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Fused and Vaulted Nasals of Tyrannosaurid Dinosaurs: Implications for Cranial Strength and Feeding Mechanics
Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics ERIC SNIVELY, DONALD M. HENDERSON, and DOUG S. PHILLIPS Snively, E., Henderson, D.M., and Phillips, D.S. 2006. Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics. Acta Palaeontologica Polonica 51 (3): 435–454. Tyrannosaurid theropods display several unusual adaptations of the skulls and teeth. Their nasals are fused and vaulted, suggesting that these elements braced the cranium against high feeding forces. Exceptionally high strengths of maxillary teeth in Tyrannosaurus rex indicate that it could exert relatively greater feeding forces than other tyrannosaurids. Areas and second moments of area of the nasals, calculated from CT cross−sections, show higher nasal strengths for large tyrannosaurids than for Allosaurus fragilis. Cross−sectional geometry of theropod crania reveals high second moments of area in tyrannosaurids, with resulting high strengths in bending and torsion, when compared with the crania of similarly sized theropods. In tyrannosaurids trends of strength increase are positively allomeric and have similar allometric expo− nents, indicating correlated progression towards unusually high strengths of the feeding apparatus. Fused, arched nasals and broad crania of tyrannosaurids are consistent with deep bites that impacted bone and powerful lateral movements of the head for dismembering prey. Key words: Theropoda, Carnosauria, Tyrannosauridae, biomechanics, feeding mechanics, computer modeling, com− puted tomography. Eric Snively [[email protected]], Department of Biological Sciences, University of Calgary, 2500 University Drive NW, Calgary, Alberta T2N 1N4, Canada; Donald M. Henderson [[email protected]], Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, Alberta T0J 0Y0, Canada; Doug S. -
Ways of Seeing Animals Documenting and Imag(In)Ing the Other in the Digital Turn
InMedia The French Journal of Media Studies 8.1. | 2020 Ubiquitous Visuality Ways of Seeing Animals Documenting and Imag(in)ing the Other in the Digital Turn Diane Leblond Electronic version URL: http://journals.openedition.org/inmedia/1957 DOI: 10.4000/inmedia.1957 ISSN: 2259-4728 Publisher Center for Research on the English-Speaking World (CREW) Electronic reference Diane Leblond, “Ways of Seeing Animals”, InMedia [Online], 8.1. | 2020, Online since 15 December 2020, connection on 26 January 2021. URL: http://journals.openedition.org/inmedia/1957 ; DOI: https:// doi.org/10.4000/inmedia.1957 This text was automatically generated on 26 January 2021. © InMedia Ways of Seeing Animals 1 Ways of Seeing Animals Documenting and Imag(in)ing the Other in the Digital Turn Diane Leblond Introduction. Looking at animals: when visual nature questions visual culture 1 A topos of Western philosophy indexes animals’ irreducible alienation from the human condition on their lack of speech. In ancient times, their inarticulate cries provided the necessary analogy to designate non-Greeks as other, the adjective “Barbarian” assimilating foreign languages to incomprehensible birdcalls.1 To this day, the exclusion of animals from the sphere of logos remains one of the crucial questions addressed by philosophy and linguistics.2 In the work of some contemporary critics, however, the tenets of this relation to the animal “other” seem to have undergone a change in focus. With renewed insistence that difference is inextricably bound up in a sense of proximity, such writings have described animals not simply as “other,” but as our speechless others. This approach seems to find particularly fruitful ground where theory proposes to explore ways of seeing as constitutive of the discursive structures that we inhabit. -
The-Smartest-Dinosaur.Pdf
BY “DINO” DON LESSEM ILLUSTRATIONS BY JOHN BINDON a LERNER PUBLICATIONS COMPANY / MINNEAPOLIS To Emily Lessem, my favorite niece Text copyright © 2005 by Dino Don, Inc. Illustrations copyright © 2005 by John Bindon Photographs courtesy of: Dino Don, Inc., p. 12; Dr. Philip Currie, Royal Tyrrell Museum of Palaeontology, Drumheller, Alberta, Canada, p. 13; Photographed by Robert Fillion. Reproduced with permission of the Canadian Museum of Nature, Ottawa, Canada, p. 30; Animals, Animals © OSF/BARTLETT, D&J, p. 31. All rights reserved. International copyright secured. No part of this book may be reproduced, stored in a retrieval system, or transmitted in any form or by any means—electronic, mechanical, photocopying, recording, or otherwise—without the prior written permission of Lerner Publications Company, except for the inclusion of brief quotations in an acknowledged review. This book is available in two editions: Library binding by Lerner Publications Company, a division of Lerner Publishing Group Soft cover by First Avenue Editions, an imprint of Lerner Publishing Group 241 First Avenue North Minneapolis, MN 55401 U.S.A. Website address: www.lernerbooks.com Library of Congress Cataloging-in-Publication-Data Lessem, Don. The smartest dinosaurs / by Don Lessem ; illustrations by John Bindon. p. cm. — (Meet the dinosaurs) Includes index. eISBN: 0–8225–3285–9 1. Dinosaurs—Juvenile literature. I. Bindon, John, ill. II. Title. III. Series: Lessem, Don. Meet the dinosaurs. QE861.5.L477 2005 567.9—dc22 2004011152 Manufactured in the United States of America 1 2 3 4 5 6 – DP – 10 09 08 07 06 05 MEET THE SMARTEST DINOSAURS . 4 HOW SMART WERE DINOSAURS? . -
FACT SHEET: Giganotosaurus
FACT SHEET: giganotosaurus NAME: GIGANOTOSAURUS (JIG-an-oh-toh-SAW-rus), “Giant Southern Lizard” SIZE: 45 feet long, 12 feet tall (the size of a locomotive), weighed 8 tons TYPE OF DIET: Carnivore (large dinosaurs) WHEN: Late Cretaceous period (100 million years ago) WHERE: Argentina (South America) When the fossilized skeleton of Giganotosaurus was discovered in 1995, many people got very excited. It was one of the largest known carnivores ever to walk the earth. Giganotosaurus means "giant southern lizard" because it was found in Argentina, South America. Only 70 percent of the skeleton was found, but it was enough to re-build this giant creature. It was about 45 feet long: that’s about the size of a locomotive. It was five feet longer than Tyrannosaurus rex and about two tons heavier. Giganotosaurus wasn’t a very intelligent dinosaur. Its brain was about the size of a banana. Giganotosaurus walked on two thick, powerful legs and had enormous jaws. Its teeth were 8 inches long and serrated like the edge of a saw. Giganotosaurus lived about 100 million years ago. Giganotosaurus may have hunted animals much larger than itself, such as the huge sauropod plant-eaters like Brontosaurus and Diplodocus. The skeleton of a 75 foot-long herbivore was found right next to the fossil of a Giganotosaurus. It had a long, skinny tail which was useful for balance. This means it could turn quickly while running and jumping. Its tail shot straight out behind it and was kept far off the ground. Several skeletons of Giganotosaurus were found together in the same place. -
Evaluating the Ecology of Spinosaurus: Shoreline Generalist Or Aquatic Pursuit Specialist?
Palaeontologia Electronica palaeo-electronica.org Evaluating the ecology of Spinosaurus: Shoreline generalist or aquatic pursuit specialist? David W.E. Hone and Thomas R. Holtz, Jr. ABSTRACT The giant theropod Spinosaurus was an unusual animal and highly derived in many ways, and interpretations of its ecology remain controversial. Recent papers have added considerable knowledge of the anatomy of the genus with the discovery of a new and much more complete specimen, but this has also brought new and dramatic interpretations of its ecology as a highly specialised semi-aquatic animal that actively pursued aquatic prey. Here we assess the arguments about the functional morphology of this animal and the available data on its ecology and possible habits in the light of these new finds. We conclude that based on the available data, the degree of adapta- tions for aquatic life are questionable, other interpretations for the tail fin and other fea- tures are supported (e.g., socio-sexual signalling), and the pursuit predation hypothesis for Spinosaurus as a “highly specialized aquatic predator” is not supported. In contrast, a ‘wading’ model for an animal that predominantly fished from shorelines or within shallow waters is not contradicted by any line of evidence and is well supported. Spinosaurus almost certainly fed primarily from the water and may have swum, but there is no evidence that it was a specialised aquatic pursuit predator. David W.E. Hone. Queen Mary University of London, Mile End Road, London, E1 4NS, UK. [email protected] Thomas R. Holtz, Jr. Department of Geology, University of Maryland, College Park, Maryland 20742 USA and Department of Paleobiology, National Museum of Natural History, Washington, DC 20560 USA. -
Spinosaurus by Guy Belleranti
Name: ______________________________ Spinosaurus By Guy Belleranti Spinosaurus was a gigantic meat-eating dinosaur. It was even longer and heavier than the famous Tyrannosaurus rex. It was one of the biggest meat-eating dinosaurs. The Spinosaurus and the Tyrannosaurus rex lived in the Cretaceous period, but they never met. T-rex lived at least 30 million years after Spinosaurus. Spinosaurus had long bony spines coming out of its back. Scientists believe these spines were connected together with skin and blood vessels into a sail-like shape. This sail might have helped Spinosaurus keep its body at the right temperature. It also added to this scary beast’s size. Like T-rex, Spinosaurus had long, strong hind legs. However, its front arms were much longer than T-rex’s. This could mean Spinosaurus sometimes walked on all fours. Spinosaurus had a long crocodile-like snout filled with sharp teeth. These teeth weren’t as large as T-rex’s. However, they would’ve been perfect for eating fish and smaller dinosaurs. Super Teacher Worksheets - www.superteacherworksheets.com Name: ______________________________ Spinosaurus By Guy Belleranti 1. Spinosaurus was.... a. the largest dinosaur that ever lived. b. much smaller than a Tyrannosaurus rex. c. one of the fastest dinosaurs. d. a meat-eating dinosaur. 2. The Spinosaurus lived during the ______________________________ period. 3. Did the Tyrannosaurus rex and the Spinosaurus ever see each other? Explain your answer. ___________________________________________________________________________________________ ___________________________________________________________________________________________ 4. The spines on the Spinosaurus' back may have been used to... a. keep it's body the right temperature b. attack other dinosaurs c. help it walk on two legs d. -
Cranial Anatomy of Allosaurus Jimmadseni, a New Species from the Lower Part of the Morrison Formation (Upper Jurassic) of Western North America
Cranial anatomy of Allosaurus jimmadseni, a new species from the lower part of the Morrison Formation (Upper Jurassic) of Western North America Daniel J. Chure1,2,* and Mark A. Loewen3,4,* 1 Dinosaur National Monument (retired), Jensen, UT, USA 2 Independent Researcher, Jensen, UT, USA 3 Natural History Museum of Utah, University of Utah, Salt Lake City, UT, USA 4 Department of Geology and Geophysics, University of Utah, Salt Lake City, UT, USA * These authors contributed equally to this work. ABSTRACT Allosaurus is one of the best known theropod dinosaurs from the Jurassic and a crucial taxon in phylogenetic analyses. On the basis of an in-depth, firsthand study of the bulk of Allosaurus specimens housed in North American institutions, we describe here a new theropod dinosaur from the Upper Jurassic Morrison Formation of Western North America, Allosaurus jimmadseni sp. nov., based upon a remarkably complete articulated skeleton and skull and a second specimen with an articulated skull and associated skeleton. The present study also assigns several other specimens to this new species, Allosaurus jimmadseni, which is characterized by a number of autapomorphies present on the dermal skull roof and additional characters present in the postcrania. In particular, whereas the ventral margin of the jugal of Allosaurus fragilis has pronounced sigmoidal convexity, the ventral margin is virtually straight in Allosaurus jimmadseni. The paired nasals of Allosaurus jimmadseni possess bilateral, blade-like crests along the lateral margin, forming a pronounced nasolacrimal crest that is absent in Allosaurus fragilis. Submitted 20 July 2018 Accepted 31 August 2019 Subjects Paleontology, Taxonomy Published 24 January 2020 Keywords Allosaurus, Allosaurus jimmadseni, Dinosaur, Theropod, Morrison Formation, Jurassic, Corresponding author Cranial anatomy Mark A. -
Have Fun with the Family
Good news! Almost every activity in our entire HAVE FUN programme is free to enjoy, so take your pick WITH THE and get there quick! Maybe you’ll become an FAMILY animator of stories from around the world. Or get up close and personal A season of surprises awaits you with a creepy crawly. in Oxford University’s Gardens, Who knows, perhaps you’ll go hunting for fantastic fossils on Planet Dinosaur! Whatever you end up Libraries & Museums. So what doing, remember to have fun, make friends and let are you waiting for? Let’s go! your mind grow. EASTER EGGSPLORATION GET SET FOR ADVENTURE saturday 31st march–sunday 15th april Download one of our fun family trails before you visit (in opening hours) or print off an activity sheet to try at home: EGGSTRAORDINARY EASTER TRAIL A Want to be a dinosaur detective? Or make Follow the eggciting egg-hunter’s trail yourself some dodo feet? Track them down at: through the museums. www.oum.ox.ac.uk/educate/famtrails.htm museum of natural history & pitt rivers museum In need of a survival guide to Ancient Egypt? Check out the Ashmolean’s 8 great trails at: sunday 1st–monday 23rd april www.ashmolean.org/family-trails (in opening hours) EASTER TRAIL 7+ Ever worn a reindeer hat or flown a carp kite? Challenge yourself to the museum’s Find out how to make your own at: Easter Trail. Help yourself from the www.prm.ox.ac.uk/family-athome Entrance Gallery. museum of the history of science GOOD STUFF TO GRAB WEEK-IN WEEK-OUT Go and grab your explorer packs from HARCOURT ARBORETUM every saturday, 11.30am–3.30pm every sunday, 2–4pm any of our information desks on site! Pick up an explorer’s backpack from the ticket office and use it to help you HANDS-ON COINS 7+ FAMILY-FRIENDLY SUNDAYS explore the 130 – acre site. -
New Tyrannosaur from the Mid-Cretaceous of Uzbekistan Clarifies Evolution of Giant Body Sizes and Advanced Senses in Tyrant Dinosaurs
New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs Stephen L. Brusattea,1, Alexander Averianovb,c, Hans-Dieter Suesd, Amy Muira, and Ian B. Butlera aSchool of GeoSciences, University of Edinburgh, Edinburgh EH9 3FE, United Kingdom; bZoological Institute, Russian Academy of Sciences, St. Petersburg 199034, Russia; cDepartment of Sedimentary Geology, Saint Petersburg State University, St. Petersburg 199178, Russia; and dDepartment of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560 Edited by Neil H. Shubin, The University of Chicago, Chicago, IL, and approved January 29, 2016 (received for review January 5, 2016) Tyrannosaurids—the familiar group of carnivorous dinosaurs in- We here report the first diagnostic tyrannosauroid from the mid- cluding Tyrannosaurus and Albertosaurus—were the apex predators Cretaceous, a new species from the Turonian (ca. 90–92 million in continental ecosystems in Asia and North America during the years ago) Bissekty Formation of Uzbekistan. This formation has latest Cretaceous (ca. 80–66 million years ago). Their colossal sizes recently emerged as one of the most important records of mid- and keen senses are considered key to their evolutionary and eco- Cretaceous dinosaurs globally (9–11). Possible tyrannosauroid logical success, but little is known about how these features devel- specimens from the Bissekty Formation were reported more than oped as tyrannosaurids evolved from smaller basal tyrannosauroids a half century ago (12), and, more recently, several isolated fossils that first appeared in the fossil record in the Middle Jurassic (ca. 170 were assigned to the group (9, 13), but none of these has been million years ago). -
A Century of Spinosaurs - a Review and Revision of the Spinosauridae
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Queen Mary Research Online A century of spinosaurs - a review and revision of the Spinosauridae with comments on their ecology HONE David William Elliott1, * HOLTZ Thomas Richard Jnr2 1 School of Biological and Chemical Sciences, Queen Mary University of London, London, E1 4NS, UK 2 Department of Geology, University of Maryland, College Park, MD 20742 USA Abstract: The spinosaurids represent an enigmatic and highly unusual form of large tetanuran theropods that were first identified in 1915. A recent flurry of discoveries and taxonomic revisions of this important and interesting clade had added greatly to our knowledge, however, spinosaur body fossils are generally rare and most species are known from only limited skeletal remains. Their unusual anatomical adaptations to the skull, limbs and axial column all differ from other large theropods and point to an unusual ecological niche and a lifestyle intimately linked to water. Keywords: Theropoda, Megalosauroidea, Baryonychinae, Spinosaurinae, palaeoecology E-mail: [email protected] 1 Introduction The Spinosauridae is an enigmatic clade of large and carnivorous theropods from the Jurassic and Cretaceous that are known from both Gondwana and Laurasia (Holtz et al., 2004). Despite their wide temporal and geographic distribution, the clade is known primarily from teeth and the body fossil record is extremely limited (Bertin, 2010). As such, relatively little is known about this group of animals, although their unusual morphology with regard to skull shape, dentition, dorsal neural spines and other features mark them out as divergent from the essential bauplan of other non-tetanuran theropods (Fig 1). -
Fact Sheet for Dinosaur Brain Cooling Story EMBARGOED for RELEASE WEDNESDAY, 16 OCTOBER 2019, 2 PM EDT
Fact sheet for dinosaur brain cooling story EMBARGOED FOR RELEASE WEDNESDAY, 16 OCTOBER 2019, 2 PM EDT Published article Porter, W. R., and L. M. Witmer. 2019. Vascular patterns in the heads of dinosaurs: evidence for blood vessels, sites of thermal exchange, and their role in physiological thermoregulatory strategies. Anatomical Record Authors Lead author: Ruger Porter, PhD, Assistant Professor of Instruction, Ohio University Heritage College of Osteopathic Medicine Athens, OH. USA [email protected] Second author: Lawrence Witmer, PhD, Professor of Anatomy, Chang Professor of Paleontology, Ohio University Heritage College of Osteopathic Medicine Athens, OH. USA [email protected] Phone: 740-591-7712 Ohio University Communications & Marketing: Jim Sabin, 740-593-0858, [email protected] Website with downloadable resources https://people.ohio.edu/witmerl/dinosaur_brain-cooling_strategies.htm Common Language Summary Keeping a cool head – Different dinosaurs evolved different cooling strategies. For more than a decade, a team of Ohio University researchers has been studying how large-bodied dinosaurs survived potentially fatal overheating. New research from this team, drawing on advanced imaging techniques and sophisticated quantitative analyses, has discovered evidence of often vast networks of blood vessels in the skulls of dinosaurs that were likely used to cool blood destined for the brain. It turns out that different groups of large dinosaurs emphasized different sites of evaporative cooling in the head. For example, armored ankylosaurs like Euoplocephalus used the nasal cavity as an air conditioner, whereas long-necked sauropods like Diplodocus and Camarasaurus also used the mouth to cool blood, probably through panting. The predatory theropod dinosaurs, such as T. rex and Majungasaurus, however, evolved a very different brain-cooling strategy, involving a highly vascular air sinus in the snout through which air was pumped by movements of the jaws. -
BBC Annual Report and Accounts 2012
PART 2 THE BBC EXECUTIVE’S REVIEW AND ASSESSMENT Drama Sherlock, starring Benedict Cumberbatch and Martin Freeman, returned for a second series in January 2012. CONTENTS AND SUBJECT INDEX Part 2 BBC Executive contents Managing the business Overview 2-28 Chief Operating Officer’s review 2-1 Director-General’s introduction 2-29 Working together 2-2 Understanding the BBC’s finances Governance 2-4 Performance by service 2-40 Executive Board 2-8 Television 2-42 Risks and opportunities 2-9 Radio 2-44 Governance report 2-10 News 2-47 Remuneration report 2-11 Future Media 2-52 Audit Committee report 2-12 Nations & Regions 2-55 Fair trading report Delivering our strategy Managing our finances 2-14 Distinctiveness and quality 2-58 Chief Financial Officer’s review 2-15 The best journalism in the world 2-59 Summary financial performance 2-16 Inspiring knowledge, music 2-60 Financial overview and culture 2-68 Collecting the licence fee 2-17 Ambitious drama and comedy 2-69 Looking forward with confidence 2-20 Outstanding children’s content 2-70 Auditor’s report 2-21 Content that brings the nation 2-71 Glossary and communities together 2-72 Contact us/More information 2-22 Value for money 2-23 Serving all audiences 2-26 Openness and transparency Subject Index Part 1 Part 2 Board remuneration 1-9/1-32 2-48 Commercial strategy 1-8 2-36 Complaints 1-3/1-19 2-55 Delivering Quality First 1-4/1-6 2-14 Digital switchover – 2-25 Distribution 1-17 2-25 Editorial priorities – 2-14 Editorial standards 1-3/1-18 2-38 Efficiency 1-6 2-59/2-61 Equality and diversity