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Home , Brood II, Cicadettini, Cicadidae, Okanagana canadensis

John R. Cooley Department of Ecology and Evolutionary Biology The University of Connecticut Storrs, CT USA 06269

Teaching Interests: Active learning, peer groups, technological engagement.

Research Interests: Speciation, secondary contact zones, sexual selection, GIS, species distribution modeling.

Current positions: Researcher and Adjunct Faculty, Department of Ecology and Evolutionary Biology, The University of Connecticut, Storrs, CT. Visiting Professor of Integrative Sciences, College of Integrative Sciences, Wesleyan University, Middletown CT.

Positions and Teaching:

Wesleyan University (Visiting Assistant of Integrative Sciences) CIS 150: The Art of Scientific Writing (2016, 2017, 2018). CIS 520: Academic Writing and Publishing (2016, 2017).

The University of Connecticut (Assistant Professor in Residence, Instructional Specialist, Adjunct Professor) EEB 2202: Evolution and Human Diversity (2016). EEB 2208: Conservation Biology (2018) EEB 2214: Biology of the Vertebrates (2004-2005). EEB 2244: General Ecology (2011, 2013, 2016). EEB 2244W: Writing section for General Ecology (2014, 2015). EEB 2245: Evolutionary Biology (2007). EEB 2245W: Writing section for Evolutionary Biology (2005-2007, 2014, 2015). EEB 2293WC: Methods of Field Ecology (2005-2006). BIO 1102: Foundations of Biology (2005-2006, 2016). BIO 1108: Principles of Biology II (2004-2006). BIO 1108: Laboratory for Introductory Biology (2012). BIO 1109: Topics in Modern Biology (Honors; 2014). BIO 2289: Introduction to Undergraduate Research (Honors; 2014, 2015, 2018).

The Ohio State University (Visiting Assistant Professor) BIOL 114: Form, Function, Diversity and Ecology (2010). EEOB 370: Extinction (2010). EEOB 405.01: Diversity and Systematics of Organisms (2009).

1 Yale University (Lecturer) E&EB 115a: Conservation Biology (2008). E&EB 220a: General Ecology (2008). E&EB 230a: Field Ecology (2008). E&EB 255b: Biology of the Invertebrates (2009). E&EB 256Lb: Laboratory for Biology of the Invertebrates (2009).

The University of Rhode Island (Adjunct Professor) Bio 272: General Evolution (2011).

Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan (Visiting Professor) Seminar in Field Ecology Research (2010).

Corinthian Colleges (Adjunct Instructor) EVS 1001: Environmental Science (2011-2012).

The University of Michigan (Graduate Student Teaching Assistant) BIO 154: Introduction to Biology. BIO 195: Introduction to Biology. BIO 305: Genetics. BIO 494: Evolution and Human Behavior.

Education:

M.B.A. School of Business, The University of Connecticut, Storrs, CT USA. Degree concentration in marketing and data analytics. Awards: Beta Gamma Sigma, Dean’s Review. (2014)

Postdoctoral Researcher Department of Ecology and Evolutionary Biology, The University of Connecticut, Storrs, CT USA. Advisor: Dr. Chris Simon. Project: NSF-DEB 99-82039 “Allochronic speciation, reproductive character displacement, and sexual selection in periodical (Magicicada spp.)”

Ph.D. Department of Biology, The University of Michigan, Ann Arbor, MI USA. Advisor: Dr. Richard D. Alexander. Thesis: Sexual behavior in North American cicadas of the genera Magicicada and . Awards: Regents’ Fellowship. (1999)

M.S. Department of Biology, The University of Michigan, Ann Arbor, MI USA. Research: Sexual behavior of syrphid flies in Colorado alpine tundra. (1992)

2 B.S. Department of Biology, Yale University, New Haven, CT USA. Advisors: Drs. Rob DeSalle and Charles Remington. Thesis: Biochemical approaches to phylogeny among the orders Hymenoptera, Coleoptera, Neuroptera, and Megaloptera. Awards: Honors, Summa Cum Laude, Phi Beta Kappa. (1990)

Publications: Cooley, J. R., D. C. Marshall, and K. B. R. Hill. 2018. A specialized fungal parasite (Massospora cicadina) hijacks the sexual signals of (: : Magicicada). Scientific Reports 8: 1432. Fujisawa, T., T. Koyama, S. Kakishima, J. R. Cooley, C. Simon, J. Yoshimura, and T. Sota. 2018. Triplicate parallel life cycle divergence despite gene flow in periodical cicadas. Communications Biology 1:26. Marshall, D. C., M. Moulds, K. B. R. Hill, M. H. Villet, B. Price, E. Wade, C. L. Owen, G. Goemans, K. Marathe, V. Sarkar, J. R. Cooley, K. Kunte, and C. Simon. 2018. A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa. Accepted. Marshall, D. C., K. B. R. Hill, and J. R. Cooley. 2018. Multimodal life cycle variation in 13- and 17-year periodical cicadas (Magicicada spp.). Journal Of The Kansas Entomological Society. Accepted. Kritsky, G., R. Troutman, D. Mozgai, C. Simon, S. M. Chiswell, S. Kakishima, T. Sota, J. Yoshimura, and J. R. Cooley. 2017. Evolution and Geographic Extent of a Surprising Northern Disjunct Population of 13-year Brood XXII (Hemiptera: Cicadidae, Magicicada). The American Entomologist 63: E15-E20. Koyama, T., H. Ito, T. Fujisawa, H. Ikeda, S. Kakishima, J. R. Cooley, C. Simon, J. Yoshimura, and T. Sota. 2016. Genomic divergence and lack of introgressive hybridization between two 13-year periodical cicadas supports life-cycle switching in the face of climate change. Mol. Ecol. 25: 5543-5556. Cooley, J. R., G. Kritsky, D. C. Marshall, K. B. R. Hill, G. J. Bunker, M. L. Neckermann, J. Yoshimura, J. E. Cooley, and C. Simon. 2016. A GIS-based map of periodical cicada Brood XIII in 2007, with notes on adjacent populations of Broods III and X (Hemiptera: Magicicada spp.). The American Entomologist 62:241-246. Cooley, J. R., C. Simon, C. Maier, D. C. Marshall, J. Yoshimura, S. M. Chiswell, M. D. Edwards, C. W. Holliday, R. Grantham, J. D. Zyla, R. L. Sanders, M. L. Neckermann, and G. J. Bunker. 2015. The distribution of periodical cicada (Magicicada) Brood II in 2013: Disjunct emergences suggest complex origins. The American Entomologist 61:245-251. Marshall, D. C., K. B. R. Hill, M. Moulds, D. Vanderpool, J. R. Cooley, A. B. Mohagan, and C. Simon. 2015. Inflation of molecular clock rates and dates: Biogeography, molecular phylogenetics, and diversification of a global cicada radiation from Australasia (Hemiptera: Cicadidae: ). Syst. Biol. 65(1): 16-34

3 Ito, H., S. Kakishima, T. Uehara, S. Morita, T. Koyama, T. Sota, J. R. Cooley, and J. Yoshimura. 2015. Evolution of periodicity in periodical cicadas. Scientific Reports 5:14094. Koyama, T., H. Ito, S. Kakishima, J. Yoshimura, J. R. Cooley, C. Simon, and T. Sota. 2015. Geographic body size variation in the periodical cicadas Magicicada: implications for life cycle divergence and local adaptation. Journal of Evolutionary Biology 28: 1270- 1277. Cooley, J. R. 2015. The distribution of periodical cicada (Magicicada) Brood I in 2012, with new, previously unreported populations (Hemiptera: Cicadidae). The American Entomologist 61: 52-57. Hughes, Derke A., Richard A. Katz, Albert H. Nuttall, and John R. Cooley. Bio-acoustic Wave Energy Transducer. The United States Of America As Represented By The Secretary Of The Navy, assignee. Patent US 8,611,190 B1. 17 Dec. 2013. Print. Sota, T., S. Yamamoto, J. R. Cooley, K. B. R. Hill, C. Simon, and J. Yoshimura. 2013. Independent divergence of 13- and 17-y life cycles among three periodical cicada lineages. PNAS 110 (17): 6919-6924. Cooley, J.R., Marshall, D.C., Richards, A.F., Alexander, R.D., Irwin, M.D., Coelho, J.R. and Simon, C. 2013. The distribution of periodical cicada Brood III in 1997, with special emphasis on Illinois (Hemiptera: Magicicada spp.). The American Entomologist 59: 9-14. Cooley, J. R., M. L. Neckermann, G. Bunker, D. C. Marshall, and C. Simon. 2013. At the limits: Habitat suitability modeling of Northern 17-year periodical cicada extinctions (Hemiptera: Magicicada spp.). Global Ecology and Biogeography 22: 410-421. Nariai, Y., S. Hayashi, S. Morita, Y. Umemura, K. Tainaka, T. Sota, J. R. Cooley, and J. Yoshimura. 2011. Life cycle shifting by gene introduction under an Allee effect in periodical cicadas. PLoS ONE 6(4): e18347. doi:10.1371/journal.pone.0018347. Cooley, J. R., G. Kritsky, M. J. Edwards, J. D. Zyla, D. C. Marshall, K. B. R. Hill, G. Bunker, M. L. Neckermann, R. Troutman, J. Yoshimura, and C. Simon. 2011. Periodical cicadas (Magicicada spp.): A GIS-based map of Broods XIV in 2008 and “XV” in 2009. The American Entomologist 57 (3): 144-151. Marshall, D. C., J. R. Cooley, and K. B. R. Hill. 2011. Developmental Plasticity of Life- Cycle Length in Thirteen-Year Periodical Cicadas (Hemiptera: Cicadidae). Annals of the Entomological Society of America. 104(3): 443-450. Marshall, D. C., K. B. R. Hill, J. R. Cooley, C. Simon. 2011. Hybridization, Mitochondrial DNA , and Prediction of the Early Stages of Reproductive Isolation: Lessons from New Zealand Cicadas of the Genus Kikihia. Systematic Biology 60(4): 482-502. Cooley, John R. and P. Fonseca. 2010. Exploiting the bioacoustics of cicadas to enhance the efficiency of sound transmission: Methodology and findings. US Navy-NUWC internal report, 19 March 2010. Smits, A., J. R. Cooley, E. Westerman. 2010. Twig to Root: Eggnest density and underground nymph distribution in a periodical cicada (Hemiptera: L.). Entomologica Americana 116: 73-77.

4 Cooley, J. R., G. Kritsky, M. J. Edwards, J. D. Zyla, D. C. Marshall, K. B. R. Hill, R. Krauss, and C. Simon. 2009. The Distribution of Periodical Cicada in 2004. The American Entomologist 55(2): 106-112. Tanaka, Y., J. Yoshimura, C. Simon, J. R. Cooley, and K. Tainaka. 2009. The Allee effect in the selection for prime-numbered cycles in periodical cicadas. PNAS 106 (22): 8975- 8979. Marshall, D. C., K. Slon, J. R. Cooley, K. B. R. Hill, C. Simon. 2008. Steady Plio-Pleistocene diversification and a 2-Million Year sympatry threshold in a New Zealand cicada radiation. Molecular Phylogenetics and Evolution 48: 1054-1066. Fontaine, K. M, J. R. Cooley, and C. Simon. 2007. Evidence for paternal leakage in hybrid periodical cicadas (Hemiptera: Magicicada spp.). PLoS One 9: e892 (DOI: pone.0000892). Bunker, G. J., M. L. Neckermann, and J. R. Cooley. 2007. New northern records of chloromera (Hemiptera: Cicadidae) in Connecticut. Transactions of the American Entomological Society 133: 357-361. Cooley, J. R. 2007. Decoding asymmetries in reproductive character displacement. Proceedings of the Academy of Natural Sciences of Philadelphia 156: 89-96. Cooley, J. R., D. Marshall, C. Simon, and K. Hill. 2006. Reconstructing Asymmetrical Reproductive Character Displacement In A Periodical Cicada Contact Zone. Journal of Evolutionary Biology 19: 855-868. Hill, K. B. R., D. C. Marshall, and J. R. Cooley. 2005. Crossing Cook Strait: Probable human transportation of two New Zealand cicadas Kikihia scutellaris (Walker) and K. ochrina (Walker) (Hemiptera: : Cicadidae) from the North Island to the South Island. New Zealand Entomologist 28: 71-80. Cooley, J. R., D. C. Marshall, and C. Simon. 2004. The historical contraction of periodical cicada Brood VII (Hemiptera: Cicadidae: Magicicada). Journal of the New York Entomological Society 112: 198-204. Cooley, J. R., and D. C. Marshall. 2004. Thresholds or comparisons: Mate choice criteria and sexual selection in a periodical cicada, Magicicada septendecim (Hemiptera: Cicadidae). Behaviour 141: 647-673. Cooley, J. R. 2004. Asymmetry and mating success in a periodical cicada, Magicicada septendecim (Hemiptera: Cicadidae). Ethology 110: 1-17. Cooley, J. R., C. Simon, and D. C. Marshall. 2003. Temporal separation and speciation in periodical cicadas. BioScience 53: 151-157. Marshall, D. C., J. R. Cooley, and C. Simon. 2003. Holocene climate shifts, life-cycle plasticity, and speciation in periodical cicadas: A comment and reply to Cox and Carlton. Evolution 57: 433-437. Cooley, J. R., and D. C. Marshall. 2001. Sexual signaling in periodical cicadas, Magicicada spp. Behaviour 138: 827-855. Cooley, J. R. 2001. Long-range acoustical signals, phonotaxis, and risk in the sexual pair- forming behaviors of and O. rimosa (Hemiptera: Cicadidae). Ann. Ent. Soc. Amer. 94: 755- 760. Cooley, J. R., C. Simon, D. C. Marshall, K. Slon, and C. Ehrhardt. 2001. Allochronic speciation and reproductive character displacement in periodical cicadas supported

5 by mitochondrial DNA, song pitch, and abdominal sternite coloration data. Molecular Ecology 10:661-672. Marshall, D. C., and J. R. Cooley. 2000. Reproductive character displacement and speciation in periodical cicadas, and a new 13- year species, Magicicada neotredecim. Evolution 54: 1313-1325. Cooley, J. R., G. S. Hammond, and D. C. Marshall. 1998. The effects of enamel paint marks on the behavior and survival of the periodical cicada, Magicicada septendecim (L.) (Homoptera) and the lesser migratory grasshopper, Melanoplus sanguinipes (F.) (Orthoptera). Great Lakes Entomologist 31: 161-168. Alexander, R. D., D. C. Marshall, and J. R. Cooley. 1997. Evolutionary Perspectives on Insect Mating. Pp. 1- 31 in: B. Crespi , J. Choe, eds. The evolution of mating systems in and arachnids. (Cambridge University Press). Marshall, D. C., J. R. Cooley, R. D. Alexander, and T. E. Moore. 1996. New records of Michigan Cicadidae (Homoptera) with notes on the use of songs to monitor range changes. Great Lakes Entomologist 29: 165- 169. Cooley, J. R. 1995. Floral Heat Rewards and Direct Benefits to Insect Pollinators. Ann. Ent. Soc. Amer. 88: 576-579. Cooley, J. R. 1995. Functional hypotheses concerning the mating behavior of Lytta nuttalli Say (Coleoptera: Meloidae) in the Front Range of central Colorado. Coleopterists Bulletin 49(2): 132.

Websites: Magicicada.org (http://magicicada.org/magicicada.php) Cicada Central (http://hydrodictyon.eeb.uconn.edu/projects/cicada/cc.php) Michigan Cicadas (http://insects.ummz.lsa.umich.edu/fauna/Michigan_Cicadas/)

Invited Presentations: Cooley, J. R. 2017. The singing insects in your backyard. Lorquin Entomological Society, Los Angeles CA, 17 November 2017. Cooley, J. R. 2017. Catching periodical cicadas in the act: Range shifts in the southern Appalachians (with a timely addendum about why Zombie Cicadas are to be avoided!). University of Maryland Department of Entomology seminar series, 27 October 2017. Cooley, J. R. 2015. Insect Singers. Bugfest, North Carolina Museum of Natural Sciences, 19 September 2015. Cooley J. R. 2015. Understanding Periodical Cicadas. Fontenelle Forest Nature Center, 7 June 2015. Cooley, J. R. 2014. Periodical Cicada Brood II. Connecticut State Museum of Natural History lecture series, 29 March 2014. Cooley, J. R., and David Rothenberg. 2013. Bug Music: The Marvelous Magicicada species. Cary Institute of Ecosystem Studies, Millbrook NY 7 June 2013. Cooley, J. R., and David Rothenberg. 2013. Cicada Summer Nature Week Program. Mohonk Mountain House, New Paltz, NY 3 -5 June 2013.

6 Cooley, J. R. 2013. Cicadas: The Ultimate Science Street Fair. World Science Festival, Washington Square Park, 2 June 2013. Cooley, J. R., Rothenberg, David, and Zuk, Marlene. 2013. Cicada Serenades: Music, Mating, and Meaning. World Science Festival, New York Botanical Garden, June 1 2013. Cooley, J. R., Fisher, Helen, and Zuk, Marlene. 2013. The Joy of Six-Legged Sex: An evening of insect courtship and cocktails. World Science Festival, Staten Island Museum NY, 31 May 2013. Cooley, J. R. 2012. Periodical Cicadas. Walk-through demonstration for the Friends of the Brushy Hills, Lexington VA 26 May 2012. Cooley, J. R. 2011. The Evolution of Mating Isolation in a Periodical Cicada Contact Zone. University of Nebraska-Omaha, Graduate Seminar Series in Biology, 9 February 2011. Cooley, J. R. 2010. The Evolution of sex roles in cicadas. Shizuoka University Seminar Series, Hamamatsu Japan, 24 November 2010. Cooley, J. R. 2009. Listening to the hidden diversity around us. Department of Evolution, Ecology, and Organismal Biology Fall Seminar series, The Ohio State University, 5 November 2009. Cooley, J. R. 2009. Cicadas: Food for Thought. Yale University Department of Anthropology Colloquium Series, 26 March 2009. Cooley, J. R. 2008. Cicadas: The enigmatic insects. American Physical Society, Ridge NY, 10 October 2008. Cooley, J. R. 2004. Symposium co-organizer, “The science and the wonder of periodical cicadas” Entomological Society of America 2004 annual meeting, Salt Lake City UT. Cooley, J. R., 2002. Allochronic Speciation in Periodical Cicadas. Meeting of the New York Entomological Society, the American Museum of Natural History, 18 November 2002. Cooley, J. R., 2002. Signals, Sex roles, and Risks in cicadas: How cicadas find mates. 418th meeting of the Connecticut Entomological Society, Yale University, 18 October 2002. Cooley, J. R. 2001. Subtle signals, potent message. The function and exploitation of wing-flick signals in Magicicada. Monte Lloyd Memorial Symposium, Department of Ecology and Evolution, The University of Chicago, 13 May 2001. Cooley, J. R. 2000. Periodical Cicadas: Insights from exceptional insects. The University of Connecticut Department of Ecology and Evolutionary Biology Seminar Series, Spring 2000. Cooley, J. R. 1999. Mating behavior in cicadas: Contrasts between periodical and non- periodical species. Invited lecture, The University of Michigan Biological Station summer 1999 lecture series. Cooley, J. R., and D. C. Marshall. 1999. Deciphering and picturing sounds. The University of Michigan Exhibit Museum and Alumni Association lecture series, “The Nature of Art and Science,” Spring 1999.

7 Cooley, J. R., and D. C. Marshall. 1998. Reproductive Character Displacement, Life cycle Evolution, and a new periodical cicada species, Magicicada neotredecim. University of Michigan Museum of Zoology invited seminar, Fall 1998.

Funding: • Co-PI, NSF-DEB-16-55891 “Exploring Endosymbiont and Complexity in the Family Cicadidae.” • Collaborator, NSF-DEB 09-55849 “Systematics and the biogeography of the family Cicadidae.” • Co-PI, NUWC-Newport: “Novel Bio-acoustic approach to improved sonar transmission power level generation with USW Application.” DOD-DARPA • PI, National Geographic Society: “Making modern maps of 2007-2009 Magicicada emergences” • Co-PI, NSF-DEB 07-20664: “Systematics and biogeography of Australian Cicadettini and their relatives worldwide” • Subcontract Co-PI, NSF-PEET 05-29679: “PEET: Auchenorrhyncha new experts” Funded through New York State Museum, Cultural Education Center, Albany, NY. • Postdoctoral Researcher, NSF-DEB 99-82039 “Allochronic speciation, reproductive character displacement, and sexual selection in periodical cicadas (Magicicada spp.)” • University of Michigan Regents’ Fellowship • T. Roosevelt Memorial Fund: “Lek behavior of high alpine syrphid flies” • Hinsdale Fellowship, University of Michigan Museum of Zoology • Walker Fellowship, University of Michigan Museum of Zoology • Colorado Mountain Club research grant • University of Colorado Mountain Research Station research grant • University of Michigan Block Grant

Media and Outreach Featured presenter, World Science Festival, NY 29 May- 2 June 2013 Mohonk Mountain House Nature Week, New Paltz NY 3-5 June 2013 Cary Institute of Ecosystem Studies Lecture Series, Millbrook NY 7 June 2013 Interviewed for Lexington VA News-Gazette May 2012. Interviewed for CBS Early Show, 2 June 2011. Interviewed for NPR’s “Science Friday” Air date 21 May 2011. Interviewed for WSIL-TV local news, 30 April 2011. Interviewed for BBC-TV: “The Code” Filmed 2011, air date Fall 2011. Brown, R. 2011. The South endures the buzz of the latest “cicada year.” New York Times, (New York), May 25; p. 14 sec. A. Biemer J. 2007. Cicada brood is scientists' map quest. Chicago Tribune, (Chicago), June 23; p. 1 sec. 1. Botts B. 2007. Cicada Love. Chicago Tribune, (Chicago), April 22; p. 1 sec. 15. Brown K. 1999. Listen, we’re different. New Scientist 163: 32-5.

8 Clack K. M. 2003. Return of the cicadas. Wilkes Journal-Patriot, (North Wilkesboro, NC), June 6; p. 1 sec. 1. Dell K. 2004. They're Baaack! Time. May 24 Vol. 163: 21 p. 62-3. Karmel T. 2005. Hunt for cicadas drives enthusiasts buggy. The Willimantic Chronicle, (Willimantic, CT), July 15; p. 5 sec. 1. Keith J. 2004. The cicadas are coming. The American Gardener. May/June p. 47 Kirk, Michael. 2008. Biologists track, map cicada distribution across nation. The UCONN Advance, September 8 2008. Knudsen E. 2006. Students, professors debate clicker use in classroom. The UConn Daily Campus, (Storrs, CT), February 20; p. 1 sec. 1. Milius, S. 2000. Cicada Subtleties. Science News 157: 408-10 Reuter A. 1999. Whale's wails, cicada's chants intrigue scientists and artist. The Ann Arbor News, (Ann Arbor, MI), April 22; p. 1 sec. D. Smith, Charlotte F. 2008. Insects' loud song isn't bugging Connecticut scientist. Charleston (WV) Daily Mail, 19 June 2008. Weiss, Cindy. 2007. UCONN Scientists map mass emergence of cicadas in Midwest. The UCONN Advance, June 18, 2007.

Consultant: Consultant (as Totoket Partners LLC) for UCONN Entrepreneurship and Innovation Consortium: Provided launch and marketing materials for a University-wide initiative. Consultant (as Totoket Partners LLC) for Diverse Supplier Development Academy (www.dsdact.org): Provided curriculum and logistical support for a start-up nonprofit designed to increase the participation of minority-, women-, and veteran- owned businesses in contracting. Consultant for Partridge Films: “The World of Wildlife” Series 2, Program 31: Seasons and Cycles. By Mark Hammond. Consultant for Pixcom Productions, Montreal: “Insectia III: Outlaws”. Air Date 18-05- 2000. Paola D’Agnolo, Production Coordinator. Consultant for Japan Tele-Work LTD: Series “Our Wonderful Spaceship”. A film by Keiko Mori Yamada, with Mitsuhito Saito, Yasuo Kasugai, and Mitsuhiko Imamori. Consultant for HGTV: “The secret lives of Gardens: Life in an Oak tree” Margaret Pickles, producer. Consultant for BBC-TV: “Time Machine” Air Date: Summer 2004 (UK), Anushcka De Rohan, producer. Consultant for BBC-TV: David Attenborough series, “Life in the Undergrowth” Episode 2. Tim Green, producer. Consultant for CBS-TV CSI Season 3 Episode 6, Episode 305 “The Execution of Catherine Willows.” Written by Carol Mendelsohn and Elizabeth Devine.

University Service: UCONN MBA Graduate Student Accelerator Program Self-Study, Fall 2013. OSU Ethics education in Science and Engineering Study Group, OSU (2009-2010)

9 Personal Response System subcommittee of the UCONN IT planning committee (2006- 2007) Represented UCONN EEB department at Science, Engineering and Health Professions Collaborative Symposium for recruitment of minority students (2006-2007).

Professional Societies: Entomological Society of America, Connecticut Entomological Society, Michigan Entomological Society, Society for the Study of Evolution, Society of Systematic Biologists, Phi Beta Kappa, Sigma Xi, Beta Gamma Sigma

Collaborators (Other than those in publications list): Thomas Buckley, LandCare New Zealand. Geoff Chambers, Victoria University, Wellington, New Zealand. Paulo Fonseca, University of Lisbon, Lisbon Portugal. Derke Hughes, US Navy NUWC Newport Rhode Island. Max Moulds, The Australian Museum, Sydney, Australia.

Peer Reviews: Annals, Entomological Society of America, Bioscience, Ecology Letters, Entomological News, Entomological Society of Washington, Environmental Entomology, Evolution, Invertebrate Systematics, Journal of Ethology, Journal of Applied Entomology, Journal of Biogeography, Journal of New Music Research, Nature: Scientific Reports, NSF, Proceedings, Royal Society, Systematics and Biodiversity, Zoology, Zootaxa.

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