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Gymnosperms the MESOZOIC: ERA of GYMNOSPERM DOMINANCE
Chapter 24 Gymnosperms THE MESOZOIC: ERA OF GYMNOSPERM DOMINANCE THE VASCULAR SYSTEM OF GYMNOSPERMS CYCADS GINKGO CONIFERS Pinaceae Include the Pines, Firs, and Spruces Cupressaceae Include the Junipers, Cypresses, and Redwoods Taxaceae Include the Yews, but Plum Yews Belong to Cephalotaxaceae Podocarpaceae and Araucariaceae Are Largely Southern Hemisphere Conifers THE LIFE CYCLE OF PINUS, A REPRESENTATIVE GYMNOSPERM Pollen and Ovules Are Produced in Different Kinds of Structures Pollination Replaces the Need for Free Water Fertilization Leads to Seed Formation GNETOPHYTES GYMNOSPERMS: SEEDS, POLLEN, AND WOOD THE ECOLOGICAL AND ECONOMIC IMPORTANCE OF GYMNOSPERMS The Origin of Seeds, Pollen, and Wood Seeds and Pollen Are Key Reproductive SUMMARY Innovations for Life on Land Seed Plants Have Distinctive Vegetative PLANTS, PEOPLE, AND THE Features ENVIRONMENT: The California Coast Relationships among Gymnosperms Redwood Forest 1 KEY CONCEPTS 1. The evolution of seeds, pollen, and wood freed plants from the need for water during reproduction, allowed for more effective dispersal of sperm, increased parental investment in the next generation and allowed for greater size and strength. 2. Seed plants originated in the Devonian period from a group called the progymnosperms, which possessed wood and heterospory, but reproduced by releasing spores. Currently, five lineages of seed plants survive--the flowering plants plus four groups of gymnosperms: cycads, Ginkgo, conifers, and gnetophytes. Conifers are the best known and most economically important group, including pines, firs, spruces, hemlocks, redwoods, cedars, cypress, yews, and several Southern Hemisphere genera. 3. The pine life cycle is heterosporous. Pollen strobili are small and seasonal. Each sporophyll has two microsporangia, in which microspores are formed and divide into immature male gametophytes while still retained in the microsporangia. -
Cephalotaxus
Reprinted from the Winter 1970 issue of t'he THE AMERICA HORTICULTURAL \t{AGAZIl\'E Copyright 1970 by The American Horticultural Society, Inc. Cephalotaxus Source of Harringtonine, A Promising New Anti..Cancer Alkaloid ROBERT E. PERDUE, JR.,l LLOYD A. SPETZMAN,l and RICHARD G. POWELL2 The plumyews (Cephalotaxus) are yew-like evergreen trees and shrubs. The genus includes seven species native to southeastern Asia from Japan and Korea to Taiwan and Hainan, and west through China to northeastern India. Two species are in cultivation in the United States, C. harringtoniaJ (Fig. 1 & 2) of which there are several varieties (one often listed as C. drupacea) , and C. fortunii (Fig. 3). The cultivars are shrubs up to about 20 feet in height; most have broad crowns. The linear and pointed leaves are spirally arranged or in two opposite ranks. The upper sur Fig. 1. Japanese plumyew (Cephalo face is dark shiny green with a conspicu taxus harringtonia var. drupacea), an ous mid-rib; the lower surface has a evergreen shrub about 6 ft. high, at broad silvery band on either side of the the USDA Plant Introduction Station, mid-rib. These bands are made up of Glenn Dale, Maryland. This photo conspicuous white stomata arranged in graph was made in 1955. The plant is numerous distinct lines. Leaf length is now about 7 ft. high, but the lower variable, from about one inch in varie branches have been severely pruned ties of C. harringtonia to three or four to provide material for chemical re inches in C. fortunii. The leaves are search. -
Torreya Taxifolia
photograph © Abraham Rammeloo Torreya taxifolia produces seeds in 40 Kalmthout Arboretum ABRAHAM RAMMELOO, Curator of the Kalmthout Arboretum, writes about this rare conifer that recently produced seed for the first time. Torreya is a genus of conifers that comprises four to six species that are native to North America and Asia. It is closely related to Taxus and Cephalotaxus and is easily confused with the latter. However, it is relatively easy to distinguish them apart by their leaves. Torreya has needles with, on the underside, two small edges with stomas giving it a green appearance; Cephalotaxus has different rows of stomas, and for this reason the underside is more of a white colour. It is very rare to find Torreya taxifolia in the wild; it is native to a small area in Florida and Georgia. It grows in steep limestone cliffs along the Apalachicola River. These trees come from a warm and humid climate where the temperature in winter occasionally falls below freezing. They grow mainly on north-facing slopes between Fagus grandifolia, Liriodendron tulipifera, Acer barbatum, Liquidambar styraciflua and Quercus alba. They can grow up to 15 to 20 m high. The needles are sharp and pointed and grow in a whorled pattern along the branches. They are 25 to 35 mm long and stay on the tree for three to four years. If you crush them, they give off a strong, sharp odour. The health and reproduction of the adult population of this species suffered INTERNATIONAL DENDROLOGY SOCIETY TREES Opposite Torreya taxifolia ‘Argentea’ growing at Kalmthout Arboretum in Belgium. -
Conifer Quarterly
Conifer Quarterly Vol. 24 No. 4 Fall 2007 Picea pungens ‘The Blues’ 2008 Collectors Conifer of the Year Full-size Selection Photo Credit: Courtesy of Stanley & Sons Nursery, Inc. CQ_FALL07_FINAL.qxp:CQ 10/16/07 1:45 PM Page 1 The Conifer Quarterly is the publication of the American Conifer Society Contents 6 Competitors for the Dwarf Alberta Spruce by Clark D. West 10 The Florida Torreya and the Atlanta Botanical Garden by David Ruland 16 A Journey to See Cathaya argyrophylla by William A. McNamara 19 A California Conifer Conundrum by Tim Thibault 24 Collectors Conifer of the Year 29 Paul Halladin Receives the ACS Annual Award of Merits 30 Maud Henne Receives the Marvin and Emelie Snyder Award of Merit 31 In Search of Abies nebrodensis by Daniel Luscombe 38 Watch Out for that Tree! by Bruce Appeldoorn 43 Andrew Pulte awarded 2007 ACS $1,000 Scholarship by Gerald P. Kral Conifer Society Voices 2 President’s Message 4 Editor’s Memo 8 ACS 2008 National Meeting 26 History of the American Conifer Society – Part One 34 2007 National Meeting 42 Letters to the Editor 44 Book Reviews 46 ACS Regional News Vol. 24 No. 4 CONIFER QUARTERLY 1 CQ_FALL07_FINAL.qxp:CQ 10/16/07 1:45 PM Page 2 PRESIDENT’S MESSAGE Conifer s I start this letter, we are headed into Afall. In my years of gardening, this has been the most memorable year ever. It started Quarterly with an unusually warm February and March, followed by the record freeze in Fall 2007 Volume 24, No 4 April, and we just broke a record for the number of consecutive days in triple digits. -
Pinus Serotina (Pond Pine) Is a Tree of the Southeastern Coastal Plain of United States
PinusPinus serotinaserotina pondpond pinepine by Dr. Kim D. Coder, Professor of Tree Biology & Health Care Warnell School of Forestry & Natural Resources Pinus serotina (pond pine) is a tree of the Southeastern Coastal Plain of United States. It was first described as a species in 1803. Historically this tree has been taxonomically associated with pitch pine Pinus rigida. Other scientific names for this tree have been Pinus rigida var. serotina (1868) and Pinus rigida subs. serotina (1939). The scientific name means a “pine with late opening cones.” Other common names include swamp pine, marsh pine, bay pine, flatwoods pine, and pocosin pine. Pinus serotina grows along the Coastal Plain from Southern New Jersey into central Florida and central Alabama. In Georgia, pond pine grows below the fall line. See Georgia range map figure. Pinus serotina grows in bogs, moving water swamps, flatwoods, savannahs, and low scrub areas including poorly drained wetland sites with widely fluctuating water tables. Unlike many other pines, pond pine tolerates and reproduces in areas with high organic matter contents. Site drainage shifts species regen- eration toward loblolly pine. Pond pine is fire tolerant and sprouts after fire damage, but is disfigured by hot fires. Pinus serotina is a medium sized tree 40-65 feet tall (maximum of 90 feet). It grows to a diam- eter of 1-2.2 feet (maximum of 3 feet). Crown form is irregular, ragged, and thin, with many tufts of needles sprouting on stems. Crown shape is usually flat-topped with many stubby and gnarled branches among longer branches. Cones held for many years lead to the appearance of a cone-crowded crown. -
Plant Classification
Plant Classification Vascular plants are a group that has a system Non-Vascular plants are low growing plants of tubes (roots, stems and leaves) to help that get materials directly from their them transport materials throughout the surroundings. They have small root-like plant. Tubes called xylem move water from structures called rhizoids which help them the roots to the stems and leaves. Tubes adhere to their substrate. They undergo called phloem move food from the leaves asexual reproduction through vegetative (where sugar is made during propagation and sexual reproduction using photosynthesis) to the rest of the plant’s spores. Examples include bryophytes like cells. Vascular plants reproduce asexually hornworts, liverworts, and mosses. through spores and vegetative propagation (small part of the plant breaks off and forms a new plant) and sexually through pollen (sperm) and ovules (eggs). A gymnosperm is a vascular plant whose An angiosperm is a vascular plant whose seeds are not enclosed in an ovule or fruit. mature seeds are enclosed in a fruit or The name means “naked seed” and the ovule. They are flowering plants that group typically refers to conifers that bear reproduce using seeds and are either male and female cones, have needle-like “perfect” and contain both male and female leaves and are evergreen (leaves stay green reproductive structures or “imperfect” and year round and do not drop their leaves contain only male or female structures. during the fall and winter. Examples include Angiosperm trees are also called hardwoods pine trees, ginkgos and cycads. and they have broad leaves that change color and drop during the fall and winter. -
PHYLOGENETIC RELATIONSHIPS of TORREYA (TAXACEAE) INFERRED from SEQUENCES of NUCLEAR RIBOSOMAL DNA ITS REGION Author(S): Jianhua Li, Charles C
PHYLOGENETIC RELATIONSHIPS OF TORREYA (TAXACEAE) INFERRED FROM SEQUENCES OF NUCLEAR RIBOSOMAL DNA ITS REGION Author(s): Jianhua Li, Charles C. Davis, Michael J. Donoghue, Susan Kelley and Peter Del Tredici Source: Harvard Papers in Botany, Vol. 6, No. 1 (July 2001), pp. 275-281 Published by: Harvard University Herbaria Stable URL: http://www.jstor.org/stable/41761652 Accessed: 14-06-2016 15:35 UTC REFERENCES Linked references are available on JSTOR for this article: http://www.jstor.org/stable/41761652?seq=1&cid=pdf-reference#references_tab_contents You may need to log in to JSTOR to access the linked references. Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at http://about.jstor.org/terms JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. Harvard University Herbaria is collaborating with JSTOR to digitize, preserve and extend access to Harvard Papers in Botany This content downloaded from 128.103.224.4 on Tue, 14 Jun 2016 15:35:14 UTC All use subject to http://about.jstor.org/terms PHYLOGENETIC RELATIONSHIPS OF TORREYA (TAXACEAE) INFERRED FROM SEQUENCES OF NUCLEAR RIBOSOMAL DNA ITS REGION Jianhua Li,1 Charles C. Davis,2 Michael J. Donoghue,3 Susan Kelley,1 And Peter Del Tredici1 Abstract. Torreya, composed of five to seven species, is distributed disjunctly in eastern Asia and the eastern and western United States. -
B.Sc Botany (Sub.) I Group: a General Characters of Gymnosperm
1 B.Sc Botany (Sub.) I Group: A General Characters of Gymnosperm Gymnosperms are a small group of plants comprising only 70 genera and 725 living species. The word Gymnosperm was used by the Greek botanists Theophrastus in 300 B.C. for the plants with unprotected seeds. Gymnosperms are naked seeded plants. The gymnosperms are characterized by the following features: i. It shows the distinct alternation of generations. ii. Sporophytic generation is the dormant phase of the life cycle. The main plants are sporophyte and the gametophytes are dependent on it throughout. iii. The sporophytic plants are usually tall, woody, perennial trees or shrubs and differentiated into root, stem and leaves. iv. Root- Root is usually well developed tap root system. The stele in root is diarch or polyarch. v. Stem- stems are usually branched but unbranched in Cycas. vi. The leaves may be compound as in Cycas or simple as in Pinus. Internal structure: i. The vascular bundles in stem are arranged in a ring. The bundles are conjoint, collateral and open. ii. The secondary growth is effected by a cambial layer which produces secondary xylem and secondary phloem on the inner and outer side respectively. iii. The secondary wood forming distinct annual ring. Spring wood: It is made up of layer tracheids with the walls and layer lumen. Autumn wood: It is made up of compactly arranged smaller tracheids with thick walls and smaller lumen. iv. The wood may be manoxylic or pycnoxylic and may be monoxylic or polyxylic. Dr. Sanjeev Kumar Vidyarthi, dept. of Botany, Dr. L.K.V.D. -
A Phylogenetic Study
92 Agnes Robertson. THE TAXOIDE/E; A PHYLOGENETIC STUDY. BY AGNES ROBERTSON, D.SC. Quain Student in Botany ; University CoUege, London. [WITH PLATE I.] I.—INTRODUCTION. ARLY in 1904, at the suggestion of Professor F. W. Oliver, I E began to make a study of the Taxoidex. At that time the the minute structure and development of the reproductive organs of the group were not known with any completeness except in Taxus, but in the course of the last three years several papers have appeared on the subject, culminating quite recently in Professor Ai A. Lawson's interesting study of Cephalotaxus, (see Literature Citations, 16, 17, 18, 19, 26). We are now acquainted with at least the main outlines of the reproductive processes of these plants, and a considerable mass of information has accumulated as to their floral morphology, anatomy, etc., so that it seems as if the moment has come to enquire what bearing this knowledge may have upon our ideas as to the phylogeny of the group. I have made some attempt to do this in the following pages. I should like to take this opportunity of thanking Professor F. W. Oliver for the help and encouragement which he has constantly given me in the study of the group. Before entering on any general discussion I have a correction to make and a few observations to record, 11.—OBSERVATIONS. (a). Torreya californica. Torr. I wish to begin these notes by correcting an error into which I fell in a paper on Torreya californica (17) published in this journal in 1904. -
Fusarium Torreyae (Sp
HOST RANGE AND BIOLOGY OF FUSARIUM TORREYAE (SP. NOV), CAUSAL AGENT OF CANKER DISEASE OF FLORIDA TORREYA (TORREYA TAXIFOLIA ARN.) By AARON J. TRULOCK A THESIS PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORIDA IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE UNIVERSITY OF FLORIDA 2012 1 © 2012 Aaron J. Trulock 2 To my wife, for her support, patience, and dedication 3 ACKNOWLEDGMENTS I would like to thank my chair, Jason Smith, and committee members, Jenny Cruse-Sanders and Patrick Minogue, for their guidance, encouragement, and boundless knowledge, which has helped me succeed in my graduate career. I would also like to thank the Forest Pathology lab for aiding and encouraging me in both my studies and research. Research is not an individual effort; it’s a team sport. Without wonderful teammates it would never happen. Finally, I would like to that the U.S. Forest Service for their financial backing, as well as, UF/IFAS College of Agriculture and Life Science for their matching funds. 4 TABLE OF CONTENTS page ACKNOWLEDGMENTS .................................................................................................. 4 LIST OF TABLES ............................................................................................................ 6 LIST OF FIGURES .......................................................................................................... 7 ABSTRACT ..................................................................................................................... 8 -
A Global Analysis of the Distribution and Conservation Status Of
Journal of Biogeography (J. Biogeogr.) (2015) 42, 809–820 SYNTHESIS Fighting their last stand? A global analysis of the distribution and conservation status of gymnosperms Yann Fragniere1,Sebastien Betrisey2,3,Leonard Cardinaux1, Markus Stoffel4,5 and Gregor Kozlowski1,2* 1Natural History Museum Fribourg, CH-1700 ABSTRACT Fribourg, Switzerland, 2Department of Biology Aim Gymnosperms are often described as a marginal and threatened group, and Botanic Garden, University of Fribourg, members of which tend to be out-competed by angiosperms and which therefore CH-1700 Fribourg, Switzerland, 3Conservation Biogeography, Department of preferentially persist at higher latitudes and elevations. The aim of our synthesis Geosciences, University of Fribourg, CH-1700 was to test these statements by investigating the global latitudinal and elevational Fribourg, Switzerland, 4Dendrolab.ch, distribution of gymnosperms, as well as their conservation status, using all extant Institute of Geological Sciences, University of gymnosperm groups (cycads, gnetophytes, ginkgophytes and conifers). 5 Bern, CH-3012 Bern, Switzerland, Institute Location Worldwide. for Environmental Sciences, Climatic Change and Climate Impacts, University of Geneva, Methods We developed a database of 1014 species of gymnosperms containing CH-1227 Carouge, Switzerland latitudinal and elevational distribution data, as well as their global conservation status, as described in the literature. The 1014 species comprised 305 cycads, 101 gnetophytes, the only living representative of ginkgophytes, and 607 conifers. Generalized additive models, frequency histograms, kernel density estimations and distribution maps based on Takhtajan’s floristic regions were used. Results Although the diversity of gymnosperms decreases at equatorial lati- tudes, approximately 50% of the extant species occur primarily between the tropics. More than 43% of gymnosperms can occur at very low elevations (≤ 200 m a.s.l.). -
Name That Gymnosperm
A B C D This tree is found frequently This evergreen is found throughout This species is found at drier and This Wyoming tree is a remnant of throughout the eastern half of the state. The species is known for lower elevations compared to some the last ice age and is found exclu- Wyoming ranging from the Black often having serotinous cones that of the other trees pictured. This tree sively in the Black Hills of Wyoming Hills to the Laramie Mountains and only open to release seeds when the shares its name with the town in and South Dakota. It grows at higher the Bighorns. The tree is known cone is heated. Once the limbs are central Wyoming. elevations along riparian or wet areas for its ability to withstand the heat removed, this tree makes an excel- in its native habitat. of wildfires because of its thick, lent pole for teepees because of its reddish-colored bark. long, slender trunk. NAME THAT GYMNOSPERM Wyoming is host to many conifer (gymnosperm or “naked in hot, dry, and low elevations while others are found in cold and seed” plants including conifer, cycads, and ginkos) tree species. high elevations. Cones are an excellent tool that can be used for The cones in this quiz and their parent trees are found at different identification of evergreens. Match the tree to the photo. Good geographical locations across the Cowboy State. Some are found luck and keep an eye out for these trees this year! E F G H Known for having extremely flexible This species is probably better known This tree is found in most high- This tree grows at high elevations limbs, this tree is often found brav- as an ornamental in Wyoming but is elevation forests of Wyoming.