsely setose. Metatar- rather conical prothorax, impunctate and very gla- somere shorter than the following three tarsomeres. brous proepisternum, moderately elongate, almost Male genitalia (Fig. 22). Aedeagus very wide, parallel-sided elytra, impunctate elytral striae, and moderately elongate, very asymmetrically narrowed elongate, parallel-sided curved left apex of the ae- apicad; lower surface in basal half concave, then deagus with obtuse tip. almost straight. Apical half of lower surface slightly concave. Apex moderately elongate, parallel-sided, Description conspicuously curved left, tip obtusely rounded. Both parameres moderately elongate, the left one Measurements. Length: 12.8 mm; width: 3.45 mm. stouter than the right one; both with rather short, Ratios. Length/width of pronotum: 1.05; base/ narrow, asetose apical part. apex of pronotum: 1.2; width pronotum/head: 1.35; Female gonocoxites. Unknown. length/width of elytra: 2.0; length/width of protibia: Variation. Unknown. M: 3.3; length/width of metatibia: M; 5.25. Colour (Fig. 128). Unicolourous black. Middle Distribution. Not recorded. and hind legs rufous, paler than the anterior leg. Head (Fig. 128). Eye fairly large, laterally mod- Collecting circumstances. Not recorded. erately projected, orbit raletively long, oblique. Cly peus with moderately deep, almost quadrangular excision. Labrum 7-setose. Clypeus divided from Clivina regularis Sloane frons by a deep and wide sulcus; upper surface Figs 23, 60, 87 without transverse sulci, rather sparsely punctate. Surface without microreticulation, glossy. Clivina regularis Sloane, 1896a: 238. – Sloane 1896a: 227; Pronotum (Fig. 128). Elongate, conical, dorsally Csiki 1927: 510; Moore et al. 1987: 76; Lorenz 2005: gently convex. Lateral margin straight and oblique, 143. faintly concave in apical third. Apex slightly concave. Examined types. Lectotype (by present designation): Marginal channel narrow; median line well im- M (head missing, male genitalia partly destroyed), New

233 England / Clivina regularis Sl. Cotype / PARATYPE Elytra (Fig. 87). Moderately elongate, but not (blue) (ANIC). absolutely parallel-sided, slightly widened apicad, Type locality. “New England”, New South Wales. convex but on disk depressed. Basal angle little produced. Striation complete, but lateral striae less Other material (6 ex.). QLD: 1 M, Queensland AUS- distinct. Striae deep, becoming slightly shallower TRALIA Challenger Exp / Clivina regularis Sl. det. K. towards apex, barely punctate. Intervals slightly Kult / Collectio Karel Kult Coll.A.Dostal,1989 convex, impunctate, without or with very weak (CDW); 1 W, Oakey Q. 25 1 1982 P. Alsopp / T 5436 (QD- transverse strioles, without microreticulation, very PIB); 1 M, C 2189 (QDPIB); 1 W, Queensland Challenger glossy. Epipleura narrow, basal sulcus shallow. Exp. 85-44 / Clivina regularis Sl. det. K. Kult (NHM); 2 MM (defect), New England (CBM, MMS). Metathoracic wings. Fully developed. Lower surface. Proepisternum impunctate, and Diagnosis. Rather small to medium sized, dark almost without strioles, remarkably glabrous, with piceous to black species with wide and shallow, faint isodiametric microreticulation. The whole lower quadrangular clypeal excision, not sulcate, moder- surface with fine, isodiametric microreticulation. ately densely punctate head, dark rufous middle and Metepisternum elongate, c. 2 × as long as wide, in hind legs, slightly oblique lateral pronotal margin, posterior half with a deep longitudinal sulcus. Ab- impunctate and glabrous proepisternum, elongate, dominal sterna impunctate. but not completely parallel-sided elytra, and short, Legs. Protibia moderately narrow and elongate, obtusely triangular apex of the aedeagus that is in the male narrower than in the female. Longitudinal slightly curved left. sulcus on upper surface distinct; lower surface with a row of 6-8 stout setae along basal part of inner Description margin. Mesotibia moderately elongate, dorsal rim Measurements. Length: 9.2-11.9 mm; width: 2.65- crenulate, with several rows of setae on the dorsal 3.3 mm. Ratios. Length/width of pronotum: 1.05- surface; subapical spur elongate. Metatibia fairly 1.11; base/apex of pronotum: 1.22-1.27; width elongate, rather sparsely setose. Metatarsomere pronotum/head: 1.31-1.40; length/width of elytra: shorter than the following three tarsomeres. 1.93-2.02; length/width of protibia: M: 2.7-2.85, Male genitalia (Fig. 23). Aedeagus wide, mod- W: 2.35-2.45; length/width of metatibia: M; 5.7-5.8, erately elongate, very asymmetrically narrowed W: 5.7-5.8. apicad; lower surface in basal half gently concave, Colour (Fig. 87). Unicolourous dark piceous to then almost straight. Apical half of lower surface black. Middle and hind legs dark rufous, slightly gently concave. Apex moderately elongate, slightly paler than the anterior leg. curved left, gently narrowed, at tip obtusely convex. Head (Fig. 87). Eye fairly large, laterally mod- Both parameres rather stout, the left one considerably erately projected, orbit comparatively elongate, stouter than the right one; both with rather short, oblique-convex. Clypeus with rather shallow, wide, narrow, asetose apical part. almost quadrangular excision, though the lateral Female gonocoxites (Fig. 60). Gonocoxites rather margins of the excision being rather oblique. Labrum narrow but fairly short, little curved, with obtusely 7-setose. Clypeus not divided from frons; upper transverse apex; with one elongate, rather stout seta surface smooth, without transverse sulci, moder- basally at lateral surface and a smaller seta below, ately densely but finely punctate. Surface without two elongate setae on the ventral surface of gonocox- microreticulation, glossy. ite 1, two stout setae at middle of the medio-dorsal Pronotum (Fig. 87). Rather elongate, slightly surface, without a nematiform seta near apex. Lateral narrowed apicad, dorsally gently convex. Lateral plate basally with one elongate seta, apically with margin straight and oblique, faintly concave in api- two shorter seta. cal third. Apex distinctly concave. Marginal channel Variation. Some variation noted in body size narrow; median line well impressed, impunctate. and in relative length of prothorax. Basal sulcus narrow. Anterior marginal seta situ- Distribution. ated in front of apical fifth. Basal groove fairly deep, North-eastern NSW, south-eastern elongate, linear, straight, impunctate, deepest pos- QLD. teriorly. Disk with or without some fine, transverse Collecting circumstances. Not recorded. strioles in middle, extremely finely punctate, without microreticulation, very glossy.

234 elegans group Clivina elegans Putzeys Figs 24, 61, 79, 88 Diagnosis. Rather small to large species with shallow to moderately deep, more or less quadrangular Clivina elegans Putzeys, 1862: 44. – Putzeys 1866: 36; 1867: clypeal excision; hind body almost always widened 179; Sloane 1896a: 231, 233; Sloane 1905a: 731; 1916: posteriad or oval shaped; clypeus 5- or more com- 605; Csiki 1927: 502; Moore et al. 1987: 70; Lorenz monly 7-setose, but in some species the number of 2005: 142. setae varies from 2-8. The aedeagus usually has an Examined types. Holotype: M, Austral. Sommerset / arrow-shaped, or club-shaped, or claw-shaped, or C.E.By. / Soc. Ent. Belg. Coll. Putzeys / det Putzeys spatulate apex, except C. robusta Sloane, the aedeagus Clivina elegans Putz. / TYPE (red) (IRSNB). of which matches the aedeagi of the procera group. Type locality. “Somerset”, North Queensland. Several species possess shortened metathoracic wings and a short metepisternum. Other material (52 ex.). QLD: Lockerbie N.Cape York, This species group is less homogeneous in body Q. Jan.’58 Darlingtons (CBM, MCZ); Lockerbie, Cape shape and structure than the procera group, and is York, N.Qld. 10.-15.VI.1969 G. B. Monteith UQIC Reg. herein subdivided into subgroups of apparently #90497 (QMB); Lockerbie, N.Q. 7-14.Apr.1977 A. & M. more closely related species. Walford-Huggins / WALFORD-HUGGINS COLLEC- TION Carnegie Museum Accession 35338 / Clivina Distribution. 33 species and one subspecies. The elegans Putz. det. B. P. Moore ’86 (CMP); Lockerbie, Q. majority is distributed in northern Australia. 3.iv.64 I.F.B.Common & M.S. Upton (ANIC); Lockerbie (N.Qld) 16.Xii.14 D.J.Rogers (QMB); 3km E Locker- bie, Cape York, N.Qld. 9-23 Mar 1987 G.B. Monteith elegans subgroup rainforest pitfall traps (CBM, QMB); 3km E Lockerbie, Cape York, N.Qld. Jan.30-Feb.4, 1975 G.B. Monteith Rainforest Pitfall trap No. 1/3, 2/10 (QMB); Lockerbie Diagnosis. Rather large species with conical protho- Area, Cape York, N.Qld. 13-27 Apr., 1973 G.B. Mon- rax and elongate-oval elytra; clypeal excision wide teith (QMB); Bamaga Cape York, N.Qld. 18-25 Mar and shallow; clypeus 5- or 7-setose; aedeagus with 1987 G.B. Monteith (QMB); Bamaga Qld M. Golding elongate apical stalk and small, club-shaped apex; / March’84 / Golding/Powell Collection donated 12 in most species the metathoracal wings are reduced, Feb 2002 (WAM); AUSTRALIA: CYP Bamaga.II.1994 Most of the 7 species occur in northern Australia, G. A. Wood at light (DPIM); Prince of Wales Is., Tor- only one in south-western QLD. res Strait, N. Qld. 6-15.ii.1975 E. Cameron / Strand vegetation (QMB); Qld: 10°36.0'Sx142°18.2'E Horn Is. Quarry RF. 60m. 1-8Jan2008. open forest. G.Monteith & The Clivina elegans complex K.Aland. dung & fungus trap 15483, 15493 (QMB); Qld: 10°36.6'Sx142°17.7'E Horn Is. Airport. 60m. 1-3Apr2008. K.Aland. open forest. dung & mushroom pitfall. 10m Three populations of quite similarly shaped and 15954 (QMB); Thursday Is. NQ. 19.3.75 W.WALSH. / structured species with elongate, oviform elytra, WALFORD-HUGGINS COLLECTION Carnegie Mu- more or less distinctly arrow-shaped apex of the seum Accession 35338 / Clivina procera Putzeys [W det. aedeagus, and serrulate margin of one of the folds by A.Walford-Huggins] / Clivina elegans Putz. det. in the internal sac occur in the northern half of the M.Baehr’95 (CMP); Cape York Eigner (QMB); Cape Cape York Peninsula. Two of these already have York Queensland / Lea has not / Clivina Cape York received names as separate species. The third, un- (QMB); Murray (?) Australia / Collectio Karel Kult named population which occurs between the ranges (CDW); Queensland Gehr. W. Müller Vermächtnis of these species, is also intermediary in some charac- (SMTD). – AUS: Bunawang J.J.F. / Clivina elegans Putz. ters. Therefore the question arises which taxonomic Id. by T. G. Sloane (ANIC). status should be attributed to the three populations, Diagnosis. Rather large to large, black species with i.e., whether all three populations should be regarded wide and rather shallow, quadrangular clypeal ex- separate species or subspecies of a single species. cision, glabrous, not sulcate head, usually 5-setose This is a matter of opinion, but for easier use, and labrum, black legs, convexly oblique lateral prono- because the characterization of subspecies is much tal margin, impunctate proepisternum, elongate, more difficult than that of species, for the present I oviform elytra, finely punctate elytral striae, sym- prefer to regard all three populations distinct species, metric, obtusely arrow shaped apical part of the being aware, however, that they are closely related. aedeagus, and strongly serrate margin of one fold Perhaps application of molecular techniques may in the internal sac. clarify the question in future.

235 Description Male genitalia (Fig. 24). Aedeagus large, length > 3 mm; moderately wide, elongate, almost regularly Measurements. Length: 11.8-17.5 mm; width: 2.9- narrowed apicad; lower surface in basal fourth con- 4.7 mm. Ratios. Length/width of pronotum: 1.09- cave, in middle almost straight to gently concave, in 1.18; base/apex of pronotum: 1.36-1.42; width apical fourth concave again. Apical part very nar- pronotum/head: 1.34-1.52; length/width of elytra: row, almost symmetric; apex short, wide, obtusely 2.02-2.27; length/width of protibia: M: 3.7-3.8, W: 3.4- trigonal to arrow-shaped, tip rounded. Lower surface 3.5; length/width of metatibia: M: 7.2-7.4, W: 6.5-6.7. not or little concave. One of the folds in the internal Colour (Fig. 88). Unicolourous black. Legs black. sac strongly denticulate on the upper surface. Both Head (Figs 79, 88). Eye moderately large, later- 1 parameres very elongate, the left one considerably ally projected, but orbit rather elongate, c. /3 of length stouter than the right one; both with elongate, very of eye, oblique-convex. Clypeus with wide and rather narrow, unisetose or asetose apical part; setae if shallow, almost quadrangular excision, though the present very short, situated right at tip. lateral margins of the excision markedly oblique. Female gonocoxites (Fig. 61). Gonocoxites rather Labrum usually 5-setose, rarely 7-setose. Clypeus narrow but fairly short, little curved, with obtusely not or shallowly divided from frons; upper surface transverse apex; with one elongate, rather stout seta rather convex, glabrous, without any sulci, finely and basally at lateral surface and a smaller seta below, sparsely punctate. Vertex with a shallow transverse two elongate setae on the ventral surface of gono- sulcus. Surface without microreticulation, glossy. coxite 1, four setae at middle of the medio-dorsal Pronotum (Fig. 88). Elongate, considerably nar- surface, and a tiny nematiform seta near apex. Lateral rowed apicad, dorsally gently convex. Lateral margin plate basally with one elongate seta, apically with oblique and convex. Apex slightly concave. Marginal one or two shorter seta. channel narrow; median line impressed but shallow, Variation. This is a very variable species, in impunctate. Basal sulcus narrow. Anterior marginal body size, shape of prothorax and elytra, and dis- seta situated at apical fifth. Basal groove shallow, tinctness of the punctation of the elytral striae; the elongate, linear, straight, impunctate. Disk with some aedeagus, however, is very similar. Specimens from fine, transverse strioles in middle, very sparsely and Horn Island (Torres Strait Islands) are large and extremely finely punctate, with very faint, superfi- compact, and they almost lack the punctation of the cial, isodiametric microreticulation, glossy. striae. Specimens from Lockerbie near the tip of CYP Elytra (Fig. 88). Elongate, oval, widened towards usually are also large and rather compact, but the middle, convex but on disk depressed. Basal angle striae are distinctly and rather coarsely punctate. little produced. Striation incomplete, 6th stria very Most specimens from Bamaga, slightly more to the shallow, 7th stria usually absent. Median striae deep, south, are rather small, with narrow prothorax and becoming slightly shallower towards apex, moder- slender and elongate, or very elongate, elytra, and ately finely punctate at least in basal half. Intervals usually with coarse punctation of the striae. Because slightly convex, impunctate, without or with very some of these character states are somewhat mixed weak transverse strioles, with fine though distinct, within the populations, no taxonomical subdivision isodiametric microreticulation, glossy. Epipleura of the species has been attempted. narrow, basal sulcus barely indicated. Metathoracic wings. Almost completely re- Distribution. Tip of Cape York Peninsula and Torres duced. Strait Islands, extreme northern QLD. Lower surface. Proepisternum impunctate, almost not striolate, with faint isodiametric micro- Collecting circumstances. Largely unrecorded. reticulation. The whole lower surface with fine, Single specimens were collected at light, in pitfall isodiametric microreticulation. Metepisternum trap or “fungus traps”, in rain forest, open forest short, c. 1.25 × as long as wide. Abdominal sterna and strand vegetation. The species does not seem impunctate. to be definitively hygrophilous. Legs. Protibia very narrow and elongate, in the male slightly more than in the female. Longitudinal sulcus on upper surface barely visible; lower surface Clivina interposita, spec. nov. with a row of 3 stout setae along basal part of inner Figs 25, 89 margin. Mesotibia moderately elongate, dorsal rim Examined types. Holotype: M, 11.51S 142.38 QLD crenulate, with two rows of few setae on the dorsal 12km SSE Heathlands 26Jan.-1.Mar.1992 P.Feehney surface; subapical spur elongate. Metatibia elongate, FIT #2 / F.I.T. ANIC 1241 closed forest #2 (ANIC). – rather sparsely setose. Metatarsomere 1 almost as Paratypes: 2 MM, 6 WW, 11.51S 142.38 QLD 12km SSE long as the following three tarsomeres. Heathlands 26Jan.-1.Mar.1992 P.Feehney FIT #2 /

236 F.I.T. ANIC 1241 closed forest #2 (ANIC, CBM); seta situated about at apical fourth. Basal groove shal- 2 WW, 11.51S 142.38 QLD 12km SSE Heathlands 2-22 low, elongate, linear, straight, impunctate. Disk with Mar.1992 P.Feehney FIT #2 / F.I.T. ANIC 1241 closed some fine, irregularly transverse strioles in middle, forest #2 (ANIC); 3 MM, 4 WW, 11.51S 142.38 QLD very sparsely and extremely finely punctate, with 12km SSE Heathlands 15-26 Jan 1992 T.A.Weir, very faint, superficial, isodiametric microreticula- I.D.Naumann / F.I.T. ANIC 1221 closed forest #2 tion, glossy. (ANIC, CBM); 1 W, 11.45S 142.35E Heathlands QLD Elytra (Fig. 89). Rather elongate, oval, consider- Jan.-Feb.1993 J.Stuart pitfall traps / unburnt grass- ably narrowed towards base, convex but on disk land (ANIC); 1 W, 11.45S 142.35E, Heathlands QLD depressed. Basal angle little produced. Striation 1 - 6 Apr 1993 by hand P. Zborowski (ANIC); 1 W, complete, but 7th stria usually fine. Median striae 11.37E 142.48E QLD 5 km WNW Captain Billy Land- deep, becoming slightly shallower towards apex, ing 2 Apr 1993 at light monsoon forest P.Zborowski moderately coarsely punctate at least in basal half. In- (ANIC); 2 MM, 4 WW, ABRS Area 2, 142°45'E, 11°40'S, tervals slightly convex, impunctate, without or with Dividing Range, 15 km W. of Captain Billy Creek, very weak transverse strioles, with very fine, slightly Cape York Pen. N. Qld. 5-12.ii.1976 G.B.Monteith superficial, isodiametric microreticulation, glossy. (CBM, QMB); 1 W, QLD: 13.8km S 3 ways to Bamaga Epipleura narrow, basal sulcus barely indicated. & Heathlands; 18-22.III.1992; G. Cassis pitfall; rain- Metathoracic wings. Almost completely re- forest / Clivina sp. det. B. P. Moore 1999 (AMS). duced. Lower surface. Proepisternum impunctate, with Etymology. The name refers to the intermediate or without some fine dorso-ventral strioles, with geographical range and also the intermediate mor- faint isodiametric microreticulation. The whole phology between C. elegans and C. kershawi. lower surface with fine, isodiametric microreticula- Diagnosis. Rather large, black species with wide tion. Metepisternum short, < 1.25 × as long as wide. and rather shallow, quadrangular clypeal excision, Abdominal sterna impunctate. glabrous, sulcate head, usually 7-setose labrum, Legs. Protibia very narrow and elongate, in the black legs, convexly oblique lateral pronotal margin, male slightly more than in the female. Longitudinal impunctate proepisternum, elongate, oviform elytra, sulcus on upper surface barely visible; lower surface punctate elytral striae, symmetric, obtusely arrow with a row of 3-4 stout setae along basal part of in- shaped apical part of the aedeagus, and faintly ser- ner margin. Mesotibia moderately elongate, dorsal rate margin of one fold in the internal sac. rim crenulate, with two rows of few setae on the dorsal surface; subapical spur elongate. Metatibia Description elongate, rather sparsely setose. Metatarsomere 1 slightly shorter than the following three tarsomeres. Measurements. Length: 11.7-13.2 mm; width: 3.3- Male genitalia (Fig. 25). Aedeagus compara- 3.6 mm. Ratios. Length/width of pronotum: 1.03- tively small, length < 2.5 mm; moderately wide, 1.08; base/apex of pronotum: 1.43-1.45; width elongate, almost regularly narrowed apicad; lower pronotum/head: 1.44-1.54; length/width of elytra: surface in basal fourth concave, in middle almost 1.94-2.02; length/width of protibia: M: 3.7-3.8, W: 3.3- straight to gently concave, in apical fourth concave 3.5; length/width of metatibia: M: 7.1-7.5, W: 6.4-6.7. again. Apical part very narrow, almost symmetric; Colour (Fig. 89). Unicolourous black. Legs black. apex short, rather narrow, slightly knobbed, tip Head (Fig. 89). Eye moderately large, laterally rounded. Lower surface not or little concave. One 1 projected, but orbit rather elongate, c. /3 of length of of the folds in the internal sac distinctly but faintly eye, oblique-convex. Clypeus with wide and rather denticulate on the upper surface. Both parameres shallow, almost quadrangular excision, though the very elongate, the left one considerably stouter lateral margins of the excision markedly oblique. than the right one; both with elongate, very narrow, Labrum usually 7-setose, rarely 5-setose. Clypeus unisetose or asetose apical part; setae if present very divided from frons by a deep sulcus; upper surface short, situated right at tip. rather convex, glabrous, without any sulci, finely Female gonocoxites. Very similar to those of and sparsely punctate. Vertex with a shallow trans- C. elegans Putzeys. verse sulcus. Surface without microreticulation, Variation. Some variation noted in length of very glossy. elytra, shape and relative length of pronotum, and Pronotum (Fig. 89). Elongate, considerably nar- degree of punctation of the elytral striae. rowed apicad, dorsally gently convex. Lateral margin oblique and convex. Apex slightly concave. Marginal Distribution. Northern Cape York Peninsula south channel narrow; median line impressed but shallow, of Jardine River, i. e. the area around Heathlands, impunctate. Basal sulcus narrow. Anterior marginal northern QLD.

237 Collecting circumstances. Most specimens were Labrum usually 7-setose, rarely 5- or 6-setose. Cly collected in rain forest and monsoon forest, mainly peus divided from frons by a fairly deep sulcus; in pitfall traps, one specimen in “unburnt grass- upper surface rather convex, glabrous, without any land”. The species does not seems to be definitively sulci, variously punctate. Vertex with a shallow hygrophilous. transverse sulcus. Surface without microreticulation, very glossy. Pronotum (Fig. 90). Elongate, considerably nar- Clivina kershawi Sloane rowed apicad, dorsally gently convex. Lateral margin Figs 26, 90 oblique and convex. Apex slightly concave. Marginal channel narrow; median shallow, impunctate. Basal Clivina kershawi Sloane, 1916: 605. – Csiki 1927: 506; Moore sulcus narrow. Anterior marginal seta situated about et al. 1987: 72; Lorenz 2005: 143. at apical fourth. Basal groove absent or barely indi- Examined types. Holotype: M, Coll. by F. G. Kershaw cated. Disk with some fine, irregularly transverse Nat. Mus. Victoria Claudie R. 11.13 Queensland / Cliv- strioles in middle, very sparsely and extremely finely ina kershawi Sl. Id. by T. G. Sloane / Type / Type (red) punctate, with very faint, superficial, isodiametric / Del by T. G. Sloane Esq. Rae 12.6.16 / HOLOTYPE microreticulation, glossy. T-16912 Clivina kershawi Sloane (red) (NMV). – Para- Elytra (Fig. 90). Elongate, oval, but little nar- types: 1 M, same data / Cotype (green) / PARATYPE rowed towards base, convex but on disk depressed. T-16913 Clivina kershawi Sloane (blue) (NMV); 1 (sex?, Basal angle little produced. Striation complete, but only hind body left, but genitalia destroyed), same data 7th stria usually fine. Median striae deep, becoming / Co-type / PARATYPE (blue) (ANIC). slightly shallower towards apex, rather finely punc- Type locality. “Claudie River”, North Queensland. tate at least in basal half. Intervals slightly convex, impunctate, without or with very weak transverse Other material (5 ex.). QLD: 1 M, 2 WW, Cape York Pen. strioles, with very fine, slightly superficial, isodia- Iron Range, NQ 14-21.Apr.1977 WALFORD-HUGGINS metric microreticulation, glossy. Epipleura narrow, / WALFORD-HUGGINS COLLECTION Carnegie Mu- seum Accession 35338 / Clivina elegans Putzeys [Series basal sulcus barely indicated. det. By A.Walford-Huggins] (CMP); 1 W, AUSTRAL- Metathoracic wings. Almost completely reduced. IA: Queensland N NW Cairns, Iron National Park Lower surface. Proepisternum impunctate, al- 12°42.61'S, 143°17.53'E Cook Hut Camp env. 12-18. most without strioles, glabrous, with faint isodiamet- XI.2010, L. Hovorka (CBM); 1 M, Gordon’s Mine Area, ric microreticulation. The whole lower surface with Iron Range, N.Qld. 12.-18.ii.1976 G.B.Monteith Rainfor- fine, isodiametric microreticulation. Metepisternum est (QMB). short, < 1.25 × as long as wide. Abdominal sterna impunctate. Diagnosis. Rather large, black species with wide Legs. Protibia very narrow and elongate, in the and rather shallow, quadrangular clypeal excision, male slightly more than in the female. Longitudinal glabrous, sulcate head, usually 7-setose labrum, sulcus on upper surface barely visible; lower surface black legs, convexly oblique lateral pronotal margin, with a row of 3 stout setae along basal part of inner impunctate proepisternum, elongate, moderately margin. Mesotibia moderately elongate, dorsal rim oviform elytra, little punctate elytral striae, sym- crenulate, with two rows of few setae on the dorsal metric, obtusely arrow shaped apical part of the surface; subapical spur elongate. Metatibia elongate, aedeagus, and faintly serrate margin of one fold in rather sparsely setose. Metatarsomere 1 shorter than the internal sac. the following three tarsomeres. Description Male genitalia (Fig. 26). Aedeagus moderately large, length 2.7-2.8 mm; moderately wide, elongate, Measurements. Length: 12.2-14.1 mm; width: 3.3- almost regularly narrowed apicad; lower surface 3.95 mm. Ratios. Length/width of pronotum: 1.06- in basal fourth concave, in middle almost straight 1.13; base/apex of pronotum: 1.51-1.56; width to gently concave, in apical fourth concave again. pronotum/head: 1.40-1.47; length/width of elytra: Apical part very narrow, almost symmetric; apex 2.03-2.11; length/width of protibia: M: 3.7-3.8, W: 3.3- short, rather narrow, slightly knobbed, tip rounded. 3.4; length/width of metatibia: M: 7.3-7.5, W: 6.5-6.6. Lower surface not or little concave. One of the folds Colour (Fig. 90). Unicolourous black. Legs black. in the internal sac distinctly but faintly denticulate Head (Fig. 90). Eye moderately large, laterally on the upper surface. Both parameres very elongate, 1 projected, but orbit rather elongate, c. /3 of length of the left one considerably stouter than the right one; eye, oblique-convex. Clypeus with wide and rather both with elongate, very narrow, unisetose or asetose shallow, almost quadrangular excision, though the apical part; setae if present fairly elongate, situated lateral margins of the excision markedly oblique. right at tip.

238 Female gonocoxites. Very similar to those of groove barely indicated. Disk with some fine, ir- C. elegans Putzeys. regularly transverse strioles in middle, apparently Variation. Some variation noted in length of impunctate, with very faint, superficial, isodiametric prothorax and elytra. microreticulation, glossy. Elytra (Fig. 91). Moderately elongate, oval, wid- Distribution. Iron Range, central Cape York Pen- est in middle, convex but on disk depressed. Basal insula, northern QLD. angle little produced. Striation incomplete, 6th and 7th stria not or barely visible. three or four median Collecting circumstances. Little recorded; one speci- striae at least in basal half impressed and punctate, men was collected in rain forest. The species does becoming shallower towards apex. Intervals almost not seem to be definitively hygrophilous. depressed, impunctate, without or with very weak transverse strioles, with very fine, slightly superficial, isodiametric microreticulation, glossy. 3rd interval Clivina variseta, spec. nov. with 5-7 discal punctures. Epipleura narrow, basal Figs 27, 91 sulcus barely indicated. Examined types. Holotype: M, Cairns / Lea has not / Metathoracic wings. Almost completely re- Cairns dist. J. A. Anderson / Clivina Cairns (QMT234904). duced. – Paratype: 1 W, AUS15, QLD36, c.25 km ene. Bramwell Lower surface. Proepisternum impunctate, with Jct. Rdh., 115m, 11°56'21.9"S, 142°38'07.2"E, 10.5.2015, or without some very fine dorso-ventral strioles, M.Baehr (CBM). with faint isodiametric microreticulation. The whole lower surface with fine, isodiametric microreticula- Etymology. The name refers to the variable number of tion. Metepisternum short, < 1.25 × as long as wide. elytral punctures and setae, Abdominal sterna impunctate. Diagnosis. Large, black species with wide and Legs. Protibia very narrow and elongate, in the rather shallow, quadrangular clypeal excision, gla- holotype extremely narrow, perhaps due to some brous, sulcate head, 7-setose labrum, black legs, short abrasion of the external teeth. Longitudinal sulcus prothorax with convexly oblique lateral margin, on upper surface shallow but distinct; lower sur- impunctate proepisternum, elongate, oviform elytra face with a row of 3-5 stout setae along basal part with 5-7 discal punctures, fairly impressed, finely of inner margin. Mesotibia moderately elongate, punctate elytral striae, and symmetric, obtusely ar- dorsal rim crenulate, with two rows of rather few row shaped apical part of the aedeagus. setae on the dorsal surface; subapical spur elongate. Metatibia elongate, rather sparsely setose. Metatarsus Description extremely elongate, metatarsomere 1 shorter than the following three tarsomeres. Measurements. Length: 17.0-17.8 mm; width: 5.3- Male genitalia (Fig. 27). Aedeagus moderately 5.4 mm. Ratios. Length/width of pronotum: 0.99- large, length 2.7-2.8 mm; narrow, elongate, slightly 1.01; base/apex of pronotum: 1.55-1.58; width asymmetrically narrowed apicad; lower surface pronotum/head: 1.55-1.57; length/width of elytra: almost regularly concave. Apical part very narrow, 1.83-1.90; length/width of protibia: M: 5.0, W: 3.9; almost symmetric; apex short, comparatively nar- length/width of metatibia: M: 7.3, W: 6.7. row, knobbed to slightly arrow-shaped, tip rounded. Colour (Fig. 91). Unicolourous black. Legs black. Lower surface barely concave. Both parameres Head (Fig. 91). Eye large, laterally well project- elongate, the left one considerably stouter than the ed, orbit short, oblique-convex. Clypeus with wide right one; both with short, narrow, unisetose apical and rather shallow, almost quadrangular excision, part; setae very short, situated right at tip. though the lateral margins of the excision markedly Female gonocoxites. Very similar to those of oblique. Labrum 7-setose. Clypeus divided from C. elegans Putzeys. frons by a fairly deep sulcus; upper surface rather Variation. Some variation noted in distinct- convex, irregularly impressed, sparsely and very ness of the elytral striae which are shallower in the finely punctate.. Surface without microreticulation, holotype. very glossy. Pronotum (Fig. 91). Short and wide, consider- Distribution. Cape York Peninsula, northern QLD. ably narrowed apicad, dorsally rather convex. The holotype is labelled “Cairns district” which does Lateral margin oblique and slightly convex. Apex not automatically mean the immediate vicinity of concave. Marginal channel narrow; median line very Cairns, but at the time when it was written, it could shallow, impunctate. Basal sulcus narrow. Anterior just so mean Cape York Peninsula. marginal seta situated about at apical fourth. Basal

239 Collecting circumstances. The paratype was col- isodiametric microreticulation, rather glossy. Three lected in a pitfall trap exposed near a muddy pit in scutellary punctures present on either side. 3rd stria a disturbed area in tropical woodland. with 14, 5th stria with 11-13, 7th stria with 13-15 small punctures. Epipleura narrow, basal sulcus slightly impressed and punctate. Clivina triseriata, spec. nov. Metathoracic wings. Much reduced. Figs 92, 135 Lower surface. Proepisternum impunctate, with some very fine dorso-ventral strioles, in lower part Examined types. Holotype: W, Australia, NT, Mal- also with faint, short, transverse strioles; with faint abinbandju Billabong, 13 km s. Jabiru, 2.-3.11.1984, M. isodiametric microreticulation. The whole lower Baehr (CBM). surface with fine, isodiametric microreticulation. Etymology. The name refers to the biseriate punctation Metepisternum rather short, c. 1.25 × as long as wide. of the elytra, i. e. the presence of series of setiferous Abdominal sterna impunctate. rd th th punctures on 3 , 5 , and 7 intervals. Legs. Protibia very narrow and elongate. Lon- Diagnosis. Large, elongate, black species with gitudinal sulcus on upper surface only near apex wide and shallow, quadrangular clypeal excision, distinct; lower surface transversely striolate, with a glabrous, barely sulcate head, 7-setose labrum, black row of 7-8 stout setae along basal part of inner mar- legs, trapezoidal prothorax with very oblique lateral gin. Mesotibia elongate, dorsal rim barely crenulate, margin, impunctate proepisternum, elongate, mark- with two rows of many setae on the dorsal surface; edly oviform elytra, and presence of three series of subapical spur elongate. Metatibia very elongate, numerous setiferous punctures on the elytra. rather sparsely setose. Metatarsus very elongate, metatarsomere 1 shorter than the following three Description tarsomeres. Male genitalia. Unknown. Measurements. Length: 15.7 mm; width: 5.0 mm. Female gonocoxites. Very similar to those of Ratios. Length/width of pronotum: 0.98; base/ C. elegans Putzeys. apex of pronotum: 1.65; width pronotum/head: Variation. Unknown. 1.62; length/width of elytra: 1.90; length/width of protibia: W: 3.8; length/width of metatibia: W: 6.6. Distribution. Kakadu NP, northern NT. Known Colour (Fig. 92). Unicolourous black. Legs black. only from type locality. Head (Fig. 92). Eye large, laterally well projected, orbit very short, oblique. Clypeus with wide Collecting circumstances. The holotype was col- and very shallow, almost quadrangular excision, lected in a pitfall trap near the shore of a billabong. though the lateral margins of the excision oblique. Labrum 7-setose. Mandibles short. Clypeus not divided from frons; upper surface rather convex, Clivina inopinata, spec. nov. sparsely, moderately coarsely punctate. Surface Figs 28, 129 without microreticulation, glossy. Pronotum (Fig. 92). Moderately elongate, trap- Examined types. Holotype: M, QLD: 18°09'Sx144°19'E Mount Surprise. 18-29 Jan 2005. A.Ewart. 450m. at light. ezoidal, remarkably narrowed apicad, dorsally rather 12187 (QMT234905). convex. Lateral margin very oblique but almost straight. Apex gently concave. Marginal channel Etymology. The latin name means “surprising” and narrow; median line well impressed, impunctate. refers to the type locality Mt. Surprise. Basal sulcus narrow. Anterior marginal seta situated Diagnosis. Fairly large, black species with wide about at apical third. Basal groove deep, compara- and rather shallow, quadrangular clypeal excision, tively short, linear, impunctate. Disk with some fine, glabrous, not sulcate head, 5-setose labrum, mod- irregularly transverse strioles in middle, impunctate, erately elongate mandible, rufo-piceous legs, short, with very fine but distinct, isodiametric microreticu- trapezoidal prothorax with convexly oblique lateral lation, moderately glossy. margin, impunctate proepisternum, oviform elytra, Elytra (Fig. 92, 135). Soldered together. Elongate, punctate elytral striae, and symmetric, spatulate apex narrow at base, oval shaped, convex but on disk of the aedeagus. depressed. Basal angle little produced. Striation th almost complete, but 7 stria weak. Striae rather Description deep, apicad slightly shallower, almost impunctate. Intervals slightly convex, impunctate, without Measurements. Length: 13.4 mm; width: 4.2 mm. transverse strioles, with fine, slightly superficial, Ratios. Length/width of pronotum: 1.02; base/

240 apex of pronotum: 1.62; width pronotum/head: in apical half with two sharp ridges, surface between 1.54; length/width of elytra: 1.90; length/width of the ridges concave. Both parameres elongate, the left protibia: M: 4.8; length/width of metatibia: M: 8.0. one considerably stouter than the right one; both Colour (Fig. 129). Unicolourous black. Legs with elongate, very narrow, unisetose apical part; rufo-piceous. setae rather short, situated right at tip. Head (Fig. 129). Eye large, laterally well pro- Female gonocoxites. Unknown. jected, orbit very short, oblique. Clypeus with wide Variation. Unknown. and rather shallow, almost quadrangular excision, though the lateral margins of the excision oblique. Distribution. North QLD, west of Atherton Table- Labrum 5-setose. Mandibles moderately elongate. land. Known only from type locality. Clypeus not divided from frons; upper surface Collecting circumstances. The holotype was sam- rather convex, glabrous, rather sparsely and very pled at light. finely punctate. Surface without microreticulation, very glossy. Pronotum (Fig. 129). Rather short and wide, fairly trapezoidal, considerably narrowed apicad, Clivina profundestriolata, spec. nov. dorsally rather convex. Lateral margin oblique and Figs 62, 93 convex. Apex gently concave. Marginal channel narrow; median line faintly impressed, impunctate. Examined types. Holotype: W, Barcaldine, C.Qld. 10 Basal sulcus narrow. Anterior marginal seta situated Feb. 1981 M.S. & B.J.Moulds / Clivina nr. ovipennis Sl. Set. B.P.Moore’81 (ANIC). – Paratype: 1 W, AUS- about at apical fourth. Basal groove very weak, TRALIA. sw Queensland 12km W of Windorah; 110m linear. Disk with some fine, irregularly transverse 28-31.i.2011; sanddunes 25°21'S, 142°32'E; St.Jakl lgt. strioles in middle, apparently impunctate, with faint, (CBM). superficial, isodiametric microreticulation, glossy. Elytra. Moderately elongate, wide at base, not Etymology. The name refers to the strong and deep completely parallel sided, convex but on disk de- strioles on the pronotum. pressed. Basal angle little produced. Striation almost Diagnosis. Fairly large, black species with wide complete, but 7th stria only in basal half visible. Striae but rather deep, quadrangular clypeal excision, punctate, more coarsely in basal half. Intervals slight- glabrous, barely sulcate head, 5-setose labrum, ly convex, impunctate, without transverse strioles, short mandible, rufo-piceous legs, short prothorax with fine, distinct, isodiametric microreticulation, with convexly oblique lateral margin, impunctate moderately glossy. Epipleura narrow, basal sulcus proepisternum, slightly oviform elytra, and deep, slightly impressed and punctate. punctate elytral striae. Metathoracic wings. Fully developed. Lower surface. Proepisternum impunctate, with Description some very fine dorso-ventral strioles, in lower part also with faint, short, transverse strioles; with faint Measurements. Length: 12.9-13.8 mm; width: 4.4- isodiametric microreticulation. The whole lower 4.6 mm. Ratios. Length/width of pronotum: 0.92- surface with fine, isodiametric microreticulation. 0.94; base/apex of pronotum: 1.62-1.66; width Metepisternum moderately elongate, slightly < 2 × pronotum/head: 1.65-1.67; length/width of elytra: as long as wide. Abdominal sterna impunctate. 1.75-1.80; length/width of protibia: W: 3.8; length/ Legs. Protibia very narrow and elongate. Lon- width of metatibia: W: 7.65-7.7. gitudinal sulcus on upper surface barely visible; Colour (Fig. 93). Unicolourous black. Middle lower surface with a row of 4 stout setae along basal and hind legs rufous to piceous, slightly darker than part of inner margin. Mesotibia elongate, dorsal rim the anterior leg. crenulate, with two rows of rather few setae on the Head (Fig. 93). Eye large, laterally well pro- dorsal surface; subapical spur elongate. Metatibia jected, orbit very short, oblique. Clypeus with wide very elongate, rather sparsely setose. Metatarsus but rather deep, almost quadrangular excision. extremely elongate, metatarsomere 1 shorter than Labrum 5-setose. Mandibles rather short. Clypeus the following three tarsomeres. not divided from frons; upper surface rather convex, Male genitalia (Fig. 28). Aedeagus rather wide, moderately densely, finely punctate. Surface without elongate, asymmetrically narrowed apicad; lower microreticulation, very glossy. surface in basal third concave, then convex, towards Pronotum (Fig. 93). Short and wide, consider- apex deeply convex and almost vertically curved ably narrowed apicad, dorsally rather depressed. down. Apex short, rather wide, spatulate and at Lateral margin oblique and convex. Apex gently lower surface denticulate, tip rounded. Lower surface concave. Marginal channel narrow; median line

241 faintly impressed, impunctate. Basal sulcus nar- gracilipes subgroup row. Anterior marginal seta situated about at apical fourth. Basal groove weak, linear, impunctate. Disk Diagnosis. Moderately large species with conical with some fine, irregularly transverse strioles in mid- prothorax and short-oval elytra; clypeal excision dle, very sparsely and extremely finely punctate, with wide and shallow; clypeus 5- or 7-setose; aedeagus extremely faint and superficial traces of isodiametric with small, club-shaped apex: metathoracal wings microreticulation, very glossy. fully developed. The most striking character is the Elytra (Fig. 93). Rather short, wide at base, narrow and elongate mandible. Two species and one slightly oval, convex but on disk depressed. Basal subspecies occur in northern Australia. angle little produced. Striation almost complete, but 7th stria only in basal half visible. Striae in basal half deep, apicad increasingly shallower, punctate, Clivina gracilipes Sloane more coarsely in basal half. Intervals slightly convex, impunctate, with or without fine transverse strioles, Clivina gracilipes Sloane, 1896a: 247. with fine, slightly superficial, isodiametric micro- Note. This characteristically shaped species occurs reticulation, rather glossy. Epipleura narrow, basal in two populations which are slightly differently sulcus slightly impressed and punctate. shaped and occupy widely separated areas: the Metathoracic wings. Fully developed. nominate form northern Australia from north- Lower surface. Proepisternum impunctate, with western QLD to the Kimberleys in extreme northern some very fine dorso-ventral strioles, in lower part WA, the other population the northern half of Cape also with faint, short, transverse strioles; with faint York Peninsula. In view of their different body shape isodiametric microreticulation. The whole lower and the disjunct distribution, and because only surface with fine, isodiametric microreticulation. females are so far known of the population from Metepisternum moderately elongate, slightly < 2 × north Queensland, they are provisionally described as long as wide. Abdominal sterna impunctate. as subspecies. Legs. Protibia very narrow and elongate. Longi- tudinal sulcus on upper surface barely visible; lower surface with a row of 5-6 stout setae along basal Clivina gracilipes gracilipes Sloane part of inner margin. Mesotibia elongate, dorsal rim Figs 29, 63, 94 crenulate, with two rows of rather few setae on the dorsal surface; subapical spur elongate. Metatibia Clivina gracilipes Sloane, 1896a: 247. – Sloane 1896a: 228, very elongate, rather sparsely setose. Metatarsus 231; Csiki 1927: 505; Moore et al. 1987: 71; Lorenz elongate, metatarsomere 1 shorter than the following 2005: 143. three tarsomeres. Examined types. Holotype: W, (defect, head and abdo- Male genitalia. Unknown. men lacking), Gulf C.F. 24/4/93 1040 / C. gracilipes, Sl Female gonocoxites (Fig. 62). Gonocoxites rather Burketown Disct. (C.F.) / HOLOTYPE Clivina gracilipes narrow and moderately elongate, little curved, with Sl. PJD (ANIC). rather acute apex; with one elongate, rather stout seta basally at lateral surface and a smaller seta below, Type locality. “Burketown District”, North Queens- two elongate setae on the ventral surface of gono- land. coxite 1, two longer and one shorter setae at middle Other material (112 ex.). NT: 100ml. E.of Kununurra, of the medio-dorsal surface, and a tiny nematiform W.A., light trap, 27.3.66, J.A.Mahon (ANIC, CBM); seta near apex. Lateral plate basally with one elongate 12.51S 132.47E 10km N by E of Mt.Cahill, 21.v.73, E. seta, apically with two or three shorter seta. G.Matthews (ANIC); Keep Creek, 29 Mar.1984 at light Variation. Little variation noted. K. & E. Carnaby (ANIC); Howard Springs 12 May 1981 J.T. Doyen coll. (ANIC); Katherine , XII.57 leg.H.Demarz Distribution. Central and south-western Queens- (ZSM); Mataranka 1 Mar 1967 M.S. Upton (ANIC); Oc- tober Ck. on Borroloola Rd 7.iv.1976 T.Weir. (ANIC); land. 12.48S 132.42E Nourlangie Creek 8km N of Mt.Cahill, Collecting circumstances. Largely unrecorded; 21.v.73, at light, E.G.Matthews (ANIC); 12.48S 132.42E the paratype was sampled in “sanddunes”, most Nourlangie Creek 8km N of Mt.Cahill, 21.v.73, T.Weir probably at light. & T.Angeles / 12370 (NTD); 12.49S 132.44E Nourlangie Creek 6km NE. of Mt.Cahill, 2.v.73, T.Weir & T.Angeles / 12396 (NTD); 12.51S 132.47E 8km E by N of Mt.Cahill, 22.v.73, T.Weir & T.Angeles / 12334 (NTD); 12.20S 131.19E Nabarlek Dam, 15km S by W. of Nimbuwah Rock, 2.vi.73, at light, E.G.Matthews (ANIC); 12.06S

242 133.04E Cooper Creek, 19km E by SW. of Mt.Borradaile, Banka env., road to Tennant Creek, 12-14.i.2009, St. Jakl 5.vi.73, M.S.Upton (ANIC); 12.06S 133.04E Cooper lgt. (CJP); 40km W of Katherine, 63m, 13°15'S, 130°44'E Creek, 19km E by SW. of Mt.Borradaile, 31.vi.73, at 28.11.2008, Sv. Bilý leg. (CBM, NMPC); 3/06 Kakadu NP light, E.G.Matthews (ANIC); 12.06S 133.04E Cooper Muirella, 10.06.2006 40m S12°51'15" E132°45'16" Light Ck., 19km E by SW. of Mt.Borradaile, 5.vi.73, T.Weir lg Berger-Dostal (CDW); Katherine, at light, 6-10.ii.68 & A.Allwood / 12300 (NTD); 12.06S 133.04E Cooper E. Matthews (ANIC). – WA: 15-16.iv.2009, Hidden Val- Ck., 19km E by SW. of Mt.Borradaile, 5.vi.73, T.Weir ley NP, Kununurra 15°46S 128°44E 64 m alt. St. Jakl lgt. & A.Allwood / 12300, 12414 (NTD); 12.06S 133.04E (CJP); 12m alt. Home Valley Station 14°42'S 127°51'E; Cooper Ck., 19km E by SW. of Mt.Borradaile, 31.vi.73, 15-18.iv.2009 nr. Pentecost river; S.Jakl lgt. (CBP); Fitz- T.Weir & N.Forrester / 12467 (NTD); 12.57S 132.32E roy Crossing, 125:35E, 18:10S. 17 Mar., 23 Mar 1984, at Jim Jim Ck. 19km WSW. of Mt.Cahill, 19.v.73, T.Weir & light K. & E. Carnaby (ANIC); Wyndham 20.iv.1975 K. T.Angeles / 11284, 11291-4 (NTD); 12.54S 132.32E Jim & E. Carnaby (ANIC); Kimberley, 3m, Wyndham env., Jim Airstrip 1.v.1973, T.Weir & T.Angeles / 11289-90 15°31'S 128°10'E, 20-21.iv.2009, St. Jakl lgt. (CJP). (NTD); 12.36S 132.52E Magela Ck. 1km NNW. of Mudg- inberry H.S., 25.v.1973, T.Weir & N.Forrester (ANIC, Diagnosis. Medium sized to fairly large, black spe- NTD 12230); 12.25S 132.58E 1km N. of Cahills Crossing cies with wide and rather shallow, quadrangular (East Alligator R.) 29.v.1973 T.Weir & N.Forrester / clypeal excision, glabrous, not sulcate head, usually 11243 (NTD); 12.31S 132.54E 9km N by E of Mudg- 5-setose labrum, narrow and elongate mandible, inberry H.S., 28.v.1973 T.Weir & N.Forrester / 11233, rufous to dark piceous legs, short, trapezoidal 35 (NTD); Milner Lagoon nr. Dunmara 16.41S 133.25E prothorax with convexly oblique lateral margin, M.V.Light 2 Apr 1981 M.Malipatil & J.Hawkins (NTD); impunctate proepisternum, oviform elytra, deep, Kakadu NP c. 1km S of Arnhem Hwy on Pine Creek punctate median elytral striae, and symmetric, Rd. M.V.Light 25-30 Mar 1980 M.B.Malipatil (NTD); knobbed apex of the aedeagus. Junction of Arnhem Hwy&Oenpelli Road M.V.Light 26-27 Jun 1980 M.B.Malipatil (NTD); (am Stuart Hwy.) Description ca. 25 km N Elliott (Bauplatz) 20.IV.2011 LF; H=274m S17°23'59.9" E133°27'12.2" leg. Michael Langer (CBM); Measurements. Length: 9.5-13.4 mm; width: 3.15- Kakadu 10-14.I.2004 Lgt. R. Novák (CKZ); Kakadu N.P. 4.5 mm. Ratios. Length/width of pronotum: 0.94- 22-25/3/93 Cooinda at light/al lume Leg. L. Toledano 0.96; base/apex of pronotum: 1.51-1.61; width / Clivina gracilipes Sl. det.M.Baehr’96 (CBM, CTV, CGT); pronotum/head: 1.42-1.59; length/width of elytra: Bachelor 26-27/3/93 at light/al lume Leg. L. Toledano Clivina gracilipes Sl. det.M.Baehr’96 (CBM); 70km SW of 1.76-1.80; length/width of protibia: M: 4.7-4.9, W: 3.5- Mataranka, 14-15.1.09, 15°19'S 132°50'E 190m, Sv. Bilý 3.7; length/width of metatibia: M: 8.0-8.3, W: 7.5-7.6. leg. (CBM, NMPC); 190m alt. 70km SW of Mataranka Colour (Fig. 94). Unicolourous black. Legs ru- 15°19'S 132°50'E 14-16.I.2009 St Jakl (CBM, CBP, CJP); fous to dark piceous, in latter case tibiae usually 190m alt. 70km SW of Mataranka 15°19'S 132°50'E 22-23. slightly paler than femora. XII.2008 L.Hovorka lgt. (CBM, CBP); Coll. C. Lecourt Head (Fig. 94). Eye large, laterally well project- 01/2000 Darwin (CBP); Litchfield NP Adelaide River; ed, orbit short, oblique-convex. Clypeus with wide 29.V.2000 leg.M.Langer (CBP); AUS13, NT25, c.12 km and rather shallow, almost quadrangular excision, wnw. Elliott, 204m, 17°37'06"S, 133°28'31"E, 17.4.2013, though the lateral margins of the excision markedly M.Baehr (CBM); AUS13, NT23, 5 km s. Larrimah, Bir- oblique. Labrum most commonly 5-setose, rarely dum Crk., 184m, 15°32'02"S, 133°22'33"E, 16.4.2013, 3- or 7-setose. Mandibles narrow and very elongate, M.Baehr (CBM); 17m alt, Kakadu NP, Ubirr env, 12°25'S 132°57'E 25-26.iv.2009, St. Jakl lgt. (CBM, CJP); 15.45S regularly curved, with very acute tip. Clypeus not 129.06E GPS Keep River Xing 1 km ENE Jarmarm Keep divided from frons; upper surface rather convex, River Nat.Pk. 25-27 May 2001 T.Weir, P.Bouchard / at glabrous, rather sparsely and finely punctate. Surface light open forest (ANIC); 12.22S 133.01E 6km SW by without microreticulation, very glossy. S of Oenpelli, 30.v.1973 T.Weir & N.Forrester / 11250 Pronotum (Fig. 94). Short and wide, markedly (NTD); East Alligaror R. 10 ml.s.W.Oenpelli Misson, trapezoidal, considerably narrowed apicad, dorsally 22.vi.1971 A.Allwood & T.Angeles / Clivina sp. / rather convex. Lateral margin oblique and slightly 11229 (NTD); King R. Coll by W.Melliman 24. 12. 16 convex. Apex almost straight. Marginal channel / Clivina gracilipes Sl. Det. by T.G.Sloane 3. 17 / COL- narrow; median line slightly impressed, impunc- 5324-5 (NMV); King R. Coll by W.Melliman 24. 12. 16 tate. Basal sulcus narrow. Anterior marginal seta / Clivina gracilipes Sl. Det. by T.G.Sloane (Legs darker situated about at apical fourth. Basal groove very than type) (ANIC); Oenpili (unreadable) P.Cahill leg 3. 12. 19 / Conspecific with Clivina gracilipes S. Id by T G. weak, linear, very elongate. Disk with some fine, Sloane / COL-5323 (NMV); Kakadu N.P. 22-25/3/93 irregularly transverse strioles in middle, apparently Leg Leech / Clivina gracilipes Sl. det.M.Baehr’96 (CBM); impunctate, with very faint, superficial, isodiametric Kakadu N.P. 22-25/3/93 Leg Leech / Cooinda / Clivina microreticulation, glossy. sp. Det. P. M. Giachino 1995 (CGT); 318m alt. 14°53'S Elytra (Fig. 94). Rather short, wide at base, oval, (wrong latitude, must be 18°53'S!), 132°01'E, Banka widest in middle, convex but on disk depressed.

243 Basal angle little produced. Striation incomplete, 1st- Collecting circumstances. Several specimens were 6th striae in basal half distinct, 7th stria not or barely collected at light, usually near rivers, larger creeks, visible, from second third outer striae succeedingly, and lagoons therefore the apical third laterally estriate. Striae near base distinctly punctate, the four or five median striae punctate at least in basal half. Intervals slightly Clivina gracilipes longior, subspec. nov. convex, impunctate, without transverse strioles, with fine, distinct, isodiametric microreticulation, Examined types. Holotype: W, Iron Range , Cape York moderately glossy. Epipleura narrow, basal sulcus Pen., N.Qld. 26-31 May, 1971 G. B. Monteith / UQIC more or less impressed. Reg. #90544 (QMT234906). – Paratypes: 1 W, 6903 N.Qu Metathoracic wings. Fully developed. / ? gracilipes Sl. immature / I.6865 Clivina gracilipes Sl.? immature Queensland (SAMA); 1 W, AUSTRALIEN Lower surface. Proepisternum impunctate, with VICTORIA Cap York. / Sammlung H. HESSE SMNS some very fine dorso-ventral strioles, in lower part 1995 (SMNS); 2 WW, Stewart R., Q. Jan.-Feb. 1927 Hale also with faint, short, transverse strioles; with faint & Tindale (SAMA No. 25-033728, 25-033653); 1 W, Iron isodiametric microreticulation. The whole lower Range Cape York, Q. Jan.’58 Darlingtons (MCZ); 1 W, surface with fine, isodiametric microreticulation. Silver Plains Cape York, Q. May-June ’58 Darlingtons Metepisternum rather short, c. 1.5 × as long as wide. / W / gracilipes Sl. det.’59 Darlington / Clivina gracilipes Abdominal sterna impunctate. Sl. (MCZ); 1 M, Silver Plains Cape York, Q. May-June ’58 Legs. Protibia very narrow and elongate, less Darlingtons / M / return / gracilipes (MCZ); 1 W, Silver so in females. Longitudinal sulcus on upper surface Plains Cape York, Q. Jan.’58 Darlingtons (CBM); 1 M, absent; lower surface with a row of 3-5 stout setae found dead in dry sand place / Coen, C.York V. 9 ’82, along basal part of inner margin. Mesotibia elongate, Q. / Australia, Harvard Ex., Darlington / ovalipennis (ll ovipennis) Sl. det. 59 Darlington / Clivina ovalipennis dorsal rim barely crenulate, with two rows of rather Sl. (MCZ). few setae on the dorsal surface; subapical spur short. Metatibia very elongate, rather sparsely setose. Meta- Etymology. The name refers to the longer elytra as tarsus extremely elongate, metatarsomere 1 shorter compared with C. g. gracilipes Sloane, 1896. than the following three tarsomeres. Diagnosis. Distinguished from the nominate sub- Male genitalia (Fig. 29). Aedeagus rather wide, species by significantly longer elytra and longer elongate, regularly narrowed apicad; lower surface elytral striae. in basal third concave, in apical part straight, only near apex slightly concave. Apical part almost sym- Description metrically triagonal; apex short, straight, narrow, gently knobbed, tip slightly turned down, obtusely Measurements. Length: 11.4-12.9 mm; width: 3.7- rounded. Lower surface in apical half with two sharp 4.15 mm. Ratios. Length/width of pronotum: 0.92- ridges, surface between the ridges concave. Both 0.96; base/apex of pronotum: 1.47-1.54; width parameres very elongate, the left one considerably pronotum/head: 1.43-1.53; length/width of elytra: stouter than the right one; both with elongate, very 1.89-1.91; length/width of protibia: W: 3.5-3.6; narrow, uni- or bisetose apical part; setae very short, length/width of metatibia: W: 7.5-7.6. situated right at tip. Colour. As in the nominate subspecies. Female gonocoxites (Fig. 63). Gonocoxites mod- Head. As in the nominate subspecies. erately stout and rather short, little curved, with Pronotum. Much as in the nominate subspecies, obtuse apex; with one elongate, rather stout seta but usually less narrowed apicad. basally at lateral surface and a smaller seta below, Elytra. Much as in the nominate subspecies, but two elongate setae on the ventral surface of gono- elytra decidedly longer and the striae longer, the coxite 1, three setae at middle of the medio-dorsal median ones almost reaching the apex. surface, and a tiny nematiform seta near apex. Lateral Metathoracic wings. Fully developed. plate basally with one elongate seta, apically with Lower surface. As in the nominate subspecies. three shorter seta. Legs. As in the nominate subspecies. Variation. Considerable variation noted in body Male genitalia. Unknown. size, also in shape of the pronotum and coarseness Female gonocoxites. As in the nominate sub- of the punctation of the elytral striae. species. Variation. Some variation noted in shape of Distribution. North-western QLD, northern NT, the pronotum and degree of the punctation of the northeastern Kimberleys, WA. elytral striae.

244 Distribution. Northern half of Cape York Peninsula, W.A., light trap, 27.3.66, J.A.Mahon (ANIC); Kakadu north QLD. N.P. 22-25/3/93 Cooinda at light/al lume Leg. L. Toledano (CTV); Kakadu N.P. Cooinda 25-26/12/96 at Collecting circumstances. Not recorded. light/al lume Leg. L.Toledano, R.Olivieri (CBM, CTV); Kakadu N.P. 22-25/3/93 Leg. Leech (CSM); 19.24S 135.58E 15 km SW of Alroy Downs H.S. 10.IV.1975 Key Clivina marginata (Putzeys) & Balderson (ANIC); 190m alt. 70km SW of Mataranka Figs 30, 64, 95 15°19'S 132°50'E 14-16.I.2009 St Jakl (CJP); Stuart Hwy c.25 km N Elliott, 274m S17°23'59.9",E 133°27'12.2" Scolyptus marginatus Putzeys, 1868: 8. – Sloane 1896a: 228, 20.iv.2011, Michael Langer (CBM); 10 km S of Banka 246; Csiki 1927: 493; Moore et al. 1987: 72; Lorenz Banka 18°52'S, 134°04'E, 318 m, 20.12.2008, Sv. Bilý 2005: 143. leg. (NMPC); Victoria Hwy Lily Creek, 150 km E Ku- nunurra, 50 m, 12.8. 1999 Hendrich leg, coll. Loc. 3/103 Examined types. Lectotype (by present designation): (CBM); 12.31S 132.54E 9km N by E of Mudginberry H.S., M, Rockhampton / Rockhampton (Australia) Coll. 28.v.1973 T.Weir & N.Forrester / 11249 (NTD). – WA: Castelnau / Scolyptus marginatus Putz. M (red) / SYNTY- Wyndham 20.iv.1975 K. & E. Carnaby (ANIC); Fitzroy PUS Scolyptus marginatus Putzeys, 1868 (red) (MCSN). River, at light 16 March 1980 K. & E. Carnaby (ANIC); – Paralectotypes: 1 W, Rockh (red) / Rockhampton Fitzroy River 16 Apr 1976 Ex. Carnaby Coll. (ANIC); (Australia) Coll. Castelnau / Scolyptus marginatus P. Fitzroy Crossing 125:35E, 18:10S 17 Mar. 1984, at light K. / SYNTYPUS Scolyptus marginatus Putzeys, 1868 (red) & E. Carnaby (ANIC); Willare Bridge on Fitzroy River. (MCSN); 1 M, Rockh (red) / Scolyptus marginatus P. / 20 Feb. 1985, at light K. & E. Carnaby (ANIC); S15.64975 Soc. Ent. Belg. Coll. Putzeys / Type (red) / Syntype E128.71519 light trap in plowed field FWI. Kununurra, (red) (IRSNB). 25.iii.2005 C.Y.Norwood / Agriculture (Dept.) Western Type locality. “Rockhampton”, Queensland. Australia 122168 (WADAK). – AUS: B. Mus. Godeffroy, No. 8279 (ANIC); Agriculture (Dept.) Western Australia Other material (65 ex.). QLD: Rockhampton PJD Q. 117923 (WADAK); Wyandotte 17- 9- 46 / S.R.E. Brock (ANIC); Rockhampton. Q 20.2.43 E.Vallis (?) E.Sutton Collection (ANIC). (QMB); Rockhampton Mar’58, Q.Darlingtons / marginata Pz. det.’69 Darlington topotype / Clivina marginata Diagnosis. Medium sized to fairly large, black Putz. (MCZ); Mackay 2. 99 / Clivina marginata Putz. species with wide pale margin of the elytra, shal- (ANIC); Mackay 1. 00 / C. French’s Coll. 5.11.08 / Cliv- low, quadrangular clypeal excision, glabrous, not ina marginata Putz. Queensland / COL-13774 (NMV); sulcate head, 7-setose labrum, narrow and elongate Townsville / B.Malkin 1.1945 / Clivina sp. (long tarsi) mandible, yellow legs, short, trapezoidal prothorax det. 48 Darlington (USNM); 17°54'28"S/141°08'45"E, with convexly oblique lateral margin, impunctate Rd. to Broadwater @ Blackbull Ck. 7m. 31.xii.2007, K. proepisternum, oviform elytra, deep, finely punctate Will AUS2007.xii.31, 5 1866 (EMEB); Iron Range Cape median elytral striae, and symmetric, knobbed apex York Jan.’58 Darlingtons (ANIC, MCZ); 7851 N.Qd. of the aedeagus. / 493 Clivina marginatus Putz.Townsville Q / I.6864 Clivina marginata Putz Queensland (SAMA); 473 / 8279 Clivina marginata Putz. Townsville. Q (SAMA); margi- Description nata Putz. immature / Townsville Feb. 09 F. P. Dodd Measurements. Length: 10.2-14.5 mm; width: 3.3- Clivina marginata / 6629 Ptz (immature) Queensland 4.6 mm. Ratios. Length/width of pronotum: 0.89- (SAMA); v.Townsville, Q. Mar.58 Darlingtons (MCZ); 0.97; base/apex of pronotum: 1.48-1.58; width Pt. Denison / Clivina marginata. Putz. Port Denison (MMS); Dawson R. / Scolyptus marginatus. Putz. Daw- pronotum/head: 1.43-1.47; length/width of elytra: son River (MMS); Cooktown / C. marginata Putz. det. 1.82-1.88; length/width of protibia: M: 5.3-5.6, W: 4.1- K. Kult 1948 (CDW); 65 km SW Mt.Garnet 18 Mar 1974 4.3; length/width of metatibia: M; 8.8-9.0, W: 7.4-7.7. A & M. Walford-Huggins / Walford-Huggins Col- Colour (Fig. 95). Black, but lateral margin of the lection Carnegie Museum / Clivina obscuripes (Black.) elytra, from 6th interval, yellow to red. Legs yellow [Series det. by A.Walford-Huggins] / Clivina marginata to pale red. Putz. det.M.Baehr’95 (CMP); Walker Ck., 25 ml. S.E. Head (Fig. 95). Eye large, laterally well project- of Karumba, 28.v.1972 G.B. & S.R.Monteith / UQIC ed, orbit short, oblique-convex. Clypeus with wide Reg.’90390 (QMB); Cairns NQ. 1/59. GB. / Clivina and rather shallow, almost quadrangular excision, ’ marginata Putz. det. B. P. Moore 69. / marginata Putz. though the lateral margins of the excision markedly 3170 / J.G. Brooks Bequest, 1976 (ANIC); 14 Mile Ck., oblique. Labrum 7-setose. Mandibles narrow and N. of Normanton, N.Qld. 15 Mar. 1985 at light K. & E. Carnaby (ANIC). – NT: Coll. G. Lecourt 01/00 elongate, regularly curved, with acute tip. Clypeus Katherine (CBP, CKZ); Katherine XII.57 leg.H.Demarz not divided from frons; upper surface rather convex, / Clivina marginata Putz. det.M.Baehr’95 (CBM, ZSM); rather densely, finely punctate, with very fine, rather m 50 Victoria River Road House 20-21/24-25.III.1996 superficial microreticulation, moderately glossy. P.M. Giachino leg. (CGT); 100ml. E.of Kununurra,

245 Pronotum (Fig. 95). Short and wide, markedly at lateral surface and a smaller seta below, two trapezoidal, considerably narrowed apicad, dorsally elongate setae on the ventral surface of gonocoxite rather convex. Lateral margin oblique and slightly 1, three setae at middle of the medio-dorsal surface, convex. Apex slightly concave. Marginal channel and a tiny nematiform seta near apex. Lateral plate narrow; median line slightly impressed, impunctate. basally with one elongate seta, apically with one to Basal sulcus narrow. Anterior marginal seta situ- three shorter seta. ated about at apical fourth. Basal groove distinct, Variation. Reasonable variation is noted in body linear, impunctate. Disk with some fine, irregularly size, shape of pronotum, and pilosity of parameres transverse strioles in middle, apparently impunctate, which may bear two setae at apex, or one at apex and with fine but distinct, isodiametric microreticulation, another removed from the apex at the lower surface. moderately glossy. One male from Rockhampton has a rather aber- Elytra (Fig. 95). Moderately elongate, wide at rant apex of the aedeagus: it is straight but not bent base, oval, widest in middle, convex but on disk down as usual and as in the holotype from Rock- depressed. Basal angle little produced. Striation hampton, and it is slightly sinuate when seen from incomplete, 1st-6th striae in basal half distinct, 7th laterally. This aedeagus is regarded an individual stria not or barely visible, from second third on outer variation. striae increasingly faded, therefore the apical third laterally estriate. Striae near base distinctly punctate, Distribution. North-eastern and northern QLD, the four or five median striae punctate at least in basal northern NT, Kimberleys in northern WA. half. Intervals slightly convex, impunctate, without Collecting circumstances. Several specimens were transverse strioles, with fine, distinct, isodiametric collected at light, usually near rivers, larger creeks, microreticulation, moderately glossy. Epipleura nar- and lagoons. row, basal sulcus more or less impressed. Metathoracic wings. Fully developed. Lower surface. Proepisternum impunctate, with oblonga subgroup some very fine dorso-ventral strioles, in lower part also with faint, short, transverse strioles; with faint Diagnosis. Medium sized to large species with isodiametric microreticulation. The whole lower slightly conical prothorax and moderately oval surface with fine, isodiametric microreticulation. elytra; clypeal excision wide and shallow; clypeus Metepisternum rather short, c. 1.5 × as long as wide. 7-setose; orbit large, oblique, eye comparatively little Abdominal sterna impunctate. protruded; apex of aedeagus not arrow- or club- Legs. Protibia very narrow and elongate, less shaped: metathoracal wings reduced, elytra soldered so in females. Longitudinal sulcus on upper surface together. Two species distributed in south-eastern absent; lower surface with a row of 4-5 stout setae QLD and eastern NSW. This group seems to be in- along basal part of inner margin. Mesotibia elongate, termediary between the procera and elegans groups. dorsal rim barely crenulate, with two rows of rather few setae on the dorsal surface; subapical spur rather short. Metatibia very elongate, rather sparsely setose. Clivina oblonga (Putzeys) Metatarsus extremely elongate, metatarsomere 1 Figs 31, 65, 96 shorter than the following three tarsomeres. Male genitalia (Fig. 30). Aedeagus moderately Scolyptus oblongus Putzeys, 1873: 10. – Blackburn 1890: wide, elongate, regularly narrowed apicad; lower 1249; Sloane 1896a: 227, 233, 247; 1905a: 731; Csiki surface in basal third concave, in apical part straight, 1927: 502; Moore et al. 1987: 73; Lorenz 2005: 143. towards apex curved down. Apical part almost sym- Scolyptus abbreviatus Putzeys, 1873: 11. – Blackburn 1890: metrically triagonal; apex short, straight, fairly wide, 1249; Sloane 1896a: 227, 231, 235, 238; 1905a: 733; knobbed to obtusely triangularly arrow-shaped, tip 1916: 606; Csiki 1927: 496; Moore et al. 1987: 73; obtusely triangular. Lower surface in apical half Lorenz 2005: 143. concave. Both parameres very elongate, the left one Examined types. Lectotype of oblonga (by present considerably stouter than the right one; both with designation): M, Scol. n. sp. Sidney (Dhn) / oblongus. P. elongate, very narrow, bisetose or even trisetose / Type (red) / Soc. Ent. Belg. Coll. Putzeys / Syntype apical part; setae comparatively elongate, situated (red) / det Putzeys Scolyptus oblongus Putz. (IRSNB). right at tip or one seta situated on the lower surface – Paralectotype: 1 M (probable), same data (IRSNB). – slightly removed from tip. Lectotype of abbreviata (by present designation): M, 43 Female gonocoxites (Fig. 64). Gonocoxites nar- / C. Dhn Queensland Wide Bay. / abbreviatus. P. / Type row and elongate, with slightly obtuse apex; with (red) / Soc. Ent. Belg. Coll. Putzeys / Syntype (red) / one more or less elongate, rather stout seta basally det Putzeys Scolyptus abbreviatus Putz. (IRSNB).

246 Type localities. Of oblonga: “Sydney”, New South Elytra (Fig. 96). Soldered together. Rather elon- Wales. – Of abbreviata: “Wide Bay”, South Queensland. gate, rather narrow at base, oval shaped, convex Other material (17 ex.). QLD: Noosa Headlnd 8.ii. 59 but on disk slightly depressed. Basal angle little th L.S.Dillon / Clivina abbreviata Putz. det B.P.Moore’69 produced. Striation complete, but 7 stria weak. / Clivina onlonga Putz. comp. w. types B.P.Moore’84 Striae deep, apicad barely shallower, impunctate. (ANIC); Noosa Nat. Park 2.9.1956 J. Balderson (ANIC); Intervals slightly convex, impunctate, without Wide Bay / 4268 Clivina abbreviata Putz. Queensland transverse strioles, with fine, distinct, isodiametric (SAMA); abbreviata Putz.! / Wide Bai Clivina / Com- microreticulation, moderately glossy. Epipleura pared with T Y P E K. Kult 1946 / Collectio Karel Kult narrow, basal sulcus shallow, irregularly punctate. (CDW); Wide Bay / K12756 / C. oblonga (AMS); Wide Metathoracic wings. Reduced. Bay / Coll. Castelnau / Scolyptus oblongus Ptz. t. Putzeys Lower surface. Proepisternum impunctate, with 1913 (MCSM); K12756 / Clivina abbreviata Putz. Id. by T. some fine dorso-ventral strioles, in lower part also G. Sloane (AMS); Wide Bay / Coll. Castelnau (MCSN); with fine, short, transverse strioles; with faint isodia- Wide Bay / Coll. Castelnau / Comte a Putz. e (unreadable) (MCSN); 93595 / Masters / Nov. Holl. Queensld metric microreticulation. The whole lower surface (NHM); Queens-land Wide Bay (ZMHB); Bu…(unread- with fine, isodiametric microreticulation. Metepis- able) ? River Spencer / S. abbreviatus Putz. (unreadable) ternum short, c. 1.25 × as long as wide. Abdominal 27/7/92 (ANIC). – AUS: abbreviata Sl. / Collectio Karel sterna impunctate. Kult (CDW); Collectio Karel Kult (CBM). Legs. Protibia narrow and elongate. Longitu- dinal sulcus on upper surface deep; lower surface Diagnosis. Medium sized to fairly large, black spe- with a row of 6 stout setae along basal part of inner cies with wide and shallow, quadrangular clypeal margin. Mesotibia rather short, dorsal rim crenu- excision, barely sulcate head, 7-setose labrum, rufo- late, with two rows of several setae on the dorsal piceous legs, prothorax with slightly oblique lateral surface; subapical spur very elongate. Metatibia margin, impunctate proepisternum, fairly elongate, elongate, rather sparsely setose. Metatarsus elongate, oviform elytra, deep, impunctate elytral striae, and metatarsomere 1 slightly shorter than the following spatulate apex of the aedeagus. three tarsomeres. Male genitalia (Fig. 31). Aedeagus rather wide, Description fairly elongate, slightly asymmetrically narrowed Measurements. Length: 10.7-14.1 mm; width: 3.1- apicad; lower surface in basal fourth concave, in 4.05 mm. Ratios. Length/width of pronotum: 0.99- middle straight, apex slightly curved down. The 1.01; base/apex of pronotum: 1.31-1.35; width very apex rather wide, slightly spatulate, straight, pronotum/head: 1.40-1.45; length/width of elytra: tip obtusely transverse. Lateral margins of lower 1.91-1.96; length/width of protibia: M: 3.1-3.3, W: 3.1- surface gently ridge-like, surface between slightly 3.2; length/width of metatibia: M: 6.0-6.1, W: 5.6-5.8. concave. Both parameres very elongate, the left one Colour (Fig. 96). Unicolourous black. Legs rufo- considerably stouter than the right one; both with piceous. elongate, very narrow, uni- or bisetose apical part; Head (Fig. 96). Eye fairly large, laterally well setae moderately short, situated right at tip, or, if a 1 projected, orbit rather elongate, c. /3 of length of eye, second seta is present, this is situated on the lower regularly oblique. Clypeus with wide and very shal- surface slightly removed from the apex. low, almost quadrangular excision, though the lateral Female gonocoxites (Fig. 65). Gonocoxites fairly margins of the excision oblique. Labrum 7-setose. narrow but rather short, little curved, with obtusely Mandibles short. Clypeus divided from frons by a transverse apex; with one elongate, rather stout seta shallow sulcus; upper surface rather convex, sparsely basally at lateral surface and a smaller seta below, punctate. Surface without microreticulation, glossy. two elongate setae on the ventral surface of gono- Pronotum (Fig. 96). Moderately elongate, wide coxite 1, three setae at middle of the medio-dorsal at apex, slightly narrowed apicad, dorsally gently surface, without a nematiform seta near apex. Lateral convex. Lateral margin slightly oblique but almost plate basally with one elongate seta, apically with straight. Apex gently concave. Marginal channel three to four shorter seta. narrow; median line well impressed, impunctate. Variation. Some variation noted in body size Basal sulcus narrow. Anterior marginal seta situ- and in shape of the pronotum. ated about at apical fourth. Basal groove shallow, elongate, linear, impunctate. Disk with some fine, Distribution. Eastern NSW, south-eastern QLD. Of irregularly transverse strioles in middle, sparsely this species, only old records are available. and very finely punctate, with very fine but distinct, Collecting circumstances. Not recorded. isodiametric microreticulation, moderately glossy.

247 Clivina robusta Sloane Coleoptera-Australia; Walker (OUM); Gosford Carter Figs 32, 66, 130 / Clivina robusta Sl. (AMS K 149370); Gosford Carter / elegans Putz. det. K. Kult (NHM); Gosford Carter Clivina robusta Sloane, 1905a: 731. – Csiki 1927: 510; Moore (WADAP 5194); Clivina elegans Putz. v. robusta Sl. (Gos- et al. 1987: 76; Lorenz 2005: 144. ford) / Pres. by HW Cox Sydney 14.2.18 / COL-5283 Ceratoglossa foveiceps Macleay, 1863 (non Clivina foveiceps (NMV); Gosford Carter / Clivina robusta Sl / Collectio Putzeys, 1846). – Putzeys 1866: 33; Blackburn 1890: Karel Kult (CDW); Gosford N.S.W. H. W. Cox / Pres. 1249; Sloane 1896a: 233, 235; 1905a: 731; Csiki 1927: by HW Cox Sydney 14.2.18 / COL-5278 (NMV); Austr. 502; Moore et al. 1987: 76; Lorenz 2005: 144. Gosford / elegans Putz. 47 det. K. Kult / Collectio Karel Clivina macleayana, nom. nov. for the preoccupied name Kult (CDW); Woy Woy 9.38. C.D. / F.E. Wilson Collec- C. foveiceps (Macleay. 1863). tion / COL-5275 (NMV); Woy Woy NSW. Sept. 1908 C. Deane / COL-5272 (NMV); Woy Woy 30.8.1975 / Clivina Examined types. Lectotype of robusta (by present des- procera Putz. Det. B.P.Moore 1999 (AMS K 147963); Woy ignation): M, Gosford 10.02. HJC / 571 Clivina elegans, Woy 9.08 C.D. / UQIC Reg. #90753 (QMB); Nora Head, Putz ? / HOLOTYPE Clivina robusta Sl. PJD (ANIC). Woy Woy, N.S.W. 25th November, 1984 J.Balderson – Paralectotypes: 2 MM, 1 W, Gosford Carter / Clivina (ANIC); Hunt. Riv. / C. French’s Coll. 5.11.08. / 917 robusta Sl. / PARATYPE (blue) (ANIC-MMS). – Lecto- / Clivina elegans Putz. Id. by T. G. Sloane / Det. by type of foveiceps (by present designation): W (abdomen T.G. Sloane Esq Rec 12.6.16 / COL-5279 (NMV); North destroyed), N. S. Wales / Clivina foveiceps, Macl. Rich- Sydney / H. J. Carter Coll. (NMV); K 17711 / North mond R. (ANIC-MMS). Sydney 5-04 HJC (AMS); Gosford 9.03 HJC / 511 Clivina elegans Putz. / COL-5282 (NMV); New ? (unreadable) Type localities. Of robusta: “Gosford”, New South N. Sydney G. E. Bryant 13.X.08 / elegans Putz. Exchange Wales. – Of foveiceps: “Richmond River”, New South with British Museum / Clivina elegans Putz. (MCZ); Wales. Gosford 19.02 HJC / H. J. Carter Coll. / COL-14025 Note. Two additional specimens are labelled “SYN- (NMV); Gosford N.S.W. H.W.Cox / Pres by H. W. Cox TYPE / Clivina foveiceps, Macl.” and “SYNTYPE”. 14.2.14 / COL 14088 (NMV); Gosford N.S.W. H.W.Cox However, they do not belong to C. foveifrons, i.e. / Clivina robusta Sl N. S. Wales / E. T. Smith Collection C. robusta, but to C. procera Putzeys. / COL 14416-7 (NMV); Gosford N.S.W. H.W.Cox / Clivina elegans Putz. var. obtusata Sln. / C. oblonga Putz. Other material (127 ex.). VIC: Swan Hill C. Oke / Cliv- (= robusta Sl.) det. K. Kult 1948 (DEI); Gosford N.S.W. ina oblonga Putz. Id. by T. G. Sloane / COL-5273 (NMV); H.W.Cox (DEI); N.W.Cox (Wyong) / Coll. Hacker Austral Melbour / Clivina oblonga Putz. det. K. Kult (DEI); Wyong Sydney district N.S.W. J.J.W. Aug.1903 (NHM); Australie Victoria / Cl. elegans Ptz. / C. oblonga / 6192 M Clivina oblonga Putz. / Ex misc. Coleoptera 24 Putz. det. K. Kult 1948 / Collectio Karel Kult (CDW); Coleoptera-Australia; Walker (OUM); Wyong (ANIC); AUSTRALIEN VICTORIA Cape York / Sammlung H. Gosford N.S.W. H.W.Cox / Clivina robusta Sln. / C. ob- Hesse SMNS 1995 (SMNS). – NSW: 6136 Ourimbah / longa Putz. (= robusta Sl.) det. K. Kult 1948 / Coll. Kraatz oblonga, Putz. / elegans, Putz. (SAMA); Clivina elegans (DEI); Sydney / C. elegans Putz. / E. W. Ferguson Col- Putz Ourimbah N.S.W. / I.6862 Clivina elegans Putz N. lection / Clivina elegans Putz. (ANIC); K. W. Ferguson S. Wales (SAMA); Clivina elegans Putz Ourimbah N.S.W. Collection / Ourimbah / C. elegans Putz. (ANIC); Palm / COL-18904 (NMV); Ourimbah Clivina elegans Putz. Beach HJC. 1.16 (ANIC); Toronto J.W. Filmer leg. (ZSM); Id. by T. G. Sloane / T93788 / F.E.Wilson Collection Toronto N.S.W. / Clivina elegans Putz. Id. by T. G. Sloane / COL-5276 (NMV); Ourimbah J.G.S. 27.11.04 (ANIC); (WADAP 5192); Wyee 9:9:1978 D. A. Doolan (AMS K 511 / Ourimbah J.G.S. Clivina elegans Putz. Id. by T. G. 255336-7); Wyee 9:6:1973 D. A. Doolan / D. A. Doolan Sloane (QMB); Ourimbah (SAMA 25-033406, 033610); Collection / Clivina procera Putz. Det. B.P.Moore 1999 Ourimbah N.S.W. 6-10-09 C. Gibbons. / Clivina elegans (AMS); Wyee 7:7:1973 D. A. Doolan / D. A. Doolan / Clivina procera Putz. Det. B.P.Moore 1999 (AMS K Collection / Clivina procera Putz. Det. B.P.Moore 1999 147955-6); Ourimbah 17.3.1963 / Clivina procera Putz. (AMS); Wyee 9:9:1978 D. A. Doolan / D. A. Doolan Col- Det. B.P.Moore 1999 (AMS K147954); Ourimbah Syd- lection / Clivina procera Putz. Det. B.P.Moore 1999 (AMS ney district J.J.W. Sep.1903 Ex misc. Coleoptera 24 K 147961); Newport N.Sydney G. E. Bryant 13.X.08 / G. Coleoptera-Australia; Walker (OUM); Ourimbah Syd- Bryant Coll. 1919-147 / C. elegans Putz. det. K. Kult 1948 ney district J.J.W. Oct.1903 / Ex J. J. Walker bequest 1939 / Collectio Karel KUlt (CDW); Newport N.Sydney G. (OUM); Clivina elegans Putzeys (Ourimbah) (Wadap E. Bryant 13.X.08 / G. Bryant Coll. 1919-147 / elegans 5193); Ourimbah 12.XI.1903 / W. W. Froggatt Collec- / Collectio Karel KUlt (CDW); N.S.Wales / Scolyptus tion (ANIC); Ourimbah J. Armstrong (MCZ); Gosford oblongus Putz. Burrawang (MMS); (unreadable) N. S. / Clivina elegans Putz. (Gosford) (MMS); Gosford, NSW Wales IV 31 Dr. K. K. Spence / K. K. Spence Collection UQIC Reg. #90760-1 (QMB); Gosford Carter / UQIC / Clivina sp. det. B. P. Moore 1999 (AMS); Narrabeen Reg. #90363 / Clivina procera Gosford (QMB); Gosford 8.02 (Wadap 5195); Brisbane Water NP, - 2km E of Carter / Clivina elegans Putz. (ANIC); Gosford Carter / Mt. Kariong, 33°27'50"S, 151°17'04"E, 1-15 Dec 1999 UQIC Reg. #90363 / Clivina robusta Sl. HJC det. / Clivina M.Gray, G.Millidge & H.Smith, pitfalls HS NTH SYD. -6 robusta Sloane / COL 18905 (NMV); Gosford 10.02 HJC (AMS K 266326); Brisbane Water NP, Girrakool-Patonga / 494 W Clivina oblonga Putz. / Ex misc. Coleoptera 24 Walking Track, 33°31'41"S, 151°16'55"E, 1-15 Dec 1999

248 M.Gray, G.Millidge & H.Smith, pitfalls HS NTH SYD. Description -5 (AMS K 266324); Brisbane Water NP, 2km E of Mt. Kariong, 33°27'50"S, 151°17'04"E, 1-15 Dec 1999 M.Gray, Measurements. Length: 12.5-17.2 mm; width: 3.3- G.Millidge & H.Smith, pitfalls HS NTH SYD. -6 (AMS 4.6 mm. Ratios. Length/width of pronotum: 1.02- K 266325-6); Wangi Point Lake Macquarie Nov.1976 1.08; base/apex of pronotum: 1.27-1.32; width K.R.Norris (ANIC); 33.12 S 151.36 E Geebung Camp, pronotum/head: 1.36-1.42; length/width of elytra: Lake Dunmoorah, 15 Nov. 1983 D.Rentz, M.Harvey 1.86-2.0; length/width of protibia: M: 4.0-4.4, W: 3.2- Stop 47 (ANIC); Newcastle HJC 11.33 / Clivina elegans 3.3; length/width of metatibia: M; 6.5-6.7, W: 5.5-5.7. Putz. (ANIC); Newcastle HJC 11.33 / Clivina robusta Colour (Fig. 130). Unicolourous black. Legs dark Sl. (ANIC); Morisset N.S.W. J. Sedlacek / COL 14016 piceous to black. (NMV); Morisset N.S.W. J. Sedlacek lgt. / Clivina sp. / Head (Fig. 130). Eye comparatively small, later- Clivina ovipennis SL. det. Balkenohl X.97 (CBP); Dorrigo N.S.W. Jan. 1931 C. Oke / COL 13672-4 (NMV); Coffs ally moderately projected, orbit elongate, almost as Harbour N.S.W H. W. Cox / Pres. by HW Cox Sydney long as the eye, regularly oblique. Clypeus with wide 14.2.18 / COL-5281 (NMV); Newcastle IX.1957 / Clivina and shallow, almost quadrangular excision, though sp. (USNM); Woy-Woy 9.08 CD. / Clivina robusta Sl. the lateral margins of the excision oblique. Labrum Woy Woy N.S.W. (QMB); Location. Nords Wharf Al- 7-setose. Mandibles short. Clypeus divided from titude 6’ Date April 68 Collector. G. Purnell / ANIC frons by a fairly deep sulcus; upper surface rather Univ. of New England Coll. Donated 1983 (ANIC); convex, not sulcate, rather densely but finely punc- Clivina procera NSW / UQIC Reg. # 90751 (QMB); New tate. Neck with a distinct transverse sulcus. Surface South Wales UQIC Reg. # 90757 (QMB); N. S. Wales / with or without traces of isodiametric microreticula- W. W. Ferguson Collection (ANIC); N. S. Wales / W. W. tion, rather glossy. Ferguson Collection / Clivina procera Putz. N. S. Wales Pronotum (Fig. 130). Moderately elongate, (ANIC); N.S.Wales / Coll. Franklin Müller / Clivina procera / det. Scolyptus procerus (DEI). – AUS: H. J. Carter slightly narrowed apicad, dorsally rather depressed. Coll. / COL-5274 (NMV); H. J. Carter Coll. / Scolyptus Lateral margin oblique but little convex. Apex gen- oblongus (Putz) / COL-5271 (NMV); Morisset (?) Oct- tly concave. Marginal channel narrow; median line Mar 37 E.Stoyles / E. Sutton Collection (QMB); Nara… well impressed, impunctate. Basal sulcus narrow. (unreadable) 20/10/89 (ANIC); K 17711 (AMS); Na 7/96 Anterior marginal seta situated slightly in front of / C. oblonga Putz. / Ex misc. Coleoptera 24 Coleoptera- apical fourth. Basal groove very shallow or barely Australia; Walker (OUM); N.W.Austr. / Clivina oblonga perceptible, elongate, linear, impunctate. Disk with Putz. (ANIC-MMS); 193 / C. oblonga Putz. = C. foveiceps some fine, irregularly transverse strioles in middle, Macl. comp. with type / Clivina elegans Putz. Id. by T. G. sparsely, finely punctate, with very fine but distinct, Sloane (ANIC); Clivina obscuripes / COL 5398 (NMV); N. isodiametric microreticulation, moderately glossy. S. Wales / 2238 / biplagiata / Clivina elegans var. obtusata Elytra. Soldered together. Rather elongate, rath- N.S.Wales / COL 5277 (NMV); 2232 / Bowring 63-47* er narrow at base, oval shaped, convex but on disk (NHM); Clivina oblonga Putz. det. K. Kult / Collectio Karel Kult (CDW). slightly depressed. Basal angle little produced. Stria- tion complete, striae deep, apicad barely shallower, Notes. Sloane (1905a) misidentified Clivina elegans impunctate to finely punctate in basal half. Intervals Putzeys, 1862, and synonymized Clivina oblonga convex, impunctate, without transverse strioles, (Putzeys, 1873) and Clivina foveiceps (Macleay, 1863) with fine, distinct, isodiametric microreticulation, under that name. moderately glossy. Epipleura narrow, basal sulcus Two of the three syntypes of C. foveiceps do not shallow, irregularly punctate. belong to that species but to C. procera Putzeys. Metathoracic wings. Reduced. Clivina foveiceps (Macleay, 1863) is a junior homo- Lower surface. Proepisternum impunctate, with nym of C. foveiceps Putzeys, 1846 and therefore is fine dorso-ventral strioles, in lower part also with herein renamed Clivina macleayana, nom. nov. fine, short, transverse strioles; with faint isodiametric microreticulation. The whole lower surface with fine, Diagnosis. Medium sized to rather large, black isodiametric microreticulation. Metepisternum short species with wide and shallow, quadrangular cly and very small, little longer than wide. Abdominal peal excision, barely sulcate head, 7-setose labrum, sterna impunctate. black legs, apicad narrowed, depressed prothorax Legs. Protibia narrow and elongate, less so in with oblique-convex lateral margin, impunctate females. Longitudinal sulcus on upper weak or proepisternum, fairly elongate, oviform elytra, deep, absent; lower surface with a row of 5-6 stout setae impunctate elytral striae, and narrow, remarkably along basal part of inner margin. Mesotibia rather sinuate apex of the aedeagus. short, dorsal rim crenulate, with two rows of several setae on the dorsal surface; subapical spur very elongate. Metatibia elongate, rather sparsely setose.

249 Metatarsus rather elongate, metatarsomere 1 slightly Note. Moore et al. (1987) noted the above specimen shorter than the following three tarsomeres. as the holotype. Actually it is a syntype, because in Male genitalia (Fig. 32). Aedeagus rather wide, the description (Sloane 1905, p. 733) an additional elongate, in apical third remarkably sinuate; lower female is explicitly noted. But apparently this speci- surface gently concave, but near apex slightly bis- men is lost. inuate. Apical part narrow, sinuate, tip curved left, Other material (4 ex.). NT: 1 M, Pine Creek District slightly obtuse. Both parameres rather elongate, large N. Territory / procera (MCZ). – AUS: 1 M (defect), ex. and stout, the left one considerably stouter than the J. J. Walker bequest 1939 / Ex Misc. Coleoptera 21 right one; both with fairly elongate, narrow, asetose Coleoptera-Australia; Walker (OUM); 2 MM (defect) apical part. (ANIC, CBM). Female gonocoxites (Fig. 66). Gonocoxites narrow and elongate, with slightly obtuse apex; with one Diagnosis. Medium sized, black species with wide elongate, rather stout seta basally at lateral surface and shallow, quadrangular clypeal excision, slightly and a smaller seta below, two elongate setae on the sulcate, moderately densely punctate head, rufous ventral surface of gonocoxite 1 and one additional middle and hind legs, rather conical prothorax, seta in middle of gonocoxite 2, four setae at middle of impunctate and very glabrous proepisternum, elon- the medio-dorsal surface, and usually a tiny nemati- gate but not absolutely parallel-sided elytra, finely form seta near apex in some specimens, however, punctate elytral striae, and symmetric, arrow-shaped only the pit is present. Lateral plate basally with one apex of the aedeagus. elongate seta, apically with three to four shorter seta. Variation. Considerable variation noted in body Description size, also in relative length of elytra, shape of pro- Measurements. Length: 11.4-12.5 mm; width: 3.35- thorax, and degree of elytral striation. 3.7 mm. Ratios. Length/width of pronotum: 1.0-1.02; base/apex of pronotum: 1.28-1.34; width pronotum/ Distribution. Eastern VIC, eastern NSW. The re- head: 1.39-1.45; length/width of elytra: 1.95-1.97; cords from Cape York and “NW.Austr” certainly length/width of protibia: M: 3.9-4.1; length/width are erroneous. of metatibia: M; 7.4-7.5. Collecting circumstances. Little recorded. Single Colour (Fig. 97). Unicolourous black. Middle specimens were sampled at light and in pitfall traps. and hind legs rufous, paler than the piceous anterior leg. Head (Fig. 97). Eye large, laterally well pro- obliquicollis subgroup jected, orbit short, oblique-convex. Clypeus with rather shallow, wide, almost quadrangular excision, Diagnosis. Medium sized with moderately conical, though the lateral margins of the excision being laterally slightly convex prothorax but elongate, rather oblique. Labrum 7-setose. Clypeus divided almost parallel-sided elytra; clypeal excision rather from frons by a deep sulcus; frons anteriorly with wide and shallow; clypeus 7-setose; eye large, pro- some faint transverse sulci, moderately densely but truded; apex of aedeagus arrow- or club-shaped: inconspicuously punctate. Surface without microre- metathoracal wings complete. ticulation, glossy. The fife species occur in northern and central Pronotum (Fig. 97). Moderately elongate, nar- Australia. In the shape of the elytra this group rowed apicad, dorsally gently convex. Lateral mar- matches the procera group. gin straight to slightly convex, and oblique, faintly concave in apical third. Apex slightly concave. Mar- ginal channel narrow; median line well impressed, Clivina obliquicollis Sloane impunctate. Basal sulcus narrow. Anterior marginal Figs 33, 97 seta situated at apical fifth. Basal groove shallow, elongate, linear, straight, impunctate. Disk with some Clivina obliquicollis Sloane, 1905a: 732. – Sloane 1916: 606; fine, transverse strioles in middle, extremely finely Csiki 1927: 508; Moore et al. 1987: 73; Lorenz 2005: punctate, with very fine, superficial, isodiametric 143. microreticulation, glossy. Examined types. Lectotype (by present designation): Elytra (Fig. 97). Moderately elongate, not ab- M, Clivina obliquicollis, Sl. Carnot Bay N.W.A. / HOLO- solutely parallel-sided, slightly widened apicad, TYPE Clivina obliquicollis Sl PJD (ANIC). convex but on disk depressed. Basal angle little produced. Striation complete, but lateral striae less Type locality. “Carnot Bay”, northwestern Australia. distinct. Striae deep, becoming slightly shallower

250 towards apex, punctate at least in basal two thirds. lgt. (CBP); 1 W, AUSTRALIA NT 10 km S of Banka Intervals slightly convex, finely and sparsely punc- Banka. 18°52'S, 132°04'E, 316 m, 12-13.1.2009, Sv. Bilý tate, without or with weak transverse strioles, with leg. (CBM); 1 W, Ti-Tree N.T. 30/12/1982 D.P.Carne At very faint, superficial, isodiametric microreticulation, Light (ANIC); 1 W, 18.59S 134.12 E NT South Attack Ck glossy. Epipleura narrow, basal sulcus indistinct, 16 Mar. 1983 I.Archibald & D.Percival (NTD); 1 W, NT, 23km S of Tennant Creek 24 Feb. 1982 G.R. Brown at shallowly punctate. head light (NTD); 1 W, NT Three Ways N of Tennant Ck. Metathoracic wings. Fully developed. 17 March 1983 I.Archibald & D.Percival (CBM). Lower surface. Proepisternum impunctate, and almost without strioles, remarkably glabrous, with Etymology. The name refers to the locality of the holo- faint isodiametric microreticulation. The whole lower type, Banka Banka in central NT. surface with fine, isodiametric microreticulation. Diagnosis. Rather large to large, black species Metepisternum moderately elongate, c. 2 × as long with rather shallow, quadrangular clypeal exci- as wide, in posterior half with a deep longitudinal sion, sulcate, densely punctate head, red femora sulcus. Abdominal sterna impunctate. on all legs, convexly oblique lateral pronotal mar- Legs. Protibia very narrow and elongate, in the gin, impunctate proepisternum, elongate, almost male more than in the female. Longitudinal sulcus parallel-sided elytra, finely punctate elytral striae, on upper surface shallow; lower surface with a row and asymmetric apical part of the aedeagus with of 6-8 stout setae along basal part of inner margin. large, club-shaped tip. Mesotibia moderately elongate, dorsal rim crenulate, with several rows of setae on the dorsal surface; Description subapical spur elongate. Metatibia elongate, rather sparsely setose. Metatarsomere 1 almost as long as Measurements. Length: 11.6-14.9 mm; width: 3.3- the following three tarsomeres. 4.5 mm. Ratios. Length/width of pronotum: 0.93-1.0; Male genitalia (Fig. 33). Aedeagus rather wide, base/apex of pronotum: 1.37-1.43; width pronotum/ moderately elongate, asymmetrically narrowed head: 1.40-1.50; length/width of elytra: 2.06-2.11; apicad; lower only near base concave, then straight, length/width of protibia: M: 3.0-3.1, W: 2.9-3.0; towards apex slightly curved down. Apical half of length/width of metatibia: M; 6.8-6.9, W: 6.5-6.9. lower surface slightly concave. Apical part very Colour (Fig. 131). Unicolourous black. Femora asymmetric, deeply concave at the left side; apex of all legs rufous, tibiae darker. wide, arrow shaped, and denticulate on the lower Head (Fig. 131). Eye large, laterally markedly surface; arrow symmetrically situated, tip obtusely projected, orbit very short, oblique. Clypeus with triangular. Both parameres elongate, the left one wide and rather shallow, almost quadrangular exci- stouter than the right one; both with short, wide, sion. Labrum 7-setose. Clypeus divided from frons by uni- or bisetose apical part; setae rather elongate, a deep sulcus; upper surface with several transverse situated right at tip; but left paramere with an ad- sulci, densely and rather coarsely punctate. Surface ditional seta at the lower surface near apex. without microreticulation, glossy. Female gonocoxites. Unknown. Pronotum (Fig. 131). Rather short, considerably Variation. Little variation noted in the punctation narrowed apicad, dorsally gently convex. Lateral of the elytral striae. margin oblique and convex. Apex slightly concave. Marginal channel narrow; median line well im- Distribution. Largely unrecorded, the holotype pressed, impunctate. Basal sulcus narrow. Anterior collected in north-western WA. marginal seta situated slightly behind apical fifth. Ba- sal groove shallow, elongate, linear, straight, impunc- Collecting circumstances. Not recorded. tate. Disk with some fine, transverse strioles in middle, more or less densely punctate, with very faint, superficial, isodiametric microreticulation, glossy. Clivina bankae, spec. nov. Elytra. Elongate, almost parallel-sided, barely Figs 34, 67, 131 widened apicad, convex but on disk depressed. Basal angle little produced. Striation complete, but lateral Examined types. Holotype: M, AUSTRALIA NT 318m striae less distinct. Striae deep, becoming slightly alt. 14°53'S (wrong longitude, must be 18°53'S!), 132°01'E, shallower towards apex, very finely punctate at least Banka Banka env., road to Tennant Creek, 12-14.i.2009, in basal half. Intervals slightly convex, very sparsely St. Jakl lgt. (NTD). – Paratypes: 3 MM, 10 WW, same data (CBM, CBP, CJP); 1 M, AUSTRALIA NT; 190m alt. 70km punctate, without or with very weak transverse SW of Mataranka 15°19'S 132°50'E 14-16.I.2009 St Jakl strioles, with very faint, superficial, isodiametric (CBM); 2 MM, AUSTRALIA NT; 190m alt. 70km SW of microreticulation, glossy. Epipleura narrow, basal Mataranka 15°19'S 132°50'E 22-23.XII.2008 L.Hovorka sulcus fairly distinct, punctate.

251 Metathoracic wings. Fully developed. 21°41'21"S, 115°50'35"E 14.5.-29.8.2006 CALM PBS01232 Lower surface. Proepisternum impunctate, with (WAM); 1 M, Australia,WA,OYW02 29.5km ESE Peeda- some dorso-ventral and more or less distinct, short mulla Hs 21°57'00"S, 115°53'15"E 28.9.2005-13.5.2006 transversal strioles, with faint isodiametric microre- CALM PBS01244 (WAM); 1 M, Australia,WA,MBW07 ticulation. The whole lower surface with fine, isodia- 15km NE Wodjina 21°03'49"S, 118°45'39"E 23.9.2005-15. 5.2006 CALM PBS01241 (WAM); 2 WW, Australia,WA, metric microreticulation. Metepisternum elongate, MBW10 13km SSE Wodjina 21°16'47"S, 118°41'56"E c. 2.25 × as long as wide, in posterior half with a deep 23.9.2005-15.5.2006 CALM PBS01243 (WAM); 1 W, longitudinal sulcus. Abdominal sterna impunctate. Australia,WA,MBE08 4 km SE The Island Hill 21°20'25"S, Legs. Protibia narrow and elongate, in the male 119°21'36"E 12.10.2005-7.5.2006 CALM PBS01240 slightly more than in the female. Longitudinal sulcus (WAM); 1 W, Australia,WA,BORS13 20 km ESE Wheelar- on upper surface distinct; lower surface with a row ra Hill 23°24'46"S, 120°19'00"E 4.9.2005-23.5.2006 CALM of 6-8 stout setae along basal part of inner margin. PBS01239 (WAM); 1 W, Australia,WA,TCMBE06 9.5 km Mesotibia moderately elongate, dorsal rim crenulate, ESE Mt.Bruce 22°38'10"S, 118°13'32"E 1.9.2005-28.5.2006 with several rows of setae on the dorsal surface; CALM PBS01253 (WAM); 1 M, Australia,WA,OYW02 subapical spur elongate. Metatibia elongate, rather 29.5km ESE Peedamulla Hs 21°57'00"S,115°53'15"E sparsely setose. Metatarsomere 1 almost as long as 28.9.2005-13.5.2006 CALM PBS01244 (WAM); 1 W, Australia,WA, DRC08. s. Roeburne, 20.51.14.5S 118.05. the following three tarsomeres. 40.2E 12.11.2003-1.5.2004 N.Gutthrie (WAM); Male genitalia (Fig. 34). Aedeagus rather wide, 2 WW, Australia,WA,PW05. s. Fortescue, 21.03.18.8S fairly elongate, narrowed apicad; lower surface gen- 116.07.56.6E 23.11.2003-8.5.2004 N.Gutthrie (WAM); tly concave. Apical part asymmetrically narrowed; 2 MM, Fortescue R. Hamersley Range N. W. A.: W. D. apex short, markedly club-shaped and hooked at Dodd / Clivina obliquicollis Sl. mss. Id. by T. G. Sloane the lower surface; at tip convex. Internal sac in api- (SAMA); 1 M, 20km NE Mt. Clere Stn. W. Aust. 25.x.1986 cal third with three large, denticulate and rather L. A. Dyson K. T. Richards (Wadap 5243); 1 W, 23 km, sclerotized folds. Both parameres very elongate, the WSW. of Barradale, WA 30 Mar.1971 Upton & Mitch- left one considerably stouter than the narrower right ell (ANIC); 1 W, 93 COLEOPTERA R2 008 NHC AL / one; both with elongate, very narrow, uni- or bisetose Barrow Island Western Australia S. Callan 17.v.2005 apical part; setae short, situated right at tip, or on the 20°47'38"S 115°26'34"E (WADAP 140255); 1 M, Skull Springs 5-I-1971 WA J. Wombey (ANIC); 1 M, WA Cape right paramere in some specimens one seta inserted Range 22°09'55"S, 113°59'25"E 25 June 1989 cave C-83 at apex, the other at the lower surface near apex. R.Wood CR # 4218 (WAM no. 39186); 1 M, WA Cape Female gonocoxites (Fig. 67). Gonocoxites very Range 22°09'S, 113°59' 28 July 1985 E.Pryor / BES: 1446 narrow and elongate, little curved, with rather acute 4m outside entrance to cave C-118 wet pitfall on surface apex; with one elongate, rather stout seta basally (WAM no. 39187); 1 W, WA Cape Range 22°01'00"S, at lateral surface and a smaller seta below, two 114°02'37"E 30 July 1989 cave C-263 R.D.Brooks CR elongate setae on the ventral surface of gonocoxite # 3443 / Clivina sp. Det. B. P. Moore 1990 (WAM no. 1, three setae at middle of the medio-dorsal surface, 39185); 1 M, Wiluna 27-391 (WAM no. 38933); 1 M, 7.5 km and a tiny nematiform seta near apex. Lateral plate SE of BANJIWARN HS (27°42'S121°37'E) W.AUST. basally with one elongate seta, apically with two to 22-28 FEB. 1980 T.F.HOUSTON ET AL 316-10 / AT three shorter seta. LIGHT AT NIGHT / Western Australian Museum , Variation. Some variation noted in body size, Dept. of Biological Survey Site BW Camp (WAM no. 39077); 1 M, Ashburton River WA. / Nat. Mus. Vic- depth of clypeal excision, and density of head toria C. French’s Coll. 6.I.08 (NMV COL-5395); 1 W, punctation. Ashburton River WA. / After careful consideration I consider only a robust spec. of C. obliquicollis Sl. 4.5.07 Distribution. Central NT. (ANIC); 1 M, 2 WW (partly defect), Asburton River, WA (ANIC); 1 M, 1 W, Murchison District, W.A. / Nat. Mus. Collecting circumstances. Most specimens were Victoria C. French’s Coll. 6.I.08 (NMV COL-5394/7); collected at light, those from Banka Banka at the 1 W, Nickol Bay N.W.Australia DuBoulay 1869 (OUM); bank of a small river. 1 W, Nulubi H’stead WA. B.Clarke / Golding/Powell Collection donated 12 Feb 2002 (WAM no. 39191); 1 W, 8 km. SWbyW of Cane R.HS., WA. 22.07S 115.33E 31 Clivina incurvicollis, spec. nov. Mar. 1971 Upton & Mitchell (ANIC); 1 W, Roebuck B Figs 35, 68, 98 M. G. (unreadable) (IRSNB); 2 MM, Fortescue R. Ham- ersley Range N. W. A.: W. D. Dodd / Clivina obliqui- Examined types. Holotype: M, Australia,WA,PHYE08 collis Sl. mss. Id. by T. G. Sloane (SAMA); 1 M, 20km 7.5km E Warrawagine Hs 20°51'49"S, 120°45'55"E NE Mt. Clere Stn. W. Aust. 25.x.1986 L. A. Dyson K. 3.7.2005-23.5.2006 CALM PBS01247 (WAM). – Paratypes: T. Richards (Wadap 5243); 1 W, 23 km, WSW. of Bar- 1 M, same data (WAM); 1 M, 3 WW, Australia,WA,OYW09 radale, WA 30 Mar.1971 Upton & Mitchell (ANIC); 6km NE Warrambo Cr. on North West Coastal Hwy. 70 MM, 107 WW, Australia,WA06,132, Kariniji NP, 8km S

252 Visitors Centre 22.55353S, 118.45290E 714m, 21.2.2006, protibia: M: 3.1-3.4 (3.8), W: 3.3-3.45; length/width M.Baehr (ANIC, CBM, CBP, CDW, MCZ, QMB); 2 MM, of metatibia: M; 6.3-6.7 (8.1), W: 6.6-7.2. 3 WW, Australia,WA06,117 Ashburton R.,5km s.mouth Colour (Fig. 98). Unicolourous black. Middle 21.76114S, 114.94970E 1m, 19.2.2006, M.Baehr (CBM); and hind legs rufous, paler than the piceous ante- 2 MM, 1 W, Australia,WA06,101, Yalgar River, 10 km rior leg. s.Moorarie, 2595233S, 117.69858E 439m, 3.2.2006, Head (Fig. 98). Eye large, laterally well pro- M.Baehr (CBM); 3 WW, Australia,WA06,155, Millie Soak, 10 km n. Cue, 424m 27.29033S, 117.91034E 1m, 25.- jected, orbit short, oblique-convex. Clypeus with 27.2.2006, M.Baehr (CBM); 2 MM, Australia,WA06,114, rather shallow, wide, almost quadrangular excision, Cane River Cr. 22.02838S, 115.58598E 79m, 19.2.2006, though the lateral margins of the excision being M.Baehr (CBM); 1 M, Australia,WA06,120, Peter rather oblique. Labrum 7-setose. Clypeus divided Cr.,15 km ne.Robe R. Cr. 21.49982S, 116.00387E 68m, from frons by a deep sulcus; frons in anterior half 19.2.2006, M.Baehr (CBM); 1 M, Australia,WA06,135, usually with additional transverse sulci, moderately Beasley R.,Cheel Plains. 22.95945S, 116.97655E 283m, densely but inconspicuously punctate. Surface with- 18.2.2006, M.Baehr (CBM); 1 M, Australia,WA06,103, out microreticulation, glossy. Gascoyne Junction 25.04722S, 115.20700E 147m, Pronotum (Fig. 98). Rather short, narrowed api- 5.2.2006, M.Baehr (CBM); 1 W, Australia,WA06,122, cad, dorsally gently convex. Lateral margin oblique, Karratha 20.75383S, 116.81295E 41m, 11.2.2006, M.Baehr slightly convex. Apex slightly concave. Marginal (CBM); 1 M, 1 W, W.A.: CALM Pilbara Survey 20km ESE Whim Creek Hotel 20°54'36.2"S,117°58'58.1"E DRE12, channel narrow; median line well impressed, im- 14.XI.2003-13.V.2004 / CALM PBS00062 (WAM); 2 MM, punctate. Basal sulcus narrow. Anterior marginal 1 W, W.A.: CALM Pilbara Survey 3.5km S Marla Pool seta situated at apical fifth. Basal groove shallow, 21°3'55.9"S, 116°9'1.7"E DRW11, 28.XI.2003-11.V.2004 elongate, linear, straight, impunctate. Disk with / CALM RBS00069 (WAM); 1 W, W.A.: CALM Pilbara some fine, transverse strioles in middle, very finely Survey 12.5km N Nullagine 21°46'13.1"S, 120°5'30.7"E punctate, with very fine, superficial, isodiametric NW03, 17.XI.2003-19.V.2004 / CALM PBS00072 microreticulation, glossy. (WAM); 1 W, W.A.: CALM Pilbara Survey 62km E Elytra (Fig. 99). Moderately elongate, not abso- Meekatharra Outcamp 21°18'57.6"S, 121°2'53.4"E NE10, lutely parallel-sided, slightly widened apicad, con- 17.V.-12.X.2004 / CALM PBS00071 (WAM); 1 W, W.A.: vex but on disk remarkably depressed. Basal angle CALM Pilbara Survey 12.5km N Nullagine 21°46'13.1"S, little produced. Striation complete, but lateral striae 120°5'30.7"E NW03, 17.XI.2003-19.V.2004 / CALM PBS00072 (WAM); 1 W, West Aust / TYLDEN COLL., less distinct. Striae deep, becoming slightly shallower formed by Rev. W. Tylden, M.A., abt 1841-1875. Pres. towards apex, rather coarsely punctate at least in ’75 by Mrs. E. C. Tylden (OUM); 1 M, 33959 / Du Boulay basal two thirds. Intervals slightly convex, extremely / Nov. Holl. Occid. (NHM); 1 M, 2 WW, Fitzroy River WA finely and sparsely punctate, without transverse 20.iv.1976 K.&E.Carnaby (ANIC); 1 M, defect (ANIC); 1 strioles, with very faint, superficial, isodiametric M, Victoria / Nat. Mus. Victoria C. French’s Coll. 6.I.08 microreticulation, glossy. Epipleura narrow, basal (NMV COL-10511). sulcus indistinct, shallowly punctate. Etymology. The name refers to the convex, apicad Metathoracic wings. Fully developed. incurved lateral margins of the prothorax. Lower surface. Proepisternum impunctate, and almost without strioles, remarkably glabrous, with Diagnosis. Medium sized to fairly large, black spe- faint isodiametric microreticulation. The whole lower cies with wide and shallow, quadrangular clypeal surface with fine, isodiametric microreticulation. excision, sulcate, moderately densely punctate head, Metepisternum moderately elongate, c. 2 × as long rufous middle and hind legs, rather conical protho- as wide, in posterior half with a deep longitudinal rax, impunctate and very glabrous proepisternum, sulcus. Abdominal sterna impunctate. elongate but not absolutely parallel-sided elytra, Legs. Protibia narrow and elongate, in the male rather coarsely punctate elytral striae, and sym- not more than in the female. Longitudinal sulcus metric, wide, arrow-shaped apex of the aedeagus. on upper surface distinct; lower surface with a row of 6-8 stout setae along basal part of inner margin. Mesotibia moderately elongate, dorsal rim crenulate, Description with several rows of setae on the dorsal surface; subapical spur elongate. Metatibia rather elongate, Measurements (those of an aberrant male specimen rather sparsely setose. Metatarsomere 1 almost as in brackets). Length: 11.1-14.5 mm; width: 3.3- long as the following three tarsomeres. 4.35 mm. Ratios. Length/width of pronotum: 0.93- Male genitalia (Fig. 35). Aedeagus wide, mod- 0.96 (0.99); base/apex of pronotum: 1.40-1.44 (1.46); erately elongate, asymmetrically narrowed apicad; width pronotum/head: 1.42-1.48 (1.51); length/ lower only near base concave, then straight, towards width of elytra: (1.92) 1.98-2.02; length/width of apex curved down. Apical half of lower surface

253 slightly concave. Apical part asymmetric, deeply Description concave at the left side; apex wide, arrow shaped Measurements. Length: 13.0-13.2 mm; width: 3.5- with wide, convexly triangular tip, denticulate on 3.75 mm. Ratios. Length/width of pronotum: 1.01- the lower surface; arrow almost symmetrically situ- 1.04; base/apex of pronotum: 1.33-1.34; width ated. Both parameres elongate, the left one stouter pronotum/head: 1.34-1.37; length/width of elytra: than the right one; both with short, bisetose apical 2.11; length/width of protibia: M: 3.4-3.6; length/ part; upper seta elongate, situated right at tip, the width of metatibia: M; 7.4-7.5. lower seta short. Colour (Fig. 132). Unicolourous piceous. Middle Female gonocoxites (Fig. 68). Gonocoxites rather and hind legs rufous, paler than the piceous anterior leg. narrow and elongate, little curved, with slightly Head (Fig. 132). Eye large, laterally remarkably obtuse apex; with one elongate, rather stout seta ba- projected, orbit very short, oblique-convex. Clypeus sally at lateral surface and a smaller seta below, two with rather shallow, wide, almost quadrangular elongate setae on the ventral surface of gonocoxite excision. Labrum in the holotyoe 7-setose, in the 1, four to five setae at middle of the medio-dorsal paratype 5-setose. Clypeus divided from frons by a surface, without a nematiform seta near apex. Lateral deep sulcus; frons in anterior half with or without plate basally with one elongate seta, apically with another shallow transverse sulcus, with moderately three to four shorter seta. dense, rather coarse punctures. Surface without Variation. This is a rather variable species with microreticulation, glossy. respect to shape of pronotum and elytra, surface Pronotum (Fig. 132). Moderately elongate, rath- structure and microreticulartion, and relative length er quadrate, little narrowed apicad, dorsally remark- of protibia and metatibia. A large male from Mur- ably depressed. Lateral margin straight, slightly chison District has particularly narrow and elongate oblique. Apex slightly concave. Marginal channel pro- and metatibiae, which by far surpass the aver- narrow; median line well impressed, impunctate. age range within the species. However, because the Basal sulcus narrow. Anterior marginal seta situated male genitalia of all dissected specimens, taken from at apical sixth. Basal groove shallow, elongate, linear, several disjunct localities, are similarly shaped, the straight, impunctate. Disk with some fine, transverse aberrant specimens are yet included in the species. strioles in middle, finely but rather densely punctate, with finest traces of very superficial, isodiametric Distribution. North-western WA: from about Mee microreticulation, very glossy. katharra to Fitzroy Crossing in the southern Kim- Elytra. Elongate, parallel-sided, barely widened berleys. apicad, convex but on disk remarkably depressed. Collecting circumstances. Many specimens col- Basal angle little produced. Striation complete, but lected in pitfall traps near ponds and billabongs, lateral striae less distinct. Striae deep, becoming or at light. slightly shallower towards apex, very finely punctate-crenulate in basal two thirds. Intervals slightly convex, impunctate, with fine transverse strioles, Clivina platynota, spec. nov. with extremely faint and superficial, isodiametric Figs 36, 132 microreticulation, very glossy. Epipleura narrow, basal sulcus indistinct, shallowly punctate. Examined types. Holotype: M, 8 km SWbyW of Cane R. Metathoracic wings. Fully developed. HS., WA. 22.07S 115.33E 31 Mar 1971 Upton & Mitchell Lower surface. Proepisternum impunctate, and (ANIC). – Paratype: 1 M, same data (CBM). almost without strioles, remarkably glabrous, with Etymology. The name refers to the depressed surface faint isodiametric microreticulation. The whole of the prothorax. lower surface with fine, isodiametric microreticulation. Metepisternum elongate, almost 2.5 × as long Diagnosis. Medium sized, piceous species with as wide, in posterior half with a deep longitudinal wide and rather shallow, quadrangular clypeal ex- sulcus. Abdominal sterna impunctate. cision, sulcate, moderately densely punctate head, Legs. Protibia narrow and elongate. Longitudi- markedly protruded eye, rufous middle and hind nal sulcus on upper surface distinct; lower surface legs, rather quadrate, dorsally depressed prothorax, with a row of 6-8 stout setae along basal part of inner impunctate and very glabrous proepisternum, elon- margin. Mesotibia moderately elongate, dorsal rim gate, almost parallel-sided elytra, finely punctate ely- crenulate, with several rows of setae on the dorsal tral striae, and rather spatulate apex of the aedeagus. surface; subapical spur elongate. Metatibia elongate, rather sparsely setose. Metatarsomere 1 almost as long as the following three tarsomeres.

254 Male genitalia (Fig. 36). Aedeagus rather wide, with additional transverse sulci, moderately densely moderately elongate, asymmetrically narrowed punctate. Surface without microreticulation, glossy. apicad; lower only near base concave, then straight, Pronotum (Fig. 99). Rather short, convexly nar- towards apex slightly curved down. Apical half of rowed apicad, dorsally rather convex. Lateral margin lower surface slightly concave. Apical part asym- oblique, distinctly convex. Apex slightly concave. metric, deeply concave at the left side; apex wide, Marginal channel narrow; median line well im- arrow shaped with wide, somewhat spatulate tip, pressed, impunctate. Basal sulcus narrow. Anterior denticulate on the lower surface; arrow slightly marginal seta situated at apical fifth. Basal groove asymmetrically situated. Both parameres elongate, shallow, elongate, linear, straight, impunctate. Disk the left one stouter than the right one; both with short, with some fine, transverse strioles in middle, very bisetose apical part; upper seta elongate, situated finely punctate, with very fine traces of superficial, right at tip, the lower seta short. isodiametric microreticulation, glossy. Female gonocoxites. Unknown. Elytra (Fig. 99). Elongate, parallel-sided, barely Variation. Little variation noted. widened apicad, convex but on disk depressed. Basal angle little produced. Striation complete, but lateral Distribution. North-western WA, south of Great striae less distinct. Striae deep, becoming slightly Sandy Desert. Known only from type locality. shallower towards apex, finely punctate at least in basal half. Intervals slightly convex, extremely Collecting circumstances. Not recorded. finely and sparsely punctate, with or without fine transverse strioles, with very faint, superficial, isodiametric microreticulation, glossy. Epipleura Clivina rectipennis, spec. nov. narrow, basal sulcus indistinct, shallowly punctate. Figs 69, 99 Metathoracic wings. Fully developed. Examined types. Holotype: W, IRRUNYTJU ROCK- Lower surface. Proepisternum impunctate, and HOLE HINCKLEY RANGE WA 26.07 S, 128.58 E 19-21 almost without strioles, remarkably glabrous, with JAN 1990 T.F. HOUSTON & M.S. HARVEY 759-11 / AT faint isodiametric microreticulation. The whole lower LIGHT (MV) AT NIGHT (WAM No. 39180). – Paratype: surface with fine, isodiametric microreticulation. 1 W, Ayers Rock NT 23 Apr 1981 D.P.Carne At light Metepisternum elongate, c. 2.25 × as long as wide, (ANIC). in posterior half with a deep longitudinal sulcus. Abdominal sterna impunctate. Etymology. The name refers to the straight, parallel lateral margins of the elytra. Legs. Protibia narrow and elongate. Longitudi- nal sulcus on upper surface distinct; lower surface Diagnosis. Medium sized to fairly large, black spe- with a row of 6-8 stout setae along basal part of inner cies with wide and rather shallow, quadrangular margin. Mesotibia moderately elongate, dorsal rim clypeal excision, sulcate, moderately densely punc- crenulate, with several rows of setae on the dorsal tate head, rufous middle and hind legs, rather conical surface; subapical spur elongate. Metatibia elongate, prothorax, impunctate and glabrous proepisternum, rather sparsely setose. Metatarsomere 1 almost as elongate, parallel-sided elytra, and finely punctate long as the following three tarsomeres. elytral striae. Male genitalia. Unknown. Female gonocoxites (Fig. 69). Gonocoxites nar- Description row and elongate, little curved, with rather acute apex; with one elongate, rather stout seta basally Measurements. Length: 11.8-14.1 mm; width: 3.4- at lateral surface and a smaller seta below, two 4.0 mm. Ratios. Length/width of pronotum: 0.96- elongate setae on the ventral surface of gonocoxite 0.97; base/apex of pronotum: 1.37-1.40; width 1, two elongate setae at middle of the medio-dorsal pronotum/head: 1.39-1.43; length/width of elytra: surface, and a tiny nematiform seta near apex. Lateral 2.10-2.13; length/width of protibia: W: 3.0-3.2; plate basally with one elongate seta, apically with length/width of metatibia: W: 6.9-7.2. three shorter seta. Colour (Fig. 99). Uniformly pale red. Mouth Variation. Considerable variation noted in body parts and antenna pale red, middle and hind legs size. yellow to pale red, anterior leg slightly darker. Head (Fig. 99). Eye large, laterally remarkably Distribution. Southern central NT, adjacent extreme projected, orbit very short, oblique-convex. Clypeus eastern WA. with rather shallow, wide, almost quadrangular excision. Labrum 7-setose. Clypeus divided from Collecting circumstances. Little recorded. Both frons by a deep sulcus; frons in anterior half usually specimens were collected at light.

255 brevisterna subgroup gular excision. Labrum 7-setose. Mandibles short. Clypeus divided from frons by a sulcus; upper Diagnosis. Medium sized to large species with surface rather convex, with an irregularly triagonal short, conical prothorax and usually rather short, sulcus in middle of frons, rather densely punctate. oval-shaped elytra; clypeal excision wide and shal- Surface without microreticulation, glossy. low; clypeus 7-setose; eye large, usually very much Pronotum (Fig. 100). Moderately elongate, trap- protruded; apex of aedeagus arrow- or club-shaped: ezoidal, narrowed apicad, dorsally gently convex. metathoracal wings reduced and elytra commonly Lateral margin oblique and usually slightly convex. soldered together. 10 species occur in northern Apex gently concave. Marginal channel narrow; Australia. median line well impressed, impunctate. Basal sulcus narrow. Anterior marginal seta situated about at apical third. Basal groove shallow, elongate, Clivina brevisterna Sloane linear, impunctate. Disk with some fine, irregularly Figs 37, 100 transverse strioles in middle, impunctate, with very fine but distinct, isodiametric microreticulation, Clivina brevisterna Sloane, 1916: 606. – Sloane 1917: 406; moderately glossy. Csiki 1927: 498; Moore et al. 1987: 68; Baehr 1987: Elytra (Fig. 100). Soldered together. Moder- 190; Lorenz 2005: 142. ately elongate, rather narrow at base, oval shaped, Examined types. Holotype of brevisterna: sex? (only convex, on disk barely depressed. Basal angle little hindbody, but genitalia destroyed), Pine Creek Disct. produced. Striation complete, striae deep, apicad N. Ter. (C.F. 09) Type / Clivina brevisterna, Sl. Type / slightly shallower, coarsely punctate. Intervals HOLOTYPE Clivina brevisterna Sl. PJD (ANIC). convex, impunctate, with or without fine transverse Type locality. “Pine Creek”, Northern Territory. strioles, with fine, slightly superficial, isodiametric microreticulation, moderately glossy. Epipleura Other material (9 ex.). NT: 2 WW, Kakadu Nat. Park, narrow, basal sulcus barely indicated. CSIRO Biodiversity Survey, Site 14, 13.73101 S, 132.77066 Metathoracic wings. Reduced. E, S. Oberprieler, Feb-Mar 2014 (CBM); 1 M, 20 km N.T. Lower surface. Proepisternum impunctate, Ry. w. Pt. Darwin G. Sc….(unreadable) (SAMA 033658); without or with some extremely fine dorso-ventral 1 W, Pt Darwin / Clivina brevisterna Sl. Id. by T. G. Sloane strioles; with faint isodiametric microreticulation. / F. E. Wilson Collection (NMV COL-5199); 1 W, Pt. Darwin / H. J. Carter Coll. p. 20.4.22. (NMV COL-5200); The whole lower surface with fine, isodiametric mi- 1 M, N. Territory / H. J. Carter Coll. p. 20.4.22. / Clivina croreticulation. Metepisternum short and small, little brevisterna Sl. mss. (NMV COL-5198); 1 M, Northern longer than wide. Abdominal sterna impunctate. Territory (ANIC); 2 WW, N. Territory (WADAP 5184-5). Legs. Protibia narrow and elongate, in females slightly less so. Longitudinal sulcus on upper surface Diagnosis. Medium sized, black species with mod- deep; lower surface with a row of 3-4 stout setae erately shallow, quadrangular clypeal excision, along basal part of inner margin. Mesotibia rather slightly sulcate head, 7-setose labrum, dark piceous short, dorsal rim little crenulate, with two rows of few to black legs, apicad narrowed prothorax with setae on the dorsal surface; subapical spur elongate. oblique lateral margin, impunctate proepisternum, Metatibia elongate, rather sparsely setose. Metatarsus moderately elongate, oviform elytra, deep, coarsely elongate, metatarsomere 1 slightly shorter than the punctate elytral striae, and wide, club-shaped apex following three tarsomeres. of the aedeagus. Male genitalia (Fig. 37). Aedeagus wide, fairly elongate, asymmetrically narrowed apicad; lower Description surface in basal half gently concave, then almost Measurements. Length: 11.3-12.8 mm; width: 3.5- straight, apical part turned down. Apical half of 3.8 mm. Ratios. Length/width of pronotum: 0.97- lower surface deeply concave. Apex short, markedly 1.02; base/apex of pronotum: 1.42-1.45; width club-shaped and hooked at the lower surface, club pronotum/head: 1.52-1.53; length/width of elytra: short and wide, at tip convex. Both parameres mod- 1.81-1.86; length/width of protibia: M: 3.5-3.8, erately elongate, the left one considerably stouter W: 3.25-3.4; length/width of metatibia: M; 6.4-6.5, than the right one; both with rather short, narrow, W: 6.35-6.5. uni- or bisetose apical part; setae very short, situated Colour (Fig. 100). Unicolourous black. Legs dark at or very near to apex. piceous to black. Female gonocoxites. Very similar to those of Head (Fig. 100). Eye large, laterally markedly C. major Sloane. projected, orbit very short, almost perpendicular. Variation. Little variation noted. Clypeus with moderately shallow, almost quadran-

256 Distribution. Northern part of NT. / Conspecific with Clivina brevisterna Sl. (WADAP 5183). – AUS: 2 MM, AUSTRALIEN v. Müller 1892 (SMNS). Collecting circumstances. Not recorded. Diagnosis. Medium sized to large, black species with wide and shallow, quadrangular clypeal exci- Clivina major Sloane (stat. nov.) sion, slightly sulcate head, 7-setose labrum, black Figs 38, 70, 101 legs, trapezoidal prothorax with oblique-convex lateral margin, impunctate proepisternum, elongate, Clivina brevisterna major Sloane, 1917: 406. – Csiki 1927: oviform elytra, deep, moderately punctate elytral 499; Moore et al. 1987: 68; Lorenz 2005: 142. striae, and wide, club-shaped apex of the aedeagus. Examined types. Lectotype of brevisterna major (by present designation): M, King R. N.T. Nat. Mus. Victoria Description Clivina brevisterna Coll. by W. McLennan 22.12.15 / (Sl.) Measurements. Length: 14.2-16.8 mm; width: 4.1- var. major Sl. Id. by T G. Sloane / SYNTYPE T-16907 5.0 mm. Ratios. Length/width of pronotum: 1.0-1.01; Clivina brevisterna var. major Sloane (red) / Type (red) (NMV). base/apex of pronotum: 1.51-1.58; width pronotum/ head: 1.51-1.56; length/width of elytra: 1.91-1.98; Paralectotypes: 2 MM, 2 WW, same data / SYNTYPE length/width of protibia: M: 4.3-5.0, W: 3.7-4.0; T-16903-06 (red) (NMV); 1 M, same data / PARATYPE length/width of metatibia: M; 7.3-7.7, W: 7.5-7.6. (blue) (ANIC). Colour (Fig. 101). Unicolourous black. Legs Type locality. “King River”, Northern Territory. black. Head (Fig. 101). Eye large, laterally markedly . Note. C major was described as a variation of C. bre- projected, orbit short, almost perpendicular. Clypeus visterna Sloane, 1916, and therefore was synonymized with shallow, almost quadrangular excision. Labrum with that species and on labels considered a “large 7-setose. Mandibles short. Clypeus divided from variation”. However, Sloane himself stated the strik- frons by a deep sulcus; upper surface rather convex, ing differences in body size, relative length of the with or without a second transverse sulcus and an elytra, and punctation of the elytral striae, and thus irregularly triagonal sulcus in middle of frons; rather clearly expressed that this “variation” is different sparsely punctate. Surface without microreticula- from C. brevisterna s. str. Therefore, according to the tion, glossy. rules of ICZN (articles 45f,g), this “variation” is to be Pronotum (Fig. 101). Moderately elongate, trap- regarded a subspecies. Because it is not possible, per ezoidal, narrowed apicad, dorsally gently convex. definitionem, that two subspecies of the same spe- Lateral margin oblique and slightly convex. Apex cies occur sympatrically, C. brevisterna major herein gently concave. Marginal channel narrow; median is raised to the status of a species. line well impressed, impunctate. Basal sulcus nar- Other material (32 ex.). NT: 5 MM, 14 WW, King R. N.T. row. Anterior marginal seta situated about at apical 24-12-15 / Coll’ by W. Mc-Lemman Rev. (?) leg. H. L. third. Basal groove fairly deep, elongate, linear, im- White 14-10-16 (NMV COL-5201, 14019, 23, 26, 28, 29- punctate. Disk with some fine, irregularly transverse 37-41, 56; 14031, 39 CBM); 1 M, King R. N.T. 24-12-15 / strioles in middle, impunctate or almost so, with ’ Coll by W. Mc-Lemman Rev. (?) leg. H. L. White 14-10- very fine but distinct, isodiametric microreticulation, 16 / ovipennis by Sl. key, P.J.D. (MCZ); 1 W, King R. N.T. moderately glossy. 24-12-15 / Coll’ by W. Mc-Lemman Rev. (?) leg. H. L. White 14-10-16 / ovipennis Sl. (tr. with mastersi ?) det.’59 Elytra (Fig. 101). Soldered together. Rather Darlington (MCZ); 1 M, 1 W, 24.12.15 Nat. Mus. Victoria elongate, narrow at base, oval shaped, convex, on King R. N.T coll. by W.McLennan (ANIC); 1 W, King R. disk barely depressed. Basal angle little produced. N.T Nat. Mus. Victoria Coll. by W. McLemman 24.12.15 Striation complete, striae deep, apicad shallower, in / Det. by T. G. Sloane to be Clivina brevisterna var. major basal half moderately coarsely punctate. Intervals Rec. 3-17 (DEI); 1 (?sex, defect), King R. N.T Nat. Mus. convex, impunctate, with or without fine transverse Victoria Coll. by W. McLemman 24-12-15 (ANIC); 1 W, strioles, with fine, slightly superficial, isodiametric Caught on flooded arm King R. N.T by W. Mclennan microreticulation, moderately glossy. Epipleura 24-12-15 / Clivina sp.? probably a large form of brevis- narrow, basal sulcus barely indicated. terna Sl. Id. by T. G. Sloane / ?C. brevisterna Sl. det. K. Metathoracic wings. Reduced. Kult 1948 (DEI); 1 M, NT. 12.816°S 131.569°E 30kmW of Lower surface. Proepisternum impunctate, Adelaide R. 30m 21-22Feb2012 G. Monteith. open forest. dung trap. 35341 (QMB); 1 M, Adelaide River N. Aust. without or with some extremely fine dorso-ventral J. Farr / Clivina brevisterna Sl. (NMV COL-5203); 1 M, strioles; with faint isodiametric microreticulation. Northern Territory / Clivina brevisterna Sl. var. major The whole lower surface with fine, isodiametric mi- SL. Id. by T. G. Sloane / F. E. Wilson Collection (NMV croreticulation. Metepisternum short and small, little COL-5202). – WA: 1 W, Kellerberrin W. Australia J. Clark longer than wide. Abdominal sterna impunctate.

257 Legs. Protibia narrow and elongate, in females N.T. / Clivina mastersi Sl. Id. by T. G. Sloane (SAMA); less so. Longitudinal sulcus on upper surface deep; N.T. Fogg Dam Area12.34S 131.18E pit trap no. 5 rain lower surface with a row of 5-6 stout setae along forest Nov.1980 N.T. Field Nat. Club (CBM, NTD); basal part of inner margin. Mesotibia rather short, Howard Springs N.T. 22.iii.68 B.P.Moore (ANIC, CBM); dorsal rim little crenulate, with two rows of few NT: 12.393°S130.9191°E Vanderlin Drive, Darwin 22- 23Feb2012 G.Monteith. open forest mushroom trap setae on the dorsal surface; subapical spur elongate. 35359 (QMB). – AUS: C. mastersi Sl. Comp. with type Metatibia very elongate, rather sparsely setose. Meta- 7/7/21 (ANIC); Australia / Clivina mastersi Sl.Bänninger tarsus elongate, metatarsomere 1 slightly shorter det. 1944 / Collectio Karel Kult Coll. A.Dostal, 1999 than the following three tarsomeres. (CDW). Male genitalia (Fig. 38). Aedeagus wide, fairly elongate, slightly asymmetrically narrowed apicad; Diagnosis. Large, black species with wide and lower surface in basal third gently concave, in middle rather shallow, quadrangular clypeal excision, barely gently convex, apical part curved down. Apical half sulcate head, 7-setose labrum, black legs, trapezoi- of lower surface slightly concave. Apex short, mark- dal prothorax with convex-oblique lateral margin, edly club-shaped and hooked at the lower surface, impunctate proepisternum, elongate, oviform elytra, club short and wide; at tip convex. Both parameres deep, almost impunctate median elytral striae, and elongate, the left one considerably stouter than the short, club-shaped apex of the aedeagus. right one; both with very elongate, narrow, uni or bisetose apical part; setae short, situated right at apex. Description Female gonocoxites (Fig. 70). Gonocoxites wide Measurements. Length: 15.4-21.1 mm; width: 4.5- and rather short, little curved, with rather acute apex; 6.0 mm. Ratios. Length/width of pronotum: 0.98- with one elongate, stout seta basally at lateral surface 1.05; base/apex of pronotum: 1.50-1.55; width and a smaller seta below, four elongate setae on the pronotum/head: 1.50-1.57; length/width of elytra: ventral surface of gonocoxite 1, two setae at middle 1.96-2.07; length/width of protibia: M: 4.6-4.8, W: 3.9- of the medio-dorsal surface, without a nematiform 4.0; length/width of metatibia: M: 7.5-7.8, W: 6.7-6.8. seta near apex. Lateral plate basally with one elongate Colour (Fig. 102). Unicolourous black. Legs dark seta, apically apparently without setae. piceous to black. Variation. Some variation noted in body size, Head (Fig. 102). Eye fairly large, laterally well 1 surface structure of head, shape of prothorax, projected, orbit rather short, c. /4 of length of eye, length of elytra, and degree of punctation of the oblique. Clypeus with wide and rather shallow, elytral striae. almost quadrangular excision. Labrum 7-setose. Mandibles short. Clypeus not or indistinctly divided Distribution. Northern parts of NT. The record from from frons; upper surface rather convex, densely but south-western WA is certainly erroneous. very finely punctate, frons in middle with a shallow Collecting circumstances. Little recorded. One fovea. Surface without microreticulation, glossy. specimen “caught on flooded arm King R.”, another Pronotum (Fig. 102). Moderately elongate, con- in “open forest, dung trap”. vexly trapezoidal, strongly narrowed apicad, dorsally gently convex. Lateral margin oblique-convex. Apex gently concave. Marginal channel narrow; median line well impressed, impunctate. Basal sulcus Clivina mastersi Sloane narrow. Anterior marginal seta situated slightly in Figs 39, 71, 102 front of apical third. Basal groove elongate, linear, Clivina mastersi Sloane, 1896a: 242. – Sloane 1896a: 228, impunctate, situated in a wide depression. Disk with 245; Csiki 1927: 508; Moore et al. 1987: 72; Lorenz some fine, irregularly transverse strioles in middle, 2005: 143. impunctate, with very fine, slightly superficial, isodiametric microreticulation, rather glossy. Examined types. Holotype: fragment (sex?, only hind Elytra (Fig. 102). Soldered together. Elongate, body left, but genitalia destroyed), Port Darwin N. Ter- rather narrow at base, oval shaped, convex but on ritory / HOLOTYPE (red) (ANIC-MMS). disk slightly depressed. Basal angle little produced. Type locality. “Port Darwin”, Northern Territory. Striation incomplete, only the median four or five striae distinct in basal half. Median striae deep, im- Other material (37 ex.). NT: Pt. Darwin / K. K. Spence Collection. / C. mastersi (?) Sloane id. K.K.Spence / punctate. Apical half or third almost striate. Median Clivina sp. Det. B.P. Moore 1999 (AMS); Port Darwin intervals in basal half convex, impunctate, without N. Territory / Griffith Collection Id. by A. M. Lea transverse strioles, with very fine, slightly superfi- (SAMA 25-033667-69, 75-81); Port Darwin N. Territory cial, isodiametric microreticulation, rather glossy. (SAMA 25-033-607); G. F. Hill Alligator Riv. Darwin. Epipleura narrow, basal sulcus barely perceptible.

258 Metathoracic wings. Reduced. Etymology. The name refers to the range of the spe- Lower surface. Proepisternum impunctate, with cies, Cobourg Peninsula in the northernmost part of or without some fine dorso-ventral strioles; with Northern Territory. faint isodiametric microreticulation. The whole Diagnosis. Large, black species with wide and lower surface with fine, isodiametric microreticula- rather shallow, quadrangular clypeal excision, tion. Metepisternum short, c. 1.25 × as long as wide. barely sulcate head, 7-setose labrum, black legs, Abdominal sterna impunctate. trapezoidal prothorax with oblique lateral margin, Legs. Protibia very narrow and elongate, less impunctate proepisternum, elongate, oviform elytra, so in females. Longitudinal sulcus on upper surface deep, almost impunctate elytral striae, and elongate, deep; lower surface with a row of 3-5 stout setae spatulate apex of the aedeagus. along basal part of inner margin. Mesotibia rather elongate, dorsal rim barely crenulate, with two rows Description of few setae on the dorsal surface; subapical spur moderately elongate. Metatibia very elongate, rather Measurements. Length: 17.8-18.9 mm; width: 5.25- sparsely setose. Metatarsus elongate, metatarsomere 5.4 mm. Ratios. Length/width of pronotum: 1.0-1.01; 1 slightly shorter than the following three tarsomeres. base/apex of pronotum: 1.51-1.54; width pronotum/ Male genitalia (Fig. 39). Aedeagus wide, fairly head: 1.51-1.54; length/width of elytra: 1.93-1.98; elongate, narrowed apicad; lower surface in basal length/width of protibia: M: 4.5-4.65; length/width third gently concave, then almost straight, but apex of metatibia: M: 6.9-7.2. suddenly turned down. Apical half of lower surface Colour (Fig. 103). Unicolourous black. Legs with two longitudinal, parallel ridges, the surface black. between deeply concave. Apical part slightly asym- Head (Fig. 103). Eye fairly large, laterally well 1 metrically narrowed; apex short, markedly club- projected, orbit rather short, < /4 of length of eye, shaped and hooked at the lower surface, club short oblique. Clypeus with wide and rather shallow, and wide; at tip convex. Both parameres elongate, almost quadrangular excision. Labrum 7-setose, in the left one considerably stouter than the narrower one specimen 6-setose, one paralateral seta lacking. right one; both with elongate, very narrow, asetose Mandibles short. Clypeus distinctly divided from apical part, frons; upper surface rather convex, sparsely punctate. Female gonocoxites (Fig. 71). Gonocoxites rather Surface without microreticulation, glossy. narrow and elongate, little curved, with rather acute Pronotum (Fig. 102). Moderately elongate, apex; with one elongate, fairly stout seta basally at markedly trapezoidal, strongly narrowed apicad, lateral surface and a smaller seta below, two elongate dorsally gently convex. Lateral margin oblique but setae on the ventral surface of gonocoxite 1, three straight. Apex gently concave. Marginal channel or four setae at middle of the medio-dorsal surface, narrow; median line well impressed, impunctate. without a nematiform seta near apex. Lateral plate Basal sulcus narrow. Anterior marginal seta situ- basally with one elongate seta, apically apparently ated slightly in front of apical third. Basal groove without setae. rather deep, elongate, linear, impunctate. Disk with Variation. Considerable variation noted in body some fine, irregularly transverse strioles in middle, size, also in relative length of prothorax and elytra, extremely finely and sparsely punctate, with very and in degree of elytral striation. fine, extremely superficial, isodiametric microreticulation, glossy. Distribution. Northern NT. Elytra (Fig. 103). Soldered together. Elongate, rather narrow at base, oval shaped, convex but on Collecting circumstances. Little recorded. One disk slightly depressed. Basal angle little produced. specimen collected in “pit trap in rain forest”, another Striation almost complete, but the 7th stria very weak. in “mushroom trap”. Striae deep, impunctate or very finely punctate, becoming slightly shallower apicad. Most intervals convex, impunctate, without transverse strioles, with Clivina cobourgiana, spec. nov. extremely fine and superficial, barely perceptible, Figs 40, 103 isodiametric microreticulation, glossy. Epipleura narrow, basal sulcus barely perceptible. Examined types. Holotype: M, N.T. Murgenella 11.34 S Metathoracic wings. Reduced. 132.52 E 3-7 Feb. 1987 pitfall trap P. Horner / burnt Eucalyptus tetrodonta forest (NTD). – Paratype: 2 MM, Lower surface. Proepisternum impunctate, with 11.09 S 132.09 E Black Point Coburg Pen. N.T. 15-23 or without some fine dorso-ventral strioles; with Feb. 1977 T.A. Weir (ANIC, CBM). faint isodiametric microreticulation. The whole lower surface with fine, isodiametric microreticula-

259 tion. Metepisternum short, c. 1.25 × as long as wide. Ent. Belg. Coll. Putzeys / Type (red) / Syntype (red) Abdominal sterna impunctate. (IRSNB). – Holotype of propinqua (= interstitialis Sloane, Legs. Protibia very narrow and elongate. Longi- 1896): W (probable, abdomen destroyed), C. interstitialis, d tudinal sulcus on upper surface deep; lower surface Sl. N. Queensl . (C.F.) (ANIC). with a row of 6-7 stout setae along basal part of Type localities. Of nyctosyloides: “Rockhampton”, inner margin. Mesotibia rather elongate, dorsal rim Queensland. – Of propinqua: “Cooktown”, Queensland. barely crenulate, with two rows of few setae on the Other material (25 ex.). QLD: 8 WW, Townsville Aus- dorsal surface; subapical spur moderately elongate. tralia / BMalkin Jan 1945 / Clivina nyctosyloides Putz. Metatibia elongate, rather sparsely setose. Metatarsus det Darlington ‘48 (CBM, MCZ, USNM); 1 W, Townsville elongate, metatarsomere 1 slightly shorter than the Australia / nyctosyloides det Darlington ‘48 / Collectio following three tarsomeres. Karel Kult Coll.A.Dostal,1989 (CDW); 1 W, Towns- Male genitalia (Fig. 40). Aedeagus wide, fairly ville Qld Feb 01 F. P. Dodd / C. nyctosyloides Putz. elongate, very asymmetrically narrowed apicad; Townsville P. FPD (ANIC); 1 W, Townsville Q. T G. S. lower surface in basal third concave, then markedly 6-06 (ANIC); 1 W, Pt. Denison / Clivina nyctosyloides Putz. convex, apical part suddenly turned down. Apical Port Denison (MMS); 1 W, Cooktown C. French (QMB); half of lower surface with two longitudinal ridges 1 W, Endevour R. N.S. Wales / oblongicollis Putz. (ZISP); which form a triangle, the surface between the ridges 1 W, End. Riv. / Nat. Mus. Victoria C. French’s Coll. 6.I.08 / Clivina nyctosyloides Putz. Queensland (NMV COL- deeply concave. Apex rather elongate, markedly 5391); 1 W, Endeavour R. / Nat. Mus. Victoria C. French’s spatulate and hooked at the lower surface, the club Coll. 6.I.08 (NMV COL-5392); 1 W, Coen R. Q. W. D. elongate and asymmetrically curved left, at tip ob- Dodd (SAMA 25-033673); 1 W, Cairns dist. F. P. Dodd tusely convex. Both parameres elongate, the left one / I.6872 Clivina nyctosyloides Putz. Id. by T. G. Sloane considerably stouter than the right one; both with (SAMA); 1 M, 3 WW, Mutchilba, N. Q. Nov. 1933 A. D. elongate, narrow, asetose or unisetose apical part; Selby / F. E. Wilson Collection (NMV COL-13676-9); 1 W, setae, if present, very short and situated right at apex. AUSTRALIA N Queensland 5km S of Prairie, 44km E Female gonocoxites. Unknown. of Hughenden,19-21.I.2011, 433m 20°53'S, 144°36'E; St. Variation. Little variation noted in striation of Jakl lgt. (CJP); 1 W, AUSTRALIA, C. Queensland PODDY the elytra and distinctness of punctation of the striae. CREEK, 90 km W Winton 23-24.I.2011, 198 m alt.S 22°12' E 142°12' St Jakl lgt. (CBM); 1 M, Australia Qld. Norman- ton Lamond / 15-i-2000, / E. Gowing-Scopes Collection Distribution. Cobourg Peninsula, extreme northern (NHM); 1 M, Edungalba CQ 1-12-46 E Adams E Sutton / NT. E. Sutton Collection don.-Dec.1964. (QMB). Collecting circumstances. Little recorded. Holo- Notes. According to Moore et al. (1987) only the type collected in “pitfall trap in burnt Eucalyptus holotype of C. nyctosyloides exists, but this is a mis- tetrodonta forest”. interpretation of a note in the description about an aberrantly coloured specimen (Putzeys 1868: 12). Moreover, Moore certainly had not seen any type Clivina nyctosyloides Putzeys material. The mentioned note, therefore, reveals that Figs 41, 72, 104 more than a single specimen was available for the description. Accordingly, a lectotype is designated Clivina nyctosyloides Putzeys, 1868: 10. – Sloane 1896a: from the three existing type specimens. The men- 181, 227, 231, 239, 242; 1905a: 732, 733; 1907: 349; 1916: tioned aberrant specimen is the female paralectotype. 606; Csiki 1927: 508; Moore et al. 1987: 73; Baehr 1987: C. interstitialis Sloane was thought by Sloane to 190; Lorenz 2005: 143. be identical with, or only a minor variation of, C. nyc- Clivina interstitialis Sloane, 1896a: 241. – Sloane 1896a: 227; 1905a: 733; Csiki 1927: 508; Moore et al. 1987: 73; tosyloides Putzeys. Therefore Sloane (1905, p. 733) Lorenz 2005: 143. synonymized both species. Although the types of Clivina propinqua nom. nov. = interstitialis Sloane, 1896a the two species differ rather in body size and are (non C. interstitialis Kolbe, 1883). also somewhat different in the shape of their elytra, a number of specimens exist along the eastern coast Examined types. Lectotype of nyctosyloides (by pre- of Queensland which are intermediary in body size sent designation): M, Rockhampton / Rockhampton and shape between both. Therefore I think that this (Australia) Coll. Castelnau / C. nyctosyloides P. (red) / SYNTYPUS Clivina nyctosyloides Putzeys, 1868 (red) synonymy is justified. (MCSN). – Paralectotypes: 1 W, Rockhampton / Rock- Moore et al. (1987) give “Port Darwin, N. T.” as hampton (Australia) Coll. Castelnau / SYNTYPUS type locality of C. interstitialis. This is an error, as Clivina nyctosyloides Putzeys, 1868 (red) (MCSN); 1 M, can be taken as well from the description as from Rockhampton / C. nyctosyloides P. Rockh. M / Soc. the label of the holotype.

260 Because the name interstitialis Sloane, 1896 is strioles; with faint isodiametric microreticulation. preoccupied by C. interstitialis Kolbe, 1883, it is The whole lower surface with fine, isodiamet- renamed “propinqua”. ric microreticulation. Metepisternum moderately short, c. 1.5 × as long as wide. Abdominal sterna Diagnosis. Medium sized, black species with wide impunctate. and shallow, quadrangular clypeal excision, barely Legs. Protibia narrow and elongate, in females sulcate head, 7-setose labrum, piceous to black legs, less so. Longitudinal sulcus on upper surface deep; trapezoidal prothorax with oblique lateral margin, lower surface with a row of 4-5 stout setae along impunctate proepisternum, rather short, oviform basal part of inner margin. Mesotibia rather short, elytra, deep, coarsely punctate elytral striae, abso- dorsal rim crenulate, with two rows of elongate lutely glabrous dorsal surface, and short and wide, setae on the dorsal surface; subapical spur elongate. arrow-shaped, asymmetric apex of the aedeagus. Metatibia elongate, rather sparsely setose. Metatarsus fairly elongate, metatarsomere 1 slightly shorter than Description the following three tarsomeres. Measurements. Length: 9.6-13.0 mm; width: 3.1- Male genitalia (Fig. 41). Aedeagus wide, fairly 4.05 mm. Ratios. Length/width of pronotum: 0.90- elongate, slightly asymmetrically narrowed apicad; 0.97; base/apex of pronotum: 1.56-1.67; width lower surface in basal third gently concave, then pronotum/head: 1.56-1.66; length/width of elytra: almost straight, apical part barely down. Apical 1.80-1.85; length/width of protibia: M: 3.75-3.85, half of lower surface gently concave. Apex short, W: 3.4-3.5; length/width of metatibia: M; 6.3-6.4, markedly arrow-shaped, arrow short and rather W: 5.5-5.7. wide, triangular, asymmetrically situated, at tip Colour (Fig. 104). Unicolourous black. Legs obtusely convex. Both parameres stout, the left one piceous to black. considerably stouter than the right one; both with Head (Fig. 104). Eye large, laterally markedly rather short to very short, trisetose apical part, that projected, orbit very short, almost perpendicular. in the left paramere is almost not existent; the two Clypeus with wide and very shallow, almost quad- longer setae situated right at the apex, the shorter rangular excision, though the lateral margins of the one slightly below apex. excision oblique. Labrum 7-setose. Mandibles short. Female gonocoxites (Fig. 72). Gonocoxites mod- Clypeus not or indistinctly divided from frons. Up- erately wide and rather short, little curved, with per surface rather convex, not sulcate, very sparsely slightly convex, transverse apex; with one elongate, punctate. Surface without microreticulation, very stout seta basally at lateral surface and a smaller seta glossy. below, two elongate setae on the ventral surface of Pronotum (Fig. 104). Rather short and wide, gonocoxite 1, two setae at middle of the medio-dorsal trapezoidal, markedly narrowed apicad, dorsally surface, without a nematiform seta near apex. Lateral gently convex. Lateral margin oblique and slightly plate basally with one elongate seta, apically with convex. Apex gently concave. Marginal channel five to six setae. narrow; median line shallow, impunctate. Basal Variation. Some variation noted in body size, sulcus narrow. Anterior marginal seta situated surface structure of head, and shape of prothorax. about at apical third. Basal groove shallow or absent, elongate, linear, impunctate. Disk with some fine, ir- Distribution. Northern and eastern QLD. regularly transverse strioles in middle, sparsely and Collecting circumstances. Little recorded. Two extremely finely punctate, without microreticulation, specimens were collected at light, according to the very glossy. collector. Elytra (Fig. 104). Free. Rather short, narrow at base, oval shaped, convex, on disk barely depressed. Basal angle little produced. Striation complete, Clivina ovalior, spec. nov. striae deep, even apicad barely shallower, coarsely Fig. 105 punctate. Intervals convex, impunctate, without transverse strioles, without microreticulation, very Examined types. Holotype: W, AUSTRALIA QLD glossy. Epipleura narrow, basal sulcus shallowly MOSSMAN 27/28.XII. (MCSV). punctate. Metathoracic wings. In most specimens reduced Diagnosis. Medium sized, black species with wide 2 to about /3 of normal length. and shallow, quadrangular clypeal excision, barely Lower surface. Proepisternum impunctate, sulcate head, 7-setose labrum, black legs, apicad without or with some extremely fine dorso-ventral narrowed prothorax with oblique lateral margin,

261 impunctate proepisternum, rather short, oviform rather sparsely setose. Metatarsus fairly elongate, elytra, deep, coarsely punctate elytral striae, and metatarsomere 1 slightly shorter than the following finely microreticulate pronotum and elytra. three tarsomeres. Male genitalia. Unknown. Etymology. The name refers to the more oval-shaped elytra as compared with C. nyctosyloides Putzeys, 1868. Female gonocoxites. Very similar to that of C. nyctosyloides Putzeys. Description Variation. Unknown.

Measurements. Length: 13.1 mm; width: 4.0 mm. Distribution. North-eastern QLD. Known only Ratios. Length/width of pronotum: 0.96; base/ from type locality. apex of pronotum: 1.46; width pronotum/head: 1.53; length/width of elytra: 1.82; length/width of Collecting circumstances. Not recorded. protibia: W: 3.8; length/width of metatibia: W: 6.9. Colour (Fig. 105). Unicolourous black. Legs black. Clivina nitescens, spec. nov. Head (Fig. 105). Eye large, laterally markedly Figs 43, 106 projected, orbit very short, almost perpendicular. Clypeus with wide and shallow, almost quadran- Examined types. Holotype: M, AUS13, NT21, T/O Stu- gular excision, though the lateral margins of the art Hwy./Edith River Rd., 145m, 14°11'10"S, 132°02'04"E, excision oblique. Labrum 7-setose. Mandibles short. 15.4.2013, M.Baehr (NTD). – Paratypes: 1 M, Scarites n. sp. Clypeus divided from frons. Upper surface rather Westaustralien 53. Schmeltz / Clivina oblongicollis Putz. convex, not sulcate, sparsely punctate. Surface with- (ZISP); 1 W, 3/06 N-Austr, NT, Kakadu NP Muirella, 10.06.2006 40m S12°51'15"' E132°45'16" Light lg Berger- out microreticulation, glossy. Dostal (CBM). Pronotum (Fig. 105). Rather short and wide, moderately narrowed apicad, dorsally gently convex. Diagnosis. Rather small to medium sized, black Lateral margin oblique and slightly convex. Apex species with wide and shallow, quadrangular clypeal gently concave. Marginal channel narrow; median excision, barely sulcate head, 7-setose labrum, black line shallow, impunctate. Basal sulcus narrow. An- legs, trapezoidal prothorax with oblique-convex lat- terior marginal seta situated about at apical third. eral margin, impunctate proepisternum, rather short, Basal groove shallow, elongate, linear, impunctate. oviform elytra, deep, coarsely punctate elytral striae, Disk with some fine, irregularly transverse strioles absolutely glabrous dorsal surface, and elongate, in middle, sparsely and extremely finely punctate, arrow-shaped, asymmetric apex of the aedeagus. with very fine though distinct microreticulation, Etymology. The name refers to the very glossy dorsal rather glossy. surface. Elytra (Fig. 105). Soldered together. Rather short, narrow at base, oval shaped, convex, on disk barely Description depressed. Basal angle little produced. Striation complete, striae deep, but in apical half shallower, in Measurements. Length: 10.3-11.7 mm; width: 3.0- basal half rather coarsely punctate. Intervals convex, 3.6 mm. Ratios. Length/width of pronotum: 0.92- impunctate, with some fine, transverse strioles, with 0.95; base/apex of pronotum: 1.54-1.60; width very fine and rather superficial microreticulation, pronotum/head: 1.56-1.61; length/width of elytra: fairly glossy. Epipleura narrow, basal sulcus barely 1.84-1.92; length/width of protibia: M: 3.45-3.5, indicated. W: 3.25; length/width of metatibia: M; 6.3-6.35, W: 6.2. Metathoracic wings. Reduced. Colour (Fig. 106). Unicolourous black. Legs Lower surface. Proepisternum impunctate, with piceous to black. some extremely fine dorso-ventral strioles; with faint Head (Fig. 106). Eye large, laterally markedly isodiametric microreticulation. The whole lower projected, orbit very short, almost perpendicular. surface with fine, isodiametric microreticulation. Clypeus with wide and very shallow, almost quad- Metepisternum short, slightly longer than wide. rangular excision, though the lateral margins of the Abdominal sterna impunctate. excision oblique. Labrum 7-setose. Mandibles short. Legs. Protibia narrow and elongate. Longitu- Clypeus not or indistinctly divided from frons. Up- dinal sulcus on upper surface deep; lower surface per surface rather convex, not sulcate, very sparsely with a row of 5 stout setae along basal part of inner punctate. Surface without microreticulation, very margin. Mesotibia rather short, dorsal rim crenu- glossy. late, with two rows of elongate setae on the dorsal Pronotum (Fig. 106). Rather short and wide, surface; subapical spur elongate. Metatibia elongate, trapezoidal, markedly narrowed apicad, dorsally

262 rather convex. Lateral margin oblique and slightly Collecting circumstances. The holotype and one convex. Apex gently concave. Marginal channel paratype were collected at light near billabongs narrow; median line shallow, impunctate. Basal and ponds. sulcus narrow. Anterior marginal seta situated about at apical third. Basal groove shallow or absent, elongate, linear, impunctate. Disk with some fine, Clivina ovalipennis Sloane irregularly transverse strioles in middle, extremely Fig. 107 sparsely and finely punctate, without microreticulation, very glossy. Clivina ovipennis Sloane, 1896a: 244 (non Clivina ovipennis Elytra (Fig. 106). Free. Rather short, rather wide Chaudoir, 1846). – Sloane 1896a: 228, 244; 1905a: 733; at base, slightly oval shaped, convex, on disk slightly Csiki 1927: 509; Moore et al. 1987: 74; Lorenz 2005: depressed. Basal angle little produced. Striation 143. Clivina ovalipennis Sloane, 1905a: 733 (nom. nov. for complete, striae deep, even apicad barely shallower, Clivina ovipennis Sloane, 1896). – Csiki 1927: 509; coarsely punctate. Intervals convex, impunctate, Moore et al. 1987: 74; Lorenz 2005: 143. without transverse strioles, without microreticulation, very glossy. Epipleura narrow, basal sulcus Examined types. Holotype: W (probable, head and d shallowly punctate. abdomen lacking), C. ovipennis, Sl. N. Queensl (C.F.) / Metathoracic wings. Fully developed. HOLOYTPE Clivina ovipennis Sl PJD (ANIC). Lower surface. Proepisternum impunctate, Type locality. “North Queensland”. without or with some fine dorso-ventral strioles; with finest traces only of isodiametric microreticu- Note. Because the type of C. ovalipennnis is badly lation. The lower surface without or with traces of defect and no specimens of similar shape of the fine, isodiametric microreticulation. Metepisternum elytra have been found in the available material, moderately elongate, > 1.5 × as long as wide. Ab- this species is regarded quite doubtful. dominal sterna impunctate. Other material. None. Legs. Protibia narrow and elongate, in the female slightly less so. Longitudinal sulcus on upper surface Diagnosis. Medium sized, black species with pi- distinct; lower surface with a row of 4-6 stout setae ceous legs, trapezoidal prothorax with oblique- along basal part of inner margin. Mesotibia rather convex lateral margin, impunctate proepisternum, short, dorsal rim crenulate, with two rows of elongate elongate, oviform elytra, deep, coarsely punctate ely- setae on the dorsal surface; subapical spur elongate. tral striae, and finely microreticulate dorsal surface. Metatibia elongate, rather sparsely setose. Metatarsus fairly elongate, metatarsomere 1 slightly shorter than Description the following three tarsomeres. Measurements. Length: c. 12.6 mm; width: 4.05 mm. Male genitalia (Fig. 43). Aedeagus rather wide, Ratios. Length/width of pronotum: 1.0; base/apex moderately elongate, almost symmetrically nar- of pronotum: 1.48; width pronotum/head: ?; length/ rowed apicad; lower surface in basal third concave, width of elytra: 1.85; length/width of protibia: then straight, towards apex very slightly curved W: c. 3.4; length/width of metatibia: W: c. 6.5. down. Apical half of lower surface with two longi- Colour (Fig. 107). Unicolourous black. Legs tudinal ridges which are arranged in a triangle, the piceous. surface between concave. Apex wide, arrow shaped, Head. Lacking. and denticulate on the lower surface; arrow elongate, Pronotum (Fig. 107). Moderately short, strongly asymmetrically situated, tip obtusely angulate. Both narrowed apicad, dorsally gently convex. Lateral parameres elongate, the left one stouter than the margin oblique and slightly convex. Apex gently right one; both with short, bisetose apical part; setae concave. Marginal channel narrow; median line very rather elongate, situated right at tip. shallow, impunctate. Basal sulcus narrow. Anterior Female gonocoxites. Very similar to that of marginal seta situated about at apical third. Basal C. nyctosyloides Putzeys. groove shallow, very elongate, linear, impunctate. Variation. Little variation noted in length and Disk with some fine, irregularly transverse strioles shape of elytra and in distinctness of the microre- in middle, barely punctate, with very fine but super- ticulation on proepisternum and abdomen. ficial, isodiametric microreticulation, rather glossy. Elytra (Fig. 107). Soldered together. Rather elon- Distribution. NorthernNT, probably also northern gate, rather narrow at base, oval shaped, convex, on WA. disk barely depressed. Basal angle little produced. Striation complete, but 7th stria very weak. Striae

263 deep, median striae in apical third, lateral striae CALM Pilbara Survey 53km NNE Whim Creek Hotel, in apical half fading, apex barely striate. Striae in 20°25'48.6"S, 118°3'50.3"E DRE03, 14.XI.2003-12.V.2004 basal half coarsely punctate. Intervals in basal half / CALM PBS00060 (CBM, WAM); 6 WW, W-Austral- convex, impunctate, without transverse strioles, with ia, DRE07, sw. Pt.Hedland, 20.28.54.6S 117.59.35.1E, very fine, superficial, isodiametric microreticulation, 15.11.2003-13.5.2004, leg.Guthrie (CBM, WAM); 3 WW, W-Australia, DRC08, w. Roeburne, 20.51.14.5S moderately glossy. Epipleura narrow, basal sulcus 117.05.40.2E, 12.11.2003-11.5.2004, leg. Guthrie (WAM); barely indicated. 1 W, W-Australia, DRW08, w. Mt.Brockman, 22.34.33.2S Metathoracic wings. Reduced. 116.22.13.3E, 28.11.2003-11.5.2004, leg. Guthrie Lower surface. Proepisternum impunctate, with (WAM); 1 W, W-Australia, MBW08, 11 km N Wood- some fine dorso-ventral strioles; with faint isodia- gina, 21°04'18"S 118°40'47"E, 23.9.2005-15.5.2006, CALM metric microreticulation. The whole lower surface PBS01242 (WAM). with fine, isodiametric microreticulation. Metepi Etymology. The name refers to the stout and heavy sternum short, < 1.25 × as long as wide. Abdominal body. sterna impunctate. Legs. Protibia narrow and elongate, the lateral Diagnosis. Medium sized, black species with wide teeth apparently rather worn. Longitudinal sulcus on and shallow, quadrangular clypeal excision, mul- upper surface barely indicated; lower surface with tisulcate head, 7-setose labrum, black legs, very a row of 7-8 stout setae along basal part of inner wide, trapezoidal prothorax with very oblique lat- margin. Mesotibia rather short, dorsal rim crenulate, eral margin, impunctate proepisternum, short and with two rows of several setae on the dorsal surface; compact, oviform elytra, deep, moderately punctate subapical spur elongate. Metatibia elongate, rather elytral striae, and obtusely arrow-shaped apex of sparsely setose. Metatarsus elongate, metatarsomere the aedeagus. 1 slightly shorter than the following three tarsomeres. Male genitalia. Unknown. Description Female gonocoxites. Destroyed. Measurements. Length: 12.0-14.2 mm; width: 3.95- Variation. Unknown. 4.9 mm. Ratios. Length/width of pronotum: 0.86- 0.90; base/apex of pronotum: 1.50-1.56; width Distribution. North QLD, without exact locality. pronotum/head: 1.52-1.59; length/width of elytra: Collecting circumstances. Not recorded. 1.72-1.78; length/width of protibia: M: 3.4-3.5, W: 3.15-3.2; length/width of metatibia: M; 6.7-6.8, W: 5.9-6.1. Clivina pachysoma, spec. nov. Colour (Fig. 108). Unicolourous black. Legs Figs 43, 73, 108 black. Head (Fig. 108). Eye large, laterally well pro- Examined types. Holotype: M, WA: CALM Pilbara jected, orbit short, very oblique. Clypeus with wide Survey 11km ESE Marda Pool, 21°3'20.4"S, 116°16'6"E and very shallow, almost quadrangular excision, DRW07, 27.XI.2003-11.V.2004 / CALM PBS00067 though the lateral margins of the excision oblique. (WAM). – Paratypes: 6 WW (2 defect), WA: CALM Pilbara Labrum 7-setose. Mandibles short. Clypeus divided Survey 11km ESE Marda Pool, 21°3'20.4"S, 116°16'6"E DRW07, 27.XI.2003-11.V.2004 / CALM PBS00067 (CBM, from frons by a shallow sulcus; upper surface rather WAM); 1 M, WA: CALM Pilbara Survey 6km SE Marda convex, with several transverse sulci, more or less Pool, 21°4'11.8"S, 116°12'15.5"E DRW10, 28.XI.2003- sparsely punctate. Surface without microreticula- 11.V.2004 / CALM PBS00068 (WAM); 1 M, 6 WW (2 de- tion, glossy. fect), WA: CALM Pilbara Survey 11km ESE Marda Pool, Pronotum (Fig. 108). Short and very wide, mark- 21°3'20.4"S, 116°16'6"E DRW07, 27.XI.2003-11.V.2004 edly narrow at apex, strongly narrowed apicad, / CALM PBS00067 (CBM, WAM); 2 WW, WA: CALM dorsally rather convex. Lateral margin very oblique Pilbara Survey 13km ESE Marda Pool, 21°3'42.9"S, but only slightly convex. Apex concave. Marginal 116°15'54.2"E DRW06, 27.XI.2003-11.V.2004 / CALM channel narrow; median line very fine, impunctate. PBS00066 (WAM); 3 MM, 2 WW (2 defect), WA: CALM Pil- Basal sulcus narrow. Anterior marginal seta situated bara Survey 3.5km S Marda Pool, 21°3'55.9"S, 116°9'1"E behind apical fourth. Basal groove barely indicated. DRW11, 28.XI.2003-11.V.2004 / CALM PBS00069 (CBM, WAM); 1 M, 4 WW, WA: CALM Pilbara Survey 3.5km Disk with some fine, irregularly transverse strioles in N Karratha Station, 20°51'14.1"S, 116°40'7.9"E DRW05, middle, barely punctate, with or without finest traces 28.XI.2003-11.V.2004 / CALM PBS00064 (WAM); 8 WW, of isodiametric microreticulation, glossy. (3 defect), WA: CALM Pilbara Survey 13km W Wick- Elytra (Fig. 108). Soldered together. Short and ham, 20°41'18.3"S, 117°0'24.9"E DRC11, 13.XI.2003- compact, rather wide at base, moderately oval 9.V.2004 / CALM PBS00059 (CBM, WAM); 2 WW, WA: shaped, dorsally convex but on disk slightly de-

264 pressed. Basal angle little produced. Striation com- Clivina froggatti Sloane plete, but 7th stria weak. Striae deep, apicad shal- Fig. 109 lower, in basal half more or less coarsely punctate, apicad increasingly impunctate. Intervals slightly Clivina froggatti Sloane, 1896b: 278. – Sloane 1916: 604; convex, impunctate, without transverse strioles, Csiki 1927: 505; Moore et al. 1987: 71; Lorenz 2005: without microreticulation, glossy. Epipleura narrow, 143. basal sulcus coarsely punctate. Examined types. Lectotype (by present designation): Metathoracic wings. Reduced. W, C. froggatti, Sl King’s Sound / PARATYPE (blue) Lower surface. Proepisternum impunctate, with (ANIC). some fine dorso-ventral strioles; with faint isodiamet- Type locality. “Kings Sound”, northwestern Australia. ric microreticulation. The whole lower surface with fine, isodiametric microreticulation. Metepisternum Other material. No additional material available. short, c. 1.25 × as long as wide. Abdominal sterna Diagnosis. Small, rufo-piceous species with very impunctate. wide and shallow, clypeal excision, large eye, slightly Legs. Protibia narrow and elongate, less so in sulcate head, 5-setose labrum, rufous legs, moder- females. Longitudinal sulcus on upper surface deep; ately depressed, apicad slightly narrowed prothorax lower surface with a row of 4 stout setae along basal with slightly oblique lateral margin, impunctate part of inner margin. Mesotibia rather short, dorsal proepisternum, rather short, oviform elytra, deep, rim crenulate, with two rows of several setae on punctate elytral striae, and finely microreticulate the dorsal surface; subapical spur very elongate. pronotum but glossy elytra. Metatibia elongate, rather sparsely setose. Metatarsus elongate, metatarsomere 1 slightly shorter than the Description following three tarsomeres. Male genitalia (Fig. 43). Aedeagus wide, elon- Measurements. Length: 7.7 mm; width: 2.3 mm. Ra- gate, almost symmetrically narrowed apicad; lower tios. Length/width of pronotum: 1.06; base/apex of surface in basal third concave, in apical part straight, pronotum: 1.25; width pronotum/head: 1.37; length/ towards apex curved down. Apex short, rather wide, width of elytra: 1.83; length/width of protibia: W: 2.5; obtusely arrow-shaped and denticulate at the lower length/width of metatibia: W: 5.7. surface, at tip obtusely rounded. Lower surface in Colour (Fig. 109). Rufopiceous. Legs rufous. apical half gently concave. Both parameres rather Head (Fig. 109). Eye large, laterally well pro- elongate, the left one considerably stouter than the jected, orbit small, very oblique. Clypeus with very right one; both with elongate, narrow, trisetose apical wide and shallow, in middle straight excision, part; setae very short, two setae situated right at tip, though the lateral margins of the excision are very one shorter seta close to tip. oblique. Labrum 5-setose. Mandibles short. Clypeus Female gonocoxites (Fig. 73). Gonocoxites rather divided from frons by a deep sulcus; upper surface narrow and elongate, little curved, with rather acute rather convex, laterally with some short sulci, ante- apex; with one elongate, stout seta basally at lateral riorly rather densely, posteriorly sparsely punctate, surface and a smaller seta below, three elongate setae without perceptible microreticulation, glossy. on the ventral surface of gonocoxite 1, two setae at Pronotum (Fig. 109). Fairly elongate, wide, wide middle of the medio-dorsal surface, without a ne- at apex, little narrowed apicad, dorsally moderately matiform seta near apex. Lateral plate basally with convex. Lateral margin slightly oblique and very one elongate seta, apically with three setae. slightly convex. Apex concave. Marginal channel Variation. Some variation noted in body size and narrow; median line deep, impunctate. Basal sulcus degree of punctation of the elytral striae. narrow. Anterior marginal seta situated at apical fourth. Basal groove very shallow, slightly oblong. Distribution. Pilbara, north-western WA. Disk with some fine, irregularly transverse strioles in middle, finely but densely punctate, with finest Collecting circumstances. Almost all specimens and extremely superficial traces of about isodiametric were collected in pitfall traps. microreticulation, glossy. Elytra (Fig. 109). Soldered together. Moderately short, rather wide at base, moderately oval shaped, dorsally convex but on disk slightly depressed. Basal angle little produced. Striation complete. Striae deep, apicad barely shallower, rather coarsely punctate, even apicad. Intervals convex, impunctate, without

265 transverse strioles, without microreticulation, glossy. Other material (4 ex.). QLD: 1 W, Kennedy River, 30 km Epipleura narrow, basal sulcus punctate. W of “Fairview” , Qld 15°35' 144°03' 24 Dec 1984 G. and Metathoracic wings. Reduced. A. Daniels / UQIC #90554 (QMB); 2 WW, Coen R, Q. W. Lower surface. Proepisternum impunctate, with D. Dodd (SAMA 25-033672); 1 W, Cooktown V. d. P. / some fine dorso-ventral strioles, with faint isodiamet- Australia. Brit. Mus. 1921–125. / Clivina hackeri Sl. Id. by T. g. Sloane / Collectio Karel Kult Coll.A.Dostal, ric microreticulation. The whole lower surface with 1999 (CDW). fine, isodiametric microreticulation. Metepisternum short, c. 1.25 × as long as wide. Abdominal sterna Diagnosis. Rather small, very compact, black spe- impunctate. cies with wide and shallow, quadrangular clypeal Legs. Protibia rather narrow and elongate. excision, transversely sulcate head, 5- or 7-setose Longitudinal sulcus on upper surface deep; lower labrum, black legs, remarkably trapezoidal protho- surface with a row of 5 stout setae along basal part rax with very oblique lateral margin, impunctate of inner margin. Mesotibia rather short, dorsal rim proepisternum, short, convex, very compact, ovi- crenulate, with two rows of rather few setae on the form elytra, and very deep, rather coarsely punctate dorsal surface; subapical spur very elongate. Metati- elytral striae. bia moderately elongate, rather sparsely setose. Metatarsus moderately elongate, metatarsomere 1 Description slightly shorter than the following three tarsomeres. Measurements. Length: 8.7-10.4 mm; width: 3.15- Male genitalia. Unknown. 3.7 mm. Ratios. Length/width of pronotum: 0.86- Female gonocoxites. Not dissected due to the 0.87; base/apex of pronotum: 1.68-1.92; width fragility of the specimen. pronotum/head: 1.69-1.82; length/width of elytra: Variation. Unknown. 1.53-1.57; length/width of protibia: W: 3.35-3.5; length/width of metatibia: W: 5.6-5.65. Distribution. South-western Kimberleys, northern Colour (Fig. 110). Unicolourous black. Legs WA. black. Collecting circumstances. Unrecorded. Head (Fig. 110). Eye large, laterally well projected, orbit rather short, oblique. Clypeus with wide and very shallow, almost quadrangular excision, hackeri subgroup though the lateral margins of the excision oblique. Labrum 7-setose, rarely 5-setose. Mandibles short. Diagnosis. Rather small to medium sized species Clypeus transversely impressed in middle, divided with short and very wide, markedly conical protho- from frons by a shallow sulcus; upper surface rather rax and short and compact, short-oval elytra; elytral convex, more or less densely punctate. Surface with striae deeply impressed; clypeal excision wide and fine but distinct, isodiametric microreticulation, shallow; clypeus usually 7-setose; eye large, usually rather dull. laterad very much protruded; apex of aedeagus ar- Pronotum (Fig. 110). Short and very wide, mark- row- or club-shaped: metathoracal wings reduced edly narrow at apex, strongly narrowed apicad, and elytra soldered together. Four species occur in dorsally rather convex. Lateral margin very oblique northern Australia. but only slightly convex. Apex slightly concave. Marginal channel narrow; median line very fine, impunctate. Basal sulcus narrow. Anterior marginal Clivina hackeri Sloane seta situated at apical third. Basal groove absent. Fig. 110 Disk with some fine, irregularly transverse strioles in middle, impunctate, with distinct, isodiametric Clivina hackeri Sloane, 1907: 348. – Csiki 1927: 505; Moore microreticulation, dull. et al. 1987: 71; Baehr 1987: 190; Lorenz 2005: 143. Elytra (Fig. 110). Soldered together. Short and Examined types. Holotype: W (probable, abdomen very compact, rather narrow at base, markedly oval lacking), Coen 18/1/06 / HOLOTYPE Clivina hackeri shaped, dorsally remarkably convex. Basal angle Sl PJD (ANIC). – Paratypes: 2 WW, Coen District Cape little produced. Striation complete, even 7th stria York Queensland H. Hacker / Nat. Mus. Victoria. deeply impressed. Striae very deep, apicad little French’s Coll. 5.11.08. / Cotype (blue) / Clivina hackeri shallower, in basal half or two thirds more or less N. Queensld. CoType / PARATYPE T-16010/11 Clivina coarsely punctate, apicad increasingly impunctate. hackeri Sloane (blue) (NMV). Intervals markedly convex, impunctate, without Type locality. “Coen”, North Queensland. transverse strioles, with distinct, isodiametric microreticulation, dull. 3rd stria commonly unilaterally

266 with only 3 setiferous punctures. Epipleura narrow, Dodd 09 / Clivina hackeri Sl. Id. by T. G. Sloane / Pres basal sulcus barely indicated. by T G Sloane Esq 12-6-16 (NMV COL-5326). Metathoracic wings. Reduced. Diagnosis. Rather small, very compact, black spe- Lower surface. Proepisternum impunctate, with cies with wide and shallow, quadrangular clypeal some fine dorso-ventral strioles, with faint isodia- excision, transversely sulcate head, 7-setose labrum, metric microreticulation. The whole lower surface black legs, remarkably trapezoidal prothorax with with fine, isodiametric microreticulation. Metepis- very oblique lateral margin, impunctate proepister- ternum very short, as long as wide. Abdominal num, short, convex, very compact, oviform elytra, sterna impunctate. very deep, rather coarsely punctate elytral striae, Legs. Protibia narrow and elongate. Longitudi- very large, foveate discal punctures, and club-shaped nal sulcus on upper surface deep; lower surface with apex of the aedeagus. a row of 4 stout setae along basal part of inner margin. Mesotibia short, dorsal rim markedly crenulate, with Description two rows of few setae on the dorsal surface; subapical spur very stout. Metatibia elongate, rather sparsely Measurements. Length: 7.7-9.7 mm; width: 2.75- setose. Metatarsus elongate, metatarsomere 1 slightly 3.8 mm. Ratios. Length/width of pronotum: 0.84- shorter than the following three tarsomeres. 0.92; base/apex of pronotum: 1.75-1.82; width Male genitalia. Unknown. pronotum/head: 1.72-1.80; length/width of elytra: Female gonocoxites. Very similar to those of 1.53-1.64; length/width of protibia: M: 3.2-3.25, C. demarzi Baehr. W: 3.0-3.3; length/width of metatibia: M; 5.6-5.7, Variation. Considerably variation noted in shape W: 5.6-5.7. of the pronotum, particularly in relative width of Colour (Fig. 111). Unicolourous black. Legs base and apex; also some variation in degree of black. punctation of the elytral striae, in the number of Head (Fig. 111). Eye rather large, laterally well discal punctures, and the number of labral setae. projected, orbit short, oblique. Clypeus with wide and very shallow, almost quadrangular excision, Distribution. Northern QLD from base to mid CYP. though the lateral margins of the excision oblique. Labrum 7-setose. Mandibles short. Clypeus more Collecting circumstances. Not recorded. or less distinctly divided from frons by a shallow sulcus; upper surface rather convex, without transverse sulci, sparsely punctate. Surface with fine but Clivina demarzi Baehr distinct, isodiametric microreticulation, dull. Figs 44, 74, 111 Pronotum (Fig. 111). Short and very wide, markedly narrow at apex, strongly narrowed apicad, dor- Clivina demarzi Baehr, 1987: 187. – Lorenz 2005: 142. sally rather depressed. Lateral margin very oblique Examined types. Holotype: M, Austral. North T., Kath- but only slightly convex. Apex slightly concave. Basal erine, XII.57, leg. H. Demarz (ZSM). – Paratype: 1 W, angles angulate but at tip slightly obtuse. Marginal Austral. North. T., Berry Springs, XII.57, leg. H. Demarz channel narrow; median line very fine, impunctate. (CBM). Basal sulcus narrow. Anterior marginal seta situated at apical third. Basal groove barely indicated. Disk Type localities. “Katherine” and “Berry Springs”, both with some fine, irregularly transverse strioles in in northern part of Northern Territory. middle, barely punctate, with distinct, isodiametric Other material (12 ex.). NT: 2 MM, 1 W, Australia Dar- microreticulation and dull, or with only traces of win 20.VI.1994 (CBM, CHP); 1 W, N.T., A. N. Andersen microreticulation and rather glossy. Kakadu Nat. Park Kapalga fire exp’t forest comp’t 8P Elytra (Fig. 111). Soldered together. Short and 11.89 (ANIC); 1 W, N.T., A. N. Andersen Kakadu Nat. very compact, more or less narrow at base, rather Park Kapalga fire exp’t forest comp’t 15P 12.93 (ANIC); oval shaped, dorsally remarkably convex. Basal 2 WW, 12.40 S 132.54 E Jabiru NT 5-9- Jul. 1977 R.Pengilly angle little produced. Striation complete, even 7th pitfall trap (NTD); 1 W, AUSTRALIA: N.T. Casuarina stria deeply impressed. Striae very deep, apicad lit- Beach to Darwin W. D. Sumlin / William D. Sumlin tle shallower, more or less coarsely punctate, apicad III Entomological Expedition to Australia 1978-1979 (USNM); 1 W, 3.6.13 G. F. Hill Koolpinjah Darwin, N.T. increasingly impunctate. Intervals markedly convex, / Clivina hackeri Sl. Id. by T. G. Sloane (SAMA); 1 W, Pt. impunctate, without transverse strioles, with distinct, Darwin Australia / Clivina hackeri Sl. Id. by T. G. Sloane isodiametric microreticulation, dull. Setiferous punc- rd / Collectio Karel Kult Coll.A.Dostal,1989 (CDW); tures on 3 stria deep, foveate, rarely unilaterally 1 W, Pt. Darwin F.E. Dodd / Clivina hackeri Sl. / H. J. with 5 punctures. Epipleura narrow, basal sulcus Carter Coll. P. 20.4.22. (NMV COL-5327); 1 W, Pt. Darwin barely indicated.

267 Metathoracic wings. Reduced. W. D. Sumlin / William D. Sumlin III Entomological Lower surface. Proepisternum impunctate, with Expedition to Australia 1978-1979 (USNM); 1 W, West. some fine dorso-ventral strioles; with faint isodiamet- Austr. J. Clark (DEI); 1 W, Daly R. N.T. H. Wesselman ric microreticulation. The whole lower surface with (SAMA 25-033674). fine, isodiametric microreticulation. Metepisternum Etymology. The name refers to the very stout and short, as long as wide. Abdominal sterna impunctate. convex elytra. Legs. Protibia narrow and elongate. Longitu- Diagnosis. Rather small, very compact, black spe- dinal sulcus on upper surface deep; lower surface cies with wide and shallow, quadrangular clypeal with a row of 4 stout setae along basal part of inner excision, multistriolate head, 5- or 7-setose labrum, margin. Mesotibia rather short, dorsal rim crenulate, black legs, remarkably trapezoidal prothorax with with two rows of few setae on the dorsal surface; very oblique lateral margin, markedly striolate pro- subapical spur stout. Metatibia elongate, rather notum, impunctate proepisternum, short, convex, sparsely setose. Metatarsus elongate, metatarsomere very compact, oviform elytra, very deep, rather 1 slightly shorter than the following three tarsomeres. coarsely punctate elytral striae, wider and more Male genitalia (Fig. 44). Aedeagus wide, rather convex odd intervals, and wide, arrow-shaped apex short, asymmetrically narrowed apicad; lower of the aedeagus. surface even near base little concave, then almost straight, but apex suddenly turned down. Apical Description half of lower surface gently concave. Apex short, markedly club-shaped and denticulate at the lower Measurements. Length: 8.8-10.1 mm; width: 4.4- surface, club short and wide, at tip convex. Both 4.6 mm. Ratios. Length/width of pronotum: 0.86- parameres short, the left one considerably stouter 0.88; base/apex of pronotum: 1.57-1.70; width than the right one; both with very short, bisetose pronotum/head: 1.65-1.70; length/width of elytra: apical part; both setae situated right at apex, the 1.61-1.73; length/width of protibia: M: 2.75, W: 2.8- upper seta being longer than the lower one. 2.9; length/width of metatibia: M: 5.4, W: 5.1-5.2. Female gonocoxites (Fig. 74). Gonocoxites nar- Colour (Fig. 112). Unicolourous black. Legs row and elongate, little curved, with convex apex; black. with one elongate, stout seta basally at lateral surface Head (Fig. 112). Eye rather large, laterally well and a smaller seta below, two elongate setae on the projected, orbit short, oblique. Clypeus with wide ventral surface of gonocoxite 1, two setae at middle and very shallow, almost quadrangular excision, of the medio-dorsal surface, without a nematiform though the lateral margins of the excision oblique. seta near apex. Lateral plate basally with one elongate Labrum 7-setose, rarely 5-setose. Mandibles short. seta, apically with three or four setae. Clypeus divided from frons by a sulcus; upper sur- Variation. Considerably variation noted in body face rather convex, with transverse sulci and several size, shape of prothorax and elytra, degree of mi- irregular strioles, more or less sparsely punctate. croreticulation on the pronotum, punctation of the Surface with fine, slightly superficial, isodiametric elytral striae, and number of discal punctures. The microreticulation, moderately glossy. most distinctive difference is the dull resp. quite Pronotum (Fig. 112). Short and very wide, mark- glossy surface of the pronotum. Because in other edly narrow at apex, strongly narrowed apicad, dor- characters states the specimens do not significantly sally rather depressed. Lateral margin very oblique differ and because they occur in the same area, this but only slightly convex. Apex slightly concave. Basal is believed an intraspecific variation. angles angulate but at tip slightly obtuse. Marginal channel narrow; median line fine, impunctate. Basal Distribution. Northern part of NT, from Katherine sulcus narrow. Anterior marginal seta situated at or to Kakadu NP. behind apical third. Basal groove barely indicated. Disk with many irregularly transverse and ramified Collecting circumstances. Not recorded. strioles, barely punctate, with very distinct, isodiametric microreticulation, markedly dull. Elytra (Fig. 112). Soldered together. Short and Clivina crassipennis, spec. nov. very compact, moderately narrow at base, fairly oval Figs 45, 112 shaped, dorsally remarkably convex, particularly Examined types. Holotype: M, AUSTRALIA: n WA in apical half. Basal angle little produced. Striation th Kununurra 22.XII.1991-5.1.1992 R.I. Storey / Clivina sp. complete, even 7 stria deeply impressed. Striae very n. near hackeri Sl. det. B.P. Moore 1999 (QMT234907). deep, apicad little shallower, more or less coarsely – Paratype: 2 WW, same data (CBM, QDPIB); 1 W, AUS- punctate, apicad increasingly impunctate. Intervals TRALIA: W.A. 25.6kmE Kununurra 11 January 1979 markedly convex, odd intervals wider than the even

268 ones, and higher, near apex markedly raised. Apical Etymology. The name is a patronym and refers to the part remarkably precipitous. Intervals impunctate, collector, J. Horák. with or without fine transverse strioles, with fine, Diagnosis. Rather small, very compact, black spe- slightly superficial, isodiametric microreticulation, cies with wide and shallow, quadrangular clypeal rather glossy. Setiferous punctures on 3rd stria in- excision, slightly sulcate head, 7-setose labrum, black conspicuous. Epipleura narrow, basal sulcus barely legs, remarkably trapezoidal prothorax with very indicated. oblique lateral margin, impunctate proepisternum, Metathoracic wings. Reduced. short, remarkably convex, very compact, oviform Lower surface. Proepisternum impunctate, with elytra, and very deep, odd arranged, moderately some fine dorso-ventral strioles; with faint isodiamet- punctate elytral striae. ric microreticulation. The whole lower surface with fine, isodiametric microreticulation. Metepisternum Description short, as long as wide. Abdominal sterna impunctate. Legs. Protibia fairly narrow and elongate, simi- Measurements. Length: 8.4 mm; width: 2.9 mm. Ra- lar in both sexes. Longitudinal sulcus on upper tios. Length/width of pronotum: 0.87; base/apex of surface deep; lower surface with a row of 4-5 stout pronotum: 1.70; width pronotum/head: 1.57; length/ setae along basal part of inner margin. Mesotibia width of elytra: 1.67; length/width of protibia: W: 3.1; rather short, dorsal rim crenulate, with two rows length/width of metatibia: W: 6.4. of few setae on the dorsal surface; subapical spur Colour (Fig. 113). Unicolourous black. Legs stout. Metatibia fairly elongate, rather sparsely se- black, only tarsi rufous. tose. Metatarsus elongate, metatarsomere 1 slightly Head (Fig. 113). Eye large, laterally markedly shorter than the following three tarsomeres. projected, orbit very short, very oblique. Clypeus Male genitalia (Fig. 45). Aedeagus rather wide, with wide and very shallow, almost quadrangular fairly elongate, slightly asymmetrically narrowed excision, though the lateral margins of the excision apicad; lower surface in basal half gently concave, oblique. Labrum 7-setose, but the setae median of the then straight, but apical part suddenly curved down. lateral ones very short resp. even absent. Mandibles Apical half of lower surface gently concave. Apex short. Clypeus divided from frons by a sulcus; up- short, arrow-shaped, arrow wide and slightly asym- per surface rather convex, laterally with some fine metrically situated, at the lower surface denticulate, transverse sulci, rather sparsely punctate. Middle at tip convex. Both parameres elongate, the left one of frons with an oval-shaped impression. Surface considerably stouter than the right one; both with with traces of isodiametric microreticulation that is moderately elongate, narrow, bisetose apical part; more distinct on clypeus and anterior part of frons, both setae situated at the very apex, the upper one rather glossy. being slightly longer. Pronotum (Fig. 113). Short and very wide, Female gonocoxites. Ver similar to those of strongly narrowed apicad, dorsally very convex. C. demarzi Baehr. Lateral margin very oblique and moderately convex. Variation. Little variation noted in the specimens Apex concave. Marginal channel narrow; median line from northern WA. The female specimen from Daly very fine, impunctate. Basal sulcus narrow. Anterior River has a less triangular pronotum, less distinct marginal seta situated slightly in front of apical third. strioles on head and pronotum, less distinct micro- Basal groove very shallow, linear, impunctate. Disk reticulation on the pronotum, and less differently with few very fine, irregularly transverse strioles shaped odd elytral intervals. It is tentatively attrib- in middle, sparsely and extremely finely punctate, uted to the species. with fine and somewhat superficial, but distinct isodiametric microreticulation, moderately glossy. Distribution. North-eastern Kimberleys, WA, Elytra (Fig. 113). Soldered together. Short and north-western NT. compact, rather narrow at base, oval shaped, dorsally remarkably convex. Basal angle little produced. Stria- Collecting circumstances. Not recorded. tion complete, but 7th stria weak. Striae deep, apicad not shallower, moderately coarsely punctate. Inter- vals very convex, impunctate, without transverse Clivina horaki, spec. nov. strioles, without microreticulation, glossy. Striation Fig. 113 unique: 4th and 5th striae combined at basal two fifth, and interrupted. 6th stria oblique in apical half and Examined types. Holotype: W, WEST AUSTRALIA th th th 18km NNE of BROOME 17°46.98.1'S 122°17.16.2'E 17.- apparently becoming the 4 stria. 5 and 6 striae 26.I.2012 J.Horák leg. (WAM). again appearing in apical third, but united at their anterior and posterior ends. 3rd and 7th intervals

269 towards apex more convex that the other intervals, Sl. Id. by T. G. Sloane / Australia. Brit. Mus. 1921–125. the 7th interval enclosing all others at the very apex. (NHM); 1 M, 1 W, AUSTRALIA / Darwini Sl. det. K. Kult Epipleura narrow, basal sulcus coarsely punctate. / Collectio Karel Kult Coll.A.Dostal, 1999 (CDW). Metathoracic wings. Reduced. Diagnosis. Small, rufo-piceous species with very Lower surface. Proepisternum impunctate, with wide and shallow, clypeal excision, rather small some fine dorso-ventral strioles; with faint isodia- eye, barely sulcate head, 5-setose labrum, rufous metric microreticulation. The whole lower surface legs, depressed, apicad little narrowed prothorax with fine, isodiametric microreticulation. Metepis- with slightly oblique lateral margin, impunctate ternum short, about as long as wide. Abdominal proepisternum, fairly elongate, oviform elytra, deep, sterna impunctate. punctate elytral striae, glossy surface, and claw- Legs. Protibia narrow and elongate. Longitudi- shaped apex of the aedeagus. nal sulcus on upper surface fairly deep; lower surface with a row of 4 stout setae along basal part of inner Description margin. Mesotibia rather short, dorsal rim crenulate, with two rows of few setae on the dorsal surface; Measurements. Length: 5.7-6.5 mm; width: 1.65- subapical spur stout. Metatibia elongate, rather 1.85 mm. Ratios. Length/width of pronotum: 1.05- sparsely setose. Metatarsus elongate, metatarsomere 1.09; base/apex of pronotum: 1.27-1.29; width 1 slightly shorter than the following three tarsomeres. pronotum/head: 1.37-1.45; length/width of elytra: Male genitalia. Unknown. 1.86-1.94; length/width of protibia: M: 3.1-3.2, W: 2.9- Female gonocoxites. Very similar to those of 3.0; length/width of metatibia: M; 5.5-5.6, W: 5.25-5.3. C. demarzi Baehr. Colour (Fig. 114). Rufo-piceous. Legs rufous. Variation. Unknown. Head (Fig. 114). Eye comparatively small, later- 2 ally well projected, but orbit elongate, c. /5 of length Distribution. South-western Kimberley Division, of eye, oblique. Clypeus with very wide and shal- northern WA. low, in middle straight excision, though the lateral margins of the excision are very oblique. Labrum Collecting circumstances. Not recorded. 5-setose. Mandibles short. Clypeus divided from frons by a deep sulcus; upper surface rather convex, almost esulcate, sparsely punctate, without or with darwini subgroup fine traces of isodiametric microreticulation, glossy. Pronotum (Fig. 114). Fairly elongate, wide, wide Diagnosis. Small species with short, rather conical at apex, slightly narrowed apicad, dorsally rather prothorax and moderately elongate, oval-shaped depressed. Lateral margin slightly oblique and very elytra; clypeal excision wide and shallow; clypeus slightly convex. Apex concave. Marginal channel 5-setose; eye laterad little protruded; apex of aedea- narrow; median line fine but distinct, impunctate. gus claw-shaped: metathoracal wings reduced and Basal sulcus narrow. Anterior marginal seta situated elytra soldered together. The three species occur in at or behind apical fourth, according to length of northern NT. prothorax. Basal groove barely indicated. Disk with some fine, irregularly transverse strioles in middle, Clivina darwini Sloane sparsely and finely punctate, without microreticula- Figs 46, 75, 114 tion, glossy. Elytra (Fig. 114). Soldered together. Moderately Clivina darwini Sloane, 1916: 604. – Csiki 1927: 501; Moore short, rather wide at base, moderately oval shaped, et al. 1987: 69; Lorenz 2005: 142. dorsally convex but on disk slightly depressed. Ba- sal angle little produced. Striation complete. Striae Examined types. Holotype: fragment (not sexed, only deep, apicad barely shallower, moderately punctate, ’ hind body left), Pt. Darwin pond. (?) 08 / Clivina darwi- even apicad. Intervals convex, impunctate, without ni. Sl. Type / HOLOTYPE Clivina darwini Sl PJD (ANIC). transverse strioles, without microreticulation, glossy. Type locality. “Port Darwin”, Northern Territory. Epipleura narrow, basal sulcus barely perceptible. Metathoracic wings. Reduced. Other material (9 ex.). NT: 1 M, 1 W, 12.40S 132.00E NT Wildman R. Cashew Farm 21 Dec. 1989 W.Houston ex Lower surface. Proepisternum impunctate, with light intercept trap (CBM, ANIC); 1 M, N.T., A. N. Andersen some fine dorso-ventral strioles, with faint isodiamet- Kakadu Nat. Park Kapalga-A2 (11) 6-2-87 pitfall (ANIC); ric microreticulation. The whole lower surface with 1 W, N.T.O. Price Douglas Daly N.T. PWCNT Study Feb fine, isodiametric microreticulation. Metepisternum 1999 (ANIC); 2 MM (1 defect), Pt. Darwin W.Dodd 09 short, c. 1.25 × as long as wide. Abdominal sterna (ANIC); 1 W, Pt. Darwin W.Dodd 09 / Clivina darwini impunctate.

270 Legs. Protibia narrow and elongate, slightly B. P. Moore’86 (ANIC); 1 M, Burnside Stn. Brocks Creek less so in females. Longitudinal sulcus on upper N.T. 23/5/32 T- Campbell (ANIC); 1 W, Clivina macleayi surface deep; lower surface with a row of 4-5 stout Sl. N. Territory (ANIC-MMS). setae along basal part of inner margin. Mesotibia Diagnosis. Small, black species with very wide and rather short, dorsal rim crenulate, with two rows shallow, clypeal excision, strongly sulcate head, of few setae on the dorsal surface; subapical spur 5-setose labrum, black legs, short, moderately con- very elongate. Metatibia moderately elongate, rather vex, apicad little narrowed prothorax with slightly sparsely setose. Metatarsus moderately elongate, oblique lateral margin, impunctate proepisternum, metatarsomere 1 slightly shorter than the following rather short, slightly oviform elytra, deep, impunc- three tarsomeres. tate elytral striae, glossy surface, and claw-shaped Male genitalia (Fig. 46). Aedeagus very wide, apex of the aedeagus. fairly elongate, very asymmetrically narrowed apicad; lower surface in basal third gently concave, then Description almost straight, near apex suddenly angulate. Apical half of lower surface gently concave. Apical part Measurements. Length: 7.8-8.8 mm; width: 2.25- very narrow, remarkably deeply concave at the left 2.65 mm. Ratios. Length/width of pronotum: 0.97- side; apex elongate, directed left, at tip claw-shaped. 0.98; base/apex of pronotum: 1.25-1.33; width Both parameres very short and stout, the left one pronotum/head: 1.44-1.48; length/width of elytra: considerably stouter than the right one; both with 1.82-1.90; length/width of protibia: M: 3.25-3.3, very short, acute, bisetose apical part; setae situated W: 2.75-2.9; length/width of metatibia: M; 5.2, at the very apex, the upper seta being longer than the W: 5.0-5.1. lower one. Both parameres longitudinally striped. Colour (Fig. 115). Black. Legs dark piceous to Female gonocoxites (Fig. 75). Gonocoxites nar- black. row and elongate, little curved, with fairly acute Head (Fig. 115). Short and wide. Eye com- apex; with one elongate, stout seta basally at lateral paratively small, laterally well projected, but orbit 2 surface and a smaller seta below, two elongate setae elongate, c. /5 of length of eye, oblique. Clypeus with on the ventral surface of gonocoxite 1 and one seta wide and very shallow, in middle straight excision, in the middle of gonocoxite 2, four setae at middle of though the lateral margins of the excision are very the medio-dorsal surface, without a nematiform seta oblique. Labrum 5-setose. Mandibles short. Clypeus near apex. Lateral plate basally with one elongate divided from frons by a deep sulcus; upper surface seta, apically with one seta. rather convex, with markedly raised temporal ridge Variation. Some variation noted in length of and a deep sulcus inside, and a mediad and posteriad pronotum and elytra. incurved sulcus on either side, which meets in the middle in a longitudinal sulcus. Surface sparsely Distribution. Northern part of NT. punctate, without or with finest traces of isodiametric Collecting circumstances. Largely unrecorded, but microreticulation, glossy. two specimens were collected in light intercept trap, Pronotum (Fig. 115). Rather short, wide, wide one in pitfall trap. at apex, slightly narrowed apicad, dorsally gently convex. Lateral margin slightly oblique and very slightly convex. Apex concave. Marginal channel Clivina macleayi Sloane narrow; median line rather deep, impunctate. Basal Figs 47, 115 sulcus narrow. Anterior marginal seta situated at or slightly in front of apical third. Basal groove barely Clivina macleayi Sloane, 1896a: 236. – Sloane 1896a: 227; indicated. Disk with some fine, irregularly transverse 1896b: 279; Csiki 1927: 507; Moore et al. 1987: 72; strioles in middle, very sparsely, finely punctate, with Lorenz 2005: 143. more or less distinct, finest and superficial traces of Examined types. Lectotype (by present designation): isodiametric microreticulation, glossy. M, Roper Riv. / 33 / Clivina macleayi, Sl. / PARATYPE Elytra (Fig. 115). Soldered together. Moderately (blue) (ANIC). short, rather wide at base, moderately oval shaped, Type locality. “Roper River”, Northern Territory. dorsally convex but on disk slightly depressed. Ba- sal angle little produced. Striation complete. Striae Other material (5 ex.). NT: 2 WW (1 defect); E Al- deep, apicad barely shallower, impunctate. Intervals ligator Riv. NT 9.XII.82. Walford-Huggins / WAL- convex, impunctate, without transverse strioles, FORD-HUGGINS COLLECTION Carnegie Museum without or with very fine, superficial, isodiametric Accession 35338 / Clivina macleayi Sloane (Series det. by A. Walford-Huggins) (CMP); 1 W, E Alligator Riv. microreticulation, glossy. Epipleura narrow, basal NT 9.XII.82. Walford-Huggins / Clivina macleayi Sl. det. sulcus barely perceptible.

271 Metathoracic wings. Reduced. Diagnosis. Small, black species with very wide and Lower surface. Proepisternum impunctate, with shallow, clypeal excision, not sulcate head, 5-setose some fine dorso-ventral strioles; with faint isodiamet- labrum, black legs, moderately elongate, rather ric microreticulation. The whole lower surface with depressed, apicad rather narrowed prothorax with fine, isodiametric microreticulation. Metepisternum oblique lateral margin, impunctate proepisternum, short, slightly < 1.25 × as long as wide. Abdominal rather short, slightly oviform elytra, deep, impunc- sterna impunctate. tate elytral striae, glossy surface, and claw-shaped Legs. Protibia narrow and elongate, slightly less apex of the aedeagus. so in females. Longitudinal sulcus on upper surface deep; lower surface with a row of 5-6 stout setae Description along basal part of inner margin. Mesotibia rather short, dorsal rim strongly crenulate, with two rows Measurements. Length: 7.7-9.1 mm; width: 2.2- of few setae on the dorsal surface; subapical spur 2.6 mm. Ratios. Length/width of pronotum: 1.02- very elongate. Metatibia moderately elongate, rather 1.07; base/apex of pronotum: 1.32-1.40; width sparsely setose. Metatarsus moderately elongate, pronotum/head: 1.49-1.63; length/width of elytra: metatarsomere 1 slightly shorter than the following 1.89-1.91; length/width of protibia: M: 3.25-3.4, three tarsomeres. W: 2.7-3.05; length/width of metatibia: M: 5.35-5.5, Male genitalia (Fig. 47). Aedeagus moderately W:5.2-5.3. wide, rather elongate, slightly asymmetrically nar- Colour (Fig. 116). Black. Legs dark piceous to rowed apicad; lower surface near base concave, then black. straight, near apex suddenly angulate. Apical half of Head (Fig. 116). Short and wide. Eye com- lower surface gently concave. Apical part narrow, paratively small, laterally well projected, but orbit 2 almost regularly triangular; apex elongate, narrow, elongate, c. /5 of length of eye, oblique. Clypeus straight, but apicad directed left, at tip claw-shaped, with wide and very shallow, in middle straight tip acute. Both parameres moderately elongate, the excision, though the lateral margins of the excision left one considerably stouter than the right one; are very oblique. Labrum 5-setose. Mandibles short. both with very short, acute, bisetose apical part; Clypeus only laterally divided from frons by a shal- setae remarkably elongate; the longer seta situated low impression; upper surface rather convex, with at the very apex, the shorter one at the lower surface moderately raised temporal ridge, barely sulcate. near the apex. Surface sparsely punctate, without or with finest Female gonocoxites. Very similar to those of traces of isodiametric microreticulation, glossy. C. horneri, spec. nov. Pronotum (Fig. 116). Fairly elongate, wide, nar- Variation. Some variation noted in shape of row at apex, considerably narrowed apicad, dorsally pronotum, length of elytra, and degree of microre- gently convex. Lateral margin oblique, straight. ticulation on pronotum and elytra. Apex concave. Marginal channel narrow; median line rather deep, impunctate. Basal sulcus narrow. Distribution. Northern part of NT. Anterior marginal seta situated at or slightly in front of apical third. Basal groove barely indicated. Disk Collecting circumstances. Not recorded. with some fine, irregularly transverse strioles in middle, very sparsely, finely punctate, without or with finest and very superficial traces of isodiametric Clivina horneri, spec. nov. microreticulation, glossy. Figs 48, 76, 116 Elytra (Fig. 116). Soldered together. Moderately short, rather wide at base, moderately oval shaped, Examined types. Holotype: M, N.T. Murgenella 11.28S 132.51E 2-6 Feb. 1987 pitfall trap P. Horner / burnt dorsally convex but disk slightly depressed. Basal Eucalyptus tetradonta forest (NTD). – Paratypes: 3 MM, angle little produced. Striation complete. Striae 5 WW, same data (CBM, NTD); 3 WW, Murgenella 11.34S deep, apicad barely shallower, impunctate. Intervals 132.52E 3-7 Feb. 1987 pitfall trap P. Horner / burnt convex, impunctate, without transverse strioles, Eucalyptus tetradonta forest (NTD); 2 WW, Murgenella without or with very fine, superficial, isodiametric 11.34S 132.52E 30 Jan.-3 Feb. 1987 pitfall trap P. Horner microreticulation, glossy. Epipleura narrow, basal / unburnt Eucalyptus tetradonta forest (NTD); 1 W, 11.09 S sulcus barely perceptible. 132.09 E Black Point Coburg Pen. N.T. 15-23. Feb 1977 Metathoracic wings. Reduced. T.A. Weir (ANIC); 1 M (defect), 1 W, N.T., A. N. Kakadu Lower surface. Proepisternum impunctate, with Nat. Park Kapalea-B1 (10) 7-1-86 Pitfall (ANIC). some fine dorso-ventral strioles; with faint isodiamet- Etymology. The name is a patronym and refers to the ric microreticulation. The whole lower surface with collector, P. Horner. fine, isodiametric microreticulation. Metepisternum

272 short, slightly < 1.25 × as long as wide. Abdominal sulcaticeps group sterna impunctate. Legs. Protibia rather narrow and elongate, Clivina vixsulcata, spec. nov. slightly less so in females. Longitudinal sulcus on Figs 77, 133 upper surface rather shallow; lower surface with a row of 5-6 stout setae along basal part of inner Examined types. Holotype: W, AUS15, QLD32, More- margin. Mesotibia rather short, dorsal rim strongly head R. c. 100 km nw. Laura, 45m, 15°01'33.4"S, 143°40' crenulate, with two rows of few setae on the dorsal 01.1"E, 4.5.2015, M.Baehr (QMT234908). surface; subapical spur very elongate. Metatibia Diagnosis. Distinguished from the other species of moderately elongate, rather sparsely setose. Meta- the sulcaticeps group by combination of large body tarsus moderately elongate, metatarsomere 1 slightly size, short and wide prothorax, and lesser number shorter than the following three tarsomeres. of frontal sulci (c. 10). Male genitalia (Fig. 48). Aedeagus wide, rather elongate, very asymmetrically narrowed apicad; Etymology. The name refers to the comparatively low lower surface gently concave, near apex suddenly number of frontal sulci. angulate. Apical half of lower surface slightly concave. Apical part very narrow, remarkably deeply Description concave at the left side; apex elongate, triangular, Measurements. Length: 11.4 mm; width: 3.65 mm. markedly directed left, at tip claw-shaped, tip acute. Ratios. Length/width of pronotum: 1.18; base/ Both parameres very short and stout, the left one apex of pronotum: 1.36; width pronotum/head: considerably stouter than the right one; both with 1.37; length/width of elytra: 2.42; length/width of extremely short, acute, bisetose apical part; the longer protibia: 2.0; length/width of metatibia: 5.4. seta situated at the very apex, the shorter one at the Colour (Fig. 133). Black, labrum -piceous, palpi lower surface near the apex. and antenna rufous, lower surface almost black, Female gonocoxites (Fig. 76). Gonocoxites wide anterior leg black, median and posterior legs rufous. and short, little curved, with obtuse apex; with one Head (Fig. 133). Moderately narrow. Frons elongate, stout seta basally at lateral surface and a convex, upper surface of head behind clypeus smaller seta below, two elongate setae on the ventral with c. 10 moderately coarse, somewhat irregular, surface of gonocoxite 1, three setae at middle of the longitudinal ridges and sulci. Paraorbital ridge medio-dorsal surface, without a nematiform seta elongate and coarse, at posterior end even slightly near apex. Lateral plate basally with one elongate raised, anteriorly united with the sharp ridge on the seta, apically with one seta. supraantennal plate, but with shallow interruption. Variation. Some variation noted in body size, Paraorbital sulcus inside of paraorbital ridge elon- length of prothorax, and degree of microreticulation gate, deep, linear. Eye moderately large, laterad fairly of the surface. produced, orbit short, oblique. Clypeus gently concave, no interruption between central part and wing Distribution. Kakadu NP and Cobourg Peninsula, of clypeus visible, wing obtusely quadrangular but northernmost NT. slightly oblique, faintly projected, laterally slightly Collecting circumstances. Most specimens were oblique; with a slight incision between wing and collected in “pitfall trap / burnt Eucalyptus tetradonta supraantennal plate. Supraantennal plate laterally forest”. gently convex, with a conspicuous, sharp, elongate ridge. Crescent-shaped ridge on base of clypeus very distinct, wide, almost transverse, with a deep sulcus New species from groups mentioned behind. Laterally of ridge with a deep, semicircular in parts 1 and 2 of the revision furrow that separates the lateral plate behind from clypeus and anterior part of frons. Labrum straight, Some new species belonging to species groups men- 5-setose. Mandible elongate, acute, markedly curved, tioned in parts 1 and 2 of the revision, which could right mandible not carinate in middle. Mentum with not be included in those parts, are subsequently elongate, triangular, acute tooth. Base of mentum described. For these species the respective parts of 6-setose, setae on mentum very elongate. Antenna the key in part 2 are repeated and expanded. short, median antennomeres slightly wider than long. Upper surface impunctate, without visible microreticulation, surface moderately glossy. Pronotum (Fig. 133). Elongate, dorsally very convex, markedly narrowed apicad, decidedly

273 conical, but the very apex barely incurved. Lateral stout, dorsal rim strongly crenulate, with a row of margin barely incised at apical third. Basal angle 4-5 very stout setae in apical part, a series of setae completely rounded, without any interruption or along median surface, with elongate and very stout boss at angle. Apical angle distinctly projected, subapical spur, apex posteriorly slightly produced. acute, apex gently concave, somewhat v-shaped, Metatibia fairly elongate, rather sparsely setose, with not margined. Lateral margin distinct, lateral sulcus a row of c. 5 setae in apical half of lateral surface, anteriorly well developed, deep, becoming shallower without or with a single seta on median surface, and towards base. Anterior transverse sulcus and median with markedly elongate apical spur. Metatarsomere line both deeply impressed, anterior sulcus slightly 1 about as long as the following two tarsomeres. removed from apex. Basal sulcus narrow and rather Male genitalia. Unknown. shallow. Anterior marginal seta situated about at Female gonocoxites (Fig. 77). Gonocoxites nar- apical fifth, posterior marginal seta situated slightly row and elongate, little curved, with rather acute inside behind the basal angle. Basal groove oblong, apex; with one very elongate, fairly stout seta basally shallow but posteriorly deeper, slightly oblique. at lateral surface and a smaller seta below, two elon- Dorsal surface with finest traces only of extremely gate setae on the ventral surface, two elongate setae shallow transverse furrows, without perceptible at middle of medio-dorsal surface, but without a microreticulation, but with extremely, almost invis- nematiform seta near apex. Lateral plate basally with ible, sparse punctures. Proepisternum not visible one elongate seta, apically with four shorter setae. behind basal angle. Variation. Unknown. Elytra. Elongate and parallel-sided, dorsally convex, very gently oblique to the not produced Distribution. Lower Cape York Peninsula, north- basal angle. 4th stria meeting 5th stria by a short eastern QLD. Known only from type locality. transverse ridge. Striae complete, deep, punctate- Collecting circumstances. Holotype collected at crenulate. Intervals convex, very glossy, without light near the bank of Morehead River. any microreticulation, with extremely fine, almost invisible, very sparse punctures. 7th interval gently carinate behind humerus. 3rd stria 4-setose. Epipleura Revised key to the species of the narrow, basally with an elongate sulcus consisting sulcaticeps group of the genus Clivina of several coarse punctures. Lower surface. Proepisternum densely and 2. Body size smaller, < 9.5 mm; eye very depressed, coarsely punctate, also with some irregular, short laterad little produced...... 3a. transverse strioles, with distinct isodiametric microreticulation. Prosternum with dense though – Body size larger, > 11.4 mm; eye less depressed, slightly finer microreticulation, in particular laterally laterad distinctly produced (Fig. 133)...... 3. behind base rather rugose, in basal part also coarsely 3. Head with c. 10 sulci; pronotum with wider base, punctate. Prosternal process wide, not bordered, ratio width base/apex 1.36 (Fig. 133); n.QLD: intercoxal part fairly wide. Metepisternum very elon- CYP...... vixsulcata, spec. nov. gate, > 3 × as long as wide, in anterior half sparsely but coarsely punctate. Abdominal sterna with fine – Head with c. 14 sulci; pronotum with narrower microreticulation but laterally coarsely punctate. base, ratio width base/apex 1.10. ne.NT...... weanyanae Legs. Profemur elongate, upper margin little ...... Baehr, 2015 convex, lower surface depressed, posteriorly slightly 3a. Head and anterior half of pronotum not micro- carinate; posterior surface with two elongate ridges reticulate, surface glossy; elytra without traces and deep elongate sulci between. Protibia short and of microreticulation; size generally smaller, body very wide; irregularly 4-dentate though the upper length 6.5-8.4 mm; head with 8-12 sulci. n.NT. tooth only boss-like; teeth elongate in comparison ...... sulcaticeps Sloane, 1923 to tibia though clumsy; longitudinal sulcus on up- – Head and anterior half of pronotum finely, per surface elongate but shallow; lower surface de- distinctly microreticulate, surface dull; elytra pressed, smooth, but with isodiametric microreticu- with traces of microreticulation; size larger, body lation, with a longitudinal ridge basally near inner length 9.5 mm; head with c. 14 sulci. ne.QLD. . margin, and a very stout transverse ridge also near ...... densesulcata Baehr, 2008 inner margin in middle, and with a row of 6 stout setae along basal part of inner margin. Mesotibia

274 emarginata group frons, frons with a distinct, circular to slightly longitudinal median fovea, also with a lateral, slightly semirubra subgroup incurved sulcus. Surface of frons gently raised and almost even. Clypeal suture, median fovea and lat- Diagnosis. Distinguished from other subgroups of eral sulcus coarsely punctate, posterior part of frons the emarginata group by combination of lobate but not more sparsely punctate. microreticulation absent, dentate posterior margin of profemur, narrow basal surface glossy. sulcus of pronotum, short, dorsally convex protho- Pronotum (Fig. 117). Moderately elongate, wide, rax, and punctate head; and from the most similar laterally parallel, dorsally convex, not narrowed api- lobipes subgroup by wide head, 5-setose labrum, cad. Lateral margin straight, usually slightly incised very parallel-sided prothorax, shorter metatibia, and at apical third. Basal angle with a very slight incision spatulate, laterally dentate apex of the aedeagus. and a faint protuberance. Apical angle not produced, Distribution. The single species C. semirubra, spec. apex almost straight, not margined. Anterior trans- nov. from north-western Queensland. verse sulcus and median line deeply impressed, impunctate. Basal sulcus narrow. Anterior marginal seta situated at very close to apex. Basal groove Clivina semirubra, spec. nov. short, rather shallow, somewhat irregular, coarsely Figs 49, 117 punctate. Disk on either side coarsely punctate in a longitudinal stripe; without microreticulation except Examined types. Holotype: M, AUS15, QLD13, Flin- near base, glossy. ders R. 37 km wsw. Normanton, 12 m, 17°52'35.4"S, Elytra (Fig. 117). Elongate, parallel-sided, barely 140°47'06.2"E, 18.4.2015, M.Baehr (QMT234909). – Para- widened apicad, convex but dorsally somewhat types: 3 MM, 3 WW, same data (CBM, CBP, CDW). depressed, basal angle not produced. Striae deep, Diagnosis. Fairly large, elongate and parallel-sided, barely weaker laterally and towards apex, coarsely dorsally moderately convex species, with black head punctate-crenulate. Intervals convex and with fine and pronotum but rufous elytra, 5-setose labrum, transverse wrinkles, impunctate, without microre- and spatulate, laterally dentate apex of the aedeagus. ticulation, glossy. Epipleura narrow, basally with few punctures or a shallow sulcus. Etymology. The name refers to the colouration: dark Lower surface. Proepisternum impunctate, with head and pronotum, rufous elytra. many coarse transverse strioles, with distinct, rugose, Description isodiametric microreticulation, dull. Prosternum microreticulate but almost impunctate, in anterior Measurements. Length: 6.9-7.7 mm; width: 1.75- part rugose. Metasternum with more or less dense 2.0 mm. Ratios. Length/width of pronotum: 1.05- transverse strioles. Metepisternum elongate, > 2.5 × 1.07; base/apex of pronotum: 1.10-1.13; width as long as wide, with a shallow, longitudinal, punc- pronotum/head: 1.15-1.19; length/width of elytra: tate furrow and some transverse strioles. Abdominal 2.14-2.19; length/width of protibia: 2.45-2.6; length/ sterna impunctate, with isodiametric microreticula- width of metatibia: 5.2-5.4. tion which is more superficial at apex. Colour (Fig. 117). Head and pronotum black, Legs. Profemur stout, moderately wide, upper elytra more or less dark rufous. Palpi and antenna margin convex, lower surface depressed, with an- more or less pale rufous, anterior leg dark piceous, gulate anterior margin; posterior margin well pro- median and posterior legs rufous, but tibiae slightly duced and evenly convex in middle. Protibia wide, darker than femora. 4-dentate but 4th tooth small, other teeth elongate; Head (Fig. 117). Wide, dorsally gently convex, longitudinal sulcus on upper surface distinct. Metati- quite even. Frontal sulcus deep, sinuate, punctate. bia moderately elongate. 1st tarsomere of metatarsus Eye large, laterally well projected, orbit very short, shorter than the following three tarsomeres. forming an almost right angle with neck. Labrum Male genitalia (Fig. 49). Aedeagus rather wide, 5-setose. Clypeus in middle gently concave, coarsely moderately elongate, asymmetrically narrowed margined, with distinct though obtuse lateral tooth. apicad; lower surface concave throughout. Apex Wing of clypeus distinctly separated from centre, rather wide, spatulate and obtusely arrow-shaped, moderately protruded, obtusely quadrangular, not laterally and at its lower surface dentate, tip obtusely surpassing the clypeal tooth, bearing a shallow rounded. Internal sac in apical third with three incision between wing and supraantennal plate. large, denticulate and rather sclerotized folds. Both Crescent-shaped ridge on clypeus shallow. Antenna parameres large and stout, the left one considerably short, median antennomeres slightly wider than stouter than the right one; both with short but acute, long. Clypeus more or less distinctly divided from asetose apex.

275 Female gonocoxites. Very similar to those of base/apex of pronotum: 1.24; width pronotum/ C. punctaticeps Putzeys, see fig. 157 in Baehr (2015). head: 1.23-1.27; length/width of elytra: 1.98-2.0; Variation. Little variation noted, apart from length/width of protibia: 2.6-2.7; length/width of some variation of density of punctation of head metatibia: 5.4-5.5. and pronotum. Colour (Fig. 118). Upper surface rufous, posterior part of head in the holotype slightly darker; Distribution. North-western QLD. Known only elytra with a sutural spot which covers three median from type locality. intervals. Palpi and antenna pale red, legs yellow, the anterior leg slightly darker. Under surface dirty Collecting circumstances. Collected at light at the yellow to red. bank of Flinders River. Head (Fig. 118). Rather short and wide, dorsally gently convex, slightly uneven. Frontal sulcus rather deep, sinuate, barely punctate. Eye large, laterally Recognition well projected, orbit short, almost perpendicular. Clypeus in middle almost straight, coarsely mar- According to shape of profemur and narrow basal gined, with rather short, obtuse lateral tooth that sulcus of the pronotum couplet 55. in the key in Baehr barely surpasses the wing. Wing of clypeus well (2015) is reached, which has to be changed as below: separated from centre, rounded, bearing a distinct 55. Labrum 5-setose; body size large, length > 6.9 mm incision between wing and supraantennal plate. and elytra elongate, ratio l/w > 2.14 and elytra Crescent-shaped ridge on clypeus barely indicated. without dark spot and aedeagus with spatulate, Antenna short, median antennomeres globular, laterally dentate apex (Fig. 49). nw.QLD...... slightly wider than long. Clypeus barely divided ...... semirubra, spec. nov. from frons, frons with an about circular, fairly deep – Labrum 7-setose, not all other character states median fovea, surface of frons gently raised, even. present together...... 55a. Surface sparsely punctate in and around the frontal fovea, punctures moderately coarse, microreticula- 55a. = 55. in Baehr (2015). tion absent, surface glossy. Pronotum (Fig. 118). Short and wide, slightly narrowed apicad, lateral margins straight but faintly heterogena group oblique, barely incised at apical third, dorsal surface rather convex. Basal angle with distinct protuber- oodnadattae subgroup ance and incision. Apical angle little produced, apex almost straight, not margined. Anterior transverse Clivina infans, spec. nov. sulcus well impressed, situated close to apex, im- Figs 50, 118 punctate. Median line fairly deep, barely crenulate. Basal sulcus narrow. Anterior marginal seta situ- Examined types. Holotype: M, AUS15, QLD13, Flin- ated at apical fourth. Basal groove very shallow, ders R. 37 km wsw. Normanton, 12 m, 17°52'35.4"S, short. Dorsal surface in middle with several fine, 140°47'06.2"E, 18.4.2015, M.Baehr (QMT234910). – Para- irregular, transverse strioles, impunctate, with fine, type: 1 M, same data (CBM). rather superficial, isodiametric microreticulation, Diagnosis. Small, rufous species with dark median moderately glossy. elytral spot, large, protruded eye and short orbit, Elytra (Fig. 118). Rather short, barely widened short, faintly microreticulate pronotum and elytra, apicad, convex but dorsally depressed, slightly and very narrow apical part of the aedeagus and incurved to the not produced basal angle. Lateral small, hook-shaped apex. Distinguished from other margin in basal fourth barely concave. Striae deep, species of the group by small size and structure of barely weaker laterally and towards apex, rather the aedeagus. Distinguished from C. communis Baehr, coarsely punctate. Intervals convex, with sparse, fine 2015 which has a quite similar aedeagus, by lesser transverse wrinkles, impunctate, with fine, superfi- size and presence of microreticulation. cial, isodiametric microreticulation, rather glossy. Epipleura basally with some coarse punctures. Etymology. The name refers to the small size of the Lower surface. Proepisternum impunctate, with species as compared with its nearest relatives. rather fine transverse strioles, with distinct isodia- Description metric microreticulation, dull. Prosternum finely microreticulate but almost impunctate, in anterior part Measurements. Length: 4.2-4.3 mm; width: 1.2- slightly rugose. Metasternum with fine transverse 1.25 mm. Ratios. Length/width of pronotum: 1.0; strioles. Metepisternum elongate, c. 2 × as long as

276 wide, with a slightly punctate furrow. Abdominal – Body size < 3.35 mm; apex of aedeagus, when sterna impunctate, with isodiametric microreticula- known, not triangular...... 156. tion which is more superficial at apex. 122. Eye depressed with distinct, oblique orbit (B15 Legs. Profemur fairly stout, moderately wide, fig. 295); pronotum and elytra usually not upper margin convex, lower surface depressed, microreticulate; if pronotum indistinctly mi- with angulate anterior margin. Protibia moderately croreticulate, elytra strongly microreticulate elongate; 4-dentate but 4th tooth small, other teeth (C. mareebae Baehr, 2015); if elytra slightly elongate; longitudinal sulcus on upper surface shal- microreticulate, elytra < 2 × as long as wide low. Metatibia fairly elongate. 1st tarsomere of meta- (C. baloghi Baehr, 2015) (questionable species tarsus shorter than the following three tarsomeres. under both couplets)...... 123. Male genitalia (Fig. 50). Aedeagus rather narrow, fairly elongate, strongly narrowed apicad; lower – Eye laterad markedly protruded, orbit short, surface gently but evenly concave. Apical part almost perpendicular or almost so (Fig. 118); micro- symmetric; apex straight, elongate, very narrow, reticulation of pronotum and elytra various. . slightly widened near tip, shortly hooked. Internal ...... 137. sac in apical third with three large, denticulate and 137. Prothorax short, ratio l/w < 1.0...... 138. rather sclerotized folds. Both parameres short and very stout, the left one considerably stouter than the – Pronotum longer, ratio l/w always > 1.0; if narrower right one; both with extremely short, at tip ratio but slightly > 1.0 head barely punctate acute acute, asetose apical part. and/or pronotum microreticulate; aedeagus Female gonocoxites. Unknown. various. not in c.WA...... 139. Variation. Apart from some variation of colour 138. Prothorax apicad distinctly narrowed; head in which partly may be due to age of specimens, little and around frontal fovea coarsely punctate, variation noted. eye large, but orbit distinct and slightly oblique (B15 fig. 367); pronotum and elytra not micro- Distribution. North-western QLD. Known only reticulate; apex of aedeagus short, convex, from type locality. laterally distinctly denticulate (B15 fig. 65). Collecting circumstances. Collected at light at the c.WA...... trapezicollis Baehr, 2015 bank of Flinders River. – Prothorax apicad barely narrowed; head only in front of frontal fovea with some coarse punctures, eye large with short orbit (Fig. 118); Recognition pronotum and elytra microreticulate; aedeagus with very narrow apical part, and narrow and According to body size, length of elytra, presence hook-shaped apex (Fig. 50), or aedeagus un- of microreticulation on pronotum and elytra, and known...... 138a. shape of aedeagus couplet 120 in the key in Baehr 138a. Pronotum coarsely punctate; aedeagus un- (2015) is reached which has to be changed as below: known. ne.QLD: Atherton Tableland...... 120. Comparatively small species, body length ...... quadraticollis Baehr, 2015 usually < 4.5 mm, when larger, either eye lat- – Pronotum impunctate; aedeagus with very erad little protruded and orbit distinct and narrow apical part and narrow and hook- oblique, or pronotum and elytra microreticu- shaped apex (Fig. 50). nw.QLD...... late and apex of aedeagus unarmed (when ...... infans, spec. nov. doubtful, try both couplets); elytra commonly rather depressed...... 121. – Usually larger, less depressed species, body australica subgroup length usually > 5 mm; eye laterad protruded and orbit very short, perpendicular, or if orbit Clivina gerstmeieri planior, subspec. nov. more or less oblique, body size large, >5.3 mm; Examined types. Holotype: W, AUS15, QLD11, Nor- apex of aedeagus almost always lancet-shaped man R. 20 km sse. Normanton, 17m, 17°51'17.4"S, or denticulate...... 164. 141°08'12.5"E, 16.4.2015, M.Baehr (QMT234911). 121. Body size usually > 3.7 mm, if slightly less Diagnosis. Distinguished from all other subspecies (3.5 mm), either eye depressed and orbit dis- of C. gerstmeieri Baehr by combination of absolutely tinct or apex of aedeagus triangular...... 122. glabrous and at the same time rather intersected elytral intervals, and depressed pronotum and elytra.

277 Etymology. The name refers to the pronotum and elytra 107a. Pronotum and elytra dorsally convex; elytral which are more depressed than in all other subspecies intervals barely intersected. Interior of SA, of C. gerstmeieri. s.NSW, sc.NT, sw.QLD...... gerstmeieri laevior Baehr, 2015 Description – Pronotum and elytra dorsally rather depressed; Measurements. Length: 4.7 mm; width: 1.15 mm. Ra- elytral intervals rather intersected. nw.QLD. . tios. Length/width of pronotum: 1.16; base/apex of ...... gerstmeieri planior, subspec. nov. pronotum: 1.15; width pronotum/head: 1.25; length/ 108. Pronotum slightly more depressed; colour dark width of elytra: 2.25; length/width of protibia: 2.7; piceous; eye slightly smaller and laterad less length/width of metatibia: 5.15. protruded, ratio width of pronotum/head Colour. Uniformly rufo-piceous. > 1.23; frontal fovea large and deep, corru- Head. Much as in the nominate subspecies, but gated, laterally of fovea with a deep sulcus on eye comparatively well produced laterad. Median either side. sw.WA ...... frontal fovea rather deep, punctate...... gerstmeieri newnhamensis Baehr, 2015 Pronotum. Much as in the nominate subspecies, but comparatively elongate and more depressed. – Pronotum slightly more convex; colour black; Elytra. Much as in the nominate subspecies, eye slightly larger and laterad more protruded, but dorsally less convex and slightly longer and ratio width of pronotum/head < 1.23; frontal narrower. Intervals without any traces of microre- fovea usually shallow, not corrugated, without ticulation, rather intersected, very glossy. or with less deep lateral sulcus. ne.WA: KID . Lower surface. As in the nominate subspecies...... gerstmeieri septemtrionalis Baehr, 2015 Legs. Much as in the nominate subspecies. Metatibia comparatively short. subgroup Male genitalia. Unknown. atridorsis Female gonocoxites. As in other subspecies, see Clivina moretona, spec. nov. fig. 211 in Baehr (2015). Variation. Unknown. Examined types. Holotype: W, AUS15, QLD35, Wenlock R.at Moreton Tel. Stn., 156m, 12°27'36.0"S, 142°38'37.2"E, Distribution. North-western QLD. Known only 9.5.2015, M.Baehr (QMT234912). – Paratypes: 2 WW, same from type locality. data (CBM). Collecting circumstances. Holotype collected at Diagnosis. Small, rufous species, characterized by light at the bank of Normanton River. rather elongate, parallel-sided prothorax, rather elongate elytra, and glossy, not microreticulate surface.

Recognition Etymology. The name refers to the type locality, More- ton Telephone Station at Wenlock River. Revised key to the subspecies of C. gerstmeieri: Description 106. Elytra with distinct microreticulation, intervals not much intersected, at the average slightly Measurements. Length: 3.5-3.6 mm; width: 0.95- shorter, ratio l/w 2.09-2.17. wc.NT, c.WA, in 1.0 mm. Ratios. Length/width of pronotum: 1.10- the north to the Pilbara ...... 1.13; base/apex of pronotum: 1.18-1.21; width ...... gerstmeieri gerstmeieri Baehr, 1989 pronotum/head: 1.28-1.32; length/width of elytra: 2.08-2.12; length/width of protibia: 2.45-2.65; – Elytra without or with indistinct microreticu- length/width of metatibia: 5.5-5.7. lation, intervals either much intersected, or Colour. Head and pronotum uniformly rufous, glabrous, at the average slightly longer, ratio elytra very slightly paler. Palpi, antenna, and legs l/w 2.15-2.31. interior of SA, sc.NT, ne.WA: pale red, the anterior leg usually very slightly darker. KID, sw.WA, nw.QLD...... 107. Under surface rufous. 107. Elytral intervals without any traces of micro- Head. Of average size, dorsally gently con- reticulation. Interior of SA, s.NSW, sc.NT, vex. Frontal sulcus deep, sinuate, impunctate. Eye sw.QLD, nw.QLD...... 107a. large, laterally well projected, orbit short, slightly oblique. Clypeus in middle gently concave, coarsely – Elytral intervals usually with fine traces of margined, with extremely short, obtuse lateral microreticulation, rather intersected. ne.WA: tooth. Wing of clypeus shallowly separated from KID, sw.WA...... 108. centre, moderately protruded, rounded, bearing a

278 shallow incision between wing and supraantennal Distribution. North QLD, north Cape York Penin- plate. Crescent-shaped ridge on clypeus indistinct. sula. Known only from type locality. Antenna short, median antennomeres globular, Collecting circumstances. Collected at light at the slightly wider than long. Clypeus not divided from bank of Wenlock River. frons. Frons with a more or less distinct, slightly oblong median fovea, surface of frons almost even. Posterior part of clypeus, and frons around the fo- Recognition vea with sparse, mixed fine and coarser punctures, microreticulation absent, surface glossy. According to shape of clypeus and to body size, Pronotum. Rather elongate, barely narrowed couplet 184 in the key in Baehr (2015) is reached apicad, almost parallel-sided, lateral margin straight, which has to be changed as below: barely incised at apical third, dorsal surface convex. Basal angle not perceptibly incised, without protu- 184. Body size very small, < 3.6 mm...... 185. berance. Apical angle little produced, apex almost – Body size larger, > 4.0 mm, commonly larg- straight, not margined. Anterior transverse sulcus er...... 191. well impressed, situated close to apex, barely crenulate. Median line deep. Basal sulcus narrow. Anterior 185. Prothorax longer and dorsally convex, ratio marginal seta situated about at apical fifth. Basal l/w > 1.07; aedeagus without many extremely groove shallow, linear, sparsely punctate. Dorsal elongate spines (B15 figs 121, 143), or with surface with very few inconspicuous, transverse elongate spines only at apex (B15 fig. 142). .... strioles, with sparse to fairly dense, fine punctures, ...... 186. without microreticulation except at the very base, – Prothorax shorter, markedly depressed, ratio very glossy. l/w < 1.05; aedeagus with many extremely Elytra. Rather elongate, parallel, barely wid- elongate spines (B15 figs 144, 145)...... 188. ened apicad, convex but dorsally depressed, basal angle not produced. Lateral margin in basal 186. Prothorax longer, ratio lw > 1.10, and more fourth barely concave. Striae well impressed, barely parallel-sided, ratio width base/apex < 1.21; weaker laterally and towards apex, fairly coarsely elytra longer, ratio l/w > 2.08; aedeagus at apex punctate-crenulate. Intervals slightly convex, with with many rather elongate spines (B15 fig. 142), few transverse wrinkles, impunctate, without mi- or eadeagus unknown...... 186a. croreticulation, glossy. Epipleura narrow, basally – Prothorax shorter, ratio lw < 1.09, and less with few punctures. parallel-sided, ratio width base/apex > 1.29; Lower surface. Proepisternum impunctate, with elytra shorter, ratio l/w < 2.06; aedeagus with- rather coarse transverse strioles, with distinct iso- out elongate spines (B15 figs 121, 143)...... diametric microreticulation, dull. Prosternum finely ...... 187. microreticulate, in anterior part more or less rugose. 186a. Body size smaller, length 3.0 mm; elytra Metasternum with more or less distinct transverse longer, ratio l/w 2.14; aedeagus at apex with strioles. Metepisternum elongate, slightly < 2.25 × as many rather elongate spines (B15 fig. 142). long as wide, with a more or less distinct, longitudi- n.NT...... multispinosa Baehr, 2015 nal, punctate furrow. Abdominal sterna impunctate, with isodiametric microreticulation which is more – Body size larger, length > 3.5 mm; elytra superficial at apex. shorter, ratio l/w < 2.12; aedeagus unknown. Legs. Profemur fairly stout, moderately wide, ne.QLD. n.CYP...... moretona, spec. nov. upper margin convex, lower surface depressed, 187. Elytra apicad widened and dorsally rather with angulate anterior margin. Protibia rather short depressed; prothorax slightly longer, ratio l/w th and wide; 4-dentate but 4 tooth small, other teeth 1.11; aedeagus with laterally distinctly dentate elongate; longitudinal sulcus on upper surface dis- apex (B15 fig. 143). nc. NT...... st tinct. Metatibia elongate. 1 tarsomere of metatarsus ...... hovorkai Baehr, 2015 slightly shorter than the following three tarsomeres. Male genitalia. Unknown. – Elytra rather parallel-sided and dorsally con- Female gonocoxites. Very similar to those of vex; prothorax slightly shorter, ratio l/w < 1.09; C. tuberculiforns Blackburn, see fig. 219 in Baehr aedeagus with laterally obtuse apex (Fig. 121). (2015). n.QLD, n.NT, n.WA...... Variation. Little variation noted only in density ...... tuberculifrons Blackburn, 1890 of punctures on the pronotum.

279 Clivina laevigata, spec. nov. slightly narrowed apicad, lateral margin straight Figs 51, 119 but slightly oblique, slightly incised at apical third, dorsal surface moderately convex. Basal angle with a Examined types. Holotype: M, AUS15, QLD44, More- slight incision and a faint protuberance. Apical angle head R. c. 100 km nw. Laura, 45m, 15°01'33.4"S, 143°40' not or very little produced, apex almost straight, 01.1"E, 14.5.2015, M.Baehr (QMT234913). – Paratypes: not margined. Anterior transverse sulcus well 3 MM, 5 WW, same data (ANIC, CBM, CBP); 2 MM, 6 WW, impressed, situated close to apex, barely crenulate. AUS15, QLD39, Wenlock R., 13 km ene. T/0 →Iron Median line deep. Basal sulcus narrow. Anterior Range, 13°05'40.6"S, 142°56'45.3"E, 126m,11.5.2015, marginal seta situated about at apical fourth. Basal M.Baehr (CBM, CDW); 1 M, 1 W, AUS15, QLD4, Walsh R., 1 km e. Mutchilba, 462m, 17°08'21.5"S, 145°13'58.4"E, groove very shallow, linear, impunctate. Dorsal 12.4.2015, M.Baehr (CBM); 1 W, AUS15, QLD31, Ken- surface with a few shallow, irregularly transverse nedy R., 14 km nw. Fairview, 105m, 15°25'19.9"S, strioles, with very sparse, fine punctures, without 144°11'16.9"E, 3.+6.5.2015, M. Baehr (CBM); 1 W, AUS15, microreticulation except in basal third, glossy. QLD41, Myall Ck., 49 km e. Weipa, 21m, 12°39'16.7"S, Elytra (Fig. 119). Moderately elongate, paral- 142°16'31.5"E, 12.5.2015, M. Baehr (CBM). lel, barely widened apicad, convex but dorsally depressed, basal angle not produced. Lateral margin Diagnosis. Rather small to medium sized, uni- in basal fourth barely concave. Striae well impressed, colourous rufous species, characterized by fairly barely weaker laterally and towards apex, fairly elongate prothorax, glossy, not microreticulate head coarsely punctate-crenulate. Intervals slightly convex and pronotum, and large, at tip obtuse, at the lower and with some transverse wrinkles, impunctate, surface strongly hook-shaped apex of the aedeagus. with very faint and superficial, about isodiametric Etymology. The name refers to the glabrous head and microreticulation, rather glossy. Epipleura narrow, pronotum. basally with some punctures. Lower surface. Proepisternum impunctate, with Description rather fine transverse strioles, with distinct isodiametric microreticulation, dull. Prosternum finely Measurements. Length: 4.6-5.3 mm; width: 1.3- microreticulate, in anterior part more or less rugose. 1.45 mm. Ratios. Length/width of pronotum: 1.08- Metasternum with more or less distinct transverse 1.12; base/apex of pronotum: 1.29-1.34; width strioles. Metepisternum elongate, slightly < 2.25 × as pronotum/head: 1.32-1.34; length/width of elytra: long as wide, with a more or less distinct, longitudi- 2.0-2.05; length/width of protibia: 1.08-1.12; length/ nal, punctate furrow. Abdominal sterna impunctate, width of metatibia: 5.5-5.7. with isodiametric microreticulation which is more Colour (Fig. 119). More or less pale rufous, superficial at apex. elytra unicolourous. Palpi, antenna, and legs pale Legs. Profemur fairly stout, moderately wide, red, the anterior leg usually very slightly darker. upper margin convex, lower surface depressed, Under surface rufous. with angulate anterior margin. Protibia rather short Head (Fig. 119). Of average size, dorsally gently and wide; 4-dentate but 4th tooth small, other teeth convex, slightly uneven. Frontal sulcus deep, sinu- elongate; longitudinal sulcus on upper surface dis- ate, impunctate. Eye large, laterally well projected, tinct. Metatibia elongate. 1st tarsomere of metatarsus orbit short, slightly oblique. Clypeus in middle almost as long as the following three tarsomeres. gently concave, coarsely margined, with extremely Male genitalia (Fig. 51). Aedeagus rather nar- short, obtuse lateral tooth. Wing of clypeus shal- row, moderately elongate, almost symmetrically lowly separated from centre, moderately protruded, narrowed apicad; lower surface straight over most rounded, bearing a shallow incision between wing of its length, only near apex curved down. Apex and supraantennal plate. Crescent-shaped ridge rather elongate and wide, spatulate, at its lower on clypeus fairly distinct. Antenna short, median surface strongly dentate, tip rounded. Internal sac in antennomeres globular, slightly wider than long. apical third with three large, denticulate and rather Clypeus indistinctly divided from frons by a shal- sclerotized folds. Both parameres large and stout, low, in middle usually interrupted sulcus; frons with the left one considerably stouter than the right one; a distinct, about circular to slightly oblong median both with very short but acute, asetose apex. fovea, surface of frons almost even. Posterior part of Female gonocoxites. Very similar to those of clypeus, clypeal suture, and frons around the fovea C. tuberculiforns Blackburn, see fig. 219 in Baehr sparsely to moderately densely, fairly coarsely punc- (2015). tate, rest of surface with sparse and fine punctures, Variation. Little variation noted in body size, microreticulation absent, surface glossy. length of prothorax, and degree of microreticulation Pronotum (Fig. 119). Moderately elongate, in basal part of the pronotum.

280 Distribution. North QLD, from Atherton Tableland sulcus; frons with a distinct, about circular to slightly to mid Cape York Peninsula. oblong median fovea, surface of frons almost even. Posterior part of clypeus, clypeal suture, and frons Collecting circumstances. All specimens collected around the fovea rather densely, very coarsely punc- at light, at or near the banks of rivers. tate, rest of surface with sparse and fine punctures, microreticulation distinct though slightly superficial, isodiametric, surface moderately glossy. Clivina uncinata, spec. nov. Pronotum (Fig. 120). Moderately elongate, Figs 52, 120 slightly narrowed apicad, lateral margin straight but slightly oblique, slightly incised at apical third, Examined types. Holotype: M, AUS15, QLD44, More- dorsal surface moderately convex. Basal angle with a head R. c. 100 km nw. Laura, 45m, 15°01'33.4"S, 143°40' slight incision and a faint protuberance. Apical angle 01.1"E, 14.5.2015, M.Baehr (QMT234914). – Paratypes: not or very little produced, apex almost straight, 1 M, 2 WW, same data (CBM, CBP, CDW); 1 M, AUS15, QLD39, Wenlock R., 13 km ene. T/0 →Iron Range, not margined. Anterior transverse sulcus well 13°05'40.6"S, 142°56'45.3"E, 126m,11.5.2015, M.Baehr impressed, situated close to apex, barely crenulate. (CBM); 1 M, 1 W, AUS15, QLD31, Kennedy R., 14 km nw. Median line deep. Basal sulcus narrow. Anterior Fairview, 105m, 15°25'19.9"S, 144°11'16.9"E, 3.+6.5.2015, marginal seta situated about at apical fourth. Basal M. Baehr (CBM); 2 WW, AUS15, QLD41, Myall Ck., 49 km groove very shallow, linear, impunctate. Dorsal e. Weipa, 21m, 12°39'16.7"S, 142°16'31.5"E, 12.5.2015, surface with several shallow, irregularly transverse M. Baehr (CBM); 1 M, AUS15, QLD36, c.25 km ene. strioles, with very sparse, fine punctures, with Bramwell Jct. Rdh., 115m, 11°56'21.9"S, 142°38'07.2"E, distinct, isodiametric microreticulation, rather dull. 10.5.2015, M.Baehr (CBM); 1 W, AUS15, QLD42, Coen Elytra (Fig. 120). Rather short, almost paral- R., 2 km n. Coen, 213m, 13°55'30.9"S, 143°11'48.6"E, lel, slightly widened apicad, convex but dorsally 13.5.2015, M. Baehr (CBM). depressed, basal angle not produced. Lateral mar- Diagnosis. Rather small, unicolourous rufous spe- gin in basal fourth barely concave. Striae well im- cies, characterized by moderately elongate protho- pressed, barely weaker laterally and towards apex, rax, microreticulate head and pronotum, strongly moderately coarsely punctate-crenulate. Intervals microreticulate elytra, and rather small but strongly slightly convex and with some transverse wrin- hook-shaped apex of the aedeagus. kles, impunctate, with rather distinct but slightly superficial, about isodiametric microreticulation, Etymology. The name refers to the strongly hook- moderately glossy. Epipleura narrow, basally with shaped apex of the aedeagus. some punctures. Lower surface. Proepisternum impunctate, Description with rather fine transverse strioles, with distinct iso- Measurements. Length: 4.4-4.7 mm; width: 1.25- diametric microreticulation, dull. Prosternum finely 1.35 mm. Ratios. Length/width of pronotum: 1.06- microreticulate, in anterior slightly more rugose. 1.08; base/apex of pronotum: 1.19-1.23; width Metasternum without distinct transverse strioles. pronotum/head: 1.28-1.32; length/width of elytra: Metepisternum elongate, slightly > 2 × as long as 0.98-2.03; length/width of protibia: 1.06-1.08; wide, with a more or less distinct, longitudinal, length/width of metatibia: 5.2-5.4. punctate furrow. Abdominal sterna impunctate, Colour (Fig. 120). More or less pale rufous, with isodiametric microreticulation which is more elytra unicolourous. Palpi, antenna, and legs pale superficial at apex. red, the anterior leg usually very slightly darker. Legs. Profemur fairly stout, moderately wide, Under surface rufous. upper margin convex, lower surface depressed, with Head (Fig. 120). Of average size, frons in middle angulate anterior margin. Protibia rather short and depressed. Frontal sulcus deep, sinuate, impunctate. wide; 4-dentate but 4th tooth small, other teeth elon- Eye large, laterally well projected, orbit short, slightly gate; longitudinal sulcus on upper surface distinct. oblique. Clypeus in middle gently concave, coarsely Metatibia rather elongate. 1st tarsomere of metatarsus margined, with extremely short, obtuse lateral tooth. almost as long as the following three tarsomeres. Wing of clypeus shallowly separated from centre, Male genitalia (Fig. 52). Aedeagus rather nar- moderately protruded, rounded, bearing a shallow row, moderately elongate, slightly asymmetrically incision between wing and supraantennal plate. narrowed apicad; lower surface gently concave Crescent-shaped ridge on clypeus fairly distinct. An- over most of its length, apical third slightly curved tenna short, median antennomeres globular, slightly down and deeply concave. Apex small and short, at wider than long. Clypeus indistinctly divided from its lower surface strongly dentate, tip triangularly frons by a shallow, in middle usually interrupted rounded. Internal sac in apical third with three large,

281 denticulate and rather sclerotized folds. Both para- Pronotum (Fig. 134). Moderately elongate, meres short but very stout, the left one considerably slightly narrowed apicad, lateral margin slightly stouter than the right one; both with very short but oblique and faintly convex, slightly incised at apical acute, asetose apex. third, dorsal surface moderately convex. Basal angle Female gonocoxites. Very similar to those of with a very faint incision and protuberance. Apical C. tuberculiforns Blackburn, see fig. 219 in Baehr angle very little produced, apex almost straight, (2015). not margined. Anterior transverse sulcus well Variation. Very little variation noted. impressed, situated close to apex, barely crenulate. Median line deep. Basal sulcus narrow. Anterior Distribution. North QLD, Cape York Peninsula. marginal seta situated about at apical fourth. Basal groove only indicated by a few rather coarse, lon- Collecting circumstances. All specimens collected gitudinal arranged punctures. Dorsal surface with at light, at or near the banks of rivers. a few faint, irregularly transverse strioles, with very sparse, fine punctures, without microreticulation, very glossy. Clivina foveifrons, spec. nov. Elytra. Rather short, parallel, barely widened Fig. 134 apicad, convex but dorsally depressed, basal angle Examined types. Holotype: W, AUS15, QLD34, Stewart not produced. Lateral margin in basal fourth barely R. c. 30 km sse. Coen, 123m, 14°08'20.6"S, 143°16'21.3"E, concave. Striae well impressed, barely weaker lat- 8.5.2015, M.Baehr (QMT234915). erally and towards apex, rather coarsely punctate. Intervals slightly convex, without transverse wrin- Diagnosis. Medium sized, pale red species, char- kles, impunctate, without microreticulation, very acterized by moderately elongate prothorax, glossy glossy. Epipleura narrow, basally with a few shallow surface, impunctate head and barely punctate prono- impressions. tum, and a very large and deep median frontal fovea. Lower surface. Proepisternum impunctate, with Etymology. The name refers to the deep median fovea rather fine transverse strioles, with distinct iso- on the frons. diametric microreticulation, dull. Prosternum finely microreticulate, in anterior slightly more rugose. Description Metasternum without distinct transverse strioles. Metepisternum elongate, slightly > 2 × as long as Measurements. Length: 4.9 mm; width: 1.3 mm. Ra- wide, with a shallow, longitudinal, punctate furrow. tios. Length/width of pronotum: 1.06; base/apex of Abdominal sterna impunctate, with isodiametric pronotum: 1.36; width pronotum/head: 1.41; length/ microreticulation which is more superficial at apex. width of elytra: 2.01; length/width of protibia: 2.45; Legs. Profemur fairly stout, moderately wide, length/width of metatibia: 5.2. upper margin convex, lower surface depressed, Colour (Fig. 134). Head and pronotum pale red, with angulate anterior margin. Protibia rather short elytra slightly paler, unicolourous. Palpi, antenna, and wide; 4-dentate but 4th tooth small, other teeth and anterior leg pale red, median and posterior legs elongate; longitudinal sulcus on upper surface dis- dark yellow. Under surface pale red. tinct. Metatibia moderately elongate. 1st tarsomere Head (Fig. 134). Of average size, dorsally gently of metatarsus only as long as the following two convex. Frontal sulcus deep, sinuate, impunctate. Eye tarsomeres. large, laterally well projected, orbit short, slightly Male genitalia. Unknown. oblique. Clypeus in middle gently concave, coarsely Female gonocoxites. Very similar to those of margined, with extremely short, obtuse lateral C. tuberculiforns Blackburn, see fig. 219 in Baehr tooth. Wing of clypeus shallowly separated from (2015). centre, moderately protruded, rounded, bearing a Variation. Unknown. shallow incision between wing and supraantennal plate. Crescent-shaped ridge on clypeus well raised, Distribution. North QLD, central Cape York Pen- slightly triangular. Antenna short, median antenno- insula. Known only from type locality. meres globular, slightly wider than long. Clypeus divided from frons by a fairly deep sulcus. Frons Collecting circumstances. Collected at light at the with a large and deep, slightly oblong median fovea, bank of Stewart River. surface of frons almost even. Surface impunctate and without microreticulation, very glossy.

282 Clivina normanbyensis, spec. nov. Elytra (Fig. 121). Rather short, almost parallel, Figs 53, 121 little widened apicad, convex but dorsally depressed, basal angle not produced. Lateral margin in basal Examined types. Holotype: M, AUS15, QLD29, East Nor- fourth barely concave. Striae well impressed, barely manby R., c. 60 km wsw. Cooktown, 159m, 15°45'31.4"S, weaker laterally and towards apex, moderately 145°00'56.1"E, 30.4.-1.5.2015, M. Baehr (QMT234916). coarsely punctate. Intervals slightly convex, with Diagnosis. Medium sized, pale red species, char- several short, transverse wrinkles, impunctate, with acterized by moderately elongate prothorax, rather fine and somewhat superficial, isodiametric micro- coarsely punctate, microreticulate head, glossy pro- reticulation, moderately glossy. Epipleura narrow, notum but microreticulate elytra, and small, obtusely basally with a few shallow impressions. arrow-shaped apex of the aedeagus. Lower surface. Proepisternum impunctate, with rather fine transverse strioles, with distinct iso- Etymology. The name refers to the type locality, East diametric microreticulation, dull. Prosternum finely Normanby River, north Queensland. microreticulate, in anterior slightly more rugose. Metasternum without distinct transverse strioles. Description Metepisternum elongate, slightly > 2 × as long as Measurements. Length: 4.95 mm; width: 1.4 mm. Ra- wide, with a shallow, longitudinal, punctate furrow. tios. Length/width of pronotum: 1.05; base/apex of Abdominal sterna impunctate, with isodiametric pronotum: 1.26; width pronotum/head: 1.36; length/ microreticulation which is more superficial at apex. width of elytra: 1.97; length/width of protibia: 2.75; Legs. Profemur fairly stout, moderately wide, length/width of metatibia: 5.5. upper margin convex, lower surface depressed, with Colour (Fig. 121). Head and pronotum pale red, angulate anterior margin. Protibia rather short and th elytra slightly paler, unicolourous. Palpi, antenna, wide; 4-dentate but 4 tooth small, other teeth elongate; and anterior leg pale red, median and posterior legs longitudinal sulcus on upper surface distinct. Metati- st dark yellow. Under surface pale red. bia moderately elongate. 1 tarsomere of metatarsus Head (Fig. 121). Of average size, dorsally gently slightly longer than the following two tarsomeres. convex. Frontal sulcus deep, sinuate, impunctate. Male genitalia (Fig. 53). Aedeagus moderately Eye large, laterally well projected, orbit rather short, wide, rather 5 elongate, almost symmetrically slightly oblique. Clypeus in middle gently concave, narrowed apicad; lower surface gently concave coarsely margined, with extremely short, obtuse lat- throughout. Apex short, rather narrow, rounded eral tooth. Wing of clypeus shallowly separated from arrow-shaped, at its lower surface dentate, tip obtuse. centre, moderately protruded, rounded, bearing a Internal sac in apical third with three large, denticu- shallow incision between wing and supraantennal late and rather sclerotized folds. Both parameres plate. Crescent-shaped ridge on clypeus barely indi- large and stout, the left one considerably stouter than cated. Antenna short, median antennomeres globu- the right one; both with short but acute, asetose apex. lar, slightly wider than long. Clypeus not divided Female gonocoxites. Unknown. from frons. Frons with a small, shallow, circular Variation. Unknown. median fovea, surface of frons almost even. Surface rather sparsely, but coarsely punctate and with Distribution. North QLD, at the base of Cape York superficial, isodiametric microreticulation, glossy. Peninsula. Known only from type locality. Pronotum (Fig. 121). Moderately elongate, Collecting circumstances. Collected at light at the slightly narrowed apicad, lateral margin straight bank of East Normanby River. but slightly oblique, slightly incised at apical third, dorsal surface moderately convex. Basal angle with a very faint incision and protuberance. Apical angle Recognition very little produced, apex almost straight, not margined. Anterior transverse sulcus well impressed, According to shape of clypeus, body size, and unicol- situated close to apex, barely crenulate. Median line ourous elytra couplet 202. in the key in Baehr (2015) deep. Basal sulcus narrow. Anterior marginal seta is reached which has to be followed on as below: situated about at apical fourth. Basal groove shallow, linear, consisting of a row of longitudinal arranged 202. Body length > 5.0 mm (variable or doubtful punctures. Dorsal surface with a few faint, irregularly species under both couplets)...... 203. transverse strioles, in apical half with sparse, rather – Body length < 4.95 mm; if > 4.8 mm, either eye fine punctures and without microreticulation, in depressed or frons with a large and deep me- basal third with fine, rather superficial, isodiametric dian fovea or elytra distinctly microreticu- microreticulation, glossy. late...... 212.

283 203. Prothorax and elytra elongate, ratio l/w of 208. Prothorax ratio l/w < 1.09 and eye large and prothorax 1.14, ratio l/w of elytra 2.12; eye orbit almost perpendicular and aedeagus with rather depressed, orbit large, oblique (B15 fig. short, arrow-shaped apex, both parameres with 374); aedeagus rather wide, with slightly asym- narrow and elongate apex (B15 fig. 126), or metrical, arrow-shaped apex (B15 fig. 131). aedeagus unknown...... 209. n.NT...... laevicollis Baehr, 2015 – Prothorax ratio l/w various and eye slightly – Prothorax and elytra shorter, ratio l/w of smaller and orbit slightly oblique and aedeagus prothorax < 1.12, usually less, ratio l/w of elytra either with longer, arrow-shaped apex or with < 2.08, usually less; eye laterad protruded, spatulate apex, both parameres with very short orbit short, slightly oblique to perpendicular; apex (Fig. 51; B15 figs 133, 135), or aedeagus aedeagus differently shaped (Fig. 51; B15 figs unknown, if latter, ratio l/w of prothorax 122, 126, 127, 132-135)...... 204. > 1.11...... 210. 204. Elytra short, ratio l/w 1.93-1.97; pronotum 209. Head with moderately dense and coarse punc- impunctate; elytra not microreticulate; proti- tures; pronotum dorsally moderately convex; bia narrow and elongate, ratio l/w > 2.9; ae- elytra without microreticulation, usually with deagus with very narrow apical part, apex a dark median spot; aedeagus with short, ar- hook-shaped (B15 fig. 122). n. and c.QLD, n.NT, row-shaped apex, both parameres with narrow n.WA...... atridorsis Sloane, 1905 and elongate apex (B15 fig. 126). nw.QLD, – Elytra longer, ratio l/w > 1.98, usually more; n.NT, n.WA...... toledanoi Baehr, 2015 pronotum usually punctate, if impunctate, – Head with dense and coarse punctures; pro- elytra distinctly microreticulate; protibia short- notum dorsally markedly convex; elytra with er and wider, ratio l/w usually < 2.5; aedeagus fine microreticulation, without dark median different (Fig. 51; B15 figs 126, 127, 132-135). . spot; aedeagus unknown. n.WA...... 205...... convexior Baehr, 2015 205. Head completely and coarsely punctate; elytra 210. Elytra finely microreticulate; aedeagus with coarsely microreticulate; apical part of aedea- spatulate apex (Fig. 51; B15 fig.135), or un- gus odd-shaped, apex strongly hook-shaped known...... 211. (B15 fig. 127). n.NT...... – Elytra not microreticulate; eye much produced, ...... paucidentata Baehr, 2015 orbit very short, only slightly oblique; aedeagus – Head only partly and less coarsely punctate; with triangular arrow-shaped apex (B15 fig. microreticulation of elytra various, if coarse, 133). n.WA...... marlgu Baehr, 2015 apex of aedagus somewhat arrow-shaped (B15 211. Body size larger, 5.8 mm; head finely punctate; fig. 126); aedeagus different (Fig. 51; B15 figs prothorax decidedly narrowed apicad (B15 fig. 126, 132-135)...... 206. 345); elytra short, ratio l/w 1.93; apex of ae- 206. Pronotum and elytra perceptibly microreticu- deagus markedly curved down, asymmetri- late; apex of aedeagus symmetrically spatulate cally spatulate apex (B15 fig. 135). n.NT: near (B15 figs 132, 134)...... 207. Mataranka...... finitima Baehr, 2015 – Pronotum not microreticulate, elytra microre- – Body size smaller, < 5.3 mm; head coarsely ticulate or not; apex of aedeagus arrow-shaped punctate; prothorax barely narrowed apicad or asymmetrically spatulate (Fig. 51; B15 figs (B15 fig. 375); elytra longer, ratio l/w > 2.0; 126, 133, 135), or unknown...... 208. apex of aedeagus either far less curved down (Fig. 51), or aedeagus unknown...... 211a. 207. Microreticulation on pronotum and elytra distinct and fairly coarse; lower surface of 211a. Eye smaller, laterad less produced, orbit aedeagus regularly concave, apex not mark- larger, distinctly oblique; head with coarser edly turned down, less spatulate (B15 fig. 132). punctation; pronotum longer, more parallel- n.WA...... discrepans Baehr, 2015 sided, ratio width base/apex 1.21 (B15 fig. 375); – Microreticulation on pronotum and elytra aedeagus unknown. n.NT: Kakadu NP...... superficial and difficult to see; lower surface ...... gubarae Baehr, 2015 of aedeagus in middle straight, apex mark- – Eye larger, laterad more produced, orbit edly turned down, spatulate (B15 fig. 134). smaller, far less oblique; head with less coarse n.NT...... spatulata Baehr, 2015 punctation; pronotum shorter, less parallel- sided, ratio width base/apex > 1.29 (Fig. 119);

284 apex of aedeagus moderately curved down, 216. Eye moderately large and laterad moderately asymmetrically spatulate (Fig. 51). ne.QLD. ... produced, orbit distinctly oblique (B15 fig...... laevigata, spec. nov. 376)...... 217. 212. Elytra coarsely microreticulate; pronotum – Eye large and laterad almost semicircularly distinctly microreticulate...... 213. protruded, orbit almost perpendicular (B15 fig. 377)...... 222. – Elytra less coarsely or not microreticulate; pronotum not or superficially microreticulate; 222. Head short and wide, deeply imbedded into if microreticulation on elytra fairly distinct, prothorax, coarsely punctate; eye laterad mark- pronotum not microreticulate and elytral in- edly protruded (B15 fig. 377); pronotum and tervals very convex...... 215. elytra not microreticulate; aedeagus with short and wide, arrow-shaped apex that bears re- 213. Prothorax and elytra shorter, ratio l/w of markably distinct lateral teeth (B15 fig. 141). prothorax < 1.08, ratio l/w of elytra < 2.05...... n.WA: KID...... breviceps Baehr, 2015 ...... 213a. – Head not as deeply imbedded into prothorax; – Prothorax and elytra longer, ratio l/w of pro- head usually not as distinctly punctate; eye thorax > 1.10, ratio l/w of elytra > 2.10; aedea- size and microreticulation of pronotum and gus unknown...... 214. elytra various; aedeagus different (Figs 51, 53; 213a. Body size larger, > 4.9 mm; prothorax shorter, B15 figs 121, 122, 139)...... 223. ratio l/w < 1.05; aedeagus with somewhat 223. Either pronotum or elytra perceptibly micro- spatulate apex (B15 fig. 132). n.WA...... reticulate; aedeagus not with symmetric, knob- ...... discrepans Baehr, 2015 shaped apex (Figs 51, 53; B15 figs 122, 139). ... – Body size smaller, < 4.7 mm; prothorax slight- ...... 224. ly longer, ratio l/w > 1.06; aedeagus with small, – Pronotum and elytra not at all microreticulate; but on lower side remarkably hook-shaped aedeagus with symmetric, knob-shaped apex apex (Fig. 52). ne.QLD...... (B15 fig. 121), or unknown...... 226...... uncinata, spec. nov. 224. Pronotum impunctate, finely microreticulate 214. Prothorax impunctate, narrower in relation to or not; elytra not microreticulate; aedeagus head, ratio width pronotum/head 1.22; elytra with very narrow apical part, apex slightly on disk distinctly depressed, microreticulation hooked (B15 fig. 122). ce. and n.QLD, n.NT, on elytra slightly less coarse. ne.QLD...... n.WA...... atridorsis Sloane, 1905 ...... languida Baehr, 2015 – Pronotum punctate, not microreticulate; elytra – Prothorax sparsely though coarsely punctate, finely microreticulate; aedeagus differently wider in relation to head, ratio width prono- shaped...... 225. tum/head 1.27 (B15 fig. 346); elytra on disk convex, microreticulation on elytra coarser. 225. Pronotum coarsely punctate; elytra shorter, nw.QLD...... scabra Baehr, 2015 ratio l/w < 1.95; aedeagus with asymmetric, obtusely rounded apex (B15 fig. 139). n.QLD, 215. Frons with large and very deep median fovea n.NT...... nourlangie Baehr, 2015 (Fig. 134), aedeagus unknown...... 215a. – Pronotum finely punctate; elytra longer, ratio – Frons at most with shallow frontal fovea; ae- l/w > 1.97; aedeagus with spatulate or small, deagus variously shaped...... 216. obtusely arrow-shaped apex (Figs 51, 53). 215a. Prothorax and elytra slightly longer, ratios l/w n.QLD...... 225a. prothorax 1.08, l/w elytra 2.03; eye laterad 225a. Pronotum and elytra longer, ratio l/w of pro- more protruded, therefore prothorax nar- notum > 1.08, l/w of elytra > 2.0; aedeagus with rower as compared with head, ratio width large, spatulate apex (Fig. 51). . laevigata, spec. prothorax/ head 1.32. nw.NT...... nov...... fossulata Baehr, 2015 – Pronotum and elytra shorter, ratio l/w of – Prothorax and elytra slightly shorter, ratios pronotum 1.05, l/w of elytra 1.97; aedeagus l/w prothorax 1.06, l/w elytra 2.01; eye laterad with small, obtusely arrow-shaped apex less protruded, therefore prothorax wider as (Fig. 53)...... normanbyensis, spec. nov. compared with head, ratio width prothorax/ head 1.41. ne.QLD, c.CYP...... foveifrons, spec. nov.

285 Appendix

Summary of measurements and ratios of the species of the genus Clivina mentioned in present paper. N = number of specimens measured; L = length in mm; l/w pr = ratio length/width of pronotum; b/a pr = ratio width of base/ width of apex of pronotum; pr/h = ratio width of pronotum/width of head; l/w el = ratio length/width of elytra; l/w pt = ratio length/width of protibia; l/w mt = ratio length/width of metatibia. *, measurements included in measurements of males. Especially measurements of the protibia can rather vary according to the age of the specimen and the degree of abrasion of the lateral teeth.

M/W M W NL l/w pr b/a pr pr/h l/w el l/w pt l/w mt l/w pt l/w mt procera M/W 6 12.2-16.7 1.02-1.05 1.26-1.33 1.33-1.40 2.01-2.06 4.0-4.15 5.6-5.8 3.3-3.35 5.5-5.8 obscuripes M/W 6 10.0-13.5 1.02-1.03 1.27-1.33 1.30-1.33 2.0-2.6 3.5-3.65 6.0-6.1 3.0-3.25 5.5-5.7 subrufipes M/W 6 10.6-13.5 0.99-1.02 1.22-1.26 1.25-1.33 2.04-2.08 3.7-3.8 5.6-6.1 3.0-3.15 5.6-6.0 cooindae M/W 6 9.3-12.7 1.01-1.04 1.26-1.30 1.27-1.32 1.99-2.07 3.65-3.8 6.2-6.25 3.25-3.35 6.0 sinuicola M/W 6 10.1-15.9 0.99-1.01 1.22-1.33 1.28-1.35 1.98-2.07 3.4-3.8 6.25-6.6 3.4-3.5 5.4-5.8 thoracica M 1 14.5 1.11 1.27 1.32 2.07 4.35 6.7 g. gemina M/W 6 11.3-13.5 1.02-1.05 1.23-1.27 1.26-1.34 2.05-2.10 3.5-3.8 6.3-6.8 2.75-2.85 5.8-6.1 g. nigripes M/W 6 11.7-13.9 1.01-1.03 1.29-1.35 1.34-1.48 2.03-2.09 3.35-3.6 6.5-6.7 2.7-2.8 6.1-6.2 carnabyi M/W 6 15.2-16.6 1.02-1.04 1.32-1.36 1.39-1.44 2.09-2.15 3.3-3.5 6.35-6.5 3.0-3.1 5.9-6.0 windjanae M/W 6 12.0-13.4 0.97-1.03 1.32-1.36 1.30-1.38 2.03-2.07 3.4-3.5 6.2-6.25 2.8-3.0 6.1-6.2 mahoni M/W 2 15.5-16.4 1.03-1.08 1.33-1.36 1.36-1.39 2.09-2.11 4.5 5.7 3.4 5.6 montisbelli W 1 12.1 1.06 1.20 1.22 2.22 2.8 7.3 victoriae M/W 5 12.3-13.3 0.99-1.01 1.21-1.25 1.28-1.32 2.00-2.05 2.85-3.25 5.7-5.9 2.75 5.55 ryaceki M/W 6 15.2-18.9 1.00-1.03 1.25-1.30 1.34-1.38 2.06-2.12 4.15-4.2 5.8-6.1 3.3-3.5 5.7-5.9 heros M/W 6 18.2-23.9 0.98-1.05 1.30-1.34 1.36-1.44 2.13-2.18 3.9-4.05 6.2-6.5 2.95-3.15 6.4-6.6 goldingi M 1 16.5 1.04 1.35 1.39 c. 2.08 3.3 5.8 newcastleana M/W 6 11.2-14.4 0.96-1.00 1.25-1.28 1.33-1.39 2.00-2.05 3.8-4.0 6.2-6.4 2.9-3.2 5.8-6.2 glabripennis M/W 6 13.7-17.0 0.98-1.01 1.29-1.31 1.37-1.41 2.02-2.06 3.35 6.7 2.85-3.1 6.1-6.3 micans M/W 4 12.9-15.6 1.04-1.06 1.16-1.24 1.42-1.50 2.0-2.03 3.25-3.4 6.0-6.1 2.5 5.1 rugosifrons M/W 6 14.1-15.9 1.01-1.06 1.26-1.33 1.32-1.38 2.02-2.08 3.15-3.25 6.1-6.2 2.9-3.05 5.7-6.1 monilicornis M/W 6 9.1-13.0 1.06-1.09 1.35-1.37 1.49-1.52 1.93-2.02 3.1-3.2 5.1-5.2 2.8-2.85 4.9-5.0 conicollis M/W 6 10.6-14.5 1.10-1.13 1.40-1.51 1.47-1.52 2.06-2.12 3.35-3.5 5.25-5.5 2.9-3.1 5.2-5.3 dubia M 1 12.8 1.05 1.25 1.35 2.0 3.3 5.25 regularis M/W 6 9.3-11.9 1.05-1.11 1.22-1.27 1.31-1.40 1.93-2.02 2.7-2.85 5.7-5.8 2.35-2.45 5.7-5.8 elegans M/W 6 12.8-17.5 1.09-1.16 1.36-1.42 1.44-1.52 2.02-2.08 3.7-3.8 7.2-7.4 3.4-3.5 6.5-6.8 interposita M/W 5 11.7-13.2 1.03-1.08 1.43-1.45 1.44-1.54 1.94-2.02 3.7-3.8 7.1-7.5 3.3-3.5 6.4-6.7 kershawi M/W 5 12.2-14.1 1.06-1.13 1.51-1.56 1.40-1.47 2.03-2.11 3.7-3.8 7.3-7.5 3.3-3.4 6.5-6.6 variseta M/W 2 17.0-17.8 0.99-1.01 1.55-1.58 1.55-1.57 1.83-1.90 5.0 7.3 3.9 6.7 triseriata W 1 15.7 0.98 1.65 1.62 1.90 3.8 6.6 inopinata M 1 13.4 1.0 1.62 1.54 1.88 5.0 7.3 profundestriolata W 2 12.9-13.8 0.92-0.94 1.62-1.66 1.65-1.67 1.75-1.80 3.8 7.65-7.7 g. gracilipes M/W 6 9.5-13.4 0.94-0.96 1.51-1.61 1.42-1.59 1.76-1.80 4.7-4.9 8.0-8.3 3.5-3.7 7.5-7.6 g. longior W 4 11.4-12.9 0.92-0.96 1.47-1.54 1.43-1.53 1.88-1.91 3.5-3.6 7.5-7.6 marginata M/W 6 10.2-14.5 0.89-0.97 1.48-1.58 1.43-1.47 1.82-1.88 5.3-5.6 8.8-9.0 4.1-4.3 7.4-7.7 oblonga M/W 6 10.7-14.1 0.99-1.01 1.31-1.35 1.40-1.45 1.91-1.96 3.1-3.3 6.0-6.1 3.1-3.2 5.6-5.8 robusta M/W 6 12.5-17.2 1.02-1.08 1.27-1.32 1.36-1.42 1.86-2.00 4.0-4.4 6.5-6.7 3.2-3.3 5.5-5.7 obliquicollis M 4 11.4-12.5 1.00-1.02 1.28-1.34 1.39-1.45 1.95-1.97 3.9-4.1 7.4-7.5 bankae M/W 6 11.6-14.9 0.93-1.00 1.37-1.43 1.40-1.50 2.06-2.13 3.0-3.1 6.8-6.9 2.9-3.0 6.5-6.9 incurvicollis M/W 6 11.1-14.5 0.93-0.99 1.40-1.46 1.42-1.51 1.93-2.02 3.1-3.8 6.3-8.1 3.3-3.45 6.6-7.2 platynota M 2 13.0-13.2 1.01-1.04 1.33-1.34 1.34-1.37 2.11 3.4-3.5 7.4-7.5 rectipennis W 2 11.8-14.1 0.96-0.97 1.37-1.40 1.39-1.43 2.10-2.13 3.0-3.2 6.9-7.2 brevisterna M/W 6 11.3-12.8 0.97-1.02 1.42-1.45 1.52-1.53 1.82-1.86 3.5-3.8 6.4-6.5 3.25-3.4 6.35-6.5

286 M/W M W NL l/w pr b/a pr pr/h l/w el l/w pt l/w mt l/w pt l/w mt major M/W 6 14.2-16.8 1.00-1.01 1.51-1.58 1.51-1.56 1.91-1.98 4.3-5.0 7.3-7.7 3.7-4.0 7.5-7.6 mastersi M/W 6 15.4-21.1 0.98-1.05 1.50-1.55 1.50-1.57 1.96-2.07 4.6-4.8 7.5-7.8 3.9-4.0 6.7-6.8 cobourgiana M 3 17.8-18.9 1.00-1.01 1.51-1.54 1.51-1.54 1.93-1.98 4.5-4.65 6.9-7.2 nyctosyloides M/W 6 9.6-13.0 0.90-0.97 1.56-1.67 1.56-1.66 1.80-1.85 3.75-3.85 6.3-6.4 3.4-3.5 5.5-5.7 ovalior W 1 13.1 0.96 1.46 1.53 1.81 3.8 6.9 nitescens M/W 3 10.3-11.7 0.92-0.95 1.54-1.60 1.56-1.61 1.84-1.92 3.45-3.5 6.3-6.35 3.25 6.2 ovalipennis W 1 c. 12.6 1.0 1.48 - 1.85 c. 3.4 c. 6.5 pachysoma M/W 6 12.0-14.2 0.86-0.90 1.50-1.56 1.52-1.59 1.72-1.78 3.4-3.5 6.7-6.8 3.15-3.2 5.9-6.1 froggatti W 1 7.7 1.06 1.25 1.37 1.83 2.5 5.7 hackeri W 4 8.7-10.4 0.86-0.87 1.68-1.92 1.69-1.82 1.53-1.57 3.35-3.5 5.6-5.65 demarzi M/W 6 7.7-9.7 0.84-0.92 1.75-1.82 1.72-1.80 1.53-1.64 3.2-3.25 5.6-5.7 3.0-3.3 5.6-5.7 crassipennis M/W 6 8.8-10.1 0.86-0.88 1.57-1.70 1.65-1.70 1.61-1.73 2.75 5.4 2.8-2.9 5.1-5.2 horaki W 1 8.4 0.87 1.70 1.57 1.67 3.1 6.4 darwini M/W 6 5.7-6.5 1.05-1.09 1.27-1.29 1.37-1.52 1.86-1.94 3.1-3.2 5.5-5.6 2.9-3.0 5.25-5.3 macleayi M/W 5 7.8-8.8 0.87-0.98 1.25-1.33 1.44-1.48 1.82-1.90 3.25-3.3 5.2 3.75-2.9 5.0-5.1 horneri M/W 6 7.7-9.1 1.02-1.07 1.32-1.40 1.49-1.63 1.89-1.91 3.25-3.4 5.35-5.5 2.7-3.05 5.2-5.3 vixsulcata W 1 11.4 1.18 1.36 1.37 2.32 2.0 5.4 semirubra M/W 6 6.9-7.7 1.05-1.07 1.10-1.13 1.15-1.19 2.14-2.19 2.45-2.6 5.2-5.4 ** infans M 2 4.2-4.3 1.0 1.24 1.23-1.27 1.98-2.0 2.6-2.7 5.4-5.5 g. planior W 1 4.7 1.16 1.15 1.25 2.25 2.7 5.15 laevigata M/W 6 4.6-5.3 1.08-1.12 1.29-1.34 1.32-1.34 2.0-2.05 2.35-2.55 5.5-5.7 ** moretona W 3 3.5-3.6 1.10-1.13 1.18-1.21 1.28-1.32 2.08-2.12 2.45-2.65 5.5-5.7 uncinata M/W 6 4.4-4.7 1.06-1.08 1.19-1.23 1.28-1.32 0.98-2.03 2.55-2.75 5.2-5.4 ** foveifrons W 1 4.9 1.06 1.36 1.41 2.01 2.45 5.2 normanbyensis M 1 4.95 1.05 1.26 1.36 1.97 2.75 5.5

Homonymy of Clivina burlirschi Baehr, 2015 a dozen species of the genus Geoscaptus (Macleay, 1863)), as well as the presence of but very few small Clivina bulirschi Baehr, 2015 was described in the 2nd hygrophilous species (only the genus Setodyschirius part of the revision of the Australian Clivinina (Baehr Fedorenko, 1996 with about a dozen species). In 2015). Dostal (2015) also described a Clivina (Physo- other continents the niche of very small hygrophil- clivina) bulirschi Dostal, 2015 from West Africa. Be- ous scaritine species is occupied by usually many cause this latter species was printed a month earlier species of the genus Dyschirius s. l. Bonelli, 1813 and than the Australian species, this is a junior homonym a number of related genera, and the niche of large and is herewith renamed C. bulirschiana, spec. nov. hygrophilous species is occupied by many species of the large genus Scarites s. l. Fabricius, 1775 and a great number of related genera. Thus, it seems that Remarks in Australia both niches are mainly occupied by species of the genus Clivina, and indeed, the Australian As already mentioned in part 2 of the revision, the Clivina fauna includes some of the smallest species Australian Clivina fauna differs from those of all other of this genus, as well as the largest. continents in the extreme differences in body size and Explication of the very large number of species perhaps also in the very large number of species, at in a prevailing dry continent seems to be more dif- least in relation to the small size of the continent, as ficult. However, the oscillations of climate during compared with other continents. Moreover, this large Pleistocene and late Tertiary in Northern and West- number of mainly hygrophilous species is surprising ern Australia repeatedly united fairly humid areas for a largely dry continent. Indeed, semiarid and arid and dissected these again by dry corridors, which regions in Australia cover relatively more space than might have caused repeated invasions of species and in any other continent. subsequent dissection of their ranges. This probably The very large differences in body size may be a produced quite rapid separation of species and thus, consequence of the presence of but very few large, it may be one, or even the most important, reason for hygrophilous scaritine species in Australia (only half the very high species number and diversity.

287 References Moore, B. P., Weir, T. A. & Pyke, J. E. 1987. Rhysodidae and Carabidae. Pp. 17-320 in: Zoological Catalogue Baehr, M. 1987. Clivina demarzi sp. n., a new flightless of Australia 4, Canberra (Australian Government Clivina from the Northern Territory of Australia Publishing Service). (Insecta, Coleoptera, Carabidae, Scaritinae). Spixi- Putzeys, J. 1846. Monographie des Clivina et genres voi- ana 10: 187-190. sins précédée d’un tableau synoptique des genres – – 2008. The Australian Clivinini 1. The genera Ancus de la tribu des Scaritides. Mémoirs de la Société des Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Sciences de Liège 2: 521-663. Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara – – 1862. Postscriptum ad Clivindarum monographiam Sloane, Syleter Andrewes, the subgenera Paraclivina atque de quibusdam aliis. Mémoirs de la Société des Kult, Semiclivina Kult, and the atrata-, biplagiata-, Sciences de Liège 18: 1-78. brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, – – 1866. Révision des Clivinides d’Australie. Stettiner denticollis-, grandiceps-, incerta-, lobata-, obliquata-, ob- Entomologische Zeitschrift 27: 33-42. soleta, orbitalis-, planiceps-, sulcaticeps-, tranquebaria-, – – 1867. Révision générale des Clivinides. Annales de and wurargae-groups of the genus Clivina Latreille. la Société Entomologique de Belgique 10: 1-242. With a note on a record of the genus Parathlibops – – 1868. Supplément à la révision générale des Cli- Basilewsky (Scapterini) (Carabidae, Scaritinae). vinides. Annales de la Société Entomologique de Coleoptera 12: 1-220. Belgique 11: 7-22. – – 2015. Revision of the Australian Clivinini 2. The – – 1873. Deuxième supplément à la révision géné- ambigua-, bataviae-, bullata-, cava-, emarginata-, hetero- rale des Clivinides. Annales de la Société Ento- gena-, impressiceps-, and sloanei-groups of the genus mologique de Belgique 16: 10-18. Clivina Latreille, the new genus Rubidiclivina, and Sloane, T. G. 1896a. On the Australian Clivinides (Fam. additions to the 1st part (Coleoptera, Carabidae, Carabidae). Revision of the Australian species of the Scaritinae). Entomologische Blätter und Coleoptera genus Clivina with the description of a new genus 111: 59-446. Clivinarchus. Proceedings of the Linnean Society of Blackburn, T. 1890. Notes on Australian Coleoptera, New South Wales 21: 143-257. with descriptions of new species. Part V. Proceed- – – 1896b. Appendix to the Australian Clivinides (Fam. ings of the Linnean Society of New South Wales Carabidae). The Clivinides of King’s Sound and its (2) 4: 1247-1276. vicinity. Proceedings of the Linnean Society of New Csiki, E. 1927. Carabidae: Carabinae II. In: Schenkling, S. South Wales 21: 275-280. (ed.). Coleopterorum Catalogus I, part 92, pp. 315- – – 1905. Revisional notes on Australian Carabidae. Part 622, Berlin (W. Junk). I. Proceedings of the Linnean Society of New South Dostal, A. 2015. Two new species of Clivina Latreille, Wales 29: 699-733. 1802 (Coleoptera: Carabidae: Scaritinae) from Af- – – 1907. Studies in Australian Entomology. No. XV. rica. Zeitschrift der Arbeitsgemeinschaft Österrei- New genera and species of Carabidae, with some chischer Entomologen 67: 27-37. notes of synonymy (Clivinini, Scaritini, Cuneipec- Lorenz, W. 2005. Systematic list of extant ground beetles tini, Trigonotomini and Llebiini). Proceedings of the of the world (Insecta Coleoptera “Geadephaga”: Linnean Society of New South Wales 32: 346-381. Trachypachidae and Carabidae incl. Paussinae, – – 1916. New Australian species of Carabidae belong- Cicindelinae. Rhysodidae). 530 pp., 2. ed., Tutzing ing to the tribe Scaritini (Coleoptera). Proceedings (printed by the author). of the Linnean Society of New South Wales 41: Macleay, W. J. 1863. On the Scaritidae of New Holland. 597-630. Transactions of the Entomological Society of New – – 1917. Carabidae from tropical Australia (New gen- South Wales 1: 55-74. era and species, notes and synonymy, and synoptic tables. Tribes Scaritini, Harpalini, Odacanthini, Lebiini, and Helluonini). Proceedings of the Lin- nean Society of New South Wales 42: 406-438.

288 Systematic list of the species mentioned in present paper with information about their distribution sulcaticeps group elegans group vixsulcata, spec. nov. n.QLD: CYP elegans subgroup elegans Putzeys, 1862 n.QLD: tip of CYP, heterogena group Torres Strait Is. lobipes subgroup interposita, spec. nov. n.QLD: n.CYP semirubra, spec. nov. nw.QLD kershawi Sloane, 1916 n.QLD: ce.CYP oodnadattae subgroup variseta, spec. nov. n.QLD: CYP infans, spec. nov. nw.QLD triseriata, spec. nov. n.NT inopinata, spec. nov. n.QLD australica subgroup profundestriolata, spec. nov. c. + sw.QLD gerstmeieri planior, subspec. nov. nw.QLD gracilipes subgroup gracilipes gracilipes Sloane, 1896 tuberculifrons subgroup nw.QLD, n.NT, n.WA: KID laevigata, spec. nov. n.QLD: CYP gracilipes longior, subspec. nov. uncinata, spec. nov. n.QLD: CYP n.QLD: CYP moretona, spec. nov. n.QLD: n.CYP marginata (Putzeys, 1868) n.QLD, n.NT, n.WA foveifrons, spec. nov. n.QLD: CYP normanbyensis, spec. nov. n.QLD oblonga subgroup oblonga (Putzeys, 1873) e.NSW, se.QLD procera group = abbreviata (Putzeys, 1873) procera Putzeys, 1866 e.SA, VIC, NSW, e.QLD robusta Sloane, 1905 e.NSW = prominens Putzeys, 1866 (syn. nov.) = macleayana (nom. nov.) obscuripes (Blackburn, 1890) n. + nw.NT = foveiceps (Macleay, 1863) subrufipes, spec. nov. n.NT: Kakadu NP obliquicollis subgroup cooindae, spec. nov. n.NT, ne.QLD obliquicollis Sloane, 1905 nw.WA: KID sinuicola, spec. nov. nw.QLD, ne.NT bankae, spec. nov. nc.NT thoracica, spec. nov. nw.QLD incurvicollis, spec. nov. nw.WA: Pilbara + sw.KID gemina gemina, spec. nov. nw.NT, ne.WA: ne.KID platynota, spec. nov. nw.WA: Pilbara gemina nigripes, subspec. nov. rectipennis, spec. nov. sc.NT, ce.WA n.WA: s. + w.KID brevisterna subgroup carnabyi, spec. nov. n.WA: s. + w.KID brevisterna Sloane, 1916 n.NT windjanae, spec. nov. n.WA: sw.KID major Sloane, 1917 (stat. nov.) mahoni, spec. nov. nw.NT n.NT montisbelli, spec. nov. n.WA: w.KID mastersi Sloane, 1896 n.NT victoriae, spec. nov. nw.NT cobourgiana, spec. nov. n.NT: Cobourg Pen. ryaceki, spec. nov. n. + nw.NT, ne.WA: ne.KID nyctosyloides Putzeys, 1868 ne.QLD heros, spec. nov. n.WA: KID = propinqua (nom. nov.) goldingi, spec. nov. n.NT = interstitialis Sloane, 1896 newcastleana, spec. nov. nc.NT ovalior, spec. nov. ne.QLD glabripennis, spec. nov. sw.QLD, ne.SA nitescens, spec. nov. n.NT, ? n.WA micans, spec. nov. w.QLD, ce.NT ovalipennis Sloane,1905 n.QLD rugosifrons, spec. nov. n.QLD, ne.NT (= ovipennis Sloane, 1896) monilicornis Sloane, 1896 n.QLD, n.NT pachysoma, spec. nov. nw.WA: Pilbara conicollis, spec. nov. n.QLD: CYP, n.NT froggatti Sloane,1896 n.WA: w.KID dubia, spec. nov. AUS hackeri subgroup regularis Sloane, 1896 ne.NSW, se.QLD hackeri Sloane, 1907 n.QLD demarzi Baehr, 1987 n.NT crassipennis, spec. nov. n.NT, n.WA: ne.KID horaki, spec. nov. n.WA: sw.KID darwini subgroup darwini Sloane, 1916 n.NT macleayi Sloane, 1896 n.NT horneri, spec. nov. n.NT

289 1 2 3

4 5 6

7 8 9 Figs 1-9. Aedeagus (left side and apex from below), parameres. Scale bars: 1 mm. 1. Clivina procera Putzeys, 1866. 2. C. obscuripes (Blackburn, 1890). 3. C. subrufipes, spec. nov. 4. C. cooindae, spec. nov. 5. C. sinuicola, spec. nov. 6. C. thoracica, spec. nov. 7. C. gemina gemina, spec. nov. 8. C. gemina nigripes, subspec. nov. 9. C. carnabyi, spec. nov.

290 10 11 12

13 14 15

16 17 18 Figs 10-18. Aedeagus (left side and apex from below), parameres. Scale bars: 1 mm. 10. C. windjanae, spec. nov. 11. C. mahoni, spec. nov. 12. C. victoriae, spec. nov. 13. C. ryaceki, spec. nov. 14. C. heros, spec. nov. 15. C. goldingi, spec. nov. 16. C. newcastleana, spec. nov. 17. C. glabripennis, spec. nov. 18. C. micans, spec. nov.

291 19 20 21

22 23 24

25 26 27 Figs 19-27. Aedeagus (left side and apex from below), parameres. Scale bars: 1 mm. 19. C. rugosifrons, spec. nov. 20. C. monilicornis Sloane, 1896. 21. C. conicollis, spec. nov. 22. C. dubia, spec. nov. 23. C. regularis Sloane, 1896. 24. C. ele gans Putzeys, 1862. 25. C. interposita, spec. nov. 26. C. kershawi Sloane, 1916. 27. C. variseta, spec. nov.

292 28 29 30

31 32 33

34 35 36 Figs 28-36. Aedeagus (left side and apex from below), parameres. Scale bars: 1 mm. 28. C. inopinata, spec. nov. 29. C. gracilipes gracilipes Sloane, 1896. 30. C. marginata (Putzeys, 1868). 31. C. oblonga (Putzeys, 1873). 32. C. robusta Sloane, 1905. 33. C. obliquicollis Sloane, 1905. 34. C. bankae, spec. nov. 35. C. incurvicollis, spec. nov. 36. C. platynota, spec. nov.

293 37 38 39

40 41 42

43 44 45 Figs 37-45. Aedeagus (left side and apex from below), parameres. Scale bars: 1 mm. 37. C. brevisterna Sloane, 1916. 38. C. major Sloane, 1917. 39. C. mastersi Sloane, 1896. 40. C. cobourgiana, spec. nov. 41. C. nyctosyloides Putzeys, 1868. 42. C. nitescens, spec. nov. 43. C. pachysoma, spec. nov. 44. C. demarzi Baehr, 1987. 45. C. crassipennis, spec. nov.

294 46 47 48

49 50 51

52 53 Figs 46-53. Aedeagus (left side and apex from below), parameres. Scale bars: 1 mm. 46. C. darwini Sloane, 1916. 47. C. macleayi Sloane, 1896. 48. C. horneri, spec. nov. 49. C. semirubra, spec. nov. 50. C. infans, spec. nov. 51. C. laevigata, spec. nov. 52. C. uncinata, spec. nov. 53. C. normanbyensis, spec. nov.

295 54 55 56 57

58 59 60 61

62 63 64 65 Figs 54-65. Right gonocoxites (ventral view). Scale bars: 0.5 mm. 54. Clivina procera Putzeys, 1866. 55. C. windjanae, spec. nov. 56. C. heros, spec. nov. 57. C. newcastleana, spec. nov. 58. C. rugosifrons, spec. nov. 59. C. conicollis, spec. nov. 60. C. regularis Sloane, 1896. 61. C. elegans Putzeys, 1862. 62. C. profundestriolata, spec. nov. 63. C. gracilipes gracilipes Sloane, 1896. 64. C. marginata (Putzeys, 1868). 65. C. oblonga (Putzeys, 1873).

296 66 67 68 69

70 71 72 73

74 75 76 77 Figs 66-77. Right gonocoxites (ventral view). Scale bars: 0.5 mm. 66. C. robusta Sloane, 1905. 67. C. bankae, spec. nov. 68. C. incurvicollis, spec. nov. 69. C. rectipennis, spec. nov. 70. C. major Sloane, 1917. 71. C. mastersi Sloane, 1896. 72. C. nyctosyloides Putzeys, 1868. 73. C. pachysoma, spec. nov. 74. C. demarzi Baehr, 1987. 75. C. darwini Sloane, 1916. 76. C. horneri, spec. nov. 77. C. vixsulcata, spec. nov.

297 78 79 Figs 78-79. Clypeus excision. Clivina procera Putzeys, 1866. 79. C. elegans Putzeys, 1862.

80 81 82 83

84 85 86 87 Figs 80-87. Habitus. Body lengths in brackets. 80. Clivina procera Putzeys, 1866 (14.5 mm). 81. C. obscuripes (Black- burn, 1890) (11.8 mm). 82. C. cooindae, spec. nov. (11.5 mm). 83. C. gemina gemina, spec. nov. (12.8 mm). 84. C. montis belli, spec. nov. (12.1 mm). 85. C. glabripennis, spec. nov. (15.7 mm). 86. C. micans, spec. nov. (15.6 mm). 87. C. regularis Sloane, 1896 (10.8 mm).

298 88 89 90 91

92 93 94 95

96 97 98 99 Figs 88-99. Habitus. Body lengths in brackets. 88. C. elegans Putzeys, 1862 (15.5 mm). 89. C. interposita, spec. nov. (12.8 mm). 90. C. kershawi Sloane, 1916 (13.5 mm). 91. C. variseta, spec. nov. (17.0 mm). 92. C. triseriata, spec. nov. (15.7 mm). 93. C. profundestriolata, spec. nov. (13.8 mm). 94. C. gracilipes gracilipes Sloane, 1896 (12.5 mm). 95. C. margi nata (Putzeys, 1868) (13.5 mm). 96. C. oblonga (Putzeys, 1873) (12.2 mm). 97. C. obliquicollis Sloane, 1905 (12.0 mm). 98. C. incurvicollis, spec. nov. (13.1 mm). 99. C. rectipennis, spec. nov (11.8 mm).

299 100 101 102 103

104 105 106 107

108 109 110 Figs 100-110. Habitus. Body lengths in brackets. 100. C. brevisterna Sloane, 1916 (12.2 mm). 101. C. major Sloane, 1917 (15.5 mm). 102. C. mastersi Sloane, 1896 (17.8 mm). 103. C. cobourgiana, spec. nov. (18.9 mm). 104. C. nyctosyloides Putzeys, 1868 (12.2 mm). 105. C. ovalior, spec. nov. (13.1 mm). 106. C. nitescens, spec. nov. (11.7 mm). 107. C. ovali pennis Sloane,1905 (c. 12.6 mm). 108. C. pachysoma, spec. nov. (13.2 mm). 109. C. froggatti Sloane,1896 (7.7 mm). 110. C. ha ckeri Sloane, 1907 (9.3 mm).

300 111 112 113 114

115 116 117 118

119 120 121 Figs 111-121. Habitus. Body lengths in brackets. 111. C. demarzi Baehr, 1987 (9.0 mm). 112. C. crassipennis, spec. nov. (9.6 mm). 113. C. horaki, spec. nov. (8.4 mm). 114. C. darwini Sloane, 1916 (6.2 mm). 115. C. macleayi Sloane, 1896 (8.3 mm). 116. C. horneri, spec. nov. (8.5 mm). 117. C. semirubra, spec. nov. (7.2 mm). 118. C. infans, spec. nov. (4.2 mm). 119. C. laevigata, spec. nov (5.0 mm). 120. C. uncinata, spec. nov. (4.5 mm). 121. C. normanbyensis, spec. nov. (4.95 mm).

301 122 123 124

125 126 127

128 129 130 Figs 122-130. Head and pronotum. 122. Clivina thoracica, spec. nov. 123. C. ryaceki, spec. nov. 124. C. heros, spec. nov. 125. C. newcastleana, spec. nov. 126. C. rugosifrons, spec. nov. 127. C. conicollis, spec. nov. 128. C. dubia, spec. nov. 129. C. inopinata, spec. nov. 130. C. robusta Sloane, 1905.

302 131 132

133 134 Figs 131-134. Head and pronotum. 131. C. bankae, spec. nov. 132. C. platynota, spec. nov. 133. C. vixsulcata, spec. nov. 134. C. foveifrons, spec. nov.

135 Fig. 135. Clivina triseriata, spec. nov. Base of elytra.

303 Alphabetical checklist of the species mentioned in present paper

Genus Clivina Latreille monilicornis Sloane, 1896 ...... p. 230 Subgenus Clivina Latreille monisbelli, spec. nov...... p. 221 bankae, spec. nov...... p. 251 moretona, spec. nov...... p. 278 brevisterna Sloane, 1916 ...... p. 256 newcastleana, spec. nov...... p. 226 carnabyi, spec. nov...... p. 218 nitescens, spec. nov...... p. 262 cobourgiana, spec. nov...... p. 259 normanbyensis, spec. nov...... p. 283 conicollis, spec. nov...... p. 231 nyctosyloides Putzeys, 1868 ...... p. 260 cooindae, spec. nov...... p. 213 = propinqua (nom. nov.) crassipennis, spec. nov...... p. 268 = interstitialis Sloane, 1896 darwini Sloane, 1916 ...... p. 270 obliquicollis Sloane,1905 ...... p. 250 demarzi Baehr, 1987 ...... p. 267 oblonga (Putzeys, 1873) ...... p. 246 dubia, spec. nov...... p. 233 = abbreviata (Putzeys, 1873) elegans Putzeys, 1862 ...... p. 235 obscuripes (Blackburn, 1890) ...... p. 210 foveifrons, spec. nov...... p. 282 ovalior, spec. nov...... p. 261 froggatti Sloane, 1896 ...... p. 265 ovalipennis Sloane,1905 gemina gemina, spec. nov...... p. 216 (= ovipennis Sloane, 1896) ...... p. 263 gemina nigripes, subspec. nov...... p. 218 pachysoma, spec. nov...... p. 264 gerstmeieri planior, subspec. nov...... p. 277 platynota, spec. nov...... p. 254 glabripennis, spec. nov...... p. 228 procera Putzeys, 1866 ...... p. 207 goldingi, spec. nov...... p. 225 = prominens Putzeys, 1866 (syn. nov.) gracilipes gracilipes Sloane, 1896 ...... p. 242 profundestriolata, spec. nov...... p. 241 gracilipes longior, subspec. nov...... p. 244 rectipennis, spec. nov...... p. 255 hackeri Sloane, 1907 ...... p. 266 regularis Sloane, 1896 ...... p. 233 heros, spec. nov...... p. 224 robusta Sloane, 1905 ...... p. 248 horaki, spec. nov...... p. 269 = macleayana (nom. nov.) horneri, spec. nov...... p. 272 = foveiceps (Macleay, 1863) incurvicollis, spec. nov...... p. 252 rugosifrons, spec. nov...... p. 229 infans, spec. nov...... p. 276 ryaceki, spec. nov...... p. 223 inopinata, spec. nov...... p. 240 semirubra, spec. nov...... p. 275 interposita, spec. nov...... p. 236 sinuicola, spec. nov...... p. 215 kershawi Sloane, 1916 ...... p. 238 subrufipes, spec. nov...... p. 212 laevigata, spec. nov...... p. 280 thoracica, spec. nov...... p. 216 macleayi Sloane, 1896 ...... p. 271 triseriata, spec. nov...... p. 240 mahoni, spec. nov...... p. 220 uncinata, spec. nov...... p. 281 major Sloane, 1917 (stat. nov.) ...... p. 257 variseta, spec. nov...... p. 239 marginata (Putzeys, 1868) ...... p. 245 victoriae, spec. nov...... p. 222 mastersi Sloane,1896 ...... p. 258 vixsulcata, spec. nov...... p. 273 micans, spec. nov...... p. 228 windjanae, spec. nov...... p. 219

304