The Carboniferous Floating Forest — an Extinct Pre-Flood Ecosystem
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Contents Part III. the LAST FIFTY THOUSAND YEARS
STATE OF MICHIGAN Insects ................................................................... 60 DEPARTMENT OF CONSERVATION Worms And Others ................................................ 61 P. J. Hoffmaster, Director The Pleasant Peninsulas .............................................62 Man and his Towns ......................................................69 THEY NEED NOT VANISH A DISCUSSION OF THE NATURAL RESOURCES In Conclusion .................................................................71 OF MICHIGAN Part III. THE LAST FIFTY THOUSAND YEARS "THE GOOD EARTH" Edited by HELEN M. MARTIN ir, sunlight, water, and soil are essential for the Acontinuance of life—plant, animal, and human life, from contributions by on the earth. Of these four basic requirements, the soil Shirley W. Allen, Geo. C. S. Benson, University of is most directly subject to the care and management of Michigan man. However, the soil has frequently been the object of man's most careless use and abuse. It is, therefore, Stannard B. Bergquist, L. R. Schoenmann, H. C. most fitting that the eminent soil scientist, A. F. Beeskaw, J. H. Kraemer, W. F. Morofsky, J. A. Porter, E. Gustafson, should begin his book on soils and soil C. Sackrider, Michigan State College management with: G. S. Mclntire, H. M. Martin, O. F. Poindexter, C. F. "During his existence upon the earth, man has depended upon Welch, Department of Conservation the soil, either directly or indirectly, for the production of the materials used by him for food and clothing and, in part, for the M. G. Adams, Stream Control Commission production of those used for fuel and shelter as well. Grains, Frank DuMond, Public Museum, Grand Rapids fruits, and vegetables that serve him as food grow directly on the soil. Cotton and flax yield materials that are made into Lynn Heatley, High School, Midland. -
RI Equisetopsida and Lycopodiopsida.Indd
IIntroductionntroduction byby FFrancisrancis UnderwoodUnderwood Rhode Island Equisetopsida, Lycopodiopsida and Isoetopsida Special Th anks to the following for giving permission for the use their images. Robbin Moran New York Botanical Garden George Yatskievych and Ann Larson Missouri Botanical Garden Jan De Laet, plantsystematics.org Th is pdf is a companion publication to Rhode Island Equisetopsida, Lycopodiopsida & Isoetopsida at among-ri-wildfl owers.org Th e Elfi n Press 2016 Introduction Formerly known as fern allies, Horsetails, Club-mosses, Fir-mosses, Spike-mosses and Quillworts are plants that have an alternate generation life-cycle similar to ferns, having both sporophyte and gametophyte stages. Equisetopsida Horsetails date from the Devonian period (416 to 359 million years ago) in earth’s history where they were trees up to 110 feet in height and helped to form the coal deposits of the Carboniferous period. Only one genus has survived to modern times (Equisetum). Horsetails Horsetails (Equisetum) have jointed stems with whorls of thin narrow leaves. In the sporophyte stage, they have a sterile and fertile form. Th ey produce only one type of spore. While the gametophytes produced from the spores appear to be plentiful, the successful reproduction of the sporophyte form is low with most Horsetails reproducing vegetatively. Lycopodiopsida Lycopodiopsida includes the clubmosses (Dendrolycopodium, Diphasiastrum, Lycopodiella, Lycopodium , Spinulum) and Fir-mosses (Huperzia) Clubmosses Clubmosses are evergreen plants that produce only microspores that develop into a gametophyte capable of producing both sperm and egg cells. Club-mosses can produce the spores either in leaf axils or at the top of their stems. Th e spore capsules form in a cone-like structures (strobili) at the top of the plants. -
Gene Expression Data Support the Hypothesis That Isoetes Rootlets Are True Roots and Not Modifed Leaves Alexander J
www.nature.com/scientificreports OPEN Gene expression data support the hypothesis that Isoetes rootlets are true roots and not modifed leaves Alexander J. Hetherington1,2, David M. Emms1, Steven Kelly1 & Liam Dolan1,3* Rhizomorphic lycopsids are the land plant group that includes the frst giant trees to grow on Earth and extant species in the genus Isoetes. Two mutually exclusive hypotheses account for the evolution of terminal rooting axes called rootlets among the rhizomorphic lycopsids. One hypothesis states that rootlets are true roots, like roots in other lycopsids. The other states that rootlets are modifed leaves. Here we test predictions of each hypothesis by investigating gene expression in the leaves and rootlets of Isoetes echinospora. We assembled the de novo transcriptome of axenically cultured I. echinospora. Gene expression signatures of I. echinospora rootlets and leaves were diferent. Furthermore, gene expression signatures of I. echinospora rootlets were similar to gene expression signatures of true roots of Selaginella moellendorfi and Arabidopsis thaliana. RSL genes which positively regulate cell diferentiation in roots were either exclusively or preferentially expressed in the I. echinospora rootlets, S. moellendorfi roots and A. thaliana roots compared to the leaves of each respective species. Taken together, gene expression data from the de-novo transcriptome of I. echinospora are consistent with the hypothesis that Isoetes rootlets are true roots and not modifed leaves. Te frst giant (> 50 m) trees to grow on Earth, the arborescent clubmosses, were tethered to the ground by rooting structures termed stigmarian systems whose homology has been debated for more than 150 years1–9. Stigmarian rooting systems consisted of two components, a central axis (rhizomorph) on which developed large numbers of fne axes (rootlets). -
Pennsylvanian Exposures in the White Breast Recreation Area, Marion County, Iowa ______
Iowa Geological & Water Survey - GSI PENNSYLVANIAN EXPOSURES IN THE WHITE BREAST RECREATION AREA, MARION COUNTY, IOWA ___________________________________________________ John P. Pope, Adrian E. Goettemoeller, and Raymond R. Anderson Geological Society of Iowa Sponsored by the Department of Natural Resources Iowa Geological and Water Survey ______________________________________ April 20, 2013 Guidebook 91 i Guidebook 91 Cover photo shows the field trip stop area, Pennsylvanian exposure on the south-facing shore of the White Breast Recreation area at Lake Red Rock. ii Iowa Geological & Water Survey - GSI PENNSYLVANIAN EXPOSURES IN THE WHITE BREAST RECREATION AREA, MARION COUNTY, IOWA John P. Pope Northwest Missouri State University Dept. Geography/Geology 800 University Drive Maryville, MO 64468-6001 [email protected] Adrian E. Goettemoeller 712 York Court Plattsmouth, NE 68048 [email protected] Raymond R. Anderson Iowa Dept. Natural Resources Geological Survey Bureau Iowa City, IA 52242-1319 [email protected] with contributions by Greg A. Ludvigson Kansas Geological Survey 1930 Constant Avenue Lawrence, KS 66047-3724 [email protected] April 20, 2013 Geological Society of Iowa Guidebook 91 Geological Society of Iowa Sponsored by the Department of Natural Resources Iowa Geological and Water Survey Additional Copies of this Guidebook or other GSI Guidebooks May be Ordered from the IGWS Publication page at https://programs.iowadnr.gov/igspubs/listPubs.aspx iii Guidebook 91 iv Iowa Geological & Water Survey - GSI TABLE OF CONTENTS Pennsylvanian Exposures in the White Breast Recreation Area, Marion County, Iowa Introduction to the Field Trip Raymond R. Anderson ................................................................................................................ 1 References ........................................................................................................................... 1 Overview of Lake Red Rock and the White Breast Recreation Area Raymond R. -
Chians in Nova Scotia,. by J. \V. DAWSON, LL.D., F.R.S
440 Dawson-New Oarboniferous Batmchians of Nova Scotia. ART. XLVIII.-On a Recent Di~covery of Oarbo?Jifero1.l.s Batra chians in Nova Scotia,. by J. \V. DAWSON, LL.D., F.R.S. 1. General Remarks. THE erect Sigillarire enclosed in the f!andstone overlying coal-group 15 of Section XV, Division 4 of the SOIl.th Joggins section, are perhaps the most remarkable repositories ever dis covered of the remains of Paleozoic land animals. As I have shown in discussing their character in my memoirs on the South Joggins Coal FOl'mation,* and my "Acadian Geology," some of these trees became em bedded in sandy deposits, and being rendered hollow by decay of their inner bark and the crumbling of their woody axes, remained for a long time as open holes or pits, gradually filling with vegetable debris and the wash of rains and land floods. They thus became places of habitation for land snails and millepedes, and pit-falls into which the smaller batrachians, prowling for prey among the undergrowth of the coal forest, fell and were unable to extri cate themselves. In this way the successive layers of deposit became stored with skeletons of batrachians which they have retained in an admirable st.ate of preservation. Onlrone sandstone at the Joggins is known to contain these reptiliferous trees, though erect Sigillarire are known at. more than sixty different levels, and many of these erect stumps have been broken up in the hope of making such discoveries. In the past summer, however, shells of Pupa vetustu were found bv Mr. -
The Carboniferous Evolution of Nova Scotia
Downloaded from http://sp.lyellcollection.org/ by guest on September 27, 2021 The Carboniferous evolution of Nova Scotia J. H. CALDER Nova Scotia Department of Natural Resources, PO Box 698, Halifax, Nova Scotia, Canada B3J 2T9 Abstract: Nova Scotia during the Carboniferous lay at the heart of palaeoequatorial Euramerica in a broadly intermontane palaeoequatorial setting, the Maritimes-West-European province; to the west rose the orographic barrier imposed by the Appalachian Mountains, and to the south and east the Mauritanide-Hercynide belt. The geological affinity of Nova Scotia to Europe, reflected in elements of the Carboniferous flora and fauna, was mirrored in the evolution of geological thought even before the epochal visits of Sir Charles Lyell. The Maritimes Basin of eastern Canada, born of the Acadian-Caledonian orogeny that witnessed the suture of Iapetus in the Devonian, and shaped thereafter by the inexorable closing of Gondwana and Laurasia, comprises a near complete stratal sequence as great as 12 km thick which spans the Middle Devonian to the Lower Permian. Across the southern Maritimes Basin, in northern Nova Scotia, deep depocentres developed en echelon adjacent to a transform platelet boundary between terranes of Avalon and Gondwanan affinity. The subsequent history of the basins can be summarized as distension and rifting attended by bimodal volcanism waning through the Dinantian, with marked transpression in the Namurian and subsequent persistence of transcurrent movement linking Variscan deformation with Mauritainide-Appalachian convergence and Alleghenian thrusting. This Mid- Carboniferous event is pivotal in the Carboniferous evolution of Nova Scotia. Rapid subsidence adjacent to transcurrent faults in the early Westphalian was succeeded by thermal sag in the later Westphalian and ultimately by basin inversion and unroofing after the early Permian as equatorial Pangaea finally assembled and subsequently rifted again in the Triassic. -
Retallack 2021 Coal Balls
Palaeogeography, Palaeoclimatology, Palaeoecology 564 (2021) 110185 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Modern analogs reveal the origin of Carboniferous coal balls Gregory Retallack * Department of Earth Science, University of Oregon, Eugene, Oregon 97403-1272, USA ARTICLE INFO ABSTRACT Keywords: Coal balls are calcareous peats with cellular permineralization invaluable for understanding the anatomy of Coal ball Pennsylvanian and Permian fossil plants. Two distinct kinds of coal balls are here recognized in both Holocene Histosol and Pennsylvanian calcareous Histosols. Respirogenic calcite coal balls have arrays of calcite δ18O and δ13C like Carbon isotopes those of desert soil calcic horizons reflecting isotopic composition of CO2 gas from an aerobic microbiome. Permineralization Methanogenic calcite coal balls in contrast have invariant δ18O for a range of δ13C, and formed with anaerobic microbiomes in soil solutions with bicarbonate formed by methane oxidation and sugar fermentation. Respiro genic coal balls are described from Holocene peats in Eight Mile Creek South Australia, and noted from Carboniferous coals near Penistone, Yorkshire. Methanogenic coal balls are described from Carboniferous coals at Berryville (Illinois) and Steubenville (Ohio), Paleocene lignites of Sutton (Alaska), Eocene lignites of Axel Heiberg Island (Nunavut), Pleistocene peats of Konya (Turkey), and Holocene peats of Gramigne di Bando (Italy). Soils and paleosols with coal balls are neither common nor extinct, but were formed by two distinct soil microbiomes. 1. Introduction and Royer, 2019). Although best known from Euramerican coal mea sures of Pennsylvanian age (Greb et al., 1999; Raymond et al., 2012, Coal balls were best defined by Seward (1895, p. -
Inferring the Evolutionary Reduction of Corm Lobation in Isoëtes Using Bayesian Model- Averaged Ancestral State Reconstruction
UC Berkeley UC Berkeley Previously Published Works Title Inferring the evolutionary reduction of corm lobation in Isoëtes using Bayesian model- averaged ancestral state reconstruction. Permalink https://escholarship.org/uc/item/39h708p5 Journal American journal of botany, 105(2) ISSN 0002-9122 Authors Freund, Forrest D Freyman, William A Rothfels, Carl J Publication Date 2018-02-01 DOI 10.1002/ajb2.1024 Peer reviewed eScholarship.org Powered by the California Digital Library University of California RESEARCH ARTICLE BRIEF COMMUNICATION Inferring the evolutionary reduction of corm lobation in Isoëtes using Bayesian model- averaged ancestral state reconstruction Forrest D. Freund1,2, William A. Freyman1,2, and Carl J. Rothfels1 Manuscript received 26 October 2017; revision accepted 2 January PREMISE OF THE STUDY: Inferring the evolution of characters in Isoëtes has been problematic, 2018. as these plants are morphologically conservative and yet highly variable and homoplasious 1 Department of Integrative Biology, University of California, within that conserved base morphology. However, molecular phylogenies have given us a Berkeley, Berkeley, CA 94720-3140, USA valuable tool for testing hypotheses of character evolution within the genus, such as the 2 Authors for correspondence (e-mail: [email protected], hypothesis of ongoing morphological reductions. [email protected]) Citation: Freund, F. D., W. A. Freyman, and C. J. Rothfels. 2018. METHODS: We examined the reduction in lobe number on the underground trunk, or corm, by Inferring the evolutionary reduction of corm lobation in Isoëtes using combining the most recent molecular phylogeny with morphological descriptions gathered Bayesian model- averaged ancestral state reconstruction. American from the literature and observations of living specimens. -
The Joggins Cliffs of Nova Scotia: B2 the Joggins Cliffs of Nova Scotia: Lyell & Co's "Coal Age Galapagos" J.H
GAC-MAC-CSPG-CSSS Pre-conference Field Trips A1 Contamination in the South Mountain Batholith and Port Mouton Pluton, southern Nova Scotia HALIFAX Building Bridges—across science, through time, around2005 the world D. Barrie Clarke and Saskia Erdmann A2 Salt tectonics and sedimentation in western Cape Breton Island, Nova Scotia Ian Davison and Chris Jauer A3 Glaciation and landscapes of the Halifax region, Nova Scotia Ralph Stea and John Gosse A4 Structural geology and vein arrays of lode gold deposits, Meguma terrane, Nova Scotia Rick Horne A5 Facies heterogeneity in lacustrine basins: the transtensional Moncton Basin (Mississippian) and extensional Fundy Basin (Triassic-Jurassic), New Brunswick and Nova Scotia David Keighley and David E. Brown A6 Geological setting of intrusion-related gold mineralization in southwestern New Brunswick Kathleen Thorne, Malcolm McLeod, Les Fyffe, and David Lentz A7 The Triassic-Jurassic faunal and floral transition in the Fundy Basin, Nova Scotia Paul Olsen, Jessica Whiteside, and Tim Fedak Post-conference Field Trips B1 Accretion of peri-Gondwanan terranes, northern mainland Nova Scotia Field Trip B2 and southern New Brunswick Sandra Barr, Susan Johnson, Brendan Murphy, Georgia Pe-Piper, David Piper, and Chris White The Joggins Cliffs of Nova Scotia: B2 The Joggins Cliffs of Nova Scotia: Lyell & Co's "Coal Age Galapagos" J.H. Calder, M.R. Gibling, and M.C. Rygel Lyell & Co's "Coal Age Galapagos” B3 Geology and volcanology of the Jurassic North Mountain Basalt, southern Nova Scotia Dan Kontak, Jarda Dostal, -
U.S. GEOLOGICAL SURVEY BULLETIN 21 Cover
rf Predictive Stratigraphic Analysis- - Concept and Application u.s. GEOLOGICAL SURVEY BULLETIN 21 Cover. Calcic paleo-Vertisol underlying the resistant transgressive marine limestone Little Stone Gap Member of the Hinton Formation (Upper Mississippian) in southwestern West Virginia. This paleosol is indicative of a relatively dry climate when evapotranspira- tion exceeded rainfall for more than 6 months out of the year. The light-gray color at the level of the photograph scale (center) is the result of gleying (bleaching) after burial. A calcified root system, located in the proximity of the scale, branches downward and sug gests a well-developed root system for a plant whose stem may have been up to 15 centi meters in diameter. Numerous mineralized fossil roots at this level indicate that land plants were very well adapted to seasonally dry conditions in nonwaterlogged environ ments by Late Mississippian time. Cross-cutting fractures, known as mukkara structures and caused by seasonal expansion (wet) and contraction (dry), are visible throughout the outcrop beneath the resistant limestone layer except where interrupted or destroyed by paleoroot systems. Predictive Stratigraphic Analysis Concept and Application Edited by C. Blaine Cecil and N. Terence Edgar U.S. GEOLOGICAL SURVEY BULLETIN 2110 A collection of extended abstracts of papers presented at two workshops on the title subject UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1994 U.S. DEPARTMENT OF THE INTERIOR BRUCE BABBITT, Secretary U.S. GEOLOGICAL SURVEY GORDON P. EATON, Director For sale by U.S. Geological Survey, Information Services Box 25286, Federal Center, Denver, CO 80225 Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply endorsement by the U.S. -
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Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Pangea B and the Late Paleozoic Ice Age ⁎ D.V. Kenta,b, ,G.Muttonic a Earth and Planetary Sciences, Rutgers University, Piscataway, NJ 08854, USA b Lamont-Doherty Earth Observatory of Columbia University, Palisades, NY 10964, USA c Dipartimento di Scienze della Terra 'Ardito Desio', Università degli Studi di Milano, via Mangiagalli 34, I-20133 Milan, Italy ARTICLE INFO ABSTRACT Editor: Thomas Algeo The Late Paleozoic Ice Age (LPIA) was the penultimate major glaciation of the Phanerozoic. Published compi- Keywords: lations indicate it occurred in two main phases, one centered in the Late Carboniferous (~315 Ma) and the other Late Paleozoic Ice Age in the Early Permian (~295 Ma), before waning over the rest of the Early Permian and into the Middle Permian Pangea A (~290 Ma to 275 Ma), and culminating with the final demise of Alpine-style ice sheets in eastern Australia in the Pangea B Late Permian (~260 to 255 Ma). Recent global climate modeling has drawn attention to silicate weathering CO2 Greater Variscan orogen consumption of an initially high Greater Variscan edifice residing within a static Pangea A configuration as the Equatorial humid belt leading cause of reduction of atmospheric CO2 concentrations below glaciation thresholds. Here we show that Silicate weathering CO2 consumption the best available and least-biased paleomagnetic reference poles place the collision between Laurasia and Organic carbon burial Gondwana that produced the Greater Variscan orogen in a more dynamic position within a Pangea B config- uration that had about 30% more continental area in the prime equatorial humid belt for weathering and which drifted northward into the tropical arid belt as it transformed to Pangea A by the Late Permian. -
81 Vascular Plant Diversity
f 80 CHAPTER 4 EVOLUTION AND DIVERSITY OF VASCULAR PLANTS UNIT II EVOLUTION AND DIVERSITY OF PLANTS 81 LYCOPODIOPHYTA Gleicheniales Polypodiales LYCOPODIOPSIDA Dipteridaceae (2/Il) Aspleniaceae (1—10/700+) Lycopodiaceae (5/300) Gleicheniaceae (6/125) Blechnaceae (9/200) ISOETOPSIDA Matoniaceae (2/4) Davalliaceae (4—5/65) Isoetaceae (1/200) Schizaeales Dennstaedtiaceae (11/170) Selaginellaceae (1/700) Anemiaceae (1/100+) Dryopteridaceae (40—45/1700) EUPHYLLOPHYTA Lygodiaceae (1/25) Lindsaeaceae (8/200) MONILOPHYTA Schizaeaceae (2/30) Lomariopsidaceae (4/70) EQifiSETOPSIDA Salviniales Oleandraceae (1/40) Equisetaceae (1/15) Marsileaceae (3/75) Onocleaceae (4/5) PSILOTOPSIDA Salviniaceae (2/16) Polypodiaceae (56/1200) Ophioglossaceae (4/55—80) Cyatheales Pteridaceae (50/950) Psilotaceae (2/17) Cibotiaceae (1/11) Saccolomataceae (1/12) MARATTIOPSIDA Culcitaceae (1/2) Tectariaceae (3—15/230) Marattiaceae (6/80) Cyatheaceae (4/600+) Thelypteridaceae (5—30/950) POLYPODIOPSIDA Dicksoniaceae (3/30) Woodsiaceae (15/700) Osmundales Loxomataceae (2/2) central vascular cylinder Osmundaceae (3/20) Metaxyaceae (1/2) SPERMATOPHYTA (See Chapter 5) Hymenophyllales Plagiogyriaceae (1/15) FIGURE 4.9 Anatomy of the root, an apomorphy of the vascular plants. A. Root whole mount. B. Root longitudinal-section. C. Whole Hymenophyllaceae (9/600) Thyrsopteridaceae (1/1) root cross-section. D. Close-up of central vascular cylinder, showing tissues. TABLE 4.1 Taxonomic groups of Tracheophyta, vascular plants (minus those of Spermatophyta, seed plants). Classes, orders, and family names after Smith et al. (2006). Higher groups (traditionally treated as phyla) after Cantino et al. (2007). Families in bold are described in found today in the Selaginellaceae of the lycophytes and all the pericycle or endodermis. Lateral roots penetrate the tis detail.