Herminium Monorchis (L.) R
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Notes on Identification Works and Difficult and Under-Recorded Taxa
Notes on identification works and difficult and under-recorded taxa P.A. Stroh, D.A. Pearman, F.J. Rumsey & K.J. Walker Contents Introduction 2 Identification works 3 Recording species, subspecies and hybrids for Atlas 2020 6 Notes on individual taxa 7 List of taxa 7 Widespread but under-recorded hybrids 31 Summary of recent name changes 33 Definition of Aggregates 39 1 Introduction The first edition of this guide (Preston, 1997) was based around the then newly published second edition of Stace (1997). Since then, a third edition (Stace, 2010) has been issued containing numerous taxonomic and nomenclatural changes as well as additions and exclusions to taxa listed in the second edition. Consequently, although the objective of this revised guide hast altered and much of the original text has been retained with only minor amendments, many new taxa have been included and there have been substantial alterations to the references listed. We are grateful to A.O. Chater and C.D. Preston for their comments on an earlier draft of these notes, and to the Biological Records Centre at the Centre for Ecology and Hydrology for organising and funding the printing of this booklet. PAS, DAP, FJR, KJW June 2015 Suggested citation: Stroh, P.A., Pearman, D.P., Rumsey, F.J & Walker, K.J. 2015. Notes on identification works and some difficult and under-recorded taxa. Botanical Society of Britain and Ireland, Bristol. Front cover: Euphrasia pseudokerneri © F.J. Rumsey. 2 Identification works The standard flora for the Atlas 2020 project is edition 3 of C.A. Stace's New Flora of the British Isles (Cambridge University Press, 2010), from now on simply referred to in this guide as Stae; all recorders are urged to obtain a copy of this, although we suspect that many will already have a well-thumbed volume. -
Evolutionary Consequences of Dioecy in Angiosperms: the Effects of Breeding System on Speciation and Extinction Rates
EVOLUTIONARY CONSEQUENCES OF DIOECY IN ANGIOSPERMS: THE EFFECTS OF BREEDING SYSTEM ON SPECIATION AND EXTINCTION RATES by JANA C. HEILBUTH B.Sc, Simon Fraser University, 1996 A THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY in THE FACULTY OF GRADUATE STUDIES (Department of Zoology) We accept this thesis as conforming to the required standard THE UNIVERSITY OF BRITISH COLUMBIA July 2001 © Jana Heilbuth, 2001 Wednesday, April 25, 2001 UBC Special Collections - Thesis Authorisation Form Page: 1 In presenting this thesis in partial fulfilment of the requirements for an advanced degree at the University of British Columbia, I agree that the Library shall make it freely available for reference and study. I further agree that permission for extensive copying of this thesis for scholarly purposes may be granted by the head of my department or by his or her representatives. It is understood that copying or publication of this thesis for financial gain shall not be allowed without my written permission. The University of British Columbia Vancouver, Canada http://www.library.ubc.ca/spcoll/thesauth.html ABSTRACT Dioecy, the breeding system with male and female function on separate individuals, may affect the ability of a lineage to avoid extinction or speciate. Dioecy is a rare breeding system among the angiosperms (approximately 6% of all flowering plants) while hermaphroditism (having male and female function present within each flower) is predominant. Dioecious angiosperms may be rare because the transitions to dioecy have been recent or because dioecious angiosperms experience decreased diversification rates (speciation minus extinction) compared to plants with other breeding systems. -
European Glacial Relict Snails and Plants: Environmental Context of Their Modern Refugial Occurrence in Southern Siberia
bs_bs_banner European glacial relict snails and plants: environmental context of their modern refugial occurrence in southern Siberia MICHAL HORSAK, MILAN CHYTRY, PETRA HAJKOV A, MICHAL HAJEK, JIRI DANIHELKA, VERONIKA HORSAKOV A, NIKOLAI ERMAKOV, DMITRY A. GERMAN, MARTIN KOCI, PAVEL LUSTYK, JEFFREY C. NEKOLA, ZDENKA PREISLEROVA AND MILAN VALACHOVIC Horsak, M., Chytry, M., Hajkov a, P., Hajek, M., Danihelka, J., Horsakov a,V.,Ermakov,N.,German,D.A.,Ko cı, M., Lustyk, P., Nekola, J. C., Preislerova, Z. & Valachovic, M. 2015 (October): European glacial relict snails and plants: environmental context of their modern refugial occurrence in southern Siberia. Boreas, Vol. 44, pp. 638–657. 10.1111/bor.12133. ISSN 0300-9483. Knowledge of present-day communities and ecosystems resembling those reconstructed from the fossil record can help improve our understanding of historical distribution patterns and species composition of past communities. Here, we use a unique data set of 570 plots explored for vascular plant and 315 for land-snail assemblages located along a 650-km-long transect running across a steep climatic gradient in the Russian Altai Mountains and their foothills in southern Siberia. We analysed climatic and habitat requirements of modern populations for eight land-snail and 16 vascular plant species that are considered characteristic of the full-glacial environment of central Europe based on (i) fossil evidence from loess deposits (snails) or (ii) refugial patterns of their modern distribu- tions (plants). The analysis yielded consistent predictions of the full-glacial central European climate derived from both snail and plant populations. We found that the distribution of these 24 species was limited to the areas with mean annual temperature varying from À6.7 to 3.4 °C (median À2.5 °C) and with total annual precipitation vary- ing from 137 to 593 mm (median 283 mm). -
The Genus Habenaria (Orchidaceae) in Thailand INTRODUCTION
THAI FOR. BULL. (BOT.), SPECIAL ISSUE: 7–105. 2009. The genus Habenaria (Orchidaceae) in Thailand HUBERT KURZWEIL1 ABSTRACT. The taxonomy of the Thai species of the largely terrestrial orchid genus Habenaria Willd. is reviewed. Forty-six species are recognised. H. humidicola Rolfe, H. poilanei Gagnep. and H. ciliolaris Kraenzl. are newly recorded for Thailand based on a single collection each, although the identifi cation of the latter two is uncertain. An aberrant specimen of H. viridifl ora (Rottler ex Sw.) Lindl. is pointed out. H. erichmichaelii Christenson is reduced to synonymy under H. rhodocheila Hance. Several diffi cult and geographically widespread species complexes are identifi ed and the need for future studies of all of the available material over the entire distribution range is emphasized. Based on the herbarium and spirit material examined here the following distribution pattern emerged: about 53 % of all collections of Thai Habenaria species were made in northern Thailand (although this may partly be due to collector’s bias) and about 15 % in north-eastern Thailand, while only between 4.5 and 7.5 % come from each of the other fl oristic regions of the country. In addition, an assessment of the conservation status has been made in all species. The present study will form the basis for a later contribution to the Flora of Thailand. KEY WORDS: Habenaria, Orchidaceae, Thailand, conservation, identifi cation, morphology, systematics. INTRODUCTION Habenaria Willd. is a largely terrestrial orchid genus placed in subfamily Orchidoideae (Pridgeon et al., 2001). The genus currently accounts for about 600 species making it by far the largest in the subfamily. -
Species-Rich Grassland and National Vegetation Classification Virtual Practical
EDUCATION RESOURCE Species-rich Grassland and National Vegetation Classification Virtual Practical This virtual species-richness assessment and NVC practical uses real images that were taken through June, July and August. These are a record of species’ presence in 1x1m quadrats, but not percentage cover. Your task is to identify the species to determine if this is species-rich grassland and to classify the NVC community. You need to: • Identify the species in each image using the list below and on the next page and identification guides, for example The Wild Flower Key by Francis Rose or New Flora of the British Isles (fourth edition) by Clive Stace. • Play the video presentation and pause to examine every picture in each quadrat. Images are taken to emphasise one species, but sometimes they contain more than one. You can record all the species that you can identify from each image. • Use the data collection form to show species presence in each quadrat by putting a tick or a ‘P’ for presence in each quadrat where a species was present. • To determine whether the grassland classifies as ‘species-rich), add up the species richness of each quadrat (excluding negative indicator species as listed in the recording form) and find the mean. • To find the NVC classification, use British Plant Communities Volume 3 Grasslands and Montane Communities (Ed. J Rodwell) and the keys to the vegetation of calcicolous and mestrophic grasslands. Assess which community this sample fits into. Do you agree with this assessment and why? • Use the MAVIS software to classify the habitat type. -
Irregular Flowering Patterns in Terrestrial Orchids: Theories Vs Empirical Data
Web Ecolog y 2: 75–82. Irregular flowering patterns in terrestrial orchids: theories vs empirical data Pavel Kindlmann and Zuzana Balounová Kindlmann, P. and Balounová, Z. 2001. Irregular flowering patterns in terrestrial or- chids: theories vs empirical data. – Web Ecol. 2: 75–82. Empirical data on many species of terrestrial orchids suggest that their between-year flowering pattern is extremely irregular and unpredictable. A long search for the reason has hitherto proved inconclusive. Here we summarise and critically review the hypoth- eses that were put forward as explanations of this phenomenon: irregular flowering was attributed to costs associated with sexual reproduction, to herbivory, or to the chaotic behaviour of the system represented by difference equations describing growth of the vegetative and reproductive organs. None of these seems to explain fully the events of a transition from flowering one year to sterility or absence the next year. Data on the seasonal growth of leaves and inflorescence of two terrestrial orchid species, Epipactis albensis and Dactylorhiza fuchsii and our previous results are then used here to fill gaps in what has been published until now and to test alternative explanations of the irregular flowering patterns of orchids. P. Kindlmann ([email protected]), Faculty of Biological Sciences, Univ. of South Bohemia, Brani šovská 31, CZ-370 05 České Bud ějovice, Czech Republic (present address: Ecologie des populations et communautés, Institut National Agronomique Paris-Grignon, 16 Rue Claude Bernard, F-75231 Paris Cedex 05, France). – Z. Balounová, Faculty of Biological Sciences and Faculty of Agriculture, Univ. of South Bohemia, Brani šovská 31, CZ-370 05 České Budějovice, Czech Republic. -
Orchid Observers
Phenology of UK Plants Orchids and Zooniverse Mark Spencer & Kath Castillo Department of Life Sciences Natural History Museum Agrimonia eupatoria Robbirt & al. 2011 and UK specimens of Ophrys sphegodes Mill NHM Origins and Evolution Initiative: UK Phenology Project • 20,000 herbarium sheets imaged and transcribed • Volunteer contributed taxonomic revision, morphometric and plant/insect pollinator data compiled • Extension of volunteer work to extract additional phenology data from other UK museums and botanic gardens • 7,000 herbarium sheets curated and mounted • Collaboration with BSBI/Herbaria@Home • Preliminary analyses of orchid phenology underway Robbirt & al. (2011) . Validation of biological collections as a source of phenological data for use in climate change studies: a case study with the orchid Ophrys sphegodes. J. Ecol. Brooks, Self, Toloni & Sparks (2014). Natural history museum collections provide information on phenological change in British butterflies since the late-nineteenth century. Int. J. Biometeorol. Johnson & al. (2011) Climate Change and Biosphere Response: Unlocking the Collections Vault. Bioscience. Specimens of Gymnadenia conopsea (L.) R.Br Orchid Observers Phenology of UK Plants Orchids and Zooniverse Mark Spencer & Kath Castillo Department of Life Sciences Natural History Museum 56 species of wild orchid in the UK 29 taxa selected for this study Anacamptis morio Anacamptis pyramidalis Cephalanthera damasonium Coeloglossum viride Corallorhiza trifida Dactylorhiza fuchsii Dactylorhiza incarnata Dactylorhiza maculata Dactylorhiza praetermissa Dactylorhiza purpurella Epipactis palustris Goodyera repens Gymnadenia borealis Gymnadenia conopsea Gymnadenia densiflora Hammarbya paludosa Herminium monorchis Neotinea ustulata Neottia cordata Neottia nidus-avis Neottia ovata Ophrys apifera Ophrys insectifera Orchis anthropophora Orchis mascula Platanthera bifolia Platanthera chlorantha Pseudorchis albida Spiranthes spiralis Fly orchid (Ophrys insectifera) Participants: 1. -
Revealing the Decline and Expansion of Orchids of NW European Russia Petr G
European Journal of Environmental Sciences 7 Revealing thE dEcline aNd ExPansioN of oRchidS of NW EuRopeaN Russia Petr G. efimov Herbarium, Komarov Botanical institute of the russian Academy of Sciences, Prof. Popov str. 2, Saint-Petersburg, 192288, russia. e-mail: [email protected], [email protected];t el. and fax: +78123460839 AbstrAct A new approach to reveal the dynamics of taxa on incompletely investigated territories was developed. The decline/expansion rate of orchids in the North Western part of European Russia (Leningrad, Novgorod and Pskov Regions, an area approx. 195,000 km2) was estimat- ed using this method. The method is based on comparison of numbers of grid cells where a certain species was recorded in various time intervals using specially designed software. More than 9000 records were used, however the territory remained insufficiently and uneven- ly studied both spatially and over time. The study revealed a statistically significant (p < 0.01) decrease for Coeloglossum viride, Corallorhiza trifida, Cypripedium calceolus, Gymnadenia conopsea, Herminium monorchis, Malaxis monophyllos, Neotinea ustulata and Orchis militaris and a significant increase for Dactylorhiza baltica, D. fuchsii and Platanthera chlorantha. In several taxa, the trend was changed over the time. Of them, Gymnadenia conopsea displayed significant decline only since the middle of XX century, and Orchis militaris and Epipactis atrorubens decreased significantly only in the end of XIX and the beginning of XX century. The reasons for these patterns of dynamics were discussed. Parallels between the dynamics of orchids and land use in different periods of time in Russia are provided. Keywords: Orchidaceae, dynamics of plant species, North-West European Russia Introduction territory covers an area approx. -
Kent-Botany-2019.Pdf
0 1 Kent Botany 2019 Contents Page Introduction 1 Corrections to Kent Botany 2018 8 Plant records: selection criteria and recorders 8 Plant records for East Kent (vice county 15) 10 Plant records for West Kent (vice county 16) 27 References 35 Compiled by Geoffrey Kitchener (January 2020, web version 1) Front cover: Raphanus raphanistrum subsp. maritimus R (Sea Radish) at Minster (Sheppey). Photo 4 July 2019, © Lliam Rooney Introduction Kent Botany 2019 is the tenth report in the Kent Botany series, reporting on current botanical developments in the county. It represents a significant milestone, as 2019 brings to an end ten seasons of recording by the Kent Botanical Recording Group, founded in March 2010. It is also the end of the Botanical Society of Britain & Ireland’s (BSBI) date class 2010-19, a ten-year period for the assemblage of records which may be compared with previous ten-year date classes, to identify trends in the distribution of our flora. In addition, it is the end of the BSBI’s Atlas 2020 project, which seeks to map the current status of the British and Irish flora, following up the last mapping (Preston et al., 2002), twenty years before. This report is issued primarily as a web version, maintained on the Kent page of the BSBI website, https://bsbi.org/kent, and this should be regarded as the definitive version. The text, substantially the same, is also published as hard copy within the Kent Field Club (KFC) Bulletin. Highlights Highlights for 2019 included the following: Juncus ranarius (Frog Rush) has been restored to the Kent flora, with the discovery of a population at Crossness; Juncus x surrejanus (J. -
Global Relationships Between Plant Functional Traits and Environment in Grasslands
GLOBAL RELATIONSHIPS BETWEEN PLANT FUNCTIONAL TRAITS AND ENVIRONMENT IN GRASSLANDS EMMA JARDINE A thesis submitted in partial fulfilment of the requirements for the degree of Doctor of Philosophy The University of Sheffield Department of Animal and Plant Sciences Submission Date July 2017 ACKNOWLEDGMENTS First of all I am enormously thankful to Colin Osborne and Gavin Thomas for giving me the opportunity to undertake the research presented in this thesis. I really appreciate all their invaluable support, guidance and advice. They have helped me to grow in knowledge, skills and confidence and for this I am extremely grateful. I would like to thank the students and post docs in both the Osborne and Christin lab groups for their help, presentations and cake baking. In particular Marjorie Lundgren for teaching me to use the Licor, for insightful discussions and general support. Also Kimberly Simpson for all her firey contributions and Ruth Wade for her moral support and employment. Thanks goes to Dave Simpson, Maria Varontsova and Martin Xanthos for allowing me to work in the herbarium at the Royal Botanic Gardens Kew, for letting me destructively harvest from the specimens and taking me on a worldwide tour of grasses. I would also like to thank Caroline Lehman for her map, her useful comments and advice and also Elisabeth Forrestel and Gareth Hempson for their contributions. I would like to thank Brad Ripley for all of his help and time whilst I was in South Africa. Karmi Du Plessis and her family and Lavinia Perumal for their South African friendliness, warmth and generosity and also Sean Devonport for sharing all the much needed teas and dub. -
Chromosome Numbers and Polyploidy Events in Korean Non-Commelinids Monocots: a Contribution to Plant Systematics
pISSN 1225-8318 − Korean J. Pl. Taxon. 48(4): 260 277 (2018) eISSN 2466-1546 https://doi.org/10.11110/kjpt.2018.48.4.260 Korean Journal of REVIEW Plant Taxonomy Chromosome numbers and polyploidy events in Korean non-commelinids monocots: A contribution to plant systematics Tae-Soo JANG* and Hanna WEISS-SCHNEEWEISS1 Department of Biological Science, College of Bioscience and Biotechnology, Chungnam National University, Daejeon 34134, Korea 1Department of Botany and Biodiversity Research, University of Vienna, A-1030 Vienna, Austria (Received 4 June 2018; Revised 9 September 2018; Accepted 16 December 2018) ABSTRACT: The evolution of chromosome numbers and the karyotype structure is a prominent feature of plant genomes contributing to or at least accompanying plant diversification and eventually leading to speciation. Polyploidy, the multiplication of whole chromosome sets, is widespread and ploidy-level variation is frequent at all taxonomic levels, including species and populations, in angiosperms. Analyses of chromosome numbers and ploidy levels of 252 taxa of Korean non-commelinid monocots indicated that diploids (ca. 44%) and tet- raploids (ca. 14%) prevail, with fewer triploids (ca. 6%), pentaploids (ca. 2%), and hexaploids (ca. 4%) being found. The range of genome sizes of the analyzed taxa (0.3–44.5 pg/1C) falls well within that reported in the Plant DNA C-values database (0.061–152.33 pg/1C). Analyses of karyotype features in angiosperm often involve, in addition to chromosome numbers and genome sizes, mapping of selected repetitive DNAs in chro- mosomes. All of these data when interpreted in a phylogenetic context allow for the addressing of evolution- ary questions concerning the large-scale evolution of the genomes as well as the evolution of individual repeat types, especially ribosomal DNAs (5S and 35S rDNAs), and other tandem and dispersed repeats that can be identified in any plant genome at a relatively low cost using next-generation sequencing technologies. -
Does Seed Modification and Nitrogen Addition Affect Seed Germination of Pulsatilla Grandis?*
ENVIRONMENTAL SCIENCES DOES SEED MODIFICATION AND NITROGEN ADDITION AFFECT SEED GERMINATION OF PULSATILLA GRANDIS?* M. Bochenková1, P. Karlík2, M. Hejcman1, P. Jiras1 1 Czech University of Life Sciences Prague, Faculty of Environmental Sciences, Department of Ecology, Czech Republic 2 Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Department of Forest Ecology, Czech Republic Pulsatilla grandis is an endangered species in the Czech Republic and is protected in whole Europe because the number of its populations is declining. One of the possible causes is the deposition of atmospheric nitrogen. In our research, we investigated how nitrogen concentrations and seed appendage removal directly affect the species’ seed germination.Seeds were allowed to –1 germinate under laboratory conditions in water solutions of NH4NO3 ranging in concentration from 0 to 4239 mg N l . They were able to germinate up to the concentration of 848 mg N l–1 even when covered with mycelium, which supports the idea that they can tolerate being strongly infected by fungi. We also found a significant positive effect of seed appendage removal on seed germination. Seeds without appendages germinated, on the average, with 11% greater probability, compared to seeds with appendages. We conclude that the germination of P. grandis is not directly affected by high N concentrations in rain wa- ter, which can range from 10 to 13 mg N l–1 near large cities. Surprisingly, low concentrations of N (up to 34 mg N l–1) might even slightly support the seed germination of P. grandis. The negative effect of N deposition on seeds is indirect and acts in conjunction with the absence of management at localities.