135 125 115 105 130 120 110 Age Period Cretaceous Chronostratigraphy Epoch Early Standard ICS Aptian Albian Stage 129.97 133.9 125.0 112.0 Valanginian Hauterivian Barremian Lt. Albian M. Albian E. Albian Substage Lt. Haut. E. Haut. E. Val. Lt. Val. E. Apt. - M. Apt. - Lt. Apt. - E. Barrem. Lt. Barrem. Bedoulian Gargasian Clansayesian magnetic Polarity Geo- M10N M"-1"r M12A M11A M"-2" M"-3" (C34 (C34 (C34 M12 M11 M10 r set r set C34 M0r M9 M8 M7 M6 M5 M3 M1 n) n) n)

"Cretaceous Normal Super-Chron M-Sequence ("Cretaceous Quiet Zone")" Craspedodiscus discofalcatus Prodeshayesites fissicostatus Parancyloceras bidentatum/ Haplocrioceras fissicostatum Douvilleiceras mammillatum Paracrioceras denckmanni Paracrioceras rarocinctum Parahoplites nutfieldiensis Milanowskia speetonensis Cheloniceras martinoides Ancyloceras inexum/S. Simanocyloceras stolleyi Deshayesites deshayesi Speetoniceras inversum Leymeriella tardefurcata Hypacanthoplites jacobi Dicostella tuberculatum Tropaeum bowerbanki Mortoniceras inflatum Endemoceras regale Olcostephanus spp. Ende. amblygonium Euhoplites loricatus Hoplites dentatus Euhoplites lautus Crasp. gottschei Prodichotomites Paracr. elegans Ende. noricum Paran. scalare Desh. forbesi Dichotomites pingue Boreal C HRONOS Dichotomites bidichotomoides Prodichotomites polytomus Prodeshayesites obsoletus Parahoplites nutfieldiensis Hypacanthoplites anglicus Mojsisovicsia subdelaruei Protohoplites puzosianus Cheloniceras parinodum Deshayesites callidiscus Hysteroceras varicosum Anahoplites intermedius Prodichotomites ivanovi Tropaeum subarcticum Otohoplites raulinianus Prodich. hollwedensis Deshayesites grandis Simbirskites variabilis Dufrenoyia transitoria Otohoplites bulliensis Dimorphoplites niobe Cheloniceras buxtorfi Leymeriella regularis Cleoniceras floridum Diploceras cristatum Chelon. meyendorffi Euhop. meandrinus Callihoplites auritus Anahoplites daviesi Leym. schrammeni Pseudosonneratia Leym. acuticostata Simbir. marginatus Nolaniceras nolani Sonneratia kitchini Dich. triptychoites Euhoplites nitidus Prodeshay. bodei Hypa. rubricosus Sonn. perinflata Lyelliceras lyelli Chelon. gracile Hoplites spathi Deshay. kiliani Chelon. debile Deshay. fittoni Hyst. orbignyi Dich. crassus steinmanni Ammonites

Creator Precambrian

Cen Paleozoic Mesozoic ScaLe Time R Same asBoreal(NWEurope) Kotetishvilia compressissima Busnardoites campylotoxus Lyticoceras nodosoplicatus Ancyloceras vandenheckii Hemihoplites feraudianus Pseudothurmannia ohmi Deshayesites deshayesi Saynoceras verrucosum Pleisiospitidiscus ligatus Criosarasinella furcillata Hypacanthoplites jacobi Acanthodiscus radiatus Parahoplites melchioris Neocomites peregrinus Tirnovella pertransiens subnodosocostatum Deshay. oglanlensis Colchidites sarasini Gerhardtia sartousi Balearites balearis Nolaniceras nolani Nicklesia pulchella Dufrenoyia furcata Subsaynella sayni Pseudocrioceras Crioceratites loryi Epicheloniceras Spitidiscus hugii Imerites giraudi Coronites darsi Deshay. weissi waagenoides Nick. nicklesi Tethyan Olcostephanus (J.)jeannoti Saynoceras verrucosum Teschenites callisdiscus Criosarasinella furcillata Neocomites peregrinus Olcostephanus nicklesi Hypacanthoplites Crioceratites loryi Karakaschiceras Karakaschiceras nodosocostatum Diodochoceras pronecostatum campylotoxus Busnardoites biassalense rubricosus Sequences Global Barr1 Barr2 Barr3 Barr4 Barr5 Barr6 Ha1 Va6 Va7 Va5 Va4 Ha4 Va3 Ha7 Ap6 Ap1 Ap2 Ap3 Ap4 Ap5 Al 1 Al 2 Al 3 Al 4 Al 5 Al 6 Al 7 Al 8 Al 9 Va2 Ha3 Ha5 138.2 136.8 136.2 135.7 135.4 134.6 133.6 132.5 130.1 129.6 129.1 128.6 128.4 127.8 126.2 125.0 124.6 124.0 121.0 109.2 108.2 107.9 106.4 105.1 102.3 117.9 112.4 110.5 111.4 111.0 Sequences Cycles T-R Major T-R Cycles Rotalipora subticinensis Globigerinelloides blowi Globuligerina hoterivica Ticinella praeticinensis Hedbergella planispira Ticinella bejaouaensis Rotalipora ticinensis Hedbergella similis Hedbergella sigali/ Globigerinelloides Globigerinelloides Leupoldina cabri Planktonic Foraminifers Ticinella primula Hedbergella gorbachikae ferreolensis delrioensis Zones algeriana Globigerinelloides blowi Leupoldina cabri(rare) Ticinella praeticinensis Ticinella bejaouaensis Rotalipora ticinensis Hedbergella similis Hedbergella sigali/ Globigerinelloides Leupoldina cabri Zonal Markers Ticinella primula (consistent) delrioensis algeriana Globuligerina oxfordiana,G.stellapolaris, G. caucasica,megamonica Rotalipora subticinensis Ticinella bejaouaensis Globigerinelloides Leupoldina cabri algeriana Hauerina (Wellmannella) antiqua Lenticulina schreiteri,Haplophragmium Arenobulimina macfadyeni,D. Boreal ZonalMarkers Haplophragmium aequale Coronotalites bartensteini Coronotalites intercedens Citharinella pinnaeformis Citharina harpa,C.seitzi Gavelinella cenomanica D. (Marssonella)kummi Gavelinella sigmoicosta Simplorbites papyracea Lenticulina saxonica Gavelinella barremiana Conorotalites aptiensis Gavelinella intermedia Hoeglundia carpenteri Gavelinella brielensis inconstans erectum P. subfusiformis gradata Hauerina (Wellmannella) antiqua Hechtina praeantiqua,C.seitzi Lenticulina (Gaudriyadhella) Haplophragmium aequale, Arenobulimina macfadyeni Gavelinella sigmoicosta, Hoeglundina carpenteri Gavelinella barremiana Hoeglundina chapmani Conorotalites aptiensis Epistomina caracolla Lenticulina schreiteri Gaudrynia dividens Saracenaria valanginiana Citharina harpa ouachensis Other BorealForaminifers E. (Pseudotextulariella)cretosa Saracenaria (Grimsdaleinella) spinosa, Gaudryinadividens, Pleurostomella praebulbosa Citharina sparsicostata,T. Arenobulimina chapmani, Hoeglundina chapmani, Quinqueloculina antiqua Conorboides lamplughi Gaudryinella sherlocki Epistomina spinulifera Textularia bettenstaedti Textularia chapmani Citharina acuminata Eggerellina mariae Epistomina ornata Gavelinella baltica Citharinella karreri Citharina discors C. pseudodiscors hauteriviana, M.bettenstaedi D. moorbergensis P. schloenbachi D. praeoxycona S. bettenstaedti Valvulineria sp. pseudoroemeri Lenticulina heiermanni, Lenticulina ouachensis C. chapmani Lenticulina hauteriviana C. d'orbignyi V. procera D. hechti P. prima S. lata ouachensis Smaller BenthicForaminifers Citharinella karreri,Conorboides E. (Pseudotextulariella)cretosa, hauteriviana, Lenticulinaguttata Lenticulina saxonica,Citharina Saracenaria (Grimsdaleinella) Lenticulina hauteriviana Citharina sparsicostata Lenticulina heiermanni Epistomina spinulifera Epistomina ornata Epistomina hechti T. pseudoroemeri T. emslandensis P. subfusiformis H. reticulata S. asperula lamplughi P. prima spinosa discors P. subnodosa,O.aff.brotzeni, Gavelinella barremianas.s. Gavelinella barremianas.l. Osangularia schloenbachi Conorotalites bartensteini Gavelinella sigmoicosta Lenticulina eichenbergi Gavelinella intermedia H. vocontianus D. hauteriviana D. ouachensis Tethyan markers G. brotzeni Updated byJamesG.Ogg(PurdueUniversity) Gavelinella sigmoicosta Gavelinella barremiana D. (Epistomina)hechti Spirillina neocomiana D. hauteriviana O. aff.brotzeni D. ouachensis F. bidentata H. vocontianus D. praeoxycona,Gavelinellaaff. Other Tethyan markers Plectorecurvoididae alternans D. plummeri,S.complanata Coronotalites intercedens G. eichenbergi bartensteini, D.zedlerae D. (Marssonella)cf.kummi Lenticulina ouachensis Textularia bettenstaedti Conorotalites aptiensis H. nonioninoides barremania G. flandrini S. carinata praecursor Saracenaria (Grimsdaleinella) Lenticulina (Gaudriyadhella) Lenticulina eichenbergi L. o.bartensteini D. praeoxycona M. reticulosa D. zedlerae ouachensis L. besairlei G. flandrini S. carinata M. sardoa spinosa Mesozoic andCenozoicSequenceChronostratigraphicFrameworkofEuropeanBasinsin:DeGraciansky, P.- C.,Hardenbol,J.,Jacquin,Th.,Vail, P. R.,andFarley, M.B.,eds.;MesozoicandCenozoicSequenceStratigraphyofEuropeanBasins,SEPMSpecialPublication60. Orbitolina (Mesorbitolina.)texana, Rectodictyoconus giganteus, D. pyrenaicus,I.hungarica Campanellula capuensis Praereticulinella cuvillieri Praereticulinella reicheli Palorbitolina lenticularis Simplorbitolina manasi Praeorbitolina cormyi A. reicheli(Adriatic) O. cuvillieri S. conulus Larger BenthicForaminifers (Adriatic) Zonal Markers Rectodictyoconus giganteus,O. Praereticulinella cuvillieri(+ A. Campanellula capuensis Pfenderina neocomiensis V. camposaurii(Adriatic) Praereticulinella reicheli Palorbitolina lenticularis Simplorbitolina manasi Praeorbitolina cormyi reicheli in Adriatic) D. pyrenaicus P. salevensis S. conulus C.? favrei (Adriatic) cuvillieri Based, inpart,on:CENOZOIC-MESOZOICBIOCHRONOSTRATIGRAPHY: JANHARDENBOL,JACQUESTHIERRY, MARTINB.FARLEY, THIERRY JACQUIN,PIERRE-CHARLESDEGRACIANSKY, AND PETERR.VAIL,1998. Other LargerBenthic and GabiOgg pachymarginalis, I. O. (M.)parva,D. O. debelmasi,P. jourdanensis, U. Foraminifers alpillensis rubiensis V. aldanchellamiliani P. tunesiana(Adriatic) P. jourdanensis,U. pachymarginalis I. rubiensis,D. N. friburgensis O. debelmasi O. (M.)parva alpillensis Early Cretaceous Cretaceous Early CC3 CC4 CC5 CC6 CC7 CC8 CC9 to: Zones a b a b a b a b a b a NC10 NC3 NC4 NC5 NC6 NC7 NC8 NC9 GEOLOGIC TIMESCALE2004 a b d e a b a b a b a b a c c Flabellites oblongus(consistent) Rucinolithus terebrodentarius, Prediscosphaera columnata Axopodorhabdus albianus Rhagodiscus gallagheri Eiffellithus monechiae Eiffellithus turriseiffelii Tranolithus orionatus Hayesites irregularis Hayesites albiensis Lithraphidites bollii Eiffellithus windii Eiffellithus striatus Eprolithus floralis Ruc. windleyae Zonal Markers Lithraphidites bollii Micrantholithus hoschulzii Calcicalathina oblongata Tubodiscus verenae Crucielipsis cuvillieri Speetonia colligata Conusphaera rothii Micranthololithus speetonensis,R. Zeugrhabdotus trivectis Cribrosphaerella ehrenbergii Rhadodiscus achlyostaurion Braarudosphaera africana Nannoconus truitti(acme) Rhagodiscus hamptonii Zeugrhabdotus scutula Rhagodiscus angustus Acaenolithus viriosus Other Nannofossils Stoverius achylosum Radiolithus planus dekaenelii Calcareous Nannofossils N. regularis N. bucheri Nannoconus truitti(acme) Prediscosphaera spinosa Retecapsa angustiforata Seribiscutum primitivum NANNOCONID CRISIS Nannoconus steinmanii Eiffellithus primus,C.deflanderi Eiffellithus striatus Rucinolithus wisei Eiffellithus windii (Gradstein, F.M., Ogg,J.G.,Smith, A.G., etal.,2004)and Farhania varolii Zones BC25a BC25b BC10 BC11 BC12 BC13 BC14 BC15 BC16 BC17 BC18 BC19 BC20 BC21 BC22 BC23 BC24 BC26 BC4 BC5 BC6 BC7 BC8 BC9 (103-138 Ma time-slice) Ma (103-138 Zeugrhabdotus scutula(acme) Lithraphidites moray-firthensis Prediscosphaera columnata Tegulalithus septentrionalis Boreal ZonalMarkers Axopodorhabdus albianus Rhagodiscus gallagheri Nannoconus abundans Eiffellithus monechiae Nannoconus borealis Tranolithus orionatus Eprolithus? antiquus Farhania varolii Triquetrorhabdulus shetlandensis Micranthololithus speetonensis Zeugrhabdotus scutula(acme) Lithraphidites moray-firthensis Tegulalithus septentrionalis Clepsilithus maculosus Ceratolithina bicornuta Nannoconus borealis Eprolithus? antiquus Eiffellithus striatus Farhania varolii Other BorealNannofossils Influx ofCrucielipsiscuvilleri&Speetonia Zeugrhabdotus diplogrammus, Hayesites irregularis,Flabellites Ceratolithina bicornuta,Owenia Rucinolithus terebrodentarius Braarudosphaera africana, pseudoseptentrionalis Repagulum parvidentatum Perissocyclus plethotretus Nannoconus truitti(acme) Gartnerago praeobliquum Eiffellithus striatus Seribiscutum primitivum, Tequlalithus tessellatus, Cretarhabdus inaequalis Zeugrhabdotus scutula, Tegumentum octiformis Rucinolithus windleyae Crucicribrum anglicum Acaenolithus viriosus Ceratolithina hamata Eiffellithus windii Conusphaera rothii Acaenolithus sp.1 Radiolithus planus Eprolithus floralis, Nannoconus Zeug. trivectis oblongus colligata hillae Diazomatolithus lehmanii(boreal) Rhagodiscus pseudoangustus Cyclagelosphaera margerelii Zeugrhabdotus noeliae(acme), Crucibiscutum salebrosum Nannoconus truitti(acme) Micrantholithus hoschulzii Stradnerlithus silvaradius Nannoconus abundans Braloweria boletiformis Tubodiscus verenae Crucielipsis cuvillieri Rhagodiscus asper Speetonia colligata Conusphaera rothii Micrantholithus (acme) (acme) D. nan. S. cret. S. terr. (un-na C. int. P. ali. oper. med) Zones O. The CONCISEGEOLOGICTIMESCALE mag. vet. N. K. sim. B. long. C. ret. V. P. ana. P. cret. Nelchinopsis kostromiensis,Muderongia Pseudoceratium anaphrissum Carpodinium obliquicostatum Palaeoperidinium cretaceum Batioladinium longicornutum Cribroperidinium intricatum Subtilisphaera terrula Systematophora cretacea Odontochitina operculata Hapsocysta peridictya Ovoidinium diversum Endoceratium turneri Muderongia staurota Pterodinium aliferum Discorsia nanna Zonal Markers crucis Kleithriasphaeridium corrugatum Gonyaulacysta teicha, A. eilema, Pseudoceratium anaphrissum Batioladinium longicornutum Nelchinopsis kostromiensis Lagenorhytis delicatula,M. North Atlantic Systematophora cretacea, Canningia cf.reticulata(of Pseudoceratium pelliferum Heslertonia heslertonensis Muderongia staurota Pterodinium aliferum Duxbury, 1977) macwhaei group Kleithriasphaeridium simplicispinum Surculosphaeridium? longifurcatum Hystrichosphaeridium arborispinum P. spinocristatum,P. polymorphum Ophiobolus sp. A ofDavey(1979) Cyclonephelium membraniphorum Kleithriasphaeridium? sarmentum, Batioladinium varigranosum, Stiphrosphaeridium antophorum Exochosphaeridium phragmites Surculosphaeridium trunculum, Nematosphaeropsis scala Exiguisphaera phragma, Oligosphaeridium abaculum, Carpodinium granulatum,C.? Pervosphaeridium truncatum, Kleithriasphaeridium loffrense T. speetonensis,C.parva,P. Cribroperidinium confossum C.tabulata,Exiguisphaera Ellipsoidictyum imperfectum Psaligonyaulax deflandrei, Canningia cf.reticulata(of Epelidosphaeridia spinosa Subtilisphaera perlucida, Dinopterygium cladoides, Apteodinium granulatum, Ellipsodinium rugulosum, Litosphaeridium arundum Muderongia asymmetrica Gonyaulacysta cassidata Leptodinium cancellatum Gonyaulacysta fastigiata Xiphophoridium alatum, Gonyaulacysta teicha, Nexosispinum vetusculum, A. Wallodinium anglicum, Spiniferites? dentatus, Coronifera oceanica, Xenascus ceratioides Spiniferites? pterotus Histiocysta palla, Duxbury, 1977) O. scabrosum A. perforatum C. trycherium solocispinum P. spinosum Other Dinocysts cornutum plectilis group Florentinia mantellii C. asymmetricum neistosa Cribroperidinium confossum Dinoflagellate Cysts Muderongia tabulata, arborispinum, Druggidiumjubatum S. primaevus,Kleithriasphaeridium Ophiiobolus sp. A ofDavey(1979), Lagenadinium? membranoidium, Muderongia simplex, Gochteodinia villosamultifurcata Callaiosphaeridium tricheryum C. compta,Tubotuberella apatela,C. Oligosphaeridiumabaculum T. speetonensis,Valensiella Carpodinium granulatum,P. Batioladinium varigranosum Ctenidodinium elegantulum Gardodinium trabeculosum C. validum,Exiguisphaera Nexosispinum vetusculum Nematosphaeropsis scala Gonyaulacysta fastigiata, Achomosphaera? neptuni Hystrichodinium furcatum Carpodinium granulatum Aprobolocysta neistosa, Exiguisphaera plectilis Hystrichosphaeridium Spiniferites? dentatus magna, C.?cornutum Sirmiodinium grossii Druggidinium spp., Muderongia crucis Systematophora palmula Cerbia tabulata porosispinum polymorphum tetracantha T. ciliatum I. distincta Foucheria modesta phragma speciosum Zones M. aust. V. mag. K. corr. (Ogg,J.G.,Ogg,G.,andGradstein,F.M., 2008) kostromiensis, Druggidiumdeflandrei, Muderongia australis,Nelchinopsis Kleithriasphaeridium corrugatum Prolixosphaeridium parvispinum Litosphaeridium siphoniphorum Palaeoperidinium cretaceum Palaeohystrichophora cf. Rhynchodiniopsis aptiana Odontochitina operculata Ellipsodinium rugulosum Subtilisphaera perlucida Aprobolocysta eilema Subtilisphaera terrula Muderongia staurota Discorsia nanna Valensiella magna infusorioides Zonal Markers Kleithriasphaeridium corrugatum Prolixosphaeridium parvispinum Druggidium apicopaucicum Pseudoceratium pelliferum Rhynchodiniopsis aptiana Muderongia macwhaei Nelchinopsis kostromiensis Aprobolocysta eilema Biorbifera johnewingii Subtilisphaera terrula Endoceratium turneri Muderongia staurota Valensiella magna Lithodinia stoverii Dichadogonyaulax bensonii Druggidium spp. Cerbia tabulata Tethys Other Tethyan Dinocysts Prolixosphaeridium sp. A ofMonteil Protoellipsodinium spinocristatum Apteodinium maculatumgrande Druggidium rhabdoreticulatum Batioladinium longicornutum, Cribroperidinium? tenuiceras Heslertonia heslertonensis Psaligonyaulax deflandrei Hystrichodinium furcatum (1993), D.?deflandrei Xiphophoridium alatum Ovoidinium diversum Coronifera oceanica Lithodinia stoverii P. polymorphum O. scabrosum S. primaevus P. spinosum C. tabulata C. parva Apteodinium maculatumgrande Oligosphaeridium? asterigerum Druggidium rhabdoreticulatum Batioladinium longicornutum Cribroperidinium confossum Tehamadinium dodekovae Heslertonia heslertonensis Druggidium deflandrei, Dapsilidinium warrenii Muderongia crucis P. polymorphum S. primaevus Foucheria modesta Zones Blank Blank Blank Blank Blank E1b G1 G2 E2 F2 F3 F1 Un-Zoned A. dendroacanthos,C.triangulare, euganea, C.messinae,T. conica D. obesa, A. horridus,T. veneta Pseudodictyomitra lodogaensis A. irregularis,T. pulchra, C. Radiolarians D. gracilis,R.euganeum, T. conica,X.spicularius pseudomacrocephala Pseudodictyomitra Mita spoletoensis Mita gracilis Zonal Markers C. biopodium,infundibuliforme B. helenae,P. tythopora Sphaerostylus lanceola A. umbilicata T. conica P. boesii Acroteuthis kemperi Neohibolites minor Aulacoteuthis spp. Neohibolites clava NW Europe Acroteuthis acrei [unnamed zone] Praeoxyteuthis brunsvicensis Neohibolites Neohibolites Neohibolites Neohibolites Neohibolites Neohibolites Neohibolites oxycaudatus acmonoides Acroteuthis praeultimus Oxyteuthis Oxyteuthis Oxyteuthis strombecki germanica jaculoides wollemani depressa Hibolites minumus inflexus ewaldi pugio Belemnites Duvalia emerici Duvalia lata Tethys gervaisiana Duvalia R. gryphoides,T. transversa,R. ammonia, M.varians,P. sulcata D. pyrenaica,delphinensis M. virginiae,P. paradoxa,P. A.? marticensis,T. carinata santanderensis P. verneuili P. cantabricus,P. A.? blumenbachi Lovetchenia P. tubiconcha O. rhodanica Western Europe A.? darderi C. douvillei H. almerae H. lamberti R. renevieri E. rousseli E. plicatus C. choffati E. katzeri varians sp. grypholdes, A.? blumenbachi, A. virginiae, M.varians,P. varians, O. rhodanica,T. transversa,R. marticensis, R.ammonia,P. E. plicatus,"P." cantabricus paradoxa, C.douvillei,M. A.? darderi,E.rousseli valdensis, M.eurystoma, H. luci,V. tombecki,M. Lovetchenia P. tubiconcha R. renevieri H. almerae H. lamberti T. carinata P. sulcata E. katzeri valangiensis sp. Rudists murgensis, Himeraelitessp. B. petersi,O.nicolinae, Sellaea spp.,E.plicatus Lovetchenia sp. E. lyratus Periadriatic Lovetchenia sp.,B.petersi,O. nicolinae, O.murgensis,G. costatus H. hechti,C.frankei,P. triplicata P. hannoverana Boreal Ostracodes R. gr. luermannae P. (K.)nodigera R. bekumensis P. intermedia M. harrisiana R. reticulata S. tricostata S. bilobata B. gaultina V. blanda S. hammi H. hechti,S.bilobata,P. triplicata S. praethorensis,C.frankei P. (K.)nodigera R. bekumensis P. intermedia S. tricostata B. gualtina S. hammi Ostracodes Tethyan Ostracodes C. frankei,P. triplicata,P. V. reicheli C. rectangularis P. bedoulensis rodewaldensis S. jonesiana S. extranea C. aptensis S. bilobata P. inversa H. derooi S. frankei,P. rodewaldensis P. inversa,S.bilobata, extranea, P. triplicata P. bedoulensis K. humilis H. derooi Axcidiella cruciata Clypeator grovesii trivolvis triquetra trivolvis restricta Zones Atopochara Atopochara steinhauseri lusitanicus maillardii Globator Sponsored, inpart,by: Charophytes trochiliscoides, A. trivolvistriqueta Cl. grovesiigautieri,P. adnatus, A. triquetra,T. calcitrapa,G.m. Clypeator grovesiilusitanicus, Atopochara trivolvisrestricta Atopochara trivolvis Embergerella cruciata,P. Zonal Markers H. neimongolensis paucibracteatus H. neimongolensis Globator spp. Western EuropeHorizons pouryae, L.?danilovae,P. fragilis, C. nigra,T. marsicana,C.lyrata S. genevensi,melitae, A. S. polyreme,muehlbergi A. hispanica,H.urladanisi H. lepina,D.undulata,T. M. verae,D.hauteriviana H.? paucicalcarea H. cylindrica tripolitanus M. elitzae S. hasi Calcareous Algae S. genevensis,H.?paucicalcarea polyreme, P. fragilis,C.nigra,S. H. urladanasi, A. hispanica marsicana, S.melitae, johnsoni, C.marteli,R.piailensis, A. fouryae,C.lyrata,T. P. jurassica,S.annulata, D. hauteriviana L.? danilovae H. cylindrica muehlbergi M. elitzae M. verae T.? neocomensis S.hasi M.? praturloni C.solkani Periadriatic Horizons C. somalica,lyrata,T. marsicana, C.velici H. dinarica L.? danilovae,S. H. catenaeformis H. urladanasi biokovensis C. lyrata,T. margicana,C. S.? somalica,S.melitae, S. annulata,C.solkani H. catenaeformis S. biokovensis H. dinarica S. melitae velici Stage Aptian Albian 129.97 133.9 112.0 125.0 September 2008 Valanginian Hauterivian Barremian 135 125 105 115 130 120 110 Age