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Ent12 3 329 346 Grichanov.PM6 Russian Entomol. J. 12(3): 329346 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2003 New Afrotropical Sciapodinae (Diptera: Dolichopodidae) with some new synonymy Íîâûå âèäû ìóõ-çåëåíóøåê Sciapodinae (Diptera: Dolichopodidae) èç òðîïè÷åñêîé Àôðèêè è íåñêîëüêî íîâûõ ñèíîíèìîâ I.Ya. Grichanov È.ß. Ãðè÷àíîâ All-Russian Institute of Plant Protection, Russian Academy of Agricultural Sciences, Podbelskogo 3, St.Petersburg-Pushkin 189620 Russia. E-mail: [email protected] Âñåðîññèéñêèé íàó÷íî-èññëåäîâàòåëüñêèé èíñòèòóò çàùèòû ðàñòåíèé ÐÀÑÕÍ, øîññå Ïîäáåëüñêîãî 3, Ñ.-Ïåòåðáóðã Ïóøêèí 196608 Ðîññèÿ. KEY WORDS: Diptera, Dolichopodidae, Sciapodinae, new species, new synonym, new combination, new name, key, Tropical Africa. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: Diptera, Dolichopodidae, Sciapodinae, Òðîïè÷åñêàÿ Àôðèêà, îïðåäåëèòåëü, íîâûå âèäû, íîâûå ñèíîíèìû. ABSTRACT. A new abundant material has been â Ïàðèæå è íåêîòîðûõ äðóãèõ ìóçååâ.  ñòàòüå îïè- examined mainly from the collections of the National ñàíû 13 íîâûõ âèäîâ: Mascaromyia alexisi sp.n., M. Museum of Natural History in Paris. Descriptions of 13 bebourensis sp.n., M. brooksi sp.n., M. cummingi sp.n., new species are given, and three species are replaced. M. grimaldii sp.n., M. loici sp.n., M. tatyanae sp.n., M. Mascaromyia alexisi sp.n., M. bebourensis sp.n., M. michaeli sp.n., Amblypsilopus reunionensis sp.n. è A. brooksi sp.n., M. cummingi sp.n., M. grimaldii sp.n., M. takamaka sp.n. ñ îñòðîâà Ðåþíüîí, A. ambila sp.n. è A. loici sp.n., M. tatyanae sp.n., M. michaeli sp.n., Amblyp- ranomafana sp.n. ñ Ìàäàãàñêàðà, Chrysosoma (Chry- silopus reunionensis sp.n. and A. takamaka sp.n. are sosoma) nguemba sp.n. èç Êàìåðóíà. Chrysosoma described from Reunion, A. ambila sp.n. and A. ranoma- (Mesoblepharia) albocrinitatum Curran, 1925 âïåð- fana sp.n. from Madagascar, Chrysosoma (Chrysoso- âûå îòìå÷àåòñÿ äëÿ Êàìåðóíà. Sciapus nubilis Parent, ma) nguemba sp.n. from Cameroon. Chrysosoma (Me- 1935 ïåðåìåùåí â ðîä Dytomyia Bickel, 1994, Sciapus soblepharia) albocrinitatum Curran, 1925 is recorded aenescensVanschuytbroeck, 1952 â ðîä Ethiosciapus for Cameroon for the first time. Sciapus nubilis Parent, Bickel, 1994, à Agonosoma micantifrons Speiser, 1910 1935 is transferred to the genus Dytomyia Bickel, 1994, â ðîä Plagiozopelma Enderlein, 1912. Ïðåäëîæåíû Sciapus aenescens Vanschuytbroeck, 1952 to Ethios- ñëåäóþùèå ñèíîíèìû: Mascaromyia leptogaster ciapusBickel, 1994, andAgonosoma micantifronsSpeis- (Thomson, 1869) (=Psilopus librativertex Lamb, 1922); er, 1910 to Plagiozopelma Enderlein, 1912. The follow- Bickelia parallela (Macquart, 1842) (=Sciapus guerini ing pairs of species are synonymized: Mascaromyia Parent, 1935; =Bickelia subparallela Grichanov, 1996); leptogaster (Thomson, 1869) (=Psilopus librativertex Ethiosciapus latipes (Parent, 1929) (=Sciapus aenescens Lamb, 1922); Bickelia parallela (Macquart, 1842) Vanschuytbroeck, 1952); Chrysosoma (Chrysosoma) (=Sciapus guerini Parent, 1935; =Bickelia subparallela vividum Becker, 1923 (=Chrysosoma tarsiciliatum Grichanov, 1996); Ethiosciapus latipes (Parent, 1929) Parent, 1929); Chrysosoma (Kalocheta) villiersi (=Sciapus aenescens Vanschuytbroeck, 1952); Chry- (Vanschuytbroeck, 1970) (=Kalocheta collarti Parent, sosoma (Chrysosoma) vividum Becker, 1923 (=Chry- 1933, nec Curran, 1927; =Chrysosoma (Kalocheta) sosoma tarsiciliatum Parent, 1929); Chrysosoma alberti Meuffels & Grootaert, 1999). Ïðåäëîæåíî òàê- (Kalocheta) villiersi (Vanschuytbroeck, 1970) æå íîâîå íàçâàíèå Condylostylus erroneus Grichanov, (=Kalocheta collarti Parent, 1933, nec Curran, 1927; =Chrysosoma (Kalocheta) alberti Meuffels & Groot- nom. nov. äëÿ Condylostylus imitans Curran, 1926. aert, 1999). A new name Condylostylus erroneus Grich- Ïðèâåäåí îïðåäåëèòåëü èçâåñòíûõ âèäîâ ðîäà Ma- anov, nom. nov. is proposed for Condylostylus imitans scaromyia Bickel, 1994.  ðåçóëüòàòå èññëåäîâàíèÿ Curran, 1926. A key to known species of Mascaromyia ÷èñëî àôðîòðîïè÷åñêèõ âèäîâ ïîäñåìåéñòâà äîñ- Bickel, 1994 is also given. Presently 13 genera and 225 òèãëî 225, îáúåäèíåííûõ â 13 ðîäîâ. Afrotropical species of Sciapodinae are known. Introduction ÐÅÇÞÌÅ. Èññëåäîâàí íîâûé ìàòåðèàë èç êîë- ëåêöèé Íàöèîíàëüíîãî ìóçåÿ åñòåñòâåííîé èñòîðèè The world fauna of sciapodine genera was recently Printed in 2004. revised by Bickel [1994]. Afrotropical species of the 330 I.Ya. Grichanov subfamily were reviewed by Grichanov [1998, 1999], species of the genus Dytomyia are known from Australia Negrobov and Grichanov [1998]. Treating material from and New Guinea and five species from Madagascar. the rich collections of the National Museum of Natural Parentia is speciose in Australia and adjacent islands History, Paris [MNHP] and the Swedish Museum of with about 60 known species [Bickel, 1994]. It is the Natural History, Stockholm (NHRS), I have found a new dominant sciapodine element in the New Zealand fauna, abundant material on the subfamily Sciapodinae. Prima- showing its possible Gondwanan origin. Afrotropical ry types of 28 Afrotropical species and paratypes of species of the genus are confined to southern Africa. additional four species have been examined in the so- Bickelia and Mascaromyia are genera endemic of the called Collection of Macquart, Collection of Parent and western Indian Ocean islands. Monotypic Bickelia and General Collection (MNHP), Collections of Thomson, Mascaromyia leptogaster are distributed on Mauritius, Loew and Speiser (NHRS). All these are listed below. Seychelles and Chagos Archipelago. The other Mascar- Lectotypes have been designated for some species. omyia species are endemic of certain islands. The fauna of Descriptions of 13 new species, some new records for Mauritius comprises 12 species; Reunion numbers 9, known African species are given here, and three species Seychelles 5 species; only one species have been recorded are replaced. Mascaromyia alexisi sp.n., M. bebouren- from Rodriguez. The real number of species of the genus sis sp.n., M. brooksi sp.n., M. cummingi sp.n., M. grim- may be doubled following a special expedition covering aldii sp.n., M. loici sp.n., M. michaeli sp.n., M. tatyanae yet unexplored islands. Mascaromyia is the typical sp.n., Amblypsilopus reunionensis sp.n. and A. takama- example of adaptive radiation with Bickelia parallela ka sp.n. are described from Reunion, A. ambila sp.n. and being older colonist and M. leptogaster more recent, A. ranomafana sp.n. from Madagascar, Chrysosoma while the other species of the genus are neo-endemics. (Chrysosoma) nguemba sp.n. from Cameroon. Sciapus nubilis Parent, 1935 is transferred to the genus Dytomy- Deposition of types of the new species in the collec- ia Bickel, 1994, Sciapus aenescens Vanschuytbroeck, tions of the National Museum of Natural History, Paris 1952 to Ethiosciapus Bickel, 1994, and Agonosoma (MNHP) and the Royal Belgian Institute for Natural micantifrons Speiser, 1910 to Plagiozopelma Ender- Sciences, Brussels (ISNB) is mentioned under the new lein, 1912. The following pairs of species are synony- names. The other material examined is deposited in the mized: Mascaromyia leptogaster (Thomson, 1869) Museum of Natural History in Stockholm (NHRS) and (=Psilopus librativertex Lamb, 1922); Bickelia paralle- the Zoological Museum in Lund University (MZLU). la (Macquart, 1842) (=Sciapus guerini Parent, 1935; Diagnoses are given for species not listed in the previous =Bickelia subparallela Grichanov, 1996); Ethiosciapus reviews [Grichanov, 1998, 1999; Negrobov & Gricha- latipes (Parent, 1929) (=Sciapus aenescens Vanschuyt- nov, 1998]. They include usually key characters and broeck, 1952); Chrysosoma (Chrysosoma) vividum some important features that were missed in original Becker, 1923 (=Chrysosoma tarsiciliatum Parent, 1929); descriptions. The relative lengths of the podomeres are Chrysosoma (Kalocheta) villiersi (Vanschuytbroeck, representative ratios and not measurements (1 mm = 78). 1970) (=Kalocheta collarti Parent, 1933, nec Curran, Hypopygia removed from the dry specimens are placed 1927; =Chrysosoma (Kalocheta) alberti Meuffels & after alkalisation into glycerol and mounted on the same Grootaert, 1999). A new name Condylostylus erroneus pin in a cavity of polymer film covered with a piece of Grichanov, nom.n. is proposed for Condylostylus imi- adhesive tape. Listing material examined, I use here tans Curran, 1926. A key to known species of Mascar- slash ( / ) to separate labels on one pin and square brackets omyia Bickel, 1994 is also given. [...] to insert my personal remarks. Bibliography includes works published after the Catalogue of the Diptera of the Afrotropical Region [Dyte & Smith, 1980]. Now 13 genera and 225 Afrotropical species of Sciapodinae are presently known. It is the largest doli- chopodid subfamily in the Region. The number of Afro- Taxonomy tropical species belonging to sciapodine genera are as follows: Chrysosoma Guérin-Méneville, 1831 (67), TRIBE MESORHAGINI (SCHINER) BICKEL Amblypsilopus Bigot, 1888 (48), Mascaromyia Bickel, 1994 (28), Condylostylus Bigot, 1859 (19), Plagiozo- Genus Mesorhaga Schiner, 1868 pelma (17), Gigantosciapus Grichanov, 1997 (13), 1. Mesorhaga pauliani Vanschuytbroeck Ethiosciapus (9), Mesorhaga Schiner, 1868 (7), Bicke- liolus Grichanov, 1996 (6), Dytomyia (5), Parentia Mesorhaga pauliani Vanschuytbroeck, 1952: 137. Hardy, 1935 (4), Bickelia Grichanov, 1996 (1) and Type material examined. [$], Holotype [red label] / I. S. Sciapus Zeller, 1842 (1). These predatory flies are Madagascar, Fort dAmbohitantely, December 1947, R.P. / P. distributed throughout the continent
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