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Some Notes on Species of the Genera Bachia and Scolecosaurus

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Some Notes on Species of the Genera Bachia and Scolecosaurus SOME NOTES ON SPECIES OF THE GENERA BACHIA AND SCOLECOSAURUS by Dr. L. D. BRONGERSMA Rijksmuseum van Natuurlijke Historie, Leiden With 4 textfigures Our herpetological collections contain only five specimens referable to the genera Bachia and Scolecosaurus, while the genus Bachia is represented in the Amsterdam Zoological Museum by eight specimens. Scanty though this material may be, some of the specimens proved to be of sufficient interest to justify the publication of these notes. At present thirteen species of Bachia 1) are known, viz., the twelve spe- cies listed in a key by Burt & Burt (1931, pp. 315-316) and Bachia anomala Roux (1929, p. 31). Perhaps Apatelus bresslaui Amaral (1935) must be referred to Bachia too. Roux (1929), Burt & Burt (1931) and Loveridge (1933) published notes on the variation of several characters, and these authors showed that the variation is greater than had been previously sup- posed by Ruthven (1925). This is also apparent from some of the fol- lowing notes. Bachia schlegeli (Dum. & Bibr.) (fig. 1) Chalcides Schlegeli Duméril & Bibron, Erp. gén., vol. 5, 1839, p. 457. Chalcides schlegelii, Fitzinger, Systema Reptilium, 1843, p. 22. Microdactylus schlegeli, Boettger, 22. & 23. Ber. Offenbach. Ver. Naturk., 1883, p. 150. Microdactylus gracilis Tschudi (M. S.), in Duméril & Bibron, Erp. gén., vol. 5, 1839, p. 457; Gray, Cat. Liz. Brit. Mus., 1845, p. 47. Cophias tridactylus, Boulenger, Cat. Liz. Brit. Mus., vol. 2, 1885, p. 418 (part.). Bachia tridactylus, Ruthven, Proc. Boston Soc. Nat. Hist., vol. 38, no. 3, 1925, p. 108 (part.). Herpetochalcis heteropus Boettger, Ber. Offenbach. Ver. Naturk., 1883, p. 150. 1) I have not followed Amaral (1935, pp. 249-250) who divides the genus into two genera Cophias and Chalcides. It escaped Amaral's notice that Cophias Fitzinger, 1843, is preoccupied by Cophias Merrem, 1820, while Chalcides Daudin, 1801 or 1802 (an X = 22. IX. 1801—21. IX. 1802), is preoccupied by Chalcides Laurenti, 1768. 238 L. D. BRONGERSMA Cophias heteropus, Boulenger, Cat. Liz. Brit. Mus., vol. 2, 1885, p. 418. Cophias boettgeri, Boulenger, Cat. Liz, Brit. Mus., vol. 3, 1887, pp. 507, 508. Bachia boettgeri, Burt & Burt, Bull. Am. Mus. Nat. Hist., vol. 61, 1931, p. 316; Parker, Proc. Zool. Soc. Lond., 1935, pp. 517,527. Bachia heteropus, Ruthven, Proc. Boston Soc. Nat. Hist., vol. 38, no. 3, 1925, p. 108. ι ex., $, ?, type, "Calcutta", don. Tschudi, Mus. Leiden, Herp. reg. no. 3580. Chalcides schlegeli Dum. & Bibr. has been referred to the synonymy of Cophias tridactylus (= Bachia tridactyla (Daud.)) by Boulenger (1885, p. 418) and by other authors after him. A re-examination of the type Fig. ι. Bachia schlegeli (Dum. & Bibr.), type, a, left side view of head; b, lower view of head ; c, left fore limb ; d, dorsal scales ; e, anal region. X 7. proved that this point of view is not correct. Bachia schlegeli (Dum. & Bibr.) is a distinct species of which Bachia boettgeri (Blgr.) is almost certainly a synonym. Ruthven (1925, p. 108, note 14) mentions that according to Duméril & Bibron the type of schlegeli has 25 longitudinal rows of scales (round the NOTES ON BACHIA AND SCOLECOSAURUS 239 body). This is an error, however, for Duméril & Bibron (1839, p. 457) wrote that on the back and sides ("du dos et des flancs,,) the scales were placed in 25 longitudinal rows ; on p. 458 they mention 26 rows. Moreover they mention that the type has 8 longitudinal rows of ventral scales. Ruth­ ven (1. c.) apparently supposed that these eight ventral rows were included in the number of 25 scale rows, but this is not the case, as is shown by the re­examination of the type. The type has only two supraoculars; the third supraocular recorded in literature, in my opinion, is a temporal scale. The fore limb bears three clawed digits, but on the hindlimb I find only two digits, unless a rather large scale on the outer border of the foot must be considered as a very much reduced digit. Using the key published by Burt & Burt (1931, pp. 315­316) the type of schlegeli must be referred to the cophias group on account of its quadran­ gular scales ; as an interparietal is present, and as it has more than 30 scale rows the key leads to Bachia boettgeri (Blgr.). With boettgeri it agrees in the number of scale rows (Boettger, 1883, p. 150; Parker, 1935, pp. 517­ 518), in the number of supraoculars, and in the number of digits. The type of boettgeri (Boettger, 1883, p. 150) has two pairs of preanal pores; these are absent in the type of schlegeli, but this may be due to a difference in sex. The greatest difference is found in the arrangement of the preanals. While the type of boettgeri has six preanals, of which the two median are placed one behind the other, with a pair on each side of these, the type of schlegeli has only three preanals (fig. 1 e). This difference may be due to individual variation, for the number and shape of the preanals is subject to rather wide variations, as is shown for Bachia cophias (Schn.) in fig. 3. In the suboculars the type of boettgeri differs by having the posterior sub­ ocular the longest, while in schlegeli the opposite is the case. A comparison of the types may bring to light differences, which are not apparent from the original description by Boettger (1883, PP­ Ι50"Ι51)» but for the present I believe these specimens to be conspecific. Redescription of the type of Bachia schlegeli (Dum. & Bibr.) : Rostral six­sided subtrapezoid, visible from above. Frontonasal six­sided; of the lateral sides the anteriors are much longer than the posteriors ; it is broadest posteriorly, its greatest breadth equalling its length. Prefrontals absent. Frontal almost pentagonal in outline, its posterior tip truncate, forming a short transverse suture with the interparietal; its length twice that of the frontonasal, and twice its greatest breadth. Parietals about equal in length to the frontal; the greatest breadth of each is contained once and a half in its length. An interparietal completely separating the parietals ; its 240 L.D.BRONGERSMA length equal to twice its breadth; it forms a short transverse suture with the frontal; its posterior border rounded; the lateral sides converging anteriorly. Nostril bordered below and posteriorly by the first upper labial ; visible from above. Anteriorly of the nostril, the first labial and the nasal have fused. Loreal about quadrangular, in contact with the frontonasal; these two shields separating the frontal from the nasal. A small preocular; two suboculars, the anterior longest and in contact with the third to fifth upper labials. Two supraoculars, second largest; three supraciliaries, first longest, third largest. Six upper labials, fourth smallest and below the centre of the eye ; second and third in contact with the loreal. Upper labials separated from the parietals. Five large temporals and two smaller ones; these latter more or less form a continuation of the row of upper labials. Parietals in contact with interparietal, frontal, 2nd supraocular, 3rd supra- ciliary, 2 temporals and 2 occipitais (the medial of which is also in contact with the posterior border of the interparietal). Five lower labials, 4th lar­ gest. Symphysial quadrangular, its lateral borders slightly converging pos­ teriorly. Postmental seven-sided. First pair of chinshields five-sided, forming a median suture; shields of the second pair separated by a pair of anterior gulars ; a third pair of much smaller chinshields. Eight enlarged gulars in a transverse row behind the chinshields and anterior gulars. None of the chinshields reaching the oral border. Ventral scales broader than dor­ sals; between the fore limbs a pair of enlarged pectorals, their length equal to once and a half that of the adjoining ventrals; 34 transverse rows of ventrals from the pectorals to the preanals (these not included). Ven­ trals in 8 longitudinal series, their length not quite twice their breadth; 7 transverse rows of scales from the gulars to the pectorals. Dorsal scales rectangular, juxtaposed, in regular transverse rows; 46 of these transverse rows from the occiput to the base of the hind limb; the dorsal scales are about twice as long as broad. The number of scales round the middle of the body varies from 32 to 34. At the level of the 17th transverse row of ventrals, i.e., about midway between pectorals and preanals, the number of scales round the body is 32 ; this same number is found at the level of the 19th and 21st transverse rows of ventrals; at the level of the 18th and 20th rows of ventrals it is 33, and at the 22nd and 23rd it is 34 ; at the level of the 30th row of ventrals (i. е., 5 rows in front of the preanals) the number of scales is 35, and at the level of the 31st and 32nd row it is 37. Anteriorly of the 17th transverse row of ventrals the number of scales becomes less, viz., 31 scales at the level of the 10th row of ventrals. The outer row of lateral scales is partly hidden in a groove which extends along the anterior half of the sides ; these scales are very small, and often they are reduced to two triangles which meet with their tips. NOTES ON BACHIA AND SCOLECOSAURUS 241 Three rectangular preanals in a transverse series, the outer scutes with rounded borders. Their length equals 3 times their breadth. No preanal pores. Colour : Brownish above, lighter below. Tail above with four light stripes, the median pair rather indistinct; the lateral pair is also distinct on the posterior part of the back. Measurements in mm Length of head and body 66 Length of tail (broken) 43+ Length of head 7 Tip of snout to fore limb Distance from fore limb to hind limb qgfyí Length of fore limb 4 Length of hind limb 4 Bachia anomala Roux (fig.
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