Contributions to the Palaeozoic and Mesozoic of the Himalaya
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A New Middle Cambrian Trilobite with a Specialized Cephalon from Shandong Province, North China
Editors' choice A new middle Cambrian trilobite with a specialized cephalon from Shandong Province, North China ZHIXIN SUN, HAN ZENG, and FANGCHEN ZHAO Sun, Z., Zeng, H., and Zhao, F. 2020. A new middle Cambrian trilobite with a specialized cephalon from Shandong Province, North China. Acta Palaeontologica Polonica 65 (4): 709–718. Trilobites achieved their maximum generic diversity in the Cambrian, but the peak of morphological disparity of their cranidia occurred in the Middle to Late Ordovician. Early to middle Cambrian trilobites with a specialized cephalon are rare, especially among the ptychoparioids, a group of libristomates featuring the so-called “generalized” bauplan. Here we describe an unusual ptychopariid trilobite Phantaspis auritus gen. et sp. nov. from the middle Cambrian (Miaolingian, Wuliuan) Mantou Formation in the Shandong Province, North China. This new taxon is characterized by a cephalon with an extended anterior area of double-lobate shape resembling a pair of rabbit ears in later ontogenetic stages; a unique type of cephalic specialization that has not been reported from other trilobites. Such a peculiar cephalon as in Phantaspis provides new insights into the variations of cephalic morphology in middle Cambrian trilobites, and may represent a heuristic example of ecological specialization to predation or an improved discoidal enrollment. Key words: Trilobita, Ptychopariida, ontogeny, specialization, Miaolingian, Paleozoic, Longgang, Asia. Zhixin Sun [[email protected]], Han Zeng [[email protected]], and Fangchen Zhao [[email protected]] (cor responding author), State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palae ontology and Center for Excellence in Life and Palaeoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; University of Chinese Academy of Sciences, Beijing 100049, China. -
Northern England Serpukhovian (Early Namurian)
1 Northern England Serpukhovian (early Namurian) 2 farfield responses to southern hemisphere glaciation 3 M.H. STEPHENSON1, L. ANGIOLINI2, P. CÓZAR3, F. JADOUL2, M.J. LENG4, D. 4 MILLWARD5, S. CHENERY1 5 1British Geological Survey, Keyworth, Nottingham, NG12 5GG, United Kingdom 6 2Dipartimento di Scienze della Terra "A. Desio", Università degli Studi di Milano, Via 7 Mangiagalli 34, Milano, 20133, Italy 8 3Instituto de Geología Económica CSIC-UCM; Facultad de Ciencias Geológicas; 9 Departamento de Paleontología; C./ José Antonio Novais 228040-Madrid; Spain 10 4NERC Isotope Geosciences Laboratory, British Geological Survey, Keyworth, 11 Nottingham, NG12 5GG, United Kingdom 12 5British Geological Survey, Murchison House, Edinburgh, United Kingdom 13 14 15 Word count 7967 16 7 figs 17 1 table 18 67 references 19 RUNNING HEADER: NAMURIAN FARFIELD GLACIATION REPONSE 1 20 Abstract: During the Serpukhovian (early Namurian) icehouse conditions were initiated 21 in the southern hemisphere; however nearfield evidence is inconsistent: glaciation 22 appears to have started in limited areas of eastern Australia in the earliest Serpukhovian, 23 followed by a long interglacial, whereas data from South America and Tibet suggest 24 glaciation throughout the Serpukhovian. New farfield data from the Woodland, 25 Throckley and Rowlands Gill boreholes in northern England allow this inconsistency to 26 be addressed. δ18O from well-preserved late Serpukhovian (late Pendleian to early 27 Arnsbergian) Woodland brachiopods vary between –3.4 and –6.3‰, and δ13C varies 28 between –2.0 and +3.2‰, suggesting a δ18O seawater (w) value of around –1.8‰ 29 VSMOW, and therefore an absence of widespread ice-caps. The organic carbon δ13C 30 upward increasing trend in the Throckley Borehole (Serpukhovian to Bashkirian; c. -
PHYLOGENY and DISTRIBUTION by LARS E
[Papers in Palaeontology, 2019, pp. 1–17] CAMBRIAN RHYNCHONELLIFORM NISUSIOID BRACHIOPODS: PHYLOGENY AND DISTRIBUTION by LARS E. HOLMER1,2 , MOHAMMAD-REZA KEBRIA-EE ZADEH3, LEONID E. POPOV4, MANSOUREH GHOBADI POUR2,4,5,J.JAVIERALVARO 6, VACHIK HAIRAPETIAN7 and ZHIFEI ZHANG1 1Shaanxi Key Laboratory of Early Life & Environments, State Key Laboratory for Continental Dynamics, Northwest University, Xi’an, 710069, China 2Department of Earth Sciences, Palaeobiology, SE-752 36, Uppsala, Sweden; [email protected] 3Department of Geology, Payame Noor University, Semnan, Iran; [email protected] 4Department of Earth Sciences, National Museum of Wales, Cathays Park, Cardiff, CF10 3NP, UK; [email protected] 5Department of Geology, Faculty of Sciences, Golestan University, Gorgan, 49138-15739, Iran; [email protected] 6Instituto de Geociencias (CSIC-UCM), Dr. Severo Ochoa 7, Madrid, 28040, Spain; [email protected] 7Department of Geology, Khorasgan (Isfahan) Branch, Islamic Azad University, PO Box 81595-158, Isfahan, Iran; [email protected] Typescript received 18 July 2018; accepted in revised form 6 November 2018 Abstract: A comprehensive review and phylogenetic analy- an early separation of the lineages of spinose and non-spi- sis of genera and species presently assigned to the rhyn- nose nisusiids. The non-spinose nisusiids probably evolved chonelliform superfamily Nisusioidea and family Nisusiidae in Laurentia by the end of Cambrian Series 4. The new suggests that this short-lived but important group of bra- nisusiid genus Bellistrophia is described. The new species chiopods first appeared in peri-Gondwana during the second Nisusia multicostata represents the first documented rhyn- half of the Cambrian Series 2, before going extinct by the chonelliform (kutorginide) brachiopod from the Miaolingian end of Drumian times. -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2018/08 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age GSSP GSSP GSSP GSSP EonothemErathem / Eon System / Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon Erathem / Era System / Period GSSA age (Ma) present ~ 145.0 358.9 ± 0.4 541.0 ±1.0 U/L Meghalayan 0.0042 Holocene M Northgrippian 0.0082 Tithonian Ediacaran L/E Greenlandian 152.1 ±0.9 ~ 635 Upper 0.0117 Famennian Neo- 0.126 Upper Kimmeridgian Cryogenian Middle 157.3 ±1.0 Upper proterozoic ~ 720 Pleistocene 0.781 372.2 ±1.6 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian Callovian 1000 Quaternary Gelasian 166.1 ±1.2 2.58 Bathonian 382.7 ±1.6 Stenian Middle 168.3 ±1.3 Piacenzian Bajocian 170.3 ±1.4 Givetian 1200 Pliocene 3.600 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Devonian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.63 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Proterozoic Neogene Sinemurian Langhian 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- 2050 Burdigalian Hettangian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 27.82 Gorstian -
1 Correlation of the Base of the Serpukhovian Stage
Correlation of the base of the Serpukhovian Stage (Carboniferous; Mississippian) in northwest Europe GEORGE D. SEVASTOPULO* & MILO BARHAM✝ *Department of Geology, Trinity College Dublin, Dublin 2, Ireland ✝Milo Barham, Department of Applied Geology, Curtin University of Technology, GPO Box U1987, Perth, WA 6845, Australia Author for correspondence: [email protected] Running head: Correlation base Serpukhovian northwest Europe Abstract - The Task Group charged with proposing the GSSP for the base of the Serpukhovian Stage (Mississippian: Lower Carboniferous) is likely to use the global First Appearance Datum (FAD: evolutionary first appearance) of the conodont Lochriea ziegleri in the lineage Lochriea nodosa-L. ziegleri for the definition and correlation of the base of the stage. It is important to establish that the FOD (First Occurrence Datum) of L. ziegleri in different basins is essentially penecontemporaneous. Ammonoids provide high-resolution biostratigraphy in the late Mississippian but their use for international correlation is limited by provincialism. However, it is possible to assess the levels of diachronism of the FOD of L. ziegleri in sections in northwest Europe using ammonoid zones. Published compilations of conodont distribution in the Rhenish Slate Mountains of Germany show the FOD of L. ziegleri in the Emstites novalis Biozone (upper part of the P2c zone of the British/Irish ammonoid biozonation) but L. ziegleri has also been reported as occurring in the Neoglyphioceras spirale Biozone (P1d zone). In the Yoredale Group of northern England, the FOD of L. ziegleri is in either the P1c or P1d zone. In NW Ireland, the oldest records of both L. nodosa and L. ziegleri are from the Lusitanoceras granosum Biozone (P2a). -
Chemostratigraphic Correlations Across the First Major Trilobite
www.nature.com/scientificreports OPEN Chemostratigraphic correlations across the frst major trilobite extinction and faunal turnovers between Laurentia and South China Jih-Pai Lin 1*, Frederick A. Sundberg2, Ganqing Jiang3, Isabel P. Montañez4 & Thomas Wotte5 During Cambrian Stage 4 (~514 Ma) the oceans were widely populated with endemic trilobites and three major faunas can be distinguished: olenellids, redlichiids, and paradoxidids. The lower–middle Cambrian boundary in Laurentia was based on the frst major trilobite extinction event that is known as the Olenellid Biomere boundary. However, international correlation across this boundary (the Cambrian Series 2–Series 3 boundary) has been a challenge since the formal proposal of a four-series subdivision of the Cambrian System in 2005. Recently, the base of the international Cambrian Series 3 and of Stage 5 has been named as the base of the Miaolingian Series and Wuliuan Stage. This study provides detailed chemostratigraphy coupled with biostratigraphy and sequence stratigraphy across this critical boundary interval based on eight sections in North America and South China. Our results show robust isotopic evidence associated with major faunal turnovers across the Cambrian Series 2–Series 3 boundary in both Laurentia and South China. While the olenellid extinction event in Laurentia and the gradual extinction of redlichiids in South China are linked by an abrupt negative carbonate carbon excursion, the frst appearance datum of Oryctocephalus indicus is currently the best horizon to achieve correlation between the two regions. Te international correlation of the traditional lower–middle Cambrian boundary has been exceedingly difcult primarily due to apparent diachroniety of the datum species used to defne the boundary refecting the endemic faunas. -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2014/02 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age Erathem / Era System / Period GSSP GSSP age (Ma) GSSP GSSA EonothemErathem / Eon System / Era / Period EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon GSSP age (Ma) present ~ 145.0 358.9 ± 0.4 ~ 541.0 ±1.0 Holocene Ediacaran 0.0117 Tithonian Upper 152.1 ±0.9 Famennian ~ 635 0.126 Upper Kimmeridgian Neo- Cryogenian Middle 157.3 ±1.0 Upper proterozoic Pleistocene 0.781 372.2 ±1.6 850 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian 1000 Callovian 166.1 ±1.2 Quaternary Gelasian 2.58 382.7 ±1.6 Stenian Bathonian 168.3 ±1.3 Piacenzian Middle Bajocian Givetian 1200 Pliocene 3.600 170.3 ±1.4 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.62 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Langhian Sinemurian Proterozoic Neogene 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- Hettangian 2050 Burdigalian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 28.1 Gorstian Oligocene Upper Norian 427.4 ±0.5 2500 Rupelian Wenlock Homerian -
Kimmigj CV 20190807
Kimmig Julien Karl Franck Kimmig Curriculum vitae 2895 Pauling Ave Apt. 334 Richland, WA 99354 Phone: 785-766-2386 Email: [email protected] Education Ph.D. in “Geology” 2014 University of Saskatchewan (Canada) Thesis title: Taxonomy, taphonomy and paleocology of a new Burgess Shale-type Lagerstätte from the Mackenzie Mountains, Northwest Territories, Canada. M.Sc. in “Biology” 2009 Topic: Advanced Methods in Taxonomy and Biodiversity Imperial College London & Natural History Museum London (United Kingdom) B.Sc. in “Geosciences” 2008 Georg-August-Universität Göttingen (Germany) Professional experience Research Affiliate since August 2019 Division of Invertebrate Paleontology University of Kansas Collections Manager (Assistant Researcher) June 2016 – July 2019 Division of Invertebrate Paleontology University of Kansas Research Associate July 2015 – June 2019 University of Saskatchewan Postdoctoral Researcher October 2014 – June 2015 Supervisor: Brian Pratt University of Saskatchewan Teaching Assistant September 2011 – September 2014 University of Saskatchewan Adjunct Researcher September 2009 – May 2011 Supervisor: Alison Longbottom The Natural History Museum London Museum and Research Assistant December 2007 – September 2008 Supervisor: Mike Reich Geowissenschaftliches Museum Göttingen 1 Kimmig Selected fieldwork (Total ~ 160 days) Cambrian, Utah and Idaho, 2019 10 days Spence Shale, Wellsville Mountains, Utah, 2018 5 days Spence Shale, Wellsville Mountains, Utah, 2017 15 days Ravens Throat River, Mackenzie Mountains, Canada, 2012 28 days -
Paleogeographic Maps Earth History
History of the Earth Age AGE Eon Era Period Period Epoch Stage Paleogeographic Maps Earth History (Ma) Era (Ma) Holocene Neogene Quaternary* Pleistocene Calabrian/Gelasian Piacenzian 2.6 Cenozoic Pliocene Zanclean Paleogene Messinian 5.3 L Tortonian 100 Cretaceous Serravallian Miocene M Langhian E Burdigalian Jurassic Neogene Aquitanian 200 23 L Chattian Triassic Oligocene E Rupelian Permian 34 Early Neogene 300 L Priabonian Bartonian Carboniferous Cenozoic M Eocene Lutetian 400 Phanerozoic Devonian E Ypresian Silurian Paleogene L Thanetian 56 PaleozoicOrdovician Mesozoic Paleocene M Selandian 500 E Danian Cambrian 66 Maastrichtian Ediacaran 600 Campanian Late Santonian 700 Coniacian Turonian Cenomanian Late Cretaceous 100 800 Cryogenian Albian 900 Neoproterozoic Tonian Cretaceous Aptian Early 1000 Barremian Hauterivian Valanginian 1100 Stenian Berriasian 146 Tithonian Early Cretaceous 1200 Late Kimmeridgian Oxfordian 161 Callovian Mesozoic 1300 Ectasian Bathonian Middle Bajocian Aalenian 176 1400 Toarcian Jurassic Mesoproterozoic Early Pliensbachian 1500 Sinemurian Hettangian Calymmian 200 Rhaetian 1600 Proterozoic Norian Late 1700 Statherian Carnian 228 1800 Ladinian Late Triassic Triassic Middle Anisian 1900 245 Olenekian Orosirian Early Induan Changhsingian 251 2000 Lopingian Wuchiapingian 260 Capitanian Guadalupian Wordian/Roadian 2100 271 Kungurian Paleoproterozoic Rhyacian Artinskian 2200 Permian Cisuralian Sakmarian Middle Permian 2300 Asselian 299 Late Gzhelian Kasimovian 2400 Siderian Middle Moscovian Penn- sylvanian Early Bashkirian -
2009 Geologic Time Scale Cenozoic Mesozoic Paleozoic Precambrian Magnetic Magnetic Bdy
2009 GEOLOGIC TIME SCALE CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST. HIST. ANOM. ANOM. (Ma) CHRON. CHRO HOLOCENE 65.5 1 C1 QUATER- 0.01 30 C30 542 CALABRIAN MAASTRICHTIAN NARY PLEISTOCENE 1.8 31 C31 251 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 70.6 254 2A PIACENZIAN 32 C32 L 630 C2A 3.6 WUCHIAPINGIAN PLIOCENE 260 260 3 ZANCLEAN 33 CAMPANIAN CAPITANIAN 5 C3 5.3 266 750 NEOPRO- CRYOGENIAN 80 C33 M WORDIAN MESSINIAN LATE 268 TEROZOIC 3A C3A 83.5 ROADIAN 7.2 SANTONIAN 271 85.8 KUNGURIAN 850 4 276 C4 CONIACIAN 280 4A 89.3 ARTINSKIAN TONIAN C4A L TORTONIAN 90 284 TURONIAN PERMIAN 10 5 93.5 E 1000 1000 C5 SAKMARIAN 11.6 CENOMANIAN 297 99.6 ASSELIAN STENIAN SERRAVALLIAN 34 C34 299.0 5A 100 300 GZELIAN C5A 13.8 M KASIMOVIAN 304 1200 PENNSYL- 306 1250 15 5B LANGHIAN ALBIAN MOSCOVIAN MESOPRO- C5B VANIAN 312 ECTASIAN 5C 16.0 110 BASHKIRIAN TEROZOIC C5C 112 5D C5D MIOCENE 320 318 1400 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 326 6 C6 APTIAN 20 120 1500 CALYMMIAN E 20.4 6A C6A EARLY MISSIS- M0r 125 VISEAN 1600 6B C6B AQUITANIAN M1 340 SIPPIAN M3 BARREMIAN C6C 23.0 345 6C CRETACEOUS 130 M5 130 STATHERIAN CARBONIFEROUS TOURNAISIAN 7 C7 HAUTERIVIAN 1750 25 7A M10 C7A 136 359 8 C8 L CHATTIAN M12 VALANGINIAN 360 L 1800 140 M14 140 9 C9 M16 FAMENNIAN BERRIASIAN M18 PROTEROZOIC OROSIRIAN 10 C10 28.4 145.5 M20 2000 30 11 C11 TITHONIAN 374 PALEOPRO- 150 M22 2050 12 E RUPELIAN -
Middle Cambrian Trilobites (Miaolingian, Ehmaniella Biozone) from the Telt Bugt Formation of Daugaard-Jensen Land, Western North Greenland
BULLETIN OF THE GEOLOGICAL SOCIETY OF DENMARK · VOL. 68 · 2020 Middle Cambrian trilobites (Miaolingian, Ehmaniella Biozone) from the Telt Bugt Formation of Daugaard-Jensen Land, western North Greenland JOHN S. PEEL Peel, J.S. 2020. Middle Cambrian trilobites (Miaolingian, Ehmaniella Biozone) from the Telt Bugt Formation of Daugaard-Jensen Land, western North Greenland. Bulletin of the Geological Society of Denmark, vol. 68, pp. 1–14. ISSN 2245-7070. https://doi.org/10.37570/bgsd-2020-68-01 A small fauna of middle Cambrian trilobites is described from the upper Telt Bugt Geological Society of Denmark Formation of Daugaard-Jensen Land, western North Greenland, and the formation https://2dgf.dk is formally defined.Blainiopsis holtedahli and Blainiopsis benthami, originally described from the equivalent Cape Wood Formation of Bache Peninsula, Nunavut, Canada, are Received 9 October 2019 documented in an assemblage assigned to the Ehmaniella Biozone (Topazan Stage of Accepted in revised form North American usage), Miaolingian Series, Wuliuan Stage, of the international stan- 5 February 2020 dard. Two new species are proposed: Ehmaniella sermersuaqensis and Clappaspis tupeq. Published online 9 March 2020 Keywords: Laurentia, North Greenland, Cambrian, Miaolingian (Wuliuan), tri- © 2020 the authors. Re-use of material is lobites. permitted, provided this work is cited. Creative Commons License CC BY: John S. Peel [[email protected]], Department of Earth Sciences (Palaeobiology), https://creativecommons.org/licenses/by/4.0/ Uppsala University, Villavägen 16, SE-75236 Uppsala, Sweden. The first Cambrian fossils from the Nares Strait setti 1951; Cooper et al. 1952; Palmer & Halley 1979). region, the narrow waterway separating northern- Several of the lithostratigraphic and biostratigraphic most Greenland and Canada, were collected from problems recognised in Poulsen’s (1927) studies were Bache Peninsula (Fig. -
Aquatic Stem Group Myriapods Close a Gap Between Molecular Divergence Dates and the Terrestrial Fossil Record
Aquatic stem group myriapods close a gap between molecular divergence dates and the terrestrial fossil record Gregory D. Edgecombea,1, Christine Strullu-Derriena,b, Tomasz Góralc,d, Alexander J. Hetheringtone, Christine Thompsonf, and Markus Kochg,h aDepartment of Earth Sciences, The Natural History Museum, London SW7 5BD, United Kingdom; bInstitut de Systématique, Evolution, Biodiversité, UMR 7205, Muséum National d’Histoire Naturelle, 75005 Paris, France; cImaging and Analysis Centre, The Natural History Museum, London SW7 5BD, United Kingdom; dCentre of New Technologies, University of Warsaw, 02-097 Warsaw, Poland; eDepartment of Plant Sciences, University of Oxford, Oxford OX1 2JD, United Kingdom; fDepartment of Natural Sciences, National Museums Scotland, Edinburgh EH1 1JF, United Kingdom; gSenckenberg Society for Nature Research, Leibniz Institution for Biodiversity and Earth System Research, 60325 Frankfurt am Main, Germany; and hInstitute for Evolutionary Biology and Ecology, University of Bonn, 53121 Bonn, Germany Edited by Conrad C. Labandeira, Smithsonian Institution, National Museum of Natural History, Washington, DC, and accepted by Editorial Board Member David Jablonski February 24, 2020 (received for review November 25, 2019) Identifying marine or freshwater fossils that belong to the stem (2–5). Despite this inferred antiquity, there are no compelling groups of the major terrestrial arthropod radiations is a long- fossil remains of Myriapoda until the mid-Silurian and no hexa- standing challenge. Molecular dating and fossils of their pancrus- pods until the Lower Devonian. In both cases, the oldest fossils tacean sister group predict that myriapods originated in the can be assigned to crown group lineages (Diplopoda in the case of Cambrian, much earlier than their oldest known fossils, but Silurian myriapods and Collembola in the case of Hexapoda) and uncertainty about stem group Myriapoda confounds efforts to the fossils have morphological characters shared by extant species resolve the timing of the group’s terrestrialization.