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A New Fossil Species of Attagenus Latreille Foss. Rec., 23, 95–104, 2020 https://doi.org/10.5194/fr-23-95-2020 © Author(s) 2020. This work is distributed under the Creative Commons Attribution 4.0 License. A new fossil species of Attagenus Latreille (Coleoptera: Dermestidae) in Rovno and Baltic ambers, with a brief review of known fossil beetles from the Rovno amber Lagerstätte Andris Bukejs1, Jiríˇ Háva1,2, and Vitalii I. Alekseev3,4 1Institute of Life Sciences and Technologies, Daugavpils University, Vien¯ıbas 13, Daugavpils, 5401, Latvia 2Private Entomological Laboratory and Collection, Rýznerova 37, 252 62 Úneticeˇ u Prahy, Prague-West District, Czech Republic 3Shirshov Institute of Oceanology, Russian Academy of Sciences, Nahimovskiy prospekt 36, Moscow, 117997, Russia 4Kaliningrad Regional Amber Museum, Marshal Vasilevskii Square 1, Kaliningrad, 236016, Russia Correspondence: Andris Bukejs ([email protected]) Received: 1 March 2020 – Revised: 11 April 2020 – Accepted: 22 April 2020 – Published: 14 May 2020 Abstract. Based on two specimens originating from Eocene Mänd et al., 2018; Kypke and Solodovnikov, 2018). All these Rovno and Baltic ambers, Attagenus (Aethriostoma) gedani- fossil resins are succinites, and all originate from geograph- cissimus sp. nov. is described, illustrated and compared with ically proximate localities of central and eastern Europe the related fossil Mesozoic species A. (Aethriostoma) turo- (from the southern Baltic area, eastern Germany, and south- nianensis Peris et Háva, 2016. The common beetle species ern Belarus and northern Ukraine) (Perkovsky et al., 2010a; for Baltic and Rovno ambers suggests the exceptional tem- Weitschat and Wichard, 2010; Rappsilber, 2016; Bogri et al., poral and geographical closeness of palaeoecosystems that 2018). All these fossil resins are rich in bioinclusions (Spahr, produced both amber deposits. An updated checklist of 1981; Poinar, 1992; Perkovsky et al., 2003, 2010b, 2012; Coleoptera known from Rovno amber (57 species belong- Weitschat and Wichard, 2002; Perkovsky, 2016a; Alekseev, ing to 20 families) is compiled and provided with a bibli- 2017; Rappsilber and Wendel, 2019; etc.). Unfortunately, the ography of the original descriptions. The registered Rovno biodiversity of these ambers is unevenly studied: the known amber beetle assemblage contains only eight species that beetle assemblage of Baltic amber is more than 8 times larger also occur in Baltic amber (14 %), while the similarity at than the known beetle assemblage of Rovno amber and more the generic level is 56 % at the moment. The relationship be- as than 30 times larger than the known beetle assemblage of tween the Rovno and Baltic amber deposits and subjective- Bitterfeld amber (Alekseev, 2013, 2017; Bukejs et al., 2016). ness of present-day results of the beetle assemblage research Differences in volume of knowledge make data statistically are briefly discussed (urn:lsid:zoobank.org:pub:289B5A43- incorrect and unreliable and also cannot provide any argu- C57A-4B75-8A15-6E61F4AFCD81). ments concerning the discussion about the age of these am- bers. The Dermestidae (Coleoptera) contains about 1680 valid taxa worldwide (Háva, 2015) and fossil representatives of 1 Introduction this family are relatively frequent in ambers. Taxonomic and morphological diversity of Cretaceous Dermestidae suggests The problem of mutual affinity in the triad of the main Euro- an ancient origin of this group with some lineages showing pean Palaeogene fossil resins (Baltic amber, Bitterfeld amber remarkable evolutionary stasis for almost 100 million years and Rovno amber) is still under discussion in the 20th cen- (Deng et al., 2017). In total, nine extinct species of the genus tury (Hoffeins and Hoffeins, 2003; Perkovsky et al., 2007; Attagenus Latreille, 1802 from fossil resin are documented to Weitschat, 2008; Szwedo and Sontag, 2009; Sontag and date. Five species belonging to the nominative subgenus have Szadziewski, 2011; Wolfe et al., 2016; Dunlop et al., 2018; Published by Copernicus Publications on behalf of the Museum für Naturkunde Berlin. 96 A. Bukejs et al.: A new fossil species of Attagenus Latreille (Coleoptera: Dermestidae) been described previously from the Eocene Baltic amber: A. antennal club with three compact antennomeres, abdomen hoffeinsorum Háva, Prokop et Hermann, 2006; A. balticus with five visible ventrites and elytra not shortened. Háva, Prokop et Hermann, 2008; A. obesus Háva, Prokop et The specimens considered here were assigned to the sub- Hermann, 2008; A. yantarnyi Háva et Bukejs, 2012; and A. genus Aethriostoma within Attagenus based on the combi- gorskii Háva, 2014. Three species of nominative subgenus nation of the following characters: (1) disc of metaventrite have been described from the Cenomanian Burmese am- nearly twice as wide as long and (2) body strongly convex ber: A. burmiticus Cai, Hava et Huang, 2017; A. lundi Háva and widely oval (in Attagenus s. str. disc of metaventrite et Damgaard, 2017; and A. secundus Deng, Slipi´ nski,´ Ren nearly twice as long as wide, and body less convex and more et Pang, 2017. One species of the subgenus Aethriostoma narrowly obovate). Motschulsky, 1858 is known from the Turonian New Jersey amber: A. turonianensis Peris et Háva, 2016. Attagenus (Aethriostoma) gedanicissimus sp. nov. In the current paper, the second extinct species of the urn:lsid:zoobank.org:act:4CE59EC6-79B0-43D9- subgenus Aethriostoma is described and illustrated from B7DD-602EF4C3AA21 early Cenozoic amber (Rovno and Baltic). Data about the Coleoptera described from Rovno amber are summarized Figs. 1–2 and briefly discussed. Type material 2 Material and methods Holotype: 6278 (ex. coll. Jonas Damzen JDC 8531) (MAIG); The material examined is deposited in the following collec- Rovno amber; adult, sex unknown. Complete beetle is in- tions: cluded in subtriangular, transparent amber piece, with dimen- – the Museum of Amber Inclusions, University of Gdansk´ sions of 25 mm × 19 mm × 16 mm and maximum thickness (Poland, MAIG); of 6 mm. Syninclusions consist of one specimen of Nemato- cera (Diptera) and numerous small to minute organic parti- – the private collection of Anders Damgaard (Holstebro, cles. Denmark, ADC), subsequently deposited in the Zoolog- Paratype: “AlDlo 558” (ADC); Baltic amber; adult, sex ical Museum, University of Copenhagen, Denmark. unknown. Complete beetle is included in transparent amber The amber pieces were polished by hand, allowing improved piece, with dimensions of 23 mm × 21 mm and maximum views of the included specimens, and they were not subjected thickness of 5 mm. Syninclusion consists of one specimen to any supplemental fixation. of Nematocera (Diptera). Observations of the specimens were made using a Nikon SMZ® 745T stereomicroscope. The photographs were taken Type strata using a Canon 70D® camera with a macro lens (Canon MP-E Rovno amber, upper Eocene (holotype); Baltic amber, mid- 65 mm). Extended depth of field at high magnifications was dle Eocene (paratype). achieved by combining multiple images from a range of focal planes using Helicon Focus® v. 6.0.18 software. Measure- Etymology ments were taken using an ocular micrometer (expressed in millimetres). The epithet of the new species is the Latin adjective gedani- cus in the superlative and is formed from Gedanum, the Latin 3 Systematic palaeontology name of Gdansk´ where the specimen is deposited. Family Dermestidae Latreille, 1807 Differential diagnosis Subfamily Attageninae Casey, 1900 Attagenus (Aethriostoma) gedanicissimus sp. nov. differs from the single known fossil representative of the subgenus, Tribe Attagenini Casey, 1900 Attagenus (Aethriostoma) turonianensis Peris et Háva, 2016 Genus Attagenus Latreille, 1802 (USA; Late Cretaceous: Turonian; New Jersey amber) in the following characters: antennomere 11 shorter than anten- Aethriostoma Subgenus Motschulsky, 1858 nomeres 9–10 combined; pronotum strongly transverse, 3× Remarks as wide as long; abdominal sutures almost straight; and larger body size; while in A. turonianensis antennomere 11 as long The studied amber beetles show the combination of charac- as antennomeres 9–10 combined; pronotum transverse, 2:1× ters unequivocally corresponding to Attagenini within Atta- as wide as long; abdominal sutures concave; and distinctly geninae: prosternum not forming “collar”, mouthparts free, smaller body size (1.81 mm). Foss. Rec., 23, 95–104, 2020 www.foss-rec.net/23/95/2020/ A. Bukejs et al.: A new fossil species of Attagenus Latreille (Coleoptera: Dermestidae) 97 Figure 1. Attagenus (Aethriostoma) gedanicissimus sp. nov., holotype, 6278 (MAIG): (a) habitus, dorsal view; (b) habitus, ventral view; (c) habitus, right lateral view. Scale bars D 0.5 mm. www.foss-rec.net/23/95/2020/ Foss. Rec., 23, 95–104, 2020 98 A. Bukejs et al.: A new fossil species of Attagenus Latreille (Coleoptera: Dermestidae) Figure 2. Attagenus (Aethriostoma) gedanicissimus sp. nov.: (a) details of forebody, ventral view, holotype, 6278 (MAIG); (b) head and pronotum, lateral view, holotype, 6278 (MAIG); (c) habitus, dorsal view, paratype AlDlo 558 (ADC). Scale bars D 0.5 mm. Abbreviations: a1–a11 are antennomeres 1–11 respectively. Foss. Rec., 23, 95–104, 2020 www.foss-rec.net/23/95/2020/ A. Bukejs et al.: A new fossil species of Attagenus Latreille (Coleoptera: Dermestidae) 99 The new Eocene species A. (Aethriostoma) gedanicissimus bisinuated, with subtriangular process medially. Metepister- sp. nov. differs from extant representatives of the subgenus num wide, 2:6× as long as wide anteriorly. in the unicolour
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