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Coleoptera, Staphylinidae) of Myanmar Part 1: a Checklist of Species

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Coleoptera, Staphylinidae) of Myanmar Part 1: a Checklist of Species Bull. Natl. Mus. Nat. Sci., Ser. A, 46(3), pp. 129–140, August 21, 2020 Inventory Studies on the Subfamily Pselaphinae (Coleoptera, Staphylinidae) of Myanmar Part 1: A Checklist of Species Shûhei Nomura1 and Mu Mu Aung2 1 Department of Zoology, National Museum of Nature and Science, 4–1–1 Amakubo, Tsukuba, Ibaraki 305–0005, Japan E-mail: [email protected] 2 Forest Research Institute, Forest Department, Ministry of Natural Resources and Environmental Conservation, Yezin, Nay Pyi Taw, Myanmar E-mail: [email protected] (Received 24 April 2020; accepted 24 June 2020) Abstract The pselaphine fauna of Myanmar is reviewed. One hundred and twenty-one species of the subfamily Pselaphinae and their distributions are listed. Key words : Pselaphinae, Staphylinidae, inventory, taxonomy, check list, Myanmar. Maya (Goniaceritae) are still valid. 1) A historical review of the pselaphine fauna L. W. Schaufuss (1877) described 29 pselaph- of Myanmar ine species from Bangkok, Thailand. Blattný The pselaphine fauna of Myanmar has been (1925) recorded some of the Schaufuss’s species studied since nineteenth century. Motschulsky also from Burma on the basis of Helfer’s Collec- (1851) described 53 pselaphine species from tion. L. W. Schaufuss (1882, etc.) and A. Raffray “Ind. Or.” abbreviated from “Indes orientale”, (1883, etc.) described some pselaphine species which is known to indicate Tenasserim, South from Burma (＝Myanmar). However, the detail Myanmar. This work was done on the basis of information of the type localities is unknown. Helfer’s Collection originally preserved in the In Besuchet (1986), four genera defined by Natural History Museum, Prague, Czech (NMP). Blattný (1925) and one genus by Motschulsky These pselaphine specimens were collected at (1851) were synonymized as follows: Raffrayella Tenasserim in the southern part of Myammer. Blattný, 1925 was synonymized with Batraxis More than 70 years later, Blattný (1925) Reitter, 1881; Herminiella Blattný, 1925 with described 8 new genera, 41 new species on the Odontalgus Raffray, 1877; Indiella Blattný, 1925 basis of the same collection. From the Blattný’s with Sognorus Reitter, 1881; Glyptosoma taxa, a new species of a new genus, Microph- Motschulsky, 1851 with Tmesiphorus Le Conte, thartus luridus was excluded from pselaphines, 1849; and Novissimus Blattný, 1925 with Tyrus because the new genus was synonymous with the Aubé, 1833. genus Edaphus of the subfamily Euaesthetinae. Besuchet (1999) suggested that three species Additionally, the other five new genera were syn- of the genus Metaxis Motschulsky, 1851 onymized with the known genera of Pselaphinae. described from Tenasserim, Myanmar as “Ind. Two genera, Philotrimium (Euplectitae) and Or.” should be combined with the genus Tyrus. Novissimus helferi Blattný, 1925 was synony- © 2020 National Museum of Nature and Science mized with Tyrus corpulentus (Motschulsky, 130 Shûhei Nomura and Mu Mu Aung Pselaphine Species from Myanmar 1 131 1851) described as a species of Metaxis by the Figs. 1–4 are deposited in the following muse- same paper. Besuchet (2008) reviewed some ums, in which the abbreviations are shown in genera and species of the subfamily Pselaphinae, parentheses. but no correction was provided for pselaphine Museum National d’Histoire Naturelle, Paris, fauna of Myanmar. Raffray Collection (MNHN) S. A. Kurbatov and G. Cuccodoro revised the National Museum of Prague, Czech, Natural His- genus Pareuplectops established by Jeannel tory, Helfer Collection (NMP) (1957) belonging to the tribe Proterini (supertribe National Museum of Nature and Science, Tsuku ba, Goniaceritae). As the result, they synonymized Japan (NMNS) P. novissimus with P. tenasserimi, both of which were described by Blattný (1925) as members of 3) A checklist of the pselaphine species known the genus Epiplectus Raffray (Euplectitae, from Myanmar Trichonychini). Nomura (2004) reported two species from Supertribe EUPLECTITAE Yangon on the basis of the specimens collected Tribe Bythinoplectini by Mr. Yoshiyasu Kusakabe. Subtribe Bythinoplectina Nomura and Idris (2008) recorded twelve new distributions of pselaphine species from Yangon Neozethopsus brachyurus (Motschulsky, 1851: on the basis of Kusakabe’s Collection. 500) Tenasserim (“Ind. or.”, as Euplectus). Nomura (2010) described Octomicrus kusaka- Neozethopsus nonplusulta (Blattný, 1925: 183) bei from Yangon, which was first species of the Tenasserim (Blattný, 1925). tribe Dimerini from Myanmar. Rhinozethus helferi (Blattný, 1925: 184) Tenas- Recently, Yin & Li (2015) described a new serim (Blattný, 1925) (Fig. 1A–B). species, Smetanabatrus ghecu from Carin Asciuii Zethopsus dohrni Raffray, 1883: 248 Birmanie Ghecù, Shan State, which was also recorded (Fig. 1C–D). from Doi Suthep, Thailand by Yin & Cuccodoro Zethopsus infuscatus Motschulsky, 1851: 500 (2018). Tenasserim (“Ind. or.”, as Euplectus). After the works shown above, more than one Zethopsus inperfectus Blattný, 1925: 184 Tenas- hundred species of the subfamily Pselaphinae serim (Blattný, 1925). have been known from Myanmar. In the present Zethopsus opacus Schaufuss, 1887: 12 Tenas- study, correct scientific names of these species, serim (Blattný, 1925); Bangkok, Thailand references and the recorded localities are herein (Fig. 1E–F). (＝Zethopsus westwoodi Schaufuss, listed up. 1882: 113 synonymised by Coulon, 1989). Pachyzethopsus blattnyi Coulon, 1989: 194 Tenasserim. 2) Abbreviations of depository The type and identified specimens shown in Fig. 1. Type and identified specimens of pselaphine species from Myanmar 1/4. A. syntype (NMP) of Zethopsus helferi Blattný, 1925 (＝Rhinozethus helferi (Blattný, 1925)); B. ditto, type and identification labels; C. Zethopsus dohrni Raffray, 1883, syntype (MNHN); D. ditto, type labels; E. Zethopsus opacus Schaufuss, 1887 from Siam, identified by A. Raffray (MNHN); F. ditto, identification labels; G. Octomicrus kusakabei Nomura, 2010, holotype (NMNS); H. ditto, type labels; I. Euplectus kali Blattný, 1925, syntype (NMP); J. ditto, type labels; K. Microplectus exiguus Raffray, 1898 from Singapore, syntype (MNHN); L. ditto, type labels; M. Chaetorhopalus pallipes Blattný, 1925, syntype (NMP); N. ditto, type labels; O. Euplectodina hipposideros (Schaufuss, 1877), identified by Raffray? (MNHN); P. Philotrimium abdominale (Motschulsky, 1851), identi- fied by Blattný (NMP). 132 Shûhei Nomura and Mu Mu Aung Pselaphine Species from Myanmar 1 133 Tribe Dimerini 500) Tenasserim (“Ind. or.”, as Euplectus). Octomicrus (Octomicrus) kusakabei Nomura, Subtribe Trimiina 2010: 235 Yangon (Fig. 1G–H). Philotrimium abdominale (Motschulsky, 1851: “ ” Tribe Euplectini 496) Tenasserim ( Ind. or. , as Trimium) (Fig. 1P). Euplectus caligula Blattný, 1925: 188 Tenas- serim. Supertribe Batrisitae Euplectus elongatus Motschulsky, 1851: 501 Tribe Batrisini “ ” Tenasserim ( Ind. or. ). Subtribe Batrisina Euplectus kali Blattný, 1925: 189 Tenasserim (Fig. 1I –J). Arthromelus trilunatus (Raffray, 1896: 245) Bir- Euplectus pulcher Motschulsky, 1851: 499 manie (as Batrisocenus) (Fig. 2A–B). Tenasserim (“Ind. or.”). Batriplica plicata (Motschulsky, 1851: 489) Euplectus uniformis Blattný, 1925: 188 Tenas- Tenasserim (“Ind. or.”, as Batrisus) (Fig. serim. 2C–D). (＝Batriplica dohrni Schaufuss, 1887: 137 Birmanie) (Fig. 2E–F). Tribe Trichonychini Batriplica fossulata (Motschulsky, 1851: 489) “ ” Subtribe Bibloplectina Tenasserim ( Ind. or. , as Batrisus). Batrisocenus anticus Blattný, 1925: 197 Tenas- Microplectus exiguus Raffray, 1898: 264 Tenas- serim. serim (Blattný, 1925); Tenasserim (Fig. Batrisocenus anthicus (Motschulsky, 1851: 487) 1K–L); Singapore. Tenasserim (“Ind. or.”, as Batrisus). Batrisocenus aubei Blattný, 1925: 197 Tenas- Subtribe Bibloporina serim. Batrisocenus birmanus (Schaufuss, 1882: 395) Chaetorhopalus pallipes Blattný, 1925: 186 Birmanie (Fig. 2G–H). Tenasserim (Fig. 1M–N). Batrisocenus franci Blattný, 1925: 200 Tenas- serim. Subtribe Panaphantina Batrisocenus lecontei (Motschulsky, 1851: 486) Tenasserim (“Ind. or.”, as Batrisus). Euplectodina hipposideros (Schaufuss, 1877: 12) Batrisocenus macrocerus (Motschulsky, 1851: Tenasserim (Blattný, 1925), Yangon (Nomura, 487) Tenasserim (“Ind. or.”, as Batrisus). 2004); Bangkok, Thailand, Vietnam, Java, Batrisocenus mariae Blattný, 1925: 201 Tenas- Sumatra (Fig. 1O). serim. Euplectodina breviusculus (Motschulsky, 1851: Batrisocenus nitidulus (Motschulsky, 1851: 489) Fig. 2. Type and identified specimens of pselaphine species from Myanmar 2/4. A. Arthromelus trilunatus (Raf- fray, 1896), syntype (MNHN); B. ditto, type labels; C. Batriplica plicata (Motschulsky) lectotype (?) desig- nated by I. Löbl (MNHN); D. ditto labels; E. Batriplica dohrni (Schaufuss, 1887) identified by A. Raffray, as the lectotype of Batriplica plicata (Motschulsky, 1851) designated by I. Löbl (MNHN); F. ditto, labels; G. Batrisocenus birmanus (Schaufuss, 1882), syntype (MNHN); H. ditto, type labels; I. Batrisocenus nitidulus (Motschulsky, 1851), identified specimen by Blattný (NMP); J. Batrisocenus reichenbachi (Motschulsky, 1851), identified specimen by Blattný (NMP); K. Batrisocenus tenasserimi Blattný, 1925, syntype (NMP); L. ditto, type labels; M. Batristerus faustus (Raffray, 1896), syntype (MNHN); N. ditto, type labels; O. Cratna venusta Blattný, 1925, syntype (NMP); P. ditto, type labels. 134 Shûhei Nomura and Mu Mu Aung Pselaphine Species from Myanmar 1 135 Tenasserim (“Ind. or.”, as Batrisus) (Fig. 2I). Supertribe Goniaceritae Batrisocenus quaestus (Schaufuss, 1887: 139) Tribe Arnyllini Birmanie. Batrisocenus reichenbachii
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