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105-123, 2000 Demre, Turkey, with (Gastropoda, Pulmonata BASTERIA, 64: 105-123, 2000 The land snails of the between Kaş area and Demre, southwestern Turkey, with special reference to Albinaria (Gastropoda, Pulmonata, Clausiliidae) Eike Neubert Sautierstrasse 47, D 79104 Freiburg, Germany; [email protected] Aydin Örstan 13348 Cloverdale Place, Germantown, Maryland 20874, USA; [email protected] & Francisco Welter-Schultes Institut fur Zoologie und Anthropologic der Universitat, Berliner Str. 28, D 37073 Gottingen, Germany; [email protected] The Kaş and Demre southwestern 40 x 10 km region extending between (Vilayet Antalya, an for studies Turkey) provides interesting area on land snails. It is one of the less well-known regions ofTurkey in terms of land snail distribution.Most of the landscape consists of limestone In this outcrops. area 32 species of land snails were found, 14 ofwhich live exclusively in the coastal and suburban area of Demre, suggesting that they were introduced. Among the 18 autochthonous of A. species are four species of Albinaria, one which is described as latelamellaris A. anatolica is and exhibits spec. nov.; very common a considerable degree of geographical variation in shell characters. Key words: Gastropoda, Albinaria, dispersal, introduced species, distribution. biogeography, morphological variation, Turkey. INTRODUCTION The last general approach to the land snails of Turkey was a monumental work by Schutt (1993, 1996), criticized mainly for the limited value of its systematic parts (Hausdorf, Bank 1994; Riedel, 1995; et al., 1997). These were claimed to have resulted in unreliable numbers for Another concerned the species many groups. objection poor quality of the distributional data. evaluations of Turkish land snails based Schutt Biogeographical on (1996) (e.g. Cook, 1997) face the difficulty ofcombining the species compositions of better surveyed regions with those from studied For poorly areas. instance, the data concerning the southeastern of the of the regions present-day Turkey largely represent our knowledge at beginning 20th whereas is much better century, western Turkey surveyed. Analysing species numbers and distribution patterns of Schutt's different regions without consideration of these 106 BASTERIA, Vol. 64, No. 4-6, 2000 shortcomings is expected to yield historically rather than biogeographically meaningful results. The only solid base for biogeographical and phylogeographical evaluations of Asia Minor and should be the and local of adjacent regions complete regional ranges the involved. species This knowledge is unlikely to be provided by one person's efforts; it will have be achieved with contributions researchers to step by step, by many (Bank et al., 1997). The 40 10 km between and is x area extending Ka§ Demre, Vilayet Antalya (fig. 1) of one the poorest known regions in terms of mollusc distribution. The last surveys on the probably best-known land snail family ofTurkey (Zonitidae) recorded several species at Ka§ and west of Ka§, on the islands of Kastelorizo (Meis, Megisti) and Kekova, at Demre and of Demre Myra, east (Riedel, 1982, 1987, 1995), but nothing was reported from the hinterlandbetween Ka§ and Demre. Some of the data are based on expeditions conducted 100 150 The situation or years ago. same is observed in Albinaria (Clausiliidae) and Isaurica lycia (Martens, 1889) (Helicidae) (Nordsieck, 1993; Subai, 1994). The Greek island of which group Kastelorizo, consists of seven small islands off Ka§, was investi- & but from gated by Poulakakis, Vardinoyannis Mylonas (1997), apart some general the results statements were not published, so that the study did not contribute much to our knowledge of the land snails of that region. The initiated present study was after 36 shells of land snails were found in an exca- vation of a late Bronze off between and age shipwreck Uluburun, a rocky peninsula Ka§ Demre. Since the of the snail finds interpretation shipwreck required a detailed knowl- of the distribution of the snails of the edge adjacent region, the area was thoroughly checked in 1998 for land snails. In fact, four shells of the shipwreck belonged to endemic Fig. 1. Map of the study area with collection sites. The site of the Uluburun shipwreck is indicated. Neubert, Orstan & Welter-Schultes: Land snails of SW. Turkey 107 and characteristic of the Uluburun viz. of and species area, Metafruticicola spec, Megisti Zonites caricus (Roth, 1839). They are considered not to have belonged to the ship's inventory, but to have reached the shipwreck site independently (Welter-Schultes, in press). MATERIAL AND METHODS In September 1998 F. Welter-Schultes collected land snail samples at the following 64 sites between Kas and Demre (fig. 1). The material is stored in the Haus der Natur, Cismar, Germany. #1 Road Kas-Finike, 0.5 km W. Agullu (crossroads Yenikoy), =Bkm E Kas, 21.9.1998 #2 Road Kas-Finike, Ikm E. crossroads Yenikoy, 21.9.1998 #3 Road Kas-Finike, 0.7 km E. crossroads Kasaba-Akcay-Elmah 21.9.1998 #4 Road Ka§-Finike, 2km E. crossroads Kasaba = 2.5 km W crossroads Kihcli-Ucagiz, 22.9.1998 #5 Road Kas-Finike, 2km W. crossroads Kihcli-Ucagiz, 22.9.1998 #6 Road Kas-Finike, Ikm VV. crossroads Kihcli-Ucagiz, 22.9.1998 #7 Road Ka§-Finike, 0.4 km S. crossroads Kihch-Ucagiz, 22.9.1998 #8 Road Kas-Kilich-Ucagiz, 4.5 km SE. crossroads road Kaf-Finike, 22.9.1998 #9 Road Ka§-Kihch-Ucagiz, 6km SE. crossroads road Kas-Finike, 22.9.1998 #lO Road Ka§ -Kihch-Ucagiz, 8 km SSE. crossroads road Kas-Finike, 22.9.1998 #ll 1.5 km NNW Kihch, = 10.5 direct km E. Ka§, 22.9.1998 #l2 2 km SVV Bogazcik, = 9 direct km ESE. Ka§, 22.9.1998 #l3 3 km SVV Bogazcik, = 8.5 direct km ESE. Ka§, 22.9.1998 #l4 Ruins 2.5 km SSW Bogazcik, = 8.5 direct km south ESE. Kas, 22.9.1998 #l5 0.8 km Wof ruins 2.5 km SSW Bogazcik, = 8 direct km south ESE. Ka§, = 1.5 km S. Qam Dagi- Mountain, 22.9.1998 #l6 W outlet of a valley Bkm SE. Ka§ at the coast, = 1.5 km NE. Kdrmen Adasi, 22.9.1998 #l7 7.5 km SE. Ka§, rocks at the coast, = 1 km N Kormen Adasi, 22.9.1998 #lB W fringe of the bay 7.5 km SE. Ka§ at the coast, = 1 km W Kormen Adasi, = 1 km N. Cilpacik Adasi, 23.9.1998 #l9 Bay E. of Ince Burun, 7.5 km south SE Ka§, 1.5 km E Uluburun peninsula ridge, 23.9.1998 #2O Bay immediately E. of the Uluburun peninsula, = 7 km SSE Ka§, 23.9.1998 #2l Ruins Wof the bay immediately E. of the Uluburun peninsula, = 7km SSE Kas, 23.9.1998 #22 Uluburun peninsula ridge, summit SW. of the eastern bay, 6.5 km SSE Ka§, 23.9.1998 #23 Uluburun peninsula ridge, 7 km SSE. Kas, 23.9.1998 #24 Uluburun peninsula ridge, southernmost high rocks in dark maccia, 7.5 km SSE. Ka§, 23.9.1998 #25 Eastern slopes of the Uluburun peninsula, 7.2 km SSE. Kas, at two thirds ridge altitude, 23.9.1998 Road S. the = 5.5 km SSE. Ka 23.9.1998 #26 Bayindir, 0.8 km N. end of road, 5, #27 Village ruins 4km S. Bayindir, = 5 km SE. Kas, Okcubldiigii, 24.9.1998 #2B 2.5 km S. Bayindir, = 4.5 km SE. Kas, wall alongside the backroad, 24.9.1998 #29 S. end of the village of Bayindir, = 3 km ESE. Ka|, 24.9.1998 #3O Road Ka§-Bayindir, 2 direct km SE. Kas, 24.9.1998 #3l Road Kas-Bayindir, 1.2 direct km SE. Kas, 24.9.1998 #32 0.4 km S. Ka§, rocks at Buyiik gakil Beach, 24.9.1998 #33 Road Kas-Finike, 2 km SiE. crossroads Kihcli-Ucagiz, 25.9.1998 #34 Road Ka§-Finike, 3.5 km ENE. crossroads Kihcli-Ucagiz, = 0.5 km W. Sanlar, = 1 km E. crossroads Gokceyazi, 25.9.1998 108 BASTERIA, Vol. 64, No. 4-6, 2000 Orculella ignorata Rupestrella rhodia Mastus rossmaessleri Zonites caricus Albinaria anatolica Albinaria lycica A. latelamellaris A. myrensis schuberti Metafruticicola Metafruticicola sp. Megisti Isaurica lycia Helix cincta complex Introduced Granopupa granum species Fig. 2. Distribution of some land snail species between Megisti and Finike. Mubert, Orstan & Welter-Schultes: Land snails of SW. Turkey 109 = #35 Road Ka§-Finike, 4.5 km E. crossroads Kihch-U<;agiz, 7.5 km W. Yavi, = 0.5 km E. Sanlar, 25.9.1998 #36 Road Ka§-Finike, 7 km E. crossroads Kihch-Ucagiz, = 5 km W. Yavi, = 0.3 km E Kiziloren, 25.9.1998 #37 Road Ka§-Finike, 1 km W. crossroads Kyaneai, = 3 km VV. Yavi, 25.9.1998 #3B Road Ka§-Finike, crossroads Kyaneai, = 2 km W. Yavi, = 8 km W. Davazlar, 25.9.1998 #39 Road Kas-Finike, 3 km VV. Davazlar, 25.9.1998 #4O Road Kas-Finike, 1.5 km W. Davazlar, 26.9.1998 #4l Road 3km E. Ka§-Finike, pass Davazlar, 26.9.1998 #42 Road Ka§-Finike, 5 km E Davazlar, = 1.5 km VV. Gurses, = 0.5 km E Qakalbayat, 26.9.1998 #43 Road Ka§-Finike, VV.-entrance of village Gurses, 10 road-km NVV Demre, 26.9.1998 #44 Road Ka§-Finike, 2 km ESE. Gurses, = 8 road-km NVV Demre, 26.9.1998 #45 New Road Ka§-Finike, 4 road-km \V. Demre (crossroads Cayagzi), = 2 km SVV. Sura, = 0.3 km S ruins, 26.9.1998 #46 Rocks = = 18.5 direct km VVSVV. 26.9.1998 at the port of Cayagzi, near Demre Kale, Finike, 18.5 km #47 Ruins at the beach of Cayagzi, near Demre = Kale, = direct VVSVV. Finike, 26.9.1998 #4B Rocks NVV.
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