Alberta Palaeontological Society Bulletin, Volume 33, No. 3
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Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
Paleobiology of Archaeohippus (Mammalia; Equidae), a Three-Toed Horse from the Oligocene-Miocene of North America
PALEOBIOLOGY OF ARCHAEOHIPPUS (MAMMALIA; EQUIDAE), A THREE-TOED HORSE FROM THE OLIGOCENE-MIOCENE OF NORTH AMERICA JAY ALFRED O’SULLIVAN A DISSERTATION PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORIDA IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY UNIVERSITY OF FLORIDA 2002 Copyright 2002 by Jay Alfred O’Sullivan This study is dedicated to my wife, Kym. She provided all of the love, strength, patience, and encouragement I needed to get this started and to see it through to completion. She also provided me with the incentive to make this investment of time and energy in the pursuit of my dream to become a scientist and teacher. That incentive comes with a variety of names - Sylvan, Joanna, Quinn. This effort is dedicated to them also. Additionally, I would like to recognize the people who planted the first seeds of a dream that has come to fruition - my parents, Joseph and Joan. Support (emotional, and financial!) came to my rescue also from my other parents—Dot O’Sullivan, Jim Jaffe and Leslie Sewell, Bill and Lois Grigsby, and Jerry Sewell. To all of these people, this work is dedicated, with love. ACKNOWLEDGMENTS I thank Dr. Bruce J. MacFadden for suggesting that I take a look at an interesting little fossil horse, for always having fresh ideas when mine were dry, and for keeping me moving ever forward. I thank also Drs. S. David Webb and Riehard C. Hulbert Jr. for completing the Triple Threat of Florida Museum vertebrate paleontology. In each his own way, these three men are an inspiration for their professionalism and their scholarly devotion to Florida paleontology. -
View of What Is Now Known About
A PHASE OF THE UPPER DEVONIAN OF WESTERN HIGHLAND COUNTY, VIRGINIA By Helen F. Pulver A. B., Berea College, 1946 A thesis submitted to the Faculty of Oberlin College in partial fulfillment of the requirements for the Degree of Master of Arts in the Department of Geology 1948 PLATE I Chemung formation, Alleghany Mountain locality, 3145 A fossiliferous bed of arenaceous shale, containing predominantly Douvillina cayuta; Schizophoria stria tula in the upper and lower right hand corners, and a part of Tylothyris mesacostalis and Camarotoeehia eximia in the center. PLATE 1 PREFACE '!his paper is a study of the Brallier and Chemung formations - of west Highland County, Virginia. '!he limits of the Brallier forma t.ion are determined on the basis ot faunal and lithological evidence, and correlations made with format.ions in New York, Pennsylvania, Maryland and West Virginia. 'Ihe field work was carried on during the lat ter part of the summer of 1947 and the identification and correlations worked out in the winter of 1947 and the spring of 1948. I wish to thank Mr. Minor B. Long and 1fr. Bruce Weber for help in measuring .the section of the field. I am. grateful to Dr. E. C. Stumm for his interest and original suggestion of the problem. ~ Dr. C.W• .Carlston I should like to express my thanks for valuable suggestions in the field and in the preparation of this manuscript. Above all I should. like to express my deep appreeiation to Dr. Fred Foreman under whose direction this problem was completed, for his continued encouragement and constant assistance. -
Brachiopods from the Mobarak Formation, North Iran
GeoArabia, 2011, v. 16, no. 3, p. 129-192 Gulf PetroLink, Bahrain Tournaisian (Mississippian) brachiopods from the Mobarak Formation, North Iran Maryamnaz Bahrammanesh, Lucia Angiolini, Anselmo Alessandro Antonelli, Babak Aghababalou and Maurizio Gaetani ABSTRACT Following detailed stratigraphic work on the Mississippian marlstone and bioclastic limestone of the Mobarak Formation of the Alborz Mountains in North Iran, forty-eight of the most important brachiopod taxa are here systematically described and illustrated. The ranges of the taxa are given along the Abrendan and Simeh Kuh stratigraphic sections, located north of Damgham. The examined brachiopod species date the base of the Mobarak Formation to the Tournaisian, in absence of age-diagnostic foraminifers. Change in brachiopod settling preferences indicates a shift from high energy, shallow-water settings with high nutrient supply in the lower part of the formation to quieter, soft, but not soppy substrates, with lower nutrient supply in the middle part of the Mobarak Formation. Brachiopod occurrence is instead scanty at its top. The palaeobiogeographic affinity of the Tournaisian brachiopods from North Iran indicates a closer relationship to North America, Western Europe and the Russian Platform than to cold-water Australian faunas, confirming the affinity of the other biota of the Alborz Mountains. This can be explained by the occurrence of warm surface-current gyres widely distributing brachiopod larvae across the Palaeotethys Ocean, where North Iran as other peri- Gondwanan blocks acted as staging-posts. INTRODUCTION The Mississippian Mobarak Formation of the Alborz Mountains (North Iran) has been recently revised by Brenckle et al. (2009) who focused mainly on its calcareous microfossil biota and refined its biostratigraphy, chronostratigraphy and paleogeography. -
The Spence Shale Lagerstätte: an Important Window Into Cambrian Biodiversity
Downloaded from http://jgs.lyellcollection.org/ by guest on September 24, 2021 Accepted Manuscript Journal of the Geological Society The Spence Shale Lagerstätte: an Important Window into Cambrian Biodiversity Julien Kimmig, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff & Bruce S. Lieberman DOI: https://doi.org/10.1144/jgs2018-195 Received 31 October 2018 Revised 21 February 2019 Accepted 28 February 2019 © 2019 The Author(s). This is an Open Access article distributed under the terms of the Creative Commons Attribution 4.0 License (http://creativecommons.org/licenses/by/4.0/). Published by The Geological Society of London. Publishing disclaimer: www.geolsoc.org.uk/pub_ethics Supplementary material at https://doi.org/10.6084/m9.figshare.c.4423145 To cite this article, please follow the guidance at http://www.geolsoc.org.uk/onlinefirst#cit_journal Downloaded from http://jgs.lyellcollection.org/ by guest on September 24, 2021 The Spence Shale Lagerstätte: an Important Window into Cambrian Biodiversity 1* 1,2 1,3 4 1,2 Julien Kimmig , Luke C. Strotz , Sara R. Kimmig , Sven O. Egenhoff & Bruce S. Lieberman 1Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA 2 Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence, KS, USA 3Pacific Northwest National Laboratory, Richland, WA 99354, USA 4Department of Geosciences, Colorado State University, Fort Collins, CO 80523, USA *Correspondence: [email protected] Abstract: The Spence Shale Member of the Langston Formation is a Cambrian (Miaolingian: Wuliuan) Lagerstätte in northeastern Utah and southeastern Idaho. It is older than the more well- known Wheeler and Marjum Lagerstätten from western Utah, and the Burgess Shale from Canada. -
An Early Miocene Dome-Skulled Chalicothere
PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, NY 10024 Number 3486, 45 pp., 26 ®gures, 8 tables October 27, 2005 An Early Miocene Dome-Skulled Chalicothere from the ``Arikaree'' Conglomerates of Darton: Calibrating the Ages of High Plains Paleovalleys Against Rocky Mountain Tectonism ROBERT M. HUNT, JR.1 CONTENTS Abstract ....................................................................... 2 Introduction .................................................................... 2 Geologic Setting ................................................................ 3 Systematic Paleontology ........................................................ 12 Age of the Dome-Skulled Chalicothere ........................................... 28 The ``Arikaree'' Conglomerates of N.H. Darton ................................... 32 Post-Laramide Evolution of the Rocky Mountains ................................. 39 Acknowledgments ............................................................. 42 References .................................................................... 42 1 Department of Geosciences, W436 Nebraska Hall, University of Nebraska, Lincoln, NE 68588-0549 ([email protected]). Copyright q American Museum of Natural History 2005 ISSN 0003-0082 2 AMERICAN MUSEUM NOVITATES NO. 3486 ABSTRACT Fragmentary skeletal remains discovered in 1979 in southeastern Wyoming, associated with a mammalian fauna of early Hemingfordian age (;18.2 to 18.8 Ma), represent the oldest known occurrence of dome-skulled chalicotheres -
Hipparion” Cf
©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien Beitr. Paläont., 30:15-24, Wien 2006 Hooijer’s Hypodigm for “ Hipparion” cf. ethiopicum (Equidae, Hipparioninae) from Olduvai, Tanzania and comparative Material from the East African Plio-Pleistocene by Miranda A rmour -Chelu 1}, Raymond L. Bernor 1} & Hans-Walter Mittmann * 2) A rmour -C helu , M., Bernor , R.L. & M ittmann , H.-W., 2006. Hooijer’s Hypodigm for “ Hipparion” cf. ethiopicum (Equidae, Hipparioninae) from Olduvai, Tanzania and comparative Material from the East African Plio-Pleistocene. — Beitr. Palaont., 30:15-24, Wien. Abstract cranialen Elemente die Hooijer zu diesem Taxon gestellt hat, auf die er sich bezogen hat oder die in irgendeiner We review here the problematic history of the nomen Beziehung dazu stehen, haben wir wiedergefunden. Selbst “Hipparion”cf. ethiopicum and Hooijer’s efforts to bring zusätzliche Fundstücke aus zeitgleichen Horizonten haben some taxonomic sense to the later Pliocene - Pleistocene wir miteinbezogen, in der Absicht, die Gültigkeit von Eu hipparion record. We review his reasoning, and the shifts rygnathohippus cf.“ethiopicum" und seines Verwandten in taxonomic allocation made by him and other equid Eurygnathohippus cornelianus und weiterer Formen, von researchers during the 1970’s. We have relocated many denen Hooijer geglaubt hat, dass sie in einem evolutionären of the postcranial specimens attributed by Hooijer to Zusammenhang mit „ Hipparion“ cf. ethiopicum stehen, “Hipparion” cf. ethiopicum, as well as other specimens zu testen. Wir machen statistische und vergleichende which he referred to, or related to this species. We have Analysen um dieses Hypodigma zu klären. also considered additional specimens from contempo raneous horizons, in order to reevaluate the efficacy of Eurygnathohippus cf “ethiopicum” and its apparent rela 1. -
Palaeoecology of the Early Cambrian Sinsk Biota from the Siberian Platform
Palaeogeography, Palaeoclimatology, Palaeoecology 220 (2005) 69–88 www.elsevier.com/locate/palaeo Palaeoecology of the Early Cambrian Sinsk biota from the Siberian Platform Andrey Yu. Ivantsova, Andrey Yu. Zhuravlevb,T, Anton V. Legutaa, Valentin A. Krassilova, Lyudmila M. Melnikovaa, Galina T. Ushatinskayaa aPalaeontological Institute, Russian Academy of Sciences, ul. Profsoyuznaya 123, Moscow 117997, Russia bA´rea y Museo de Paleontologı´a, faculdad de Ciences, Universidad de Zaragoza, C/ Pedro Cerbuna, 12, E-50009, Zaragoza, Spain Received 1 February 2002; accepted 15 January 2004 Abstract The Sinsk biota (Early Cambrian, Botoman Stage, Siberian Platform) inhabited an open-marine basin within the photic zone, but in oxygen-depleted bottom waters. Its rapid burial in a fine-grained sediment under anoxic conditions led to the formation of one of the earliest Cambrian Lagerst7tte. All the organisms of the biota were adapted to a life under dysaerobic conditions. It seems possible that the adaptations of many Cambrian organisms, which composed the trophic nucleus of the Sinsk Algal Lens palaeocommunity to low oxygen tensions allowed them to diversify in the earliest Palaeozoic, especially during the Cambrian. Nowadays these groups comprise only a negligible part of communities and usually survive in settings with low levels of competition. Nonetheless, the organization of the Algal Lens palaeocommunity was not simple, it consisted of diverse trophic guilds. The tiering among sessile filter-feeders was well developed with the upper tier at the 50 cm level. In terms of individuals, the community was dominated by sessile filter-feeders, vagrant detritophages, and diverse carnivores/scavengers. The same groups, but in slightly different order, comprised the bulk of the biovolume: vagrant epifaunal and nektobenthic carnivores/ scavengers, sessile filter-feeders, and vagrant detritophages. -
Geological Survey
DEPARTMENT OF THE INTERIOR UNITED STATES GEOLOGICAL SURVEY ISTo. 162 WASHINGTON GOVERNMENT PRINTING OFFICE 1899 UNITED STATES GEOLOGICAL SUEVEY CHAKLES D. WALCOTT, DIRECTOR Olf NORTH AMERICAN GEOLOGY, PALEONTOLOGY, PETROLOGY, AND MINERALOGY FOR THE YEAR 1898 BY FRED BOUG-HTOISr WEEKS WASHINGTON GOVERNMENT PRINTING OFFICE 1899 CONTENTS, Page. Letter of transmittal.......................................... ........... 7 Introduction................................................................ 9 List of publications examined............................................... 11 Bibliography............................................................... 15 Classified key to the index .................................................. 107 Indiex....................................................................... 113 LETTER OF TRANSMITTAL. DEPARTMENT OF THE INTERIOR, UNITED STATES GEOLOGICAL SURVEY. Washington, D. C., June 30,1899. SIR: I have the honor to transmit herewith the manuscript of a Bibliography and Index o'f North American Geology, Paleontology, Petrology, and Mineralogy for the Year 1898, and to request that it be published as a bulletin of the Survey. Very respectfully, F. B. WEEKS. Hon. CHARLES D. WALCOTT, Director United States Geological Survey. 1 I .... v : BIBLIOGRAPHY AND INDEX OF NORTH AMERICAN GEOLOGY, PALEONTOLOGY, PETROLOGY, AND MINERALOGY FOR THE YEAR 1898. ' By FEED BOUGHTON WEEKS. INTRODUCTION. The method of preparing and arranging the material of the Bibli ography and Index for 1898 is similar to that adopted for the previous publications 011 this subject (Bulletins Nos. 130,135,146,149, and 156). Several papers that should have been entered in the previous bulletins are here recorded, and the date of publication is given with each entry. Bibliography. The bibliography consists of full titles of separate papers, classified by authors, an abbreviated reference to the publica tion in which the paper is printed, and a brief summary of the con tents, each paper being numbered for index reference. -
Prospects for Rewilding with Camelids
Journal of Arid Environments 130 (2016) 54e61 Contents lists available at ScienceDirect Journal of Arid Environments journal homepage: www.elsevier.com/locate/jaridenv Prospects for rewilding with camelids Meredith Root-Bernstein a, b, *, Jens-Christian Svenning a a Section for Ecoinformatics & Biodiversity, Department of Bioscience, Aarhus University, Aarhus, Denmark b Institute for Ecology and Biodiversity, Santiago, Chile article info abstract Article history: The wild camelids wild Bactrian camel (Camelus ferus), guanaco (Lama guanicoe), and vicuna~ (Vicugna Received 12 August 2015 vicugna) as well as their domestic relatives llama (Lama glama), alpaca (Vicugna pacos), dromedary Received in revised form (Camelus dromedarius) and domestic Bactrian camel (Camelus bactrianus) may be good candidates for 20 November 2015 rewilding, either as proxy species for extinct camelids or other herbivores, or as reintroductions to their Accepted 23 March 2016 former ranges. Camels were among the first species recommended for Pleistocene rewilding. Camelids have been abundant and widely distributed since the mid-Cenozoic and were among the first species recommended for Pleistocene rewilding. They show a range of adaptations to dry and marginal habitats, keywords: Camelids and have been found in deserts, grasslands and savannas throughout paleohistory. Camelids have also Camel developed close relationships with pastoralist and farming cultures wherever they occur. We review the Guanaco evolutionary and paleoecological history of extinct and extant camelids, and then discuss their potential Llama ecological roles within rewilding projects for deserts, grasslands and savannas. The functional ecosystem Rewilding ecology of camelids has not been well researched, and we highlight functions that camelids are likely to Vicuna~ have, but which require further study. -
Fossil Flora of the John Day Basin, Oregon, and to Request Its Publication As a Bulletin of the Survey
Bulletin No. 204 Series 0, Systematic Geology and Paleontology, 58 DEPARTMENT OF THE INTERIOR UNITED STATES GEOLOGICAL SURVEY CHARLES D. WALCOTT, DIRECTOR FOSSIL FLORA JOHN DAT BASIN OREGON" BY FRANK MALL KNOWLTON WASHINGTON GOVERNMENT P 11 IN TING OFFICE 1002 CONTENTS. Page. Letter of transrnittal...................................................... 7 Introduction............................................................. 9 Geographic location and topographic features .............................. 10 History of exploration in the John Day Basin.............................. 11 Geological features of the John Day Basin ................................. 14 History of geological investigation in the region ........................ 14 Pre-Cretaceous rocks ................................................. 16 Knoxville and Chico beds ............................................ 17 Clarno formation..................................................... 17 John Day series...................................................... 18 Columbia Elver lava ................................................. 19 Mascall formation .................................................... 19 Kattlesnake formation...'............................................. 19 River terraces......................................................... 20 Localities for fossil plants in the John Day Basin........................... 20 Systematic description of species........................................... 21 Species excluded from this work ......................................... -
Biodiversity Patterns Across the Late Paleozoic Ice Age
Palaeontologia Electronica palaeo-electronica.org Biodiversity patterns across the Late Paleozoic Ice Age Barbara Seuss, Vanessa Julie Roden, and Ádám T. Kocsis ABSTRACT The Late Palaeozoic Ice Age (LPIA, Famennian to Wuchiapingian) witnessed two transitions between ice- and greenhouse conditions. These alternations led to drastic alterations in the marine system (e.g., sea-level, habitat size, sea-surface temperature) forcing faunal changes. To reassess the response of the global marine fauna, we ana- lyze diversity dynamics of brachiopod, bivalve, and gastropod taxa throughout the LPIA using data from the Paleobiology Database. Diversity dynamics were assessed regarding environmental affinities of these clades. Our analyses indicate that during the LPIA more taxa had an affinity towards siliciclastic than towards carbonate environ- ments. Deep-water and reefal habitats were more favored while grain size was less determining. In individual stages of the LPIA, the clades show rather constant affinities towards an environment. Those bivalves and brachiopods with an affinity differ in their habitat preferences, indicating that there might have been little competition among these two clades. Origination and extinction rates are similar during the main phase of the LPIA, whether environmental affinities are considered or not. This underlines that the LPIA marine fauna was well adapted and capable of reacting to changing environ- mental and climatic conditions. Since patterns of faunal change are similar in different environments, our study implies that the changes in faunal composition (e.g., diversity loss during the LPIA; strong increase of brachiopod diversity during the Permian) were influenced by the habitat to only a minor degree but most likely by yet unknown abiotic factors.