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INTRODUCTION Including Hypecoum L
Available Online at http://www.journalajst.com ASIAN JOURNAL OF SCIENCE AND TECHNOLOGY Asian Journal of Science and Technology ISSN: 0976-3376 Vol. 12, Issue, 04, pp.11653-11662, April, 2021 RESEARCH ARTICLE FLORAL DIVERSITY WITHIN PAPAVERACEAE, FUMARIACEAE AND HYPECOACEAE *Wafaa K. Taia ARTICLE INFOAlexandria UniversityABSTRACT-Faculty of Science-Botany Departmen, Alexandria- Egypt Article History: Twenty seven species belonging to eight genera have been investigated in this study. These species Received 14th January, 2021 covered the three restricted families, Papaveraceae, Fumariaceae and Hypecoaceae. The floral Received in revised form characters have been examined carefully, and the herbarium sheets, flowers, stigma, fruits and pollen 20th February, 2021 grains have been photographed. The results indicated that the flower arrangement and symmetry, Accepted 19th March, 2021 stamen number, presence of style, shape of stigma, and type of fruits as well as pollen grain characters th Published online 26 April, 2021 all together proved new taxonomic division of the Papaveraceae s.l.. This investigation supports the separation of the Fumariaceae with two tribes from both the papaveraceae and Hypecoaceae. Key words: Meanwhile, the position of the Hypecoaceae, as subfamily level, under the Papaveraceae is more Floral- Fumariaceae -Hypecoaceae- acceptable. Floral morphological key has been constructed as well as phenogram show the relations Papaveraceae-Taxonomy. between these taxa using SYSTAT 13 program. A correlation analysis of nineteen most important characters has been investigated using SPSS program and an identification key has been constructed. Citation: Wafaa K.Taia, 2021. “Floral diversity within Papaveraceae, Fumariaceae and Hypecoaceae.”, Asian Journal of Science and Technology, 12, (04), 11653-11662. -
Flora of South Australia 5Th Edition | Edited by Jürgen Kellermann
Flora of South Australia 5th Edition | Edited by Jürgen Kellermann PAPAVERACEAE (partly)1 Neville G. Walsh2 (subfam. Fumarioideae) & Jürgen Kellermann3 (family description) Herbaceous annuals or perennials, sometimes becoming shrubby as the inflorescence develops; most parts of the plant produce latex and contain alkaloids, leaves entire and often deeply dissected, pinnately or palmately compound, exstipulate. Inflorescence cymose or racemose, often a thyrse with leaf-like or membranous bracts; flowers bisexual, sepals 2 or 3, caducous; petals 4 or 6; stamens 4, 6 or numerous; ovary superior, carpels 2 or 3 or numerous (not in S.A.); in FUMARIOIDEAE : flowers either almost regular with petals in 2 whorls of differently shaped petals, stamens 4 (Hypecoum only), or flowers zygomorphic with sepals 2, in a lateral position, usually the same colour and texture as the corolla, and petals 4, in two whorls, with the 2 lateral ones being the inner ones, with the large dorsal one pouched or spurred at the base (with nectary scale), stamens 6, joined in an anterior and a posterior bundle, each consisting of 3 stamens, ovary surmounted by a style with a terminal 2- or 3-fid stigma, carpels 2, ovule 1 or more (outside S.A.); in PA P AVEROIDEAE : flowers regular, stamens numerous, ovary unilocular with numerous ovules; flowers regular, sepals caducous, stamens 4, ovary unilocular with numerous ovules. Fruit a capsule opening by valves or pores; seeds with small embryo, endosperm mealy or oily. Poppies, fumitories. The family is distributed throughout the temperate N hemisphere with some species in E Africa and S America; often grow in open areas or disturbed sites. -
Evolutionary History of Fumitories (Subfamily Fumarioideae, Papaveraceae): an Old Story Shaped by the Main Geological and Climatic Events in the Northern Hemisphere Q
Molecular Phylogenetics and Evolution 88 (2015) 75–92 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Evolutionary history of fumitories (subfamily Fumarioideae, Papaveraceae): An old story shaped by the main geological and climatic events in the Northern Hemisphere q Miguel A. Pérez-Gutiérrez a, Ana T. Romero-García a, M. Carmen Fernández b, G. Blanca a, ⇑ María J. Salinas-Bonillo c, Víctor N. Suárez-Santiago a, a Department of Botany, Faculty of Sciences, University of Granada, c/ Severo Ochoa s/n, 18071 Granada, Spain b Department of Cell Biology, Faculty of Sciences, University of Granada, c/ Severo Ochoa s/n, 18071 Granada, Spain c Department of Biology and Geology, University of Almería, c/ Carretera de Sacramento s/n, 04120 Almería, Spain article info abstract Article history: Fumitories (subfamily Fumarioideae, Papaveraceae) represent, by their wide mainly northern temperate Received 21 July 2014 distribution (also present in South Africa) a suitable plant group to use as a model system for studying Revised 30 March 2015 biogeographical links between floristic regions of the Northern Hemisphere and also the Southern Accepted 31 March 2015 Hemisphere Cape region. However, the phylogeny of the entire Fumarioideae subfamily is not totally Available online 7 April 2015 known. In this work, we infer a molecular phylogeny of Fumarioideae, which we use to interpret the bio- geographical patterns in the subfamily and to establish biogeographical links between floristic regions, Keywords: such as those suggested by its different inter- and intra-continental disjunctions. The tribe Hypecoeae Ancestral-area reconstruction is the sister group of tribe Fumarieae, this latter holding a basal grade of monotypic or few-species genera Biogeography Fumarioideae with bisymmetric flowers, and a core group, Core Fumarieae, of more specious rich genera with zygomor- Molecular dating phic flowers. -
The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development
The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development A thesis presented to the faculty of the College of Arts and Sciences of Ohio University In partial fulfillment of the requirements for the degree Master of Science Alastair R. Plant August 2013 © 2013 Alastair R. Plant. All Rights Reserved. 2 This thesis titled The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development by ALASTAIR R PLANT has been approved for the Department of Environmental and Plant Biology and the College of Arts and Sciences by Stefan Gleissberg Assistant Professor of Environmental and Plant Biology Robert Frank Dean, College of Arts and Sciences 3 ABSTRACT PLANT, ALASTAIR R., M.S., August 2013, Plant Biology The Regulation of Ontogenetic Diversity in Papaveraceae Compound Leaf Development Director of Thesis: Stefan Gleissberg The leaf is almost ubiquitous throughout land plants but due to its complex and flexible developmental program is highly morphologically variable between taxa. Description of the functions of regulatory genes key to leaf development in different evolutionary lineages allows the study of changes in developmental mechanisms through evolutionary time as a means for anatomical and morphological diversification. The roles of homologs of CINCINNATA-like TCP family genes, ARP genes, and Class I KNOX genes were investigated in two members of the Papaveraceae, a basal eudicot lineage positioned in between major angiosperm groups, by phylogenetic analysis, in situ hybridization, expression profiling by quantitative polymerase chain reaction, and virus- induced gene silencing in Eschscholzia californica and Cysticapnos vesicaria. Expression data were similar to those for homologous genes in core eudicot species, however, some gene functions found in core eudicots were not associated with basal eudicot homologs, and so have either been gained or lost from the ancestral state. -
Department of Environment, Water and Natural Resources
Photograph: Helen Owens © Department of Environment, Water and Natural Resources, Government of South Australia Department of All rights reserved Environment, Copyright of illustrations might reside with other institutions or Water and individuals. Please enquire for details. Natural Resources Contact: Dr Jürgen Kellermann Editor, Flora of South Australia (ed. 5) State Herbarium of South Australia PO Box 2732 Kent Town SA 5071 Australia email: [email protected] Flora of South Australia 5th Edition | Edited by Jürgen Kellermann PAPAVERACEAE (partly)1 Neville G. Walsh2 (subfam. Fumarioideae) & Jürgen Kellermann3 (family description) Herbaceous annuals or perennials, sometimes becoming shrubby as the inflorescence develops; most parts of the plant produce latex and contain alkaloids, leaves entire and often deeply dissected, pinnately or palmately compound, exstipulate. Inflorescence cymose or racemose, often a thyrse with leaf-like or membranous bracts; flowers bisexual, sepals 2 or 3, caducous; petals 4 or 6; stamens 4, 6 or numerous; ovary superior, carpels 2 or 3 or numerous (not in S.A.); in FUMARIOIDEAE : flowers either almost regular with petals in 2 whorls of differently shaped petals, stamens 4 (Hypecoum only), or flowers zygomorphic with sepals 2, in a lateral position, usually the same colour and texture as the corolla, and petals 4, in two whorls, with the 2 lateral ones being the inner ones, with the large dorsal one pouched or spurred at the base (with nectary scale), stamens 6, joined in an anterior and a posterior bundle, each consisting of 3 stamens, ovary surmounted by a style with a terminal 2- or 3-fid stigma, carpels 2, ovule 1 or more; in PA P AVEROIDEAE : flowers regular, stamens numerous, ovary unilocular with numerous ovules; flowers regular, sepals caducous, stamens 4, ovary unilocular with numerous ovules. -
Bbyct-133 Plant Ecology and Taxonomy Nomenclature and Systems of Classification
BBYCT-133 PLANT ECOLOGY AND Indira Gandhi TAXONOMY National Open University School of Sciences Block 4 NOMENCLATURE AND SYSTEMS OF CLASSIFICATION UNIT 16 Binomial Nomenclature 109 UNIT 17 Scientific Naming of Plants: Nomenclature 120 UNIT 18 System of Classification 141 UNIT 19 Biometrics, Numerical Taxonomy 161 UNIT 20 Cladistics 172 Course Design Committee Prof. A.K. Bhatnagar (Retd.) School of Sciences, Department of Botany, IGNOU University of Delhi, Delhi-110054 Prof. M.S. Nathawat, Director, Dr. A.K. Kavathekar (Retd.) Prof. Vijayshri, Director (Ex.) Sr. Consultant, Prof. Amrita Nigam Department of Botany, Prof. Jaswant Sokhi Sri Venkateswara College, University of Delhi, New Delhi-110001 Block Preparation Team Prof. Amrita Nigam Dr. A.K. Kavathekar (Retd.) School of Sciences, IGNOU Sr. Consultant Department of Botany Prof. Jaswant Sokhi Sri Venkateswara College, School of Sciences, IGNOU University of Delhi, Dr. E. Chauhan (Unit-20) New Delhi-110001 Sr. Consultant Department of Botany Deshbandhu College, University of Delhi, New Delhi-110019 Course Coordinators: Prof. Amrita Nigam and Prof. Jaswant Sokhi Production Mr. Sunil Kumar AR(P), SOS, IGNOU Acknowledgements: Dr. Eklavya Chauhan for giving useful inputs. Sh. Manoj Kumar, Assistant for word processing and CRC preparation. Mr. Ajit Kumar for diagrams. November, 2019 Indira Gandhi National Open University, 2019 ISBN: All rights reserved. No part of this work may be reproduced in any form, by mimeograph or any other means, without permission in writing from Indira Gandhi National Open University. Further information on Indira Gandhi National Open University courses may be obtained from the University’s office at MaidanGarhi, New Delhi-110 068 or IGNOU website www.ignou.ac.in. -
Evolution of Reproductive Morphology in the Papaveraceae S.L. (Papaveraceae and Fumariaceae, Ranunculales)
® International Journal of Plant Developmental Biology ©2010 Global Science Books Evolution of Reproductive Morphology in the Papaveraceae s.l. (Papaveraceae and Fumariaceae, Ranunculales) Oriane Hidalgo • Stefan Gleissberg* Department of Environmental and Plant Biology, Ohio University, 500 Porter Hall, Athens, Ohio 45701, USA Corresponding author : * [email protected] ABSTRACT Flower bearing branching systems are of major importance for plant reproduction, and exhibit significant variation between and within lineages. A key goal in evolutionary biology is to discover and characterize changes in the genetic programming of development that drive the modification and diversification of morphology. Here we present a synopsis of reproductive architecture in Papaveraceae s.l., a lineage in which the evolution of inflorescence determinacy, flower structure and symmetry, and effloration sequence produced unique reproduc- tive syndromes. We discuss the potential of this group to study key issues on the evolution of reproductive structures, and refer to candidate gene families, choice of landmark species, and available tools for developmental genetic investigations. _____________________________________________________________________________________________________________ Keywords: effloration sequence, flower organ identity, flower symmetry, Fumariaceae, inflorescence determinacy Abbreviations: AP1, APETALA1; AP3, APETALA3; CEN, CENTRORADIALIS; CRC, CRABS CLAW; CYC, CYCLOIDEA; FLO, FLORICAULA; FUL, FRUITFULL; LFY, LEAFY; PI, PISTILLATA; TFL1, -
The Seed Atlas of Pakistan-. Papaveraceae
Pak. J. Bot., 48(3): 1035-1044, 2016. THE SEED ATLAS OF PAKISTAN-. PAPAVERACEAE RUBINA ABID,* AFSHEEN ATHER AND M. QAISER Department of Botany University of Karachi, Karachi, Pakistan *Corresponding author’s e-mail: [email protected] Abstract Numerical analysis based on seed morphological characters of 14 taxa belonging to the family Papaveraceae is carried out. Seed macro and micro morphological characters were found useful to strengthen the taxonomic decisions and trace out the phylogenetic relationship within the family Papaveraceae. Key words: Seed morphology, Papaveraceae, Numerical analysis, Pakistan. Introduction out the phylogenetic relationship of the taxa within the family Papaveraceae from Pakistan. The family Papaveraceae comprises 43 genera and 820 species mostly distributed in north temperate and tropical Materials and Methods regions (Mabberley, 2008). In Pakistan the family is represented by 14 species distributed in 6 genera viz., Mature and healthy seeds of 14 taxa of the family Argemone L., Eschscholtzia Cham., Glaucium Mill., Papaveraceae were collected from herbarium specimens. Hypecoum L., Papaver L. and Roemeria Medic (Jafri & Mostly 10 plants/species and 10 seeds/plant were studied Qaiser, 1974). Previously seed morphological studies were (Appendix-I) and examined under stereomicroscope (Nikon not only used as an additional taxonomic tool but also to XN Model), compound microscope (Nikon type 102) and predict the evolutionary relationship of the taxa at various scanning electron microscope (JSM-6380A). For scanning levels (Elisens & Tomb, 1983; Qaiser, 1987; Hufford, 1995; electron microscopy dry seeds were directly mounted on Akbari & Azizian, 2006: Ather et al., 2009; Rajbhandary & metallic stub using double adhesive tape and coated with Shrestha, 2010; Kanwal et al., 2012; Abid et al., 2013). -
Random Local Clock and Molecular Evolution Studies on Corydalis (Papaveraceae S
International Journal of Biology; Vol. 4, No. 4; 2012 ISSN 1916-9671 E-ISSN 1916-968X Published by Canadian Center of Science and Education Random Local Clock and Molecular Evolution Studies on Corydalis (Papaveraceae s. l.) Sophia Wan-Pyo Hong1 1 Natural Products Research Institute, Department of Pharmacy, Seoul National University, Seoul, South Korea Correspondence: Sophia Wan-Pyo Hong, Natural Products Research Institute, Department of Pharmacy, Seoul National University, Seoul 151-742, South Korea. Tel: 82-010-2806-7925. E-mail: [email protected] Received: April 11, 2012 Accepted: April 23, 2012 Online Published: August 22, 2012 doi:10.5539/ijb.v4n4p31 URL: http://dx.doi.org/10.5539/ijb.v4n4p31 Abstract Molecular evolutionary rate and dating studies have been carried out on nuclear ribosomal DNA (nrDNA ITS I and II) and non-coding plastid DNA (rps16 intron) nucleotide sequences from Corydalis and other related genera in the poppy family Papaveraceae s. l. Strict, relaxed, and local clock models have been compared. Although it has been suggested that the nrDNA ITS I and II regions evolved in concert, they may have evolved at different evolutionary rates under differential constraints. Based on random local clock and uncorrelated relaxed lognormal clock Bayesian analyses using BEAST v.1.6.2, the ITS II region may have evolved in a clock-like fashion. Keywords: local clock, Corydalis, molecular clock, nrDNA, rps16, Bayesian analysis 1. Introduction 1.1 Molecular Evolutionary Rate Models Methodologies for estimating molecular evolutionary -
MUELLERIA National Herbarium of Victoria, Royal Botanic Gardens Victoria
MUELLERIA National Herbarium of Victoria, Royal Botanic Gardens Victoria Title and Author pp Volume 1(1), 1955 Foreword JS. Turner 3 Preface AW. Jessep 5 New species and varieties of Stylidium from Western Australia. R. Erickson & JH. Willis 7 A new species of Eria (Orchidaceae). TE. Hunt 21 Systematic notes on Victorian Compositae - 1. (Olearia ). JH. Willis 24 The Eucalyptus species of Cavanilles. AK. Cameron 34 A new species of Pestalotiopsis (Fungi Imperfecti) on Pittosporum bicolor. AB.Court 43 Changes in the nomenclature of three Victorian monocotyledons. JH. Willis & AB Court 45 Robert Brown's Bass Strait journal of April/May, 1802 (A transcription). JH. Willis & CI. Skewes 46 Robert Brown's collectings in Victoria. JH. Willis 51 Notes on the growth of an English elm. PF. Morris 54 The present position of muscology in Victoria (a centennial review). JH. Willis 55 A remarkable lichen from arid Australia. P. Bibby 60 A bibliography of the Australian baobab. JH. Willis 61 Recent changes in the nomenclature of three Australian conifers. PF. Morris 64 Volume 1(2), 1959 Australian species of the fungal genus Cordyceps. JH. Willis 67 Orthography of certain species Epithets. JH. Willis 90 Reduction of the lichen genus Bibbya. JH. Willis 91 Notes on the vegetation of Eucla District, WA. JH. Willis 92 Plants of the Recherche Archipelago, WA. JH. Willis 97 New species and varieties of Ptilotus R. Br. (Amaranthaceae). G. Benl 101 Reinstatement of Calotis suffruticosa Domin (Compositae). GL. Davis 109 Two new Australian species of Brachycome Cass. (Compositae). GL. Davis 111 Studies in Mimosaceae Part1. AB. -
PAPAVERACEAE, the Poppy Family Herbs Or Shrubs, Often with Milky Or Colored
PAPAVERACEAE, the Popp y Family Robert F. C. Naczi Keywords: Papaveraceae, Adlumia, Argemone, Capnoides, Chelidonium, Corydalis, Dicentra, Eschscholzia, Fumaria, Glaucium, Lamprocapnos, Macleaya, Papaver, Pseudofumaria, Sanguinaria, Stylophorum, Flora, Floristics, North America http://dx.doi.org/10.21135/893275471.024 PAPAVERACEAE, the Poppy Family Herbs or shrubs, often with milky or colored sap; lvs mostly alternate, dissected or lobed; fls often large, radially or bilaterally symmetric, perfect, hypogynous; sep usu 2(3), fully or partially enclosing bud, usu caducous; pet ≥4 (rarely 0), distinct or inner pair connivent, often showy; stamens 6 or numerous; nectaries none or present as spurs at bases of filaments; pistil of 2(–4), seldom (as in Argemone and Papaver) more numerous carpels, connate to form compound, typically 1-locular ovary; ovules numerous on parietal placentae, but the latter sometimes (as in Papaver) deeply intruded as partial partitions, or ovary seldom (as in Glaucium) fully partitioned into 2 locules; fr typically a 1-locular capsule, dehiscent by partial or complete abscission of elongate valves usu alternating with placentae, the latter forming persistent replum, or valves sometimes (as in Argemone and Papaver) reduced and subapical, so that capsule opens by ring of pores, or fr sometimes dry and indehiscent (e.g., in Fumaria; capsule 2-locular with deciduous valves in Glaucium); seeds with copious endosperm and small, dicotyledonous embryo, often arillate. 44/780, mostly N. Temperate. Morphologic synapomorphies: perianth 2-merous, sep caducous, carpels connate, frs capsular, placentation parietal. Two subfamilies, Fumarioideae and Papaveroideae, occur in our area. Authors of recent floristic works often treat both of these taxa as families. -
Potential Biological Control Agents for Fumitory (Fumaria Spp.) in Australia
Potential biological control agents for fumitory (Fumaria spp.) in Australia M. Jourdan,1 J. Vitou,1 T. Thomann,1 A. Maxwell2,3 and J.K. Scott2 Summary Fumaria species are increasingly problematic in the cropping regions of southern Australia, and one fumitory, Fumaria densifloraDC, has developed populations with herbicide resistance. Consequently, the potential for biological control was assessed. Nine species of fungi were found associated with Fumaria spp. in a survey of 33 sites in southern France. According to the literature, species potentially host specific to fumitory includeCladosporium brachormium Berk. and Broome, Entyloma fumariae J. Schröt. and Peronospora affinis Rossmann. Of the insects detected on Fumaria spp. in France, the stem weevil, Sirocalodes mixtus Mulsant and Rey has potential as a biological control agent because it is thought to be host specific. None of these species were detected amongst the six pathogen spe- cies found during surveys of 64 locations in southeastern and southwestern Australia. The absence of pathogens and insects associated with Fumaria species in Australia, the lack of Fumaria spp. native to Australia, and few closely related crops or ornamental species, indicate that there are opportunities for research into the development of natural enemies for the biological control of fumitory. Keywords: Fumaria species, biological control, field surveys, fungal pathogens, arthropods. Introduction species and two subspecies: Fumaria bastardii Boreau, Fumaria capreolata L., Fumaria capreolata L. subsp. Fumitory species are weeds of many parts of the world, capreolata, F. densiflora, Fumaria indica (Hausskn.) mainly in cereal and legume cultivation, vineyards, Pugsley, Fumaria muralis Sond. ex W.D.J.Koch, Fu- wastelands and gardens, but have not been considered maria muralis Sond.